identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03AC87E7C329FFC89F35FCF39747BBA4.text	03AC87E7C329FFC89F35FCF39747BBA4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Landouria Godwin-Austen 1918	<div><p>Genus Landouria Godwin-Austen, 1918</p> <p>Type species</p> <p>Helix huttonii L. Pfeiffer, 1842 (new name for Helix orbicula Hutton &amp; Benson, 1838), by original designation.</p> <p>Type locality</p> <p>Himalaya near Simla, Mahasu, northern India (Hutton &amp; Benson 1838).</p> <p>Distribution</p> <p>The genus is also reported from northeast India, west Himalaya, Nepal, Mandalay and Shan in Myanmar (Burma), Yunnan in China (Zilch 1966; Schileyko &amp; Kuznetsov 1998; Ramakrishna et al. 2010; Budha et al. 2015).</p> <p>Schileyko &amp; Kuznetsov (1998) tentatively argued that Landouria is restricted to northern India, Nepal, northern Myanmar, and Sri Lanka. Yet, recently, species of Landouria have been reported from China (Páll-Gergely et al. 2013), Laos (Hirano et al. 2014), Timor-Leste (Köhler et al. 2018), Java in Indonesia (Nurinsiyah et al. 2019), and Thailand (Tumpeesuwan &amp; Tumpeesuwan 2019).</p> </div>	http://treatment.plazi.org/id/03AC87E7C329FFC89F35FCF39747BBA4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn	Nahok, Benchawan, Tumpeesuwan, Sakboworn, Tumpeesuwan, Chanidaporn (2021): Five new species of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from northeastern Thailand, with note on genitalia and radula morphology of Landouria diplogramma (Möllendorff, 1902). European Journal of Taxonomy 767 (1): 142-166, DOI: https://doi.org/10.5852/ejt.2021.767.1495, URL: http://dx.doi.org/10.5852/ejt.2021.767.1495
03AC87E7C329FFC49F70F9BF90D8BF13.text	03AC87E7C329FFC49F70F9BF90D8BF13.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Landouria circinata Nahok & Tumpeesuwan & Tumpeesuwan 2021	<div><p>Landouria circinata sp. nov.</p> <p>urn:lsid:zoobank.org:act: B434E62B-9FA6-43FB-B27F-A4B1B0413225</p> <p>Figs 3A, 4A, 5A, 6A–C; Tables 1–4</p> <p>Landouria sp. – Tanmuangpak et al. 2012: 7, 12, fig. 5y. — Tanmuangpak 2016: 125–126, figs 4.74– 4.75.</p> <p>Landouria sp.4 – Ounchareon 2015: 40–42, figs 4.12, 4.13, 5.1, 5.7.</p> <p>Landouria sp.2 – Yingkhamhang 2016: 21–22, 39–41, 43–45, 47–48, figs 4.3–4.4, 4.21b, 4.22c, 4.23b, 5.1b, 5.2b.</p> <p>Diagnosis</p> <p>Shell brownish-corneous, periphery angular, sharply keeled. Flagellum circinate; penis cylindrical and longer than vagina; base of gametolytic sac thick and stout. Radula with unicuspid short tongue-shaped central teeth.</p> <p>Etymology</p> <p>The Latin word ‘ circinata ’ refers to the coiled or curled flagellum which resembles the circinate vernation of a fern shoot.</p> <p>Type material</p> <p>Holotype THAILAND • shell (SH = 8.2 mm, SW = 13.8 mm, AH = 5.1 mm, AW = 5.1 mm, UW = 3.8 mm); Loei Province, Mueang District, Phu Pha Lom Forest Park; 17°33′18.03″ N, 101°52′6.71″ E; alt. 415 m; 7 Jun. 2017; B. Nahok and U. Chanlabut leg.; NHMSU-00023. (Fig. 3A)</p> <p>Paratypes THAILAND • 11 shells, 43 living specs; same collection data as for holotype; GenBank No. MN449401 and MZ435746; NHMSU-00024.</p> <p>Description</p> <p>SHELL (Fig. 3A, Table 1). Dextral, medium sized, conical-lenticular, with six slightly convex whorls. Suture shallow. Apex obtuse, depressed and low-conical. Light brown at 4 early whorls, dark brown at 2 last whorls. Sharply keeled, with pale brown band on the peripheral keel. Umbilicus very deep and wide. Aperture lip somewhat thickened, rounded rhombic.