identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
3E0B87B0C175FFAAFA0873A5DD5AFBC7.text	3E0B87B0C175FFAAFA0873A5DD5AFBC7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Freddius Takeda & Kajihara 2018	<div><p>Genus Freddius gen. nov</p> <p>Diagnosis. Cheliplanids possess hooks (sensu Karling 1983) articulated with movable nails; paired lateral auxiliary apparatuses are present between hooks.</p> <p>Type species. Freddius tricaudatus sp. nov., fixed by original designation.</p> <p>Etymology. The new masculine genus name is derived from the character name Freddy Krueger, a horror icon in the movie A Nightmare on Elm Street, alluding the resemblance between his razor-armed glove used to kill his victims and the elaborate proboscis hooks of the flatworms in the new taxon.</p></div> 	http://treatment.plazi.org/id/3E0B87B0C175FFAAFA0873A5DD5AFBC7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Takeda, Naoya;Kajihara, Hiroshi	Takeda, Naoya, Kajihara, Hiroshi (2018): A New Genus and Five New Species of Kalyptorhynchia (Platyhelminthes: Rhabdocoela) Discovered in Northern Japan. Species Diversity 23 (1): 1-11, DOI: 10.12782/specdiv.23.1, URL: http://dx.doi.org/10.12782/specdiv.23.1
3E0B87B0C175FFA8FA7E75FCD945FF74.text	3E0B87B0C175FFA8FA7E75FCD945FF74.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Freddius tricaudatus Takeda & Kajihara 2018	<div><p>Freddius tricaudatus sp. nov.</p> <p>(Figs 2–5)</p> <p>Material examined. Holotype: ICHUM 4832, adult, whole mount, 43°01′16″N, 144°50′13″E, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=144.83694&amp;materialsCitation.latitude=43.02111" title="Search Plazi for locations around (long 144.83694/lat 43.02111)">Akkeshi</a>, Hokkaido, Japan, intertidal sand, 15 June 2012. Paratypes: ICHUM 4829–4831, three adults, whole mount, same data as holotype; ICHUM 4833–4835, three adults, fluorescent phalloidin stained, same data as holotype; ICHUM 4836–4838, three adults, whole mount, from type locality, 24 June 2013.</p> <p>Description. Living animal body approximately 900 µm long and 130 µm wide. Pair of hooks present, asymmetrical in structure (Figs 2, 3), one 12–14 µm and the other 10–11 µm in length, each 3–4 µm wide and semi-cylindrical in shape, anteriorly articulated with three movable nails, 9–11 µm in length (Fig. 3); longer hook possesses a pair of additional lateral nails, 12–16 µm long, which are (i) anteriorly directed, (ii) articulated at both sides of the hook near its posterior end, (iii) longer than the anterior movable nails, and (iv) equipped with a pair of spines at their midpoint (Fig. 3). Paired auxiliary apparatuses present between hooks, tapered rod in shape, 4 µm long (Fig. 3C, F). Proboscis gland 55 µm long, 20 µm wide, situated posterior to proboscis (Fig. 2). Eyes absent (Fig. 2). One testis 100 µm long, 30 µm wide, not divided into follicles, situated posterior to proboscis gland (Fig. 2). Multiple sacs of unknown function, arranged posterior to testis (Fig. 2). Pharynx 100 µm long, 50 µm wide, posterior to sacs (Fig. 2). Mouth opening anterior to pharynx (Fig. 4A). Male copulatory organ, posterior to pharynx (Figs 2, 4), roughly tubular; anterior part bulbous and muscular; posterior part with cone-shaped cirrus spines, 20 µm long, 10 µm wide (Figs 4B, 5). Genital pore opening on ventral side of body, posterior to cirrus spines (Fig. 4). Single ovary 50 µm long, 20 µm wide, situated posterior to genital pore (Fig. 2). Pair of yolk glands 300 µm long and 25 µm wide, situated posterior to ovary (Fig. 2). Posterior end of body trifurcated, with tail-shaped serial beads in the middle (Fig. 2).</p> <p>Etymology. The specific name refers to the three tails in the new species.