taxonID	type	description	language	source
038487C8FFAEFF82FDE7FAF6FA91F832.taxon	discussion	Key to species (male) of Vestiplex from Tibet, China	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFA8FF86FD81FE0EFC36FBE4.taxon	description	urn: lsid: zoobank. org: act: 788 D 3 C 24 - ECA 9 - 471 D- 8 F 77 - F 4 D 688 CD 3 C 20 Figs 1 – 9, 86	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFA8FF86FD81FE0EFC36FBE4.taxon	diagnosis	Diagnosis Nasus elongate, apex slightly expanded. Scape and pedicel of antenna yellow; flagellar segments 1 – 4 grayish yellow, other segments pale brown, except base darker. Prescutum almost wholly reddish brown with four olive-gray striped pruinescence, median stripe broadened apically and gradually narrowing to base. Posterior margin of gonocoxite with a sharp black spine. Tergite 9 separated medially, with a pair of angular arms arising from ventral surface of tergite 9, with granulated margins, apex with a small inner projection, outer surface of arms with a pair of depressions (Fig. 9). Outer gonostylus slender, spoon-like. Lower beak of inner gonostylus absent.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFA8FF86FD81FE0EFC36FBE4.taxon	etymology	Etymology The species is named after the shape of inner gonostylus, it looks like a ‘ Magatama’, a curved commashaped bead from prehistoric Japan, called ‘ 勾玉’ in China. The name is to be treated as a noun in apposition.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFA8FF86FD81FE0EFC36FBE4.taxon	materials_examined	Material examined Holotype CHINA • ♂; Tibet, Bomi, Yigong (¼ 贡); 30 ° 16 ′ 19 ″ N, 94 ° 48 ′ 28 ″ E; 2270 m a. s. l.; 5 – 7 May 2017; Qicheng Yang leg.; light trap; CAU. Paratypes CHINA • 13 ♂♂; same collection data as for holotype; CAU • 4 ♂♂; Tibet, Motuo, 80 K; 29 ° 39 ′ 29 ″ N, 95 ° 29 ′ 24 ″ E; 2020 m a. s. l.; 8 May 2017; Qicheng Yang leg.; light trap; CAU.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFA8FF86FD81FE0EFC36FBE4.taxon	description	Description Male MEASUREMENTS. Body length 16.5 – 20.0 mm, wing length 21.0 – 24.0 mm, antenna length 2.5 – 3.0 mm (n = 18). HEAD (Figs 1, 3). Mainly brownish yellow. Occipital marking brown, elongated to vertex, broadened apically and gradually narrowing to base. Rostrum and nasus grayish yellow with brown setae, nasus elongate, apex slightly expanded. Setae on head dark brown. Antenna relatively short; scape and pedicel yellow; flagellar segments 1 – 4 grayish yellow, other segments pale brown, except base darker. Proboscis and palpus brown. THORAX (Figs 1, 3). Mainly reddish brown, heavily grayish yellow pruinescence. Pronotum brown with yellow setae, middle black. Prescutum almost wholly reddish brown with four olive-gray striped pruinescence, median stripe broadened apically and gradually narrowing to base. Scutum brown with two olive-gray pruinose spots. Scutellum reddish brown with dark gray pruinescence. Mediotergite dark yellow with yellow pruinescence and olive-gray mid-longitudinally. Thoracic pleuron dark yellow with grayish yellow pruinescence. Anepisternum, katepisternum and anepimeron dark brown at antero-upper corners. Anatergite and katatergite yellow. Parascutellum yellow. Meron dark yellow. Setae on thorax yellow. Legs with coxae and femora yellow, femora with brownish black apexes; tibiae brownish yellow with black tips. Tarsi brownish black. Setae on legs dark brown. Wing variegated brown with numerous smaller pale areas; pterostigma pale black; petiole of cell m 1 distinctly shorter than discal cell (Fig. 2). Halter with stem dark yellow, knob black. ABDOMEN (Figs 1, 3). Mainly dark yellow. Abdominal tergites each with three black stripes, middle stripe stout; lateral stripes relatively narrow. Abdominal segments 5 – 9 black. Setae on abdomen yellow. HYPOPYGIUM (Figs 4 – 9). Posterior margin of gonocoxite with a sharp black spine. Tergite 9 separated medially, with a pair of angular arms arising from ventral surface of tergite 9, with granulated margins, apex with a small inner projection, outer surface arms with a pair of depressions. Sternite 9 with a median prominence. Apex of adminiculum slim. Outer gonostylus slender, spoon-like, apex circular (Fig. 5). Inner gonostylus with big concavity; inner margin black sclerotized; lower beak absent; reversed side with big concavity (Figs 6 – 7). Female Unknown or unrecognized.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFA8FF86FD81FE0EFC36FBE4.taxon	distribution	Distribution China (Tibet).	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFA8FF86FD81FE0EFC36FBE4.taxon	discussion	Remarks The new species is similar to T. (V.) yunnanensis Alexander, 1942 from Yunnan, China, but the latter differs in the following characters: posterior margin of gonocoxite with a slightly twisted spine; outer gonostylus long and slender; lower beak of inner gonostylus sharp.