taxonID	type	description	language	source
03979A42FFC6C204FF78FA5C6157F78D.taxon	description	B. sect. Petermannia (Klotzsch 1854: 124) de Candolle (1859: 128).	en	Averyanov, Leonid V., Nuraliev, Maxim S., Konstantinov, Eugene L., Maisak, Tatiana V., Kopylov-Guskov, Yury O., Lyskov, Dmitry F., Quang, Bui Hong, Kuznetsov, Andrey N., Kuznetsova, Svetlana P., Nguyen, Khang Sinh, Shui, Yu-Min (2021): New and noteworthy species of Begonia (Begoniaceae) from Vietnam and Laos. Phytotaxa 522 (2): 75-93, DOI: 10.11646/phytotaxa.522.2.1
03979A42FFC6C204FF78FA5C6157F78D.taxon	diagnosis	Diagnosis: — The new species is most similar to B. tatianae Aver. in Averyanov et al. (2019 a: 93), differing mainly in its leaves with hairy petiole and blade, adaxial surface of blade with numerous irregular spots and without crystalloid bodies, inflorescences 2.5 – 8 cm long and as long as leaves, pistillate flowers with 5 tepals, glabrous ovary.	en	Averyanov, Leonid V., Nuraliev, Maxim S., Konstantinov, Eugene L., Maisak, Tatiana V., Kopylov-Guskov, Yury O., Lyskov, Dmitry F., Quang, Bui Hong, Kuznetsov, Andrey N., Kuznetsova, Svetlana P., Nguyen, Khang Sinh, Shui, Yu-Min (2021): New and noteworthy species of Begonia (Begoniaceae) from Vietnam and Laos. Phytotaxa 522 (2): 75-93, DOI: 10.11646/phytotaxa.522.2.1
03979A42FFC6C204FF78FA5C6157F78D.taxon	materials_examined	TYPE: — VIETNAM. Herbarium voucher specimen and photos prepared from living plants on 28 August 2020 by L. Averyanov, T. Maisak, plants originated from Ha Giang Province, Quan Ba District, Can Ty Commune, Sin Suoi Ho Village, around point 23 ° 05 ’ 44.4 ’’ N 105 ° 01 ’ 17.1 ’’ E, very steep mountain slopes composed of solid highly eroded karstic limestone at elevation 800 – 1000 m a. s. l., remnants of primary evergreen broad-leaved and coniferous (with Calocedrus rupestris and Xanthocyparis vietnamensis) humid forest, lithophytic and terrestrial ephemeroid herb, not common, 11 October 2019, L. Averyanov, Nguyen Sinh Khang, T. Maisak, VR 1575 a (holotype LE: LE 01076932 http: // en. herbariumle. ru /? t = occ & id = 43259; isotype LE: LE 01076911 http: // en. herbariumle. ru /? t = occ & id = 45234; photo epitypes LE: LE 01087472 http: // en. herbariumle. ru /? t = occ & id = 40437, LE 01087473 http: // en. herbariumle. ru /? t = occ & id = 40438).	en	Averyanov, Leonid V., Nuraliev, Maxim S., Konstantinov, Eugene L., Maisak, Tatiana V., Kopylov-Guskov, Yury O., Lyskov, Dmitry F., Quang, Bui Hong, Kuznetsov, Andrey N., Kuznetsova, Svetlana P., Nguyen, Khang Sinh, Shui, Yu-Min (2021): New and noteworthy species of Begonia (Begoniaceae) from Vietnam and Laos. Phytotaxa 522 (2): 75-93, DOI: 10.11646/phytotaxa.522.2.1
03979A42FFC3C201FF78FE8664C8FD17.taxon	description	B. sect. Petermannia (Klotzsch 1854: 124) de Candolle (1859: 128).	en	Averyanov, Leonid V., Nuraliev, Maxim S., Konstantinov, Eugene L., Maisak, Tatiana V., Kopylov-Guskov, Yury O., Lyskov, Dmitry F., Quang, Bui Hong, Kuznetsov, Andrey N., Kuznetsova, Svetlana P., Nguyen, Khang Sinh, Shui, Yu-Min (2021): New and noteworthy species of Begonia (Begoniaceae) from Vietnam and Laos. Phytotaxa 522 (2): 75-93, DOI: 10.11646/phytotaxa.522.2.1
