taxonID	type	description	language	source
03BE2125FFE1560DFF65CDD7FA86FEB3.taxon	type_taxon	Type species. Psilotreta frontalis Banks, 1899	en	WICHARD, WILFRIED, MÜLLER, PATRICK, XU, CHUN-PENG (2021): The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber. Palaeoentomology 4 (4): 331-338, DOI: 10.11646/palaeoentomology.4.4.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.8
03BE2125FFE1560DFF65CDD7FA86FEB3.taxon	diagnosis	Diagnosis (adapted to the fossils and modified after Banks, 1899, Parker & Wiggins, 1987, Oláh & Johanson, 2010). Head: Ocelli absent, antennae longer than forewings, maxillary palps five-segmented, terminal segment not annulate or flexible, labial palps three-segmented. Tibial spurs: 2 / 4 / 4. Wings: Forewings longer and narrower than hindwings. Discoidal cell usually present. R 1 running separate to wing margin, not fused with Sc or R 2. Fork I often longer than fork II; fork I sessile, fork II usually petiolate, fork III usually absent, fork IV always absent; medial cell absent, M reduced, often simple or simply forked or three-forked. Crossveins sc-r, r, s, r-m and cu-a usually present, crossveins m and cu absent. In hindwings fork I longer than fork II. Male genitalia: Inferior appendages (gonocoxites) two-segmented, harpagoes originating from apex of coxopodites or originate mesally on coxopodites. Following Oláh & Johanson (2010) genus Psilotreta includes three diagnostic species-groups: 1. Psilotreta japonica diagnostic species-group, 2. Psilotreta trimeresuri diagnosticspecies-group, 3. Psilotretafrontalis diagnostic species-group. The diagnostic species groups are distinguished on the following genitalic characters: (1) the position of the harpagoes on the coxopodites; (2) the number and size of apical spines on the harpagoes; and (3) the presence or absence of phallic parameres.	en	WICHARD, WILFRIED, MÜLLER, PATRICK, XU, CHUN-PENG (2021): The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber. Palaeoentomology 4 (4): 331-338, DOI: 10.11646/palaeoentomology.4.4.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.8
03BE2125FFE1560FFCC7C86FFE88FEDE.taxon	description	(Fig. 1)	en	WICHARD, WILFRIED, MÜLLER, PATRICK, XU, CHUN-PENG (2021): The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber. Palaeoentomology 4 (4): 331-338, DOI: 10.11646/palaeoentomology.4.4.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.8
03BE2125FFE1560FFCC7C86FFE88FEDE.taxon	materials_examined	Type material. Male, NIGP 175453, deposited in the Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences. Well preserved in mid-Cretaceous Burmese amber, Kachin State, Myanmar. Body well preserved; forewing in dorsal view, hindwing covered dorsally by forewing. Antennae and legs present. Male genitalia visible in ventral view.	en	WICHARD, WILFRIED, MÜLLER, PATRICK, XU, CHUN-PENG (2021): The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber. Palaeoentomology 4 (4): 331-338, DOI: 10.11646/palaeoentomology.4.4.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.8
03BE2125FFE1560FFCC7C86FFE88FEDE.taxon	etymology	Etymology. This new species is the first fossil Psilotreta described from the mid-Cretaceous Burmese amber.	en	WICHARD, WILFRIED, MÜLLER, PATRICK, XU, CHUN-PENG (2021): The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber. Palaeoentomology 4 (4): 331-338, DOI: 10.11646/palaeoentomology.4.4.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.8
03BE2125FFE1560FFCC7C86FFE88FEDE.taxon	diagnosis	Diagnosis. Filamentous antennae about three times longer than forewing lengths. In forewings media M reduced, here without base, apical part originating from R 4 + 5. In male genitalia harpagoes originate mesally on coxopodites. Therefore the extinct species belong very probably to the Psilotreta trimeresuri diagnostic speciesgroup (sensu Oláh & Johanson, 2010).	en	WICHARD, WILFRIED, MÜLLER, PATRICK, XU, CHUN-PENG (2021): The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber. Palaeoentomology 4 (4): 331-338, DOI: 10.11646/palaeoentomology.4.4.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.8
