taxonID	type	description	language	source
03AEE5789D4D3E2F7772D39E71AAFB63.taxon	description	Figs 1 – 2	en	Frolov, Andrey V., Akhmetova, Lilia A. (2021): Review of the Orphninae (Coleoptera: Scarabaeidae) of Sri Lanka, with description of a new species of genus Orphnus Macleay, 1819. European Journal of Taxonomy 767 (1): 40-54, DOI: https://doi.org/10.5852/ejt.2021.767.1485, URL: http://dx.doi.org/10.5852/ejt.2021.767.1485
03AEE5789D4D3E2F7772D39E71AAFB63.taxon	diagnosis	Differential diagnosis Orphnus mysoriensis is similar to O. medvedevi sp. nov. in having the pronotum with a more or less developed tubercle on the base medially and the endophallus with one group of spinules. It differs from the latter in body length (8.0 – 10.0 mm, as opposed to 4.6 – 6.0 mm in O. medvedevi sp. nov.), spinules of endophallus as a rather dispersed group of more than 10 spinules (as opposed to a compact cluster consisting of less than 10 spinules in O. medvedevi sp. nov., Fig. 1 G vs Fig. 3 G), and tubercle on the base of pronotum normally with two punctures (as opposed to always smooth tubercle in O. medvedevi sp. nov.).	en	Frolov, Andrey V., Akhmetova, Lilia A. (2021): Review of the Orphninae (Coleoptera: Scarabaeidae) of Sri Lanka, with description of a new species of genus Orphnus Macleay, 1819. European Journal of Taxonomy 767 (1): 40-54, DOI: https://doi.org/10.5852/ejt.2021.767.1485, URL: http://dx.doi.org/10.5852/ejt.2021.767.1485
03AEE5789D4D3E2F7772D39E71AAFB63.taxon	materials_examined	Type material of Orphnus mysoriensis Lectotype (here designated, Fig. 1 A – E, G) INDIA • ♂; “ Mysore / Orphnus mysoriensis Westw / TYPE WESTWOOD Trans. Ent. Soc. 4. 1846. P. 176. Coll. Hope Oxon. / TYPE COL: 484 1 / 3 Orphnus mysoriensis Westw. HOPE DEPT. OXFORD ”; OUM. Paralectotypes INDIA • 1 ♂; “ Mysore / TYPE COL: 484 2 / 3 Aegidium mysoriensis Westw. HOPE DEPT. OXFORD ”; OUM • 1 ♀; “ Mysore / TYPE COL: 484 3 / 3 Aegidium mysoriensis Westw. HOPE DEPT. OXFORD ”; OUM. Type material of Orphnus detegens Lectotype (here designated, Fig. 2 E – G) SRI LANKA • ♂; “ Type / Ceylon 59 106 / Orphnus detegens Walker. a. n. H. (type) / detegens Wlk ”; BMNH. Additional material examined SRI LANKA – Central Province • 3 ♂♂, 4 ♀♀; Kandy; 7 ° 18 ′ N, 80 ° 38 ′ E; MHNG • 2 ♂♂; same locality data as for preceding; MNHB • 7 ♂♂; same locality data as for preceding; 1 – 18 Apr. 1991; Jiri Kolibac leg.; NHMB • 1 ♀; same locality data as for preceding; 18 Mar. 1973; G. Zimmermann leg.; MNHB • 2 ♂♂; same locality data as for preceding; 1908; G. B. Longstaff leg.; OUM • 2 ♂♂, 2 ♀♀; Nalanda; 7 ° 40 ′ N, 80 ° 39 ′ E; V. De Poll leg.; MNHN • 1 ♀; same locality data as for preceding; 1889; I. Z. Kannegeter leg.; MNHN • 2 ♂♂, 1 ♀; Peradenyia; 7 ° 16 ′ N, 80 ° 35 ′ E; MHNG • 3 ♂♂; same locality data as for