taxonID	type	description	language	source
5C5187FFFFFFFF8ABD6BFF30B7CBFCEA.taxon	type_taxon	Type species. Empherium rasnitsyni sp. nov.	en	HAKIM, MARINA, HUANG, DI-YING, AZAR, DANY (2021): New fossil psocids from Cretaceous Siberian ambers (Psocodea: Trogiomorpha: Atropetae). Palaeoentomology 4 (2): 186-198, DOI: 10.11646/palaeoentomology.4.2.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.2.8
5C5187FFFFFFFF8ABD6BFF30B7CBFCEA.taxon	etymology	Etymology. Named after a derivative form of the type genus Empheria. Gender: masculine.	en	HAKIM, MARINA, HUANG, DI-YING, AZAR, DANY (2021): New fossil psocids from Cretaceous Siberian ambers (Psocodea: Trogiomorpha: Atropetae). Palaeoentomology 4 (2): 186-198, DOI: 10.11646/palaeoentomology.4.2.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.2.8
5C5187FFFFFFFF8ABD6BFF30B7CBFCEA.taxon	diagnosis	Diagnosis. Macropterous. Head with dorsal suture; ocelli present; antennae 25 - segmented (23 flagellomeres), last segment being diminutive; sensillum present on middle of second palpomere. Forewing densely setose, setae arranged on either sides of veins and dispersed in some areas of membrane (e. g., pterostigma cell and anal region); branching of M 1 and M 2 basal to branching of R 2 + 3 and R 4 + 5, branches of M long; areola postica relatively short; nodulus absent. Hind wing with Sc free; basi-radial cell present; stems of R and M fused in a short segment before vein R 1 emerging from common stem R + M; veins Rs and M bifurcated; CuA simple. Tarsi three-segmented, basal tarsomere with two rows of four spines, pretarsal claw with pulvillus and without preapical tooth. Female subgenital plate tapering apically.	en	HAKIM, MARINA, HUANG, DI-YING, AZAR, DANY (2021): New fossil psocids from Cretaceous Siberian ambers (Psocodea: Trogiomorpha: Atropetae). Palaeoentomology 4 (2): 186-198, DOI: 10.11646/palaeoentomology.4.2.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.2.8
5C5187FFFFFFFF8FBD6BFC3DB0F1FF61.taxon	description	(Figs 2 – 5)	en	HAKIM, MARINA, HUANG, DI-YING, AZAR, DANY (2021): New fossil psocids from Cretaceous Siberian ambers (Psocodea: Trogiomorpha: Atropetae). Palaeoentomology 4 (2): 186-198, DOI: 10.11646/palaeoentomology.4.2.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.2.8
5C5187FFFFFFFF8FBD6BFC3DB0F1FF61.taxon	materials_examined	Type material. Holotype specimen PIN 3426 / 34, female, mostly well-preserved (Fig. 2). Visible breaks in the amber, especially over the head area, creating a mirror effect during observation. No syninclusions observed.	en	HAKIM, MARINA, HUANG, DI-YING, AZAR, DANY (2021): New fossil psocids from Cretaceous Siberian ambers (Psocodea: Trogiomorpha: Atropetae). Palaeoentomology 4 (2): 186-198, DOI: 10.11646/palaeoentomology.4.2.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.2.8
5C5187FFFFFFFF8FBD6BFC3DB0F1FF61.taxon	etymology	Etymology. Named after Professor Alexandr Rasnitsyn.	en	HAKIM, MARINA, HUANG, DI-YING, AZAR, DANY (2021): New fossil psocids from Cretaceous Siberian ambers (Psocodea: Trogiomorpha: Atropetae). Palaeoentomology 4 (2): 186-198, DOI: 10.11646/palaeoentomology.4.2.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.2.8
5C5187FFFFFFFF8FBD6BFC3DB0F1FF61.taxon	diagnosis	Diagnosis. As for genus, by monotypy. Locality and horizon. Nizhnyaya Agapa, Taimyr Peninsula, Russia, Dolgan Formation, Cretaceous (Upper Cenomanian).	en	HAKIM, MARINA, HUANG, DI-YING, AZAR, DANY (2021): New fossil psocids from Cretaceous Siberian ambers (Psocodea: Trogiomorpha: Atropetae). Palaeoentomology 4 (2): 186-198, DOI: 10.11646/palaeoentomology.4.2.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.2.8