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFABFF88FDF8FB01FA95F8B5.taxon	description	urn: lsid: zoobank. org: act: 635 FCCE 1 - 5615 - 49 F 6 - B 272 - 82142261 A 897 Figs 10 – 20, 86	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFABFF88FDF8FB01FA95F8B5.taxon	diagnosis	Diagnosis Scape and pedicel yellow, flagellum grayish yellow. Prescutum with four grayish yellow stripes, all stripes with brown margins. Scutellum yellow with dark brown mid-longitudinal stripe. Mediotergite yellow with brown mid-longitudinal stripe. Tergite 9 separated medially, pair of angular arms arising from ventral surface of tergite 9, arms with granulated margins. Base of posterior margin of inner gonostylus without a small protrusion; upper beak slightly twisted, lower beak curved, middle concave.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFABFF88FDF8FB01FA95F8B5.taxon	etymology	Etymology The species is named after the uncinatus of the adminiculum, the Latin adjective ‘ uncinella ’ means ‘ bearing hooks, barbed’.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFABFF88FDF8FB01FA95F8B5.taxon	materials_examined	Material examined Holotype CHINA • ♂; Tibet, Bomi, Yigong; 30 ° 16 ′ 19 ″ N, 94 ° 48 ′ 28 ″ E; 2270 m a. s. l.; 5 – 7 May 2017; Qicheng Yang leg.; sweeping net; CAU. Paratypes CHINA • 12 ♂♂; same collection data as for holotype; CAU • 9 ♂♂; Tibet, Bomi, 106 K; 29 ° 50 ′ 04 ″ N, 94 ° 44 ′ 36 ″ E; 2290 m a. s. l.; 17 May 2017; Qicheng Yang leg.; light trap; CAU.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFABFF88FDF8FB01FA95F8B5.taxon	description	Description Male MEASUREMENTS. Body length 14.0 mm, wing length 18.0 – 20.0 mm, antenna length 3.0 – 3.5 mm (n = 22). HEAD (Figs 10, 12). Mainly yellow. Occipital stripe dark gray. Rostrum and nasus yellow. Setae on head brown. Scape and pedicel yellow, flagellum grayish yellow. Proboscis and palpus brownish yellow. THORAX (Figs 10, 12). Mainly yellow, heavily pruinose. Pronotum brownish yellow with yellow setae. Prescutum with four grayish yellow stripes, all stripes with brown margins, median stripe broadened apically and gradually narrowing to base, lateral stripes long-oval. Scutum yellow with four grayish yellow spots, anterior margins of anterior two spots brown. Scutellum yellow with dark brown midlongitudinal stripe. Mediotergite yellow with brown mid-longitudinal stripe. Thoracic pleuron yellow. Parascutellum yellow. Meron yellow. Setae on thorax yellow. Legs with coxae and femora yellow, femora with dark brown apices, tibiae and tarsi dark brown. Setae on legs dark brown. Wing variegated light brown; pterostigma brown; petiole of cell m 1 distinctly shorter than discal cell (Fig. 11). Halter with stem yellow, knob pale black with pale yellow apex. ABDOMEN (Figs 10, 12). Mainly yellow. Abdominal tergites each with three brown stripes. Abdominal segments 5 – 9 dark brown to black. Setae on abdomen brownish yellow. HYPOPYGIUM (Figs 13 – 17, 20). Gonocoxite elongated, separated from sternite 9, posterior margin with an unusually strong and slender black spine, directed chiefly dorsally, slightly twisted. Tergite 9 separated medially, middle of posterior margin of lobe slightly protrude; a pair of angular black arms arising from ventral surface of tergite 9, surface of arms not smooth, with granulated margin. Sternite 9 with a median prominence. Adminiculum narrow spade-like, apex slightly furcal, ventral caudal part with a small uncinate projection. Outer gonostylus anteriorly bent, posterior margin of basal half truncated (Fig. 16). Inner gonostylus with concavity at middle, posterior margin with big semicircular bulge; base of posterior margin of inner gonostylus without a small protrusion; upper beak slightly twisted, lower beak curved, middle concave (Fig. 14). Semen pump (Figs 18 – 19) as shown in figures. Aedeagus wide, basal width about 0.5 mm, gradually narrowing towards apex, and apex not bifurcated. Female Unknown or unrecognized.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFABFF88FDF8FB01FA95F8B5.taxon	distribution	Distribution China (Tibet).	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFABFF88FDF8FB01FA95F8B5.taxon	discussion	Remarks The new species is similar to T. (V.) schizophallus (Alexander, 1973), but the latter differs in the following characters: arms of tergite 9 with only inner margin roughened; gonocoxite with an additional small spine (the one near posterior margin of basistylar suture); outer gonostylus with very abundant black setae from conspicuous punctures; apex of adminiculum profoundly divided into two long parallel spines, ventral caudal part without uncinatus. It is also similar T. (V.) deserrata Alexander, 1934 and T. (V.) factiosa Alexander, 1940, but the posterior margin of inner gonostylus in the latter has no protrusion. The posterior margin of tergite 9 of T. (V.) factiosa is smooth, its arms are large and strong.