</p> <p>GENITAL SYSTEM (n = 3) (Fig. 5A, Table 4). Penis very long, cylindrical, and curled, divided into two portions of equal length, internally with parallel, transverse folds, containing a short, rather rounded verge. Epiphallus short cylindrical, thinner than the penis. Flagellum slightly crenated and curled up, resembling the circinate vernation of a fern shoot, with thin grooves internally and tubercles on its surface at the end, internally with transverse robust folds. Vas deferens is a slender, cylindrical tube, thicker at the end where it connects to free oviduct; entering flagellum basally. Vagina less than half as long as penis, internally with four thin longitudinal pilasters. Free oviduct shorter than vagina. Gametolytic sac thickened at base, stout and gradually tapering distally to form a slender distal tube ending in a balloonshaped, medium-sized sac. Prostate gland very long, uterus long and swollen.</p> <p>RADULA (n = 3). Comprises 95 transverse rows with 57–61 teeth per row; radular formula: (20)-(8–10)- 1-(8–10)-(20). Central tooth symmetric, unicuspid, tongue-shaped; lateral teeth longer than central tooth and oblique, unicuspid, tongue-shaped (Fig. 6A). Marginal teeth tricuspid with medium-sized endocone; mesocone rather large, with rounded cusp; ectocone triangular (Fig. 6B–C).</p> <p>Remarks</p> <p>Landouria circinata sp. nov. resembles L. strobiloides (see Tumpeesuwan &amp; Tumpeesuwan 2019: fig. 3) in shell morphology, L. circinata sp. nov. has a smaller shell, a shallower suture, and darker peripheral zone than L. strobiloides (Table 4, Fig. 3A). Its central and lateral radular teeth are unicuspid and tongueshaped, whereas they are lanceolate in L. strobiloides (Table 4, Fig. 6A). The unique characters of L. circinata sp. nov. are its circinate flagellum and its penis that is much longer than the vagina (Fig. 5A), whereas L. strobiloides has a strobilus-like flagellum and a penis that is shorter than the vagina.</p> <p>Distribution</p> <p>Landouria circinata sp. nov. is known only from the type locality.</p> </div>	http://treatment.plazi.org/id/03AC87E7C329FFC49F70F9BF90D8BF13	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn	Nahok, Benchawan, Tumpeesuwan, Sakboworn, Tumpeesuwan, Chanidaporn (2021): Five new species of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from northeastern Thailand, with note on genitalia and radula morphology of Landouria diplogramma (Möllendorff, 1902). European Journal of Taxonomy 767 (1): 142-166, DOI: https://doi.org/10.5852/ejt.2021.767.1495, URL: http://dx.doi.org/10.5852/ejt.2021.767.1495
03AC87E7C325FFC29F7FFEE391E1BC45.text	03AC87E7C325FFC29F7FFEE391E1BC45.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Landouria tuberculata Nahok & Tumpeesuwan & Tumpeesuwan 2021	<div><p>Landouria tuberculata sp. nov.</p> <p>urn:lsid:zoobank.org:act: 84133944-D5C9-49A3-9997-8E1C7B335938</p> <p>Figs 3B, 4B, 5B, 6D–F; Tables 1–4</p> <p>Diagnosis</p> <p>Shell moderately keeled, with numerous tiny tubercles on shell surface (Fig. 4B). Flagellum like a short protrusion with a pointed tip, penis swollen, base of gametolytic sac narrow. Radula with slender, lanceolate central teeth.</p> <p>Etymology</p> <p>The specific epithet ‘ tuberculata ’ refers to the shell sculpture consisting of numerous tiny tubercles (Fig. 4B).</p> <p>Type material</p> <p>Holotype THAILAND • shell (SH = 9.6 mm, SW = 14.4 mm, AH = 5.2 mm, AW = 5.5 mm, UW = 3.5 mm); Loei Province, Phu Rua District, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.27819&amp;materialsCitation.latitude=17.437525" title="Search Plazi for locations around (long 101.27819/lat 17.437525)">Wat Thepnimit</a>; 17°26′15.09″ N, 101°16′41.46″ E; alt. 647 m; 23 Sep. 2017; B. Nahok, K. Tanmuangpak and U. Chanlabut leg.; NHMSU-00025. (Fig. 3B)</p> <p>Paratypes THAILAND • 10 shells, 7 living specs; same collection data as for holotype; GenBank No. MN449402 and MZ435747; NHMSU-00026.