preceding; Friederichs S. G. leg.; MNHB • 2 ♂♂; same locality data as for preceding; 2 Apr. 1907; O. John leg.; ZIN • 1 ♂; same locality data as for preceding; 3 Apr. 1907; O. John leg.; ZIN • 2 ♂♂, 1 ♀; same locality data as for preceding; Apr. 1901; MNHN • 5 ♂♂; same locality data as for preceding; Apr. 1914; Friederichs S. G. leg.; MNHB • 2 ♂♂; same locality data as for preceding; Apr. 1939; J. Vinson leg.; MNHN • 1 ♀; Punduloya; 7 ° 01 ′ N, 80 ° 40 ′ E; OUM • 1 ♂, 1 ♀; Sigiriya; 7 ° 57 ′ N, 80 ° 45 ′ E; Oct. 1977; MCSN • 1 ♂, 1 ♀; same locality data as for preceding; Oct. 1977; NMPC • 1 ♂; Weragamtota; 7 ° 19 ′ N, 80 ° 59 ′ E; 13 Sep. 1953; F. Keiser leg.; NHMB. – North Central Province • 2 ♀♀; Anuradhapura; 8 ° 19 ′ N, 80 ° 24 ′ E; Friederichs S. G. leg.; MNHB • 4 ♂♂; Kala Oya; 8 ° 13 ′ N, 80 ° 06 ′ E; 7 Nov. 1983; MNHN • 1 ♂; same locality data as for preceding; 8 Nov. 1983; MNHN • 2 ♂♂; Nat. Park Wilpattu; 8 ° 25 ′ N, 80 ° 03 ′ E; 7 Oct. 1982; G. S. Medvedev leg.; ZIN • 1 ♂; Nat. Park Wilpattu, Talawila; 8 ° 25 ′ N, 80 ° 03 ′ E; 9 Oct. 1982; G. S. Medvedev leg.; ZIN • 1 ♂, 3 ♀♀; Wilpattu N. P. 17 km WNW entrance Tala Wila; 8 ° 25 ′ N, 80 ° 03 ′ E; 8 Oct. 1982; V. F. Zaitzev leg.; ZIN. – North Western Province • 2 ♂♂, 1 ♀; Puttalam; 8 ° 02 ′ N, 79 ° 50 ′ E; 1899; W. Horn leg.; MNHN • 1 ♂; Rajakadaluwa; 7 ° 39 ′ N, 79 ° 50 ′ E; 22 Aug. 1953; F. Keiser leg.; NHMB. – Sabaragamuwa Province • 2 ♀♀; Sabaragamuwa, Ratnapura; 6 ° 45 ′ N, 80 ° 30 ′ E; 20 Jan. 1970; Mussard, Besuchet, Lobl leg.; MCSN. – Southern Province • 3 ♀♀; Bentota; 6 ° 26 ′ N, 79 ° 60 ′ E; 14 Aug. 1978; P. Cabella leg.; MCSN • 1 ♂; same locality data as for preceding; 16 Aug. 1978; P. Cabella leg.; MCSN • 1 ♀; same locality data as for preceding; 20 Aug. 1978; P. Cabella leg.; MCSN • 3 ♂♂, 5 ♀♀; same locality data as for preceding; 23 Mar. 1973; G. Zimmermann leg.; MNHB • 1 ♀; Matara; 5 ° 57 ′ N, 80 ° 33 ′ E; 20 Oct. 1953; F. Keiser leg.; NHMB. – Western Province • 1 ♀; Colombo; 6 ° 56 ′ N, 79 ° 51 ′ E; MNHN • 1 ♂; same locality data as for preceding; MHNG • 1 ♂; same locality data as for preceding; Feb. 1884; MNHB • 1 ♂, 1 ♀; same locality data as for preceding; Mar. 1953; MHNG • 3 ♂♂; Hendala; 6 ° 59 ′ N, 79 ° 53 ′ E; 15 Aug. 1979; MNHN • 3 ♀♀; Kalutara; 6 ° 35 ′ N, 79 ° 58 ′ E; Mar. 1986; W. U. E. Just leg.; NMPC • 2 ♂♂; Kalutara Distr., Tebuwana; 6 ° 35 ′ N, 80 ° 03 ′ E; 13 Mar. 2000; S. Mahunka and L. Mahunka-Papp leg.; HNHM • 1 ♀; Mount Lavinia; 6 ° 50 ′ N, 79 ° 52 ′ E; 5 Apr. 1973; G. Benick leg.; MNHB • 5 ♂♂, 4 ♀♀; Wadduwa; 6 ° 40 ′ N, 79 ° 56 ′ E; V. De Poll leg.; MNHN. – Sri Lanka (no exact locality) • 9 ♂♂, 1 ♀; MNHN • 3 ♂♂, 2 ♀♀; IRSNB • 1 ♂; OUM • 1 ♂; NMPC • 2 ♂♂; ZIN • 4 ♂♂, 5 ♀♀; MNHB • 1 ♂, 1 ♀; MHNG • 1 ♀; Diener leg.; HNHM • 3 ♀♀; Friederichs S. G. leg.; MNHB • 2 ♀♀; S. Niethner