03979A42FFC3C201FF78FE8664C8FD17.taxon	diagnosis	Diagnosis: — Similar to B. rubrosetosa Aver. in Averyanov & Nguyen (2012: 20) and B. saolaensis Y. M. Shui, T. A. Le & C. T. Vu in Shui et al. (2019 a: 61), differing in glabrous stem, petiole, leaf blade, pedicel and ovary, stipules 4 – 8 × 1.5 – 3 mm, narrowly oblong-lanceolate leaf blade, tepals of staminate flowers 2 – 3.5 mm long, tepals of pistillate flowers 3 – 4.5 mm long and abaxially hairy, styles 1 mm long.	en	Averyanov, Leonid V., Nuraliev, Maxim S., Konstantinov, Eugene L., Maisak, Tatiana V., Kopylov-Guskov, Yury O., Lyskov, Dmitry F., Quang, Bui Hong, Kuznetsov, Andrey N., Kuznetsova, Svetlana P., Nguyen, Khang Sinh, Shui, Yu-Min (2021): New and noteworthy species of Begonia (Begoniaceae) from Vietnam and Laos. Phytotaxa 522 (2): 75-93, DOI: 10.11646/phytotaxa.522.2.1
03979A42FFC3C201FF78FE8664C8FD17.taxon	materials_examined	TYPE: — VIETNAM. Quang Nam Province: Nam Giang District, Song Thanh Nature Reserve, in the forest, river bank, 15 ° 33 ’ 55 ’’ N 107 ° 23 ’ 15 ’’ E, elev. 1000 m, 27 April 2019, Nuraliev M. S. 2425 (holotype MW: MW 0595740; isotypes E, MW: MW 0595741; photos LE: LE 01093019 http: // en. herbariumle. ru /? t = occ & id = 43974).	en	Averyanov, Leonid V., Nuraliev, Maxim S., Konstantinov, Eugene L., Maisak, Tatiana V., Kopylov-Guskov, Yury O., Lyskov, Dmitry F., Quang, Bui Hong, Kuznetsov, Andrey N., Kuznetsova, Svetlana P., Nguyen, Khang Sinh, Shui, Yu-Min (2021): New and noteworthy species of Begonia (Begoniaceae) from Vietnam and Laos. Phytotaxa 522 (2): 75-93, DOI: 10.11646/phytotaxa.522.2.1
03979A42FFC1C203FF78FCE26373FB6B.taxon	description	B. sect. Platycentrum (Klotzsch 1855: 243) de Candolle (1859: 134).	en	Averyanov, Leonid V., Nuraliev, Maxim S., Konstantinov, Eugene L., Maisak, Tatiana V., Kopylov-Guskov, Yury O., Lyskov, Dmitry F., Quang, Bui Hong, Kuznetsov, Andrey N., Kuznetsova, Svetlana P., Nguyen, Khang Sinh, Shui, Yu-Min (2021): New and noteworthy species of Begonia (Begoniaceae) from Vietnam and Laos. Phytotaxa 522 (2): 75-93, DOI: 10.11646/phytotaxa.522.2.1
03979A42FFC1C203FF78FCE26373FB6B.taxon	diagnosis	Diagnosis: — Similar to B. duclouxii Gagnepain (1919: 198), B. limprichtii Irmsch. in Limpricht (1922: 440) and B. setifolia Irmscher (1939: 549), differing from them by the following combination of characters: leaf blade 8.2 – 11.5 cm long, hairy peduncle, bracts 3.5 – 5 mm long, purple tepals, staminate flowers with outer tepals ca. 6 mm long and ca. 3 – 4 mm wide, actinomorphic androecium, anthers with supraconnective.	en	Averyanov, Leonid V., Nuraliev, Maxim S., Konstantinov, Eugene L., Maisak, Tatiana V., Kopylov-Guskov, Yury O., Lyskov, Dmitry F., Quang, Bui Hong, Kuznetsov, Andrey N., Kuznetsova, Svetlana P., Nguyen, Khang Sinh, Shui, Yu-Min (2021): New and noteworthy species of Begonia (Begoniaceae) from Vietnam and Laos. Phytotaxa 522 (2): 75-93, DOI: 10.11646/phytotaxa.522.2.1
