taxonID	type	description	language	source
A9374A20CE685962B8CEFEA4765A2607.taxon	diagnosis	Diagnosis. Differs from its only congener known from the Caucasus, B. lusitanus Verhoeff, 1898, mainly by the opisthomere possessing a smooth rather than striated lateral process, and a slightly rather than strongly bent anterior process.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
A9374A20CE685962B8CEFEA4765A2607.taxon	materials_examined	Material examined. 2 ♂♂, 8 ♀♀ (ZMUM), Georgia: AR Abkhazia, Pitsunda, Bzyb River Valley, meadow with a few Buxus trees, in litter, 8. IV. 1983, SIG leg.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
A9374A20CE685962B8CEFEA4765A2607.taxon	distribution	General distribution. From eastern Poland in the west to southwestern Siberia, Russia and western Kazakhstan in the east (Nefediev et al. 2014, Kime and Enghoff 2017); subendemic to the Russian [= East European] Plain (Wytwer et al. 2009; Zuev 2014).	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
7824503F8F605F0ABECD12713484DCD0.taxon	diagnosis	Diagnosis. Differs from its only congener known from the Caucasus, B. jawlowskii, mainly by the opisthomere possessing a striated rather than smooth lateral process, and a strongly rather than slightly bent anterior process.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
7824503F8F605F0ABECD12713484DCD0.taxon	materials_examined	Material examined (ZMUM). Georgia: 7 ♂♂, 1 ♀, AR Abkhazia, region of Sukhumi, Nizhnyaya Yashtukha, tobacco plantation, 26. V- 19. VI. 1981, A. Markosyan leg.; 1 ♂, Tbilisi, canyon of Vere River, 18. X. 1980, M. Kokhia leg.; 2 ♂♂, 2 ♀♀, eastern Georgia, Akhmeta District, E of Kasristskali, salty swamp, reed thickets, 7. V. 1983, V. Yanushev leg.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
7824503F8F605F0ABECD12713484DCD0.taxon	distribution	General distribution. Subcosmopolitan (Kime and Enghoff 2017, Vagalinski and Lazanyi 2018).	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
0EA6D72F9F2853A294CD29A19FE88E39.taxon	description	Fig. 1	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
0EA6D72F9F2853A294CD29A19FE88E39.taxon	materials_examined	Material examined. Azerbaijan: 1 ♂ (SMNG), Ismayilli District, S of Zərgəran, 40.7310 ° N, 48.3680 ° E, 880 m a. s. l., slope with Corylus, Clematis and some Prunus trees, stone heaps overgrown by moss, mainly in thick litter and under stones, 30. III. 2015, D. Antic and H. Reip leg.; 1 ♀ (SMNG), Ismayilli District, near Qurbanəfəndi, 1.3 road km towards Xanəgah, 40.8614 ° N, 48.1132 ° E, 640 m a. s. l., slope with Fagus and a single Populus tree, in litter, same date and collectors; 1 ♂, 3 ♀♀, 2 juv. (ZMUM), Altiagac National Park, 1050 - 1100 m, mixed broadleaved forest, litter, 20 and 26. IV. 1987, SIG and K. Eskov leg.; many ♂♂, ♀♀, juv. (ZMUM), SW of Quba, 750 m, Fagus, Quercus, Carpinus etc. forest, litter and under bark, 23. IV. 1987, SIG and K. Eskov leg.; 1 ♂, 2 ♀♀ (ZMUM), Shemakha District, Pirkuli, 1700 m, Quercus, Taxus, 22. XI. 1962, T. Perel leg. Georgia: 1 ♂, 2 ♀♀ (SMNG), Mtskheta-Mtianeti, valley of Andaki River, 2.2 km upstream of its confluence with Argun River, 42.5006 ° N, 45.2156 ° E, 1430 m a. s. l., 08. VIII. 2014, F. Walther leg.; 1 ♂, 1 ♀ (SMNG), Mtskheta-Mtianeti, 1 km downstream of Mutso, 42.5006 ° N, 45.2156 ° E, 1530 m a. s. l., 9. VIII. 2014, F. Walther leg.; 1 ♂, 1 ♀ (SMNG), NW of Stepantsminda, 42 ° 40 ' 1 " N, 44 ° 36 ' 49 " E, 2093 m a. s. l., Juniperus, 02. VII. 2019, K. Voigtlaender leg.; 2 ♂♂, 3 ♀♀ (ZMUM), Stepantsminda, 8. VI. 1974, M. Kokhia leg. Russia: 1 ♂ (SMNG 33575), Republic of Adygea, Novoprokhladnoye, 20. V. 2004, K. Voigtlaender leg.; 2 ♂♂, 1 ♀ (ZMUM), Stavropol Province, Kursavka, Acacia and Rosa hedge alongside a road, sifted litter, 30. V. 1982, SIG leg.; 2 ♀♀ (SMNG), Stavropol, Tamanskaya Dacha Forest, near Komsomolskiy Reservoir, 45.0481 ° N, 41.9567 ° E, 530 m a. s. l., deciduous forest (Quercus, Carpinus, Acer), 22. VIII. 2012, F. Walther leg.; many ♂♂, ♀♀ (ZMUM), Stavropol Province, Novoaleksandrovsk, east end, X. 2013, B. Korotyaev leg.; 4 ♂♂, 1 ♀ (ZMUM), Kabardino-Balkar Republic, Chegemsky District, upper course of Chegem River, meadow, under stones, 1500 m, 12. VII. 1986, SIG leg.; 3 ♂♂, 2 ♀♀ (SMNG), Republic of North Ossetia - Alania, Alagirsky District, Borzikau, near Fiagdon River, 1200 m a. s. l., 42 ° 50 ' 37 " N, 44 ° 18 ' 44 " E, under stones, 11. V. 2016, A. S. Sazhiev leg.; 3 ♂♂, 3 ♀♀ (ZMUM), Republic of North Ossetia-Alania, North Ossetian State Nature Reserve, Tsey Valley, 23. VII. 1984, S. Alekseev leg.; many ♂♂, ♀♀, juv. (ZMUM), Chechen Republic, Kharachoy, SE of Vedeno, 950 m, Fagus, Carpinus etc. forest, in litter, under bark and stones, 17. VII. 1986, SIG leg.; many ♂♂, ♀♀, juv. (ZMUM), Chechen Republic, Argun Valley, 5 km N of Shatoy, Corylus, Fagus, Carpinus etc. forest, 750 m, in litter, under stones and bark, 18. VII. 1986, SIG leg.; 4 ♂♂, 7 ♀♀ (ZMUM), 8 ♂♂, 10 ♀♀ (ZMUM), Republic of Dagestan, Kazbekovskiy District, Dylym, Fagus and Carpinus forest, 13. VII. 1985, K. Khajialiyev leg.; 8 ♂♂, 10 ♀♀ (ZMUM), Republic of Dagestan, Almak, Carpinus forest, 15. VII. 1985, K. Khajialiyev leg.; 6 ♂♂, 5 ♀♀ (ZMUM), Republic of Dagestan, Sergokalinskiy District, Murguk, Quercus forest, western slope, 4. III. 1985, K. Khajialiyev leg.; 10 ♂♂, 10 ♀♀, Republic of Dagestan, Kayakentskiy District, Quercus forest, 27. VII. 1985, K. Khajialiyev leg.; 4 ♂♂, 7 ♀♀, Republic of Dagestan, Botlikh Distrikt, Kharami Pass, 2150 m, subalpine meadow, under stones, 16. VII. 1986, SIG leg.; 1 ♂, 1 ♀ (HNHM), Republic of Dagestan, Kurush, 3000 m a. s. l., 8. VII. 1989, Z. Korsos leg.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
0EA6D72F9F2853A294CD29A19FE88E39.taxon	distribution	General distribution. Also known from eastern Ukraine, Crimea, and the Kursk, Orel, Voronezh, Belgorod, Rostov-on-Don, Samara, Volgograd, and Saratov regions in central and southern European Russia (Loksina 1969; Prisnyi 2001). The subspecies B. r. strandschanum (Verhoeff, 1937) occurs in southeastern Bulgaria and northeastern Greece (Lazanyi et al. 2012; Kime and Enghoff 2017). Despite its markedly disjunct distribution in relation to that of the nominate subspecies, until now, no reliable morphological characters to distinguish between the two taxa have been proposed (see also Vagalinski and Lazanyi 2018). It thus seems that the usage of molecular data is the only way to define the nomenclatural faith of B. r. strandschanum.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
A7C20728BA7658348D802957B1DF343E.taxon	description	Figs 2, 3	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
A7C20728BA7658348D802957B1DF343E.taxon	materials_examined	Material examined (ZMUM). Georgia: AR Abkhazia: 2 ♂♂, 1 juv., Gulripsh near Sukhumi, 24. X. 1953, E. Borutzky leg.; 2 ♂♂, 2 ♀♀, Ochamchira District, Jgerda, Kodorskiy Mountain Range, in litter and under bark, 21. IX. 1985, I. A. Ushakov leg.; 2 ♂♂, 3 ♀♀, Pitsunda-Myussera Nature Reserve, Myussera part, 20 - 130 m, mixed deciduous forest (Castanea, Alnus, etc.), in litter, under bark and stones, 8 - 10. IV. 1983, SIG leg.; 1 ♂, 1 ♀, Pskhu-Gumistinskiy Nature Reserve, Gumistinskiy part, cordon Nizhniy Tsumur, 25. IX. 1985, I. A. Ushakov leg.; 3 ♂♂, 1 ♀, 1 juv., Sukhum District, Tsebelda, 300 m a. s. l., Carpinus, Acer and Buxus scrub, in litter, 19. VIII. 1986, SIG leg.; 6 ♂♂, 11 ♀♀, 3 juv., Sukhum District, Lake Amtkelis (Azanta) ca. 16 km N of Tsebelda, 550 m, Alnus forest, litter and under bark, 19. VIII. 1986, SIG leg.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
A7C20728BA7658348D802957B1DF343E.taxon	diagnosis	Diagnosis. A species of Colchiobrachiulus differing from its single known congener, C. montanus sp. nov., by the larger size (both sexes> 30 mm in length and 2 mm in height, vs. males <20 mm in length and 1.3 mm in height, and females <25 mm in length and 2 mm in height in C. montanus sp. nov.), the longer epiproct, and by certain details of gonopodal structure, more specifically, by the broader apex of the promere and its distomesal process significantly outreaching the apex, vs. the same being subequal to the apex in C. montanus sp. nov.; by the apical outgrowth of the opisthomere's basoposterior process being relatively broader, with shorter branches at margin; and by the solenomere being bi- rather than unipartite in its distal section.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
A7C20728BA7658348D802957B1DF343E.taxon	distribution	General distribution. COLC-SWGC.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
D6B023DADDAC5F6B9FFDE384D67F0509.taxon	description	General description. Small to medium-sized (L (males) = 15 - 37 mm) Brachyiulini. Ommatidia present. Ozopores right on or tightly behind pro-metazonal suture at least on more anterior body rings. Epiproct well-developed, from moderately to relatively (not conspicuously) long. Male hypoproct rounded trapezoidal to semi-circular, ventrally with two distal paramedian setae. Male mandibular stipites considerably expanded, broadly rounded, without a distinct anterior / anteroventral corner. Male walking legs ventrally with two well-developed adhesive pads, one each on postfemur and tibia. Penis short and stout, with very short apical lobes and small terminal lamellae. Gonopods: In situ considerably protruding from gonopodal sinus, directed caudoventrad. Promere higher than opisthomere, elongate, bearing a slender distomesal process with a narrow channel connected with a deep distomesal groove on caudal sur face of promere, both designed to envelop the solenomere; median ridge and median groove rather poorly developed; flagellum thin, micro-dentate apically. Opisthomere rather elongate; basoposterior process weakly pronounced, mostly fused to CBO, ending in a finely branched, papillose, freely protruding, apical outgrowth; anterior process long and flattened, partly enveloping the solenomere; an apicoposterior or lateral process, and a mesomeroidal lobe absent; solenomere fine and slender, with a simple tubular structure; with long spiniform filaments along flagellum channel. Vulva: Subcylindrical, mostly symmetrical. Bursa with a distinct, more or less obtuse, postero-apical margin. Opening placed apically on bursa. Operculum subequal in height to bursa. Receptaculum seminis: central tube narrow; posterior tube long and very narrow, somewhat folded; posterior ampulla small to medium-sized.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
0A043F07F4C15996B58F4FC060CC244B.taxon	description	Figs 4, 5	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
0A043F07F4C15996B58F4FC060CC244B.taxon	materials_examined	Material examined (all from Russia, Karachay-Cherkess Republic, Teberda Biosphere Nature Reserve). Holotype: ♂ (ZMUM) (unbroken), Mount Mussa-Achitara in Dombai, 2700 - 2800 m, alpine meadow, under stones, 29. VII. 1986, SIG leg. Paratypes: 1 ♂ (ZMUM) (head to ring 6 and rest of body, gonopods prepared for SEM), 2 ♀♀ (ZMUM) (one unbroken, the other with head to ring 3 and rest of body, left vulva dissected), same collecting data as for holotype; 1 ♂ (ZMUM) (head to ring 2 (head damaged, gnathochilarium separated), rings 3 to 6, and rest of body (half-broken in the middle, gonopods dissected )), 2 ♀♀ (ZMUM) (one unbroken, the other broken into two pieces), Alibek Canyon near Dombai, 2000 - 2100 m, sparse Betula stand, litter, under stones, 25. VIII. 1986, SIG leg.; 1 ♂ (NMNHS) (head to ring 3, ring 4 to ring 6, pleurotergum 7, and rest of body, penis and gonopods dissected), 1 subad. ♂ (NMNHS) (in 2 pieces), Arkhyz part, VI-VII. 1988, A. P. Zolotarev leg.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
0A043F07F4C15996B58F4FC060CC244B.taxon	diagnosis	Diagnosis. A species of Colchiobrachiulus differing from its single known congener, C. dioscoriadis comb. nov., by its smaller size (males <20 mm in length and 1.3 mm in height, mature females <25 mm in length and 2 mm in height, vs.> 30 mm in length and 2 mm in height for both sexes in C. dioscoriadis comb. nov.), by a shorter epiproct, and by gonopodal details, more specifically by the more narrow apex of the promere and its distomesal process being subequal to the apex, vs. that same process significantly outreaching the apex in C. dioscoriadis comb. nov.; by the apical outgrowth of the opisthomere's basoposterior process being relatively more narrow, with longer branches at margin; and by the solenomere being uni- rather than bipartite in its distal section.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
0A043F07F4C15996B58F4FC060CC244B.taxon	description	Description. Measurements: holotype ♂ in S XI, 43 + 1 + T, L = 16 mm, H = 1.2 mm; paratype ♂♂ in S X, 42 - 45 + 1 + T, H = 1.15 - 1.2 mm; paratype ♀♀ in S X-XI, 42 - 45 + 1 + T, L = 18 - 22 mm, H = 1.5 - 1.85 mm. Colouration (after> 30 years of preservation in alcohol) (Fig. 4): Apparently heavily faded, mostly brown; head and collum (Fig. 4 B) with a usual colour pattern; antennae brownish; prozonae dark brown above ozopore level, each with several uneven light spots dorsally, the largest one right above ozopore; metazonae lighter brown-beige; dorsum with a blackish axial line; legs brown; pre-anal ring dark brown, contrasting to the lighter previous ring, paraprocts proximally dark brown, distally lighter. External structures: Eye patches in adults consisting of 35 - 40 ommatidia arranged in easily countable vertical rows. Vertigial, supralabral, and labral setae: two, four, and 18 - 22, respectively. Antennae 1.4 - 1.5 x as long as head in males and 1.1 - 1.2 x in females; antennomere 2> 3> 4 ~ 5> 6. Promentum of gnathochilarium separating lamellae linguales in their proximal 2 / 5 or so, each of the latter with three or four setae in a longitudinal row. Collum mostly smooth, with just two or three shallow grooves at posterolateral corners. Body rings slightly vaulted. Prozonae with very short and shallow, mostly parallel, longitudinal striae. Metazonae not very deeply, but densely striate, n Schub = 10 or 11; setae apparently mostly abraded, ca. 1 / 2 - 2 / 3 of metazonal length. Ozopores relatively large, placed right in pro-metazonal suture in more anterior body rings, and ca. 1 x their diameter in more posterior ones; sutures not sinuous in front of ozopores. Tarsus of mid-body legs ca. 1.2 x as long as tibia and 3 x as long as apical claw. Telson (Fig. 4 C): Epiproct moderately long in males, almost reaching the level of the longest paraproctal setae, considerably shorter and stouter in females, ending with a short and blunt hyaline tip turned slightly dorsad. Hypoproct roundly trapezoid, barely protruding behind rear contour of paraprocts in males, broadly rounded in females, completely concealed under paraprocts in females. Paraprocts densely covered with relatively long setae; without distinct rows of shorter setae along caudal margins. Male sexual characters: Mandibular stipites considerably expanded, protruding mostly ventrad, forming no distinct corner. Leg pair 1 compact, rounded, parallel hooks. Walking legs with crested adhesive pads, both tibial and postfemoral ones gradually reduced posteriad, but still visible until caudalmost pairs. Pleurotergum 7 ventrally forming elongated, narrowly rounded lobes (Fig. 5 A) originating from the zone around pro-metazonal suture, protruding mostly ventro-mesad behind gonopods. Penis (Fig. 5 B) short and stout, with broad and rounded apical lobes ending up in short and blunt terminal lamellae directed distad. Gonopods (5 C, D-H): Promere (Fig. 5 D, p in Fig. 5 C) considerably higher than opisthomere, slender, gradually tapering towards a narrowly rounded apex; mesal margin distally bearing a long and fine process with a pointed tip turned mesad; caudal surface with a short and very weakly pronounced median ridge, a broad and shallow median groove, and a deep and narrow distomesal groove, the latter in situ enveloping the lateral side of solenomeral apex, ending up in an even narrower channel running through distomesal process; flagellum thin, slightly longer than height of promere, with a micro-dentate apical part. Opisthomere (Fig. 5 C, E-H) relatively slender; basoposterior process shaped as a broad and flattened lobe running parallel to CBO, mostly fused to it, ending in a multibranched, coral-like, papillose, freely protruding, apical outgrowth; anterior process well-developed, long and tapering, twisted at> 90 ° around solenomere, partly enveloping the lateral and posterior sides of the latter, both being subequal in height; mesal side with a small lobe (presumably gonocoxal gland), and a rather shallow anteromesal sinus frontal to the gland; two sparse rows of very long, erect, spiniform filaments running along basomedial section of flagellum channel; solenomere very fine and slender, unipartite, sigmoid, without specialised structures apically. Female sexual characters: Leg pairs 1 (significantly) and 2 (slightly) shorter and thicker than following legs. Vulva (Fig. 5 I) rather elongated, subcylindrical, mostly symmetrical (lateral valve somewhat broader than mesal one), with an obtuse postero-apical margin; operculum as high as bursa; both bursa and operculum apically with rather large hyaline protrusions; setation moderately dense throughout. Receptaculum seminis composed of a relatively narrow and moderately long, digitiform, central tube, and a very thin posterior tube, the latter somewhat bent on its way to a medium-sized, ovoid, posterior ampulla.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
0A043F07F4C15996B58F4FC060CC244B.taxon	distribution	General distribution. NWGC.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
09EF99088F85526C8CD88056B0E2E079.taxon	materials_examined	Material examined. 1 ♂ (ZMUM), Georgia: Kutaisi District, 8 km E of Orpiri, environs of Cave Tsutskhvati, deciduous forest, rock, litter, 24. X. 1981, SIG leg.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
09EF99088F85526C8CD88056B0E2E079.taxon	distribution	General distribution. WECA?	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
93A67E5871975DCFA6D1E0F9137738EF.taxon	description	Fig. 6	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
93A67E5871975DCFA6D1E0F9137738EF.taxon	materials_examined	Material examined. 1 ♂, 2 ♀♀ (ZIN), Armenia: Lake Sevan, Elenovka [Sevan], 16. VI. 1927, A. Schelkovnikov leg.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
93A67E5871975DCFA6D1E0F9137738EF.taxon	distribution	General distribution. CECA-LECA.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