</p> <p>Remarks. The new species is undoubtedly a member of Cheliplanidae; it possesses proboscis with side pieces and a postrostral bulb, without separate lateral gland sacs, one pair of proboscis hooks, and characteristics that are shared among all the members of the family (Karling 1983); in addition, the structure of the genital organs in this species is similar to that in other members of the family. In cheliplanids, the proboscis hooks consist of a symmetrical pair of simple, curved spines, which, however, are not elaborate enough to be articulated like in the new species. The morphology of the proboscis hook in this species is so peculiar that it cannot be classified with certainty into any of the existing four genera in the family (Tyler et al. 2006 –2013). Therefore, we propose to establish a new genus in the Cheliplanidae family to accommodate this new species.</p> </div>	http://treatment.plazi.org/id/3E0B87B0C175FFA8FA7E75FCD945FF74	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Takeda, Naoya;Kajihara, Hiroshi	Takeda, Naoya, Kajihara, Hiroshi (2018): A New Genus and Five New Species of Kalyptorhynchia (Platyhelminthes: Rhabdocoela) Discovered in Northern Japan. Species Diversity 23 (1): 1-11, DOI: 10.12782/specdiv.23.1, URL: http://dx.doi.org/10.12782/specdiv.23.1
3E0B87B0C177FFAEF8D67022DEAAFE0B.text	3E0B87B0C177FFAEF8D67022DEAAFE0B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Proschizorhynchella caudociliata Takeda & Kajihara 2018	<div><p>Proschizorhynchella caudociliata sp. nov.</p> <p>(Figs 6, 7; Table 1)</p> <p>Material examined. Holotype: ICHUM 4863, adult, whole mount, 42°33′25″N, 141°55′42″E, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.92834&amp;materialsCitation.latitude=42.556942" title="Search Plazi for locations around (long 141.92834/lat 42.556942)">Mukawa</a>, Hokkaido, Japan, intertidal sand, 20 May 2012. Paratypes: ICHUM 4262, 4264, 4265, three adults, whole mounts, same data as holotype.</p> <p>Description. Living animal body approximately 3.0 mm long and 0.3 mm wide (Fig. 6). Proboscis 230–270 µm long, 70–80 µm wide (Fig. 6). Pair of black eyes situated behind proboscis (Fig. 6). Two testes, each 150–280 µm in diameter (Fig. 6). Yolk gland 750 µm long, 110 µm wide (Fig. 6A). Two adhesive girdles present; anterior one located at level of posterior end of pharynx, posterior one near caudal end; each girdle composed of six adhesive papillae arranged in regular intervals (Fig. 6A). Pharynx 220 µm long, 150 µm wide (Fig. 6). Pair of seminal vesicles, each 150–310 µm long, 10–30 µm wide, located posterior to pharynx (Fig. 6A). Male copulatory organ 160 µm long, 50 µm wide (Fig. 6A). Stylet cone shaped, 45 µm long, 12 µm wide, comprised of thin sclerotic sheet rolled up four times (Figs 6A, 7). Ovary 120 µm long, 60 µm wide, located posterior to pharynx (Fig. 6A, B). Uterus, 90 µm long, 40 µm wide, located around ovary (Fig. 6A). Common genital pore opening on ventral side of body between two adhesive girdles (Fig. 6A). Bursa 190 µm long, 80 µm wide, located around common genital pore (Fig. 6A). Large oval structure (adhesive gland?) 120 µm long, 100 µm wide, located at the caudal end (Fig. 6A). Caudal end covered with long tactile cilia (Fig. 6A).</p> <p>Etymology. The specific name refers to the long tactile cilia on the caudal region, which look like a brush.</p> <p>Remarks. Proschizorhynchella caudociliata can be distinguished from all congeners, except Proschizorhynchella papillata (Doe, 1974) and P. shibazakii sp. nov., based on the characteristics listed in Table 1. These three species, however, can be distinguished on the basis of two characteristics: (i) four coils of the copulatory stylet in P. caudociliata and three in P. papillata and P. shibazakii and (ii) a vacuolated region and long tactile hair present near the caudal end in P. caudociliata but absent in P. papillata and P. shibazakii.</p> </div>	http://treatment.plazi.org/id/3E0B87B0C177FFAEF8D67022DEAAFE0B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Takeda, Naoya;Kajihara, Hiroshi	Takeda, Naoya, Kajihara, Hiroshi (2018): A New Genus and Five New Species of Kalyptorhynchia (Platyhelminthes: Rhabdocoela) Discovered in Northern Japan. Species Diversity 23 (1): 1-11, DOI: 10.12782/specdiv.23.1, URL: http://dx.doi.org/10.12782/specdiv.23.1
