identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03B28787AB470678FCAF8558FA19F9B2.text	03B28787AB470678FCAF8558FA19F9B2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Idanotermitinae Engel 2021	<div><p>Idanotermitinae Engel, subfam. nov.</p> <p>Type genus. Idanotermes Engel, 2008.</p> <p>Diagnosis. Imago: Mastotermitids most notable for lacking ocelli, but otherwise encompassing the following additional features more typical of the family: fontanelle absent; occipital carina present; lateral cervical sclerites present but ventral cervical sclerite absent; procoxal ventral carina present; tarsi pentamerous. The genus Anisotermes Zhao et al., 2019 perhaps belongs within this group or basal among Mastotermitinae, although other characters are suggestive of a Meiatermes -grade taxon but these are likely plesiomorphies (Zhao et al., 2019).</p> </div>	http://treatment.plazi.org/id/03B28787AB470678FCAF8558FA19F9B2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	JIANG, RI-XIN;ZHANG, HONG-RUI;ELDREDGE, K. TARO;SONG, XIAO-BIN;LI, YAN-DA;TIHELKA, ERIK;HUANG, DI-YING;WANG, SHUO;ENGEL, MICHAEL S.;CAI, CHEN-YANG	JIANG, RI-XIN, ZHANG, HONG-RUI, ELDREDGE, K. TARO, SONG, XIAO-BIN, LI, YAN-DA, TIHELKA, ERIK, HUANG, DI-YING, WANG, SHUO, ENGEL, MICHAEL S., CAI, CHEN-YANG (2021): Further evidence of Cretaceous termitophily: Description of new termite hosts of the trichopseniine Cretotrichopsenius (Coleoptera: Staphylinidae), with emendations to the classification of lower termites (Isoptera). Palaeoentomology 4 (4): 374-389, DOI: 10.11646/palaeoentomology.4.4.13, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.13
03B28787AB44067AFCAF83F5FD80FF36.text	03B28787AB44067AFCAF83F5FD80FF36.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arceotermitidae Engel 2021	<div><p>Arceotermitidae Engel, fam. nov.</p> <p>Type genus. Arceotermes Engel &amp; Jiang, gen. nov.</p> <p>Diagnosis. The soldiers of this group can be distinguished from those of Mastotermitidae and Archotermopsidae by the tetramerous tarsi,smaller number of antennal articles (less than 20 articles), and absence of compound eyes (although still present and reduced in Cosmotermitinae Engel, subfam. nov., vide infra) (Figs 5, 6). The head is not dorsoventrally compressed like the soldier heads of Stolotermitidae (particularly Stolotermes Hagen,1858).Like Mastotermitidae, the pronotum is about as wide as the head capsule, while in Archotermopsidae, Hodotermitidae, and Stolotermitidae it is narrower than the head capsule. Unlike Stolotermitidae, the right marginal teeth are broad, with the second marginal tooth possessing a distinct gnathal edge (Fig. 6). The right apical tooth has a distinct constriction at its base (not merely an indentation along the outer margin but there is an actual narrowing of the mandibular diameter where the right apical tooth arises: Fig. 6). A subsidiary tooth is absent. Further distinguished from Stolotermitidae by the deeply incised mandibular teeth (rather than shallowly incised). Like Archotermopsidae and Stolotermitidae the pronotum is flat, and not weakly saddle-shaped as in Hodotermitidae. The tibiae lack prominent lateral spines (although there are some minute, outer, peg-like setae in Arceotermes), as are present in Archotermopsidae. Refer to Krishna et al. (2013) for an account of basal termite families and additional characters.</p> </div>	http://treatment.plazi.org/id/03B28787AB44067AFCAF83F5FD80FF36	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	JIANG, RI-XIN;ZHANG, HONG-RUI;ELDREDGE, K. TARO;SONG, XIAO-BIN;LI, YAN-DA;TIHELKA, ERIK;HUANG, DI-YING;WANG, SHUO;ENGEL, MICHAEL S.;CAI, CHEN-YANG	JIANG, RI-XIN, ZHANG, HONG-RUI, ELDREDGE, K. TARO, SONG, XIAO-BIN, LI, YAN-DA, TIHELKA, ERIK, HUANG, DI-YING, WANG, SHUO, ENGEL, MICHAEL S., CAI, CHEN-YANG (2021): Further evidence of Cretaceous termitophily: Description of new termite hosts of the trichopseniine Cretotrichopsenius (Coleoptera: Staphylinidae), with emendations to the classification of lower termites (Isoptera). Palaeoentomology 4 (4): 374-389, DOI: 10.11646/palaeoentomology.4.4.13, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.13