5C5187FFFFFFFF8FBD6BFC3DB0F1FF61.taxon	description	Description. Head 0.70 mm wide (Fig. 3 A); dorsal suture visible, much accentuated. Ocelli present. Compound eyes bare, as long as vertex. Antenna with 25 antennomeres, setae thin and sparse along its length, no secondary annulations; complete antenna with 23 flagellomeres, 0.01 mm wide and 0.05 – 0.06 mm long, last segment diminutive, with two small and thin apical setae pointing forward. Maxillary palpomeres four-segmented, setose, second palpomere with strong conical sensillum at middle (Fig. 3 B). Labial palpomeres two-segmented. Lacinia not visible. Thorax 0.60 mm wide; legs with tibia bearing occasional spines along its length and two apical spurs. Tarsi three-segmented (Fig. 3 C); ratio of tarsal segments (from base to apex) in fore and mid-legs 3: 1: 1, in hindlegs 4.5: 1: 1; basal tarsomere with two rows of four spines but no ctenidiobothria (Fig. 3 D); pretarsal claw without preapical tooth, pulvillus present (long and narrow basally but rounded, sphere-like, apically). Wings macropterous (Fig. 3 E); forewing (1.80 mm long and 0.50 mm wide) with marginal setae; membrane hyaline, setae arranged in two rows around veins and sparsely dispersed in some areas of membrane (e. g., pterostigma cell and anal region); Sc present, curved back and fusing with R, additional veinlet emerging from Sc and reaching costal margin; Sc’ reaching margin at 1.09 mm; pterostigma cell closed basally, 0.30 mm long and 0.10 mm wide; R 1 reaching margin at 1.30 mm from base; Rs bifurcated, R 2 + 3 reaching margin at 1.60 mm and R 4 + 5 reaching margin at apex; short crossvein r 1 - rs present; M three-branched, veins very long, M 3 separating from common stem M at 0.76 mm, stem M 1 and M 2 very short, fork M 1 and M 2 at 0.86 mm, branches M 1, M 2 and M 3 reaching margin respectively at 1.50, 1.40 and 1.18 mm; CuA bifurcated at 0.50 mm, CuA 1 and CuA 2 veins short, reaching margin respectively at 0.80 and 0.70 mm; areola postica (AP) cell free, 0.30 mm long and 0.18 mm wide; CuP and A 1 simple; A 1 weakly visible, A 2 short. Hind wing (1.38 mm long and 0.40 mm wide) without marginal setae; membrane hyaline, not setose; Sc present, short, not reaching margin; basi-radial cell present, with basal section unclear, potentially absent or weakly visible; stems R and M fused in a short segment, 0.05 mm long; R 1 branching from common stem R + M at 0.40 mm from base, reaching margin at 0.90 mm; common stem Rs as long as branches R 2 + 3 and R 4 + 5; R 2 + 3 reaching margin at 1.20 mm and R 4 + 5 at apex; M two-branched, with M 1 reaching margin at 1.20 mm; CuA simple; CuP and A not visible. Female ovipositor with dorsal and ventral valvulae not visible; external valvulae (V 3) elongated, setose (Fig. 3 F, G); subgenital plate setose, narrowed apically, with tapered segment rounded; paraprocts with a thick posterior spine each.	en	HAKIM, MARINA, HUANG, DI-YING, AZAR, DANY (2021): New fossil psocids from Cretaceous Siberian ambers (Psocodea: Trogiomorpha: Atropetae). Palaeoentomology 4 (2): 186-198, DOI: 10.11646/palaeoentomology.4.2.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.2.8
5C5187FFFFFFFF8FBD6BFC3DB0F1FF61.taxon	discussion	Remarks. Teratology and aberrations in wing venation seem very common in this family. Within this single specimen, we observe between the pair of forewings several differences that can be attributed to teratological malformations: R 4 + 5 is bifurcated apically in one wing, the shape of Sc differs from one wing to the other, the Sc veinlet is aborted mid-way in one wing, and another aborted veinlet is observed on the pterostigma of the other (Fig. 4 B).	en	HAKIM, MARINA, HUANG, DI-YING, AZAR, DANY (2021): New fossil psocids from Cretaceous Siberian ambers (Psocodea: Trogiomorpha: Atropetae). Palaeoentomology 4 (2): 186-198, DOI: 10.11646/palaeoentomology.4.2.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.2.8