03979A42FFC1C203FF78FCE26373FB6B.taxon	materials_examined	TYPE: — VIETNAM. Quang Nam Province: Nam Giang District, Song Thanh Nature Reserve, forest, river bank, 15 ° 34 ’ 12 ’’ N 107 ° 22 ’ 44 ’’ E, elev. 1040 m, 02 May 2019, Nuraliev M. S. 2463 a (holotype MW: MW 0595748; photos LE: LE 01093237 http: // en. herbariumle. ru /? t = occ & id = 81268).	en	Averyanov, Leonid V., Nuraliev, Maxim S., Konstantinov, Eugene L., Maisak, Tatiana V., Kopylov-Guskov, Yury O., Lyskov, Dmitry F., Quang, Bui Hong, Kuznetsov, Andrey N., Kuznetsova, Svetlana P., Nguyen, Khang Sinh, Shui, Yu-Min (2021): New and noteworthy species of Begonia (Begoniaceae) from Vietnam and Laos. Phytotaxa 522 (2): 75-93, DOI: 10.11646/phytotaxa.522.2.1
03979A42FFCDC20FFF78FF3664B3FC2F.taxon	description	Ku (1999: 187), Shui et al. (2002: 317, 2019 b: 218, Fig. 47 D, E), Huang & Shui (2006: 171, Fig. 49), Gu et al. (2007: 196), Shui & Chen (2017: 246). B. sect. Platycentrum (Klotzsch 1855: 243) de Candolle (1859: 134).	en	Averyanov, Leonid V., Nuraliev, Maxim S., Konstantinov, Eugene L., Maisak, Tatiana V., Kopylov-Guskov, Yury O., Lyskov, Dmitry F., Quang, Bui Hong, Kuznetsov, Andrey N., Kuznetsova, Svetlana P., Nguyen, Khang Sinh, Shui, Yu-Min (2021): New and noteworthy species of Begonia (Begoniaceae) from Vietnam and Laos. Phytotaxa 522 (2): 75-93, DOI: 10.11646/phytotaxa.522.2.1
03979A42FFCDC20FFF78FF3664B3FC2F.taxon	materials_examined	TYPE: — CHINA. [Yunnan Province:] Pingbian Xian, Timi, 1600 – 1800 m, April 1959, Exped. Univ. Yunnan 59 - 8 (YUKU [as YUNU]).	en	Averyanov, Leonid V., Nuraliev, Maxim S., Konstantinov, Eugene L., Maisak, Tatiana V., Kopylov-Guskov, Yury O., Lyskov, Dmitry F., Quang, Bui Hong, Kuznetsov, Andrey N., Kuznetsova, Svetlana P., Nguyen, Khang Sinh, Shui, Yu-Min (2021): New and noteworthy species of Begonia (Begoniaceae) from Vietnam and Laos. Phytotaxa 522 (2): 75-93, DOI: 10.11646/phytotaxa.522.2.1
03979A42FFCDC20FFF78FF3664B3FC2F.taxon	biology_ecology	Ecology and phenology: — Evergreen forests, shrublands, shady moist environments, at 500 – 1800 m a. s. l. Flowering in April – October, fruiting from July.	en	Averyanov, Leonid V., Nuraliev, Maxim S., Konstantinov, Eugene L., Maisak, Tatiana V., Kopylov-Guskov, Yury O., Lyskov, Dmitry F., Quang, Bui Hong, Kuznetsov, Andrey N., Kuznetsova, Svetlana P., Nguyen, Khang Sinh, Shui, Yu-Min (2021): New and noteworthy species of Begonia (Begoniaceae) from Vietnam and Laos. Phytotaxa 522 (2): 75-93, DOI: 10.11646/phytotaxa.522.2.1
03979A42FFCDC20FFF78FF3664B3FC2F.taxon	distribution	Distribution: — China: Yunnan Province (Pingbian); northern Vietnam: Lao Cai Province (Bat Xat Nature Reserve) and Son La Province (Ta Xua Nature Reserve). Note: — The species is reported here from Vietnam for the first time. Specimens examined: — VIETNAM. Lao Cai Province: Bat Xat District, Bat Xat Nature Reserve, Y Ty Commune, 22 ° 36 ’ 06.5 ’’ N 103 ° 39 ’ 03.2 ’’ E, elev. 1420 m, 30 July 2019, Bui Hong Quang, Tran Duc Binh VN-RU 153 (HN; MW: MW 0757744; photos LE: LE 01093238 http: // en. herbariumle. ru /? t = occ & id = 