</p> <p>Description</p> <p>SHELL (Fig. 3B, Table 1). Dextral, medium-sized, tightly coiled, with 6½ slightly convex whorls. Suture rather shallow, apex obtuse with high conical spire. Body whorl angular, with tubercles all over its surface, without distinct incised spiral lines. Colour red brown in living specimens (Fig. 4B), empty shells are paler (Fig. 3B). Umbilicus deep and wide. Aperture obliquely oval, reflexed, and slightly thickened.</p> <p>GENITAL SYSTEM (n = 3) (Fig. 5B, Table 4). Penis cylindrical, swollen in the middle part, internally with rather wavy, corrugated longitudinal pilasters and a short, hollow verge with 2–3 lobes around its opening. Epiphallus evenly cylindrical, as long as penis. Flagellum like a short protrusion with a pointed tip, without nodes, internally with longitudinal slender pilasters. Vas deferens cylindrical, narrow, entering in the lower half of the flagellum. Vagina relatively short, internally with five corrugated longitudinal pilasters. Free oviduct very short. Gametolytic sac slightly thicker at base, with a long, narrow and thin cylindrical tube, and at its distal end a small, swollen spherical sac. Prostate gland very long. Uterus long and thin.</p> <p>RADULA (n = 3). Comprises 126–130 transverse rows with 79–87 teeth per row; radular formula: (28– 30)-(11–13)-1-(11–13)-(28–30). Central tooth symmetric, slender lanceolate, and unicuspid (Fig. 6D). Lateral teeth larger than the central tooth, lanceolate, bicuspid, with large endocone and sharp cusp (Fig. 6D). Marginal teeth tricuspid, with small sharp endocone; large, slender mesocone; and small v-shaped ectocone with two cusps (Fig. 6E–F).</p> <p>Remarks</p> <p>Landouria tuberculata sp. nov. differs from other Thai species of Landouria, by the shell sculpture of numerous tiny tubercles (Fig. 4B). Lateral teeth of radula bicuspid and lanceolate (Fig. 6D–E).</p> <p>Distribution</p> <p>Landouria tuberculata sp. nov. is currently only known from the area surrounding Phu Rua National Park, Phu Rua District, Loei Province, Thailand.</p> </div>	http://treatment.plazi.org/id/03AC87E7C325FFC29F7FFEE391E1BC45	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn	Nahok, Benchawan, Tumpeesuwan, Sakboworn, Tumpeesuwan, Chanidaporn (2021): Five new species of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from northeastern Thailand, with note on genitalia and radula morphology of Landouria diplogramma (Möllendorff, 1902). European Journal of Taxonomy 767 (1): 142-166, DOI: https://doi.org/10.5852/ejt.2021.767.1495, URL: http://dx.doi.org/10.5852/ejt.2021.767.1495
03AC87E7C323FFC19F28FDB5972BBC2E.text	03AC87E7C323FFC19F28FDB5972BBC2E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Landouria trochomorphoides Nahok & Tumpeesuwan & Tumpeesuwan 2021	<div><p>Landouria trochomorphoides sp. nov.</p> <p>urn:lsid:zoobank.org:act: B7DBFB2E-E7BF-46E5-AFB5-1B45CA4072DE</p> <p>Figs 3C, 4C, 5C, 6G–I; Tables 1–4</p> <p>Diagnosis</p> <p>Shell with rather low spire, sharply keeled, with radial scaly processes (Fig. 4C). Basal portion of flagellum thickened, tapering towards tip; penis narrow and cylindrical; base of gametolytic sac narrow. Radula, with lanceolate central and lateral teeth.