taxonID	type	description	language	source
E12C8212FFCAFF9F768DFE6CFC17FAD4.taxon	description	(Figs. 1, 3)	en	Short, Andrew Edward Z. (2009): Two new species of the hygropetric genus Oocyclus from eastern India (Coleoptera: Hydrophilidae). Acta Entomologica Musei Nationalis Pragae 49 (2): 625-630, DOI: http://doi.org/10.5281/zenodo.5321656
E12C8212FFCAFF9F768DFE6CFC17FAD4.taxon	materials_examined	Type material. HOLOTYPE: male, ‘ INDIA, Meghalaya State (10) / E Khasi Hills, 11 km SW Cherra- / punjee, Laitkynsew, 21 - 24. iv. / 2008, 25 ° 13 ′ N 91 ° 39 ′ E, 810 m / Fikáček, Podskalská, Šípek lgt. ’ // ‘ seepage: wet rocks algae / blue algae / moss ca. 1.5 - 2 km / via rd. from ‘ Cherrapunjee Holid. / Resort’ in direct. Cherrapunjee, / exposed’ (NMPC). PARATYPES: 16 spec., same data as holotype (KSEM, NHMW, NMPC, USNM).	en	Short, Andrew Edward Z. (2009): Two new species of the hygropetric genus Oocyclus from eastern India (Coleoptera: Hydrophilidae). Acta Entomologica Musei Nationalis Pragae 49 (2): 625-630, DOI: http://doi.org/10.5281/zenodo.5321656
E12C8212FFCAFF9F768DFE6CFC17FAD4.taxon	diagnosis	Diagnosis. Posterolateral corners of pronotum angulate. Procoxae distinctly spinose. Raised medial glabrous region of the metaventrite extended along its entire length. Abdominal ventrites dark brown to black; finely and evenly pubescent. Elytra with detectable but often obscured rows of systematic punctures. This species shares many diagnostic characters with O. melinoventris Short & Swanson, 2005 (currently only known from Thailand), and the extended glabrous region of the metaventrite is currently known only from these two species (see SHORT & SWANSON 2005, Fig. 11). However, they are easily distinguished by the color of the ventrites, which is bright yellow in O. melinoventris while nearly black in O. armstrongorum sp. nov. Oocyclus melinoventris is also smaller in size, never exceeding 5.0 mm, while the smallest individuals of O. armstrongorum sp. nov. are 5.2 mm.	en	Short, Andrew Edward Z. (2009): Two new species of the hygropetric genus Oocyclus from eastern India (Coleoptera: Hydrophilidae). Acta Entomologica Musei Nationalis Pragae 49 (2): 625-630, DOI: http://doi.org/10.5281/zenodo.5321656
E12C8212FFCAFF9F768DFE6CFC17FAD4.taxon	description	Description. Size and form. Total body length = 5.2 – 5.6 mm. Broadly oval, moderately convex. Elytra slightly longer than wide. Color. Dorsum black, without any distinct colored iridescence. Maxillary palps yellow except apex of the apical segment, which is slightly darkened. Labial palps yellow. Mentum and stipes dark brown to black, the same color as ventral surface of head. Pseudoepipleura and tarsi reddish brown, with the remainder of the sternum and legs very dark brown to black. Ventrites uniformly dark brown, similar in color as the sternum. Head. General punctation on labrum, clypeus and frons variable in size from almost very fine to moderately coarse; dense, distance between punctures 0.5 – 1.0 × the width of one puncture. Systematic row of punctures on labrum composed of coarse, setiferous punctures laterally with center row appearing absent or with punctures reduced in size or density. Frons with an irregular row of systematic punctures mesad of each eye, bearing fine recumbent to erect setae. Clypeal systematic punctures not apparent, blending with the surrounding course ground punctation. Antennae with scape subequal to slightly longer in length to segments 2 – 5; first two segments of club subequal in length and apical segment slightly shorter than two preceding segments combined. Maxillary palps short, as long to slightly longer in length than width of labrum; apical segment ca. 1.5 × as long as penultimate. Labial palps three-fourths as long as width of mentum. Mentum