96050); Son La Province: Bac Yen District, Ta Xua Nature Reserve, forest margin, 21 ° 18 ’ 44 ’’ N 104 ° 31 ’ 49 ’’ E, elev. 1340 m, 06 October 2019, Nuraliev M. S., Kopylov-Guskov Yu. O., Lyskov D. F. 2835 (MW: MW 0757091; photos LE: LE 01093020 http: // en. herbariumle. ru /? t = occ & id = 43975); Son La Province: Phu Yen District, Ta Xua Nature Reserve, forest on slope, 21 ° 21 ’ 31 ’’ N 104 ° 39 ’ 47 ’’ E, elev. 930 m, 09 October 2019, Nuraliev M. S., Kopylov-Guskov Yu. O., Lyskov D. F. 2892 (MW: MW 0757092; photos LE: LE 01093021 http: // en. herbariumle. ru /? t = occ & id = 43976).	en	Averyanov, Leonid V., Nuraliev, Maxim S., Konstantinov, Eugene L., Maisak, Tatiana V., Kopylov-Guskov, Yury O., Lyskov, Dmitry F., Quang, Bui Hong, Kuznetsov, Andrey N., Kuznetsova, Svetlana P., Nguyen, Khang Sinh, Shui, Yu-Min (2021): New and noteworthy species of Begonia (Begoniaceae) from Vietnam and Laos. Phytotaxa 522 (2): 75-93, DOI: 10.11646/phytotaxa.522.2.1
03979A42FFCDC20EFF78FC1E6517FE37.taxon	description	Smith et al. (1986: 248, Fig. 25.23), Ku (1999: 250), Shui & Huang (1999: 18), Shui et al. (2002: 321, 2019 b: 219), Huang & Shui (2006: 224), Gu et al. (2007: 202), Shui & Chen (2017: 260), Hughes et al. (2018: 339), Wang et al. (2019: 52, Fig. 3). B. sect. Platycentrum (Klotzsch 1855: 243) de Candolle (1859: 134).	en	Averyanov, Leonid V., Nuraliev, Maxim S., Konstantinov, Eugene L., Maisak, Tatiana V., Kopylov-Guskov, Yury O., Lyskov, Dmitry F., Quang, Bui Hong, Kuznetsov, Andrey N., Kuznetsova, Svetlana P., Nguyen, Khang Sinh, Shui, Yu-Min (2021): New and noteworthy species of Begonia (Begoniaceae) from Vietnam and Laos. Phytotaxa 522 (2): 75-93, DOI: 10.11646/phytotaxa.522.2.1
03979A42FFCDC20EFF78FC1E6517FE37.taxon	materials_examined	TYPE: — CHINA. Yunnan Province: Mengzi, damp forests of the Red River, 1830 – 2130 m, February 1896, W. Hancock 468 (lectotype, designated by Shui & Huang [1999: 18]: K: K 000761493 photo!).	en	Averyanov, Leonid V., Nuraliev, Maxim S., Konstantinov, Eugene L., Maisak, Tatiana V., Kopylov-Guskov, Yury O., Lyskov, Dmitry F., Quang, Bui Hong, Kuznetsov, Andrey N., Kuznetsova, Svetlana P., Nguyen, Khang Sinh, Shui, Yu-Min (2021): New and noteworthy species of Begonia (Begoniaceae) from Vietnam and Laos. Phytotaxa 522 (2): 75-93, DOI: 10.11646/phytotaxa.522.2.1
03979A42FFCDC20EFF78FC1E6517FE37.taxon	biology_ecology	Ecology and phenology: — Forests, shady moist environments on slopes and along stream banks, at 800 – 2100 m a. s. l. Flowering in June – October, fruiting from July.	en	Averyanov, Leonid V., Nuraliev, Maxim S., Konstantinov, Eugene L., Maisak, Tatiana V., Kopylov-Guskov, Yury O., Lyskov, Dmitry F., Quang, Bui Hong, Kuznetsov, Andrey N., Kuznetsova, Svetlana P., Nguyen, Khang Sinh, Shui, Yu-Min (2021): New and noteworthy species of Begonia (Begoniaceae) from Vietnam and Laos. Phytotaxa 522 (2): 75-93, DOI: 10.11646/phytotaxa.522.2.1
