taxonID	type	description	language	source
03DEC73D856AFFC8FF3EFF109E47B0CD.taxon	description	(Figs 1, 2, 3 A, 4 A, 5 A)	en	Hoskin, Conrad J., Couper, Patrick J. (2014): Two new skinks (Scincidae: Glaphyromorphus) from rainforest habitats in north-eastern Australia. Zootaxa 3869 (1): 1-16, DOI: http://dx.doi.org/10.11646/zootaxa.3869.1.1
03DEC73D856AFFC8FF3EFF109E47B0CD.taxon	materials_examined	Material examined: Holotype: QMJ 93341, Melville Range (14 ° 16 ' 33 " S, 144 ° 29 ' 32 " E, elevation 460 m a. s. l.), Cape Melville, north-east Queensland, C. J. Hoskin & H. B. Hines, 13 December 2013. Paratypes: QMJ 93339, QMJ 93340, collection details as for holotype; QMJ 92570, QMJ 92571, Melville Range (14 ° 16 ' 33 " S, 144 ° 29 ' 32 " E, elevation 460 m a. s. l), C. J. Hoskin, 20 March 2013; QMJ 92553, QMJ 92554, Melville Range (14 ° 18 ' 55 " S, 144 ° 29 ' 50 " E, 110 m a. s. l.), C. J. Hoskin & K. Aland, 9 February 2013.	en	Hoskin, Conrad J., Couper, Patrick J. (2014): Two new skinks (Scincidae: Glaphyromorphus) from rainforest habitats in north-eastern Australia. Zootaxa 3869 (1): 1-16, DOI: http://dx.doi.org/10.11646/zootaxa.3869.1.1
03DEC73D856AFFC8FF3EFF109E47B0CD.taxon	diagnosis	Diagnosis. Glaphyromorphus othelarrni sp. nov. is diagnosed from all congeners in having: adpressed limbs in contact; more than 27 midbody scale rows; the prefontal separated from the preocular; large body size (max SVL ~ 93 mm); usually eight supralabials (with 6 th below centre of eye); more than 13 subdigital lamellae beneath 4 th finger; more than 21 lamellae beneath 4 th toe.	en	Hoskin, Conrad J., Couper, Patrick J. (2014): Two new skinks (Scincidae: Glaphyromorphus) from rainforest habitats in north-eastern Australia. Zootaxa 3869 (1): 1-16, DOI: http://dx.doi.org/10.11646/zootaxa.3869.1.1
03DEC73D856AFFC8FF3EFF109E47B0CD.taxon	etymology	Etymology. Othelarrni means ‘ He Listens’ and this was a name given to Bob Flinders, who was born in the Cape Melville area and who passed on much of the knowledge and responsibility for that country to the current generation of its Traditional Owners. The species was named by the bubu gudjin of Cape Melville, the Traditional Owners who have the responsibility to speak for the land where the species live.	en	Hoskin, Conrad J., Couper, Patrick J. (2014): Two new skinks (Scincidae: Glaphyromorphus) from rainforest habitats in north-eastern Australia. Zootaxa 3869 (1): 1-16, DOI: http://dx.doi.org/10.11646/zootaxa.3869.1.1
03DEC73D856AFFC8FF3EFF109E47B0CD.taxon	description	Measurements and scale counts of holotype QMJ 93341 (Figs 1, 2, 3 A, 4 A). SVL = 75.4 mm, AG = 37.3 mm, TL = 140 mm, L 1 = 19.9 mm, L 2 = 29.6 mm, HL = 14.5 mm, HW = 11.4 mm, NL = 13.5 mm, midbody scale rows = 28, paravertebrals = 58, lamellae 4 th toe = 25, lamellae 4 th finger = 15, supralabials = 8, supralabial below centre of eye = 6 th infralabials = 7, supraciliaries = 7. Description of type series. Data presented as range followed by mean in brackets (n = 7, unless stated otherwise). Adult measurements (mm): SVL = 75.4 – 92.9 (85.3), AG = 37.3 – 49.6 (43.9), Tail = 140.0 – 144.0 (142.0), L 1 = 18.3 – 20.7 (19.7), L 2 = 29.6 – 32.9 (30.8), HL = 14.5 – 17.0 (16.0), HW = 11.4 – 13.1 (12.1), NL = 12.5 – 15.5 (13.9) (Table 1). Adult proportions (as % SVL): AG = 49 – 53 (51), Tail = 147 – 186 (173), L 1 = 22 – 26 (24), L 2 = 33 – 41 (38), HL = 18 – 20 (19), HW = 13 – 15 (14), NL = 14 – 18 (16) (Table 1). Body: elongate. Neck broad and not well differentiated from back of head. Snout rounded in profile. Limbs moderate, pentadactyl, and overlapping when adpressed. Scalation: Scales smooth, with rounded posterior margins; 28 – 30 (mean = 28.3) rows at midbody; paravertebral scales only slightly enlarged (except enlarged nuchals) and numbering 55 – 61 (mean = 58.9) in a line between the parietals and the posterior margin of the hindlimb. Nasals moderate, well-spaced with a relatively