taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03AC87E7C329FFC49F70F9BF90D8BF13.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5514845/files/figure.png	http://doi.org/10.5281/zenodo.5514845	Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm.	Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm.	2021-09-17	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn		Zenodo	biologists	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn			
03AC87E7C329FFC49F70F9BF90D8BF13.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5514847/files/figure.png	http://doi.org/10.5281/zenodo.5514847	Fig. 4. Living adults of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov. B. L. tuberculata sp. nov. C. L. trochomorphoides sp. nov. D. L. chloritoides sp. nov. E. L. elegans sp. nov. F. L. diplogramma (Möllendorff, 1902) comb. nov. Scale bars = 10 mm.	Fig. 4. Living adults of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov. B. L. tuberculata sp. nov. C. L. trochomorphoides sp. nov. D. L. chloritoides sp. nov. E. L. elegans sp. nov. F. L. diplogramma (Möllendorff, 1902) comb. nov. Scale bars = 10 mm.	2021-09-17	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn		Zenodo	biologists	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn			
03AC87E7C329FFC49F70F9BF90D8BF13.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5514849/files/figure.png	http://doi.org/10.5281/zenodo.5514849	Fig. 5. Illustrations of the genital system of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., paratype (NHMSU-00024). B. L. tuberculata sp. nov., paratype (NHMSU-00026). C. L. trochomorphoides sp. nov., paratype (NHMSU-00028). D. L. chloritoides sp. nov., paratype (NHMSU-00030). E. L. elegans sp. nov., paratype (NHMSU-00033). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Internal sculpture of a = flagellum; b = penis; c = vagina. Scale bars = 10 mm.	Fig. 5. Illustrations of the genital system of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., paratype (NHMSU-00024). B. L. tuberculata sp. nov., paratype (NHMSU-00026). C. L. trochomorphoides sp. nov., paratype (NHMSU-00028). D. L. chloritoides sp. nov., paratype (NHMSU-00030). E. L. elegans sp. nov., paratype (NHMSU-00033). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Internal sculpture of a = flagellum; b = penis; c = vagina. Scale bars = 10 mm.	2021-09-17	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn		Zenodo	biologists	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn			
03AC87E7C329FFC49F70F9BF90D8BF13.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5514851/files/figure.png	http://doi.org/10.5281/zenodo.5514851	Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm.	Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm.	2021-09-17	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn		Zenodo	biologists	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn			
03AC87E7C325FFC29F7FFEE391E1BC45.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5514845/files/figure.png	http://doi.org/10.5281/zenodo.5514845	Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm.	Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm.	2021-09-17	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn		Zenodo	biologists	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn			
03AC87E7C325FFC29F7FFEE391E1BC45.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5514847/files/figure.png	http://doi.org/10.5281/zenodo.5514847	Fig. 4. Living adults of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov. B. L. tuberculata sp. nov. C. L. trochomorphoides sp. nov. D. L. chloritoides sp. nov. E. L. elegans sp. nov. F. L. diplogramma (Möllendorff, 1902) comb. nov. Scale bars = 10 mm.	Fig. 4. Living adults of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov. B. L. tuberculata sp. nov. C. L. trochomorphoides sp. nov. D. L. chloritoides sp. nov. E. L. elegans sp. nov. F. L. diplogramma (Möllendorff, 1902) comb. nov. Scale bars = 10 mm.	2021-09-17	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn		Zenodo	biologists	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn			
03AC87E7C325FFC29F7FFEE391E1BC45.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5514849/files/figure.png	http://doi.org/10.5281/zenodo.5514849	Fig. 5. Illustrations of the genital system of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., paratype (NHMSU-00024). B. L. tuberculata sp. nov., paratype (NHMSU-00026). C. L. trochomorphoides sp. nov., paratype (NHMSU-00028). D. L. chloritoides sp. nov., paratype (NHMSU-00030). E. L. elegans sp. nov., paratype (NHMSU-00033). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Internal sculpture of a = flagellum; b = penis; c = vagina. Scale bars = 10 mm.	Fig. 5. Illustrations of the genital system of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., paratype (NHMSU-00024). B. L. tuberculata sp. nov., paratype (NHMSU-00026). C. L. trochomorphoides sp. nov., paratype (NHMSU-00028). D. L. chloritoides sp. nov., paratype (NHMSU-00030). E. L. elegans sp. nov., paratype (NHMSU-00033). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Internal sculpture of a = flagellum; b = penis; c = vagina. Scale bars = 10 mm.	2021-09-17	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn		Zenodo	biologists	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn			
03AC87E7C325FFC29F7FFEE391E1BC45.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5514851/files/figure.png	http://doi.org/10.5281/zenodo.5514851	Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm.	Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm.	2021-09-17	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn		Zenodo	biologists	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn			