03979A42FFCDC20EFF78FC1E6517FE37.taxon	distribution	Distribution: — China: SE Yunnan; northern Vietnam: Ha Giang Province (districts Quan Ba and Vi Xuyen). Notes: — The presence of this species in Vietnam is confirmed here by specimen citation for the first time. Li et al. (2018) treated B. versicolor as a narrow endemic of southeast Yunnan. Hughes et al. (2015 –, 2018) indicated B. versicolor to inhabit northern Vietnam, and Pham et al. (2021) listed it for the country, but they have not provided any documentation of their records. The specimens Averyanov et al. VR 1450 and Averyanov et al. VR 1514 were erroneously assigned by Averyanov et al. (2020) to B. discolor R. Brown in Aiton (1813: 284), which is currently treated as a synonym of the Chinese endemic B. grandis Dryander (1791: 163) (Gu et al. 2007). In line with our re-identification of these specimens, we confirm that B. grandis is unknown from Vietnam at present time. Specimens examined: — VIETNAM. Ha Giang Province: Quan Ba District, Tung Vai Commune, Thang Village, around point 23 ° 03 ’ 04.2 ’’ N 104 ° 51 ’ 09.7 ’’ E, steep rocky slopes near mountain top composed with eroded stratified highly eroded limestone at elevation 1200 – 1450 m a. s. l., primary evergreen broad-leaved very humid forest, lithophytic herb on steep rocky slope, leaves dark green above, pale light green with unclear purplish radiating spots below, locally common, 24 April 2018, L. Averyanov, Nguyen Sinh Khang, Nguyen Tien Hiep, Nguyen Quang Hieu, Chuong Quang Ngan, T. Maisak, VR 794 (LE: LE 01050193 http: // en. herbariumle. ru /? t = occ & id = 4045); Ha Giang Province: Quan Ba District, Tung Vai Commune, Thang Village, around point 23 ° 03 ’ 12.6 ’’ N 104 ° 51 ’ 52.1 ’’ E, steep rocky slopes of river valley composed with eroded limestone mixed with sandstone at elevation 1000 – 1100 m a. s. l., remnants of primary evergreen broad-leaved very humid forest along river, terrestrial and lithophytic herb on very steep shady river slope, leaves dirty purple marked above, not common, 07 October 2019, L. Averyanov, Nguyen Sinh Khang, T. Maisak, VR 1450 (LE: LE 01066974 http: // en. herbariumle. ru /? t = occ & id = 12755; photo LE: LE 01061058 http: // en. herbariumle. ru /? t = occ & id = 13034); Ha Giang Province: Quan Ba District, Thai An Commune, Seo Lung Two Village, around point 22 ° 58 ’ 11.6 ’’ N 105 ° 05 ’ 11.3 ’’ E, very steep mountain slopes composed of stratified shale and clayey subsoil at elevation 800 – 1000 m a. s. l., primary and secondary evergreen broad-leaved humid forest, terrestrial rosulate rhizomatous herb on steep shady slope, leaves above dark green to purple, often with palmate spots, below dark purple, flowers pink, anthers yellow, occasional, 10 October 2019, L. Averyanov, Nguyen Sinh Khang, T. Maisak, VR 1514 (LE: LE 01067108 http: // en. herbariumle. ru /? t = occ & id = 12829; photos LE: LE 01070176 http: // en. herbariumle. ru /? t = occ & id = 13171); Ha Giang Province: Vi Xuyen District, Cao Bo Municipality, Tay Con Linh Nature Reserve, disturbed forest on slope, 22 ° 46 ’ 35 ’’ N 104 ° 53 ’ 15 ’’ E, elev. 1310 m, 19 October 2019, Nuraliev M. S., Kopylov-Guskov Yu. O., Lyskov D. F. 3004 (MW: MW 0757089, MW 0757090; photos LE: LE 01093022 http: // en. herbariumle. ru /? t = occ & id = 43977).	en	Averyanov, Leonid V., Nuraliev, Maxim S., Konstantinov, Eugene L., Maisak, Tatiana V., Kopylov-Guskov, Yury O., Lyskov, Dmitry F., Quang, Bui Hong, Kuznetsov, Andrey N., Kuznetsova, Svetlana P., Nguyen, Khang Sinh, Shui, Yu-Min (2021): New and noteworthy species of Begonia (Begoniaceae) from Vietnam and Laos. Phytotaxa 522 (2): 75-93, DOI: 10.11646/phytotaxa.522.2.1
