taxonID	type	description	language	source
03C7F607FFFDFFC5A75436CDEBA066B1.taxon	materials_examined	TYPE MATERIAL. Holotype ♀: [NAMIBIA]: S. W. Afr [ica], Erongo Mt., Farm Ameib, 21.45 S – 15.39 E, 23.2.1975; E-Y: 687, from under stones, leg. Endrödy & Schulze. – (TMSA).	en	Deschodt, Christian M., Davis, Adrian L. V. (2017): Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae). Annals of the Ditsong National Museum of Natural History 7 (7): 109-114, DOI: http://doi.org/10.5281/zenodo.5562912
03C7F607FFFDFFC5A75436CDEBA066B1.taxon	etymology	ETYMOLOGY. This species is named after its collecting locality.	en	Deschodt, Christian M., Davis, Adrian L. V. (2017): Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae). Annals of the Ditsong National Museum of Natural History 7 (7): 109-114, DOI: http://doi.org/10.5281/zenodo.5562912
03C7F607FFFDFFC5A75436CDEBA066B1.taxon	description	DESCRIPTION. Dark brown, convex. Head: two short and small slightly upturned clypeal teeth and one much smaller tooth medially on lower margin of clypeus. Clypeal teeth shorter than half the distance between them. Pronotum: punctate with punctures almost round medially becoming ovoid posteriorly and laterally, surface shagreened. Short, golden brown setae present at outside lateral margin. Protibia: surface shagreened, outside margin anteriorly tridentate in last one-third and smooth in first two-thirds. Median dorsal ridge present, almost parallel with outside margin. Golden brown setae present, longer and more numerous terminally. Ventral surface more or less flat, punctate. Elytra: convex. Surface shagreened, punctures on elytral interval more or less in single line, punctures on other intervals random. Punctures and striae well defined. Sterna: prosternum with clear, round punctures placed in close proximity. Mesometasternal suture distinct and straight, punctures on mesosternum larger and further apart than on metasternum. Metasternum with faint and shallow anterior pointing curve between coxae. Pygidium: surface shagreened and punctate, puncturessmalland furtherapart thantheirdiameter.	en	Deschodt, Christian M., Davis, Adrian L. V. (2017): Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae). Annals of the Ditsong National Museum of Natural History 7 (7): 109-114, DOI: http://doi.org/10.5281/zenodo.5562912
03C7F607FFFDFFC5A75436CDEBA066B1.taxon	discussion	DISCUSSION. This new species can be separated from all other species in the genus by being slightly larger and the edge between the clypeal teeth being curved and not straight as in the other species.	en	Deschodt, Christian M., Davis, Adrian L. V. (2017): Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae). Annals of the Ditsong National Museum of Natural History 7 (7): 109-114, DOI: http://doi.org/10.5281/zenodo.5562912
03C7F607FFFDFFC5A75432B1EEC86034.taxon	description	Gender of new name: masculine. DESCRIPTION. Small (7.1 – 9.7 mm long × 4.5 – 5.8 mm wide) dark brown, rounded and convex beetles. Head: with two small and short, slightly upturned clypeal teeth; one much smaller tooth medially on lower margin of clypeus. Pronotum: dark brown, convex with ovoid to round punctures. Protibia: tridentate in last one-third of outside margin, more or less smooth in first two-thirds. Elytra: strongly convex, dark brown. Sterna: mesometasternal suture straight. Scutellum not visible. Pygidium: shagreened with small punctures.	en	Deschodt, Christian M., Davis, Adrian L. V. (2017): Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae). Annals of the Ditsong National Museum of Natural History 7 (7): 109-114, DOI: http://doi.org/10.5281/zenodo.5562912
