taxonID	type	description	language	source
03E887AB59593153FF3D6CADFCFCEF5F.taxon	description	T y p e m a t e r i a l 1 syntype (lectotype by present designation), relatively well preserved (only segments 6 - 11 of left antenna and segments 2 - 11 of right antenna missing), and without attached label under. Close to the specimen, the following handwritten label is pinned on the bottom of box: " colchica Chaud. Monts Taurus 48. Nordman. " (MNHN, " Collection Chaudoir ", box no 216 " Argutor "). As the original description does not implies that there are one or more syntypes, a lectotype by inference of " holotype " for this species is designated following Article 74.6. of the International Code of Zoological Nomenclature (ICZN 1999). T y p e l o c a l i t y. " Adjara " (CHAUDOIR 1850: 78). As far as the choice of type locality was based on a presumption of Chaudoir, we have doubt about the real place of finding the holotype and thus about the actual distribution of this species. In the last paragraph of the description, CHAUDOIR (1850: 78) write: " M. de Normann l’a rapportée de son voyage en Abkhasie, en Mingrélie et dans le Gouriel; je presume qu’il l’a trouvé sur le plateau de l’Adjara, car évidemment un insecte alpin. ". The supposition for inaccurate type locality is also supported by the label data of the syntype (e. g. " Monts Taurus … ") and by data of the morphology and distribution of the related taxa; see below. M a l e g e n i t a l i a. Median lobe of aedeagus in lateral view (Figs 1 - 2) with ostium deflected to left; apical and basal parts equal in length, apical one with slanting dorsal and straight ventral margin, so that apex became pointed at tip; basal part with strongly developed and prominent ventral process; internal sac in apical position inside, with single strongly chitinised piece (visible in left lateral position). Apex of median lobe dorsally (Fig. 3) with tip deflected to left. Left paramere (apical part broken) (Figs 4 - 5) with apical and basal parts bent each other almost at right angle. After drawing, the genitalia were placed in Euparal on the same label on which the specimen is glued. T a x o n o m i c n o t e s a n d r e l a t i o n s h i p s The presence of scanty pubescence on the distal part of left antennomere 3, dense and continuous punctuation on the basal part of pronotum (which also prolongs forward along the lateral margins and reaches the anterior part of pronotum), absence of medial setiferous punctures on the hind coxae, glabrous ventral side of onychium, and presumable sickle-shaped left paramere demonstrate that this species belongs to Oreoplatysma, and do not to Cryobius. The following transfer is proposed: Pterostichus (Oreoplatysma) colchicus (CHAUDOIR 1850), new assignment of Pterostichus (Cryobius) colchicus (CHAUDOIR 1850). Details in the external morphology and morphology of both the aedeagus and of the left paramere suppose that this species is allied to several representatives of the group " koenigi ", like P. cecchiniae (JAKOBSON 1907), P. rousianus KIRSCHENHOFER 1987, and P. strasseri REITTER 1898. But, according to the best of our knowledge, no Oreoplatysma species with pubescent distal part of antennomeres 3 lives in the territories south of the Caucasus Major (except P. colchicus and P. kadleci DVO ÁK 1995), and especially in the area of Adzaria. P. kadleci from the Kop Da i (NE Turkey) is distinct from P. colchicus in the very thin and long apex of aedeagus. P. colchicus should be valid species, but further investigation looking for eventual synonymy between it and subsequently described species and for the true area of distribution is needed. D i s t r i b u t i o n.? Adzaria (KRYZHANOVSKIJ et al. 1995: 102).	en	Guéorguiev, B. (2010): The " colchicus " group of Pterostichus (Coleoptera, Carabidae). Linzer biologische Beiträge 42 (1): 659-669, DOI: http://doi.org/10.5281/zenodo.5332860
03E887AB59583155FF3D6CADFCCEEB23.taxon	description	T y p e m a t e r i a l 2, 2 syntypes (male lectotype and paralectotypes by present designation), all well preserved, and without attached labels under. Close to the four specimens, the following handwritten label is pinned on the bottom of box: " platydera Chaud. Caucase centr. 8000. i. " (MNHN, " Collection Chaudoir ", box no 216 " Argutor "); 1 syntype (paralectotype by present designation), well preserved, labelled: " 2700 " [handwritten grimy label] / " laticollis Chaud. Caucas … " [handwritten grimy label] / " SYNTYPUS Feronia laticollis Chaudoir 1846 labelled by MNHUB 2005 " [red typeset label] (MNHUB). T y p e l o c a l i t y. " Imérétie, présdeGlola " (CHAUDOIR 1848: 147). O t h e r m a t e r i a l e x a m i n e d. 1 of P. platyderus, labelled: " Coll. Germar-Schaum " [typeset white label] / " platyderus Chaud. Caucas occid. " [handwritten grimy label] (DEI); 1 paratype of P. abishirensis BELOUSOV 1991 (NMNHS). M a l e g e n i t a l i a. Median lobe of aedeagus in lateral view (Figs 6 - 7) with ostium deflected to left; apical part longer than basal part, with ventral margin slightly concave distally and apex somewhat bent downwards; basal part with strongly developed and prominent ventral process; internal sac (non-inverted state) with indistinct chitinised piece. Apex of median lobe dorsally (Fig. 8) straight with tip slightly deflected to left. Left paramere (Figs 9 - 10) sickle-shaped; basal part longer than apical one, forming widened bulb; apical part with pointed tip. The genitalia of the two males from MNHN have been