taxonID	type	description	language	source
8208B456FFC05818FCA0AF6424F285BE.taxon	type_taxon	Type species. Dakotaeschnidium spedeni sp. nov.	en	NEL, ANDRÉ (2021): Maastrichtian representatives of the dragonfly family Aeschnidiidae question the entomofaunal turnover of the early Late Cretaceous. Palaeoentomology 4 (3): 209-212, DOI: 10.11646/palaeoentomology.4.3.5, URL: http://dx.doi.org/10.11646/palaeoentomology.4.3.5
8208B456FFC05818FCA0AF6424F285BE.taxon	etymology	Etymology. Named after ‘ Dakota’, name of a native people living in North America, and Aeschnidium. Gender masculine.	en	NEL, ANDRÉ (2021): Maastrichtian representatives of the dragonfly family Aeschnidiidae question the entomofaunal turnover of the early Late Cretaceous. Palaeoentomology 4 (3): 209-212, DOI: 10.11646/palaeoentomology.4.3.5, URL: http://dx.doi.org/10.11646/palaeoentomology.4.3.5
8208B456FFC05818FCA0AF6424F285BE.taxon	diagnosis	Diagnosis. Hind wing characters only. Basal and distal subdiscoidal spaces transverse and very broad, posteriorly opened, without any angle between AAspl and AA 1 a veins; vein AA 1 a poorly defined; two rows of cells in hypertriangle; two in submedian space, and two-three in basal part of area between RP and MA; no infrasubdiscoidal space basal to basal subdiscoidal space.	en	NEL, ANDRÉ (2021): Maastrichtian representatives of the dragonfly family Aeschnidiidae question the entomofaunal turnover of the early Late Cretaceous. Palaeoentomology 4 (3): 209-212, DOI: 10.11646/palaeoentomology.4.3.5, URL: http://dx.doi.org/10.11646/palaeoentomology.4.3.5
8208B456FFC1581BFF02AA34266F809D.taxon	description	(Fig. 1) urn: lsid: zoobank. org: pub: C 625 EE 39 - 050 C- 49 CC- 9182 - 220 F 212 BCFB 4	en	NEL, ANDRÉ (2021): Maastrichtian representatives of the dragonfly family Aeschnidiidae question the entomofaunal turnover of the early Late Cretaceous. Palaeoentomology 4 (3): 209-212, DOI: 10.11646/palaeoentomology.4.3.5, URL: http://dx.doi.org/10.11646/palaeoentomology.4.3.5
8208B456FFC1581BFF02AA34266F809D.taxon	materials_examined	Material. HolotypeYPM IP 028154 (collector Ian G. Speden, 1962, IP number 28154; lot count 1; part / counterpart). Invertebrate Paleontology, Yale Peabody Museum, New Haven, USA.	en	NEL, ANDRÉ (2021): Maastrichtian representatives of the dragonfly family Aeschnidiidae question the entomofaunal turnover of the early Late Cretaceous. Palaeoentomology 4 (3): 209-212, DOI: 10.11646/palaeoentomology.4.3.5, URL: http://dx.doi.org/10.11646/palaeoentomology.4.3.5
8208B456FFC1581BFF02AA34266F809D.taxon	etymology	Etymology. Named after Ian G. Speden who collected the type specimen.	en	NEL, ANDRÉ (2021): Maastrichtian representatives of the dragonfly family Aeschnidiidae question the entomofaunal turnover of the early Late Cretaceous. Palaeoentomology 4 (3): 209-212, DOI: 10.11646/palaeoentomology.4.3.5, URL: http://dx.doi.org/10.11646/palaeoentomology.4.3.5
8208B456FFC1581BFF02AA34266F809D.taxon	diagnosis	Diagnosis. As for the genus. Wing membrane darkened.	en	NEL, ANDRÉ (2021): Maastrichtian representatives of the dragonfly family Aeschnidiidae question the entomofaunal turnover of the early Late Cretaceous. Palaeoentomology 4 (3): 209-212, DOI: 10.11646/palaeoentomology.4.3.5, URL: http://dx.doi.org/10.11646/palaeoentomology.4.3.5