2AEB39A4316F57199DAFE2802D43FC35.taxon	description	General description. Small to medium-sized Brachyiulini (L (males) = 11 - 32 mm). Ommatidia present. Ozopores right on or tightly behind pro-metazonal suture at least on more anterior body rings. Epiproct well-developed, markedly long. Male hypoproct broadly rounded, without setae. Male mandibular stipites considerably expanded, broadly rounded, anterior / anteroventral corner either indistinct or broadly rounded. Male walking legs ventrally with two well-developed adhesive pads, one each on postfemur and tibia. Penis short and stout, with indiscernible or very short apical lobes, as well as small and rounded terminal lamellae. Gonopods: In situ protruding from gonopodal sinus only with their apical parts, directed completely ventrad. Promere as high as opisthomere, with a broad base, more or less narrowing distad and bearing a slender, tapering, apicomesal process. Opisthomere from rather stout to slender; a basoposterior process mostly vestigial, i. e., present as a weakly pronounced vertical ridge running parallel to CBO, ending with a simple, lamellar, apical outgrowth protruding nearly perpendicular to CBO; anterior process fine and tapering; a lateral process absent or represented by a small lobe; a mesomeroidal lobe absent; solenomere complex, consisting of a tapering basomesal process and several other small processes and lobes apically. Vulva: Subcylindrical, mostly symmetrical. Bursa with a distinct, slightly obtuse to subrectangular, postero-apical margin. Opening placed right on top of bursa. Operculum shorter than, to subequal to, bursa. Receptaculum seminis: central tube narrow, digitiform; posterior tube very narrow, more or less folded; posterior ampulla small to medium-sized.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
33C3DA18F2BE52A9A8D9D5125BFEEA8F.taxon	description	Figs 7, 8	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
33C3DA18F2BE52A9A8D9D5125BFEEA8F.taxon	materials_examined	Material examined. Azerbaijan: 1 ♀ (SMNG), Lerik District, Hyrcan Nature Reserve, road Lǝnkǝran-Lerik at km 322, 38.7638 ° N, 48.5819 ° E, 400 m a. s. l., small valley, forest of Parrotia with some Quercus, in thick litter, 26. III. 2015, D. Antic and H. Reip leg.; 2 ♂♂, 4 ♀♀, 8 juv. (ZMUM), Lankaran, Istisu W of Astara, forest, 29. VII. 1974, SIG leg.; 6 ♂♂, 5 ♀♀, 2 juv. (ZMUM), Istisu, ca. 8 km SW of Masalli, Quercus, Acer, Carpinus etc., 80 - 140 m, in litter, under bark and stones, 19 - 20. X. 1983, SIG leg.; 2 ♂♂ (ZMUM), Yardimli, Avash, 1200 - 1500 m, 14 - 17. VI. 1996, SIG leg.; 1 ♂, 2 ♀♀ (ZMUM), Astara, Istisu, W of Astara, 100 m, 2 - 6. VI. 1996, W. Schawaller leg.; many ♂♂, ♀♀, juv. (ZMUM), ca. 6 km WSW of Astara, Quercus, Acer, Carpinus, etc., 10 - 30 m, in litter and under bark, 18. III. 1983, SIG leg.; 1 ♂, 1 ♀, 2 juv. (ZMUM), Lankaran, Alekseevka (Avrora), 24. VII. 1974, SIG leg.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
33C3DA18F2BE52A9A8D9D5125BFEEA8F.taxon	diagnosis	Diagnosis. A species of Iraniulus differing from its only known congener, I. tricornis sp. nov., by a larger body (males> 25 mm long and higher than 1.7 mm, females> 30 mm long and ca. 2.5 mm high, vs. males <20 mm long and ca. 1 mm high, females <25 mm long and ca. 1.5 mm high in I. tricornis sp. nov.), by having a proportionately longer, more slender opisthomere, and by certain details of solenomere structure: directed completely distad, with a larger basocaudal process, and with an apical part ending in two short rounded branches of equal size on the mesal side, and one slender sigmoid branch on the lateral side, vs. the same turned somewhat caudad in I. tricornis sp. nov., with a proportionately smaller basocaudal process, and with an apical part bearing three sharply pointed branches of equal size.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
33C3DA18F2BE52A9A8D9D5125BFEEA8F.taxon	distribution	General distribution. Iran, Lahijan (type locality); Azerbaijan, Lenkoran [Lankaran] (Samedov et al. 1972). Apparently, a Hyrcanian endemic.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
D646B4181C065F798324116D94B118CF.taxon	description	Figs 9, 10, 11	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
D646B4181C065F798324116D94B118CF.taxon	materials_examined	Material examined (all from Georgia, Svanetia): Holotype: ♂ (unbroken) (ZMUM), Khumpreri River (left affluent of Enguri), near Dizi, 1.5 - 2 km before the influx, ca. 1000 m a. s. l., leaf litter, 14. IX. 1986, A. Ryvkin leg. Paratypes: 6 ♂♂ (ZMUM) (one in 4 pieces, gonopods prepared for SEM, leg pair 2 and penis dissected; another in head and 2 pieces, gonopods in situ, the rest unbroken), 1 ♂ (NMNHS) (in 2 pieces with dissected gonopods), 1 ♂ (NHMD) (unbroken), 1 ♂ (IBER) (in 2 pieces, gonopods dissected, one flagellum prepared for SEM), 3 ♀♀ (ZMUM) (unbroken), 1 ♀ (NMNHS) 1 ♀ (NHMD), 1 ♀ (IBER) (all unbroken), 2 juv. (unbroken), same collecting data as for holotype; 3 ♂♂ (one in head to ring 3, ring 4 - 6, pleurotergum 7, and rest of body, gonopods and leg pair 2 dissected; the others unbroken), 2 adult ♀♀ (one in head to ring 3 and rest of body, vulvae dissected; the others unbroken), 5 juv. ♀♀ (unbroken) (all in ZMUM), ca. 50 km west of Mestia, west of Dizi, ca. 800 m a. s. l., by a waterfall, in leaf litter, 19. IX. 1986, A. Ryvkin leg.; 2 ♂♂ (one in head to ring 6 and rest of body, gonopods dissected; the other unbroken), 3 juv. (unbroken) (all in ZMUM), Mestia, 1500 m, Betula and Rhododendron on moraine, litter and under stones, 5 - 16. IX. 1986, SIG leg.; 1 ♂ (in head to ring 6 and rest of body), 2 ♀♀, 1 juv. ♂, 2 juv. ♀♀ (all in ZMUM), same locality as for holotype, 9. IX. 1986, A. Ryvkin leg.; 6 ♂♂ (one in 3 pieces, gonopods dissected, the others unbroken), 8 ♀♀ (two in 2 pieces, the rest unbroken), 1 juv. ♂, 4 juv. ♀♀ (all in ZMUM), Mestia, 1500 m a. s. l., litter and under stones, 22. X. 1979, SIG leg.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
D646B4181C065F798324116D94B118CF.taxon	diagnosis	Diagnosis. A species of Iraniulus differing from its only known congener, I. fagorum, by a smaller body: males <20 mm long and ca. 1 mm high, females <25 mm long and ca. 1.5 mm high, vs. males> 25 mm long and higher than 1.7 mm, females> 30 mm long and ca. 2.5 mm high in I. fagorum), by having a proportionately shorter and stouter opisthomere, and by details of the solenomere: turned somewhat caudad, with a proportionately smaller basocaudal process, and with an apical part bearing three sharply pointed branches of equal size, vs. the same directed completely distad in I. fagorum, with a larger basocaudal process, and with an apical part ending in two short, rounded branches of equal size on the mesal side and one slender sigmoid branch on the lateral side.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
D646B4181C065F798324116D94B118CF.taxon	description	Description. Measurements: holotype ♂ in S IX, 43 + 2 + T, L = 16 mm, H = 1.05 mm; paratype ♂♂ in S VII-X, 38 - 43 + 2 - 3 + T, L = 11 - 13 mm, H = 0.85 - 1.1 mm; paratype ♀♀ in S VII-X, 38 - 45 + 1 - 2 + T, H = 1.2 - 1.6 mm, L = 14 - 21 mm. Colouration: (after> 30 years in alcohol) (Fig. 9): mostly brown-beige; head with the usual colour pattern; antennae light brown, collum brownish, margins dark brown; prozonae dorsally with a broad, dark brown, transverse stripe next to suture, and with irregular dark brown spots just before ozopores; metazonae ochre-brown; dorsum with a continuous, blackish, axial line; legs light brown; telson brown-grey, caudal parts of paraprocts lighter. External structures: Eye patches in adults consisting of 20 - 38 ommatidia arranged in hardly traceable vertical rows. Vertigial, supralabral, and labral setae: two, four, and 17 - 20, respectively. Antennae 1.5 - 1.6 x as long as head in males and ca. 1.4 x in females; antennomere 2> 5 ≥ 3 ~ 4> 6. Gnathochilarium with promentum separating both lamellae linguales in their basal 1 / 3 - 2 / 5, each with three setae in a longitudinal row. Collum mostly smooth, with only 2 - 3 short striae near posterolateral corners. Body rings slightly vaulted. Prozonae with very short, shallow, longitudinal striae in posterior parts. Metazonae moderately deeply striated, n Schub = 7 or 8 in males and 9 or 10 in females; metazonal setae from 2 / 5 (in more anterior rings) to 1 / 2 (in more posterior rings) of metazonal length. Ozopores placed right on pro-metazonal suture in first several rings, gradually taking a more posterior position to ~ 1.5 their diameter in caudalmost rings; sutures sinuous in front of ozopores in some rings. Tarsus of mid-body legs slightly shorter than tibia and ca. 3 x as long as apical claw. Telson (Fig. 9 B): Epiproct very long, straight, ending in a fine, tapering, hyaline tip surpassing the longest paraproctal setae in both sexes; with several long setae. Hypoproct broadly rounded in both sexes, somewhat protruding behind rear margin of paraprocts in males, tightly fitting under their ventral side in females; without setae. Paraprocts covered with sparse long setae, but without distinct rows of shorter setae along caudal margins. Male sexual characters: Mandibular stipites (in Fig. 9 A) moderately expanded, forming a broadly rounded antero-ventral corner. Leg pair 1 compact and set parallel to hooks turned slightly mesad. Walking legs with crested adhesive pads, both tibial and prefemoral ones reduced towards telson, but still discernible in caudalmost legs; femora without modifications. Pleurotergum 7 ventrally forming relatively small shovel-like lobes (Fig. 10 A) originating mostly from metazona, directed largely ventrad. Penis (Fig. 10 B) stout, compact, nearly as broad as long, without differentiated apical lobes, but with broad and rounded terminal lamellae directed ventrad. Gonopods (Fig. 11): Promere (Fig. 11 B, p in Fig. 11 A) as high as opisthomere, broadest at base, somewhat narrowed distad; mesal and lateral margins gently sigmoid; apex laterally broadly rounded, mesally bearing a horn-like apicomesal process bent caudolaterad; caudal surface with a very strongly developed median ridge, a somewhat less strongly protruding lateral ridge running all the way to the top, and a relatively deep and broad median groove; a small, rounded, distomesal lobe directed mesad; flagellum somewhat longer than height of promere. Opisthomere (Fig. 11 A, C-E) relatively thick and stout; anterior process fine and tapering, bent mesocaudad; apical outgrowth of basoposterior process well-developed, protruding at nearly 90 ° to CBO; a faint lobe distolaterally; mesal side with a large membranous lobe, probably gonocoxal gland, at the base of seminal channel, and a deep and narrow anteromesal sinus just frontally to the membranous lobe; with a group of spiniform filaments in distal section of flagellum channel; solenomere consisting of a tapering basomesal process bent caudad and terminating both flagellum and seminal channels, a fine basocaudal process directed meso-caudad, and an apical part ending with three more or less equally sized and shaped pointed branches. Female sexual characters: Leg pairs 1 and 2 somewhat thicker and shorter than following legs. Vulva (Fig. 10 C) roughly cylindrical, strongly transversely compressed; bursa with a well-pronounced, slightly obtuse, postero-apical margin; opening small, positioned at the very top of bursa; operculum subequal to bursa, both bursa and operculum apically bearing large hyaline protrusions; setation scattered over vulval surface. Receptaculum seminis consisting of a narrow, finger-shaped, central tube, and a very narrow, long, somewhat folded, posterior tube leading to an ovoid posterior ampulla.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
D646B4181C065F798324116D94B118CF.taxon	distribution	General distribution. SWGC.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
C8EA91933E9F516D92320C6E98964E73.taxon	diagnosis	Diagnosis. Differs from its only congener known from the Caucasus, M. (s. str.) spathulatum (Lohmander, 1936), mainly by the colour pattern: body uniformly dark with an orange to dark red mid-dorsal line, vs. body with a light yellow to ochre dorsum divided by a black axial line in M. spathulatum; and by the promere having mostly parallel side margins, ending with a flat apex, vs. the same significantly tapering all the way to a narrowly rounded apex in M. spathulatum.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
C8EA91933E9F516D92320C6E98964E73.taxon	distribution	General distribution. Central and southwestern parts of the Balkan Peninsula, northwestern Caucasus.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
37FB26D6CA345F2EB479DF84D78FF828.taxon	materials_examined	Material examined (SMNG): Russia: RU 12 - 86, 1 ♂, 1 ♀, Republic of Adygea, Kamennomostkiy, northern end of Dakhovskaya Canyon near quarry, limestone cliffs, 44.2844 ° N, 40.1778 ° E, 430 m a. s. l., 24. VIII. 2012, F. Walther leg.; RU 12 - 051, 3 ♂♂, 1 juv., Dakhovskaya, limestone cliffs NW of, 44.2439 ° N, 40.1758 ° E, 800 m a. s. l., 13. VIII. 2012, F. Walther leg.; 33579, 1 ♂, Republic of Adygea, Mount Shibaba, 21. V. 2004, K. Voigtlaender leg.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
37FB26D6CA345F2EB479DF84D78FF828.taxon	diagnosis	Diagnosis. Differs from its only congener known from the Caucasus, M. (s. str.) Megaphyllum hercules, mainly by the colour pattern: body with a light yellow to ochre dorsum divided by a black axial line, vs. body uniformly dark with an orange to dark red mid-dorsal line in M. hercules; and by the promere significantly tapering all the way to a narrowly rounded apex, vs. the same having mostly parallel side margins, ending with a flat apex in M. hercules.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
37FB26D6CA345F2EB479DF84D78FF828.taxon	distribution	General distribution. WCIS?	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
3CE8E8A8686E5BE1AB5C1625F97C3E0F.taxon	materials_examined	Material examined. Georgia: AR Ajara: 1 ♂ (SMNG), Machakhlispiri, 2.7 km upstream of Acharistskali, 41.5278 ° N, 41.7211 ° E, 29. IX. 2011, F. Walther leg.; 6 ♂♂, 8 ♀♀, 4 juv. (ZMUM), Keda Municipality, Makhuntseti, Platanus grove, in leaf litter, 9. X. 1981, SIG leg.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
3CE8E8A8686E5BE1AB5C1625F97C3E0F.taxon	diagnosis	Diagnosis. A species of Omobrachyiulus differring from congeners mainly by the solenomere being apically bifurcate into two pointed branches of nearly equal size and shape, in combination with the complete absence of an anterior process on the opisthomere.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
3CE8E8A8686E5BE1AB5C1625F97C3E0F.taxon	distribution	General distribution. LECA.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
B2D33B9F8E9B505D9D498D7874C35D48.taxon	description	Figs 18, 19, 20	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
B2D33B9F8E9B505D9D498D7874C35D48.taxon	materials_examined	Material examined (all from Georgia). Holotype: ♂ (unbroken) (ZMUM), Manglisi, Quercus forest, 12. XI. 1984, E. Kvavadze leg. Paratypes: 2 ♂♂ (ZMUM) (one in head to ring 2, ring 3 to ring 6, pleurotergum 7, and rest of body; right antenna, leg pair 2, penis, legs 6, 7, and mid-body leg dissected, opisthomeres and left promere prepared for SEM; the other unbroken with removed hypoproct), 3 ♀♀ (ZMUM), same collecting data as for holotype; 1 ♂ (ZMUM) (partly broken in anterior section), 2 ♀♀ (ZMUM) (one unbroken, the other in 3 pieces), AR Ajara, E of Khulo, decidous forest near spring, rock, litter, 10. X. 1981, SIG leg.; 2 ♂♂ (ZMUM) (one in three pieces, with dissected gonopods, the other with a missing posterior body half), 1 ♂ (NHMD) (unbroken), 1 ♂ (NMNHS) (in two pieces), 1 ♀ (NMNHS) (unbroken), Kutaisi District, Sataplia Nature Reserve, forest litter and under stones, 25. X. 1981, SIG leg.; 1 ♂ (ZMUM) (in three pieces, with dissected gonopods), 1 ♂ (IBER) (unbroken), 5 ♀♀ (ZMUM) (unbroken), 7 juv. (ZMUM) (two unbroken, the rest in 2 or more fragments), AR Ajara, Khulo, 900 m, forest of Quercus, Abies, Alnus, litter, 11. X. 1981, SIG leg.; 2 ♂♂, 3 ♀♀ (SMNG), Georgia, Samtskhe-Javakheti, 17. IX. 2011, Bakuriani, Tskhratskaro Pass, 41.7067 ° N, 43.5058 ° E, 2080 m a. s. l., 17. IX. 2011, F. Walther leg.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
B2D33B9F8E9B505D9D498D7874C35D48.taxon	diagnosis	Diagnosis. A species of Omobrachyiulus most similar to O. hortensis by the promere significantly outreaching the opisthomere, the latter possessing a massive lobe-like basoposterior process forming two distinct corners, a basal and a distal one, and having a micro-spiculate mesal side, and a unipartite solenomere with a slender rod-like ending. Differs from O. hortensis mainly by the broad, unipartite, collar-shaped, apical outgrowth of the basoposterior process dentate at margin, that same outgrowth being clearly tripartite in the latter species, consisting of a mesal lamellar, a median fan-like, and a lateral spiniform part.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