quadrate, anterior margin slightly convex; bearing distinct scattered punctures on anterior two-thirds, some of which bear fine setae. Thorax. General punctation on elytra very dense, distance between punctures absent to 1.0 × the width of one puncture, composed of punctures of varying size, from very fine to very coarse; becoming extremely dense posteriorly, making elytra appear almost rugose. Pronotal punctation similar but distinctly less impressed. Pronotal systematic punctures distinct, 1.5 – 2.0 × the size of larger general punctation punctures and set with a fine recumbent seta; anterior and posterior series each forming an irregular row. Lateral margins of pronotum with a sparse row of setiferous punctures. Posterolateral corners of pronotum angulate, but not spinose. Sutural punctation on elytra unmodified from general punctation. Elytra with five diffuse rows of systematic punctures, which are mostly obscured by the larger ground punctures, but generally detectable. Prosternum with fine median carina along entire length, without a tooth or long setae anteriorly. Process of the mesoventrite with lateral extensions sloping evenly downward; apex set with a few fine setae. Mesoventrite with glabrous area extending to entire length, slightly narrower than width of process of the mesoventrite anteriorly, becoming evenly narrowed anteriorly. Procoxae covered with short spines and coarse long setae; mesocoxae without spines, at most with a few coarse, long setae. Protibiae with 9 – 11 spines on dorsal face. Protarsal segments 1 – 4 small, subequal in length; apical segment ca. as long as segments 1 – 4 combined. Abdomen. Ventrites with moderately dense pubescence, the longest setae shorter to subequal in length to longest setae around mesosternal glabrous area. Fifth ventrite entire, with pubescence slightly denser than ventrites 1 – 4. Aedeagus (Fig. 3) with parameres evenly curved along outer margin, slightly curving inwards apically; inner margins bisinuate. Apex of median lobe nearly subequal with apex of parameres.	en	Short, Andrew Edward Z. (2009): Two new species of the hygropetric genus Oocyclus from eastern India (Coleoptera: Hydrophilidae). Acta Entomologica Musei Nationalis Pragae 49 (2): 625-630, DOI: http://doi.org/10.5281/zenodo.5321656
E12C8212FFCAFF9F768DFE6CFC17FAD4.taxon	etymology	Etymology. It is my pleasure to name this species after my friends Dr. Andrew Armstrong and Ms. Christina Lamneck in honor of their recent marriage.	en	Short, Andrew Edward Z. (2009): Two new species of the hygropetric genus Oocyclus from eastern India (Coleoptera: Hydrophilidae). Acta Entomologica Musei Nationalis Pragae 49 (2): 625-630, DOI: http://doi.org/10.5281/zenodo.5321656
E12C8212FFCAFF9F768DFE6CFC17FAD4.taxon	biology_ecology	Biology. As is typical for nearly all members of the genus, the examined specimens of O. armstrongorum sp. nov. were collected on a rock seepage habitat that was covered in algae. A detailed account of that locality is provided by FIKÁČEK & ŠÍPKOVÁ (2009) and GENTILI & FIKÁČEK (2009).	en	Short, Andrew Edward Z. (2009): Two new species of the hygropetric genus Oocyclus from eastern India (Coleoptera: Hydrophilidae). Acta Entomologica Musei Nationalis Pragae 49 (2): 625-630, DOI: http://doi.org/10.5281/zenodo.5321656
E12C8212FFCAFF9F768DFE6CFC17FAD4.taxon	distribution	Distribution. Known only from the type locality in Meghalaya State of India.	en	Short, Andrew Edward Z. (2009): Two new species of the hygropetric genus Oocyclus from eastern India (Coleoptera: Hydrophilidae). Acta Entomologica Musei Nationalis Pragae 49 (2): 625-630, DOI: http://doi.org/10.5281/zenodo.5321656