03B28787AB44067BFF0D8249FA28F9F2.text	03B28787AB44067BFF0D8249FA28F9F2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Krishnatermitidae Engel 2021	<div><p>Krishnatermitidae Engel, fam. nov.</p> <p>Type genus. Krishnatermes Engel et al., 2016.</p> <p>Diagnosis. Krishnatermitidae are seemingly intermediate in divergence between other basal Euisoptera and Mastotermitidae (Engel et al., 2009, 2016), particularly owing to the lack of a hind wing basal suture (i. e., without defined hind wing scale, a trait otherwise found in all other Euisoptera), symplesiomorphically shared with mastotermitids. They also share the primitive features of a broad pronotum, procoxal ventral carina/keel, wholly pentamerous tarsi, a large forewing scale with all primary veins originating within the scale, and the forewing with multiple elongate radial veins encompassing a uniformly wide radial field. Krishnatermitidae differ from in the absence of ocelli (present in Mastotermitinae), presence of a subsidiary tooth (absent in Mastotermitidae, present in Archotermopsidae), shorter antennae with 14–18 antennomeres (20–32 in Mastotermitidae), absence of an occipital sulcus (present in Mastotermitidae as well as Ithytermes Sánchez-García et al., 2020), forewing M sclerotized as CuA rather than Rs (M sclerotized as Rs rather than CuA in Mastotermitidae), hind wing without anal lobe (present in Mastotermitidae), and reduced number of cercomeres, with 2–3 cercomeres (five in Mastotermitidae and Ithytermes). Additional traits are summarized by Engel et al. (2016). Krishnatermitidae lack a ventral cervical sclerite and fontanelle.</p> <p>The family presently includes only Krishnatermes and quite likely Ithytermes, although several other poorly known Cretaceous genera could potentially belong here.</p> </div>	http://treatment.plazi.org/id/03B28787AB44067BFF0D8249FA28F9F2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	JIANG, RI-XIN;ZHANG, HONG-RUI;ELDREDGE, K. TARO;SONG, XIAO-BIN;LI, YAN-DA;TIHELKA, ERIK;HUANG, DI-YING;WANG, SHUO;ENGEL, MICHAEL S.;CAI, CHEN-YANG	JIANG, RI-XIN, ZHANG, HONG-RUI, ELDREDGE, K. TARO, SONG, XIAO-BIN, LI, YAN-DA, TIHELKA, ERIK, HUANG, DI-YING, WANG, SHUO, ENGEL, MICHAEL S., CAI, CHEN-YANG (2021): Further evidence of Cretaceous termitophily: Description of new termite hosts of the trichopseniine Cretotrichopsenius (Coleoptera: Staphylinidae), with emendations to the classification of lower termites (Isoptera). Palaeoentomology 4 (4): 374-389, DOI: 10.11646/palaeoentomology.4.4.13, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.13
03B28787AB44067BFF0D84B6FE9CF8FF.text	03B28787AB44067BFF0D84B6FE9CF8FF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Melqartitermitidae Engel 2021	<div><p>Melqartitermitidae Engel, fam. nov.</p> <p>Type genus. Melqartitermes Engel et al., 2007.</p> <p>Diagnosis. The Melqartitermitidae are a peculiar Early Cretaceous group of primitive termites. Like Mastotermitidae and some other basal families, melqartitermitids have wholly pentamerous tarsi and a large flat pronotum. However, unlike Mastotermitidae the hind wing lacks an anal lobe and ocelli are lacking (absent in Mastotermitinae). Unlike Archotermopsidae a subsidiary tooth is present and the radial field is narrow, closer in form to that of Stolotermitidae (but this family has tetramerous tarsi, lacks a subsidiary tooth, and has a smaller pronotum). Like Mastotermitidae and other early diverging families, Melqartitermitidae have a large forewing scale with all primary veins originating within the scale. Aside from its unique combination of traits, the family differs from others in the primitive retention of a ventral cervical sclerite. In this respect it resembles Mylacrotermitidae Engel, fam. nov. (vide infra), from which it differs in the presence of a defined forewing basal suture.</p> </div>	http://treatment.plazi.org/id/03B28787AB44067BFF0D84B6FE9CF8FF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	JIANG, RI-XIN;ZHANG, HONG-RUI;ELDREDGE, K. TARO;SONG, XIAO-BIN;LI, YAN-DA;TIHELKA, ERIK;HUANG, DI-YING;WANG, SHUO;ENGEL, MICHAEL S.;CAI, CHEN-YANG	JIANG, RI-XIN, ZHANG, HONG-RUI, ELDREDGE, K. TARO, SONG, XIAO-BIN, LI, YAN-DA, TIHELKA, ERIK, HUANG, DI-YING, WANG, SHUO, ENGEL, MICHAEL S., CAI, CHEN-YANG (2021): Further evidence of Cretaceous termitophily: Description of new termite hosts of the trichopseniine Cretotrichopsenius (Coleoptera: Staphylinidae), with emendations to the classification of lower termites (Isoptera). Palaeoentomology 4 (4): 374-389, DOI: 10.11646/palaeoentomology.4.4.13, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.13