B2D33B9F8E9B505D9D498D7874C35D48.taxon	description	Description. Measurements: holotype ♂ in S IX, 45 + 1 + T, L = 17 mm, H = 1.4 mm; paratype ♂♂ in S IX, 44 - 46 + 1 + T, L = 16 - 20 mm, H = 1.35 - 1.55 mm; paratype ♀♀ in S IX, 44 - 46 + 1 + T, L = 17.5 - 22 mm, H = 1.7 - 1.9 mm. Colouration (most specimens considerably faded in ethanol) (Fig. 18): Head light brown with the usual dark band between eye patches, distal part of male mandibular stipites saturated dark brown, contrasting to the much lighter proximal part; body mostly light brown, lighter ventrally; prozonae dorsally with dark grey to blackish transverse stripes; each metazona with a blurred dark brown band encircling entire ring; dorsum with a black axial line. External structures: Eye patches in adults consisting of 30 - 35 ommatidia, usually arranged in easily countable vertical rows. Vertigial, supralabral, and labral setae: two, four, and 19 - 23, respectively. Antennae (in Fig. 18 B) ca. 1.6 - 1.7 x as long as head in males and 1.1 - 1.4 x in females; antennomere 2> 3 = 4 = 5> 6. Gnathochilarium with an elongated promentum, the latter separating both lamellae linguales> halfway; four setae in a longitudinal row on each lamella. Collum completely smooth. Body rings barely vaulted. Prozonae with short and very shallow longitudinal striae spread across whole surface. Metazonae moderately deeply striated, n Schub = 7 or 8 in males and 9 or 10 in females; metazonal setae rather sparse, 2 / 3 (in middle to caudal parts of body) - 3 / 4 (in frontal part of body) of metazonal length. Ozopores set tightly behind pro-metazonal suture in more anterior rings, gradually moved further backwards, to nearly equal to their diameter behind the suture in caudalmost rings; sutures straight to gently sinuous in front of ozopores. Tarsus of a mid-body leg slightly shorter than to equal to tibia, and nearly 3 x as long as apical claw. Telson (Fig. 18 C): Epiproct moderately long, stout, shorter than, or barely reaching, the level of longest paraproctal setae, ending bluntly, without a distinct, or with a poorly defined, hyaline tip; with several long setae dorsolaterally. Hypoproct (Fig. 19 A) in males very broad, subrectangular, its margin with three large median teeth and two lower protuberances laterally on each side, the latter being sometimes serrate; in females broadly rounded, tightly adhering under paraprocts; ventral surface sparsely setose. Paraprocts moderately densely setose, without distinct row of shorter setae along caudal margins. Male sexual characters: Mandibular stipites (in Fig. 18 B) strongly expanded, forming an acute anterior corner. Leg pair 1 compact rounded hooks, slightly turned against one another. Leg pair 2 and sometimes 3 slightly thicker than following pairs, adhesive pads rather weakly developed, becoming better developed towards mid-body, postfemoral ones disappearing in caudal third of body, tibial ones present until last leg pairs; femora from leg pair 5 to mid-body legs distally with a small shallow pit (arrow in Fig. 19 B). Pleurotergum 7 considerably enlarged, ventrally forming large and rounded lobes (Fig. 19 C) originating mostly from metazona, protruding caudoventrad behind gonopods. Penis (Fig. 19 D) very short, slightly broader than long, compressed anterocaudally, with very short apical lobes ending in rounded terminal lamellae directed distad. Gonopods (Figs 19 E, F, 20): In situ considerably protruding from gonopodal sinus. Promere (Fig. 20 B, p in 20 A) considerably higher than opisthomere, slender; mesal margin mostly straight, lateral margin parallel to mesal one in proximal section, distally steeply slanting towards a blunt apex; caudal surface with a short, well-pronounced, median ridge, and a broad and rather shallow median groove; with a distinct, deep, distal groove, and a smooth, rounded, distomesal lobe directed caudad; flagellum slightly longer than height of promere. Opisthomere (Figs 19 E, F, 20 A, C-E) compact and robust; basoposterior process represented by a massive lobe with a micro-spiculate mesal surface, distally forming a very broad apical outgrowth bent anteriad, tightly adhering to caudal and partly to mesal side of solenomere, with several teeth / horns at margin; anterior process shaped as a lamellar ridge with a slightly serrate anterior margin; mesomeroidal lobe broad and rounded, moderately developed; a rounded hump-like lobe on lateral side; mesal side with a large lobe (presumably gonocoxal gland) leading to the seminal channel, and a large anteromesal sinus frontally to the gland; solenomere unipartite, mostly straight, relatively broad, apically with a fine digitiform process; a group of very long, erect, spiniform filaments in proximal section of flagellum channel. Female sexual characters: Leg pairs 1 and 2 shorter, leg pair 2 also thicker, than following legs. Vulva (Fig. 19 G) mostly symmetrical (lateral valve somewhat more oblique than mesal one), rather elongate, slightly compressed on sides; bursa with an obtuse postero-apical margin; operculum shorter than bursa, both ending up with well-developed hyaline protrusions; setation denser on bursa than on operculum. Receptaculum seminis consisting of a long and mostly straight central tube with a somewhat enlarged bottom part, and a long and thin posterior tube forming several spiral twists on its way to an ovoid posterior ampulla.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
B2D33B9F8E9B505D9D498D7874C35D48.taxon	distribution	General distribution. LECA-COLC.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
A919AB30AC4C57A9948BD8175C596AE3.taxon	description	Fig. 12	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
A919AB30AC4C57A9948BD8175C596AE3.taxon	materials_examined	Material examined. Georgia: Holotype: juvenile ♀ (by monotypy) (ZMB 748), Borjomi (Karsch 1881). Armenia: 1 ♂, 1 ♀, 4 juv. (ZMUM), Byn Kulp (?), 15. VIII. 1927, M. Makaryan leg.; 3 ♂♂, 5 ♀♀, 1 juv. (ZMUM), halfway between Alaverdi and Bagratashen, Carpinus forest, in litter, 24. V. 1987, SIG and K. Eskov leg.; many ♂♂, ♀♀, juv. (ZMUM), Ijevan, deciduous bush, in litter, 16. IV. 1983, SIG leg.; 1 ♂, 1 juv. (ZMUM), Odzun, W of Alaverdi, 1500 - 1550 m, Quercus, Fagus, Carpinus etc. forest, in litter and under stones with ants, 23 - 24. V. 1987, SIG and K. Eskov leg.; 2 ♂♂, 1 ♀, 7 juv. (ZMUM), Stepanavan, 1600 - 1650, Quercus, Fagus, Carpinus etc. forest, in litter and under bark, 21 - 22. V. 1987, SIG and K. Eskov leg. Azerbaijan: 1 ♂, 1 ♀, (SMNG), Ismayilli District, S of Zərgəran, 40.7310 ° N, 48.3680 ° E, 880 m a. s. l., slope with Corylus, Clematis and some Prunus trees, stone heaps overgrown by moss, mainly in thick litter and under stones, 30. III. 2015, D. Antic and H. Reip leg.; many ♂♂, ♀♀, juv. (ZMUM), Shamakhi District, Pirkuli, near observatory, 1200 - 1250 m, Quercus, Acer, Taxus etc. forest, litter, 30. IV. 1987, SIG and K. Eskov leg.; 2 ♂♂, 3 ♀♀, 1 juv. (ZMUM), Talysh Mts, Djoni, 1500 m, 28 - 29. V. 1976, V. G. Dolin leg.; 2 ♂♂, 4 ♀♀, 3 juv. (ZMUM), NW of Bash Layisqi, ca. 20 km NNW of Sheki, 1250 m, Fagus, Carpinus, Acer etc. forest, 3. V. 1987, in litter, SIG and K. Eskov leg.; 2 ♂♂, 4 ♀♀ (ZMUM), 9 juv., ca. 5 km N of Kutkashen, 1150 - 1200 m; Fagus and Carpinus forest, in litter and rotten wood, 2. V. 1987, SIG and K. Eskov leg.; ♂, ♀ (ZMUM), Altyagach National Park, 1050 - 1100 m, mixed broadleaved forest, litter, 20 and 26. IV. 1987, SIG and K. Eskov leg.; 1 ♂ (ZMUM), Zagatala (= Zakataly) Nature Reserve, Agkemal, 1800 - 2100 m, forest, 24 - 27. V. 1981, SIG and J. Martens leg.; many ♂♂, ♀♀, juv. (ZMUM), Laza, 2200 m, 10. VII. 1985, H. Aliyev leg.; 3 ♂♂, 4 ♀♀ (ZMUM), Istisu, ca. 8 km SW of Masally, Quercus, Acer, Carpinus, etc., 80 - 140 m, litter, under bark and stones, 19 - 20. X. 1983, SIG leg.; many ♂♂, ♀♀, juv. (ZMUM), Azerbaijan, above Agsu, 120 km W of Baku, 900 m, Quercus, shrub, 22. V. 1981, SIG and J. Martens leg.; Georgia: 1 ♂, 1 ♀, Borjomi District, Akhaldaba, 1000 m, Nedzura River Valley, Picea, Carpinus and Fagus forest, in litter, under logs, 12. V. 1983, SIG leg.; many ♂♂, ♀♀, juv. (ZMUM), S of Bakuriani Pass, Tskhratskaro, 2100 m, sparse Betula forest at timberline, 13. V. 1983, SIG leg.; many ♂♂, ♀♀, juv. (ZMUM), Lagodekhi, Fagus, Carpinus etc. forest, XI. 1967, T. S. Perel leg.; many ♂♂, ♀♀, juv. (ZMUM), AR Ajara, 15 km W of Adigeni, Abies, Picea, Fagus, Acer forest, 1500 - 1700 m, litter, logs and under stones, 14 - 15. V. 1983, SIG leg.; many ♂♂, ♀♀, juv. (ZMUM), Mayakovskii District, above Sairme, 1700 - 2000, 1. V. 1985, E. Kvavadze leg.; 3 ♂♂, 5 ♀♀ (ZMUM), Babaneuri Nature Reserve, ENE of Akhmeta, near Babaneuri, 500 m, 4 - 5. V. 1987, SIG and K. Eskov leg.; 2 ♂♂ (ZMUM), Magalakhari Pass, between Akhmeta and Tianeti, Fagus and Carpinus forest, 1200 m, in litter and under bark, 6. V. 1987, SIG and K. Eskov leg.; many ♂♂, ♀♀, juv. (ZMUM), N of Kvareli, 700 - 750 m, Fagus, Carpinus, Quercus etc. forest, in litter and under bark, 4. V. 1987, SIG and K. Eskov; many ♂♂, ♀♀, juv. (ZMUM), Batsaro Nature Reserve, ca. 20 km N of Akhmeta, Fagus, Castanea etc. forest, 800 - 850 m, in litter, 5 - 6. V. 1987, SIG and K. Eskov leg.; 1 ♂, 1 ♀ (ZMUM), Mariamjvari Nature Reserve, ENE of Sagarejo, 1150 - 1250 m, Fagus, Carpinus, Acer, Pinus etc. forest, in litter, under bark and stones, 13 - 14. V. 1987, SIG and K. Eskov leg.; 1 ♂, 7 juv. (ZMUM), Saguramo Nature Reserve, NE of Mtskheta, Zedazeni, 1100 - 1200 m, Fagus, Carpinus, Acer etc. forest, litter and under bark, 20. VII. 1987, SIG and K. Eskov leg.; many ♂♂, ♀♀, juv. (ZMUM), Racha, Oni District, N of Utsera, decidous forest near spring, rock, litter, 20. IX. 1981, SIG leg.; 2 ♂♂, 2 ♀♀ (ZMUM), 45 km W Mestia, above Nakra, 1700 m, Abies, Picea etc., forest, in litter and under stones, 3. IX. 1986, SIG leg.; many ♂♂, ♀♀, juv. (ZMUM), above Sairme, 1700 - 2000 m, 1. V. 1985, E. Kvavadze leg; many ♂♂, ♀♀, juv. (SMNG), NW of Abano, 42.6115 ° N, 44.3742 ° E, 2270 m a. s. l., 06. VII. 2019, K. Voigtlaender leg.; 1 ♂ (SMNG), Georgia, Telavi, in a house garden, 41.9240 ° N, 45.4670 ° E, 09. VII. 2019, K. Voigtlaender leg.; many ♂♂ and ♀♀ (SMNG), Tusheti, Pirikita Alazani Valley, close to Girevi, 42 ° 29 ' 42.1 " N, 45 ° 28 ' 43.2 " E, stony pasture at river bank, 2010 m a. s. l, 6. IX. 2009, F. Walther leg.; many ♂♂, ♀♀, juv. (SMNG), Samtskhe-Javakheti, 500 m W of Gujarula Valley, 41.3078 ° N, 43.4672 ° E, 1010 m a. s. l, 17. IX. 2011, F. Walther leg.; 2 ♂♂, 3 ♀♀, 1 juv. (SMNG), Imereti, Zekari Pass, 41.8522 ° N, 42.8078 ° E, 2200 m a. s. l., 22. VIII. 2014, F. Walther leg.; 1 ♂ (SMNG), Khevsureti, Churo Valley, near mouth of Khone Skali River, 42 ° 34 ' 05.8 " N, 45 ° 13 ' 56.2 " E, 10 - 12. IX. 2009, F. Walther leg.; 1 ♂, 1 ♀ (SMNG), Mtshkheta-Mtianeti, 1.1 km SW of Kobi, 42.5553 ° N, 44.4973 ° E, 2060 m a. s. l., 12. VIII. 2014, F. Walther leg.; 2 ♂♂ (SMNG), Mukhli, bridge over Rioni River, 100 m from the village; 42.5436 ° N, 43.2325 ° E, 640 m a. s. l., 9. X. 2011, F. Walther leg.; 1 ♂ (SMNG), Tusheti, Diklo, 42 ° 23 ' 24.0 " N, 45 ° 40 ' 41.8 " E, 2100 m a. s. l., strongly eroded place near stream, 4. IX. 2009, collector?; 1 ♂, 3 ♀♀ (SMNG), Akhaltsikhe to Borjomi, S of Chobiskhevi, near Chobiskhevi River, 42.7651 ° N, 43.3265 ° E, mountain Fagus forest, VII. 2011, L. Mumladze leg.; 1 ♂, 2 ♀♀ (SMNG), N of Lagodekhi, Lagodekhi Nature Reserve, Lagodekhiskhevi River, mixed forest, VII. 2013, L. Mumladze leg.; Russia: 1 ♂ (SMNG), Stavropol, Tamanskaya Dacha Forest, near Komsomolskiy Reservoir, 45.0481 ° N, 41.9567 ° E, 530 m a. s. l., deciduous forest (Quercus, Carpinus, Acer), 22. VIII. 2012, F. Walther leg.; 2 ♂♂ (ZMUM), Stavropol, Russkiy Forest, 6. VII. 2013, R. V. Zuev leg. and det.; many ♂♂, ♀♀, juv. (ZMUM), Stavropol Province, E of Novopavlovsk, Quercus forest, Salix, Alnus etc. along stream, 28. V. 1982, SIG leg.; many ♂♂, ♀♀, juv. (ZMUM), Republic of Dagestan, Sergokalinskiy District, Murguk, mixed forest, near a river, 10. VIII. 1985, K. Khajialiyev leg.; many ♂♂, ♀♀, juv. (ZMUM), Republic of Dagestan, Samur Valley, Garakh ca. 35 km SW of Magaramkent, 700 - 800 m, Crataegus, Quercus, Acer, Rosa etc. scrub, in litter, 23. X. 1987, SIG leg.; 4 ♂♂, 3 ♀♀, 2 juv. (ZMUM), Republic of Dagestan, Chiragchay Valley, Khiv, 900 - 950 m, Carpinus, Fagus, Corylus, Rhododendron etc. forest., in litter and under stones, 24. X. 1987, SIG leg.; 2 ♂♂, 1 ♀, 1 juv. (HNHM), Republic of Dagestan, Kurush, 2400 m a. s. l., under stones, 9. VII. 1989, Z. Korsos leg.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
A919AB30AC4C57A9948BD8175C596AE3.taxon	diagnosis	Diagnosis. A species of Omobrachyiulus differing from congeners mainly by the opisthomere having a thumb-like apical outgrowth of the basoposterior process, bent somewhat anteriad, and by the solenomere ending with a fine, short, sharply pointed, apical process.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
A919AB30AC4C57A9948BD8175C596AE3.taxon	distribution	General distribution. Also found in Russia, Kalmykia (Golovatch and Matyukhin 2011), the Hyrcan area of Azerbaijan, northern and northwestern Iran, northeastern Turkey, and the island of Thassos, Greece (the subspecies O. C. thassensis).	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
5F498907D49D5224927DDB24CA8A282B.taxon	materials_examined	Material examined. Georgia: 1 ♂ (SMNG), Mtshkheta-Mtianeti, forest SW of Lake Bazaleti, 42.0308 ° N, 44.6653 ° E, 910 m a. s. l., 4. X. 2011, F. Walther leg.; 1 ♂ (ZMUM), Racha-Lechkhum-Kvemo Svaneti Planned National Park, 10 km NE of Shovi, Gurshevi near Mamisoni Pass, 2000 - 2200 m, Abies, Fagus, Alnus, litter and under stones, at a spring, 21. X. 1981, SIG leg.; 1 ♂ (ZMUM), AR Abkhazia, Pitsunda-Myussera Nature Reserve, Myussera part, 20 - 130 m, mixed deciduous forest (Castanea, Alnus etc.), in litter, under bark and stones, 8 - 10. IV. 1983, SIG leg. 1 ♂ (ZMUM), AR Abkhazia, environs of Lake Ritsa, in litter, mixed forest, 24. X. 1978, SIG leg.; 2 ♂♂, 6 ♀♀ (ZMUM), AR Abkhazia, Myussera Forest, 13. VI. 1978, V. Dolin leg.; 1 ♂ (ZMUM), AR Abkhazia, Ochamchira District., near Otapi, at the entrance to Cave Golova Otapa, II. 1983, S. Smirnov leg.; 1 ♂, 3 ♀♀ (ZMUM), AR Ajara, Batumi Botanical Garden, 15 - 20 m, 30. V- 7. VI. 1981, SIG and J. Martens leg. Russia: 1 ♂ (SMNG), Republic of Adygea, Lagonakskiy Ridge, limestone cliffs on the SW slope of Mount Gora Matuk, 44.1061 ° N, 39.9225 ° E, 1810 m a. s. l., 16. VIII. 2012, F. Walther leg.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
5F498907D49D5224927DDB24CA8A282B.taxon	diagnosis	Diagnosis. A species of Omobrachyiulus differing from congeners mainly by the opisthomere bearing a large, flattened, roughly diamond-shaped, apical outgrowth of the basoposterior process, and a shortly bifurcate solenomere, with the anterior branch being clavate, and the posterior one fine and pointed, bent anteriad.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
5F498907D49D5224927DDB24CA8A282B.taxon	distribution	General distribution. CAUC.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
1D8AA424F53D5D0D98AA18B0CB907110.taxon	description	Fig. 13	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
1D8AA424F53D5D0D98AA18B0CB907110.taxon	materials_examined	Material examined. Armenia: 3 ♂♂, 2 ♀♀ (ZMUM), " Prov. Zori, Mount Polat ", 30. VIII. 1925, A. Schelkovnikov leg. [probably H. Lohmander det.]. Georgia: 1 ♂ (SMNG), Khevsureti, ca. 200 m below Shatili, 42 ° 39 ' 34.8 " N, 45 ° 09 ' 55.0 " E, 1415 m a. s. l., 12 - 15. IX. 2009, F. Walther leg.; 1 ♂, 4 ♀♀ (SMNG), Guria, Chkhakoura, towards Bakhmaro, 41.8797 ° N, 42.3511 ° E, 1780 m a. s. l., 21. VIII. 2014, F. Walther leg.; many ♂♂, ♀♀, juv. (SMNG), Samtskhe-Javakheti, Mtsvanne Monastery, 41.8094 ° N, 43.3122 ° E, 840 m a. s. l., 24. VIII. 2014, F. Walther leg.; 4 ♂♂, 3 ♀♀, 3 juv. (SMNG), Samegrelo-Zemo Svaneti, 3 km NE of Ushguli, 42.9347 ° N, 43.0358 ° E, 19. VIII. 2014, F. Walther leg.; 1 ♂, 7 ♀♀ (SMNG), Samtskhe-Javakheti, Bakuriani, ca. 9 km to Tskhratskaro, Ugheltekhili Pass, 41.7070 ° N, 43.5028 ° E, 2210 m a. s. l., 22. VIII. 2014, F. Walther leg.; 1 ♂ (SMNG), Imereti, Sairme, towards Zekari Pass, 41.8572 ° N, 42.7894 ° E, 1920 m a. s. l., 22. VIII. 2014, F. Walther leg.; 1 ♂ (SMNG), Mtshketa-Mtianeti, Zhinvali towards Shatili, at rkm 41 / 65, 42.4031 ° N, 44.9281 ° E, 1090 m a. s. l., 8. VIII. 2014, F. Walther leg.; 1 ♂ (SMNG), Khashuri, Akhaldaba, Nedzvi Managed Reserve, Nedzvistskali River, 41.9082 ° N, 43.5151 ° E, humid coniferous (Picea) forest, VI. 2011, L. Mumladze leg.; 2 ♂, 1 ♀ (SMNG), AR Ajara, Goderdzis Ugheltekhili Pass, W slope, 41.6483 ° N, 42.4936 ° E, 1730 m a. s. l., 3. X. 2012, F. Walther leg.; 1 ♂, 2 ♀♀ (SMNG), Racha-Lechkhumi and Kvemo Svaneti, 3 km S of Mravaldzali, 41.8797 ° N, 42.3511 ° E, 1900 m a. s. l., 9. X. 2011, F. Walther leg.; many ♂♂, ♀♀, juv. (SNMG), Shida Kartli, Gagluantubani, Tana Valley, 24.1 road-km upstream Didi Ateni, 41.8508 ° N, 43.8658 ° E, 1290 m a. s. l., 16. IX. 2011, F. Walther leg.; 3 ♂♂, 8 ♀♀ (ZMUM), near Manglisi, Quercus forest, 12. XI. 1984, E. Kvavadze leg.; many ♂♂, ♀♀, juv. (ZMUM), Racha, Oni District, N of Utsera, deciduous forest near a spring, rock, in litter, 20. IX. 1981, SIG leg.; 1 ♂, 1 ♀ (ZMUM), Tsodoreti near Tbilisi, in litter, 24. XII. 1974, unknown collector; 2 ♂♂, 3 ♀♀ (ZMUM), Mukhura, ca. 14 km E of Tkibuli, 650 m, Castanea, Acer etc. forest, in litter, under logs, 18. X. 1987, SIG leg.; many ♂♂, ♀♀, juv. (ZMUM), Republic of South Ossetia - Alania, 15 km E of Kvaisi, Suram Mountain Ridge, Ertso Pass, ca. 1000 m, deciduous bush along spring, in litter and under stones, 20. X. 1981, SIG leg.; 2 ♂♂, 3 ♀♀, 2 juv. (ZMUM), 40 km W of Mestia, Kherkhvashi E of Nakra, 1250 - 1700 m, Quercus, Fagus, Carpinus, Picea, Abies etc. forest, litter and bark, 21. VIII- 21. IX. 1986, SIG leg.; 1 ♂, 5 ♀♀ (ZMUM), Kutaisi District, 8 km E of Orpiri, environs of Cave Tsutskhvati, deciduous forest, rock, litter, 24. X. 1981, SIG leg.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
1D8AA424F53D5D0D98AA18B0CB907110.taxon	diagnosis	Diagnosis. A species of Omobrachyiulus very similar to O. unugulis sp. nov. by the general shape of the promere and by the opisthomere (Fig. 13) having a slender, rod-like, anterior process directed distad, a broad, strongly flattened, multilobed, apical outgrowth of the basoposterior process, and the similarly finely and asymmetrically bifurcate solenomere. Differs from the latter species by the following gonopodal characters: promere with a more narrowly rounded apex; apical outgrowth of opisthomeral basoposterior process nearly symmetrical, tripartite, vs. the same being asymmetrically bilobed, forming a dentate distal part and a thumb-like lateral part in O. unugulis sp. nov.; anterior process of opisthomere equal to or slightly higher than solenomere, vs. the same being significantly shorter than solenomere in O. unugulis sp. nov.; solenomere abruptly bent caudad rather than nearly straight and directed almost completely distad; and by the male hypoproct being penta- or septidentate rather than tridentate.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
1D8AA424F53D5D0D98AA18B0CB907110.taxon	distribution	General distribution. Caucasian endemic with a broad distribution south of the Greater Caucasus watershed; not reported north of the latter.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
902B73BE746158FCBBF5A319018CC993.taxon	description	Figs 25, 26, 27	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