3E0B87B0C171FFACFA4470A0DE89FD2F.text	3E0B87B0C171FFACFA4470A0DE89FD2F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Proschizorhynchella magnoliae Takeda & Kajihara 2018	<div><p>Proschizorhynchella magnoliae sp. nov.</p> <p>(Figs 8, 9; Table 1)</p> <p>Material examined. Holotype: ICHUM 4859, adult, whole mount for fluorescent staining, 43°01′16″N, 144°50′13″E, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=144.83694&amp;materialsCitation.latitude=43.02111" title="Search Plazi for locations around (long 144.83694/lat 43.02111)">Akkeshi Bay</a>, Hokkaido, Japan, intertidal sand, 24 June 2013. Paratypes: ICHUM 4849, 4850, two adults, whole mounts, type locality, 1 July 2011; ICHUM 4851, 4852, two adults, whole mounts, type locality, 7 July 2011; ICHUM 4853–4857, five adults, whole mounts, type locality, 15 June 2012; ICHUM 4858, 4860, two adults, same data as holotype.</p> <p>Description. Living animal body approximately 1.5 mm long and 160 µm wide (Fig. 8). Proboscis 240–290 µm long, 30 µm wide; pair of proboscis glands 90 µm long, 50 µm wide (Fig. 8). Pair of black eyes situated anterior to brain (Fig. 8A, B, D). Two testes, each 90–150 µm in diameter (Fig. 8A). Pharynx 120 µm long, 170 µm wide (Fig. 8 A–C). Three adhesive girdles present; anterior one located at the level of posterior end of pharynx, middle one at level of seminal vesicles, posterior one near caudal end; each girdle composed of six adhesive papillae arranged in regular intervals (Fig. 8A). Pair of yolk glands 400–440 µm long, 120– 170 µm wide (Fig. 8A). Pair of seminal vesicles, each 220– 230 µm long, 20 µm wide, located posterior to pharynx (Figs 8A, 9A, B). Male copulatory organ tubular in shape, 120 µm long, 50 µm wide (Figs 8C, 9A), tapering toward its tip, equipped with stylet. Stylet cone shaped, 31 µm long, 13 µm wide, comprised of thin sclerotic sheets (exact number of sheets not clear but appears to be more than one), equipped with ridges and processes inside (Fig. 9C, D). Common genital pore opening on ventral side of posterior region of body, posteriorly leading to saccate common atrium, middle part of which is dorsally connected to male copulatory organ and pair of yolk glands (Fig. 9A, B). Ovary 80 µm long, 60 µm wide, connected to the end of common atrium (Fig. 9A, B).</p> <p>Etymology. The specific name is a noun in the genitive case, referring to the stylet of the new species, which resembles a flower of lily magnolia.</p> <p>Remarks. Proschizorhynchella magnoliae sp. nov. most closely resembles Proschizorhynchella helgolandica (L’Hardy, 1965), which they has (i) a pair of eyes, (ii) two testes, (iii) three adhesive girdles, (iv) a single genital pore, and (v) no cirrus spine (Table 1). In P. helgolandica, the copulatory stylet comprises a pair of elongated cuticular sheets that face each other, with a concave confronting surface; the tip of each sheet is sharply pointed. On the other hand, in P.magnoliae, the copulatory stylet takes the form of spiral sheets that are complicatedly coiled with each other, with an elaborate inner surface with numerous ridges and short spines.</p> </div>	http://treatment.plazi.org/id/3E0B87B0C171FFACFA4470A0DE89FD2F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Takeda, Naoya;Kajihara, Hiroshi	Takeda, Naoya, Kajihara, Hiroshi (2018): A New Genus and Five New Species of Kalyptorhynchia (Platyhelminthes: Rhabdocoela) Discovered in Northern Japan. Species Diversity 23 (1): 1-11, DOI: 10.12782/specdiv.23.1, URL: http://dx.doi.org/10.12782/specdiv.23.1