03B28787AB44067BFF0D8071FE71FEE6.text	03B28787AB44067BFF0D8071FE71FEE6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mylacrotermitidae Engel 2021	<div><p>Mylacrotermitidae Engel, fam. nov.</p> <p>Type genus. Mylacrotermes Engel et al., 2007.</p> <p>Diagnosis. The Mylacrotermitidae share with Melqartitermitidae Engel, fam. nov. the presence of ventral cervical sclerite, the absence of ocelli, a large pronotum, wholly pentamerous tarsi, and a large forewing scale (vide supra), but are unique among termites in the absence of a defined forewing basal suture demarcating the scale. Instead, the forewing is gnawed off like the hind wing in Mastotermitidae and some other extinct families where the chew marks and rough edge can be observed where the hind wing is removed.</p> </div>	http://treatment.plazi.org/id/03B28787AB44067BFF0D8071FE71FEE6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	JIANG, RI-XIN;ZHANG, HONG-RUI;ELDREDGE, K. TARO;SONG, XIAO-BIN;LI, YAN-DA;TIHELKA, ERIK;HUANG, DI-YING;WANG, SHUO;ENGEL, MICHAEL S.;CAI, CHEN-YANG	JIANG, RI-XIN, ZHANG, HONG-RUI, ELDREDGE, K. TARO, SONG, XIAO-BIN, LI, YAN-DA, TIHELKA, ERIK, HUANG, DI-YING, WANG, SHUO, ENGEL, MICHAEL S., CAI, CHEN-YANG (2021): Further evidence of Cretaceous termitophily: Description of new termite hosts of the trichopseniine Cretotrichopsenius (Coleoptera: Staphylinidae), with emendations to the classification of lower termites (Isoptera). Palaeoentomology 4 (4): 374-389, DOI: 10.11646/palaeoentomology.4.4.13, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.13
03B28787AB45067AFF0D8039FC5AFF37.text	03B28787AB45067AFF0D8039FC5AFF37.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arceotermes Engel & Jiang 2021	<div><p>Arceotermes Engel &amp; Jiang, gen. nov.</p> <p>Type species. Arceotermes hospitis Engel &amp; Jiang, sp. nov.</p> <p>Etymology. The new genus-group name is a combination of the Latin terms arceo (meaning, to “hinder” or “guard”) and termes (“termite”; genitive, termitis). The gender of the name is masculine.</p> <p>Diagnosis. Total body length (as preserved) 5.00 mm, head length (excluding mandibles) 2.08 mm, maximum width 1.83 mm. Like most soldier termites, the head is large and longer than broad, although the posterior temples are not tumid (tumid in Archotermopsis Desneux, 1904b) and the head is not dorsoventrally compressed (compressed in Stolotermitidae). The lateral surface of the head lacks striations (present in Archotermopsis), and the lateral margins are largely parallel. The dorsal surface of the head lacks a clearly visible Y-shaped ecdysial cleavage scar (it is difficult, though, to determine whether it is absent or merely exceedingly faint), lacks a fontanelle, and lacks ocelli. The compound eyes are absent and the antenna is composed of 17, largely moniliform articles. Mandibles are large, elongate, and gently bent ventrad toward their apices, with prominent apical and marginal teeth. Like Archotermopsidae and Hodotermitidae, the soldier mandibles have two right marginal teeth and three left marginal teeth (in addition to the apical tooth), the right marginal teeth are broad, much like that of Archotermopsis, and simple (sometimes with two points, such as in Hodotermopsis Holmgren, 1911 (Hodotermopsinae Engel, subfam. nov., vide infra), Zootermopsis laticeps (Banks, 1906), and to a lesser degree like Microhodotermes Sjöstedt, 1926). The apical teeth are long, pointed, and curved. The postmentum is greatly broadened anteriorly, with concave lateral margins tapering posteriorly. The pronotum is broad (not massive and elongate as in Ginormotermes Engel et al., 2016), and about as wide as the head, with the anterior margin broadly concave and the lateral margins faintly convex, roughly parallel, and broadly rounded posteriorly to a broadly and weakly convex posterior margin. The surface of the pronotum is flat.The tibial spur formula is 3-3-? (metatarsi not preserved in holotype), and the tarsi are tetramerous, while the pretarsus lacks an arolium. Unfortunately, the abdominal apex is damaged in the holotype of the type species and the cerci are not preserved.