902B73BE746158FCBBF5A319018CC993.taxon	materials_examined	Material examined (all from Russia). Holotype: ♂ (in two pieces, gonopods dissected, right antenna missing) (ZMUM), Republic of Adygea, Caucasian Nature Reserve, Lagonaki Plateau, Azishskiy Pass, forest, pitfall traps, 14 - 26. VIII. 2013, Yu. Chumachenko leg. Paratypes: 1 ♂, 1 ♀, 1 subad. ♂, 2 juv. ♀♀ (SMNG 34136) (the male in 2 pieces, with dissected gonopods; the female unbroken, the subadult in two pieces, the juveniles unbroken), Republic of Adygea, Mt. Koryto, by a hut, 44 ° 03 ' 53 " N, 40 ° 21 ' 02 " E, 1600 m a. s. l., open Abies forests with dense undergrowth of herbs and shrubs, sifting leaf litter, 22. VIII. 2005, K. Voigtlaender leg.; 1 ♂, 1 ♀ (SMNG 34142) (the male in 2 pieces, with dissected gonopods; the female in two pieces, right vulva dissected), Republic of Adygea, Shestakova Meadow, 44 ° 01 ' 56 " N, 40 ° 23 ' 25 " E, Abies, Acer, ferns, etc., pitfall traps, 24 - 25. VIII. 2005, K. Voigtlaender leg.; 1 ♂, 1 juv. ♀ (SMNG 34154) (the male in head to ring 6 and rest of body, gonopods and hypoproct dissected; the juvenile unbroken), Republic of Adygea, a hut on the west bank of Afonka River, 44.0019 ° N, 40.4066 ° E, at a tree base, hand collecting, 27. VIII. 2005, K. Voigtlaender leg.; 1 ♂ (NMNHS) (in head to ring 6, ring 7, and rest of body in three pieces; gonopods and hypoproct dissected), at Sakhray River, some 2 km E of Novoprokhladnoye, 44 ° 08 ' 11.4 " N, 40 ° 19 ' 04.4 " E, 635 m a. s. l., Fagus forest, in dead wood, hand collecting, 30. VIII. 2005, K. Voigtlaender leg.; 2 ♂♂ (NHMD) (one in 2 pieces, gonopods, dissected, the other in 2 pieces, gonopods missing), 5 ♀♀ (NHMD) (some partly damaged), 3 juv. (NHMD), Krasnodar Province, ca. 9 km SW of Mount Fisht, Babuk-Aul, 43 ° 53 ' 26 " N, 39 ° 49 ' 11 " E, 560 m a. s. l., Castanea and Fagus forest, sifting leaf litter, 11. VII. 2011, A. Solodovnikov leg.; 2 ♂♂ (NHMD) (one unbroken, the other in five pieces, antenna, penis, leg 3, left flange of pleurotergum 7, and hypoproct dissected, opisthomeres and left promere prepared for SEM), 17 ♀♀ (NHMD) (some partly broken, one with dissected vulvae), 28 juv. and subad. (NHMD), Russia, Krasnodar Prov., ca. 7 km SW of Mount Fisht, 43 ° 54.214 ' N, 39 ° 50.507 ' E, 1200 m, Fagus and Rhododendron forest, sifting leaf litter, 14. VII. 2011, A. Solodovnikov leg.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
902B73BE746158FCBBF5A319018CC993.taxon	diagnosis	Diagnosis. A species of Omobrachyiulus most similar to O. implicitus and O. lazanyiae sp. nov. by the overall shape of the promere, the weakly developed basoposterior process and mesomeroidal lobe of the opisthomere, and the broad, collar- or scarf-shaped, apical outgrowth of the basoposterior process which has a characteristic, wrinkled, lamellar part and partially covering the mesal side of the solenomere. Differs from these two species mainly by the apically clavate solenomere, and by the apical outgrowth of the opisthomeral basoposterior process consisting of a relatively narrow cockscomb-like distal part and a strongly developed scarf-shaped mesal part; as well as by other morphological details summarised in Table 1.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
902B73BE746158FCBBF5A319018CC993.taxon	description	Description. Measurements: holotype ♂ in S IX, 45 + 3 + T, L = 15.5 mm, H = 1.05 mm; paratype ♂♂ in S VIII-IX, 44 - 46 + 1 - 3 + T, L = 14 - 16 mm, H = 1 - 1.1 mm; paratype ♀♀ in S VIII-X, 41 - 46 + 2 - 3 + T, L = 11.9 - 19.9 mm, H = 0.85 - 1.3 mm. Colouration (Fig. 25 A): mostly brown, dorsum dark brown, prozonae dorsally with two short, transverse, blackish stripes, one each at frontal margin and near suture; metazonae with one longer, brown, transverse stripe, gradually narrowing and disappearing somewhat below ozopore level; dorsum with a blurred, blackish, axial line, this being more prominent on prozonae; with a dark brown spot above and in front of ozopore. External structures: Eye patches in adults consisting of 30 - 40 ommatidia, mostly arranged in easily countable vertical rows. Vertigial, supralabral, and labral setae: two, four (one ♂ with five), and 16 - 18, respectively. Antennae 1.3 - 1.4 x as long as head in males and 1.2 - 1.3 x in females; antennomere 2> 4 = 5 ≥ 3. Gnathochilarium with a promentum separating both lamellae linguales over ca. 2 / 5 of their length, each latter with three or four setae in a longitudinal row; male stipites parabasally each with a longitudinal row of several short and stiff setae. Collum mostly smooth, with two or three broad shallow grooves near posterolateral corners, and with several densely set, oblique, shallow striae laterally at anterior margin. Body rings barely vaulted. Prozonae with very short, fine, mostly parallel, longitudinal striae near pro-metazonal suture. Metazonae moderately deeply striated, n Schub = 8 or 9; metazonal setae 1 / 3 to nearly half of metazonal length. Ozopores set tightly behind pro-metazonal suture in more anterior rings, gradually moved further backwards, to nearly equal to their diameter behind the suture in caudalmost rings, sutures not being sinuous in front of ozopores. Tarsus of mid-body legs equal to, to slightly longer than, tibia, and nearly 3 x as long as apical claw. Telson (Fig. 25 B): Epiproct long, wedge-like, with a well-developed, slightly clavate, hyaline tip turned dorsad, considerably surpassing longest paraproctal setae. Hypoproct (Fig. 26 A) rounded subtrapezoidal to semi-circular, ventrally with two distal paramedian setae; somewhat protruding past caudal margin of paraprocts in males, tightly fitting to their ventral side in females. Paraprocts sparsely to moderately setose. Male sexual characters: Mandibular stipites weakly to moderately expanded, forming a blunt rectangular or slightly obtuse anterior corner. Leg pair 1 rather slender, parallel to slightly converging hooks. Adhesive leg pads large and crested, gradually reduced in caudal direction, postfemoral ones disappearing in caudal third of body, tibial ones discernible until caudalmost leg pairs; femora without modifications. Pleurotergum 7 ventrally forming rather elongated, distally somewhat clavate and broadly rounded lobes (Fig. 26 B) originating from the zone around pro-metazonal suture. Penis (Fig. 26 C) very short, with short diverging apical lobes and broad terminal lamellae directed caudad. Gonopods (Figs 26 D, 27): Promere (Fig. 27 A, B) rather slender, subequal in height to opisthomere, broadest at base, gradually tapering towards a narrow apex bent mesad; caudal surface with a strongly developed median ridge, and a very deep and relatively narrow median groove; with a small and not too deep distal groove, and a thick, strongly pronounced, distolateral lobe; flagellum (Fig. 27 C and f in Fig. 27 A) slightly longer than height of promere, apically with several whorls of basad pointing denticles. Opisthomere (Figs 26 D, 27 D-G) rather elongated; a basoposterior process present as a long and narrow, weakly developed lobe, distally forming a dentate cockscomb-like apical outgrowth, mesally extending into a strongly developed, soft, wrinkled, lamellar part, tightly enveloping the base of solenomere mesally and partly frontally; an anterior process absent; a mesomeroidal lobe barely discernible as a weakly developed rounded swelling slightly above opisthomeral mid-height, with a slender apicolateral part bent strongly mesad, considerably outreached by solenomere; mesal side with a large conoid lobe (presumably gonocoxal gland), and a not too deep anteromesal sinus frontal to the gland; several very long, erect and spiniform filaments parabasally along flagellum channel; solenomere relatively slender, bent caudomesad; apex somewhat clavate, consisting of two tightly contiguous, irregularly shaped, apical lobes; a minute tapering lobe bent basad and positioned subapically on mesal side. Female sexual characters: Leg pairs 1 and 2 thicker and considerably shorter than following legs. Vulva (Fig. 26 E) symmetrical, strongly compressed on sides; bursa with an obtuse postero-apical margin; operculum as high as bursa; setation sparse, present only in distal parts of bursa and operculum. Receptaculum seminis consisting of a rather narrow and short central tube, and a very narrow and short, mostly straight, posterior tube leading to a minute, ovoid, posterior ampulla.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
902B73BE746158FCBBF5A319018CC993.taxon	distribution	General distribution. NWGC.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
832812E12F0F581280A0D8C5645CE0F2.taxon	description	Figs 32, 33, 34	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
832812E12F0F581280A0D8C5645CE0F2.taxon	materials_examined	Material examined (all from Russia): Holotype: ♂ (unbroken) (ZMUM), Republic of Adygea, Caucasian Biosphere Nature Reserve, Lagonaki Plateau, Azishskiy Pass, subalpine meadow, pitfall traps, 26. VIII- 23. IX. 2013, Yu. Chumachenko leg. Paratypes: 1 ♂ (SMNG) (in 2 pieces, with dissected gonopods), Stavropol, Tamanskaya Dacha Forest, near Komsomolskiy Reservoir, 45.0481 ° N, 41.9567 ° E, 530 m a. s. l., Quercus, Carpinus, Acer, etc., 22. VIII. 2012, F. Walther leg.; 1 ♂ (SMNG 34143) (in 3 pieces, with dissected gonopods), Republic of Adygea, Shestakova Meadow, 44 ° 01 ' 56 " N, 40 ° 23 ' 25 " E, grassy area with sparse Fagus and Betula next to a brook, pitfall traps, 24 - 25. VIII. 2005, K. Voigtlaender leg.; 2 ♂♂ (ZMUM) (one in 2, the other in 3 pieces, with dissected gonopods), 1 ♂ (unbroken) (NHMD), same collecting data as for holotype; 1 ♂ (ZMUM) (in head to collum and 4 pieces, gonopods and penis dissected), 1 juv. (ZMUM), Republic of Adygea, Caucasian Biosphere Nature Reserve, Lagonaki Plateau, Instruktorskaya Cleft, forest, pitfall traps, 23. VIII- 11. IX. 2013, Yu. Chumachenko leg.; 2 ♂♂ (ZMUM) (one in two pieces with dissected gonopods, the other in head, collum to ring 2, ring 3 to ring 6, and rest of body, gonopods prepared for SEM), Stavropol, Mamayskiy Forest, 31. III. 2013, R. Zuev leg.; 1 ♂ (NMNHS) (in two pieces with dissected gonopods), Stavropol, Tamanskaya Dacha Forest, 10. IX. 2013, R. V. Zuev leg.; 1 ♂ (ZMUM) (in 2 pieces with dissected gonopods), 3 adult ♀♀ (ZMUM) (two unbroken, one in 2 pieces with dissected vulvae), 2 subadult ♀♀ (ZMUM), Stavropol, Tamanskaya Dacha Forest, Quercus, Acer, Carpinus, 01. VI. 2014, R. V. Zuev leg.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
832812E12F0F581280A0D8C5645CE0F2.taxon	diagnosis	Diagnosis. A species of Omobrachyiulus differing from congeners mainly by the apically strongly enlarged and hollow solenomere, postero-apically bearing a fine, pointed, sigmoid process, and by the well-developed lateral lobe of the opisthomere ending with a pointed process which is turned anteriad.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
832812E12F0F581280A0D8C5645CE0F2.taxon	description	Description. Measurements: holotype ♂ in S IX, 42 + 1 + T, H = 1.1 mm, L = 18.5 mm; paratype ♂♂ in S VIII-IX, 40 - 43 + 1 + T, L = 15 - 19 mm, H = 1.05 - 1.3 mm; paratype ♀♀ in S VIII-IX, 42 - 45 + 0 - 1 + T, L = 15 - 21 mm; H = 1.5 - 1.8 mm. Colouration (Fig. 32): Head and collum with the usual pattern; prozonae brown-beige, dorsally with a narrow, dark grey, transverse band near pro-metazonal suture, and with a blackish spot in front of ozopores; metazonae brownish, darker in anterior section, light ochre-brown at hind margin; dorsum with a continuous, blackish, axial line; pre-anal ring dark brown, paraprocts lighter brown-beige. External structures: Eye patches in adults consisting of 22 - 35 well-pronounced ommatidia mostly arranged in easily countable vertical rows. Vertigial, supralabral, and labral setae: two, four, and 22 - 28, respectively. Antennae (in Fig. 32 B) 1.6 - 1.7 x as long as head in males and 1.3 - 1.4 x in females; antennomere 2> 5 ≥ 3 ≥ 4> 6. Gnathochilarium with a relatively large promentum separating both lamellae linguales nearly halfway, each latter with two or three setae in a longitudinal row. Collum mostly smooth, with only tree or four striae near posterolateral corners. Body rings gently vaulted. Prozonae with very short, fine, mostly purely longitudinal striae in posterior sections. Metazonae rather deeply striated, n Schub = 6 or 7 in males and 7 or 8 in females; metazonal setae 1 / 2 - 2 / 3 of metazonal length. Ozopores touching or set barely behind pro-metazonal sutures along entire body; sutures not sinuous in front of ozopores. Tarsus of mid-body legs equal to, to slightly longer than, tibia, and ca. 3 x as long as apical claw. Telson (Fig. 32 C): Epiproct mostly straight, distally slightly turned ventrad, wedge- rather than roof-like, ending with an indistinct, blunt, hyaline tip reaching the level of longest anal setae. Hypoproct (Fig. 33 A) in males broad trapezoidal with broadly rounded corners, margin coarse, sometimes forming three small blunt denticles, considerably protruding past rear margin of paraprocts; same being broader, with an always smooth margin, barely protruding behind rear margin of paraprocts. Paraprocts covered with relatively sparse long setae; without distinct rows of shorter setae along caudal margins. Male sexual characters: Mandibular stipites (in Fig. 32 B) considerably expanded, forming a distinct, blunt or narrowly rounded, anteroventral corner. Leg pair 1 compact, parallel hooks, slightly turned mesad. Legs 2 (Fig. 33 B) and following pairs ventrally with well-developed crested adhesive pads, postfemoral ones vanishing in caudal third of body; femora 2 basoventrally with small bumps (black arrow), femora of other legs without modifications. Pleurotergum 7 ventrally forming broad shovel-like lobes (Fig. 33 C) originating mostly from metazona, directed ventromesad, touching one another behind gonopods. Penis (Fig. 33 D) nearly as long as broad, compressed antero-caudally, with short diverging apical lobes ending in rather large terminal lamellae directed distad. Gonopods (Figs 33 E, 34): Promere (Fig. 34 A) as high as opisthomere, broadest at base, abruptly narrowing after an incision at lateral margin's mid-proximal section; lateral margin distally slightly convex, mesal one very gently sigmoid; apex broad, with a nearly rectangular meso-apical corner; caudal surface with a strongly pronounced median ridge, a deep and very narrow median groove, and a deep, rounded, distal groove; flagellum somewhat longer than height of promere, apically bearing blunt denticles directed basad, arranged in two or three transverse rows. Opisthomere (Figs 33 E, 34 B-F) moderately elongated; basoposterior process moderately developed, its apical outgrowth large, forming two parts: a smaller median one, and a larger mesal one bent strongly basofrontad, covering the basomesal side of solenomere like a hood; both parts distally with several small conical denticles; an anterior process absent; mesomeroidal lobe well-developed, narrow, ridge-like in its proximal part, distally widening in a pyramidal structure with a distinct apex; lateral side with a well-developed ridge-like lobe apically forming a pointed horn-like process bent frontad; mesal side with a relatively large, subtriangular lobe (presumably gonocoxal gland), and a shallow, indistinct, anteromesal sinus; moderately long, erect, spiniform filaments present over most of flagellum channel; solenomere unipartite, apically clavate and deeply hollow, with a fine sigmoid apical process. Female sexual characters: Vulva (Fig. 33 F) rather short and compact, symmetrical; operculum slightly higher than bursa, both bursa and operculum ending up with broad and rather short hyaline protrusions; setation dense throughout. Receptaculum seminis consisting of a very large, sack-shaped, central tube, and a very long, narrow, somewhat folded, posterior tube leading to a nearly spherical posterior ampulla.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
832812E12F0F581280A0D8C5645CE0F2.taxon	distribution	General distribution. NWGC-WCIS.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
CD0F6BD106F75DF2B6E8631130287C4A.taxon	materials_examined	Material examined. Georgia: 1 ♂ (SMNG), Mtskheta-Mtianeti, Stepantsminda: mouth of river Snotskali, 42.6339 ° N, 44.6258 ° E, 1750 m a. s. l., 07. X. 2012, F. Walther leg., 1 ♂ (SMNG), Mtskheta-Mtianeti, Darial Gorge at rkm 8 / 131, 42.7039 ° N, 44.6264 ° E, 1480 m a. s. l., 10. VIII. 2014, F. Walther leg.; Russia: 3 ♂♂, 1 ♀, 6 juv. (ZMUM), Karachay-Cherkess Republic, Teberda Nature Reserve, Kizgich Canyon N of Arkhyz, near a riverine, Alnus and Betula wet forests, under bark and stones, 1450 - 1500 a. s. l., 5. VI. 1985, SIG leg.; 1 ♂ (ZMUM), Chechen Republic, Assa Valley, ca. 9 km SSW of Muzhichi, 800 m, Fagus, Alnus, Carpinus etc. forest, in litter, under bark and stones, 15. VII. 1986, SIG leg.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
CD0F6BD106F75DF2B6E8631130287C4A.taxon	diagnosis	Diagnosis. A species of Omobrachyiulus differing from congeners by the following combination of opisthomere characters: basoposterior process in the shape of a strongly pronounced lobe forming a nearly right-angled corner at base, ending with a simple and smooth apical outgrowth with a blunt tip; anterior process shaped as a stout spine; mesomeroidal lobe strongly pronounced; solenomere simple, bent strongly caudad.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
CD0F6BD106F75DF2B6E8631130287C4A.taxon	distribution	General distribution. GRCA.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
20045851659B544AADE8C329313F625C.taxon	description	Fig. 17 B, C	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
20045851659B544AADE8C329313F625C.taxon	materials_examined	Material examined. Georgia: 3 ♂♂, 2 ♀♀ (ZMUM), Mestia, 1500 m a. s. l., Betula, Rhododendron on moraine, litter and under stones, 5 and 15. IX. 1986, SIG leg.; 1 ♂ (SMNG), Mestia, below Chalaadi Glacier, 43.1118 ° N, 42.7453 ° E, Abies nordmanniana and various deciduous trees, in leaf litter, 9. VI. 2019, H. Reip leg.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
CA380A8E3E42597DBF8E4971D5A39B52.taxon	description	Fig. 17 A	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
CA380A8E3E42597DBF8E4971D5A39B52.taxon	materials_examined	Material examined. Georgia: 2 ♂♂, 2 ♀♀ (ZMUM), NE of Poti, Chaladidi, Alnus, Quercus, Fraxinus forest on swamp, in litter, 13. IV. 1983, SIG leg.; 1 ♂, 1 ♀ (ZMUM), AR Abkhazia, Sukhum District, Nizhnyaya Yashtukha, forest nursery, 29. III. 1985, A. Markossian; 1 ♂, 2 juv. (ZMUM), same place, tobacco plantation, 16. VI. 1980 and 25. VII. 1980, A. Markossian.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
CA380A8E3E42597DBF8E4971D5A39B52.taxon	diagnosis	Diagnosis. A species of Omobrachyiulus most similar to O. armatus sp. nov. by the promere significantly outreaching the opisthomere, the latter possessing a massive lobe-like basoposterior process forming two distinct corners, a basal and a distal one, and having a micro-spiculate mesal side, and a unipartite solenomere with a slender rod-like ending. Differs from O. armatus sp. nov. mainly by the clearly tripartite apical outgrowth of the basoposterior process, consisting of a mesal lamellar, a median fan-shaped, and a lateral spiniform part, vs. that same outgrowth being broad, unipartite, collar-shaped and dentate at the margins in the latter species.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
