identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
84082A4BC02DB51547F03DE5EC88FCC7.text	84082A4BC02DB51547F03DE5EC88FCC7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bryodelphax arenosus Gąsiorek 2018	<div><p>Bryodelphax arenosus new species</p> <p>(Figs. 2–4, Tables 2, 3)</p> <p>Material examined. All individuals were mature females. Holotype (slide no. MY.813.01) and 14 paratypes (slides MY.813.01–11), six additional processed for DNA extraction and sequencing. Type locality: Bako Peninsula, lowland coastal rainforest close to the Tajor Waterfall (1°43′24″N, 110°27′50″E; 106 m asl), moss from a tree, coll. Piotr Gąsiorek &amp; Artur Oczkowski, 5 th August 2017. Holotype and nine paratypes deposited in the <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.46388&amp;materialsCitation.latitude=1.7233334" title="Search Plazi for locations around (long 110.46388/lat 1.7233334)">Institute of Zoology</a> and <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.46388&amp;materialsCitation.latitude=1.7233334" title="Search Plazi for locations around (long 110.46388/lat 1.7233334)">Biomedical Research</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.46388&amp;materialsCitation.latitude=1.7233334" title="Search Plazi for locations around (long 110.46388/lat 1.7233334)">Jagiellonian University</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.46388&amp;materialsCitation.latitude=1.7233334" title="Search Plazi for locations around (long 110.46388/lat 1.7233334)">Kraków</a>, Poland (slides MY.813.01–06), three paratypes deposited in the <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.46388&amp;materialsCitation.latitude=1.7233334" title="Search Plazi for locations around (long 110.46388/lat 1.7233334)">Natural History Museum</a> of Denmark, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.46388&amp;materialsCitation.latitude=1.7233334" title="Search Plazi for locations around (long 110.46388/lat 1.7233334)">Copenhagen</a> (slides MY.813.07–09), and last two paratypes deposited in the Raffles Museum of Biodiversity Research, National University of Singapore (slides MY.813.10–11).</p> <p>Description of the new species. Adults (i.e., from the third instar onwards, with developed gonopore; measurements in Table 2).</p> <p>Body yellowish, pearly opalescent; eyes absent or not visible after preparation. Both primary and secondary clavae oval and small (Figs. 2, 3C, 4A). Cirri interni and externi with poorly developed cirrophores. Cirri A long (on average reaching ⅓ of the body length, whereas Bryodelphax typically has cirrus A /body length ratio around ¼; Figs. 2, 3A). Dorsal plates with faint intra-cuticular pillars visible only under 100× magnification with immersion oil (Figs. 3C, 4), better visible on the scapular and caudal plates (Figs. 3C, 4A, C), pillars on the posterior portion of the caudal plate are also the largest (Figs. 3C, 4C). Distinct dark ridges are present on lateral portions of the scapular, paired, and caudal plates, forming characteristic and stable ornamented pattern (Figs. 3B, C, 4). Pseudopores or pores (SEM observations are needed to verify the morphology of the cuticular depressions) minute and barely visible (visible in PCM as bright dots with blurred margins; Figs. 3C, 4), sometimes absent. Pseudopores/pores distributed unevenly, i.e., on the anterocentral portion of the scapular plate 8–21 pseudopores/pores per 100 μm 2 were observed (x=17, N=10; Fig. 4A), on the central portion of the caudal plate: 11–18 pseudopores/pores per 100 μm 2 (x=15, N=10; Fig. 4C), whereas sparse and bleak pseudopores/pores occur rarely on other, especially paired, plates. Scapular plate deeply faceted by dark ridges, with six or eight lighter rectangular areas in the central portion (Figs. 3A, C, 4A). Scapular sutures clearly separate lateral portions from the central part, extending from the base of cirrophore A to the posterior margin of the plate (Figs. 3B, C, 4A). Paired plates divided into two roughly equal anterior and posterior parts by a transverse stripe (Figs. 3, 4B). Caudal plate mildly faceted in the central part by the same dark ridges as the scapular plate (Fig. 3A), but the faceting is fainter and often poorly visible (Figs. 3C, 4C). Median plates 1 and 2 divided transversally into two unequal parts (Figs. 2, 3), and additionally the anterior portion of median 2 has a transverse suture (Fig. 3). Median plate 3 large, triangular in shape, and with a roundish posterior edge (Figs. 2, 3). Thin, non-sculptured supplementary lateral platelets present at the levels of median plates (three pairs of platelets on each body side: a pair between scapular plate and first pair of the segmental plates, a pair between paired plates, and a pair between second pair of segmental plates and caudal plate; Fig. 3B, C). Ventral cuticle smooth, without plates or intra-cuticular pillars. Papillae on legs I and IV absent or not detectable in PCM. Large, sclerotised pedal plates IV present, with irregular margin, but never forming a dentate collar (Fig. 3C, white arrowhead). External claws of all legs smooth, internal claws with minuscule bumps near their bases, which can be spurs (Fig. 3C, insert). However, they are not measurable in PCM since barely divergent from the claw branch.</p> <p>No juveniles, larvae (i.e., the second and first instars, respectively) or eggs were found.</p> <p>Remarks. All found specimens were mature females, thus it is likely that the new species is parthenogenetic.</p> <p>DNA sequences. New 18S and 28S rRNA sequences were obtained for two specimens; each fragment was represented by a single haplotype and is deposited in GenBank: MH 414953 (18S rRNA, 1112 bp long), XX000000 (28S rRNA, 688 bp long). p-distances between sequenced species of Bryodelphax ranged from 0.8% to 2.8% (18S rRNA), and from 7.5% to 19.3% (28S rRNA) (Table 3).</p> <p>Etymology. From Latin (h) arenosus = sandy, inhabiting sand. The new species was found in coastal lowland rainforest with peculiarly sandy soil (arenosols), atypical for this biome.</p> </div>	http://treatment.plazi.org/id/84082A4BC02DB51547F03DE5EC88FCC7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Gąsiorek, Piotr	Gąsiorek, Piotr (2018): New Bryodelphax species (Heterotardigrada: Echiniscidae) from Western Borneo (Sarawak), with new molecular data for the genus. Raffles Bulletin of Zoology 66: 371-381, DOI: http://doi.org/10.5281/zenodo.5358943