leg.; MNHB • 1 ♂; Templeton leg.; OUM • 1 ♂; Walker leg.; BMNH • 2 ♂♂; Wollaston leg.; OUM • 11 ♂♂; 1871; Thwaites leg.; OUM • 2 ♂♂; 1873; Thwaites leg.; OUM • 5 ♂♂; 1889; H. Fruhstorfer leg.; MNHN • 1 ♂; Mar. 1889; H. Fruhstorfer leg.; MNHN.	en	Frolov, Andrey V., Akhmetova, Lilia A. (2021): Review of the Orphninae (Coleoptera: Scarabaeidae) of Sri Lanka, with description of a new species of genus Orphnus Macleay, 1819. European Journal of Taxonomy 767 (1): 40-54, DOI: https://doi.org/10.5852/ejt.2021.767.1485, URL: http://dx.doi.org/10.5852/ejt.2021.767.1485
03AEE5789D4D3E2F7772D39E71AAFB63.taxon	description	Variation Body length of the examined specimens varied from 8.0 to 10.5 mm (males) and from 7.0 to 9.0 mm (females). Head and pronotum armature in males varied from well-developed with long lateral pronotal processes and frontoclypeal horn (Fig. 2 A) to excavated pronotum with gibbosities aside of the excavation and frontoclypeal tubercle (Fig. 2 C), with intermediate variants (Fig. 2 B).	en	Frolov, Andrey V., Akhmetova, Lilia A. (2021): Review of the Orphninae (Coleoptera: Scarabaeidae) of Sri Lanka, with description of a new species of genus Orphnus Macleay, 1819. European Journal of Taxonomy 767 (1): 40-54, DOI: https://doi.org/10.5852/ejt.2021.767.1485, URL: http://dx.doi.org/10.5852/ejt.2021.767.1485
03AEE5789D4D3E2F7772D39E71AAFB63.taxon	distribution	Distribution The species is distributed rather widely on Sri Lanka except for the eastern part (Fig. 1 H).	en	Frolov, Andrey V., Akhmetova, Lilia A. (2021): Review of the Orphninae (Coleoptera: Scarabaeidae) of Sri Lanka, with description of a new species of genus Orphnus Macleay, 1819. European Journal of Taxonomy 767 (1): 40-54, DOI: https://doi.org/10.5852/ejt.2021.767.1485, URL: http://dx.doi.org/10.5852/ejt.2021.767.1485
03AEE5789D493E227730D02B714BFE3B.taxon	description	urn: lsid: zoobank. org: act: 3 F 99 F 2 E 5 - 0 FD 6 - 4 CE 9 - A 3 AC- 53 A 7 FBA 4241 D Fig. 3	en	Frolov, Andrey V., Akhmetova, Lilia A. (2021): Review of the Orphninae (Coleoptera: Scarabaeidae) of Sri Lanka, with description of a new species of genus Orphnus Macleay, 1819. European Journal of Taxonomy 767 (1): 40-54, DOI: https://doi.org/10.5852/ejt.2021.767.1485, URL: http://dx.doi.org/10.5852/ejt.2021.767.1485
03AEE5789D493E227730D02B714BFE3B.taxon	diagnosis	Differential diagnosis Orphnus medvedevi sp. nov. is similar to O. mysoriensis in having the pronotum with more or less developed tubercle on base medially and endophallus with one group of spinules. It differs from the latter in shorter body (4.6 – 6.0 mm, as opposed to 8.0 – 10.0 mm. in O. mysoriensis), spinules of endophallus as a compact cluster consisting of less than 10 spinules (as opposed to a rather dispersed group of more than 10 spinules in O. mysoriensis, Fig. 3 G vs Fig. 1 G), and tubercle on the base of pronotum always smooth (as opposed to having normally two punctures in O. mysoriensis).	