03C7F607FFFDFFC5A75432B1EEC86034.taxon	discussion	DISCUSSION. Morphologically this new genus can easily be separated from Versicorpus by lacking serrations on the outside margins of the front tibiae and by being smaller (7.1 – 9.7 mm long) and more convex than the larger (10.1 mm) and more elongate Versicorpus. The clypeus of Namakwanus has much more pronounced clypeal teeth than Namaphilus. The known species in the genus thus currently comprise Namaphilus davisi (Deschodt & Scholtz, 2007) comb. nov., Namaphilus endroedyi (Deschodt, Davis & Scholtz, 2011) comb. nov. and Namaphilus ameibensis spec. nov.	en	Deschodt, Christian M., Davis, Adrian L. V. (2017): Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae). Annals of the Ditsong National Museum of Natural History 7 (7): 109-114, DOI: http://doi.org/10.5281/zenodo.5562912
03C7F607FFFDFFC7A58D3157EEB061AD.taxon	materials_examined	SPECIMEN EXAMINED. Holotype ♂: [NAMIBIA]: Gobabeb, Kuiseb River, Central Namib, S. W. Afr., 11. v. 1959, H. Dick Brown. – (TMSA).	en	Deschodt, Christian M., Davis, Adrian L. V. (2017): Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae). Annals of the Ditsong National Museum of Natural History 7 (7): 109-114, DOI: http://doi.org/10.5281/zenodo.5562912
03C7F607FFFDFFC7A58D3157EEB061AD.taxon	etymology	ETYMOLOGY. This species is named in honour of Professor Clarke H. Scholtz, for his extensive and lifelong contribution to entomology.	en	Deschodt, Christian M., Davis, Adrian L. V. (2017): Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae). Annals of the Ditsong National Museum of Natural History 7 (7): 109-114, DOI: http://doi.org/10.5281/zenodo.5562912
03C7F607FFFDFFC7A58D3157EEB061AD.taxon	description	DESCRIPTION. Dark brown, convex. Head: two long, distinct, slightly upturned clypeal teeth and one much smaller tooth medially on lower margin of clypeus. Slight red tinge laterally. Pronotum: punctate with round punctures of variable size, surface shagreened. Laterally clearly flattenedwith ridge in median one-third justinside of lateral margin. Ridge parallel with outside margin anteriorly and curving inward posteriorly. Lateral sides with slight red tinge. Protibia: surface shagreened, outside margin anteriorly tridentate in last one-third and slightly sinuous to dentate in first two-thirds. Outside two denticles larger than median denticle. Median dorsal ridge present, angulate at posterior denticle of outside margin. Golden brown setae present, longer terminally. Ventral surface withmedian ridge. Elytra: convex. Surface shagreened, punctures on second to fourth intervals more or less in straight double lines, punctures on all other intervals forming more or less straight single lines. Punctures and striae not well defined. Sterna: prosternum with clear round and adjacent punctures. Mesometasternal suture distinct and very slightly curved, punctures on mesosternum larger and further apart than on metasternum, surface shagreened. Pygidium: surface shagreened and punctate, puncturessmalland further apartthantheirdiameter.	en	Deschodt, Christian M., Davis, Adrian L. V. (2017): Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae). Annals of the Ditsong National Museum of Natural History 7 (7): 109-114, DOI: http://doi.org/10.5281/zenodo.5562912
03C7F607FFFDFFC7A58D3157EEB061AD.taxon	discussion	DISCUSSION. This new species can be separated from N. irishi by the tips of the clypeal horns being slightly further apart, more upturned and slightly pointing inward at the apex.	en	Deschodt, Christian M., Davis, Adrian L. V. (2017): Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae). Annals of the Ditsong National Museum of Natural History 7 (7): 109-114, DOI: http://doi.org/10.5281/zenodo.5562912
