identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
41D326060C70536D806CD1C8C0810C40.text	41D326060C70536D806CD1C8C0810C40.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liodessus picinus Balke & Suarez-Megna & Ospina-Torres & Venegas & Prieto & Hendrich 2021	<div><p>Liodessus picinus sp. nov.</p> <p>Figures 1, 2G, 3</p> <p>Type locality.</p> <p>Sumapaz National Park, Bogota, Colombia.</p> <p>Holotype.</p> <p>"Colombia, ♀; Bogota D.C., PN Sumapaz; 3,500 m; 13.xi.2018; 4.290°N; 74.207°W; Ospina, Venegas, Balke and Megna (COL_MB_2018_04) Voucher MB8416" (LIAUN).</p> <p>Paratypes.</p> <p>2 ♀♀, same data but voucher numbers MB8414, 8415 (ZSM).</p> <p>Description of holotype.</p> <p>Habitus with distinct discontinuity between pronotum and elytra (Fig. 1A), pronotum widest before base (Fig. 1A, B). Total length of beetle 2.0 mm; length without head 1.8 mm; maximum width 0.9 mm.</p> <p>Color. Very dark brown to blackish dorsally and ventrally, base of meso- and metatibia contrastingly of lighter color (Fig. 1A, B).</p> <p>Surface sculpture. Head with few setiferous punctures in front of a distinct occipital line, distinct microreticulation present except on middle of head between the eyes (Fig. 1B); posteriorly of occipital line with distinct microreticulation and few punctures. Pronotum and elytron shiny, with moderately dense and coarse setiferous punctation; pronotum with few wrinkles laterally (Fig. 1B).</p> <p>Structures. Head with distinct occipital line, with rounded clypeus. Antenna stout. Pronotum with distinct lateral bead and distinct, long and deep basal striae (Fig. 1A, B). Elytron with short basal striae, without sutural line and without basal epipleural transverse carina. Metathoracic wings not examined in holotype (in paratype: short, about half the length of elytron).</p> <p>Variation.</p> <p>One paratype is slightly larger, TL 2.1 mm. In one paratype, the elytral basal striae are not very obvious.</p> <p>Male.</p> <p>Unknown.</p> <p>Etymology.</p> <p>Picinus, black, highlighting the dark coloration of the species.</p> <p>Comparative notes.</p> <p>The species is well characterized by its small size (2.0-2.1 mm total length, shorter than most other Colombian Páramo species (Megna et al. 2019; Balke et al. 2021) except L. lacunaviridis (Balke et al. 2021)); dark coloration, discontinuous habitus, shiny surface of pronotum and elytrae, presence of distinct occipital line, distinct basal pronotal striae with short or even faint basal elytral striae.</p> <p>We assembled sequences of the Colombian species, trimming the sequence ends to avoid missing characters. We obtained a 611 base pair reference alignment to observe sequence variation. Liodessus picinus sp. nov. differs by approx. 9-11% (5′ cox1) from each of the other Colombian species and therefore forms a well-delineated clade with distinct mitochondrial cox 1 signature.</p> <p>Distinguishing characters of the other Colombian Páramo species from the new species are:</p> <p>Liodessus azufralis Megna et al., 2019: lighter coloration, brown to dark brown (Fig. 2E);</p> <p>L. bogotensis: brighter coloration (Fig. 2A, B); L. lacunaviridis Balke et al., 2021 and L. quimbaya Megna et al., 2019: female with dull surface of pronotum and elytron due to presence of fine microreticulation (e.g. Fig. 2C, F); L. quillacinga Megna et al., 2019: long, distinct basal elytral stria (Fig. 2D).</p> <p>Distribution.</p> <p>Only known from the type locality.</p> <p>Habitat.</p> <p>Shallow and exposed peatland puddles, collected with strainer out of mats of vegetation including mosses (most likely Sphagnum sp.), Ranunculus sp. and Azolla sp. (Fig. 3B-D). The species is syntopic with L. bogotensis.</p> <p>Checklist of the High Andean species of Liodessus Guignot, 1953</p> <p>1. Liodessus acollensis Guignot, 1955: Peru</p> <p>2. Liodessus alpinus Balke, Megna, Zenteno, Figueroa &amp; Hendrich, 2020b: Peru</p> <p>3. Liodessus altoperuensis Balke, Megna, Zenteno, Figueroa &amp; Hendrich, 2020a: Peru</p> <p>4. Liodessus andinus Guignot, 1957: Bolivia</p> <p>5. Liodessus azufralis Megna, Hendrich &amp; Balke, 2019: Colombia</p> <p>6. Liodessus bogotensis Guignot, 1953: Colombia</p> <p>7. Liodessus caxamarca Balke, Megna, Zenteno, Figueroa &amp; Hendrich, 2020a: Peru</p> <p>8. Liodessus hauthi Balke, Megna, Zenteno, Figueroa &amp; Hendrich, 2020b: Peru</p> <p>9. Liodessus lacunaviridis Balke, Ospina-Torres, Megna &amp; Hendrich, 2020a: Colombia</p> <p>10. Liodessus picinus sp. nov.: Colombia</p> <p>11a. Liodessus quillacinga quillacinga Megna, Hendrich &amp; Balke, 2019: Colombia</p> <p>11b Liodessus quillacinga cochaensis Megna, Hendrich &amp; Balke, 2019: Colombia</p> <p>11c. Liodessus quillacinga cumbalis Megna, Hendrich &amp; Balke, 2019: Colombia</p> <p>12. Liodessus quimbaya Megna, Hendrich &amp; Balke, 2019: Colombia</p> <p>13. Liodessus rhigos Balke, Megna, Zenteno, Figueroa &amp; Hendrich, 2020b: Peru</p> <p>14. Liodessus thespesios Balke, Megna, Zenteno, Figueroa &amp; Hendrich, 2020b: Peru</p> </div>	http://treatment.plazi.org/id/41D326060C70536D806CD1C8C0810C40	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Balke, Michael;Suarez-Megna, Yoandri;Ospina-Torres, Rodulfo;Venegas, Juan Simon;Prieto, Carlos;Hendrich, Lars	Balke, Michael, Suarez-Megna, Yoandri, Ospina-Torres, Rodulfo, Venegas, Juan Simon, Prieto, Carlos, Hendrich, Lars (2021): A new Colombian species of Liodessus diving beetles from the Paramo de Sumapaz (Coleoptera, Dytiscidae, Bidessini). ZooKeys 1059: 79-87, DOI: http://dx.doi.org/10.3897/zookeys.1059.70134, URL: http://dx.doi.org/10.3897/zookeys.1059.70134