en	Frolov, Andrey V., Akhmetova, Lilia A. (2021): Review of the Orphninae (Coleoptera: Scarabaeidae) of Sri Lanka, with description of a new species of genus Orphnus Macleay, 1819. European Journal of Taxonomy 767 (1): 40-54, DOI: https://doi.org/10.5852/ejt.2021.767.1485, URL: http://dx.doi.org/10.5852/ejt.2021.767.1485
03AEE5789D493E227730D02B714BFE3B.taxon	etymology	Etymology The new species is named after Gleb Sergeevich Medvedev (1931 – 2009), a Russian coleopterist.	en	Frolov, Andrey V., Akhmetova, Lilia A. (2021): Review of the Orphninae (Coleoptera: Scarabaeidae) of Sri Lanka, with description of a new species of genus Orphnus Macleay, 1819. European Journal of Taxonomy 767 (1): 40-54, DOI: https://doi.org/10.5852/ejt.2021.767.1485, URL: http://dx.doi.org/10.5852/ejt.2021.767.1485
03AEE5789D493E227730D02B714BFE3B.taxon	materials_examined	Type material Holotype SRI LANKA • ♂; “ Shri Lanka Vilpattu [Sri Lanka, Wilpattu] 7.10.1982 G. Medvedev ”; ZIN. Paratypes SRI LANKA • 2 ♀♀; “ Shri Lanka Vilpattu [Sri Lanka, Wilpattu] 8.10.1982 G. Medvedev ”; ZIN • 1 ♂, 6 ♀♀; “ Sri Lanka. Nat. Park Wilpattu. Talawila 9. X. 982. Medvedev ”; ZIN • 13 ♂♂, 13 ♀♀; “ Sri Lanka, Wilpattu N. P. 17 km WNW entrance Tala Wila, 8. X. 1982 V. F. Zaitzev leg. ”; ZIN.	en	Frolov, Andrey V., Akhmetova, Lilia A. (2021): Review of the Orphninae (Coleoptera: Scarabaeidae) of Sri Lanka, with description of a new species of genus Orphnus Macleay, 1819. European Journal of Taxonomy 767 (1): 40-54, DOI: https://doi.org/10.5852/ejt.2021.767.1485, URL: http://dx.doi.org/10.5852/ejt.2021.767.1485
03AEE5789D493E227730D02B714BFE3B.taxon	description	Description Male, holotype (Fig. 3 A, D – G) BODY. Length 6.0 mm. Colour uniformly brown. CLYPEUS. Wide, with convex anterior margin, rounded laterally, finely crenulate. Genae small, not protruding past eyes. Frontal suture indistinct. Clypeus with long, slender horn directed upwards and slightly curved backwards apically. Dorsal surface of head finely punctate. Labrum deeply sinuate in the middle, distinctly protruding past clypeus. PRONOTUM. With rounded sides, about 1.5 times as wide as long, with deeply concave disc, conical lateral processes aside excavation, and smooth, rounded tubercle medially near base. Anterior angles acute; posterior angles rounded, indistinct in dorsal view. Pronotum bordered on anterior margin and base. Lateral margins with long, sparse, brown setae. Sides irregularly punctate with round punctures separated by 3 – 5 puncture diameters. SCUTELLUM. Subtriangular, narrowly rounded apically, about 1 / 10 length of elytra. ELYTRA. About as long as wide, with distinct humeral humps, widest in middle, lateral