03AC87E7C323FFC19F28FDB5972BBC2E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5514845/files/figure.png	http://doi.org/10.5281/zenodo.5514845	Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm.	Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm.	2021-09-17	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn		Zenodo	biologists	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn			
03AC87E7C323FFC19F28FDB5972BBC2E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5514847/files/figure.png	http://doi.org/10.5281/zenodo.5514847	Fig. 4. Living adults of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov. B. L. tuberculata sp. nov. C. L. trochomorphoides sp. nov. D. L. chloritoides sp. nov. E. L. elegans sp. nov. F. L. diplogramma (Möllendorff, 1902) comb. nov. Scale bars = 10 mm.	Fig. 4. Living adults of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov. B. L. tuberculata sp. nov. C. L. trochomorphoides sp. nov. D. L. chloritoides sp. nov. E. L. elegans sp. nov. F. L. diplogramma (Möllendorff, 1902) comb. nov. Scale bars = 10 mm.	2021-09-17	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn		Zenodo	biologists	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn			
03AC87E7C323FFC19F28FDB5972BBC2E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5514849/files/figure.png	http://doi.org/10.5281/zenodo.5514849	Fig. 5. Illustrations of the genital system of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., paratype (NHMSU-00024). B. L. tuberculata sp. nov., paratype (NHMSU-00026). C. L. trochomorphoides sp. nov., paratype (NHMSU-00028). D. L. chloritoides sp. nov., paratype (NHMSU-00030). E. L. elegans sp. nov., paratype (NHMSU-00033). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Internal sculpture of a = flagellum; b = penis; c = vagina. Scale bars = 10 mm.	Fig. 5. Illustrations of the genital system of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., paratype (NHMSU-00024). B. L. tuberculata sp. nov., paratype (NHMSU-00026). C. L. trochomorphoides sp. nov., paratype (NHMSU-00028). D. L. chloritoides sp. nov., paratype (NHMSU-00030). E. L. elegans sp. nov., paratype (NHMSU-00033). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Internal sculpture of a = flagellum; b = penis; c = vagina. Scale bars = 10 mm.	2021-09-17	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn		Zenodo	biologists	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn			
03AC87E7C323FFC19F28FDB5972BBC2E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5514851/files/figure.png	http://doi.org/10.5281/zenodo.5514851	Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm.	Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm.	2021-09-17	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn		Zenodo	biologists	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn			
03AC87E7C320FFDF9F41FD0890C2BA73.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5514845/files/figure.png	http://doi.org/10.5281/zenodo.5514845	Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm.	Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm.	2021-09-17	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn		Zenodo	biologists	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn			
03AC87E7C320FFDF9F41FD0890C2BA73.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5514847/files/figure.png	http://doi.org/10.5281/zenodo.5514847	Fig. 4. Living adults of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov. B. L. tuberculata sp. nov. C. L. trochomorphoides sp. nov. D. L. chloritoides sp. nov. E. L. elegans sp. nov. F. L. diplogramma (Möllendorff, 1902) comb. nov. Scale bars = 10 mm.	Fig. 4. Living adults of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov. B. L. tuberculata sp. nov. C. L. trochomorphoides sp. nov. D. L. chloritoides sp. nov. E. L. elegans sp. nov. F. L. diplogramma (Möllendorff, 1902) comb. nov. Scale bars = 10 mm.	2021-09-17	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn		Zenodo	biologists	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn			
03AC87E7C320FFDF9F41FD0890C2BA73.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5514849/files/figure.png	http://doi.org/10.5281/zenodo.5514849	Fig. 5. Illustrations of the genital system of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., paratype (NHMSU-00024). B. L. tuberculata sp. nov., paratype (NHMSU-00026). C. L. trochomorphoides sp. nov., paratype (NHMSU-00028). D. L. chloritoides sp. nov., paratype (NHMSU-00030). E. L. elegans sp. nov., paratype (NHMSU-00033). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Internal sculpture of a = flagellum; b = penis; c = vagina. Scale bars = 10 mm.	Fig. 5. Illustrations of the genital system of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., paratype (NHMSU-00024). B. L. tuberculata sp. nov., paratype (NHMSU-00026). C. L. trochomorphoides sp. nov., paratype (NHMSU-00028). D. L. chloritoides sp. nov., paratype (NHMSU-00030). E. L. elegans sp. nov., paratype (NHMSU-00033). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Internal sculpture of a = flagellum; b = penis; c = vagina. Scale bars = 10 mm.	2021-09-17	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn		Zenodo	biologists	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn			
03AC87E7C320FFDF9F41FD0890C2BA73.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5514851/files/figure.png	http://doi.org/10.5281/zenodo.5514851	Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm.	Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm.	2021-09-17	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn		Zenodo	biologists	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn			