03BE2125FFE1560FFCC7C86FFE88FEDE.taxon	description	Description. Embedded in amber, conspicuous imago by long antennae, which about three times as long as elongated forewings with a length of 7 mm. Extant Psilotreta species often with antennae longer than their forewings. In forewings (7 mm in lengths) fork I about 1 / 3 times longer than fork II, discoidal cell closed. Fork II sessile, arising at junction of R 4 + 5 and crossveins. Media M reduced, simple (M 1, 2, 3, 4), without base, represented by single apical branch, originating from R 4 + 5; forks III and IV absent. Cu 1 running straight, apically bifurcated to fork V; Cu 2 simple, running unforked to wing margin. Male genitalia visible ventrally, but through very cloudy amber and difficult to recognize. Only inferior appendages can be described. In ventral view inferior appendages appear narrow, elongated and two-segmented consisting of coxopodite and harpago. Shorter harpago arising from coxopodite laterally in middle and not fully reaching apical level of coxopodite. Inferior appendages covered with relatively long hairs; spines not visible at apex of harpago. Although other structures (e. g., phallic apparatus, parameres) of male genitalia cannot be clearly described, unambiguous organization of inferior appendages suggests that Psilotreta fossilis sp. nov. likely belongs to the diagnostic species group Psilotreta trimeresuri.	en	WICHARD, WILFRIED, MÜLLER, PATRICK, XU, CHUN-PENG (2021): The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber. Palaeoentomology 4 (4): 331-338, DOI: 10.11646/palaeoentomology.4.4.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.8
03BE2125FFE1560FFCC7C86FFE88FEDE.taxon	discussion	Remarks. The wing venation of this fossil agrees largely with those of extant Psilotreta species that also have simple media (M 1, 2, 3, 4), e. g., Psilotreta baureo Malicky, 1989, Psilotreta illuan Malicky, 1989, Psilotreta enikoae Oláh & Johanson, 2010, Psilotreta kurenschikovorum Arefina-Armitage & Armitage, 2011, all extant species from Southeast Asia.	en	WICHARD, WILFRIED, MÜLLER, PATRICK, XU, CHUN-PENG (2021): The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber. Palaeoentomology 4 (4): 331-338, DOI: 10.11646/palaeoentomology.4.4.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.8
03BE2125FFE3560FFF65CFEBFB86FA16.taxon	description	(Fig. 2)	en	WICHARD, WILFRIED, MÜLLER, PATRICK, XU, CHUN-PENG (2021): The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber. Palaeoentomology 4 (4): 331-338, DOI: 10.11646/palaeoentomology.4.4.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.8
03BE2125FFE3560FFF65CFEBFB86FA16.taxon	materials_examined	Type material. Male, NIGP 175454, deposited in the Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences. Well preserved in mid Cretaceous Burmese amber, Kachin State, Myanmar. Body well preserved, but embedded in turbid and unclear amber. Head and antennae incomplete. Forewing in dorsal view, hindwing overlapping the forewing from ventrally. Legs present. Male genitalia turbid, visible in ventral view.	en	WICHARD, WILFRIED, MÜLLER, PATRICK, XU, CHUN-PENG (2021): The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber. Palaeoentomology 4 (4): 331-338, DOI: 10.11646/palaeoentomology.4.4.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.8
03BE2125FFE3560FFF65CFEBFB86FA16.taxon	etymology	Etymology. The new species is named after the Kachin State (Myanmar), where the Burmese amber is found.	en	WICHARD, WILFRIED, MÜLLER, PATRICK, XU, CHUN-PENG (2021): The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber. Palaeoentomology 4 (4): 331-338, DOI: 10.11646/palaeoentomology.4.4.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.8
03BE2125FFE3560FFF65CFEBFB86FA16.taxon	diagnosis	Diagnosis. Male bearing bipectinate antennae. In forewing forks I, II, III and V present; media M apically three-branched in M 1 + M 2 and M 3 + 4. Cu 1 running straight, apically bifurcated to fork V; Cu 2 simple, running unforked to wing margin. Male genitalia with two-segmented inferior appendages; harpago arising laterally in middle from coxopodite and overtopping coxopodite in length.	en	WICHARD, WILFRIED, MÜLLER, PATRICK, XU, CHUN-PENG (2021): The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber. Palaeoentomology 4 (4): 331-338, DOI: 10.11646/palaeoentomology.4.4.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.8