large external naris; rostral and frontonasal in moderate contact; prefrontals large, moderately to narrowly separated and not contacting 1 st preocular; frontal contacting frontonasal, prefrontals, first two supraoculars, frontoparietals and narrowly separated or in point contact with 1 st supraciliary; supraoculars 4, second the largest; supraciliaries 7 – 8 (mean = 7.3), first the largest; frontoparietals paired and distinct from interparietal; parietals in contact behind interparietal; 7 – 9 (mean = 7.9) nuchal scales; primary temporals 1; secondary temporals 2, upper largest and overlapping lower; loreals 2; preoculars 2; presuboculars 2; an enlarged subocular scale penetrating the suture between the 5 th and 6 th supralabials; supralabials 8, 6 th below centre of eye (except QMJ 93339, which has 9 on the left side, with 7 th below centre of eye); infralabials 6 – 7 (mean = 6.7); postmental contacting 2 infralabials on each side; lower eyelid scaly; ear opening round or vertically oval, without lobules and with tympanum moderately recessed; lamellae under 4 th finger 14 – 15 (mean = 14.3); lamellae under 4 th toe 22 – 25 (mean = 23.3). Colour pattern in preservative (Figs 3 A, 4 A): Dorsal ground colour light to dark brown, immaculate (QMJ 92570) or with black spots (QMJ 93339) or transverse bars anteriorly (QMJ 93341). Lateral surfaces with longitudinally aligned flecks or vertical wavy bars, which are most prominent on the neck but extend to forebody before breaking up into a series of black flecks that extend to the groin and base of tail. The upper labials are predominantly light with dark vertical bars along sutures. Venter immaculate cream except for a grey tinge on the belly and chest of some individuals. Dark grey flecking present along edge of jaw and lower neck. Colour pattern in life (Figs 1, 2, 5 A). As for preserved specimens but colours richer and appearance generally more glossy. The dorsum is distinctly copper-coloured on lighter individuals. Comparison with similar species. Glaphyromorphus othelarrni sp. nov. can only be confused with G. fuscicaudis, G. nigricaudis and G. nyanchupinta sp. nov. It is readily distinguished from all three species by its supralabial count (typically 8 with 6 th below centre of eye vs typically 7 with 5 th below centre of eye) (Fig. 4 A), the number of subdigital lamellae beneath the 4 th finger (14 – 15 vs <14) and 4 th toe (mean 23 vs means of 18 – 21), and its relatively longer limbs (L 1 / SVL: 0.22 – 0.26 vs ≤ 0.22; L 2 / SVL: 0.33 – 0.41 vs ≤ 0.34) (Table 1). It is further distinguished from G. fuscicaudis in having a proportionately larger head (HW / SVL: 0.13 – 0.15 vs 0.12 – 0.13; HL / SVL: 0.18 – 20 vs 0.16 – 0.17); shorter interlimb length (AG / SVL 0.49 – 0.53 vs 0.52 – 0.58); and generally fewer paravertebral scales (mean 59 vs 64) (Table 1). It also lacks the series of yellow dorsolateral blotches that are prominent in G. fuscicaudis (Figs 5 A, 5 C). Glaphyromorphus othelarrni sp. nov. is further distinguished from G. nigricaudis in having a proportionately shorter interlimb length (AG / SVL 0.49 – 0.53 vs 0.52 – 0.60); a more robust form (WT / SVL 0.17 – 0.22 vs 0.09 – 0.17); more midbody scale rows (28 – 0.30 vs 24 – 28); and more paravertebral scales (55 – 61 vs 51 – 56) (Table 1). It is further distinguished from G. nyanchupinta sp. nov. in being larger in all measures (e. g., SVL 74.5 – 92.9 vs 49.2 – 53.6); in having a proportionately longer tail (TL / SVL 1.47 – 1.86 vs 1.00); a more robust form (WT / SVL 0.17 – 0.22 vs 0.04 – 0.06); more midbody scale rows (28 – 30 vs 25 – 27) (Table 1); and a less patterned dorsum (dorsal pattern breaks up beyond midbody vs pattern present to hindlimbs) (Figs 5 A, 5 B), and less patterned upper labials (upper labials predominantly pale with dark sutures vs upper labials predominantly dark with a central pale dot) (Fig. 4 A, 4 B).	en	Hoskin, Conrad J., Couper, Patrick J. (2014): Two new skinks (Scincidae: Glaphyromorphus) from rainforest habitats in north-eastern Australia. Zootaxa 3869 (1): 1-16, DOI: http://dx.doi.org/10.11646/zootaxa.3869.1.1