</p> <p>Etymology</p> <p>The specific epithet ‘ trochomorphoides ’ refers to the conchological similarity between the shell of the new species and that of the land snail genus Trochomorpha Albers, 1850 (Trochomorphidae Möllendorf, 1890).</p> <p>Type material</p> <p>Holotype THAILAND • shell (SH = 10.0 mm, SW = 21.3 mm, AH = 7.4 mm, AW = 9.1 mm, UW = 6.2 mm); Loei Province, Pha Khao District, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.93433&amp;materialsCitation.latitude=17.119005" title="Search Plazi for locations around (long 101.93433/lat 17.119005)">Pha Sawan Cave</a>; 17°7′8.42″ N, 101°56′3.59″ E; alt. 359 m; 6 Oct. 2017; B. Nahok, S. Tumpeesuwan and C. Tumpeesuwan leg.; NHMSU-00027. (Fig. 3C)</p> <p>Paratypes THAILAND • 17 shells, 9 living specs; same collection data as for holotype; GenBank No. MN449403 and MZ435748; NHMSU-00028.</p> <p>Description</p> <p>SHELL (Fig. 3C, Table 1). Dextral, relatively large-sized. Whorls six, suture shallow, spire only slightly elevated. Protoconch with radially elongated scars. Body whorl sharply keeled, brownish-corneous with a brown zone at periphery, scaly processes all over shell surface (Fig. 4C). Umbilicus moderately deep and very wide. Aperture lip rhombic, peristome reflexed, and expanded.</p> <p>GENITAL SYSTEM (n = 3) (Fig. 5C, Table 4). Penis slightly longer than vagina, slender cylindrical, internally with parallel, transverse robust folds; verge absent. Epiphallus slender cylindrical with thickened end. Flagellum as long as epiphallus, basal portion thick and progressively tapering towards tip, without nodes, inner surface smooth. Vas deferens is a thin cylindrical tube, entering epiphallus apically. Vagina thinner than penis, internally with parallel, transverse robust folds. Free oviduct very short. Gametolytic sac slightly narrower at base, with a long narrow cylindrical tube, and a small, oval sac at the distal end. Prostate gland and uterus very long.</p> <p>RADULA (n = 3). Comprises 107–111 transverse rows with 65–73 teeth per row; radular formula: (24– 26)-(8–10)-1-(8–10)-(24–26). Central tooth unicuspid triangular (Fig. 6G). Lateral teeth short and broad leaf-shaped, bicuspid, larger than central tooth, with small ectocone (Fig. 6G–H). Marginal teeth asymmetric, tricuspid with small endocone; mesocone large and long, with curved margins; ectocone triangular with two to four cusps (Fig. 6H–I).</p> <p>Remarks</p> <p>Landouria trochomorphoides sp. nov. differs from other taxa by its very large, rather flat, sharply keeled shell with scaly surface (Figs 3C, 4C). Lateral teeth of radula are stout bicuspid (Fig. 6G–I), genital system is small and slim (Fig. 5C).</p> <p>Distribution</p> <p>Landouria trochomorphoides sp. nov. is known only from the type locality.</p> </div>	http://treatment.plazi.org/id/03AC87E7C323FFC19F28FDB5972BBC2E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn	Nahok, Benchawan, Tumpeesuwan, Sakboworn, Tumpeesuwan, Chanidaporn (2021): Five new species of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from northeastern Thailand, with note on genitalia and radula morphology of Landouria diplogramma (Möllendorff, 1902). European Journal of Taxonomy 767 (1): 142-166, DOI: https://doi.org/10.5852/ejt.2021.767.1495, URL: http://dx.doi.org/10.5852/ejt.2021.767.1495