03979A42FFCCC20EFF78FE14647DFC07.taxon	description	B. sect. Platycentrum (Klotzsch 1855: 243) de Candolle (1859: 134).	en	Averyanov, Leonid V., Nuraliev, Maxim S., Konstantinov, Eugene L., Maisak, Tatiana V., Kopylov-Guskov, Yury O., Lyskov, Dmitry F., Quang, Bui Hong, Kuznetsov, Andrey N., Kuznetsova, Svetlana P., Nguyen, Khang Sinh, Shui, Yu-Min (2021): New and noteworthy species of Begonia (Begoniaceae) from Vietnam and Laos. Phytotaxa 522 (2): 75-93, DOI: 10.11646/phytotaxa.522.2.1
03979A42FFCCC20EFF78FE14647DFC07.taxon	diagnosis	Diagnosis: — Remarkable in producing bulbils at junction of petiole and leaf blade; most close to B. longialata K. Y. Guan & D. K. Tian (2000: 132), differing in petiole 5 – 10 cm long, shallowly lobed leaf blade 8 – 12 cm long and wide and setose on both sides and margin, staminate flowers with pedicel 12 – 22 mm long and tepals subequal in length, pistillate flowers with pedicel 0.4 – 1.2 cm long and with 4 tepals, styles 2.2 – 3 mm long. Also the new species is similar to B. macrotoma Irmscher (1951: 41), differing in its abaxially hairy leaf blade, glabrous peduncle, abaxially glabrous outer tepals of staminate flowers and all tepals of pistillate flowers, staminate flowers with outer tepals 10 – 15 mm long and 16 – 21 mm wide, pistillate flowers with 2 or 3 styles.	en	Averyanov, Leonid V., Nuraliev, Maxim S., Konstantinov, Eugene L., Maisak, Tatiana V., Kopylov-Guskov, Yury O., Lyskov, Dmitry F., Quang, Bui Hong, Kuznetsov, Andrey N., Kuznetsova, Svetlana P., Nguyen, Khang Sinh, Shui, Yu-Min (2021): New and noteworthy species of Begonia (Begoniaceae) from Vietnam and Laos. Phytotaxa 522 (2): 75-93, DOI: 10.11646/phytotaxa.522.2.1
03979A42FFCCC20EFF78FE14647DFC07.taxon	materials_examined	TYPE: — LAO PDR. Champasak Province: at the base of Tad Tayicseau Waterfall, 15 ° 05 ’ 02.9 ’’ N 106 ° 35 ’ 28.8 ’’ E, at elevation 725 m a. s. l., lithophyte on stream rocks along shady river slopes, 07 December 2017, E. Konstantinov, T. Vongsa, K 418 (holotype LE: LE 01057897 http: // en. herbariumle. ru /? t = occ & id = 8387).	en	Averyanov, Leonid V., Nuraliev, Maxim S., Konstantinov, Eugene L., Maisak, Tatiana V., Kopylov-Guskov, Yury O., Lyskov, Dmitry F., Quang, Bui Hong, Kuznetsov, Andrey N., Kuznetsova, Svetlana P., Nguyen, Khang Sinh, Shui, Yu-Min (2021): New and noteworthy species of Begonia (Begoniaceae) from Vietnam and Laos. Phytotaxa 522 (2): 75-93, DOI: 10.11646/phytotaxa.522.2.1