03DEC73D856AFFC8FF3EFF109E47B0CD.taxon	distribution	Distribution. Known only from the Melville Range, Cape Melville, north-eastern Australia (Fig. 6). Glaphyromorphus othelarrni sp. nov. has been recorded in three areas: in the vicinity of the type locality in the western uplands (14 ° 16 ' 33 " S, 144 ° 29 ' 32 " E, 450 – 520 m a. s. l.), around the highest peak in the Melville Ra. (14 ° 16 ' 59 " S, 144 ° 29 ' 59 " E, 600 m a. s. l.), and in the lowlands at the south of the range (14 ° 18 ' 55 " S, 144 ° 29 ' 50 " E, 110 m a. s. l.). Habitat and habits. Found in rocky areas in rainforest (Fig. 7). All individuals were found where thick leaflitter had accumulated at the base of boulders or amongst boulders (e. g., Fig. 7 B). Skinks were observed active in the leaf-litter, on adjacent rock surfaces, and amongst crevices between the boulders. When pursued the skinks retreated deep into the leaf-litter or into rock crevices. Activity was greatest in the couple of hours before dusk, and during this period the skink was commonly encountered wherever there were boulders in the rainforest. Most individuals were missing at least one digit (e. g., the 5 th toe on the right hindfoot in Figure 2), and some individuals were missing all fingers or toes on a foot. The reason for this was not resolved. The other skinks found in sympatry at G. othelarrni sp. nov. sites were Eulamprus brachysoma (Lönnberg & Andersson, 1915), Bellatorias frerei (Günther, 1897), Saproscincus saltus Hoskin, 2013, an undescribed species of Carlia (Hoskin, in press), and a species of Lygisaurus (Hoskin & Hines, under investigation).	en	Hoskin, Conrad J., Couper, Patrick J. (2014): Two new skinks (Scincidae: Glaphyromorphus) from rainforest habitats in north-eastern Australia. Zootaxa 3869 (1): 1-16, DOI: http://dx.doi.org/10.11646/zootaxa.3869.1.1
03DEC73D8561FFC5FF3EFF109814B1EF.taxon	description	(Figs 3 B, 4 B, 5 B, 8, 9)	en	Hoskin, Conrad J., Couper, Patrick J. (2014): Two new skinks (Scincidae: Glaphyromorphus) from rainforest habitats in north-eastern Australia. Zootaxa 3869 (1): 1-16, DOI: http://dx.doi.org/10.11646/zootaxa.3869.1.1
03DEC73D8561FFC5FF3EFF109814B1EF.taxon	materials_examined	Material examined. Holotype: QMJ 85244, mature male with turgid opaque testes, Peach Creek (13 ° 44 ' 12 " S, 143 ° 19 ' 47 " E, elevation 530 m a. s. l.), McIlwraith Range, north-east Queensland, collected 20 July 2007 by S. Williams & C. Moritz. Paratypes: QMJ 38195, adult female with developing follicles, 17 km ENE of Mt Croll (13 º 46 ' S, 143 º 19 ' E), McIlwraith Range, collected 2 June 1979, J. W. Winter & R. G. Atherton; QMJ 66642, adult female with developing follicles, Peach Creek headwaters (13 º 44 ' 15 ", S 143 º 20 ' 20 " E, 530 m a. s. l.), McIlwraith Range, collected 25 August 1998 by K. McDonald & J. Covacevich; QMJ 70609, gravid adult female, McIlwraith Range (13 º 44 ' 01 " S, 143 º 20 ' 09 " E, 530 m a. s. l.), collected 16 August 1999, K. McDonald, A. Freeman & H. Hines.	en	Hoskin, Conrad J., Couper, Patrick J. (2014): Two new skinks (Scincidae: Glaphyromorphus) from rainforest habitats in north-eastern Australia. Zootaxa 3869 (1): 1-16, DOI: http://dx.doi.org/10.11646/zootaxa.3869.1.1
03DEC73D8561FFC5FF3EFF109814B1EF.taxon	diagnosis	Diagnosis. Glaphyromorphus nyanchupinta sp. nov. is diagnosed from all congeners in having: narrowly separated adpressed limbs (not separated by more than the length of the forelimb); more than 24 midbody scale rows; the prefontal separated from the preocular; small body size (max SVL ~ 54 mm); seven supralabials (with 5 th below centre of eye); fewer than 21 lamellae beneath 4 th toe; a strong barred body pattern extending to hindlimbs; dark supralabial scales with a central white dot; dark streaks typically present on throat.	en	Hoskin, Conrad J., Couper, Patrick J. (2014): Two new skinks (Scincidae: Glaphyromorphus) from rainforest habitats in north-eastern Australia. Zootaxa 3869 (1): 1-16, DOI: http://dx.doi.org/10.11646/zootaxa.3869.1.1