03AC87E7C320FFDF9F41FD0890C2BA73.text	03AC87E7C320FFDF9F41FD0890C2BA73.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Landouria chloritoides Nahok & Tumpeesuwan & Tumpeesuwan 2021	<div><p>Landouria chloritoides sp. nov.</p> <p>urn:lsid:zoobank.org:act: 740CA614-D17B-47A3-A719-7DFC7D2CE2D5</p> <p>Figs 3D, 4D, 5D, 6J–L; Tables 1–4</p> <p>Landouria sp.1 – Ounchareon 2015: 29–32, 44–50, 54–56, 59, 61, figs 4.2c, 4.3c, 4.4. 4.5i–l, 5.5, 5.7.</p> <p>Landouria sp.2 – Buddharaksa 2016: 37–38, 41–42, figs 4.19, 4.20, 5.1, 5.2.</p> <p>Landouria sp.4 – Yingkhamhang 2016: 25–26, 39–41, 43–45, 49–52, figs 4.7, 4.8, 4.21d, 4.22d, 5.3a, 5.4a.</p> <p>Diagnosis</p> <p>Shell shape similar to that of the genus Chloritis Beck, 1837 (Camaenidae), but with last whorl stout and usually slightly angular, peripheral keel blunt, suture deep, aperture oval and oblique. Flagellum small ovate, penis very large and stout compared to vagina; vagina and free oviduct short (Fig. 5D). Radula with lanceolate teeth.</p> <p>Etymology</p> <p>The specific epithet ‘ chloritoides ’ refers to the conchological similarity between the shell of the new species and that of the camaenid genus Chloritis.</p> <p>Type material</p> <p>Holotype THAILAND • shell (SH = 12.0 mm, SW = 19.0 mm, AH = 7.7 mm, AW = 7.3 mm, UW = 3.9 mm); Khon Kaen Province, Chum Phae District, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.96529&amp;materialsCitation.latitude=16.758055" title="Search Plazi for locations around (long 101.96529/lat 16.758055)">Wat Thep Udom Wanaram</a>; 16°45′29″ N, 101°57′55.03″ E; alt. 406 m; 9 Jun. 2017; B. Nahok, S. Tumpeesuwan and C. Tumpeesuwan leg.; NHMSU-00029. (Fig. 3D)</p> <p>Paratypes THAILAND • 8 shells, 14 living specs; same collection data as for holotype; GenBank No. MN449404 and MZ435749; NHMSU-00030.</p> <p>Other material examined</p> <p>THAILAND • 4 shells, 8 living specs; North East, Khon Kaen Province, Chum Phae District, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.9861&amp;materialsCitation.latitude=16.830496" title="Search Plazi for locations around (long 101.9861/lat 16.830496)">Pu Lup Cave</a>; 16°49′49.78″ N, 101°59′9.97″ E; alt. 336 m; NHMSU-00031.</p> <p>Description</p> <p>SHELL (Fig. 3D, Table 1). Dextral, somewhat depressed, relatively large-sized. Whorls 6½, suture deep, apex obtuse, and moderately depressed, with low conical spire. Body whorl rounded, slightly keeled, red-brown in living specimens (Fig. 4D), empty shells are paler (Fig. 3D). Umbilicus very deep and wide. Aperture almost circular, with slightly thickened and weakly expanded lip.</p> <p>GENITAL SYSTEM (n = 3) (Fig. 5D, Table 4). Penis very large and stout, longer than vagina, cylindrical, gradually tapering towards epiphallus, internally with six rather thick, corrugated longitudinal pilasters; opening of the short, grooved verge surrounded by 2–3 lobes. Epiphallus short and thick. Flagellum short, elongate ovate, without nodes, internally with three large longitudinal pilasters. Vas deferens long and slender, entering in the lower half of the flagellum. Vagina short, as long as free oviduct, internally with seven, corrugated longitudinal pilasteres varying in size. Gametolytic sac thickened at base, with a long narrow cylindrical tube, and a small oval sac at distal end. Uterus swollen, prostate gland long.</p> <p>RADULA (n = 3). Comprises 125–129 transverse rows with 75–83 teeth per row; radular formula: (26–28)- (11–13)-1-(11–13)-(26–28). Central tooth short lanceolate, symmetric, unicuspid (Fig. 6J). Lateral teeth large and elongated lanceolate (Fig. 6J). Marginal teeth lanceolate, gradually changing from unicuspid to bicuspid and finally tricuspid; endocone small; ectocone slightly oblique, with wide triangular notch and two to four lateral cusps (Fig. 6K–L).