CA380A8E3E42597DBF8E4971D5A39B52.taxon	distribution	General distribution. COLC-SWGC.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
1E3B6D74F39B5D9DA78583358A61C42A.taxon	description	Fig. 24	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
1E3B6D74F39B5D9DA78583358A61C42A.taxon	materials_examined	Material examined. Georgia: AR Abkhazia: 6 ♂♂, 7 ♀♀, 1 juv. (ZMUM), Pitsunda-Myussera Nature Reserve, Myussera part, 120 - 130 m a. s. l., mixed deciduous forest (Castanea, Alnus etc.), in litter, under bark and stones, 8 - 10. IV. 1983, SIG leg.; 1 ♂, 3 ♀♀, 1 juv. (ZMUM), Avadhara, 1600 - 1700 m a. s. l., Abies, Fagus, fern, Rubus, Galium, 18. IX. 1985, I. A. Ushakov leg.; 2 ♂♂, 3 ♀♀, 1 juv. (ZMUM), N of Lake Ritsa, Abies, Fagus, in litter, 13. IX. 1985, I. A. Ushakov leg. Russia: Republic of Adygea: 1 ♂, 2 ♀♀ (AE), near Maykop, Polkovnitskaya Ravine, 44 ° 20.72 ' N, 40 ° 11.37 ' E, pitfall traps, 01 - 17. X. 2011, Yu. Chumachenko leg.; 1 ♂, 1 juv. ♂ (AE), same place, collecting method and collector, 17. X- 01. XI. 2011; 1 ♂, 1 juv. (ZMUM), confluence of Kisha and Belaya rivers, mixed forest, ca. 200 m a. s. l., 11. VI. 2013, R. V. Zuev leg.; 5 ♂♂, 6 ♀♀, 5 juv. (ZMUM), Krasnodar Province, 5.5 km NE of Krasnaya Polyana, lower course of Achipse River, 43 ° 15 ' 25 " N, 40 ° 15 ' 25 " E, ca. 600 m a. s. l., 19 - 23. VIII. 2014, K. Makarov and A. Matalin leg.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
1E3B6D74F39B5D9DA78583358A61C42A.taxon	diagnosis	Diagnosis. A species of Omobrachyiulus most similar to O. fasciatus sp. nov. and O. lazanyiae sp. nov. by the overall shape of the promere, the weakly developed basoposterior process and mesomeroidal lobe of the opisthomere, and the broad, collar- or scarf-shaped, apical outgrowth of the basoposterior process, the latter having a characteristic, wrinkled, lamellar part partially covering the mesal side of the solenomere. Differs from these two species mainly by the very slender and tripartite solenomere apically forked into two mostly symmetrical, strongly diverging branches, with a minute ear-like lobe mesally at the base of the bifurcation; and by the distally broader promere with a very broad and deep distal groove on the caudal surface, this being a small pit in O. fasciatus sp. nov. and O. lazanyiae sp. nov.; as well as in other gonopod details summarised in a tabular key (Table 1) to these three species.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
1E3B6D74F39B5D9DA78583358A61C42A.taxon	distribution	General distribution. WECA.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
7F9BC6CC01665AA29E71859147F6DC59.taxon	description	Figs 38, 39, 40	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
7F9BC6CC01665AA29E71859147F6DC59.taxon	materials_examined	Material examined (all from Georgia). Holotype: ♂ (unbroken) (ZMUM), AR Ajara, Kintrishi Nature Reserve, mouth of Khekpara River, 2. X. 1984, E. Kvavadze leg. Paratypes: 8 ♂♂ (ZMUM) (five unbroken, one in head to ring 3, ring 4 to pleurotergum 7, and rest of body, with dissected penis and gonopods, one in head to pleurotergum 7 and rest of body, gonopods dissected, one broken in 2 pieces, one in 3 pieces), 1 ♂ (NHMD) (unbroken), 1 ♂ (NMNHS) (unbroken), 1 ♂ (IBER) (unbroken), 8 ♀♀ (ZMUM) (six unbroken, two in 2 pieces, one in head to ring 3 and rest of body, one in 2 pieces, head missing), 1 ♀ (NHMD) (unbroken), 1 ♀ (NMNHS) (unbroken), 1 ♀ (IBER) (unbroken), 2 juv. (ZMUM), same collecting data as for holotype; 1 ♂ (in head to ring 6 and rest of body), 4 ♀♀ (one broken in two pieces, the rest unbroken) (ZMUM), AR Ajara, Kintrishi Reserve, Zeraboseli, 450 - 600 m a. s. l., deciduous forest, litter and under stones, 13. X. 1981, SIG leg.; 4 ♂♂ (one in head to ring 6 + rest of body, gonopods dissected; one broken in two pieces, the rest unbroken), 2 ♀♀ (one broken in two pieces, the other unbroken), 2 juv. (ZMUM), same area, 800 m a. s. l., same date and collector; 1 ♂ (in leg to ring 6 and rest of body, gonopods dissected), 2 ♀♀ (one in 2 pieces, the other unbroken), 1 juv. (ZMUM), AR Ajara, Batumi Botanical Garden, 13. X. 1978, SIG leg.; 1 ♂ (ZMUM) (in head to ring 6, pleurotergum 7 to ring 9, and rest of body, opisthomeres and left promere prepared for SEM), AR Ajara, E of Kobuleti, 3 km SE of Chakhati, deciduous forest, near a spring, litter and under stones, 14. X. 1981, SIG leg.; 1 ♂ (ZMUM) (in head to pleurotergum 7 and rest of body, gonopods dissected), Samtskhe-Javakheti, 15 km W of Adigeni, Abieas, Picea, Fagus, Acer etc. forest, 1500 - 1700 m, litter, logs, under stones, 14 - 15. V. 1983, SIG leg.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
7F9BC6CC01665AA29E71859147F6DC59.taxon	diagnosis	Diagnosis. A species of Omobrachyiulus most similar to O. sevangensis comb. nov., O. ponticus sp. nov., and O. trochiloides sp. nov. in gonopod structure, as well as in the shape of the male telson. Differs from all these species mainly by the opisthomere having a bifid anterior process and a better developed, higher mesomeroidal lobe ending with a distinct apicolateral part.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
7F9BC6CC01665AA29E71859147F6DC59.taxon	description	Description. Measurements: holotype ♂ in S IX, 40 + 1 + T, L = 15.5 mm, H = 1.05 mm; paratypes in S VIII-IX, 39 - 42 + 1 - 2 + T, L = 13 - 16.5 mm and 19 - 24 mm, H = 1 - 1.1 mm and 1.4 - 1.6 mm, in males and females, respectively. Colouration (after> 30 years in alcohol) (Fig. 38): Head light brown with the usual dark band between eye patches; collum light brown, margins darker; Prozonae dark brown, becoming greyish on more posterior rings, with numerous lighter spots, dorsally with a blackish transverse stripe; metazonae anteriorly dark brown, lighter at hind margins; dorsum without axial line; whole body lighter on ventral side; pre-anal ring dark brown, paraprocts lighter. External structures: Eye patches in adults consisting of 30 - 35 ommatidia arranged in easily countable vertical rows. Vertigial, supralabral, and labral setae: two, four, and 15, respectively. Antennae (in Fig. 38 A) ca. 1.5 x as long as head in males and 1.3 x in females; antennomere 5> 2> 4> 3> 6. Gnathochilarium with promentum almost separating lamellae linguales halfway, each latter with three or four setae in a longitudinal row. Collum smooth, with only several small, shallow, oval grooves near posterolateral corners. Body rings very gently vaulted. Prozonae with scattered, short and shallow, mostly parallel longitudinal striae. Metazonae relatively deeply striated, n Schub = 6 or 7 (males) and 7 or 8 (females); metazonal setae ca. 3 / 4 metazonal length in most rings, this ratio becoming 1: 1 in caudalmost rings. Ozopores placed right in pro-metazonal suture in first several rings, gradually taking a more posterior position to ~ 1 / 2 of their diameter behind suture in caudalmost rings; sutures gently sinuous in front of ozopores in only some rings. Tarsus of mid-body legs 1.3 - 1.5 x as long as tibia and 3.5 - 5 x as long as apical claw. Telson (Fig. 38 C): Epiproct very long, ending in a pointed hyaline tip turned slightly to considerably ventrad, surpassing the longest paraproctal setae in males, just reaching their level in females. Hypoproct (Fig. 39 A) relatively small, tridentate in males, with the teeth protruding behind rear contour of paraprocts; rounded, completely concealed under paraprocts in females; ventral surface with six submarginal setae. Paraprocts sparsely to moderately densely setose, without distinct rows of shorter setae along caudal margins. Male sexual characters: Mandibular stipites (in Fig. 38 B) considerably expanded, protruding mostly anteriad, forming a broadly rounded anterior corner. Leg pair 1 compact rounded hooks, somewhat turned against one another. Leg pair 2 and several following pairs (Fig. 39 B) with crested adhesive pads, and with a small weakly pronounced bump proximally on femur (black arrow); tibial pads reduced posteriad, but still present until last leg pair, postfemoral ones completely disappearing in last several pairs. Pleurotergum 7 ventrally forming shovel-like lobes (Fig. 39 C) originating from the zone around pro-metazonal suture, protruding mostly ventrad, completely concealing opisthomeres from lateral view. Penis (Fig. 39 D) antero-caudally strongly compressed, with diverging apical lobes; terminal lamellae not differentiated. Gonopods (Figs 39 G-F, 40 A-E): In situ mostly concealed in gonopodal sinus, only apical part of promeres visible, approximately reaching the level of pleurotergal protrusions of pleurotergum 7, opisthomeres fully concealed between promeres and pleurotergal protrusions. Promere (Fig. 40 A, p in Fig. 39 E) slightly higher than opisthomere, mesal margin mostly straight, lateral margin parallel to the former in proximal section, distally slanting towards a narrowly rounded apex; caudal surface with a short, strongly pronounced, median ridge, a very short and narrow median groove, and a rounded, micro-squamous, distomesal lobe directed caudad; flagellum slightly longer than height of promere. Opisthomere (Figs 39 G-F, 40 B-E) relatively slender; basoposterior process weakly pronounced, ending in a tapering apical outgrowth with its tip turned anteriad; anterior process shaped as an elongated lobe running parallel to CBO, distally forked into two fine, pointed tips: a shorter, proximal one, and a longer, distal one, the former with a minute lamella at base; mesomeroidal lobe rather weakly pronounced, ending in an apicolateral part with slightly jagged edge bent anteriad; mesal side with a broad and deep anteromesal sinus, and (apparently) without a lobe at base; solenomere unipartite, straight, apically hollow; a row of rather small spiniform filaments along proximal section of flagellum channel. Female sexual characters: Leg pairs 1 and 2 somewhat thicker than, and ca. as long as, following legs. Vulva (Fig. 39 H) relatively elongated; bursa symmetrical, with a strongly obtuse postero-apical margin; side sclerites apically ending up in small hyaline protrusions; operculum significantly higher than bursa, distally thickened and bent somewhat anteriad, with a rounded apical margin, caudal surface laterally ending up with two pointed hyaline protrusions; whole bursa sparsely setose, operculum with only several setae in distal part. Receptaculum seminis consisting of a short finger-shaped central tube, and a mostly straight posterior tube of same gauge, ending in a medium-size ovoid posterior ampulla.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
7F9BC6CC01665AA29E71859147F6DC59.taxon	distribution	General distribution. LECA.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
8D3A35470ECB560CAEFE46BE165E424C.taxon	description	Figs 28, 29, 30	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
8D3A35470ECB560CAEFE46BE165E424C.taxon	materials_examined	Material examined (SMNG). Holotype: ♂ (in two pieces, gonopods, leg 6 and a mid-body leg dissected), Georgia, AR Abkhazia, Gulripsh Distr., 6 km E of Tsebelda, right bank of Jampali River, Cave Nizhnyaya Shakuranskaya, 8 - 9. XI. 1987, N. Myuge leg. Paratype: 1 ♂ (in head, collum, rings 2 - 3, ring 4, rings 7 - 8, and rest of body; leg pair 1, leg 7, a mid-body leg, right flange of ring 7, and hypoproct dissected; opisthomeres and one promere prepared for SEM), same collecting data as for holotype.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
8D3A35470ECB560CAEFE46BE165E424C.taxon	diagnosis	Diagnosis. A species of Omobrachyiulus most similar to O. implicitus and O. fasciatus sp. nov. by the overall shape of the promere, the weakly developed basoposterior process and mesomeroidal lobe of the opisthomere, and the broad, collar- or scarf-shaped, apical outgrowth of the basoposterior process, having a characteristic, wrinkled, lamellar part, partially covering the mesal side of the solenomere. Differs from those two species mainly by the solenomere which is apically slender (not clavate as in O. fasciatus sp. nov.), divided in two parallel, tightly contiguous (not strongly diverging as in O. implicitus) branches, subapically on mesal side bearing a minute lobe with several denticles, protruding perpendicular to the main axis of the solenomere.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
8D3A35470ECB560CAEFE46BE165E424C.taxon	description	Description. Measurements: holotype ♂ in S IX, 41 + 2 + T, L = 12 mm, H = 0.95 mm; paratype ♂ in S IX, 42 + 1 + T, L = 12.5 mm, H = 0.9 mm. Colouration (strongly faded from the ethanol conservation) (Fig. 28 A): mostly brownish beige; head and collum with the usual pattern; prozonae dorsally with two dark transverse stripes, one in frontal section and the other next to pro-metazonal suture, the former broader; a dark washed spot in front of ozopores; metazonae uniformly light brown-beige; dorsum with a blackish axial line. External structures: Eye patches in adults consisting of 20 - 30 ommatidia arranged in unclear vertical rows. Vertigial, supralabral, and labral setae: two, four (one ♂ with five), and 16, respectively. Antennae (in Fig. 28 A) 1.1 - 1.2 x as long as head in males; antennomere 2> 3 ≥ 4 = 5. Gnathochilarium with a very small promentum separating both lamellae linguales over ca. 1 / 4 of their length, each latter with four setae in a longitudinal row. Collum completely smooth, with just one or two faint longitudinal grooves near posterolateral corners. Body rings not vaulted. Prozonae with very short, shallow, scattered, longitudinal striae in posterior parts. Metazonae moderately deeply striated, n Schub = 7 or 8; metazonal setae from 1 / 2 (in frontal and mid-body rings) to 2 / 3 (in caudalmost rings) of metazonal length. Ozopores set tightly behind pro-metazonal suture in more anterior rings, moved slightly backwards, to ca. half their diameter behind the suture in more posterior rings, sutures being gently sinuous in front of ozopores. Tarsus of mid-body legs ca. 1.3 x as long as tibia and ca. 3 x as long as apical claw. Telson (Fig. 28 B): Epiproct rather long (approximately reaching the level of longest paraproctal setae), straight, ending bluntly without distinct hyaline tip; bearing several long setae. Hypoproct (Fig. 29 A) rounded subtrapezoidal, ventrally with two distal paramedian setae; slightly protruding past caudal margin of paraprocts. Paraprocts sparsely setose. Male sexual characters: Mandibular stipites (in Fig. 28 A) rather weakly expanded, forming a narrowly rounded anterior corner. Leg pair 1 (Fig. 29 B) markedly slender, slightly converging hooks with minute pointed tarsal remnants in the paratype (black arrow), the latter being absent in the holotype. Leg pair 2 somewhat shorter than following pairs. Adhesive pads as in O. fasciatus sp. nov. Ventral protrusions of pleurotergum 7 (Fig. 29 C) similar to those in O. fasciatus sp. nov., but insignificantly widening distally. Gonopods (Figs 29 D, 30): Promere (Fig. 30 A) rather stout, subequal to opisthomere, broadest at base, insignificantly tapering towards a broadly rounded apex; caudal surface with a pronounced median ridge, a deep and broad median groove, a small, but deep distal groove near lateral margin, and a rather weakly pronounced distolateral lobe. Opisthomere (Figs 29 D, 30 B-E) relatively slender; basoposterior process similar to that in O. fasciatus sp. nov., but with a broader apical outgrowth surrounding the proximal part of solenomere from all sides except frontally, and having a less strongly developed lamellar part; a (presumably) anterior process present as a small slender leaf running tightly contiguous with the basal part of solenomere; mesomeroidal lobe moderately pronounced, without an apicolateral part; mesal side mostly as in O. fasciatus sp. nov., but with somewhat shorter spiniform filaments at flagellum channel; basal section of seminal channel densely micro-spiculate; solenomere slender, gently sigmoid, directed distad; apex divided into two parallel, tightly contiguous branches, the mesal one bearing a minute lobe protruding meso-caudad, with several denticles at margin. Female sexual characters: unknown.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
8D3A35470ECB560CAEFE46BE165E424C.taxon	distribution	General distribution. SWGC.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
3FE2E90AD4DF5C24A222AD83804F2FFB.taxon	description	Updated general description. Small to medium-sized Brachyiulini (L (males) = 10 - 33 mm). Ommatidia present. Ozopores set tightly behind pro-metazonal suture at least on more anterior body rings. Epiproct present, varying in length. Male hypoproct either broad, dentate at margin and trapezoidal or semi-circular / semi-elliptic. Male mandibular stipites moderately to considerably expanded, forming a distinct anterior / anteroventral corner. Male pleurotergum 7 significantly bulging only in some species. Male walking legs ventrally with two well-developed adhesive pads, one each on postfemur and tibia, postfemoral pads rarely lacking in leg pair 2. Penis short and stout, usually considerably compressed anterocaudally, with short or indistinct apical lobes. Gonopods: In situ: promeres slightly to considerably protruding outside gonopodal sinus, directed completely ventrad. Promere subequal to or higher than opisthomere, mostly roughly leaf-shaped, caudal side sometimes forming small lobes, but bearing no distinct processes; flagellum usually much longer than height of promere. Opisthomere from stout to relatively slender; basoposterior process usually represented by a differently pronounced vertical lobe, apically forming a variously shaped and freely protruding outgrowth; an anterior process usually present, varying in shape and size; a mesomeroidal lobe always present, mostly rounded, varying in size and prominence; solenomere always conspicuously protruding from CBO, usually slender and tubular. Vulva: Subcylindrical. Bursa usually with a distinct, more or less obtuse, postero-apical margin. Opening narrow oval or cleft-like, positioned apically. Operculum usually somewhat longer than or subequal to, rarely shorter than, bursa. Receptaculum seminis: central tube narrow, mostly straight, forming no distinct central ampulla; posterior tube narrow, usually more or less twisted, ending in an ovoid or spherical posterior ampulla.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