03DEC73D8561FFC5FF3EFF109814B1EF.taxon	etymology	Etymology. Nyanchupinta translates as: ‘ nyanchu’ for ‘ dead leaves or mulch’ and ‘ pinta’ for ‘ covered’, referring to the lizard being hidden in the leaf-litter. The species was named by Elders of the Kaantju clan, traditional owners of the McIlwraith Range where the species lives.	en	Hoskin, Conrad J., Couper, Patrick J. (2014): Two new skinks (Scincidae: Glaphyromorphus) from rainforest habitats in north-eastern Australia. Zootaxa 3869 (1): 1-16, DOI: http://dx.doi.org/10.11646/zootaxa.3869.1.1
03DEC73D8561FFC5FF3EFF109814B1EF.taxon	description	Measurements and scale counts of holotype QMJ 85244 (Figs 3 B, 4 B, 8, 9). Mature male with turgid, opaque testes. SVL = 53.0 mm, AG = 26.4 mm, TL not measured due to partial tail loss, L 1 = 10.5 mm, L 2 = 17.0 mm, HL = 10.2 mm, HW = 7.4 mm, NL = 7.9 mm, midbody scale rows = 27, paravertebrals = 60, lamellae 4 th toe = 17, lamellae 4 th finger = 11, supralabials = 7 with 5 th below centre of eye, infralabials = 6, supraciliaries = 6 left / 7 right. Description of type series. Data presented as range followed by mean in brackets (n = 4, unless stated otherwise). Adult measurements (mm): SVL = 49.2 – 53.6 (51.8), AG = 25.4 – 29.5 (27.3), TL = 51.8 (n = 1), L 1 = 10.1 – 11.3 (10.7), L 2 = 14.3 – 17.0 (15.5), HL = 9.2 – 10.2 (9.5), HW = 6.6 – 7.4 (6.9), NL = 7.8 – 9.8 (8.5) (Table 1). Adult proportions (as % SVL): AG = 50 – 55 (53), TL = 100 (n = 1), L 1 = 20 – 21 (21), L 2 = 28 – 32 (30), HL = 18 – 19 (18), HW = 13 – 14 (13), NL = 15 – 18 (16) (Table 1). Body: elongate, tubiform. Neck broad and not well differentiated from back of head. Snout rounded in profile. Limbs short, pentadactyl, and separated by the about the length of the forelimb when adpressed. Scalation: Scales smooth with rounded posterior margins; 25 – 27 (mean = 26.5) rows at midbody; paravertebrals not transversely enlarged (except enlarged nuchals) and numbering 56 – 60 (mean = 58.5) in a line between the parietals and the posterior margin of the hindlimb. Nasals moderate, wellspaced, with a relatively large external naris; rostral and frontonasal in moderate contact; prefrontals large, moderately to very narrowly separated, and not contacting 1 st preocular; frontal contacting frontonasal, prefrontals, first two supraoculars, frontoparietals and usually in point contact with 1 st supraciliary; supraoculars 4, second the largest; supraciliaries 6 – 7 (mean = 6.5), first the largest; frontoparietals paired and distinct from interparietal; parietals in contact behind interparietal; 6 – 7 (mean = 6.3) nuchal scales; primary temporals 1; secondary temporals 2, upper largest and overlapping lower; loreals 2; preoculars 2; presuboculars 2; an enlarged subocular scale penetrating the suture between the 4 th and 5 th supralabials; supralabials 7, 5 th below centre of eye; infralabials 6; postmental contacting two infralabials on each side; lower eyelid scaly; ear opening round or vertically oval, without lobules and with tympanum moderately recessed; lamellae under 4 th finger 10 – 11 (mean = 10.8); lamellae under 4 th toe 17 – 20 (mean = 17.8); medial pair of preanal scales greatly enlarged. Colour pattern in preservative (Fig. 3 B, 4 B, 9): Dorsal ground colour mid brown with narrow, dark brown or dark grey wavy bars extending from vertebral zone to lower flanks. Pattern extends to hindlimbs but generally more prominent on anterior half of dorsum. Laterally the pattern breaks up, becoming increasingly mottled on the posterior third of the body and sides of tail. The temporal region and sides of neck marked with dark reticulations, where