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFA7FF8FFD9BFE23FDC6FC29.taxon	description	urn: lsid: zoobank. org: act: D 91 DF 543 - 8018 - 4 FD 5 - 8 E 99 - DCB 2 AAF 3 E 8 FF Figs 21 – 29, 86	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFA7FF8FFD9BFE23FDC6FC29.taxon	diagnosis	Diagnosis Vertex brownish black, relatively protruded. Prescutum virtually brownish black, black stripes distinct, all stripes with dark black margins. Scutum virtually brownish black. Scutellum dark yellow with brownish black mid-longitudinal stripe. Mediotergite reddish dark brown, dried with pale gray pruinescence and black mid-longitudinally. Thoracic pleuron dark brown with pale gray pruinescence. Posterior margin of tergite 9 with long setae, pair of weak arms arising from ventral surface of tergite 9, slender hidden bridge between them.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFA7FF8FFD9BFE23FDC6FC29.taxon	etymology	Etymology The species is named after the type locality Nayogabu Snow Mountain.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFA7FF8FFD9BFE23FDC6FC29.taxon	materials_examined	Material examined Holotype CHINA • ♂; Tibet, Bomi, Yigong; 30 ° 16 ′ 19 ″ N, 94 ° 48 ′ 28 ″ E; 2290 m a. s. l.; 7 May 2017; Qicheng Yang leg.; light trap; CAU. Paratypes CHINA • 8 ♂♂; same collection data as for holotype; CAU.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFA7FF8FFD9BFE23FDC6FC29.taxon	description	Description Male MEASUREMENTS. Body length 16.5 – 18.5 mm, wing length 20 – 22.0 mm, antenna length 4.5 mm (n = 9). HEAD (Figs 21, 23). Mainly brownish yellow. Occipital mark dark brown, vertex brownish black, relatively protruded. Rostrum and nasus dark yellow, nasus relatively long. Setae on head dark brown. Scape dark yellow, pedicel pale yellow, flagellum brownish black except first flagellar segment yellowish brown. Proboscis and palpus yellowish gray. THORAX (Figs 21 – 23). Mainly dark brown, heavily pale gray pruinescence. Pronotum brown with yellow setae. Prescutum virtually brownish black, black stripes distinct, all stripes with dark black margin. Scutum virtually brownish black. Scutellum dark yellow with brownish black mid-longitudinal stripe. Mediotergite reddish dark brown, dried with pale gray pruinescence and black mid-longitudinally. Thoracic pleuron dark brown with pale gray pruinescence. Parascutellum dark brown. Meron brown. Setae on thorax pale yellow. Legs with coxae and femora yellow, femora with brownish black apexes; tibiae and tarsi dark brown. Setae on legs dark brown. Wing variegated brown; pterostigma brown; petiole of cell m distinctly shorter than discal cell (Fig. 24). Halter with stem yellow, knob dark brown. ABDOMEN (Figs 21, 23). Mainly yellow. Abdominal tergites each with three stripes, middle stripe of tergites 1 – 2 brownish black, wide, others brown; bilateral stripes black. Abdominal segments 6 – 9 dark black. Setae on abdomen yellow. HYPOPYGIUM (Figs 25 – 29). Gonocoxite hardly elongated, unseparated from sternite 9. Tergite 9 longitudinally divided into two parts, posterior margin with long setae, a pair of weak arms arising from ventral surface of tergite 9, a slender hidden bridge between them (Fig. 28). Appendages of sternite 9 slender (Fig. 29). Apex and middle of outer gonostylus slightly wide (Fig. 26). Inner gonostylus with concavity at middle, posterior part with many bristles, lower margin with bristled raise; base of lower beak inward-rolled, enwrapped black lower beak; upper beak yellow, lower margin black, base with a slight raise (Fig. 27). Apex of adminiculum hooked. Female Unknown or unrecognized.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFA7FF8FFD9BFE23FDC6FC29.taxon	distribution	Distribution China: Tibet.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFA7FF8FFD9BFE23FDC6FC29.taxon	discussion	Remarks The new species is similar to T. (V.) zayulensis from Chayu (= Zayu) of Tibet, but the latter differs in the following characters: prescutum and scutum yellowish brown; thoracic pleuron brownish yellow; all sterno-pleurite with dark brown margin. The outer gonostylus is different from that of T. (V.) nayogabuensis sp. nov.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFA2FF90FDB3FC45FEB1FB03.taxon	description	Figs 30 – 38	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFA2FF90FDB3FC45FEB1FB03.taxon	diagnosis	Diagnosis Occipital stripe dark brown, middle wide, anterior part extended to vertex. All sterno-pleurites with dark brown margins. Tergite 9 longitudinally divided into two parts, posterior margin with long setae, and a bridge between them, a pair of angular arms arising from ventral surface of tergite 9. Inner gonostylus with concavity at middle, concavity with many bristles, lower margin of concavity with bristled raise; base of inner gonostylus inward-rolled, enwrapped black and swollen lower beak; upper beak relatively obtuse, slightly recurved.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFA2FF90FDB3FC45FEB1FB03.taxon	materials_examined	Material examined CHINA • 1 ♂; Tibet, Chayu, Longgucun (AE 古村); 28 ° 50 ′ 44 ″ N, 97 ° 29 ′ 29 ″ E; 2640 m a. s. l.; 7 Jul. 2015; Shaolin Han leg.; CAU • 2 ♂♂; Tibet, Chayu, Shuidianqiao (水ΨΚ); 28 ° 39 ′ 60 ″ N, 97 ° 28 ′ 07 ″ E; 2640 m a. s. l.; 12 Jun. 2019; Qicheng Yang leg.; sweeping net; CAU • 1 ♂; Tibet, Chayu County; 28 ° 39 ′ 49 ″ N, 97 ° 27 ′ 52 ″ E; 2337 m a. s. l.; Jun. 2020; Hui Wang leg.; CAU.