margins slightly rounded in basal half. First (sutural) stria distinct, as feebly impressed groove with row of punctures. Other stria before humeral humps as rows of round setiferous punctures, separated by more than 3 their diameters except for base of elytra. Elytral intervals covered with minute punctures. WINGS. Macropterous. LEGS. Protibiae with 3 outer teeth. Lateral margin basad of outer teeth not crenulate. Apical spur of protibia absent. Left protarsus absent. Middle and hind legs similar in shape; metafemora and metatibiae about ⅛ as long as the mesofemora and mesotibiae. Mesotibia and metatibiae somewhat triangular with 2 apical spurs, inner margin almost straight, with 1 transverse keel. Upper spur of tibiae as long as two basal tarsomeres. Claws ⅓ length of apical tarsomere. Femora almost impunctate. ABDOMEN. Ventrally irregularly punctate, pubescent, with sparse, long setae. Abdominal sternite 8 medially shorter than sternites 4 – 7 combined. Pygidium invisible from above, with slightly truncate apex in caudal view. Plectrum triangular with rounded apex, wider than long. AEDEAGUS. Parameres relatively short (0.5 length of phallobase), curved downwards, tapering apically, spear-shaped in dorsal view, with feebly visible lateral teeth (Fig. 3 D – E). Endophallus with a compact cluster of 6 spinules (Fig. 3 G). Female Female (Fig. 3 H) differs from the male in having a relatively smaller pronotum without armature, frontoclypeus without process, and prothoracic spur present. Variation Body length of the paratypes varies from 5.9 to 4.6 mm (males) and 4.7 to 6.0 mm (females). Some of the paratypes have darker elytra. Head and pronotum armature in males paratypes varies from relatively well-developed, similar to that of the holotype, to excavated pronotum with gibbosities beside the excavation and frontoclypeal tubercle (Fig. 3 C), with intermediate variants (Fig. 3 B).	en	Frolov, Andrey V., Akhmetova, Lilia A. (2021): Review of the Orphninae (Coleoptera: Scarabaeidae) of Sri Lanka, with description of a new species of genus Orphnus Macleay, 1819. European Journal of Taxonomy 767 (1): 40-54, DOI: https://doi.org/10.5852/ejt.2021.767.1485, URL: http://dx.doi.org/10.5852/ejt.2021.767.1485
03AEE5789D493E227730D02B714BFE3B.taxon	distribution	Distribution The species is known from one locality in north-western Sri Lanka (Fig. 3 I).	en	Frolov, Andrey V., Akhmetova, Lilia A. (2021): Review of the Orphninae (Coleoptera: Scarabaeidae) of Sri Lanka, with description of a new species of genus Orphnus Macleay, 1819. European Journal of Taxonomy 767 (1): 40-54, DOI: https://doi.org/10.5852/ejt.2021.767.1485, URL: http://dx.doi.org/10.5852/ejt.2021.767.1485