3E0B87B0C173FFA2FA4373C4DD88FE08.text	3E0B87B0C173FFA2FA4373C4DD88FE08.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Proschizorhynchella shibazakii Takeda & Kajihara 2018	<div><p>Proschizorhynchella shibazakii sp. nov.</p> <p>(Figs 10–14; Table 1)</p> <p>Material examined. Holotype: ICHUM 4275, adult, whole mount, 43°12′33″N, 140°51′31″E, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=140.85861&amp;materialsCitation.latitude=43.209167" title="Search Plazi for locations around (long 140.85861/lat 43.209167)">Oshoro</a>, Hokkaido, Japan, intertidal sand, 13 June 2011. Paratypes: ICHUM 4276–4278, three adults, whole mounts, same data as holotype; ICHUM 4279, 4280, two adults, serial sagittal sections, same data as holotype; ICHUM 4281, 4282, two adults, serial transverse sections, same data as holotype; ICHUM 4283, one adult, whole mount, type locality, 21 May 2012; ICHUM 4861, egg, whole mount, laid by animals collected on 1 July 2013.</p> <p>Description. Living animal body approximately 2.6 mm long and 0.5 mm wide (Figs 10, 11A). Four pairs of bristles located at slender anterior tip of body (Figs 10, 11C). Proboscis 350 µm long, 120 µm wide; pair of proboscis glands 140 µm long, 80 µm wide (Fig. 10). Pair of black eyes situated anterior to brain (Figs 10, 11C). Gut anteroposteriorly elongated. Two testes 300–330 µm in diameter (Figs 10, 11C). Pair of yolk glands 1 mm long, 180 µm wide (Figs 10, 11A). Pharynx 480 µm long, 330 µm wide (Figs 10, 11A). Two adhesive girdles present; anterior one located at level of posterior end of pharynx, posterior one near caudal end; each girdle comprised of six adhesive papillae arranged in regular intervals (Figs 10, 11B). Pair of seminal vesicles, each 620 µm long, 60 µm wide, located posterior to pharynx (Fig. 10). Male copulatory organ tubular in shape, 240 µm long, 30 µm wide, with ejaculatory duct surrounded by circular muscles and further surrounded by longitudinal muscles (Figs 10, 12A, B, 13); copulatory organ tapering toward its tip, equipped with stylet and situated in male genital canal. Stylet cone shaped, 29–31 µm long (31 µm in holotype), 7 µm wide, comprised of thin sclerotic sheet rolled up three times (Fig. 12C, D). Male genital canal opens to anterodorsal part of common atrium of the latter (Fig. 13). Uterus 90 µm long, 30 µm wide, anterior to common atrium (Figs 10, 13). Each yolk gland connected to each side of common atrium (Figs 10, 13). Common genital pore opening on ventral side of body between two adhesive girdles (Figs 10, 13). Ovary 110 µm long, 70 µm wide, anteriorly connected to posterodorsal portion of common atrium via a common oviduct (Figs 10, 13). Bursa oval in dorsal view, 250 µm long, 150 µm wide; bursal tissue divided into two (smaller anterior and larger posterior) parts by constriction; spermatids observed in posterior bursal tissue in all specimens observed; anterior bursal tissue leading forward to connect to common oviduct near ovary via narrow sperm duct (Figs 10, 13). Egg oval, 260 µm long, 200 µm wide, covered in brown shell with colorless axis (Fig. 14).</p> <p>Etymology. The specific name is a noun in the genitive case, derived from the name Mr. Kouji Shibazaki, a caretaker of Oshoro Marine Station, Hokkaido University.</p> <p>Remarks. Proschizorhynchella shibazakii can be distinguished from all congeners based on the characteristics listed in Table 1 except P. papillata. These two species, however, can be distinguished based on the shape of the male copulatory organ. The differences in morphological characteristics between P. shibazakii sp. nov. and P. papillata are (i) the number of the apical sensory bristles, which is eight in P. shibazakii and four in P. papillata; (ii) the male copulatory organ, which is narrow and tubular in P. shibazakii, and bulb shaped in P. papillata; (iii) the internal part of the circular muscles, which is thin in P. shibazakii but thick in P. papillata; (iv) the border cells which are present in P. shibazakii but absent in P. papillata; and (v) the length of the stylet, which is 29–31 µm (31 µm in holotype) in P. shibazakii and 55–57 µm in P. papillata. Proschizorhynchella shibazakii cannot be distinguished from P. caudociliata based on the characteristics listed in Table 1; however, they differ in the structure of the copulatory stylet (see Remarks for P. caudociliata).