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFA2FF90FDB3FC45FEB1FB03.taxon	description	Redescription Male MEASUREMENTS. Body length 18.5 mm, wing length 20.0 mm, antenna length 4.0 mm (n = 3). HEAD (Figs 30, 32). Mainly yellow. Occipital stripe dark brown, middle wide, anterior part extended to vertex. Rostrum and nasus yellow, nasus long. Setae on head dark brown. Scape yellow, pedicel pale yellow, flagellum yellowish brown. Proboscis and palpus yellowish dark brown. THORAX (Figs 30, 32). Mainly brownish yellow, heavily gray pruinescence. Pronotum yellowish brown, middle and both side each with brown spot. Prescutum yellowish brown with four brown stripes, all stripes with dark brown margins. Scutum yellowish brown with four dark brown spots. Scutellum yellowish brown with dark brown mid-longitudinal stripe. Mediotergite brown with dark brown midlongitudinal stripe. Thoracic pleuron brownish yellow. All sterno-pleurite with dark brown margins. Setae on thorax yellow. Legs with coxae yellow, femora with dark brown tips; tibiae and tarsi dark brown. Setae on legs dark brown. Wing variegated pale brown; pterostigma brown; petiole of cell m 1 distinctly shorter than discal cell (Fig. 31). Halter with stem yellow, knob greyish black. ABDOMEN (Figs 30, 32). Mainly yellow. Abdominal tergites each with three stripes, middle stripe brown, bilateral stripes black. Abdominal segments 6 – 9 dark black. Setae on abdomen brownish yellow. HYPOPYGIUM (Figs 33 – 38). Gonocoxite slightly elongated, separated from sternite 9. Tergite 9 longitudinally divided into two parts, posterior margin with long setae, and a bridge between them, a pair of angular arms arising from ventral surface of tergite 9. Sternite 9 with a pair of very short appendages, as shown in figure, with a long seta (Fig. 35), covered by inner gonostylus and not visible in the lateral external view. Outer gonostylus relatively small, posterior margin basally slightly wide (Fig. 34). Inner gonostylus with concavity at middle, concavity with many bristles, lower margin with bristled raise; base of inner gonostylus inward-rolled, enwrapped black and swollen lower beak; upper beak relatively obtuse, slightly recurved (Fig. 36). Female Described by Alexander (1963).	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFA2FF90FDB3FC45FEB1FB03.taxon	distribution	Distribution China: Tibet.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFBDFF97FDEBFA07FBC0FDB1.taxon	description	urn: lsid: zoobank. org: act: 4 AB 06 AB 7 - 6108 - 40 AF-A 8 C 8 - CAEDF 0226964 Figs 39 – 50, 84, 86	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFBDFF97FDEBFA07FBC0FDB1.taxon	diagnosis	Diagnosis Scape and pedicel yellow, first flagellar segment dark yellow, other flagellar segments bicolor, yellowish with dark brown base. Scutellum and mediotergite dark yellow. Gonocoxite elongated, separated from sternite 9, posterior margin with an unusually strong black spine, bent inward and horn-like.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFBDFF97FDEBFA07FBC0FDB1.taxon	etymology	Etymology The species is named after the shape of gonocoxite, the Latin adjective ‘ bucera ’ means ‘ ox-horned’.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFBDFF97FDEBFA07FBC0FDB1.taxon	materials_examined	Material examined Holotype CHINA • ♂; Tibet, Bomi, Yigong; 30 ° 16 ′ 19 ″ N, 94 ° 48 ′ 28 ″ E; 2240 m a. s. l.; 7 Jun. 2017; Qicheng Yang leg.; light trap; CAU. Allotype CHINA • ♀; same collection data as for holotype; CAU. Paratypes CHINA • 9 ♂♂; same collection data as for holotype; CAU.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFBDFF97FDEBFA07FBC0FDB1.taxon	description	Description Male MEASUREMENTS. Body length 13.0 mm, wing length 15.0 mm, antenna length 4.5 mm (n = 10). HEAD (Figs 39, 41). Mainly yellow. Occipital stripe dark brown, with brown setae. Rostrum and nasus yellow, rostrum with brown setae and nasus with dark yellow setae. Scape and pedicel yellow, first flagellar segment dark yellow, other flagellar segments bicolor yellowish with dark brown base. Proboscis and palpus grayish yellow. THORAX (Figs 39, 41). Mainly yellow, heavily pruinose. Pronotum yellow with yellow setae. Prescutum with four yellowish gray stripes, median stripe broadened apically and gradually narrowing to base, lateral stripes long-oval. Scutum yellow with four yellowish gray spots. Scutellum and mediotergite dark yellow. Thoracic pleuron yellow. Parascutellum pale yellow. Meron yellow. Setae on thorax yellow. Legs with coxae and femora yellow, femora with dark brown apices and brown middle parts; tibiae and tarsi dark brown. Setae on legs dark brown. Wing light brown; pterostigma brown; petiole of cell m 1 and discal cell almost equilong (Fig. 40). Halter with stem brown, knob dark brown with pale tip. ABDOMEN (Figs 39, 41). Mainly yellow. Abdominal tergites each with three stripes, middle stripe pale gray, lateral