03AEE5789D443E22770CD4EE7786F938.taxon	description	Fig. 4	en	Frolov, Andrey V., Akhmetova, Lilia A. (2021): Review of the Orphninae (Coleoptera: Scarabaeidae) of Sri Lanka, with description of a new species of genus Orphnus Macleay, 1819. European Journal of Taxonomy 767 (1): 40-54, DOI: https://doi.org/10.5852/ejt.2021.767.1485, URL: http://dx.doi.org/10.5852/ejt.2021.767.1485
03AEE5789D443E22770CD4EE7786F938.taxon	diagnosis	Differential diagnosis Orphnus bicolor is similar to O. parvus in having the pronotum without tubercle on base medially and endophallus with a few groups of spinules. It differs from the latter in having the parameres in dorsal view with small but distinct notches basad of the lateral teeth (Fig. 4 F), longer parameres (about 0.7 length of phallobase [Fig. 4 E] as opposed to 0.5 length of phallobase in O. parvus [Fig. 5 E]), and abdominal sternite 8 medially longer than sternites 4 – 7 combined (as opposed to sternite 8 medially as long as sternites 4 – 7 combined in O. parvus).	en	Frolov, Andrey V., Akhmetova, Lilia A. (2021): Review of the Orphninae (Coleoptera: Scarabaeidae) of Sri Lanka, with description of a new species of genus Orphnus Macleay, 1819. European Journal of Taxonomy 767 (1): 40-54, DOI: https://doi.org/10.5852/ejt.2021.767.1485, URL: http://dx.doi.org/10.5852/ejt.2021.767.1485
03AEE5789D443E22770CD4EE7786F938.taxon	materials_examined	Type material Lectotype (here designated, Fig. 4 A – F) INDIA • ♂; “ TYPE / Ind. orient. Daldorff Mus. T. Lund Geotrupes bicolor F. / Lectotype Geotrupes bicolor F. A. Frolov des. 2016 ”; ZMUKG. Paralectotype INDIA • 1 ♂; “ TYPE / Paralectotype Geotrupes bicolor F. A. Frolov des. 2016 ”; ZMUKG. Additional material examined SRI LANKA – Central Province • 1 ♂; Weragamtota; 7 ° 19 ′ N, 80 ° 59 ′ E; 13 Sep. 1953; F. Keiser leg.; NHMB. – Southern Province • 1 ♂, 1 ♀; Tissamaharana; 6 ° 17 ′ N, 81 ° 17 ′ E; 30 Oct. 1983; MNHN.	en	Frolov, Andrey V., Akhmetova, Lilia A. (2021): Review of the Orphninae (Coleoptera: Scarabaeidae) of Sri Lanka, with description of a new species of genus Orphnus Macleay, 1819. European Journal of Taxonomy 767 (1): 40-54, DOI: https://doi.org/10.5852/ejt.2021.767.1485, URL: http://dx.doi.org/10.5852/ejt.2021.767.1485
03AEE5789D443E22770CD4EE7786F938.taxon	description	Variation Body length of the examined specimens varies from 7.5 to 9.0 mm (males) and 8.6 mm (female).	en	Frolov, Andrey V., Akhmetova, Lilia A. (2021): Review of the Orphninae (Coleoptera: Scarabaeidae) of Sri Lanka, with description of a new species of genus Orphnus Macleay, 1819. European Journal of Taxonomy 767 (1): 40-54, DOI: https://doi.org/10.5852/ejt.2021.767.1485, URL: http://dx.doi.org/10.5852/ejt.2021.767.1485