03C7F607FFFFFFC0A7AB3455EBB566F9.taxon	discussion	NOTES ON THE CURRENTLY KNOWN SPECIES IN THE BYRRHIDIUM GROUP	en	Deschodt, Christian M., Davis, Adrian L. V. (2017): Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae). Annals of the Ditsong National Museum of Natural History 7 (7): 109-114, DOI: http://doi.org/10.5281/zenodo.5562912
03C7F607FFFFFFC0A7AB3455EBB566F9.taxon	discussion	The Byrrhidium group is confined to the arid west of Namibia and South Africa (Fig. 2). It currently consists of six genera and 14 described species. These species (Deschodt et al., 2007, 2011, 2016) may mostly be associated with dung middens of rock hyraxes (Procavia capensis (Pallas, 1766 )). As there are gaps in the known range of this group relative to that of rock hyraxes (Fig. 2), new related dung beetle species and genera may await discovery.	en	Deschodt, Christian M., Davis, Adrian L. V. (2017): Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae). Annals of the Ditsong National Museum of Natural History 7 (7): 109-114, DOI: http://doi.org/10.5281/zenodo.5562912
03C7F607FFF8FFC0A59330F3EC4C6788.taxon	materials_examined	Widespread in Namaqualand from the environs of Nuwerus [31.149 ° S 18.358 ° E] in the south to Steinkopf [29.266 ° S 17.734 ° E] in the north. More or less confined to the Namaqualand Klipkoppe Shrubland vegetation unit as defined by Mucina and Rutherford (2006).	en	Deschodt, Christian M., Davis, Adrian L. V. (2017): Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae). Annals of the Ditsong National Museum of Natural History 7 (7): 109-114, DOI: http://doi.org/10.5281/zenodo.5562912
03C7F607FFF8FFC0A59330F3EC4C6788.taxon	materials_examined	Collected some distance from rock hyrax middens among Namaqualand heuweltjies (Mima-like earth mounds, Lovegrove and Siegfried, 1986), but probably also associated with rock hyraxes.	en	Deschodt, Christian M., Davis, Adrian L. V. (2017): Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae). Annals of the Ditsong National Museum of Natural History 7 (7): 109-114, DOI: http://doi.org/10.5281/zenodo.5562912
03C7F607FFF8FFC0A59331A5ECF06070.taxon	materials_examined	Collected in the Richtersveld National Park [28.113 ° S 16.934 ° E] in association with rock hyraxes.	en	Deschodt, Christian M., Davis, Adrian L. V. (2017): Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae). Annals of the Ditsong National Museum of Natural History 7 (7): 109-114, DOI: http://doi.org/10.5281/zenodo.5562912
03C7F607FFF8FFC0A593367DECC860FF.taxon	materials_examined	Confined to the lower Boom River [28.013 ° S 17.063 ° E] in southern Namibia in close association with rock hyrax dung middens.	en	Deschodt, Christian M., Davis, Adrian L. V. (2017): Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae). Annals of the Ditsong National Museum of Natural History 7 (7): 109-114, DOI: http://doi.org/10.5281/zenodo.5562912
03C7F607FFF8FFC0A59336E8ECC8614A.taxon	materials_examined	Confined to the Witputs area [27.573 ° S 17.147 ° E] in southern Namibia in close association with rock hyrax dung middens.	en	Deschodt, Christian M., Davis, Adrian L. V. (2017): Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae). Annals of the Ditsong National Museum of Natural History 7 (7): 109-114, DOI: http://doi.org/10.5281/zenodo.5562912
03C7F607FFF8FFC0A5933764EC866186.taxon	materials_examined	Only recorded from the Tatasberg Mountains [28.323 ° S 17.241 ° E], Richtersveld National Park, South Africa, in close association with rock hyrax dung middens. Asingle female was collected live in the shade of boulders six months after the previous rains (C. M. D. pers. obs.).	en	Deschodt, Christian M., Davis, Adrian L. V. (2017): Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae). Annals of the Ditsong National Museum of Natural History 7 (7): 109-114, DOI: http://doi.org/10.5281/zenodo.5562912