03BE2125FFE3560FFF65CFEBFB86FA16.taxon	description	Description. Head: antennae bipectinate, longer than forewings, consisting of a long scapus, about 2 / 3 as long as head, a small pedicellus and more than 22 bipectinate flagellomeres, before antennae cut off. Each flagellomere bearing rami on both sides; rami longer than flagellomeres, basally slender and gradually expanded towards rounded apex. Maxillary palps five-segmented; 1 st segment smaller than 2 nd segment, 3 rd and 4 th segments of about equal lengths and longer than 2 nd and 5 th segments, both also of same length. Wings: forewing with a length of 5.5 mm; forewing venation with apical forks I, II, III and V present, M apically forked in fork III (M 1 + M 2) and M 3 + 4. Discoidal cell not visible, forks I and II sessile. Crossveins difficult to recognize. Male genitalia (visible ventrally): inferior appendages two-segmented. Coxopodite elongate, digitiform, in ventro-lateral view basally slender and gradually expanded towards rounded apex. Harpago originating latero-mesally on coxopodite; also basally slender and gradually expanded towards rounded apex and overtop coxopodite. Segments of inferior appendage irregularly hairy and obviously of same lengths. So far as visible: Segment X apparently covering phallic apparatus dorsally with two oblong scales. Lateral preanal appendages (or parameres?) long and straight.	en	WICHARD, WILFRIED, MÜLLER, PATRICK, XU, CHUN-PENG (2021): The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber. Palaeoentomology 4 (4): 331-338, DOI: 10.11646/palaeoentomology.4.4.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.8
03BE2125FFE3560FFF65C84BFD9EF93E.taxon	type_taxon	Type species. Palaeopsilotreta xiai Wichard & Wang, 2017 Revised diagnosis. Head: Male antennae (partially) bipectinate, about as long as forewings. Rami of bipectinate flagellomeres slim and about as long as flagellomeres. Female antennae simple, not bipectinate. Maxillary palps five-segmented, terminal segment not flexible or annulate. Ocelli absent. Frontal, antennal and posterior setose warts present. Thorax: Pronotum with two pronotal setose warts, mesoscutum bearing pair of setose warts, mesoscutellum covered by a single wart, large, domelike and ovoid. Tibial spurs: 2 / 4 / 4. Wings: In fore- and hindwings forks I, II, (III) and V present; discoidal cells open or closed, medial cell absent. In forewings media M variable: Simple (M 1, 2, 3, 4) or two-branched in M 1 + 2 and M 3 + 4 or three-branched in M 1, M 2 and M 3 + 4. Cu 2 reaching wing margin and not being reduced or absent as in other odontocerid genera. Male genitalia: Inferior appendages two-segmented, harpagoes originating from apex of coxopodites (P. xiai, P. burmanica and P. cretacea) or originating mesally on coxopodites (P. kachini and P. succini).	en	WICHARD, WILFRIED, MÜLLER, PATRICK, XU, CHUN-PENG (2021): The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber. Palaeoentomology 4 (4): 331-338, DOI: 10.11646/palaeoentomology.4.4.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.8
03BE2125FFE3560FFF65C84BFD9EF93E.taxon	discussion	Remarks. The affiliation of the genus Palaeopsilotreta to the family Odontoceridae was discussed and proposed in Wichard et al. (2020). Palaeopsilotreta is closely related to Psilotreta and is distinguished by the apomorphic, bipectinated antennae in the male, which are not verified in any species of Psilotreta. Based on the presence of five-segmented palpi, the spur formula being 2 / 4 / 4, the absence of the medial cell and the presence of wing forks I, II, (III), and V in both wings (as in Psilotreta), we can assign the fossil genus Palaeopsilotreta to Odontoceridae. Whether the genus Palaeopsilotreta is a monophyletic group of the family Odontoceridae or of a closely related family (e. g., Philorheithridae) needs in time to be investigated with further fossil findings and taxonomic data.	en	WICHARD, WILFRIED, MÜLLER, PATRICK, XU, CHUN-PENG (2021): The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber. Palaeoentomology 4 (4): 331-338, DOI: 10.11646/palaeoentomology.4.4.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.8