E12C8212FFC8FF9976D5FA92FC08FD36.taxon	description	(Figs. 2, 4)	en	Short, Andrew Edward Z. (2009): Two new species of the hygropetric genus Oocyclus from eastern India (Coleoptera: Hydrophilidae). Acta Entomologica Musei Nationalis Pragae 49 (2): 625-630, DOI: http://doi.org/10.5281/zenodo.5321656
E12C8212FFC8FF9976D5FA92FC08FD36.taxon	materials_examined	Type material. HOLOTYPE: male, ‘ INDIA, Meghalaya State (10) / E Khasi Hills, 11 km SW Cherra- / punjee, Laitkynsew, 21 - 24. iv. / 2008, 25 ° 13 ’ N 91 ° 39 ’ E, 810 m / Fikáček, Podskalská, Šípek lgt. ’ // ‘ seepage: wet rocks algae / blue algae / moss ca. 1.5 - 2 km / via rd. from ‘ Cherrapunjee Holid. / Resort’ in direct. Cherrapunjee, / exposed’ (NMPC). PARATYPES: 24 spec., same data as holotype (KSEM, NHMW, NMPC, USNM).	en	Short, Andrew Edward Z. (2009): Two new species of the hygropetric genus Oocyclus from eastern India (Coleoptera: Hydrophilidae). Acta Entomologica Musei Nationalis Pragae 49 (2): 625-630, DOI: http://doi.org/10.5281/zenodo.5321656
E12C8212FFC8FF9976D5FA92FC08FD36.taxon	diagnosis	Diagnosis. Posterolateral corners of pronotum evenly rounded. Procoxae without distinct spines. Abdominal ventrites finely and evenly pubescent. Elytra without detectable rows of systematic punctures. This species is externally very similar to O. bhutanicus SATÔ, 1979 but differs in several subtle characters: although the dorsal punctation on the elytra of both species consists of punctures of mixed sizes, in O. madidus sp. nov. the smallest punctures are much finer and smaller than the smallest punctures seen on O. bhutanticus. The largest punctures on both species are relatively similar in size. Consequently, the size range of the punctures observed in O. madidus sp. nov. is more extensive. In addition, the metafemora in O. madidus sp. nov. are slightly broader (ca. 2.1 × as long as wide, compared to ca. 2.3 × for O. bhutanicus). Additionally, the anteromedial tooth of the prosternal carina, while still small in O. madidus sp. nov., is larger than in the holotype of O. bhutanicus. Unfortunately, the aedeagus of the holotype of O. bhutanicus has been missing for more than 25 years (see SHORT & SWANSON 2005 for discussion) and so I was not able to compare these structures directly; based on the figure by SATÔ (1979: Fig. 16), the inner margin of the parameres are sinuate in that species (straight in O. madidus). SATÔ’ s (1979) drawing is not symmetrical, with the left paramere indicating a small apical constriction and the right evenly rounded; thus, I cannot determine which is correct for comparison with O. madidus sp. nov., which is closer to the former condition.	en	Short, Andrew Edward Z. (2009): Two new species of the hygropetric genus Oocyclus from eastern India (Coleoptera: Hydrophilidae). Acta Entomologica Musei Nationalis Pragae 49 (2): 625-630, DOI: http://doi.org/10.5281/zenodo.5321656
E12C8212FFC8FF9976D5FA92FC08FD36.taxon	description	Description. Size and form. Total body length = 3.7 – 4.2 mm. Oval, moderately convex. Elytra longer than wide. Color. Dorsum black; head, pronotum, and elytra with distinct greenish sheen. Maxillary and labial palps yellow; apex of maxillary palps dark brown. Mentum and stipes light to reddish brown, distinctly paler than ventral face of head. Legs, epipleura, lateral margins of prosternum, and posterior margin of each ventrite yellow to light brown, with remainder of venter slightly to moderately darker reddish brown. Head. General punctation on labrum, clypeus and frons slightly variable in size from very fine to moderately fine; densely distributed, distance between punctures 1.0 – 1.5 × the width of one puncture. Systematic row of punctures on labrum moderately dense, forming a nearly continuous lateral row set with moderately long setae, but each puncture still fairly distinguishable, not forming a groove. Frons with an irregular