03C7F607FFF8FFC0A59337A3ECB1626E.taxon	materials_examined	Only recordednear Ai-Ais [27.920 ° S 17.489 ° E] in the Fish River Canyon, Namibia.	en	Deschodt, Christian M., Davis, Adrian L. V. (2017): Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae). Annals of the Ditsong National Museum of Natural History 7 (7): 109-114, DOI: http://doi.org/10.5281/zenodo.5562912
03C7F607FFF8FFC0A593347BEC5862FD.taxon	materials_examined	Known from a single male specimen collected at Zebrafontein [27.750 ° S 16.883 ° E], about 30 km NNE of Rosh Pinah.	en	Deschodt, Christian M., Davis, Adrian L. V. (2017): Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae). Annals of the Ditsong National Museum of Natural History 7 (7): 109-114, DOI: http://doi.org/10.5281/zenodo.5562912
03C7F607FFF9FFC1A7BB32F9EEF1651E.taxon	materials_examined	Known only from the four specimens in the type series. Not collected since its description in 1987. Most probably occurs in the mountains around Windhoek [c. 22.571 ° S 17.086 ° E].	en	Deschodt, Christian M., Davis, Adrian L. V. (2017): Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae). Annals of the Ditsong National Museum of Natural History 7 (7): 109-114, DOI: http://doi.org/10.5281/zenodo.5562912
03C7F607FFF9FFC1A7BB330BEEAA6590.taxon	materials_examined	Known only from a single specimen collected in 1959, most probably from the Kuiseb River Canyon near Barrowberg Mountain [23.658 ° S 15.477 ° E] close to Gobabeb in the central Namib Desert.	en	Deschodt, Christian M., Davis, Adrian L. V. (2017): Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae). Annals of the Ditsong National Museum of Natural History 7 (7): 109-114, DOI: http://doi.org/10.5281/zenodo.5562912
03C7F607FFF9FFC1A7BB309AED346779.taxon	materials_examined	Known only from a single female specimen collected under a rock in the Erongo Mountains, Namibia.	en	Deschodt, Christian M., Davis, Adrian L. V. (2017): Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae). Annals of the Ditsong National Museum of Natural History 7 (7): 109-114, DOI: http://doi.org/10.5281/zenodo.5562912
03C7F607FFF9FFC1A7BB339DEEAC6678.taxon	materials_examined	Collected after rain in the rocky areas around Hardap Dam in Namibia.	en	Deschodt, Christian M., Davis, Adrian L. V. (2017): Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae). Annals of the Ditsong National Museum of Natural History 7 (7): 109-114, DOI: http://doi.org/10.5281/zenodo.5562912
03C7F607FFF9FFC1A7BB3075EEBE6691.taxon	materials_examined	Collected live in the Naukluft Mountains, Namibia, at least six months after the previous rain (C. M. D. pers. obs.). The holotype is currently curated by the South African National Collection of Insects [SANC] in Pretoria.	en	Deschodt, Christian M., Davis, Adrian L. V. (2017): Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae). Annals of the Ditsong National Museum of Natural History 7 (7): 109-114, DOI: http://doi.org/10.5281/zenodo.5562912
03C7F607FFF9FFC1A7BB3172EE8A67C4.taxon	materials_examined	Known only from a single female specimen collected in 1975 from under a rock in the Erongo Mountains, Namibia.	en	Deschodt, Christian M., Davis, Adrian L. V. (2017): Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae). Annals of the Ditsong National Museum of Natural History 7 (7): 109-114, DOI: http://doi.org/10.5281/zenodo.5562912
03C7F607FFF9FFC1A7BB31E1EFF16076.taxon	materials_examined	Known from a single male specimen collected from Bullsport Farm [24.133 ° S 16.366 ° E] in the Remhoogte Mountains to the northeast of the Tsondab River Canyon, Namibia.	en	Deschodt, Christian M., Davis, Adrian L. V. (2017): Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae). Annals of the Ditsong National Museum of Natural History 7 (7): 109-114, DOI: http://doi.org/10.5281/zenodo.5562912