</p> <p>The genus differs from soldiers of Krishnatermes and Ginormotermes by the tetramerous tarsi (pentamerous in those genera), and from Cosmotermes Zhao et al., 2020 by the absence of compound eyes, the long left first marginal tooth relative to the second marginal tooth, and the broader pronotum (distinctly narrower than head in Cosmotermes).</p> </div>	http://treatment.plazi.org/id/03B28787AB45067AFF0D8039FC5AFF37	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	JIANG, RI-XIN;ZHANG, HONG-RUI;ELDREDGE, K. TARO;SONG, XIAO-BIN;LI, YAN-DA;TIHELKA, ERIK;HUANG, DI-YING;WANG, SHUO;ENGEL, MICHAEL S.;CAI, CHEN-YANG	JIANG, RI-XIN, ZHANG, HONG-RUI, ELDREDGE, K. TARO, SONG, XIAO-BIN, LI, YAN-DA, TIHELKA, ERIK, HUANG, DI-YING, WANG, SHUO, ENGEL, MICHAEL S., CAI, CHEN-YANG (2021): Further evidence of Cretaceous termitophily: Description of new termite hosts of the trichopseniine Cretotrichopsenius (Coleoptera: Staphylinidae), with emendations to the classification of lower termites (Isoptera). Palaeoentomology 4 (4): 374-389, DOI: 10.11646/palaeoentomology.4.4.13, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.13
03B28787AB450675FCAF8039FB5AFDF5.text	03B28787AB450675FCAF8039FB5AFDF5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arceotermes hospitis Engel & Jiang 2021	<div><p>Arceotermes hospitis Engel &amp; Jiang, sp. nov.</p> <p>(Figs 1A, B, 5, 6)</p> <p>Holotype. Holotype soldier, QUST-INSECT-0015. In mid-Cretaceous amber from Kachin Province, northern Myanmar. Deposited in the College of Marine Science and Biological Engineering, Qingdao University of Science and Technology, Qingdao, China.</p> <p>Etymology. The specific epithet is taken from the Latin term hospitis, meaning, “host”, and refers to the hosting relationship between these termites and their inquilines, C. burmiticus.</p> <p>Diagnosis. As for the genus (vide supra).</p> <p>Description. Soldier:Total body length(as preserved) 5.00 mm; head enlarged, subrectangular, longer than wide, head length (excluding mandibles) 2.08 mm, maximum width 1.83 mm, chestnut brown, with virtually no setae, remainder of body with sparsely scattered, fine, suberect setae; head not dorsoventrally compressed, slightly wider than pronotum, lateral sides roughly parallel, posterolateral angles not tumid, lateral surface without stridulatory file, posterior border gently and broadly convex; fontanelle absent; labrum longer than wide, projecting over about 0.75× mandibular length, labrum length 0.55 mm, width 0.52 mm; anteclypeus pale (contrasting against darker coloration of labrum and postclypeus), flat, transverse, subequal in length to postclypeus; postclypeus transverse, comparatively flat; clypeus flanked by small, shallow lobes lateral to each lobe overhanding antennal insertion (as in soldiers of Archotermopsidae); mandibles heavily sclerotized, approximately equal in size and length, left mandible length 0.97 mm, slightly overlapping apically, elongate, extending well beyond apex of labrum, gently bent ventrad apically; dentition prominent, with two right marginal teeth and three left marginal teeth; right marginal teeth broad, apical teeth long, pointed, curved; subsidiary tooth lacking; compound eyes absent; antenna with 17 moniliform articles; weak occipital carina present anteriorly, disappearing by tangent with widest point of postmentum; postmentum greatly widened anteriorly, somewhat spatulate in form, lateral margins concave, tapering posteriorly.</p> <p>Pronotum broad, about as wide as head, anterior margin broadly and gently concave, lateral margins faintly convex, roughly parallel, posterolaterally broadly rounded, posterior border weakly convex, surface flat. Procoxa without carina; tibial spur formula 3-3-?; tarsi tetramerous; pretarsal claws simple; arolium absent; profemur 1.33 mm, protibia length 0.83 mm; protarsus length 0.35 mm.