row of systematic punctures mesad of each eye, bearing fine erect setae. Clypeus with a few very indistinct systematic punctures along anterolateral margins, slightly larger than surrounding punctation and bearing short setae. Antennae with scape subequal in length to segments 2 – 5; first two segments of club subequal in length and apical segment subequal in length to the two preceding segments combined. Maxillary palps short, about as long as width of labrum; segment 2 slightly bulbous, apical segment slightly longer than penultimate. Labial palps one-half to three-fourths as long as width of mentum. Mentum quadrate, anterior margin slightly convex, bearing very fine and scattered punctures. Thorax. General punctation on pronotum and elytra composed of extremely fine and coarse punctures, evenly mixed and distributed. Pronotal systematic punctures present, ca. 1.5 × the size of general punctation and set with a fine seta, sometimes partially blending with larger general punctures; anterior and posterior series each forming an irregular row. Lateral margins of pronotum set with a few sparsely distributed setiferous punctures; with punctures very small as to almost appear absent. Posterolateral corners of pronotum evenly rounded. Sutural punctation on elytra absent or unmodified from general punctation. Elytra without distinct rows of larger punctures, as they blend uniformly with general punctation; some of these rows (of systematic punctures) can be detected by the presence of fine, short setae. Prosternum with median carina along entire length, with small anteromedial tooth; without long spines or hairs anteriorly. Process of the mesoventrite with lateral extensions sloping evenly downward; apex set with a few long fine setae. Metaventrite with small oval glabrous area posteromedially, slightly longer than wide, length of glabrous area about half the total length of metaventrite. Pro- and mesocoxae densely pubescent; without distinct spines. Protibiae with 10 spines on dorsal face. Protarsal segments 1 – 4 small, subequal in length; apical segment ca. as long as segments 1 – 4 combined. Abdomen. Ventrites 1 – 5 with uniform pubescence, longest setae about as long as the setae around the metasternal glabrous area. Fifth ventrite entire. Aedeagus with parameres slightly attenuated apically; inner margins straight. Median lobe (Fig. 4) not reaching the apex of the parameres.	en	Short, Andrew Edward Z. (2009): Two new species of the hygropetric genus Oocyclus from eastern India (Coleoptera: Hydrophilidae). Acta Entomologica Musei Nationalis Pragae 49 (2): 625-630, DOI: http://doi.org/10.5281/zenodo.5321656
E12C8212FFC8FF9976D5FA92FC08FD36.taxon	etymology	Etymology. Madidus, Latin, meaning soaked or drenched, referring to the wet habitats of the species.	en	Short, Andrew Edward Z. (2009): Two new species of the hygropetric genus Oocyclus from eastern India (Coleoptera: Hydrophilidae). Acta Entomologica Musei Nationalis Pragae 49 (2): 625-630, DOI: http://doi.org/10.5281/zenodo.5321656
E12C8212FFC8FF9976D5FA92FC08FD36.taxon	biology_ecology	Biology. Collected with and presumed identical to O. armstrongorum sp. nov. (see above).	en	Short, Andrew Edward Z. (2009): Two new species of the hygropetric genus Oocyclus from eastern India (Coleoptera: Hydrophilidae). Acta Entomologica Musei Nationalis Pragae 49 (2): 625-630, DOI: http://doi.org/10.5281/zenodo.5321656
E12C8212FFC8FF9976D5FA92FC08FD36.taxon	distribution	Distribution. Known only from the type locality in Meghalaya state of India.	en	Short, Andrew Edward Z. (2009): Two new species of the hygropetric genus Oocyclus from eastern India (Coleoptera: Hydrophilidae). Acta Entomologica Musei Nationalis Pragae 49 (2): 625-630, DOI: http://doi.org/10.5281/zenodo.5321656