0D3006B627FE5A938E2FBC2376206771.taxon	materials_examined	Material examined. Georgia: 2 ♂♂ (SMNG), Akhaltsikhe to Borjomi, S of Chobiskhevi, at Chobiskhevi River, 42.7651 ° N, 43.3265 ° E, mountain Fagus forest, VII. 2011, L. Mumladze leg.; 1 ♂ (SMNG), Samegrelo-Zemo Svaneti, Nikortsminda, Skhartali, near the entrance to Sakishore Cave, 42.4420 ° N, 43.1590 ° E, Picea and Abies forest, under dead wood, 15. VI. 2019, H. Reip leg.; 1 ♂, 12 ♀♀ (ZMUM), Kherkhvashi E of Nakra, 1250 - 1700 m, Quercus, Fagus, Carpinus, Abies etc. forest, in litter and under bark, 21. VIII- 21. IX. 1986, SIG leg.; 2 ♂♂, 1 ♀ (ZMUM), Chokhatauri District near Bakhmaro, 4 km SSE of Nabeglavi, 600 m, Alnus along stream, 8. VI. 1981, SIG and J. Martens leg.; 1 ♂, 4 ♀♀, 3 juv. (ZMUM), Tbilisi, Tskhneti, 1150 - 1300 m, Fagus, Carpinus, Acer etc. forest, 16 - 18. V. 1987, SIG and K. Eskov leg.; 1 ♂, 2 ♀♀ (ZMUM), Mukhura, ca. 15 km E of Tkibuli, 700 - 800 m, Castanea, Fagus, Carpinus etc. forest, in litter, under bark and stones, 7 - 9. V. 1987, SIG and K. Eskov leg.; 2 ♂♂, 3 ♀♀ (ZMUM), environs of Manglisi, Quercus forest, 12. XI. 1984, E. Kvavadze leg.; 3 ♂♂, 9 ♀♀, 4 juv. (ZMUM), Algeti National Park, W of Manglisi, Fagus, Picea, Acer etc., 1400 - 1450 m, litter and under bark, 16 - 18. V. 1987, SIG and K. Eskov leg. Russia: 5 ♂♂, 10 ♀♀ (ZMUM), Chechen Republic, Argun Valley, 5 km N of Shatoy, Corylus, Fagus, Carpinus etc. forest, 750 m, in litter, under stones and bark, 18. VII. 1986, SIG leg.; 1 ♂ (NHMD), Karachay-Cherkess Republic, ca. 15 km SW of Teberda, 1500 m a. s. l., Fagus-Picea forest, pitfall traps, 22 - 28. VIII. 2011, A. Solodovnikov and S. Tarasov leg.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
0D3006B627FE5A938E2FBC2376206771.taxon	diagnosis	Diagnosis. A species of Omobrachyiulus closely resembling O. divaricatus and O. unugulis sp. nov. by the general shape of both pro- and opisthomere. Differs from these two species mostly by the bulging pillow-shaped, rather than antero-caudally strongly flattened, apical outgrowth of the basoposterior process, as well as by the very large (both long and broad) epiproct.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
0D3006B627FE5A938E2FBC2376206771.taxon	distribution	General distribution. CAUC.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
FA33AAF820C65286AECF2C84C63A72A7.taxon	description	Figs 41, 42, 43	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
FA33AAF820C65286AECF2C84C63A72A7.taxon	materials_examined	Material examined (ZMUM). Holotype: ♂ (in head, collum to ring 3, ring 4 to ring 6, and rest of body; leg pair 1, penis, right leg 3, gonopods, left flange of pleurotergum 7, and a mid-body leg dissected) (ZMUM), Georgia, AR Abkhazia, Pitsunda-Myussera Nature Reserve, Myussera part, 20 - 130 m, mixed deciduous forest (Castanea, Alnus, etc.), litter, under bark and stones, 8 - 10. IV. 1983, SIG leg. Paratypes: 2 ♀♀ (one unbroken, the other in head to ring 3 and rest of body, right vulva dissected), same collecting data as of the holotype.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
FA33AAF820C65286AECF2C84C63A72A7.taxon	diagnosis	Diagnosis. A species of Omobrachyiulus most similar to O. sevangensis (Lohmander, 1936) comb. nov., O. kvavadzei sp. nov., and O. trochiloides sp. nov.; sharing with O. Omobrachyiulus sevangensis and O. trochiloides sp. nov. the presence of a lamellar ridge-like anterior process of the opisthomere. Differs from the former species by the presence of a distinct, rounded, apicolateral part of the mesomeroidal lobe of the opisthomere, as well as by the solenomere considerably outreaching both anterior process and apical outgrowth of the basoposterior process, and from the latter species by the much more obscure, poorly differentiated basoposterior process ending in an unciform, rather than a fine rod-like, apical outgrowth. Resembling O. kvavadzei sp. nov. mostly by the promere having a distomesal lobe and by the mesomeroidal lobe of the opisthomere bearing an apicolateral part, but differing in other gonopod characters included in the tabular key to the Omobrachyiulus sevangensis group (Table 2).	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
FA33AAF820C65286AECF2C84C63A72A7.taxon	description	Description. Measurements: holotype ♂ in S VII, 35 + 1 + T, L = 10 mm, H = 0.9 mm; paratype ♀♀ in S VII-VIII, 34 - 35 + 2 + T, L = 11.5 - 12 mm, H = 1.2 - 1.35 mm. Colouration (Fig. 41 A): Head and collum with the usual colour pattern; body gradually lighter from dorsal to ventral side; prozonae grey, dorsally with a transverse dark brown stripe near pro-metazonal suture; metazonae light brown-beige; dorsum with a continuous whitish axial line. External structures: Eye patches in adults consisting of 20 - 23 relatively large, clearly convex ommatidia arranged in mostly easily recognisable vertical rows. Vertigial, supralabral, and labral setae: two, four, and 17, respectively. Antennae ca. 1.4 x as long as head in males and 1.1 - 1.15 x in females; antennomere 2> 5> 4 ≥ 3> 6. Gnathochilarium with a conspicuously large promentum separating lamellae linguales> halfway, the latter with three or four setae in a longitudinal row; stipites in males parabasally each with a longitudinal row of several setae, non-setose in females. Collum smooth, with a single deep groove running along entire lateral margin. Body rings somewhat vaulted. Prozonae with scattered, very short and shallow, longitudinal grooves. Metazonae rather deeply striated, n Schub = 6 or 7; metazonal setae relatively short, ca. 2 / 5 of (in more anterior rings), to equal to (in more posterior rings), metazonal length, arranged in very dense whorls. Ozopores placed right in pro-metazonal suture in more anterior rings, and tightly behind suture in more posterior ones; sutures not being sinuous in front of ozopores. Tarsus of mid-body legs 1.3 x as long as tibia and 3.8 x as long as apical claw. Telson (Fig. 41 B): Epiproct very long (considerably surpassing the longest paraproctal setae in males, somewhat less pronounced in females), straight, distally bent ventrad, ending with a pointed hyaline tip; with a few setae. Hypoproct in males narrowly trapezoidal, tridentate, the teeth blunt, protruding behind rear contour of paraprocts; the same being narrowly rounded in females. Paraprocts rather sparsely setose, without distinct row of shorter setae along caudal margins. Male sexual characters: Mandibular stipites rather weakly expanded, forming a rounded ventro-anterior corner. Leg pair 1 (Fig. 42 A) considerably converging (turned against one another) hooks with slender tibial outgrowths. Leg pair 2 and following pairs with crested adhesive pads, both tibial and postfemoral ones gradually reduced towards telson, but still visible after mid-body; femora of pairs 3 - 7 with an oval groove in the centre of a bulge. Pleurotergum 7 ventrally forming broadly rounded lobes (Fig. 42 B) originating from the zone around pro-metazonal suture, protruding mesoventrad behind gonopods. Penis (42 C) very small, subquadrate, without apical lobes and with very short, rounded, terminal lamellae directed distolaterad. Gonopods (Fig. 43): Promere (Fig. 43 B, p in Fig. 43 A) slightly exceeded in height by solenomere, slender, with nearly symmetrical mesal and lateral margins joining in a narrow, blunt apex; median ridge short and strongly pronounced; a well-developed distomesal lobe. Opisthomere slender; basoposterior process weakly pronounced, ending in a tapering apical outgrowth turned slightly anteriad; anterior process shaped as a micro-serrate lamellar ridge with a freely protruding, tapering, apical part; mesomeroidal lobe with a moderately pronounced basal part, flattened distally and ending in an apicolateral part being very similar in shape to that in O. kvavadzei sp. nov.; solenomere deeply divided into two fine, slender, parallel branches: an anterior and a posterior one, the latter being slightly longer than the former; spiniform filaments along flagellum channel either absent or too small and slanting to be detected by light microscopy. Female sexual characters: Leg pairs 1 and 2 somewhat shorter, first also thicker, than following legs. Vulva (Fig. 42 D, E) rather slender, similarly shaped to that of O. kvavadzei sp. nov., the main difference being the less strongly slanting apex of bursa, and the smaller, apically positioned opening in O. ponticus sp. nov.; both bursa and operculum moderately densely covered with long setae. Receptaculum seminis consisting of a short finger-shaped central tube, and a long, spirally twisted, posterior tube ending in an ovoid posterior ampulla with a minute pocket at bottom.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
FA33AAF820C65286AECF2C84C63A72A7.taxon	distribution	General distribution. SWGC.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
4CCB2E36BE8456A89A2F7D285177C9BD.taxon	description	Figs 21, 22, 23	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
4CCB2E36BE8456A89A2F7D285177C9BD.taxon	materials_examined	Material examined (all from Georgia). Holotype: ♂ (unbroken) (ZMUM), AR Ajara, 15 km W of Adigeni, Abies, Picea, Fagus, Acer, etc., 1500 - 1700 m, litter, logs, under stones, 14 - 15. V. 1983, SIG leg. Paratypes: 1 ♂ (ZMUM) (in head and 4 pieces, gonopods, penis, legs 2, 3, 7 and a mid-body leg dissected), 1 ♂ (NHMD) (unbroken), 1 ♂ (NMNHS) (unbroken), 2 ♀♀ (ZMUM) (one unbroken, the other in 3 pieces with dissected left vulva), 1 ♀ (NHMD) (unbroken), 1 ♀ (NMNHS) (unbroken), 1 subad. ♀ (ZMUM) (unbroken), 5 juv. (ZMUM) (two unbroken, the others fragmented), same collecting data as for holotype; 1 ♂ (ZMUM) (in 3 pieces, gonopods prepared for SEM), AR Ajara, Khulo Distr., 1800 m a. s. l., Danisparauli, Abies and fern litter, 10. X. 1981, SIG leg.; 1 ♂ (ZMUM) (in 5 pieces, with dissected gonopods), 2 ♀♀ (ZMUM) (one in 2 pieces, the other unbroken), AR Ajara, Khulo Distr., Goderdzi Pass, 2000 m, sparse Abies and Picea forest, litter, under stones and in rotten logs, 10. X. 1981, SIG leg.; 3 ♂♂ (SMNG) (two in 2 pieces with dissected gonopods, one unbroken), 1 ♂ (IBER) (in head to ring 6, pleurotergum 7 to ring 9, and rest of body, gonopods dissected), 3 ♀♀ (SMNG) (all unbroken), 1 ♀ (IBER) (unbroken), 1 juv., one head to ring 6 ♂ fragment (SMNG), Sairme Gorge, 41.9580 ° N, 42.7713 ° E, 600 m a. s. l., Fagus orientalis and Castanea sativa, in litter, 14. VII. 2013, L. Mumladze leg.; 3 ♂♂ (SMNG) (one unbroken, one in 2 pieces with intact gonopods, one in 3 pieces with dissected gonopods), Sairme Gorge, 41.8468 ° N, 42.8075 ° E, 2200 m a. s. l., Acer sp. and Betula sp., 11. VII. 2013, L. Mumladze leg.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
4CCB2E36BE8456A89A2F7D285177C9BD.taxon	diagnosis	Diagnosis. A species of Omobrachyiulus most clearly differing from congeners by the caudally flattened, serrate, elongated, apical outgrowth of the basoposterior process of opisthomere, and by the conspicuously large anterior process which is almost as high as the solenomere.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
4CCB2E36BE8456A89A2F7D285177C9BD.taxon	description	Description. Measurements: holotype ♂ in S IX, 41 + 1 + T, L = 15 mm, H = 1.3 mm; paratype ♂♂ in S IX-X, 41 - 44 + 1 + T, L = 15 - 17 mm, H = 1.25 - 1.4 mm; paratype ♀♀ in S IX-X, 38 - 42 + 1 - 2 + T, L = 15 - 19 mm, H = 1.4 - 1.65 mm. Colouration (considerably faded from ethanol) (Fig. 21): Prozonae dorsolaterally dark grey to blackish, ventrolaterally dark grey near pro-metazonal suture; metazonae dorsolaterally with a transverse dark brown stripe; ozopores surrounded by an irregular blackish spot; ventrolateral side of body mostly light brown-beige; dorsum with a black axial line. External structures: Eye patches in adults consisting of 30 - 35 ommatidia, mostly arranged in easily countable vertical rows. Vertigial, supralabral, and labral setae: two, four, and 22 - 26, respectively. Antennae (in Fig. 21 B) 1.40 - 1.55 x as long as head in males and 1.25 - 1.45 x in females; antennomere 2> 3 = 5> 4> 6. Gnathochilarium with a relatively small promentum separating lamellae linguales by <1 / 3 of their length, each latter with five setae in a longitudinal row; stipites in males basally with a kidney-shaped swelling densely covered with fine and stiff setae. Collum mostly smooth, with only several very short and fine striae near posterolateral corner. Body rings somewhat vaulted (more significantly in caudal part of body). Prozonae with very short and fine longitudinal striae in their hind sections. Metazonae rather shallowly striated, n Schub = 8 or 9 in males and 11 or 12 in females; metazonal setae 2 / 5 - 1 / 2 of metazonal length. Ozopores set tightly behind pro-metazonal suture in more anterior rings, gradually moved further back, to nearly equal to their diameter behind the suture in caudalmost rings, sutures not sinuous in front of ozopores. Tarsus of mid-body legs 1.1 - 1.2 as long as tibia and ca. 4 x as long as apical claw. Telson (in Fig. 21 C): Epiproct long (reaching the level of longest paraproctal setae), straight or slightly bent dorsad, ending with a minute hyaline tip. Hypoproct broadly rounded, ventrally with two median submarginal setae; barely protruding behind rear contour of paraprocts in males, tightly adhering to their ventral side in females. Paraprocts sparsely to moderately densely covered with long setae; no distinct rows of shorter setae along caudal margins. Male sexual characters: Mandibular stipites (in Fig. 21 B) considerably expanded, broadly rounded, protruding mostly ventrad, with either an indistinct or strongly obtuse and blunt ventro-anterior corner. Leg pair 1 compact hooks, slightly to considerably turned against one another. Leg pair 2 slightly thicker than following legs, with a weakly pronounced adhesive pad distoventrally on tibia, without pad on postfemur; following pairs with two pads, one each on postfemur and tibia, both clearly discernible until caudalmost pairs; femora without modifications. Pleurotergum 7 strongly enlarged, ventrally forming rather slender, fingertip-shaped lobes (Fig. 22 A) originating entirely from metazona, protruding ventrad behind gonopods. Penis (Fig. 22 B) somewhat longer than broad, with very short, barely discernible apical lobes, and with broad, triangular, terminal lamellae directed completely caudad. Gonopods (Figs 22 C, 23): In situ considerably protruding from gonopodal sinus. Promere (Fig. 23 B, p in Fig. 23 A) considerably higher than opisthomere, relatively slender; mesal margin more or less straight, lateral margin gently sinuous, distally both margins symmetrically slanting towards a narrowly rounded apex; caudal surface with a rather short, strongly pronounced, median ridge, a relatively deep and broad median groove, a broad distal groove, and a small, rounded, micro-squamous, distolateral lobe; flagellum somewhat longer than height of promere, distally with rows of blunt teeth directed basad. Opisthomere (Figs 22 C, 23 A, C-E) relatively elongated; basoposterior process shaped like a weakly pronounced lobe, distally forming a rather narrow apical outgrowth with a flat caudal surface and densely serrate / dentate margins; anterior process large, distally tapering, bent caudad, reaching just below tip of solenomere; mesomeroidal lobe broad and rounded, weakly pronounced, positioned slightly above opisthomeral mid-height, forming a slender apicolateral part reaching as high as anterior process, proximally fused to the latter; lateral side with a rounded hump-like lobe; mesal side with a broad and flattened lobe (presumably gonocoxal gland), and a rather deep and spacious anteromesal sinus; a group of very long, erect, spiniform filaments at proximal section of flagellum channel; solenomere unipartite, tubular, relatively narrow, bent caudad, meso-apically with a fine digitiform process. Female sexual characters: Leg pairs 1 and 2 markedly thicker, first also shorter, than following legs. Vulva (Fig. 22 D) mostly symmetrical, elongated, considerably compressed on sides; bursa with a strongly obtuse postero-apical margin; operculum slightly shorter than bursa, both parts ending with small hyaline protrusions; setation dense throughout. Receptaculum seminis consisting of a rather narrow, mostly straight, central tube, abruptly arched posteriad right before junction with a large, oblong, central ampulla, and a very narrow, somewhat folded, posterior tube leading to a rather large, pyriform, posterior ampulla.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
4CCB2E36BE8456A89A2F7D285177C9BD.taxon	distribution	General distribution. LECA.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
05723FAECB805CE5A479BA890DE3DBD4.taxon	description	Fig. 31	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
05723FAECB805CE5A479BA890DE3DBD4.taxon	materials_examined	Material examined. Syntype slide (gonopods, flange of male body ring 7) (ZMB 13622; Verhoeff prep. 3794). Russia: Republic of Adygea: 1 ♂ (SMNG 33579), Mount Shibaba, 21. V. 2004, K. Voigtlaender leg.; 4 ♂♂, 1 ♀ (SMNG 33575), at Sakhray River, 20. V. 2004, K. Voigtlaender leg.; 2 ♂♂, 5 ♀♀, 7 juv. (SMNG 33576), same collecting data.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
05723FAECB805CE5A479BA890DE3DBD4.taxon	diagnosis	Diagnosis. A species of Omobrachyiulus morphologically very similar to O. zuevi sp. nov. Differing from the latter species by having generally more slender and less strongly embossed gonopods, and by certain details of opisthomere structure, as follows: anterior process well-developed, linguiform, clearly visible from most aspects, vs. the same being vestigial, visible only in oral and oral-mesal views in O. zuevi sp. nov.; mesomeroidal lobe with a rather weakly developed apicolateral part, vs. the same being massive, nearly reaching the level of the solenomere in O. zuevi sp. nov.; solenomere apically only moderately enlarged, with a soft sigmoid process directed basad, vs. the same ending with a very broad, flower-like apex in O. zuevi sp. nov.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
05723FAECB805CE5A479BA890DE3DBD4.taxon	distribution	General distribution. WCIS.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
BD441442676D5FF987B36DC206601CDE.taxon	materials_examined	Material examined. Armenia: Elenovka [Sevan], 2 ♂♂ (ZIN) (gonopods dissected, not in the vial), 4 ♀♀ (vulvae dissected, not in the vial), 10. VIII. 1932, A. Schelkovnikov leg.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