</p> <p>Remarks</p> <p>Landouria chloritoides sp. nov. can be distinguished from other species of Landouria in Thailand by its conchological similarity to the genus Chloritis and its much larger and stout penis (Fig. 5D). The radula of L. chloritoides sp. nov. is similar to that of L. tuberculata sp. nov. but differs by its unicuspid lateral teeth (Fig. 5J) vs the bicuspid lateral teeth of L. tuberculata sp. nov.</p> <p>Distribution</p> <p>Landouria chloritoides sp. nov. is currently known only from the limestone area around Phu Pha Man National Park, Chum Phae District, Khon Kaen Province, Thailand.</p> </div>	http://treatment.plazi.org/id/03AC87E7C320FFDF9F41FD0890C2BA73	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn	Nahok, Benchawan, Tumpeesuwan, Sakboworn, Tumpeesuwan, Chanidaporn (2021): Five new species of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from northeastern Thailand, with note on genitalia and radula morphology of Landouria diplogramma (Möllendorff, 1902). European Journal of Taxonomy 767 (1): 142-166, DOI: https://doi.org/10.5852/ejt.2021.767.1495, URL: http://dx.doi.org/10.5852/ejt.2021.767.1495
03AC87E7C33EFFDE9F68FBC390AFBB6F.text	03AC87E7C33EFFDE9F68FBC390AFBB6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Landouria elegans Nahok & Tumpeesuwan & Tumpeesuwan 2021	<div><p>Landouria elegans sp. nov.</p> <p>urn:lsid:zoobank.org:act: 6A33A2BA-7197-4C1E-AEB3-D7815F8A05AC</p> <p>Figs 3E, 4E, 5E, 6M–O; Tables 1–4</p> <p>Landouria sp.3 – Yingkhamhang 2016: 23–24, 39–45, 47–48, 61, 74, figs 4.5, 4.6, 4.21–4.23c, 5.1c, 5.2c.</p> <p>Diagnosis</p> <p>Shell similar to Landouria circinata sp. nov., but with a sharper and more prominent keel at the periphery and dark brown zones on the periphery, just above and below the keel. Flagellum droplet shaped, penis long and cylindrical, vagina and free oviduct very short. Radula with elongate tongue-shaped central and lateral teeth.</p> <p>Etymology</p> <p>The specific epithet ‘ elegans ’ (Latin for ‘magnificent’) refers to the beauty of the shell.</p> <p>Type material</p> <p>Holotype THAILAND • shell (SH 10.0 = mm, SW 17.7 = mm, AH = 6.4 mm, AW = 7.1 mm, UW = 3.6 mm); Khon Kaen Province, Phu Pha Man District, Phu (mountain) <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.80186&amp;materialsCitation.latitude=16.652927" title="Search Plazi for locations around (long 101.80186/lat 16.652927)">Hua Chang</a>; 16°39′10.54″ N, 101°48′6.67″ E; alt. 305 m; 3 May 2018; B. Nahok, S. Tumpeesuwan and C. Tumpeesuwan leg.; NHMSU-00032. (Fig. 3E)</p> <p>Paratypes THAILAND • 11 shells; 9 living specs; same collection data as for holotype; GenBank No. MN449408 and MZ435751; NHMSU-00033.</p> <p>Description</p> <p>SHELL (Fig. 3E, Table1). Dextral, conical, relatively large-sized.Whorls 6. Protoconch with radially elongated wrinkles. Suture rather shallow, apex obtuse depressed, with rather high conical spire. Last whorl with a sharp keel that is distinctly bent downwards. Colour brownish-corneous with a dark brown zone bordering the keel at the periphery. Umbilicus very deep and wide. Apertural lip thickened, rounded rhombic.</p> <p>GENITAL SYSTEM (n = 3) (Fig. 5E, Table 4). Penis longer than vagina, cylindrical, internally with four corrugated, longitudinal pilasters; verge absent. Epiphallus shorter than penis, distal part (ep2) small, whereas proximal part (ep1) is swollen. Flagellum droplet-shaped, without nodes, internally with four corrugated, longitudinal pilasters. Vas deferens long, cylindrical, proximal part that connects to free oviduct somewhat thicker than distal part, entering the flagellum at its base. Vagina very short, internally with five corrugated longitudinal pilasters. Free oviduct shorter than vagina. Proximal part of gametolytic sac rather stout, but tapering to become a slender tube with large oval sac at distal end. Uterus and prostate gland very long and large.