</p> <p>Abdomen damaged as preserved.</p></div> 	http://treatment.plazi.org/id/03B28787AB450675FCAF8039FB5AFDF5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	JIANG, RI-XIN;ZHANG, HONG-RUI;ELDREDGE, K. TARO;SONG, XIAO-BIN;LI, YAN-DA;TIHELKA, ERIK;HUANG, DI-YING;WANG, SHUO;ENGEL, MICHAEL S.;CAI, CHEN-YANG	JIANG, RI-XIN, ZHANG, HONG-RUI, ELDREDGE, K. TARO, SONG, XIAO-BIN, LI, YAN-DA, TIHELKA, ERIK, HUANG, DI-YING, WANG, SHUO, ENGEL, MICHAEL S., CAI, CHEN-YANG (2021): Further evidence of Cretaceous termitophily: Description of new termite hosts of the trichopseniine Cretotrichopsenius (Coleoptera: Staphylinidae), with emendations to the classification of lower termites (Isoptera). Palaeoentomology 4 (4): 374-389, DOI: 10.11646/palaeoentomology.4.4.13, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.13
03B28787AB4A0674FCAF8363FEF2FC24.text	03B28787AB4A0674FCAF8363FEF2FC24.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cosmotermitinae Engel 2021	<div><p>Cosmotermitinae Engel, subfam. nov.</p> <p>Type genus. Cosmotermes Zhao et al., 2020.</p> <p>Diagnosis. In the soldier caste these termites can be distinguished most notable from Arceotermitinae by the presence of compound eyes, albeit reduced; the narrower pronotum; and the in the left mandible the posterior margin of left marginal tooth is shorter than or equal to the anterior margin of the second marginal tooth (the posterior margin of the left marginal tooth is longer than the anterior margin of the second marginal tooth in Arceotermes: Fig. 6). Unlike Stolotermitidae, the basal suture in alates of Cosmotermes is convex rather than straight and diagonal.</p> </div>	http://treatment.plazi.org/id/03B28787AB4A0674FCAF8363FEF2FC24	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	JIANG, RI-XIN;ZHANG, HONG-RUI;ELDREDGE, K. TARO;SONG, XIAO-BIN;LI, YAN-DA;TIHELKA, ERIK;HUANG, DI-YING;WANG, SHUO;ENGEL, MICHAEL S.;CAI, CHEN-YANG	JIANG, RI-XIN, ZHANG, HONG-RUI, ELDREDGE, K. TARO, SONG, XIAO-BIN, LI, YAN-DA, TIHELKA, ERIK, HUANG, DI-YING, WANG, SHUO, ENGEL, MICHAEL S., CAI, CHEN-YANG (2021): Further evidence of Cretaceous termitophily: Description of new termite hosts of the trichopseniine Cretotrichopsenius (Coleoptera: Staphylinidae), with emendations to the classification of lower termites (Isoptera). Palaeoentomology 4 (4): 374-389, DOI: 10.11646/palaeoentomology.4.4.13, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.13
03B28787AB4B0677FF0D83F6FE6EFF13.text	03B28787AB4B0677FF0D83F6FE6EFF13.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hodotermopsinae Engel 2021	<div><p>Hodotermopsinae Engel, subfam. nov.</p> <p>Type genus. Hodotermopsis Holmgren, 1911.</p> <p>Diagnosis. Imago right mandible with apical tooth as long as or slightly shorter than first marginal tooth (apical tooth longer than first marginal tooth in Archotermopsinae s. str.). Imago left mandible with apical short, subequal to first marginal tooth (as long as or longer than first marginal tooth, sometimes apical, first marginal, and second marginal teeth equally developed [i. e., Archotermopsis], in Archotermopsinae s. str.), second marginal tooth forming broad cutting edge, with apex projecting to or nearly to level of first marginal tooth (if second marginal tooth forms broad cutting edge in Archotermopsinae [i. e., Zootermopsis Emerson, 1933], then apex of second marginal tooth projecting to half length of first marginal tooth). Soldier with lateral spines only present on metatibia (present on all tibia in Archotermopsinae s. str.); cerci 1.5× or less length of styli (2× or more in Archotermopsinae s. str.). Otherwise typical for Archotermopsidae in presence of cryptically pentamerous tarsi (i. e., second tarsomere reduced and obscured from above giving the false impression of tetramerous tarsi from some angles) and right mandible of image with a subsidiary tooth present at base of anterior margin of first marginal tooth (i. e., between apical tooth and first marginal tooth). While Archotermopsinae tend to have one poorly distinguished inferior branch of Rs apically that parallels the main Rs branch to or just posterior to the wing apex, in Hodotermopsis there may be at times two inferior branches of Rs (Hodotermitidae have more developed, striking straight parallel inferior branches that strike toward the posterior margin (Engel et al., 2009; Krishna et al., 2013).