BD441442676D5FF987B36DC206601CDE.taxon	diagnosis	Diagnosis. A species of Omobrachyiulus most simillar to O. kvavadzei sp. nov., O. ponticus sp. nov., and O. trochiloides sp. nov. in gonopod structure, as well as in the shape of the male telson. Differs from these species mostly by details of the opisthomere: from O. kvavadzei sp. nov. by the significantly lower position of the mesomeroidal lobe devoid of a distinct apicolateral part; from O. ponticus sp. nov. by both the anterior process and the apical outgrowth of the basoposterior process being subequal to, rather than considerably exceeded by, the solenomere; and from O. trochiloides sp. nov. by having a much more obscure, poorly differentiated basoposterior process ending in an unciform, rather than a fine rod-like, apical outgrowth.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
BD441442676D5FF987B36DC206601CDE.taxon	distribution	General distribution. LECA.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
A7E646D3C74A5A2DBA1F2628BA921E6B.taxon	description	Figs 44, 45, 46	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
A7E646D3C74A5A2DBA1F2628BA921E6B.taxon	materials_examined	Material examined. Holotype: ♂ (in head to ring 6, pleurotergum 7, and rest of body; distal part of left antenna broken off, gonopods and right flange of pleurotergum 7 on permanent slide) (ZMUM), Georgia, Borjomi Nature Reserve, Baniskhevi Valley, 800 - 900 m, Picea, Fagus and Carpinus forest, in litter, logs and under stones, 12 and 16. V. 1983, SIG leg. Paratypes: 1 ♂ (in head to ring 2, ring 3 to ring 6, and rest of body; gonopods prepared for SEM, penis, leg pair 2, leg 11 and mid-body leg on permanent slide) (ZMUM), 4 adult ♀♀ (one in head to ring 3 + rest of body, left vulva dissected) (ZMUM), 2 subadult ♀♀ (ZMUM), same collecting data as of the holotype; 1 ♂ (in head, collum to ring 6, pleurotergum 7 (in pieces), and ring 8 to rest of body; right antenna, leg pair 1, left flange of pleurotergum 7, and a mid-body leg dissected) (NMNHS), 1 adult and 2 subadult ♀♀ (unbroken) (NMNHS), Georgia, Chokhatauri District, near Bakhmaro, 4 km SSE Nabeglavi, 1950 - 2020 m, Abies nordmanniana, 8. VI. 1981, SIG and J. Martens leg.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
A7E646D3C74A5A2DBA1F2628BA921E6B.taxon	diagnosis	Diagnosis. A species of Omobrachyiulus most simillar to O. sevangensis comb. nov., O. kvavadzei sp. nov., and O. ponticus sp. nov. in gonopod structure, as well as in the shape of the male telson. Easily distinguishable from all these species by the massive, well-differentiated basoposterior process of the opisthomere, ending with a very fine and slender apical outgrowth, as well as by the solenomere being unipartite rather than bifurcated apically.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
A7E646D3C74A5A2DBA1F2628BA921E6B.taxon	description	Description. Measurements: male holotype in S VIII, 39 + 2 + T, L = 17.5 mm, H = 1.1 mm; male paratypes in S VIII, 37 + 2 + T, L = 12.8 mm, H = 1 mm, and 42 + 2 + T, L = 18 mm, H = 1.2 mm; adult female paratypes in S IX-X, 40 - 46 + 0 - 1 + T; L = 21 - 24 mm, H = 1.4 - 1.95 mm; subadult female paratypes in S VIII, 36 - 42 + 2 + T. Colouration (after> 35 years in alcohol) (Fig. 44): Head brown with a broad, dark band between ommatidia; collum brown, with a much darker frontal quarter and a dark posterior margin; frontal parts of prozonae light grey to creamy, posterior parts dark grey with some brownish spots; frontal parts of metazonae dark brown, posteriorly light ochre-brown; whole ventral side of body light brownish beige, each prozona ventrolaterally with a large, darker, blurred spot; paraprocts brown-grey, lighter caudally; dorsum with a black axial line; legs yellowish, tibiae and tarsi in males brown. External structures: Eye patches in adults consisting of 28 - 35 ommatidia arranged in easily recognisable vertical rows. Vertigial, supralabral and labral setae: two, four, and 17 - 20, respectively. Antennae ca. 1.6 x as long as head in males and 1.2 x in females; antennomere 2> 3 ~ 5> 4> 6. Gnathochilarium with a promentum separating lamellae linguales in their proximal 2 / 5, the latter with three or four setae in a longitudinal row; stipites with a group of short, stiff setae basomedially near the borders with the lamellae. Collum mostly smooth, with four or five very short and shallow striae at posterolateral corners. Body rings weakly vaulted. Prozonae with scattered minute grooves. Metazonae rather shallowly striated, n Schub = 9 or 10; metazonal setae relatively short (ca. 1 / 2 of metazonal length) and erect. Ozopores placed right in pro-metazonal suture in first several rings, gradually taking a more posterior position to ~ 1 / 2 of their diameter in caudalmost rings; sutures not sinuous in front of ozopores. Tarsus of mid-body legs 1.4 - 1.5 x as long as tibia and 3.6 - 3.9 x as long as apical claw. Telson (Fig. 44 B): Epiproct very long, straight to slightly bent ventrad, ending with a pointed hyaline tip turned ventrad, considerably surpassing longest paraproctal setae in both sexes. Hypoproct in males trapezoidal, tridentate, teeth well-developed, robust, protruding behind rear contour of paraprocts; the same being broadly rounded, completely concealed below paraprocts in females; ventral surface with five submarginal setae. Paraprocts moderately densely setose (ca. 20 long setae each side), without distinct rows of shorter setae along caudal margins. Male sexual characters: Mandibular stipites moderately expanded, protruding ventro-anteriad, forming a broadly rounded anterior corner. Leg pair 1 (Fig. 45 A, B) compact hooks turned somewhat against one another, bearing minute tarsal remnants (white arrow) in some specimens. Leg pair 2 and following pairs with crested adhesive pads ventrally on tibia and postfemur, these gradually reduced towards telson, completely disappearing in caudalmost legs; femora of legs 3 - 7 with an oval groove. Pleurotergum 7 ventrally forming large rounded lobes (Fig. 45 C) originating mostly from metazona, protruding ventromesad behind gonopods. Penis (Fig. 45 D, E) somewhat longer than broad, strongly compressed anterocaudally, with diverging apical lobes and well-developed, tapering, terminal lamellae. Gonopods (Fig. 46): In situ protruding from gonopodal sinus with their apical parts, laterally not concealed by protrusions of pleurotergum 7. Promere (Fig. 46 A) as high as opisthomere, being conspicuously similar to that of O. kvavadzei sp. nov. and O. ponticus sp. nov. in overall shape, and particularly in having a short and strongly pronounced median ridge and a distomesal micro-squamous lobe; differing mostly by the broader apex. Opisthomere (Fig. 46 B-E) rather slender; basoposterior process a well-developed and massive lobe, deeply divided from CBO, ending in a small, pointed, apical outgrowth bent anteriad; anterior process shaped as a furrowed lamellar ridge, completely fused to CBO; mesomeroidal lobe strongly pronounced, positioned at base of opisthomere, without any expanded parts; solenomere markedly slender, somewhat bent posteriad; only a few scattered spiniform filaments near flagellum channel. Female sexual characters: Leg pairs 1 and 2 somewhat thicker than following legs, leg pair 3 also slightly thicker and somewhat longer than neighbouring pairs. Vulva (Fig. 45 F) very similar to that in O. ponticus sp. nov., including the external shape of the bursa, the structure of the receptaculum seminis, and the very high operculum; differs by a larger opening and a smaller central tube in relation to both posterior tube and ampulla.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
A7E646D3C74A5A2DBA1F2628BA921E6B.taxon	distribution	General distribution. LECA.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
0941EA70CE9D595E8BCE1FAFB9B2D21A.taxon	description	Figs 14, 15, 16	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
0941EA70CE9D595E8BCE1FAFB9B2D21A.taxon	materials_examined	Material examined (SMNG). Holotype: ♂ (in head to ring 6, pleurotergum 7 to ring 29, and rest of body, gonopods, left antenna, right flange of pleurotergum 7, hypoproct, leg 6, a mid-body, and an end-body leg dissected), Georgia, Samegrelo-Zemo Svaneti, Mestia, source of the valley above Lengeri, 43.0742 ° N, 42.6983 ° E, 2060 m a. s. l., 27. IX. 2012, F. Walther leg. Paratype: 1 subadult ♂ (in head and collum and rest of body), same collecting data as for holotype.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
0941EA70CE9D595E8BCE1FAFB9B2D21A.taxon	diagnosis	Diagnosis. A species of Omobrachyiulus very similar to O. divaricatus by the general shape of the promere and by the opisthomere having a slender, rod-like, anterior process directed distad, a broad, strongly flattened, multilobed, apical outgrowth of the basoposterior process, and a similarly finely and asymmetrically bifurcate solenomere. Differs from O. divaricatus by the following gonopodal characters: promere with a more broadly rounded apex; apical outgrowth of opisthomeral basoposterior process asymmetrically bilobed, forming a dentate distal part and a thumb-like lateral part, vs. the same being more or less symmetrical, tripartite in O. divaricatus; anterior process of opisthomere significantly lower rather than equal to, or higher than, solenomere; solenomere mostly straight, directed almost completely distad, vs. the same being abruptly bent caudad in O. divarictus; and by the male hypoproct being tri-, rather than penta- or septidentate.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
0941EA70CE9D595E8BCE1FAFB9B2D21A.taxon	description	Description. Measurements: holotype ♂ in S in XI, 50 + 1 + T, L = 29 mm, H = 1.95 mm. Paratype subadult ♂ in S X, 47 + 2 + T. Colouration (Fig. 14): Mostly brown-grey with purple tinges; head brown, with the usual pattern; collum mostly grey, dorsally brownish, with dark brown margins; prozonae grey, ventrolaterally with numerous light spots; metazonae grey, with a transverse dark brown stripe in posterior half, this abruptly narrowing below ozopore level, hind margins light brown-beige; dorsum with a black axial line; epiproct dark brown-grey, paraprocts somewhat lighter brown; legs light yellow. External structures: Eye patches consisting of ca. 45 ommatidia arranged in easily recognisable vertical rows. Vertigial, supralabral, and labral setae: two, four, and 18, respectively. Antennae (Fig. 14 A) ca. 1.7 x as long as head in males; antennomere 2> 5 ≥ 3> 4> 6. Gnathochilarium with promentum of moderate size, separating lamellae linguales in ca. 2 / 5 of their length, each latter with three setae in a longitudinal row. Collum mostly smooth, with several short and shallow striae at posterolateral corners. Body rings not vaulted. Prozonae with numerous minute grooves in posterior third. Metazonae relatively shallowly striated, n Schub = 10 or 11; setae (apparently) mostly abraded, rather short based on the few remaining. Ozopores set tightly behind pro-metazonal suture in more anterior rings, gradually moved further back, to nearly equal to their diameter behind the suture in caudalmost rings, sutures gently sinuous in front of ozopores in some rings. Tarsus of mid-body legs slightly shorter than tibia, and slightly over 3 x as long as apical claw. Telson (Fig. 14 B): Epiproct moderately long, straight, broad, roof-like, ending with a short and blunt hyaline tip equal to, to slightly surpassed by, the longest paraproctal setae. Hypoproct (in males) (Fig. 15 A) broadly trapezoidal, tridentate, ventrally with six submarginal setae, only the teeth protruding behind rear contour of paraprocts. Paraprocts moderately densely setose, with a row of sparse short (ca. half as long as the ' regular' ones) and stiff setae along each caudal margin. Male sexual characters: Mandibular stipites (in Fig. 14 A) considerably expanded, protruding mostly anteriad, forming a narrow anterior corner. Leg pair 1 compact parallel. Walking legs (Fig. 15 B) with well-developed crested pads, both tibial and postfemoral ones gradually reduced towards telson, but still visible in caudalmost pairs; femora without modifications. Pleurotergum 7 (Fig. 15 C) ventrally forming rather slender and rounded lobes originating from the zone around pro-metazonal suture, protruding mostly ventrad behind gonopods. Gonopods (Fig. 16): Promere (Fig. 16 B, p in Fig. 16 A) relatively slender, broadest at base, mesal margin gently concave, lateral one bulging in mid-section, both joining into a broadly rounded apex; caudal surface with a strongly pronounced and relatively long median ridge, a narrow and rather deep median groove, and a broad and deep, but indistinctly marked, distal groove; flagellum just slightly longer than height of promere. Opisthomere (Fig. 16 A, C-E) rather slender; basoposterior process shaped as a weakly pronounced, hump-like lobe, distally forming a broad, bilobed, apical outgrowth: with an apically dentate distal lobe, and a rounded thumb-like lateral lobe; anterior process well-developed, slender and rod-like, with a slightly serrate apical part; mesomeroidal lobe large, but rather weakly pronounced, flattened; mesal side with a large lobe (presumably gonocoxal gland), and a very deep anteromesal sinus; several minute spiniform filaments along flagellum channel; solenomere long and slender, significantly exceeding anterior process; apically finely and asymmetrically bifurcate, directed almost completely distad and only slightly caudad. Female sexual characters: unknown.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
0941EA70CE9D595E8BCE1FAFB9B2D21A.taxon	distribution	General distribution. SWGC.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
119161A5478155C191C058D161704B8E.taxon	description	Figs 35, 36, 37	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
119161A5478155C191C058D161704B8E.taxon	materials_examined	Material examined (ZMUM). Holotype: ♂ (in head to ring 6 and rest of body, opisthomeres dissected), Stavropol, Mamayskiy Forest, 31. III. 2013, R. Zuev leg. Paratypes: 1 ♂ (in head to ring 2, ring 3 to ring 6, pleurotergum 7 (broken into two pieces), and rest of body; penis and pleurotergum 7 dissected, gonopods prepared for SEM), 1 ♀ (in head to body ring 2 + rest of body, left vulva dissected), same collecting data as for holotype.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
119161A5478155C191C058D161704B8E.taxon	diagnosis	Diagnosis. A species of Omobrachyiulus being morphologically very similar to O. roseni (Verhoeff, 1921). Differing from the latter species by having generally stouter, more robust gonopods, and by details of the opisthomere, as follows: anterior process minute, vestigial, mostly hidden between base of solenomere and distal part of mesomeroidal lobe, vs. the same being significantly larger, clearly visible from most angles in O. roseni; mesomeroidal lobe protruding in a massive, rounded, apicolateral part, and a much smaller mesoapical part, vs. the apicolateral part being much less pronounced, nearly same size as the mesoapical one in O. roseni; solenomere with a strongly enlarged, flower-like apex, vs. the same being apically only moderately enlarged, with a soft sigmoid process directed basad in O. roseni.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
119161A5478155C191C058D161704B8E.taxon	description	Description. Measurements: holotype ♂ in S VIII or IX, 40 + 2 + T, L = 10.5 mm, H = 0.8 mm; paratype ♂ in S VIII or IX, 42 + 2 + T, L = 12 mm, H = 0.85 mm; paratype ♀ in S IX or X, 44 + 1 + T, L = 14.5 mm, H = 1.15 mm. Colouration (Fig. 35): Mostly brown-beige, frontal and caudal parts of body darker than mid-body; head with the usual pattern; antennae light brown; collum dark brown at margins; prozonae with numerous densely set yellowish oval spots surrounded by brown rings forming a reticulate pattern; dorsally with a transverse dark brown stripe; metazonae light brown to yellowish beige; whole ventral side of body light beige; dorsum with a dark brown axial line; legs very light, in males dorsally with brown tinges. External structures: Eye patches in adults consisting of ca. 30 weakly pronounced ommatidia, arranged in hardly recognisable vertical rows. Vertigial, supralabral, and labral setae: two, four, and 13 or 14, respectively. Antennae (in Fig. 35 A, B) ca. 1.3 x as long as head in males and 1.15 x in females; antennomere 2> 3 = 5> 4> 6. Gnathochilarium with relatively small promentum separating lamellae linguales in ca. 1 / 3 of their length, each latter with four setae in a longitudinal row. Collum mostly smooth, with only three or four very shallow, barely visible grooves near posterolateral corners. Body rings not vaulted. Prozonae with very short and fine, mostly parallel longitudinal striae. Metazonae rather shallowly striated, n Schub = 6 or 7; setae absent. Ozopores set tightly behind pro-metazonal suture in more anterior rings, gradually moved further back, to nearly equal to their dm behind the suture in caudalmost rings, sutures slightly (in the males) to strongly (in the female) sinuous in front of ozopores in anterior body section. Tarsus of mid-body legs as long as tibia, and slightly> 3 x as long as apical claw. Telson (Fig. 35 C): Epiproct moderately long, stout, turned somewhat ventrad in males, almost perfectly straight in females, ending with a short and blunt hyaline tip, not reaching the level of the longest paraproctal setae; with several long setae. Hypoproct semi-elliptical (more broadly rounded in the female) ventrally with two median submarginal setae, margin somewhat protruding behind rear contour of paraprocts in the males, tightly adhering to their ventral side in the female. Paraprocts sparsely covered with very long setae; no distinct rows of shorter setae along caudal margins. Male sexual characters: Mandibular stipites (in Fig. 35 A) moderately expanded, protruding mostly ventrad, forming a rounded anteroventral corner. Leg pair 1 compact hooks turned slightly to considerably against one another. Leg pair 2 and following pairs with crested adhesive pads, postfemoral ones disappearing after mid-body, tibial ones present until caudalmost legs; femora of leg pair 3 and next several pairs with an oval groove. Pleurotergum 7 ventrally forming elongated rounded lobes (Fig. 36 A) originating from the zone around pro-metazonal suture, protruding mostly mesad behind gonopods. Penis (Fig. 36 B, C) compact, stout, broader than long, with very short, contiguous, apical lobes ending in small terminal lamellae turned caudad. Gonopods (Figs 36 D, 37): Promere (Fig. 37 A) relatively broad and stout, as high as opisthomere, broadest at base, somewhat narrowing in mid-section, then widening again distally, before lateral margin abruptly slanting to join mesal margin in a narrow, blunt apex; caudal surface with a well-pronounced, strongly arched, median ridge, a relatively deep and very narrow median groove, and a broad and deep distal groove; a broad and flat, subapical denticle turned basocaudad; flagellum somewhat longer than height of promere. Opisthomere (Figs 36 D, 37 B-E) stout and robust; basoposterior process present as a weakly pronounced lobe, distally forming a broad, flattened, shield-like, apical outgrowth protruding in a median, blunt, digitiform tip at margin; anterior process very small, mostly concealed behind distal part of mesomeroidal lobe, the latter massive, very strongly pronounced, distally extending into a broad and rounded apicolateral part, and a much lower, pointed, pyramidal apicomesal part; mesal side with a rather narrow lobe (presumably gonocoxal gland), and a very deep and spacious anteromesal sinus; a row of long and erect spiniform filaments at proximal section of flagellum channel; solenomere unipartite, rather broad tubular, turned anteriad, apically strongly broadening, forming a lamellar flower-like structure with multiple, minute, spiniform filaments at margin. Female sexual characters: Leg pairs 1 and 2 slightly thicker than following legs. Vulva (Fig. 36 E) symmetrical, relatively elongate, somewhat compressed on sides; bursa with a strongly obtuse postero-apical margin; operculum as high as bursa, both parts ending with small hyaline protrusions; setation dense throughout. Receptaculum seminis consisting of a rather broad, digitiform, central tube, and a very narrow, short, mostly straight, posterior tube gradually widening into a small, drop-like, posterior ampulla.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