</p> <p>RADULA (n = 3). Comprises 110–114 transverse rows with 69–77 teeth per row; radular formula: (22–24)- (12–14)-1-(12–14)-(22–24). Central and lateral teeth unicuspid, tongue-shaped. Central tooth smaller and shorter than lateral teeth (Fig. 6M–N). Marginal teeth asymmetric, tricuspid with small endocone; mesocone large with curved margins; ectocone triangular and located at tooth base (Fig. 6N–O).</p> <p>Remarks</p> <p>Landouria elegans sp. nov. is conchologically quite similar to L. circinata sp. nov. and L. strobiloides, from which it differs by its sharper and more distinct keel with its downward bent rim (Fig. 3A vs Fig. 3E). The radula of L. elegans sp. nov. resembles that of L. circinata sp. nov., but central tooth and lateral teeth of L. elegans sp. nov. are longer and more slender than in L. circinata sp. nov. (Fig. 6A–C vs Fig. 6M–O). Epiphallus of L. elegans sp. nov. is similar to that of L. chloritoides sp. nov., but in L. elegans sp. nov. it is larger and connects with the vas deferens near the flagellum base (Fig. 5D vs Fig. 5E).</p> <p>Distribution</p> <p>Landouria elegans sp. nov. is known only from the type locality.</p> </div>	http://treatment.plazi.org/id/03AC87E7C33EFFDE9F68FBC390AFBB6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn	Nahok, Benchawan, Tumpeesuwan, Sakboworn, Tumpeesuwan, Chanidaporn (2021): Five new species of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from northeastern Thailand, with note on genitalia and radula morphology of Landouria diplogramma (Möllendorff, 1902). European Journal of Taxonomy 767 (1): 142-166, DOI: https://doi.org/10.5852/ejt.2021.767.1495, URL: http://dx.doi.org/10.5852/ejt.2021.767.1495
03AC87E7C33FFFDC9CD8FACF9704B918.text	03AC87E7C33FFFDC9CD8FACF9704B918.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Landouria diplogramma (Mollendorff 1902) Nahok & Tumpeesuwan & Tumpeesuwan 2021	<div><p>Landouria diplogramma (Möllendorff, 1902) comb. nov.</p> <p>Figs 3F, 4F, 5F, 6P–R; Tables 1–4</p> <p>Plectotropis diplogramme Möllendorff, 1902: 158–159.</p> <p>Ganesella diplogramme – Panha 1996: 36.</p> <p>Aegista (Plectotropis) diplogramma – Hemmen &amp; Hemmen 2001: 47.</p> <p>Thaitropis sp. – Sutcharit et al. 2017: 249, fig. 6–32e.</p> <p>Diagnosis</p> <p>Shell small, low-conical, weakly keeled, with a slightly elevated spire, whitish corneous with a pale brown band above periphery. Flagellum like a short protrusion, with rounded tip. Penis short swollen. Central and the first few lateral radular teeth triangular.</p> <p>Material examined</p> <p>THAILAND • 29 shells, 14 living specs; Nakhon Ratchasima Province, Pak Chong District, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.26535&amp;materialsCitation.latitude=14.615159" title="Search Plazi for locations around (long 101.26535/lat 14.615159)">Wat Thep Phithak Punnaram</a>; 14°36′54.57″ N, 101°15′55.25″ E; alt. 451 m; 23 Jun. 2017; B. Nahok and U. Chanlabut leg.; GenBank No. MN449411 and MZ435752; NHMSU-00034. (Fig. 3F)</p> <p>Description</p> <p>SHELL (Fig. 3F, Table 1). Dextral, small, depressed conical. Whorls 5½, suture rather shallow, apex obtuse and depressed, whitish corneous, with a pale brown peripheral band. Protoconch almost smooth. Umbilicus moderately deep and wide. Aperture oval oblique, with thin, solid, and weakly reflected lip.