</p> </div>	http://treatment.plazi.org/id/3E0B87B0C173FFA2FA4373C4DD88FE08	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Takeda, Naoya;Kajihara, Hiroshi	Takeda, Naoya, Kajihara, Hiroshi (2018): A New Genus and Five New Species of Kalyptorhynchia (Platyhelminthes: Rhabdocoela) Discovered in Northern Japan. Species Diversity 23 (1): 1-11, DOI: 10.12782/specdiv.23.1, URL: http://dx.doi.org/10.12782/specdiv.23.1
3E0B87B0C17DFFA1F8DE71ACDE1AF9DF.text	3E0B87B0C17DFFA1F8DE71ACDE1AF9DF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Proschizorhynchella shuttlecock Takeda & Kajihara 2018	<div><p>Proschizorhynchella shuttlecock sp. nov.</p> <p>(Figs 15, 16; Table 1)</p> <p>Material examined. Holotype: ICHUM 4846, adult, whole mount, 44°03′03″N, 141°39′46″E, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.66277&amp;materialsCitation.latitude=44.05083" title="Search Plazi for locations around (long 141.66277/lat 44.05083)">Obira</a>, Hokkaido, Japan, intertidal sand, 25 May 2012. Paratypes: ICHUM 4845, adult, whole mount, 45°29′16″N, 141°58′05″E, Soya, Hokkaido, Japan, intertidal sand, 26 May 2012; ICHUM 4847, 4848, two adults, whole mounts, same data as holotype.</p> <p>Description. Living animal body approximately 2.6 mm long, 0.6 mm wide. Proboscis 600 µm long, 100 µm wide; pair of proboscis glands 170 µm long, 110 µm wide (Fig. 15 A–C). Pair of black eyes situated behind proboscis (Fig. 15 A–C). Two testes, each 200–290 µm in diameter (Fig. 15A, B). Pharynx typical for genus, 300 µm long, 400 µm wide (Fig. 15A, B). No adhesive girdles (Fig. 15A, B). Pair of seminal vesicles, each 410–470 µm long, 30–40 µm wide, located posterior to pharynx (Fig. 15A). Stylet badminton shuttlecock shaped, 68 µm long, comprised of incomplete tube of thin sclerotic sheet with single, narrow (~ 2 µm), longitudinal gap; proximal three-quarters cone shaped, 22 µm in diameter at its base, narrowing toward its tip, with many longitudinal ridges inside; distal quarter bulbous, slightly elongated along stylet axis, 14 µm in maximum diameter, with single, transverse constriction slightly behind its widest portion; numerous, minute, wart-like projections present inside distal bulbous region (Fig. 16A, B); cirrus spine absent. Ovary 100 µm long, 70 µm wide, located posterior to pharynx (Fig. 15A). Common atrium 110 µm in diameter, leading to genital pore opening on ventral side of body located near caudal end (Fig. 15A).</p> <p>Etymology. The specific name refers to the shape of the stylet in the new species.</p> <p>Remarks. The stylet in Proschizorhynchella shuttlecock sp. nov. is badminton shuttlecock shaped, which is unique among the congeners, which mostly show a conical stylet (Table 1). Proschizorhynchella shuttlecock lacks adhesive girdles and thus resembles Proschizorhynchella faroeensis (Steinböck, 1931) and Proschizorhynchella spiracirro (Schilke, 1970). The latter two possess cirrus spines (Steinböck 1931; Schilke 1970) but P. shuttlecock does not.</p> </div>	http://treatment.plazi.org/id/3E0B87B0C17DFFA1F8DE71ACDE1AF9DF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Takeda, Naoya;Kajihara, Hiroshi	Takeda, Naoya, Kajihara, Hiroshi (2018): A New Genus and Five New Species of Kalyptorhynchia (Platyhelminthes: Rhabdocoela) Discovered in Northern Japan. Species Diversity 23 (1): 1-11, DOI: 10.12782/specdiv.23.1, URL: http://dx.doi.org/10.12782/specdiv.23.1