119161A5478155C191C058D161704B8E.taxon	distribution	General distribution. WCIS.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
A198C84EFCDF5695A5259E142CDE39F6.taxon	description	Figs 47, 48, 49	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
A198C84EFCDF5695A5259E142CDE39F6.taxon	materials_examined	Material examined (ZMUM): Holotype: ♂ (in head and 5 pieces, left antenna, penis, leg pairs 1 and 2, penis, gonopods, right flange of pleurotergum 7, mid-body leg, and hypoproct dissected), Georgia, Svanetia, mouth of Nenskra River, Lukhi, N of Khaishi, ca. 800 m a. s. l., leaf litter, 2. IX. 1986, A. Ryvkin leg. Paratype: 1 ♀ (in 2 pieces, left vulva dissected), 5 juv. (unbroken), same collecting data as for holotype.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
A198C84EFCDF5695A5259E142CDE39F6.taxon	diagnosis	Diagnosis. Differs from its only known congener, S. waltheri gen. nov., sp. nov., mainly by being on average slightly smaller, by the relatively shorter male antennae, by the position and shape of the ventral protrusions of male body pleurotergum 7, and by the following details of gonopod structure: promere proportionally broader, abruptly narrowing only apically, with the mesal process significantly exceeding the apex, vs. the same being more slender, gradually narrowing distally, and with the mesal process just slightly exceeding the apex in S. waltheri gen. nov., sp. nov.; opisthomere possessing a distomesal process vs. this being absent from S. waltheri gen. nov., sp. nov., having a broad and flat apical margin of the basoposterior process, vs. the same forming a distinct apical corner in S. waltheri sp. nov., and with the solenomere gradually narrowing all the way to the top, vs. the same being somewhat enlarged apically in S. waltheri gen. nov., sp. nov.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
A198C84EFCDF5695A5259E142CDE39F6.taxon	description	Description. Measurements: holotype in S IX, 42 + 1 + T, L = 23 mm, H = 1.6 mm; paratype ♀ in S X, 46 + 1 + T, L = 27 mm, H = 2.2 mm. Colouration (> 30 years in alcohol) (Fig. 47): Head and collum with the usual colour pattern, antennae dark brown; trunk dorsally mostly brown-grey, with a blackish axial line and with dark grey to blackish transverse stripes running along pro-metazonal suture on both pro- and metazonae, reaching down to ozopore level; contrasting lighter brown-beige below ozopore level; frontal sections of prozonae brown-beige, posteriorly with an irregular dark brown spot around ozopore; metazonae mostly lighter than prozonae, their posterior sections with a dark brown band encircling entire ring, these narrowing and lightening ventrally; hind margins ochre-brown; epiproct dorsally dark grey, laterally dark brown; hypoproct and proximal part of paraprocts mostly light brown; legs yellowish. External structures: Eye patches consisting of 27 or 28 and 37 or 38 markedly convex ommatidia, in the male and female, respectively; developmental rows easily countable. Vertigial, supralabral, and labral setae: two, four, and 20, respectively. Antennae ca. 1.35 x as long as head in the male, and 1.25 x in the female; antennomere 2> 3 ≥ 5 ≥ 4> 6. Gnathochilarium with a moderately sized promentum separating lamellae linguales by nearly half their length, each latter with four setae in a longitudinal row; male stipites basolaterally each with a faint oblong hump bearing several setae. Collum mostly smooth, with only several sparsely set, short and shallow grooves near posterolateral corners; frontolateral margin gently concave. Body rings considerably vaulted. Prozonae with densely set, very short, shallow, parallel, longitudinal striae in their hind sections. Metazonae moderately deeply striated, n Schub = 9 or 10 in the male, and 10 or 11 in the female; setae erect to somewhat slanting, from ca. 2 / 5 (in mid-body and caudal rings) to equal to metazonal length (in anteriormost rings). Ozopores set tightly behind pro-metazonal suture in more anterior rings, and ca. half their diameter behind in more posterior ones; sutures gently to considerably sinuous in front of ozopores in most rings. Tarsus of mid-body legs equal to tibia, and slightly> 3 x as long as apical claw. Telson (Fig. 47 B): Epiproct rather long (slightly exceeding the level of longest paraproctal setae in the male, just reaching it in the female) and straight, ending up with a well-developed, blunt, unciform, hyaline tip bent dorsad; with several long setae on dorsal side. Hypoproct in the male (Fig. 48 A) rounded trapezoidal, with a thickened, faintly undulate, caudal margin slightly protruding behind caudal contour of paraprocts, this being broadly rounded, thinner, and tightly adhering under paraprocts in the female; with a pair of long distal paramedian setae. Paraprocts rather sparsely covered with long setae, without rows of shorter setae at caudal margins. Male sexual characters: Mandibular stipites (Fig. 47 A) considerably expanded, protruding ventro-anteriad, forming a subrectangular, rounded, ventro-anterior corner. Leg pair 1 compact parallel hooks. Leg pair 2 slightly shorter and thicker than following legs, both tibial and postfemoral adhesive pads well-developed, crested in anterior part of body, gradually reduced in the last third, completely absent from caudalmost legs; femora without modifications. Pleurotergum 7 ventrally forming blunt and spade-shaped lobes (Fig. 48 B) originating entirely from metazona, oriented with their broad sides parallel to the body axis, protruding mostly mesad, touching one another behind gonopods. Penis (Fig. 48 C) very small, deeply hidden above coxae 2, barrel-shaped, slightly longer than broad, strongly flattened dorso-ventrally, with very short apical lobes ending in short and blunt terminal lamellae directed distad. Gonopods (Fig. 49): Promere (Fig. 49 B, C and p in Fig. 49 A) relatively slender, somewhat higher than opisthomere, leaf-shaped, with a gently convex mesal margin, and a markedly sigmoid lateral one, both joining in a narrowly rounded apex; caudal surface with a massive and long mesal process originating at mid-height, ending with a short unciform apex turned baso-anteriad, considerably exceeding the apex of main promeral body; a short, well-pronounced, median ridge, a broad and weakly defined median groove, and a short and well-pronounced distal ridge just lateral to base of mesal process; flagellum just slightly shorter than height of promere. Opisthomere (Fig. 49 A, D-F) slender; basoposterior process present as a broad, anterocaudally flattened lobe ending with a flat apical margin bent caudad and bearing short and micro-papillary branches, mesally expanding into a short lobe with a coarse margin; mesomeroidal lobe a weakly pronounced ridge; a thumb-like distomesal process originating from border between basoposterior process and solenomere, directed distomesad; anteromesally at base a well-developed, elongate lobe (presumably gonocoxal gland); solenomere unipartite, long, flattened on sides, gradually narrowing distad, gently sigmoid, caudo-apically with a very fine and sharply pointed process; a dense row of small, erect, spiniform filaments at flagellum channel's mid-height. Female sexual characters: Leg pairs 1 and 2 somewhat shorter, 1 st also slightly thicker, than following legs. Vulva (Fig. 48 D) slightly asymmetrical (lateral valve of bursa more strongly slanting towards apex compared to mesal one), rather elongate, subconical, strongly compressed on sides; bursa with a distinct, obtuse, postero-apical margin; an opening absent, with the two valves completely fused all the way to bursal apex; operculum slightly shorter than bursa, both structures ending up with ear-like hyaline protrusions, bursal ones considerably larger; setation dense throughout. Receptaculum seminis composed of a short and very narrow central tube abruptly widening into an oblong central ampulla, forming several constrictions followed by a digitiform reservoir at bottom; and a long and thin, slightly folded, posterior tube, this somewhat widening before ending in a subspherical posterior ampulla.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
A198C84EFCDF5695A5259E142CDE39F6.taxon	distribution	General distribution. SWGC.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
63312AF70C1A58F9B18B247D975052CD.taxon	diagnosis	Diagnosis. A genus of Brachyiulini differing from contribal genera by the following combination of characters: promeres positioned anteriorly and only slightly laterally in relation to opisthomeres; caudal surface of promere bearing a strongly developed mesal process; opisthomere possessing a broad and flattened basoposterior process and a faint mesomeroidal lobe, and lacking lateral and anterior processes; solenomere simple, more or less slender, ending with a sharply pointed tip; vulva subconical, bursa with completely fused valves (forming neither an opening nor a cleft), a supposed autapomorphy.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
63312AF70C1A58F9B18B247D975052CD.taxon	description	General description. Medium-sized (L (males) = 23 - 28 mm) Brachyiulini. Ommatidia present. Ozopores set tightly behind pro-metazonal suture at least on more anterior body rings. Epiproct well-developed, relatively (not conspicuously) long. Hypoproct rounded trapezoidal to semi-circular, ventrally with two distal paramedian setae. Male mandibular stipites considerably expanded, forming a distinct ventro-anterior corner. Male pleurotergum 7 significantly bulging. Male walking legs ventrally with two well-developed adhesive pads, one each on postfemur and tibia. Penis short and stout, with very short apical lobes and small terminal lamellae. Gonopods: In situ: promeres considerably protruding outside gonopodal sinus, directed partly caudad, opisthomeres visible only with their apical parts. Promere slightly higher than opisthomere, elongate, bearing a long, stout, mesal process on caudal side; flagellum slightly shorter than height of promere. Opisthomere slender; basoposterior process well-developed, broad, flattened frontocaudally; anterior and apicoposterior processes absent; mesomeroidal lobe shaped as a weakly pronounced ridge; solenomere long and slender, distally bipartite, the frontal part ending in a fine, sharply pointed tip. Vulva: Subconical. Bursa without distinct postero-apical margin. Opening or median cleft absent - the two valves completely fused to one another. Operculum subequal in height to bursa. Receptaculum seminis: central tube short and narrow, ending in a distinct central ampulla; posterior tube long and narrow, ending in a subspherical posterior ampulla.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
C891E4EE1E0E5AD4BDDA2959F415A530.taxon	description	Figs 50, 51, 52	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
C891E4EE1E0E5AD4BDDA2959F415A530.taxon	materials_examined	Material examined (SMNG): Holotype: ♂ (in head to pleurotergum 7 and rest of body, gonopods intact), Georgia: Samegrelo-Zemo Svaneti, Lebarde, Tekhuri Valley, 30 rkm from Taleri towards Lebarde [at the influx of Lebarde and Tekhuri rivers], 42.7258 ° N, 42.4703 ° E, 1260 m a. s. l., 10. X. 2011, F. Walther leg. Paratypes: 1 ♂ (in 3 pieces, leg 7, mid-body and end-body leg, hypoproct, left flange of pleurotergum 7, and penis dissected, gonopods prepared for SEM), 1 ♀ (in head to body ring 3 and rest of body, vulvae dissected, right one prepared for SEM), same collecting data as of the holotype.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
C891E4EE1E0E5AD4BDDA2959F415A530.taxon	diagnosis	Diagnosis. Differs from its only known congener, S. ryvkini gen. nov., sp. nov., mostly by being on average slightly larger, by the relatively more slender male antennae, by the position and shape of the ventral protrusions of male pleurotergum 7, and by the following details of gonopod structure: promere more slender, distally significantly narrowing, with the mesal process barely exceeding the apex, vs. the same being somewhat stouter, only apically abruptly narrowing, and with the mesal process significantly exceeding the apex in S. ryvkini gen. nov., sp. nov.; opisthomere lacking a distomesal process, vs. this being present in S. ryvkini gen. nov., sp. nov., having the basoposterior process with a distinct apical corner, vs. the same ending in a broad and flat margin in S. ryvkini gen. nov. sp. nov., and with the solenomere being somewhat enlarged apically, vs. the same narrowing all the way to the top in S. ryvkini gen. nov., sp. nov.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
C891E4EE1E0E5AD4BDDA2959F415A530.taxon	description	Description. Measurements: holotype in S X, 43 + 1 + T, L = 26 mm, H = 1.8 mm; paratype ♂ in S IX, 43 + 1 + T, L = 27.5 mm, H = 1.75 mm; paratype ♀ in S IX, 44 + 1 + T, L = 37.5 mm, H = 2.65 mm. Colouration (apparently somewhat faded) (Fig. 50): Similar to S. ryvkini gen. nov., sp. nov., but generally lighter, predominantly grey rather than brown, without significant difference between dorsal and ventral sides, and without transverse dark stripes along pro-metazonal sutures; prozonae dark grey, with numerous, densely set, light brown-beige spots arranged transversely above and below ozopore level; metazonae mostly grey, with numerous, minute, dark brown / blackish dots throughout, dorsally with a very narrow, transverse, blurred, dark brown band in posterior section; hind margins ochre-brown. External structures: Eye patches in adults vertically rather elongate, due to partial reduction of the higher developmental (vertical) rows, consisting of 21 - 35 markedly convex ommatidia. Vertigial, supralabral, and labral setae: two, four, and 16 - 18, respectively. Antennae (Fig. 50 A) 1.6 - 1.8 x as long as head in the males, and 1.25 x in the female; antennomere 2> 5 ≥ 3> 4> 6. Gnathochilarium as in S. ryvkini gen. nov., sp. nov., but lamellae linguales with three rather than four setae in a longitudinal row, and male stipites without setose hump. Body rings slightly to moderately vaulted. Male hypoproct (Fig. 51 A) very similar to that in S. ryvkini gen. nov., sp. nov., but almost perfectly semi-circular and with a densely and bluntly serrate margin, rather than apically somewhat flattened and with only several faint undulations. Tarsus of mid-body legs subequal to, to slightly longer than, tibia, and ca. 3 x as long as apical claw. Male sexual characters: Walking legs with adhesive pads still present in caudalmost pairs. Pleurotergum 7 ventrally forming rather narrow and oar-like lobes (Fig. 51 B) with a slightly concave apical margin, originating from border zone between pro- and metazona, protruding mostly ventrad behind gonopods. Penis (Fig. 51 C) similar to that in S. ryvkini gen. nov., sp. nov., but trapezoidal (widening distally) rather than barrel-shaped. All other external somatic characters as in S. ryvkini gen. nov., sp. nov. Gonopods (Figs 51 D, 52): Promere (Fig. 52 A) similar to that in S. ryvkini gen. nov., sp. nov., but more slender, distally mostly symmetrical rather than with a markedly convex lateral margin; mesal process barely, rather than considerably, surpassing the apex of promere, and ending with a finer, less strongly bent tip; median and distal ridges less strongly pronounced compared to the type species. Opisthomere (Figs 51 D, 52 C-F): lacking a distomesal process; basoposterior process narrower in comparison to S. ryvkini gen. nov., sp. nov., with a distinct apical corner rather than with a flat margin, and with a much smaller mesal lobe; mesal side with a slender lobe (presumably gonocoxal gland) and a rather deep and narrow anteromesal sinus; solenomere somewhat enlarged distally rather than narrowing; otherwise of a very similar shape to that of the type species. Female sexual characters: First two leg pairs of same condition as in S. ryvkini gen. nov., sp. nov. Vulva (Fig. 51 E) externally also very similar to the type species, mainly differing in being more elongated and with the operculum being equal to, rather than slightly shorter than, bursa. Receptaculum seminis with a somewhat wider and shorter central tube, not forming a distinct ampulla at bottom; posterior tube gradually widening towards bottom, rather than constricting to form an ampulla.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
C891E4EE1E0E5AD4BDDA2959F415A530.taxon	distribution	General distribution. SWGC.	en	Vagalinski, Boyan, Golovatch, Sergei I. (2021): The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae). ZooKeys 1058: 1-127, DOI: http://dx.doi.org/10.3897/zookeys.1058.68628, URL: http://dx.doi.org/10.3897/zookeys.1058.68628