03979A42FFC9C20AFF78FA5E65B1FEA7.taxon	description	Ku (1999: 168), Shui et al. (2002: 325, 2019 b: 220), Huang & Shui (2006: 163), Gu et al. (2007: 203, Fig. 225 5 – 9), Shui & Chen (2017: 98), Dong & Liu (2019: 37). B. sect. Platycentrum (Klotzsch 1855: 243) de Candolle (1859: 134).	en	Averyanov, Leonid V., Nuraliev, Maxim S., Konstantinov, Eugene L., Maisak, Tatiana V., Kopylov-Guskov, Yury O., Lyskov, Dmitry F., Quang, Bui Hong, Kuznetsov, Andrey N., Kuznetsova, Svetlana P., Nguyen, Khang Sinh, Shui, Yu-Min (2021): New and noteworthy species of Begonia (Begoniaceae) from Vietnam and Laos. Phytotaxa 522 (2): 75-93, DOI: 10.11646/phytotaxa.522.2.1
03979A42FFC9C20AFF78FA5E65B1FEA7.taxon	materials_examined	TYPE: — CHINA. Yunnan Province: Wenshan mt., in mixed forests, 2000 m, 17 August 1947, Feng K. M. 11323 (holotype PE; isotypes E: E 00022104 photo!, KUN).	en	Averyanov, Leonid V., Nuraliev, Maxim S., Konstantinov, Eugene L., Maisak, Tatiana V., Kopylov-Guskov, Yury O., Lyskov, Dmitry F., Quang, Bui Hong, Kuznetsov, Andrey N., Kuznetsova, Svetlana P., Nguyen, Khang Sinh, Shui, Yu-Min (2021): New and noteworthy species of Begonia (Begoniaceae) from Vietnam and Laos. Phytotaxa 522 (2): 75-93, DOI: 10.11646/phytotaxa.522.2.1
03979A42FFC9C20AFF78FA5E65B1FEA7.taxon	biology_ecology	Ecology and phenology: — Evergreen broad-leaved forests, moist environments along streams, mountain bamboo thickets, at 1000 – 2200 m a. s. l. Flowering in June – October, fruiting from August.	en	Averyanov, Leonid V., Nuraliev, Maxim S., Konstantinov, Eugene L., Maisak, Tatiana V., Kopylov-Guskov, Yury O., Lyskov, Dmitry F., Quang, Bui Hong, Kuznetsov, Andrey N., Kuznetsova, Svetlana P., Nguyen, Khang Sinh, Shui, Yu-Min (2021): New and noteworthy species of Begonia (Begoniaceae) from Vietnam and Laos. Phytotaxa 522 (2): 75-93, DOI: 10.11646/phytotaxa.522.2.1
03979A42FFC9C20AFF78FA5E65B1FEA7.taxon	distribution	Distribution: — China: Yunnan Province (Funing, Wenshan); northern Vietnam: Cao Bang Province (Phia Oac- Phia Den National Park) and Lao Cai Province (Bat Xat Nature Reserve). Note: — The species is reported here from Vietnam for the first time. Specimens examined: — VIETNAM. Lao Cai Province: Bat Xat District, Bat Xat Nature Reserve, 4 km SSE of Y Ty village, disturbed forest, 22 ° 36 ’ 46 ’’ N 103 ° 37 ’ 22 ’’ E, elev. 1980 m, 13 June 2019, Nuraliev M. S. 2726 (E, MW: MW 0757094, MW 0757095; photos LE: LE 01093024 http: // en. herbariumle. ru /? t = occ & id = 43979); Cao Bang Province: Nguyen Binh District, Phia Oac-Phia Den National Park, 11 km WSW of Nguyen Binh town, Phia Oac summit area, wet foggy mossy crooked forest with bamboo on slope, 22 ° 36 ’ 58 ’’ N 105 ° 51 ’ 49 ’’ E, elev. 1900 m, 03 October 2018, Nuraliev M. S. 2235 (E, MW: MW 0757093; photos LE: LE 01093023 http: // en. herbariumle. ru /? t = occ & id = 43978).	en	Averyanov, Leonid V., Nuraliev, Maxim S., Konstantinov, Eugene L., Maisak, Tatiana V., Kopylov-Guskov, Yury O., Lyskov, Dmitry F., Quang, Bui Hong, Kuznetsov, Andrey N., Kuznetsova, Svetlana P., Nguyen, Khang Sinh, Shui, Yu-Min (2021): New and noteworthy species of Begonia (Begoniaceae) from Vietnam and Laos. Phytotaxa 522 (2): 75-93, DOI: 10.11646/phytotaxa.522.2.1