stripes brown. Abdominal segments 5 – 9 black. Setae on abdomen brownish yellow. HYPOPYGIUM (Figs 44 – 45, 48 – 50). Gonocoxite elongated, separated from sternite 9, gonocoxite ends with an unusually strong black spine, bent inward and horn-like, ventral of spine with long setae, basal setae longest and decreases to tip, curl inward. Tergite 9 longitudinally divided into two parts, slightly connected. Sternite 9 with a median prominence, adminiculum with a pair of sharp projections on both sides. Outer gonostylus black with yellow bristles, bent, willow leaf shaped (Figs 44, 48). Inner gonostylus with a slanted bar like raise; upper beak sharp, curled inward (Figs 45, 48). Semen pump (Figs 46 – 47) as shown in figures. Aedeagus with a sclerotized suborbicular appendage at base. Female MEASUREMENTS. Body length 17.5 mm, wing length 17 mm, antenna length 3 mm (n = 1). COLORATION. Body color slightly darker than male. Antenna relatively shorter than male. Scape and pedicel yellow, flagellar segments 1 – 4 bicolor yellow with pale brown base, other segments pale brown. Brown and black part of legs paler than male. Wing same as male except knob of halter without pale tip. Abdominal tergites each with three stripes, middle stripe brown, lateral stripes black, abdominal segments 5 – 7 yellowish brown, and brown gradually deepened, 8 th segment dark brown. Other characteristics as in male (Fig. 42). OVIPOSITOR. Tergite 9 dark brown, tergite 10 shining reddish brown. Cercus brown, tip slightly upturned, outer margin with rough serration (Fig. 43).	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFBDFF97FDEBFA07FBC0FDB1.taxon	distribution	Distribution China (Tibet).	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFBDFF97FDEBFA07FBC0FDB1.taxon	discussion	Remarks The new species is similar to T. (V.) coxitalis Alexander, 1935 from Anhui and Taiwan, China, but the latter differs in the following characters: gonocoxite ends with a slightly twisted spine; outer gonostylus narrow, filiform; inner gonostylus without slanted bar-like raise; sternite 9 near caudal end on either side of midline with a conspicuous hair pencil of reddish bristles, these pencils not arising from basal tubercles (Men 2015). This species is similar to T. (V.) bicalcarata Savchenko, 1965 and T. (V.) maoershanensis Men & Young, 2017, but the outer gonostylus of the latter is not willow leaf shaped. The posterior margin of inner gonostylus with a finger-like protrusion (Men et al. 2017).	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFBAFF98FDBFFDCCFE05FB0C.taxon	description	Figs 51 – 57, 86	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFBAFF98FDBFFDCCFE05FB0C.taxon	diagnosis	Diagnosis Scape yellow, pedicel pale yellow, first flagellar segment yellowish brown, other flagellar segments dark brown. Middle of abdominal tergites 2 – 4 with pale brown spots, both sides each with brown stripe. Gonocoxite strongly elongated. Outer gonostylus brownish black, without setae except margins of apex and base with few setae. Inner gonostylus with upper beak bearing born-like elevation.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFBAFF98FDBFFDCCFE05FB0C.taxon	materials_examined	Material examined CHINA • 2 ♂♂; Tibet, Bomi, Yigong; 30 ° 16 ′ 19 ″ N, 94 ° 48 ′ 28 ″ E; 2200 m a. s. l.; 5 May 2017; Qicheng Yang leg.; light trap; CAU.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFBAFF98FDBFFDCCFE05FB0C.taxon	description	Redescription Male MEASUREMENTS. Body length 15.0 mm, wing length 17.0 mm, antenna length 4.0 mm (n = 2). HEAD (Figs 51, 53). Mainly reddish yellow. Vertex with brown stripe. Rostrum and nasus dark yellow. Setae on head dark brown. Scape yellow, pedicel pale yellow, first flagellar segment yellowish brown, other flagellar segments dark brown. Proboscis and palpus brownish yellow. THORAX (Figs 51, 53). Mainly dark yellow, heavily pruinose. Pronotum dark yellow with dark yellow setae, middle pale brown and both sides with a pale brown spot. Prescutum dark yellow with four reddish brown stripes, median stripes broadened apically and gradually narrowing to base. Scutum with four reddish brown spots. Scutellum yellow with black mid-longitudinal stripe. Mediotergite yellow. Thoracic pleuron reddish yellow. Setae on thorax yellow. Legs with coxae and femora yellow, femora with brownish black apices and middle portions, tibiae and tarsi brownish black. Setae on legs brownish black. Wing light brown; pterostigma pale brown; petiole of cell m 1 slightly shorter than discal cell (Fig. 52). Halter with stem yellow, knob pale yellow. ABDOMEN (Figs 51, 53). Mainly yellow. Middle of abdominal tergites 2 – 4 with pale brown sports, both sides each with a brown stripe. Abdominal segment 5 yellowish brown, segments 6 – 9 brownish black. Setae on abdomen yellow. HYPOPYGIUM (Figs 54 – 57). Gonocoxite strongly elongated, thick, separated from sternite 9. Tergite 9 longitudinally divided into two parts, a bridge present between them. Sternite 9 with a median prominence. Outer gonostylus sclerotized, brownish black, without setae except margins of apex and base with few setae. Inner gonostylus with setae dorsally; upper beak with a horn-like elevation bearing 2 setae at tip, base of upper beak with 2 distinct setae; inner gonostylus with setae dorsally (Fig. 55). Female See Brunetti (1911).	