03AC87E7C33EFFDE9F68FBC390AFBB6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5514845/files/figure.png	http://doi.org/10.5281/zenodo.5514845	Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm.	Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm.	2021-09-17	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn		Zenodo	biologists	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn			
03AC87E7C33EFFDE9F68FBC390AFBB6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5514847/files/figure.png	http://doi.org/10.5281/zenodo.5514847	Fig. 4. Living adults of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov. B. L. tuberculata sp. nov. C. L. trochomorphoides sp. nov. D. L. chloritoides sp. nov. E. L. elegans sp. nov. F. L. diplogramma (Möllendorff, 1902) comb. nov. Scale bars = 10 mm.	Fig. 4. Living adults of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov. B. L. tuberculata sp. nov. C. L. trochomorphoides sp. nov. D. L. chloritoides sp. nov. E. L. elegans sp. nov. F. L. diplogramma (Möllendorff, 1902) comb. nov. Scale bars = 10 mm.	2021-09-17	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn		Zenodo	biologists	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn			
03AC87E7C33EFFDE9F68FBC390AFBB6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5514849/files/figure.png	http://doi.org/10.5281/zenodo.5514849	Fig. 5. Illustrations of the genital system of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., paratype (NHMSU-00024). B. L. tuberculata sp. nov., paratype (NHMSU-00026). C. L. trochomorphoides sp. nov., paratype (NHMSU-00028). D. L. chloritoides sp. nov., paratype (NHMSU-00030). E. L. elegans sp. nov., paratype (NHMSU-00033). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Internal sculpture of a = flagellum; b = penis; c = vagina. Scale bars = 10 mm.	Fig. 5. Illustrations of the genital system of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., paratype (NHMSU-00024). B. L. tuberculata sp. nov., paratype (NHMSU-00026). C. L. trochomorphoides sp. nov., paratype (NHMSU-00028). D. L. chloritoides sp. nov., paratype (NHMSU-00030). E. L. elegans sp. nov., paratype (NHMSU-00033). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Internal sculpture of a = flagellum; b = penis; c = vagina. Scale bars = 10 mm.	2021-09-17	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn		Zenodo	biologists	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn			
03AC87E7C33EFFDE9F68FBC390AFBB6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5514851/files/figure.png	http://doi.org/10.5281/zenodo.5514851	Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm.	Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm.	2021-09-17	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn		Zenodo	biologists	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn			
03AC87E7C33FFFDC9CD8FACF9704B918.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5514845/files/figure.png	http://doi.org/10.5281/zenodo.5514845	Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm.	Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm.	2021-09-17	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn		Zenodo	biologists	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn			
03AC87E7C33FFFDC9CD8FACF9704B918.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5514847/files/figure.png	http://doi.org/10.5281/zenodo.5514847	Fig. 4. Living adults of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov. B. L. tuberculata sp. nov. C. L. trochomorphoides sp. nov. D. L. chloritoides sp. nov. E. L. elegans sp. nov. F. L. diplogramma (Möllendorff, 1902) comb. nov. Scale bars = 10 mm.	Fig. 4. Living adults of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov. B. L. tuberculata sp. nov. C. L. trochomorphoides sp. nov. D. L. chloritoides sp. nov. E. L. elegans sp. nov. F. L. diplogramma (Möllendorff, 1902) comb. nov. Scale bars = 10 mm.	2021-09-17	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn		Zenodo	biologists	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn			
03AC87E7C33FFFDC9CD8FACF9704B918.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5514849/files/figure.png	http://doi.org/10.5281/zenodo.5514849	Fig. 5. Illustrations of the genital system of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., paratype (NHMSU-00024). B. L. tuberculata sp. nov., paratype (NHMSU-00026). C. L. trochomorphoides sp. nov., paratype (NHMSU-00028). D. L. chloritoides sp. nov., paratype (NHMSU-00030). E. L. elegans sp. nov., paratype (NHMSU-00033). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Internal sculpture of a = flagellum; b = penis; c = vagina. Scale bars = 10 mm.	Fig. 5. Illustrations of the genital system of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., paratype (NHMSU-00024). B. L. tuberculata sp. nov., paratype (NHMSU-00026). C. L. trochomorphoides sp. nov., paratype (NHMSU-00028). D. L. chloritoides sp. nov., paratype (NHMSU-00030). E. L. elegans sp. nov., paratype (NHMSU-00033). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Internal sculpture of a = flagellum; b = penis; c = vagina. Scale bars = 10 mm.	2021-09-17	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn		Zenodo	biologists	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn			
03AC87E7C33FFFDC9CD8FACF9704B918.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5514851/files/figure.png	http://doi.org/10.5281/zenodo.5514851	Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm.	Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm.	2021-09-17	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn		Zenodo	biologists	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn			