8208B456FFC1581BFF02AA34266F809D.taxon	description	Description. A fragment of hind wing showing the discoidal triangle, hypertriangle, a fragment of the postdiscoidal area, the submedian space and basal part of anal area, 11.7 mm long, 21.0 mm wide; whole membrane very dark with yellow veins; part of wing basal to hypertriangle missing; hypertriangle subdivided into ca. 40 small irregular cells, 1.9 mm wide, 11.6 mm long; discoidal triangle clearly transverse, subdivided into more than 27 irregular cells, with vein MAb more than 7.0 mm long, curved, vein separating discoidal triangle from hypertriangle 3.1 mm long, and vein MP + CuA 6.6 mm long; postdiscoidal area very broad at base, with more than 13 rows of cells between MAa and MP; anal area very broad, 16.2 mm wide; only three posterior branches of AA in anal area; no infrasubdiscoidal space basal to basal subdiscoidal space; basal subdiscoidal space transverse, posteriorly opened because vein AA 1 a poorly defined, distal subdiscoidal space transverse, with a shape of acute triangle, very broad and long, subdivided into ca. 70 irregular cells, no angle between AAspl and AA 1 a veins; PsA vein clearly visible vanishing one cell basal to antero-basal angle of discoidal triangle; submedian space subdivided into more than 11 irregular cells; area between RP and MA subdivided into numerous cells disposed in three rows.	en	NEL, ANDRÉ (2021): Maastrichtian representatives of the dragonfly family Aeschnidiidae question the entomofaunal turnover of the early Late Cretaceous. Palaeoentomology 4 (3): 209-212, DOI: 10.11646/palaeoentomology.4.3.5, URL: http://dx.doi.org/10.11646/palaeoentomology.4.3.5
8208B456FFC1581BFF02AA34266F809D.taxon	materials_examined	Type locality and horizon. Fox Hills Formation, Trail City Member. USA; South Dakota; Dewey County; float, conc. 5, face on SW side of SSE-trending spur, 1.25 miles W of E end of Long’s Ridge, Long’s Ridge, and 6 miles W of Whitehorse. Maastrichtian, Late Cretaceous.	en	NEL, ANDRÉ (2021): Maastrichtian representatives of the dragonfly family Aeschnidiidae question the entomofaunal turnover of the early Late Cretaceous. Palaeoentomology 4 (3): 209-212, DOI: 10.11646/palaeoentomology.4.3.5, URL: http://dx.doi.org/10.11646/palaeoentomology.4.3.5
8208B456FFC1581BFF02AA34266F809D.taxon	discussion	Remarks. The very broad anal of this fossil is typical of that of a hind wing of an Aeschnidiidae, with a basal subdiscoidal space and a distal subdiscoidal space. The transverse discoidal triangle is also only found in this family (Fleck & Nel, 2003). The particular shape of the distal subdiscoidal space of YPM IP 028154 is also found in Gigantoaeschnidium ibericus Nel & Martinez-Delclòs, 1993, but the basal subdiscoidal space of Gigantoaeschnidium is much smaller than that of YPM IP 028154, and it has an angle between AAspl and AA 1 a veins (Fleck & Nel, 1993: fig. 63 a). Angloaeschnidium toyei Fleck & Nel, 1993 and Cooperaeschnidium durandi Fleck & Nel, 1993 have their distal subdiscoidal spaces similar to those of YPM IP 028154 but their distal subdiscoidal spaces are much smaller than that of YPM IP 028154. They also have only one row of cells in their hypertriangles. Even if Santanoptera gabbotti Martill & Nel, 1996 (Aptian, Crato Formation, Brazil) is based on a forewing, its distal subdiscoidal space is similar to that of YPM IP 028154, and it has no angle between AAspl and AA 1 a veins (Fleck & Nel, 1993: fig. 89). They also share the presence of two rows of cells in the hypertriangle, two in the submedian space, and two-three in the basal part of area between RP and MA. Nevertheless, Santanoptera has a smaller basal subdiscoidal space that is posteriorly closed by a well-defined vein AA 1 a. Despite the fact that the specimen YPM IP 028154 is reduced to the basal third of a hind wing, it has a series of characters allowing to consider that it corresponds to a new genus and species, possibly related to the South American genus Santanoptera, as they share the very particular absence of an angle between AAspl and AA 1 a veins, and presence of several rows of cells in hypertriangle, submedian space, etc.	en	NEL, ANDRÉ (2021): Maastrichtian representatives of the dragonfly family Aeschnidiidae question the entomofaunal turnover of the early Late Cretaceous. Palaeoentomology 4 (3): 209-212, DOI: 10.11646/palaeoentomology.4.3.5, URL: http://dx.doi.org/10.11646/palaeoentomology.4.3.5