03AEE5789D443E22770CD4EE7786F938.taxon	distribution	Distribution The species was described form “ East India ”. In Sri Lanka, it is known from two rather distant localities in the centre and on the southern coast (Fig. 4 I).	en	Frolov, Andrey V., Akhmetova, Lilia A. (2021): Review of the Orphninae (Coleoptera: Scarabaeidae) of Sri Lanka, with description of a new species of genus Orphnus Macleay, 1819. European Journal of Taxonomy 767 (1): 40-54, DOI: https://doi.org/10.5852/ejt.2021.767.1485, URL: http://dx.doi.org/10.5852/ejt.2021.767.1485
03AEE5789D443E26777ED3EF7000FEDF.taxon	description	Fig. 5	en	Frolov, Andrey V., Akhmetova, Lilia A. (2021): Review of the Orphninae (Coleoptera: Scarabaeidae) of Sri Lanka, with description of a new species of genus Orphnus Macleay, 1819. European Journal of Taxonomy 767 (1): 40-54, DOI: https://doi.org/10.5852/ejt.2021.767.1485, URL: http://dx.doi.org/10.5852/ejt.2021.767.1485
03AEE5789D443E26777ED3EF7000FEDF.taxon	diagnosis	Differential diagnosis Orphnus parvus is similar to O. bicolor in having the pronotum without a tubercle on base medially and endophallus with a few groups of spinules. It differs from the latter in having the parameres in dorsal view without notches basad of lateral teeth (Fig. 5 G), shorter parameres (about 0.5 length of phallobase [Fig. 5 E] as opposed to 0.7 length of phallobase in O. bicolor [Fig. 4 E]), and abdominal sternite 8 medially as long as sternites 4 – 7 combined (as opposed to sternite 8 medially distinctly longer than sternites 4 – 7 combined in O. bicolor).	en	Frolov, Andrey V., Akhmetova, Lilia A. (2021): Review of the Orphninae (Coleoptera: Scarabaeidae) of Sri Lanka, with description of a new species of genus Orphnus Macleay, 1819. European Journal of Taxonomy 767 (1): 40-54, DOI: https://doi.org/10.5852/ejt.2021.767.1485, URL: http://dx.doi.org/10.5852/ejt.2021.767.1485
03AEE5789D443E26777ED3EF7000FEDF.taxon	materials_examined	Type material of Geotrupes parvus Lectotype (here designated, Fig. 5 A – C) INDIA • ♂; “ Bengal Mai 1808 Parvus ♂ Wied. / Mus. Westerm. / TYPE ”; ZMUKK. Paralectotype INDIA • 1 ♂; “ Mus. Westerm. / TYPE ”; ZMUKK. Type material of Orphnus nanus Lectotype (here designated, Fig. 5 D – H) INDIA • 1 ♂; “ TYPE COL: 482 1 / 2 Orphnus nanus Westw. HOPE DEPT. OXFORD / Orphnus nanus Westw East Ind [...] Hearsey / TYPE WESTWOOD Trans. Ent. Soc. 4.1846. P. 177 Coll. Hope Oxon. / […] / W / LECTOTYPUS Orphnus nanus Westw. Frolov et al. 2016 ”; ZMUKK. Paralectotype INDIA • 1 ♂; “ TYPE COL: 482 2 / 2 Orphnus nanus Westw. HOPE DEPT. OXFORD / TYPE WESTWOOD Trans. Ent. Soc. 4.1846. P. 177 Coll. Hope Oxon. / Prof. Westwood’s private collection purchased from Miss Swann 1895 Collected by Gen. Hearsey in India / [9 … H] / Orphnus nanus Westw ”; ZMUKK. Additional material examined SRI LANKA • Central Province – 2 ♂♂; Kandy; 7 ° 18 ′ N, 80 ° 38 ′ E; MHNG • 1 ♀; Peradenyia; 7 ° 16 ′ N, 80 ° 35 ′ E; MHNG • 1 ♂, 1 ♀; same locality data as for preceding; Friederichs S. G. leg.; MNHB • 1 ♂; same locality data as for preceding; 3 Apr. 1907; O. John leg.; ZIN • 1 ♀; Sigiriya; 7 ° 57 ′ N, 80 ° 45 ′ E; Oct. 1977; MHNG • 2 ♀♀; Weragamtota; 7 ° 19 ′ N, 80 ° 59 ′ E; 13 Sep. 1953; F. Keiser leg.; NHMB. – North Central Province • 1 ♀; Anuradhapura; 8 ° 19 ′ N, 80 ° 24 ′ E; Friederichs S. G. leg.; MNHB • 1 ♂; Kala Oya; 8 ° 13 ′ N, 80 ° 06 ′ E; 7 Nov. 1983; MNHN • 1 ♀; same locality data as for preceding; 8 Nov. 1983; MNHN • 1 ♀; Vilpattu Nat. Park, Talawila; 8 ° 25 ′ N, 80 ° 03 ′ E; 9 Nov. 1983; G. S. Medvedev leg.; ZIN. – Southern Province • 1 ♀; 9 km SW of Yala; 6 ° 22 ′ N, 81 ° 31 ′ E; 21 Oct. 1982; V. F. Zaitzev leg.; ZIN • 8 ♂♂; Southern Province, Bentota; 6 ° 26 ′ N, 79 ° 60 ′ E; 23 Mar. 1973; G. Zimmermann leg.; MNHB • 1 ♂; Point de Galle; 6 ° 01 ′ N, 80 ° 13 ′ E; MHNG • 1 ♂; Tissamaharama; 6 ° 17 ′ N, 81 ° 17 ′ E; 8 Aug. 1978; P. Cabella leg.; MCSN • 2 ♂♂, 2 ♀♀; same locality data as for preceding; 30 Oct. 1983; MNHN • 2 ♂♂; Yala; 6 ° 22 ′ N, 81 ° 31 ′ E; 21 Oct. 1982; G. S. Medvedev leg.; ZIN. – Western Province • 1 ♂; Colombo; 6 ° 56 ′ N, 79 ° 51 ′ E; Friederichs S. G. leg.; MNHB • 1 ♂; same locality data as for preceding; Mar. 1953; G. Frey leg.; MHNG • 1 ♂, 1 ♀; Negombo; 7 ° 12 ′ N, 79 ° 50 ′ E; 23 Mar. 1973; B. Huttler leg.; NMPC • 1 ♀; Wadduwa; 6 ° 40 ′ N, 79 ° 56 ′ E; V. De Poll leg.; MNHN. – Sri Lanka (no exact locality) • 2 ♂♂, 1 ♀; OUM • 1 ♀; MHNG • 1 ♂; MNHB • 1 ♂; Friederichs S. G. leg.; MNHB • 1 ♂; Parry leg.; IRSNB • 1 ♂; S. Niethner leg.; MNHB.	en	Frolov, Andrey V., Akhmetova, Lilia A. (2021): Review of the Orphninae (Coleoptera: Scarabaeidae) of Sri Lanka, with description of a new species of genus Orphnus Macleay, 1819. European Journal of Taxonomy 767 (1): 40-54, DOI: https://doi.org/10.5852/ejt.2021.767.1485, URL: http://dx.doi.org/10.5852/ejt.2021.767.1485
03AEE5789D443E26777ED3EF7000FEDF.taxon	description	Variation Body length of the examined specimens varies from 6.1 to 8.2 mm (males) and 6.0 to 7.5 mm (females).	en	Frolov, Andrey V., Akhmetova, Lilia A. (2021): Review of the Orphninae (Coleoptera: Scarabaeidae) of Sri Lanka, with description of a new species of genus Orphnus Macleay, 1819. European Journal of Taxonomy 767 (1): 40-54, DOI: https://doi.org/10.5852/ejt.2021.767.1485, URL: http://dx.doi.org/10.5852/ejt.2021.767.1485
03AEE5789D443E26777ED3EF7000FEDF.taxon	distribution	Distribution The species is distributed rather widely throughout Sri Lanka except for the north-eastern part (Fig. 5 J).	en	Frolov, Andrey V., Akhmetova, Lilia A. (2021): Review of the Orphninae (Coleoptera: Scarabaeidae) of Sri Lanka, with description of a new species of genus Orphnus Macleay, 1819. European Journal of Taxonomy 767 (1): 40-54, DOI: https://doi.org/10.5852/ejt.2021.767.1485, URL: http://dx.doi.org/10.5852/ejt.2021.767.1485