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFBAFF98FDBFFDCCFE05FB0C.taxon	distribution	Distribution China (Tibet, Yunnan); India (Assam, Himachal Pradesh, Sikkim, Uttarakhand, Uttar Pradesh, W Bengal); Myanmar; Vietnam.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFBAFF98FDBFFDCCFE05FB0C.taxon	discussion	Remarks Here, we add new distributional data and new illustrations. Yigong Township is now the current northernmost distribution of T. (V.) himalayensis. The species was considered by Brunetti to be variable but within certain limits and can be easily recognized by the pale yellow wide ring on all the femora about as far from the tip as the width of the ring. We examined other additional material, in addition to the previously indicated: 1 ♂ (paratype) Darjiling; 6000 feet a. s. l.; 22 Sep. 1908; Brunetti (USNM) / 1 ♂ (syntype) Darjiling; 20 Oct. 1905; Brunetti (BMNH (E) # 246064). There is no difference in the characteristics of the femora of the three, but the wing veins are slightly different. It seems that the difference in the wing veins shown in Joseph’s illustrations (Joseph 1974) can be regarded as intraspecific variation. But there are also some differences in the shape of inner gonostylus as shown in the illustrations, especially the beak different from others. It is necessary to make further observations on the specimens of ZSI.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFB5FF9DFDBFFB69FBC2FA7A.taxon	description	Figs 58 – 65	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFB5FF9DFDBFFB69FBC2FA7A.taxon	diagnosis	Diagnosis Occipital stripe dark brown. Rostrum brownish yellow, nasus brown with yellow setae. Scape brownish yellow and pedicel pale yellow, first flagellar segment brownish yellow, other flagellar segments black. Scutum yellow with four brownish black spots, which are virtually confluent on each side, anterior two spots small; all spots with olive-gray pruinescence. Upper part of fore coxa brownish black. Inner gonostylus with upper beak sharp bearing, a semicircular striped raise at base; posterior crest produced backward. Adminiculum with a pair of long projections on both sides, longer than apex of phallic guide.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFB5FF9DFDBFFB69FBC2FA7A.taxon	materials_examined	Material examined CHINA • 3 ♂♂; Tibet, Yadong; 27 ° 29 ′ 52 ″ N, 88 ° 55 ′ 51 ″ E; 3200 m a. s. l.; 12 Jul. 2018; Qicheng Yang leg.; light trap; CAU.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFB5FF9DFDBFFB69FBC2FA7A.taxon	description	Redescription Male MEASUREMENTS. Body length 12.0 – 13.0 mm, wing length 15.0 mm, antenna length 3.0 mm (n = 3). HEAD (Figs 58, 60). Mainly brownish yellow. Occipital stripe dark brown. Rostrum brownish yellow, nasus brown with yellow setae. Setae on head brownish black. Scape brownish yellow, pedicel pale yellow, first flagellar segment brownish yellow, other flagellar segments black. Proboscis and palpus black. THORAX (Figs 58, 60). Mainly brownish yellow, heavily yellow and gray pruinescence. Pronotum brownish gray. Prescutum with four dark brownish black stripes, median stripes virtually confluent, broadened apically and gradually narrowing to base, lateral stripes long-oval; all stripes with olive-gray pruinescence. Scutum yellow with four dark brownish black spots, which are virtually confluent on each side, anterior two spots small, all spots with olive-gray pruinescence. Scutellum yellow with dark brown mid-longitudinal stripe. Mediotergite yellow, middlly pale gray with dark brown mid-longitudinal stripe. Scutellum and mediotergite with golden pruinescence. Thoracic pleuron with yellow pruinescence. Anepisternum and katepisternum yellow, each with a brownish black anterior lower spot. Anepimeron yellow with a small brownish black upper spot; katepimeron yellow. Meron yellow with brownish black lower part. Parascutellum yellow. Anatergite and katatergite yellow. Setae on thorax dark yellow. Legs with coxae and femora yellow, except upper part of fore coxa brownish black, femora with black apices; tibiae brownish yellow with black apices; tarsi dark black. Setae on legs black. Wing yellowish brown; pterostigma brown; petiole of cell m 1 slightly shorter than discal cell (Fig. 59). Halter with stem brown, knob grayish black with pale gray apex. ABDOMEN (Figs 58, 60). Mainly yellow. Abdominal tergites each with three brown stripes, middle stripe pale. Abdominal segments 7 – 9 brownish black. Setae on abdomen yellow except black setae on middle stripe. HYPOPYGIUM (Figs 61 – 65). Gonocoxite elongated into dagger-shaped extension, separated from sternite 9. Posterior margin of tergite 9 depressed medially, connected by a short bridge. Sternite 9, adminiculum trident-shaped, with a pair of long projections on both sides, longer than apex of phallic guide, ventral part of adminiculum with a large projection. Outer gonostylus black, with a central bulge posterior margin bearing subtriangular protrusion (Fig. 62). Inner gonostylus with upper beak sharp, base of beak with a semicircular striped raise; posterior crest produced backward (Fig. 63). Female See Brunetti (1911).	