dark bars merge together, and the upper labials are predominantly dark, each bearing a pale central dot. Venter immaculate cream except at edge of jawline and chin, where there are dark markings around the scale edges. These are least prominent in QMJ 66642 but extend to the throat in the rest of the type series. Limbs pale brown with a dark reticulated pattern. Colour pattern in life (Figs 5 B, 8). As for spirit specimens but colours richer, particularly on body bars and face, and appearance generally more glossy. Comparison with similar species. Glaphyromorphus nyanchupinta sp. nov. can only be confused with G. othelarrni sp. nov., G. fuscicaudis and G. nigricaudis. It is readily distinguished from all three species by its small body size (max SVL = ~ 54 mm vs> 85 mm), less robust form (WT / SVL 0.04 – 0.06 vs> 0.09); number of subdigital lamellae beneath the 4 th toe (17 – 20 vs generally 20 or more) (Table 1); labial pattern (supralabials predominantly dark, enclosing a central white dot vs supralabials pale with dark barring along sutures) (Fig. 4 B), lateral head and neck pattern (dark reticulations vs dark bars or spots) (Figs 3 B, 5 B); the extent of the body pattern (dark dorsal and lateral bars extend posteriorly to hindlimb vs pattern generally strongest on anterior half of body and breaking up or absent beyond midbody) (Figs 3 B, 5 B); and dark streaks typically present on the throat (Fig. 9) vs throat typically unmarked. Glaphyromorphus nyanchupinta sp. nov. is further distinguished from G. othelarrni sp. nov. in having a proportionately shorter tail (TL / SVL 1.00 vs 1.47 – 1.86); shorter limbs (L 1 / SVL: 0.20 – 0.21 vs 0.22 – 0.26; L 2 / SVL: 0.28 – 0.32 vs 0.33 – 0.41); fewer midbody scale rows (25 – 27 vs 28 – 30); fewer subdigital lamellae beneath the 4 th finger (10 – 11 vs 14 – 15); and fewer supralabial scales (7 with 5 th below centre of eye vs typically 8 with 6 th below centre of eye) (Table 1). It is further distinguished from G. fuscicaudis in having a proportionately larger head (HW / SVL: 0.13 – 0.14 vs 0.12 – 0.13; HL / SVL: 0.18 – 0.19 vs 0.16 – 0.17); fewer midbody scale rows (25 – 27 vs 28 – 30); and generally fewer paravertebral scales (mean 59 vs 64) (Table 1). It also lacks the series of yellow dorsolateral blotches that are prominent in G. fuscicaudis (Figs 5 B, 5 C). It is further distinguished from G. nigricaudis in having fewer paravertebral scales (56 – 60 vs 51 – 56) (Table 1).	en	Hoskin, Conrad J., Couper, Patrick J. (2014): Two new skinks (Scincidae: Glaphyromorphus) from rainforest habitats in north-eastern Australia. Zootaxa 3869 (1): 1-16, DOI: http://dx.doi.org/10.11646/zootaxa.3869.1.1
03DEC73D8561FFC5FF3EFF109814B1EF.taxon	distribution	Distribution. Known only from the uplands of McIlwraith Range, north-east Australia (Fig. 6). All individuals have been collected in the same area, at about 530 m elevation in the headwaters of Peach Ck. The McIlwraith Range is poorly explored and it is likely the species is more widespread in the uplands. Habitat and habits. Found in upland rainforest (Fig. 10). Individuals have been collected from under logs in rainforest. A gravid female (QMJ 70609), with two fully-developed eggs, was collected in mid August.	en	Hoskin, Conrad J., Couper, Patrick J. (2014): Two new skinks (Scincidae: Glaphyromorphus) from rainforest habitats in north-eastern Australia. Zootaxa 3869 (1): 1-16, DOI: http://dx.doi.org/10.11646/zootaxa.3869.1.1
03DEC73D8565FFC2FF3EFD7B9899B614.taxon	discussion	Group defined as having: adpressed limbs overlapping or separated by at most the length of forelimb, 24 or more midbody scale rows, and prefrontal not contacting first preocular scale.	en	Hoskin, Conrad J., Couper, Patrick J. (2014): Two new skinks (Scincidae: Glaphyromorphus) from rainforest habitats in north-eastern Australia. Zootaxa 3869 (1): 1-16, DOI: http://dx.doi.org/10.11646/zootaxa.3869.1.1