</p> <p>Repeated studies demonstrate the distinctiveness of Hodotermopsis (Bourguignon et al., 2015; Buček et al., 2019). It may be warranted to elevate the rank of this group but given the contradictory placements of several Teletisoptera in recent phylogenetic estimates (Bourguignon et al., 2015; Legendre et al., 2015; Engel et al., 2016; Buček et al., 2019), we are hesitant to make any further refinements to the classification beyond noting the special placement of the genus.</p> </div>	http://treatment.plazi.org/id/03B28787AB4B0677FF0D83F6FE6EFF13	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	JIANG, RI-XIN;ZHANG, HONG-RUI;ELDREDGE, K. TARO;SONG, XIAO-BIN;LI, YAN-DA;TIHELKA, ERIK;HUANG, DI-YING;WANG, SHUO;ENGEL, MICHAEL S.;CAI, CHEN-YANG	JIANG, RI-XIN, ZHANG, HONG-RUI, ELDREDGE, K. TARO, SONG, XIAO-BIN, LI, YAN-DA, TIHELKA, ERIK, HUANG, DI-YING, WANG, SHUO, ENGEL, MICHAEL S., CAI, CHEN-YANG (2021): Further evidence of Cretaceous termitophily: Description of new termite hosts of the trichopseniine Cretotrichopsenius (Coleoptera: Staphylinidae), with emendations to the classification of lower termites (Isoptera). Palaeoentomology 4 (4): 374-389, DOI: 10.11646/palaeoentomology.4.4.13, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.13
03B28787AB480671FCAF82F9FEBCF8B9.text	03B28787AB480671FCAF82F9FEBCF8B9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tanytermitalis Engel & Cai 2021	<div><p>Tanytermitalis Engel &amp; Cai, gen. nov.</p> <p>Type species. Tanytermitalis philetaerus Engel &amp; Cai, sp. nov.</p> <p>Etymology. The new genus-group name is derived from the generic name Tanytermes (using the genitive of L. termes as the root, termitis), type genus of the family, combined with the Latin suffix – alis (meaning, “to have the nature or quality of”). The gender of the name is masculine.</p> <p>Diagnosis. Total body length (as preserved) 5.28 mm, head length 1.12 mm, maximum width 0.68 mm. Head elongate, posteriormost section not tapering, sides roughly parallel, (slightly tapering posteriorly slightly in Tanytermes); labrum covering mandibles (i.e., mandibles not projecting beyond labral apex), apical margin comparatively straight, only slightly narrower than posterior margin, thus straight sides slightly converging apically, labrum slightly longer than clypeus (anteclypeus and postclypeus); anteclypeus transverse, with apical margin gently convex; postclypeus transverse, slightly shorter than anteclypeus along midline; both left and right mandibles with sharply-pointed and prominent apical teeth [remainder of dentition obscured from view dorsally and ventrally]; lacinia bifurcate apically (as is in most Isoptera); antenna with 18 articles, flagellum moniliform; compound eyes without anterior emargination, circular, relatively small, immediately posterior to antennal torulus (separated from torulus in Tanytermes), separated from posterior border of head by about twice compound eye diameter (separation greater in Tanytermes); ocelli absent; fontanelle absent. Pronotum broader than long (longer than wide in Tanytermes), maximal width slightly more than width of head (narrower in Tanytermes), surface flat; anterior border slightly concave (straight in Tanytermes); lateral borders parallel in anterior half, broadly rounding to posterior margin in posterior half; posterior border broadly and faintly convex (straight in Tanytermes). Tarsi tetramerous; tibial spur formula 3–3–2 (Tanytermes 2– 3–2); pretarsal claws simple, arolium absent (present in Tanytermes). Wing membranes hyaline, faintly infumate, not reticulate; veins Sc, R, and Rs heavily pigmented, M and CuA thinner and lightly pigmented; forewing scale large, overlapping hind wing scale, humeral margin almost straight (as in Tanytermes), basal suture weakly convex; all veins originating inside wing