examined. After drawing, the genitalia of the lectotype were placed in Euparal on separate transparent label pinned under the specimen, from whom they were extracted; that of the paralectotype were also put in Euparal, but placed on the same label on which the specimen is glued. The genitalia of the male from DEI are identical with those of the specimens from MNHN while these of the specimen from MNHUB have not been extracted. T a x o n o m i c n o t e s a n d r e l a t i o n s h i p s The presence of pubescence on the distal part of antennomeres 3, missing scutellar pore of the elytra, dense and continuous punctuation on the basal part of pronotum (which is also detectable along the lateral margins in the anterior part of pronotum), absence of medial setiferous punctures on the hind coxae, glabrous ventral side of onychium, and sickle-shaped shape of left paramere demonstrates that this species is member of Oreoplatysma, and do not of Cryobius. The following transfer is proposed: Pterostichus (Oreoplatysma) platyderus (CHAUDOIR 1850), new assignment of Pterostichus (Cryobius) platyderus (CHAUDOIR 1850). This species seems related to the species of the group " abishirensis " (BELOUSOV 1991, KRYZHANOVSKIJ et al. 1995: 103), above all in the missing scutellar pore of elytra and shape of aedeagus. The group in question includes two vicarious species, e. g. P. asinubas DAVIES 2004 (replacement name for P. abasinus BELOUSOV 1991) and P. abishirensis BELOUSOV 1991. P. platyderus differs easily from both species in the longer and rounder right paramere and in the different shape of aedeagus. By the shape of the pronotum, aedeagus and right paramere, P. platyderus resembles more several species of the group " koenigi ", as P. satyrus KURNAKOV 1962, P. krivensis BELOUSOV 1991 and P. rousianus. In the lateral margin of pronotum rather concave towards posterior angles and median lobe of aedeagus (dorsal view) bent to the left, the species is close to P. rousianus, but the former is distinct in aedeagus (lateral view) with concave ventral margin before apex and apex bent downwards. P. platyderus and P. krivensis are very similar in the shape of pronotum and indistinct outer basal impression as well as in the shape of the aedeagus and of the left parameres. However, the former is distinct from the latter in the absence of distinct punctuation on the head, less protruding forward anterior angles of the pronotum, absence of scutellar pore, and somewhat shorter apical part of the left paramere. Based on the original description of P. satyrus (KURNAKOV 1962), we failed to find significant differences between this species and P. platyderus, save temporae gradually slanting backward and their length almost equal to diameter of eyes as well as elytral striae somewhat less punctured in the latter. Most probably P. platyderus occupies an intermediate position between the group " abishirensis " and the couple of species P. satyrus – P. krivensis, but is evidently closer to the last two. P. platyderus should be valid species, but further taxonomic investigations should examine eventual synonymy between it and subsequently described species. D i s t r i b u t i o n. Imeretia (KRYZHANOVSKIJ et al. 1995: 102).	en	Guéorguiev, B. (2010): The " colchicus " group of Pterostichus (Coleoptera, Carabidae). Linzer biologische Beiträge 42 (1): 659-669, DOI: http://doi.org/10.5281/zenodo.5332860
03E887AB595F3155FF3D6EE9FCC9EF11.taxon	description	T y p e l o c a l i t y. " AlpesdelaTouchétie (Caucasecentraletoriental) " (CHAUDOIR 1868: 251). M a l e g e n i t a l i a. Median lobe of aedeagus in lateral view (Fig. 11) with ostium deflected to left, apical part longer than basal part and with straight ventral margin, tip of apex somewhat bent upwards; basal part with strongly developed and prominent ventral process; internal sac (inverted state) with single strongly chitinised piece. Apex of median lobe in dorsal view (Fig. 12) with tip deflected to left. Left paramere (Figs 13 - 14) sickle-shaped, regularly round in middle; basal part forming big widened bulb; apical part with round (not pointed) tip. After drawing, the genitalia were included in Euparal and put on separate transparent label pinned under the lectotype. T a x o n o m i c n o t e s a n d r e l a t i o n s h i p s The presence of pubescence on the distal part of third antennomeres, dense and continuous punctuation on the basal part of pronotum, absence of medial setiferous punctures on hind coxae, glabrous ventral side of onychium, and sickle-shaped left paramere demonstrate that this species belongs to subgenus Oreoplatysma, and not to Cryobius. Therefore, the following transfer is proposed: Pterostichus (Oreoplatysma) stomoides (CHAUDOIR 1868), new assignment of Pterostichus (Cryobius) stomoides (CHAUDOIR 1868). According to the external morphology and structure of the male genitalia, this species seems closer to the taxa from the group " daghestanus ", and especially to P. chefsuricus REITTER 1896 and P. kirschenblatti KRYZHANOVSKIJ 1988. However, the apex of aedeagus distinctly bent to left and somewhat turn upwards and different shape of the left paramere suppose that P. stomoides is distinct from the other two species. P. stomoides should be treated as valid species. D i s t r i b u t i o n. Tushetia (KRYZHANOVSKIJ et al. 1995: 102).	en	Guéorguiev, B. (2010): The " colchicus " group of Pterostichus (Coleoptera, Carabidae). Linzer biologische Beiträge 42 (1): 659-669, DOI: http://doi.org/10.5281/zenodo.5332860