</p> <p>GENITAL SYSTEM (n = 3) (Fig. 5F, Table 4). Penis divided into two short portions, the portion that connects to epiphallus is swollen and the portion that connects to atrium is constricted; internally with three corrugated longitudinal pilasters; with short, rather rounded verge. Flagellum like a short protrusion, internally with two narrow longitudinal pilasters. Vas deferens long, cylindrical, entering the epiphallus at base. Free oviduct very short. Vagina twice as long as free oviduct, internally with five thick, corrugated longitudinal pilasters. Proximal part of gametolytic sac slightly thicker than the cylindrical tube, with a medium-sized oval sax at distal end. Uterus and prostate gland long.</p> <p>RADULA (n = 3). Comprises 121–125 transverse rows with 61–69 teeth per row; radular formula: (23–25)- (7–9)-1-(7–9)-(23–25). Central tooth usually symmetric, unicuspid, triangular (Fig. 6P). Lateral teeth longer and larger than central teeth, bicuspid, ectocone small, with sharp cusp (Fig. 6Q). Marginal teeth gradually changing from bicuspid to tricuspid, small endocone; mesocone large lanceolate; ectocone triangular with two to four tiny cusps (Fig. 6R).</p> <p>Remarks</p> <p>Specimens deposited in the Leiden Museum, The Netherlands, RMNH.MOL.309867 and RMNH. MOL.309851 (https://images.app.goo.gl/PgrpqKaWuSrgjdvN9), are identified as Plectotropis diplogramme Möllendorff, 1902. These specimens appear identical to specimens from Khao See Siad Ah in Nakhon Ratchasima Province, therefore we used the name “ Landouria diplogramma (Möllendorff, 1902) ” for this species in the present study. Landouria diplogramma differs from other Thai species of Landouria by its relatively small lustrous shell without peripheral keel, but with a pale brown band present above periphery (Figs 3F, 4F). Its radular teeth are relatively short (Fig. 6P–R). Its penis and flagellum are relatively short (Fig. 5F).</p> <p>The genus Thaitropis Schileyko, 2004 was recently reported in Thailand (Schileyko 2004; Sutcharit et al. 2017). Yet, this genus was synonymized with Landouria by Nurinsiyah et al. (2019). Based on the illustration provided by Sutcharit et al. (2017: fig. 6–32e), we identify their Thai Thaitropis sp. as L. diplogramma, because it has the same shell morphology and colours: Plectotropis diplogramme Möllendorff, 1902, RMNH.MOL.309867, and L. diplogramma (Möllendorff, 1902) from Khao See Siad Ah, Nakhon Ratchasima (NHMSU-00034) (Figs 3F, 4F). Unfortunately, the whereabouts of the type specimen of Plectotropis diplogramme are unknown. Yet, it might be deposited in the collections of the Senckenberg Museum in Frankfurt, Germany where it should be looked up and checked in a further study.</p> <p>Distribution</p> <p>Landouria diplogramma was collected by local people in the mountain range known as Khao See Siad Ah, Pak Chong District, Nakhon Ratchasima Province, Thailand. Other reported localities comprise Bangkok, Hinlap, Muoklek, Kanburi (Kanchanaburi) (Möllendorff 1902).</p> </div>	http://treatment.plazi.org/id/03AC87E7C33FFFDC9CD8FACF9704B918	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Nahok, Benchawan;Tumpeesuwan, Sakboworn;Tumpeesuwan, Chanidaporn	Nahok, Benchawan, Tumpeesuwan, Sakboworn, Tumpeesuwan, Chanidaporn (2021): Five new species of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from northeastern Thailand, with note on genitalia and radula morphology of Landouria diplogramma (Möllendorff, 1902). European Journal of Taxonomy 767 (1): 142-166, DOI: https://doi.org/10.5852/ejt.2021.767.1495, URL: http://dx.doi.org/10.5852/ejt.2021.767.1495