scale, termination of CuP (claval fissure) on posterior margin just prior to basal cleavage suture; Sc terminating on costal margin in proximal quarter of wing length, at distance from basal suture about 0.75× forewing scale length; R simple, terminating at point just proximad of wing midlength (terminating at about one-third wing length in Tanytermes); Rs first branching slightly before termination of R (R shorter in Tanytermes, terminating in basal third of wing); Rs lacking inferior branches, with four superior branches (eight in Tanytermes), all simple; Rs running generally parallel to costal wing margin for entire length, forming narrow radial field, Rs separate from M along its entire length, Rs terminating prior to wing apex (as in Tanytermes); M relatively simple, running about half way between Rs and CuA, first branching slightly before wing midlength (in apical third of wing in Tanytermes), with first branch terminating at wing apex, medial field encompassing wing apex; CuA pectinate along its entire length, CuA-field encompassing nearly entire posterior margin of forewing, about 0.75× of posterior margin (about 0.85× of posterior margin in Tanytermes). Abdomen relatively narrow, cylindrical; cerci dimerous.</p> </div>	http://treatment.plazi.org/id/03B28787AB480671FCAF82F9FEBCF8B9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	JIANG, RI-XIN;ZHANG, HONG-RUI;ELDREDGE, K. TARO;SONG, XIAO-BIN;LI, YAN-DA;TIHELKA, ERIK;HUANG, DI-YING;WANG, SHUO;ENGEL, MICHAEL S.;CAI, CHEN-YANG	JIANG, RI-XIN, ZHANG, HONG-RUI, ELDREDGE, K. TARO, SONG, XIAO-BIN, LI, YAN-DA, TIHELKA, ERIK, HUANG, DI-YING, WANG, SHUO, ENGEL, MICHAEL S., CAI, CHEN-YANG (2021): Further evidence of Cretaceous termitophily: Description of new termite hosts of the trichopseniine Cretotrichopsenius (Coleoptera: Staphylinidae), with emendations to the classification of lower termites (Isoptera). Palaeoentomology 4 (4): 374-389, DOI: 10.11646/palaeoentomology.4.4.13, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.13
03B28787AB480677FF0D8231FB3CFD76.text	03B28787AB480677FF0D8231FB3CFD76.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tanytermitidae Engel 2021	<div><p>Tanytermitidae Engel, fam. nov.</p> <p>Type genus. Tanytermes Engel et al., 2007.</p> <p>Included genera. Aside from the type genus, the family includes the new genus Tanytermitalis Engel &amp; Cai, gen. nov. (vide infra).</p> <p>Diagnosis. Alate with head elongate, much like the longer alate heads of some Heterotermes Froggatt, 1897 (e. g., Heterotermes paradoxus (Froggatt, 1898), H. occiduus Hill, 1927), sub-rectangular, fontanelle absent (present in Neoisoptera) (Figs 1B, 7), Y-shaped cleavage scar absent; compound eyes small, circular (Fig. 7C); ocelli absent (present in Icoisoptera); mandibles not extending beyond apex of labrum (Fig. 7D),right mandible with two sharp marginal teeth and elongate apical tooth, without subsidiary tooth (present in Archotermopsidae and Stolotermitidae) [dentition known only for Tanytermes, obscured in Tanytermitalis, vide infra]; antenna with 14–18 articles (longer in Mastotermitidae and Teletisoptera, although some Stolotermitidae with as few as 15 articles) (Fig. 7A, B). As in all termites with the exception of some basal groups like Melqartitermitidae and Mylacrotermitidae (vide supra), Tanytermitidae lack ventral cervical sclerites (observable only in Tanytermitalis). Pronotum flat (weakly saddle shaped in Hodotermitidae), either slightly narrower or broader than head. Tibial spur formula 2–3–2 or 3–3–2 (3–3– 3 in Kalotermitidae) (Fig. 7E–G); tarsi tetramerous (i. e., second tarsomere absent—second tarsomere becomes cryptic in some Teletisoptera and then wholly absent in other clades) (Fig. 7E–G); pretarsal claws simple, arolium present or absent. Wing membranes not reticulate; Sc, R, and Rs heavily pigmented, remaining longitudinal sectors lightly pigmented; forewing scale large, overlapping hind wing scale (plesiomorphy relative to Termitidae); CuP (claval fissure) terminating on posterior wing margin prior to basal cleavage suture (i. e., inside of scale and not along suture—terminating on suture in many, but not all, basal Neoisoptera; reversed notably in Termitidae); Sc short, R1 simple, Rs running parallel to costal margin, Rs without apical parallel posterior inferiors (apical parallel posterior inferiors present in Hodotermitidae), radial field narrow, M running about halfway between Rs and CuA and sclerotized like CuA (rather than like Rs). Hind wing without anal lobe (present in Mastotermitidae). Cerci dimerous (observable only in Tanytermitalis, unknown for Tanytermes), as in Kalotermitidae.