5C5187FFFFFAFF8FBD6BFEF3B6EFFE4B.taxon	type_taxon	Type species. Eolepinotus pilosus Vishnyakova, 1975	en	HAKIM, MARINA, HUANG, DI-YING, AZAR, DANY (2021): New fossil psocids from Cretaceous Siberian ambers (Psocodea: Trogiomorpha: Atropetae). Palaeoentomology 4 (2): 186-198, DOI: 10.11646/palaeoentomology.4.2.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.2.8
5C5187FFFFFAFF8FBD6BFE62B74BF7B5.taxon	description	(Figs 6 – 8)	en	HAKIM, MARINA, HUANG, DI-YING, AZAR, DANY (2021): New fossil psocids from Cretaceous Siberian ambers (Psocodea: Trogiomorpha: Atropetae). Palaeoentomology 4 (2): 186-198, DOI: 10.11646/palaeoentomology.4.2.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.2.8
5C5187FFFFFAFF8FBD6BFE62B74BF7B5.taxon	materials_examined	Type material. Holotype specimen PIN 3603 / 1, female, badly preserved but with enough morphological features for clear observation and identification (Fig. 6). No syninclusion.	en	HAKIM, MARINA, HUANG, DI-YING, AZAR, DANY (2021): New fossil psocids from Cretaceous Siberian ambers (Psocodea: Trogiomorpha: Atropetae). Palaeoentomology 4 (2): 186-198, DOI: 10.11646/palaeoentomology.4.2.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.2.8
5C5187FFFFFAFF8FBD6BFE62B74BF7B5.taxon	etymology	Etymology. Named after the late Professor Vladimir Zherikhin. Locality and horizon. Timmerdyakh-Khaya, Yakutia, Russia, Timmerdyakh Formation, Cretaceous (Upper Cenomanian – Turonian).	en	HAKIM, MARINA, HUANG, DI-YING, AZAR, DANY (2021): New fossil psocids from Cretaceous Siberian ambers (Psocodea: Trogiomorpha: Atropetae). Palaeoentomology 4 (2): 186-198, DOI: 10.11646/palaeoentomology.4.2.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.2.8
5C5187FFFFFAFF8FBD6BFE62B74BF7B5.taxon	diagnosis	Diagnosis. Head with dorsal suture well-defined, with anterior arms of frontal sutures; compound eyes bare; antennomeres without secondary annulations; micropterous; wings veinless and setose; pretarsal claws short, no preapical tooth, no pulvilli. Differential characters: forewings narrow and elongated (round and short in Eolepinotus pilosus); tibia bearing sparse spines along its length.	en	HAKIM, MARINA, HUANG, DI-YING, AZAR, DANY (2021): New fossil psocids from Cretaceous Siberian ambers (Psocodea: Trogiomorpha: Atropetae). Palaeoentomology 4 (2): 186-198, DOI: 10.11646/palaeoentomology.4.2.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.2.8
5C5187FFFFFAFF8FBD6BFE62B74BF7B5.taxon	description	Description. Head 0.60 mm wide; dorsal suture present, very well defined; anterior arms of frontal sutures visible (Fig. 7 A). Ocelli absent. Compound eyes bare, as long as vertex. Both antennae incomplete, with at least 15 – 16 flagellomeres, no secondary annulations. Maxillary palpomeres four-segmented (Fig. 7 B), second palpomere from base with two spurs in apical half, third palpomere with one spur apically, fourth palpomere with one thinner and shorter spur at 2 / 3 of length and a sensory field of short setae at apex. Labial palpomeres two-segmented. Lacinia not visible. Thorax badly preserved, pronotum very setose; legs with thick femur in all legs; hind tibia bearing thin spines along its length (Fig. 7 C); tarsi three-segmented; basal tarsomere with two rows of small spines; pretarsal claws short without preapical tooth and no pulvillus (Fig. 7 D); wings micropterous, veinless (Fig. 7 E); forewings setose, elongated and narrowed. Abdomen partially destroyed and partially covered by debris. Female ovipositor with dorsal and ventral valvulae not visible, either reduced or absent; external valvulae clearly visible, elongated, setose (Fig. 7 F). Subgenital plate setose, possibly weakly bilobed apically; paraprocts not visible.	en	HAKIM, MARINA, HUANG, DI-YING, AZAR, DANY (2021): New fossil psocids from Cretaceous Siberian ambers (Psocodea: Trogiomorpha: Atropetae). Palaeoentomology 4 (2): 186-198, DOI: 10.11646/palaeoentomology.4.2.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.2.8