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFB5FF9DFDBFFB69FBC2FA7A.taxon	distribution	Distribution China (Tibet); India (Uttarakhand, W Bengal); Nepal (Simbhanjang Pass).	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFB0FFA1FD82F94CFA8BFE15.taxon	description	Figs 66 – 73	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFB0FFA1FD82F94CFA8BFE15.taxon	diagnosis	Diagnosis Vertex with brownish black stripe. Scape, pedicel and first flagellar segment yellow, other flagellar segments bicolor, yellowish with dark brown base. Scutellum and mediotergite dark yellow with dark gray mid-longitudinally. Prescutum dark yellow with four stripes, median stripes reddish brown, lateral stripes dark. Tergite 9 of male separated medially, with fleshy upper tergal lobes flattened, and two widened black sclerotized blades at a lower level.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFB0FFA1FD82F94CFA8BFE15.taxon	etymology	Etymology The species is named after the shape of blades of tergite 9, the Latin adjective ‘ platyphylla ’ refers to ‘ having flat leaves’.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFB0FFA1FD82F94CFA8BFE15.taxon	materials_examined	Material examined Holotype CHINA • ♂; Tibet, Bomi, Tongmai (通麦); coordinates of the town: 30 ° 06 ′ 00 ″ N, 95 ° 04 ′ 56 ″ E; 21 Aug. 2011; Lihua Wang leg.; light trap; specific geographic coordinates and altitude not recorded; CAU.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFB0FFA1FD82F94CFA8BFE15.taxon	description	Description Male MEASUREMENTS. Body length 15.0 mm, wing length 17.5 mm, antenna length 5.0 mm (n = 1). HEAD (Figs 66, 68). Mainly reddish yellow. Vertex with brownish black stripe. Rostrum and nasus dark yellow. Setae on head dark brown. Scape, pedicel and first flagellar segment yellow, other flagellar segments bicolor, yellowish with dark brown base. Proboscis brown; palpus yellowish brown. THORAX (Figs 66, 68). Mainly reddish yellow with dark yellow pruinescence. Pronotum reddish yellow with yellow setae. Prescutum dark yellow with four reddish brown stripes, median stripes broadened apically and gradually narrowing to base, lateral stripes long-oval, dark reddish brown. Scutum almost entirely brown with four olive-gray pruinose spots. Scutellum and mediotergite dark yellow with dark gray mid-longitudinally. Thoracic pleuron reddish yellow. Anepisternum pale brown. Setae on thorax yellow. Legs with coxae and femora dark yellow, femora with dark brown apices and brown middle portion, tibiae and tarsi dark brown. Setae on legs dark brown. Wing variegated light brown; pterostigma brown; petiole of cell m 1 shorter than discal cell (Fig. 67). Halter with stem yellow, knob pale yellow with dark brown lower margin. ABDOMEN (Figs 66, 68). Mainly yellow. Middle of abdominal tergites 1 – 2 with pale brown stripes, other tergites without spot. Abdominal segments 5 – 9 brown. Setae on abdomen yellow. HYPOPYGIUM (Figs 69 – 73). Gonocoxite elongated, separated from sternite 9, gonocoxite ends with a sharp black upper spine, directed chiefly upward, bottom of gonocoxite also with a black upper spine (Figs 69, 73). Posterior margin of membranous suture between tergite 9 and gonocoxite with a spiny projection. Tergite 9 separated medially, with fleshy upper tergal lobes flattened, two widened sclerotized blades at a lower level, apex of blade black and serrulate. Sternite 9 with a median prominence; apex of appendages of sternite 9 slender, basally dilated. Outer gonostylus slender, strip-shaped; stem almost without setae (Fig. 71). Inner gonostylus with a subtriangular elevation, lower beak close to base of upper beak (Fig. 72). Female Unknown or unrecognized.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFB0FFA1FD82F94CFA8BFE15.taxon	distribution	Distribution China (Tibet).	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FFB0FFA1FD82F94CFA8BFE15.taxon	discussion	Remarks The new species is similar to T. (V.) gandharva Alexander, 1951 from India (Assam) and Myanmar, but the latter differs in the following characters: dorsolateral extensions of tergite 9 long and slender, sclerotized ventral blades much less massive; the spine of gonocoxite is smaller than this new species.