</p> <p>Cerci and forewing venation somewhat similar to some Kalotermitidae, likely a reflection of the position of Tanytermitidae as closer to Icoisoptera than to more basal grade termites (Engel et al., 2009, 2016), but differing in the shorter CuA (subapical or apical termination in most Icoisoptera, terminating at apical third in Tanytermitidae). In addition, ocelli are present in imagoes of Icoisoptera, while lacking in the present family.</p> </div>	http://treatment.plazi.org/id/03B28787AB480677FF0D8231FB3CFD76	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	JIANG, RI-XIN;ZHANG, HONG-RUI;ELDREDGE, K. TARO;SONG, XIAO-BIN;LI, YAN-DA;TIHELKA, ERIK;HUANG, DI-YING;WANG, SHUO;ENGEL, MICHAEL S.;CAI, CHEN-YANG	JIANG, RI-XIN, ZHANG, HONG-RUI, ELDREDGE, K. TARO, SONG, XIAO-BIN, LI, YAN-DA, TIHELKA, ERIK, HUANG, DI-YING, WANG, SHUO, ENGEL, MICHAEL S., CAI, CHEN-YANG (2021): Further evidence of Cretaceous termitophily: Description of new termite hosts of the trichopseniine Cretotrichopsenius (Coleoptera: Staphylinidae), with emendations to the classification of lower termites (Isoptera). Palaeoentomology 4 (4): 374-389, DOI: 10.11646/palaeoentomology.4.4.13, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.13
03B28787AB4E0671FF0D87B2FE97F302.text	03B28787AB4E0671FF0D87B2FE97F302.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tanytermitalis philetaerus JIANG & ZHANG & ELDREDGE & SONG & LI & TIHELKA & HUANG & WANG & ENGEL & CAI 2021	<div><p>Tanytermitalis philetaerus Engel &amp; Cai, sp. nov.</p> <p>(Figs 1B, D, 7)</p> <p>Holotype. Holotype alate, NIGP174707. In mid- Cretaceous amber from Kachin Province, northern Myanmar. Deposited in the Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing, China.</p> <p>Etymology. The specific epithet is taken from the Greek term ΦΙΛέταΙΡΟΣ (philetairos) [composed of ΦῘÌΛΟΣ (phílos), meaning, “beloved”, and ἑταῖΡΟΣ (hetaîros), meaning, “friend”), and refers to the relationship between these termites and their inquilines.</p> <p>Diagnosis. As for the genus (vide supra).</p> <p>Description. Imago: Total body length (as preserved) 5.28 mm; length of head to apex of labrum 1.12 mm; head width 0.68 mm; compound eye diameter 0.22 mm; distance from compound eye to posterior margin 0.36 mm; pronotal medial length 0.50 mm; pronotal width 0.87 mm; metatibial length 0.58 mm; length of forewing with scale 6.72 mm; forewing width 2.24 mm. Integument dark brown, microsculpturing of head finely imbricate. Setation of head and body sparse; setae not evident on pronotum. Rs with four simple branches, Rs terminating before wing apex; M with four main branches, M terminating at wing apex, basalmost branch terminating at apical quarter of posterior wing margin; CuA pectinately branched, with at least four simple branches (remainder of CuA obscured in holotype), cubital field occupying majority of posterior wing margin.</p></div> 	http://treatment.plazi.org/id/03B28787AB4E0671FF0D87B2FE97F302	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	JIANG, RI-XIN;ZHANG, HONG-RUI;ELDREDGE, K. TARO;SONG, XIAO-BIN;LI, YAN-DA;TIHELKA, ERIK;HUANG, DI-YING;WANG, SHUO;ENGEL, MICHAEL S.;CAI, CHEN-YANG	JIANG, RI-XIN, ZHANG, HONG-RUI, ELDREDGE, K. TARO, SONG, XIAO-BIN, LI, YAN-DA, TIHELKA, ERIK, HUANG, DI-YING, WANG, SHUO, ENGEL, MICHAEL S., CAI, CHEN-YANG (2021): Further evidence of Cretaceous termitophily: Description of new termite hosts of the trichopseniine Cretotrichopsenius (Coleoptera: Staphylinidae), with emendations to the classification of lower termites (Isoptera). Palaeoentomology 4 (4): 374-389, DOI: 10.11646/palaeoentomology.4.4.13, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.13