03BE2125FFE3560AFCC7CF03FE87F889.taxon	description	(Fig. 3)	en	WICHARD, WILFRIED, MÜLLER, PATRICK, XU, CHUN-PENG (2021): The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber. Palaeoentomology 4 (4): 331-338, DOI: 10.11646/palaeoentomology.4.4.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.8
03BE2125FFE3560AFCC7CF03FE87F889.taxon	materials_examined	Type material. Male, ZFMK-TRI 000822, kept in the amber collection of the Museum Zoological Research Museum Alexander Koenig, Bonn, Germany (former Patrick Müller collection, BUB 3865); Mid-Cretaceous Burmese amber inclusion; Kachin State, Myanmar. Body well preserved, but a fracture across body impairs viewing. Forewing in dorsal view, hindwing covered dorsally by forewing. Antennae and legs present. Male genitalia visible in ventral view, but one hind leg across genitalia interferes with view.	en	WICHARD, WILFRIED, MÜLLER, PATRICK, XU, CHUN-PENG (2021): The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber. Palaeoentomology 4 (4): 331-338, DOI: 10.11646/palaeoentomology.4.4.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.8
03BE2125FFE3560AFCC7CF03FE87F889.taxon	etymology	Etymology. The new species is named after the amber in which the fossil species is embedded.	en	WICHARD, WILFRIED, MÜLLER, PATRICK, XU, CHUN-PENG (2021): The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber. Palaeoentomology 4 (4): 331-338, DOI: 10.11646/palaeoentomology.4.4.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.8
03BE2125FFE3560AFCC7CF03FE87F889.taxon	diagnosis	Diagnosis. Fossil extinct species embedded in mid- Cretaceous Burmese amber. Male bearing bipectinate antennae. In forewing forks I, II, V present, discoidal cell closed. Media M reduced, without base, originating from R 4 + 5, running to wing margin, simply forked at apex into M 1 + 2 and M 3 + 4. Cu 1 running straight, apically bifurcated to fork V (Cu 1 a + Cu 1 b). Cu 2 simple, running unforked to Cu 1 b and further (Cu 2 + Cu 1 b) straight to wing margin. In male genitalia harpago small, originating mesally on point of bifurcation of strong and slightly curved coxopodite, dorso-apically covered with irregular, dark spines.	en	WICHARD, WILFRIED, MÜLLER, PATRICK, XU, CHUN-PENG (2021): The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber. Palaeoentomology 4 (4): 331-338, DOI: 10.11646/palaeoentomology.4.4.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.8
03BE2125FFE3560AFCC7CF03FE87F889.taxon	description	Description. Head: Antennae bipectinate, longer than forewings, consisting of a strong scapus, a small pedicellus and a long flagellum, first half with 13 flagellomeres bipectinate, last half of flagellum slightly toothed and filiform. Each of bipectinate flagellomeres bearing rami on both sides; rami partially significantly longer than flagellomere, basally slender and gradually expanded towards rounded apex. Maxillary palps five-segmented; 1 st segment smaller than 2 nd segment, 3 rd longest segment, 4 th and 5 th segments of about equal length and smaller than 3 rd segment. Wings: forewing length 6 mm; apical forks I, II and V present, M without base, (pseudo-) originating in R 4 + 5 and simple forked in M 1 + 2 and M 3 + 4. Discoidal cell closed, forks I and II sessile. Male genitalia: inferior appendages two-segmented. Coxopodites in ventral view uniform, strong and slightly curved towards genital centre. Harpago originating lateromesally on coxopodite, dorso-apically covered with irregular, dark spines.	en	WICHARD, WILFRIED, MÜLLER, PATRICK, XU, CHUN-PENG (2021): The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber. Palaeoentomology 4 (4): 331-338, DOI: 10.11646/palaeoentomology.4.4.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.8