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FF8CFFA7FD8FFB65FD59FEE8.taxon	description	urn: lsid: zoobank. org: act: 6536 F 0 F 9 - 0 D 40 - 490 C-BEF 5 - 06 A 3 A 1153033 Figs 75 – 83	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FF8CFFA7FD8FFB65FD59FEE8.taxon	diagnosis	Diagnosis Proboscis relatively short. Pronotum brownish yellow with dark yellow setae. Prescutum mainly reddish yellow with four olive-brown striped pruinescence, velvety, two middle stripes virtually fused. Gonocoxite ends hook-like, not separated from sternite 9; tergite 9 with two pairs of lobes, longitudinally divided into two parts, slightly connected, pair of emarginate arms arising from ventral surface of tergite 9. Inner gonostylus with posterior crest produced backward, base of inner gonostylus and outer basal lobe of inner gonostylus rolled inward, produced depression with row of short setulae.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FF8CFFA7FD8FFB65FD59FEE8.taxon	etymology	Etymology The species is named after the type locality Motuo.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FF8CFFA7FD8FFB65FD59FEE8.taxon	materials_examined	Material examined Holotype CHINA • ♂; Tibet, Motuo, 80 K; 29 ° 39 ′ 29 ″ N, 95 ° 29 ′ 24 ″ E; 2020 m a. s. l.; 13 Jun. 2017; Qicheng Yang leg.; light trap; CAU. Allotype CHINA • ♀; same collection data as for holotype; CAU. Paratype CHINA • 1 ♂; same collection data as for holotype; CAU.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FF8CFFA7FD8FFB65FD59FEE8.taxon	description	Description Male MEASUREMENTS. Body length 12.0 mm, wing length 15.0 mm, antenna length 4.0 mm (n = 2). HEAD (Figs 74 – 75). Mainly dark yellow. Vertex with dark brown stripe. Frontal tubercle relatively high. Proboscis relatively short, rostrum and nasus dark yellow. Setae on head dark brown. Scape yellow, pedicel pale yellow, flagellum dark brown. Proboscis and palpus brownish yellow. THORAX (Figs 74 – 75). Mainly dark yellow, heavily pruinescence. Pronotum brownish yellow with dark yellow setae. Prescutum mainly reddish yellow with four olive-brown striped pruinescence, velvety, two middle stripes virtually fused. Scutum with four reddish brown spots, with olive-brown striped pruinescence. Scutellum and mediotergite yellow with pale brown mid-longitudinal stripes. Thorax yellow in lateral view. Pleura below anterior thoracic spiracle to middle with an irregular brownish black spot; anepisternum and katepisternum pale brownish yellow. Setae on thorax yellow. Legs with coxae and bases of femora pale yellow, apices of femora brown; tibiae and tarsi dark brown. Setae on legs brownish black. Wing light brown; pterostigma brown; petiole of cell m 1 shorter than discal cell (Fig. 76). Halter with stem dark yellow, knob brownish yellow. ABDOMEN (Figs 74 – 75). Mainly yellow. Abdominal dorsum with three brown longitudinal stripes, abdominal segment 6 brownish yellow, segments 7 – 9 brown. Setae on abdomen dark brown and yellow. HYPOPYGIUM (Figs 79 – 83). Gonocoxite ends hook-like, not separated from sternite 9, tergite 9 bearing two pairs of lobes, longitudinally divided into two parts, slightly connected, inner margin with dense setae; a pair of emarginate arms arising from ventral surface of tergite 9, inner margin black, other areas yellow. Sternite 9 with a median prominence, middle of sternite 9 slightly raised. Sternite 8 with abundant setae. Outer gonostylus slender and small, apex curly (Figs 79 – 81). Inner gonostylus, dorsal crest produced into fishtail, base of inner gonostylus and outer basal lobe of inner gonostylus inward-rolled, produced depression with a row of short setulae, outer basal lobe of inner gonostylus large triangular (Figs 80 – 81). Female MEASUREMENTS. Body length 13 mm, wing length 15 mm, antenna length 2.4 mm (n = 1). COLORATION. Body color slightly paler than male. Antenna shorter than male, dark yellow. Legs paler than male, tibiae and tarsi dark yellow. Abdominal tergites each with three stripes, middle stripe brown, lateral stripes brownish black, abdominal 7 – 8 segments yellowish brown, 8 th segment dark brown. Junction of each 3 – 9 segments brownish black (Fig. 77). OVIPOSITOR. Tergite 9 yellowish brown, tergite 10 yellow. Cercus dark yellow, slightly up-turned, outer margin without rough serration (Fig. 78).	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FF8CFFA7FD8FFB65FD59FEE8.taxon	distribution	Distribution China (Tibet).	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
038487C8FF8CFFA7FD8FFB65FD59FEE8.taxon	discussion	Remarks The new species is similar to T. (V.) theowaldana Alexander, 1964 from Assam, India, but the latter differs in the following characters: basal abdominal tergites yellow; gonocoxite ends sharp and not hooked; tergite 9 with a pair of larger and elongate lobes. The species may form a new species group, perhaps together with T. (V.) theowaldana. However, we have not examined the specimens of the latter species and can therefore not say with certainty if these two species share characteristics that would separate them from existing species groups.	en	Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan, Yang, Ding (2021): Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet. European Journal of Taxonomy 764 (1): 152-192, DOI: https://doi.org/10.5852/ejt.2021.764.1477, URL: http://dx.doi.org/10.5852/ejt.2021.764.1477
