taxonID	type	description	language	source
5C28AA26060CFFFDFF7DFC89018382EE.taxon	type_taxon	Type species: Hurleyella cumberlandensis new species. Habitus (Fig. 1).	en	Runyon, Justin B., Robinson, Harold (2010): Hurleyella, a new genus of Nearctic Dolichopodidae (Diptera). Zootaxa 2400 (1): 57-65, DOI: 10.11646/zootaxa.2400.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2400.1.6
5C28AA26060CFFFDFF7DFC89018382EE.taxon	diagnosis	Diagnosis. The combination of the following character states will diagnose Hurleyella in the Nearctic. Minute size, body length approximately 1.0 mm. Vertex of head not excavate. Antennal scape without dorsal setae; pedicel without medioapical thumb-like projection. Eyes with short hairs between facets. Wing with costa continuous to tip of unbranched vein M; vein R 2 + 3 short, fusing with costa about half way between termination of R 1 and R 4 + 5; veins R 4 + 5 and M 1 converging near wing apex, then nearly parallel at apex; crossvein dm-cu shorter than last part of CuA 1. Proepisternum with at most a few, small hairs. Femora without preapical setae. In Nearctic keys (Robinson 1964 a; Robinson & Vockeroth 1981), Hurleyella runs to Xanthochlorus Loew, but is easily distinguished by its minute size, dark body color, apical arista, and short wing vein R 2 + 3. In the Neotropics, Hurleyella is only likely to be confused with Microcyrtura, Microchrysotus, or Micromedetera, which also have the short wing vein R 2 + 3. Microcyrtura is distinguished from Hurleyella by the nearly straight preabdomen bearing a sharply reflexed gentital capsule in the male and by the very long arista; Microchrysotus has ornamented palpi and fore tarsi and a banded wing. The wing venation and curved abdomen of Hurleyella are very much like Micromedetera, but the hypopygium of Hurleyella is symmetrical and lacks the distinct projections asymmetrically from one side found in Micromedetera. On the basis of its wing venation, Hurleyella keys in the Manual of Central American Diptera (Bickel 2009) to Micromedetera with which it is considered to be most closely related.	en	Runyon, Justin B., Robinson, Harold (2010): Hurleyella, a new genus of Nearctic Dolichopodidae (Diptera). Zootaxa 2400 (1): 57-65, DOI: 10.11646/zootaxa.2400.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2400.1.6
5C28AA26060CFFFDFF7DFC89018382EE.taxon	description	Description. Male: Head: face pollinose, with dorsal half broadly triangular, ventral half narrowed with sides nearly parallel to slightly broadened at palpus; frontoclypeal suture distinct (Fig. 2 A). Dorsal postcranium strongly concave (Fig. 1). Eyes (Figs. 2 A, C) with short hairs between facets; ommatidia enlarged near face. Vertical setae not on elevation or tubercle; ocellar tubercle not prominent; postocellar setae very short, hair-like; postocular setae very short, sparse (Fig. 2 A). Ventral postcranial hairs (beard) and gena absent. Palpus (Fig. 2 A) small, not ornamented. Proboscis sclerotized, shining, not or slightly enlarged. Labellum with 6 sclerotized pseudotracheae, but not geminately so. Antenna (Figs. 2 A, B) short, scape without dorsal setae; pedicel with apical ring of setulae; arista apical, about as long as height of face. Thorax: scutum with posterior third distinctly flattened. 4 or 5 dorsocentral setae; acrostichal setae absent; 1 notopleural seta; 1 scutellar seta per side, no additional hairs; 1 humeral seta. Legs: not ornamented, without major setae. Coxa II without lateral seta. Coxa III with minute lateral hair. Femora without preapical setae. Wing (Fig. 3): vein R 2 + 3 short, fusing with costa about half way between termination of R 1 and R 4 + 5. Basal section of M 1 curving posteriorly to crossvein dm-cu, remainder nearly straight, directed slightly anteriorly. Vein R 4 + 5 curving toward, then nearly parallel to M 1 just before wing apex. Vein M without evident bosse alaire, the slight flexion and wing indentation typically on vein M distal to crossvein dm-cu that is present in many dolichopodids. Vein A 1 absent. Calypter without evident setae. Abdomen: cylindrical, gradually but only slightly tapering (Figs. 1, 4 A). Hypopygium (Figs. 1, 4 – 5) bulbous, borne on a broad peduncle formed by abdominal segment 7; abdominal tergite 7 setose; epandrium deeply emarginate dorsally, with basal lobes bearing setae; abdominal sternite 8 with setae; hypopygial foramen left basal; hypandrium fused to base of epandrium, divided into 2 asymmetrical lobes; surstylus large, complex and only partially attached to epandrium, with many lobes, several blade-like setae, and pair of long lateroapical membranous flanges; phallus simple, abruptly arched and projecting ventrally. Female: Similar to male. Abdominal terminalia with approximately 6 acanthophorite spines.	en	Runyon, Justin B., Robinson, Harold (2010): Hurleyella, a new genus of Nearctic Dolichopodidae (Diptera). Zootaxa 2400 (1): 57-65, DOI: 10.11646/zootaxa.2400.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2400.1.6
5C28AA26060CFFFDFF7DFC89018382EE.taxon	etymology	Etymology: Hurleyella is named in honor of the late dipterist Richard Hurley (1934 – 2008). The Latin diminutive - ella denotes the small body size of this genus.	en	Runyon, Justin B., Robinson, Harold (2010): Hurleyella, a new genus of Nearctic Dolichopodidae (Diptera). Zootaxa 2400 (1): 57-65, DOI: 10.11646/zootaxa.2400.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2400.1.6
5C28AA26060CFFFDFF7DFC89018382EE.taxon	discussion	Remarks: Because only two species are known, the division of generic and specific characters is rather arbitrary, but we have attempted to include the usual characters known to be of value for other dolichopodid genera, or those characters anticipated to be constant in Hurleyella.	en	Runyon, Justin B., Robinson, Harold (2010): Hurleyella, a new genus of Nearctic Dolichopodidae (Diptera). Zootaxa 2400 (1): 57-65, DOI: 10.11646/zootaxa.2400.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2400.1.6
5C28AA26060EFFFAFF7DFDB6008B85D5.taxon	description	Description. Male: Body length approximately 1.0 mm; wing approximately 1.0 mm x 0.4 mm. Habitus (Fig. 1). Head: dark metallic blue-green ground color nearly to wholly concealed with ochre to gray pollen. Face with ventral half very narrow (narrowest width less than width of first flagellomere), with nearly parallel sides; covered with moderately dense to dense gray to gray-ochre pollen; frons covered with dense ochre pollen. Dorsal postcranium with sparse to moderately dense ochre pollen. Vertical setae small, approximately equal to ocellar setae (subequal in length to first flagellomere); postocular setae very short, sparse, white (Fig. 2 A). Palpus gray pollinose, small, ovate, with a few black setae (Fig. 2 A). Proboscis dark yellow-brown, sclerotized, shining, not enlarged; with short white hairs at apex. Antenna (Figs. 2 A, B) wholly black; first flagellomere subtriangular with round apex, a little longer than wide, with rather long hairs; arista apical, about as long as height of face. Thorax: scutum dark metallic blue-green, covered with dense ochre pollen. Setae of thorax rather short, black; 1 presutural supra-alar seta; 2 postsutural supra-alar setae; 1 postalar seta; proepisternum with 1 small white hair. Pleura metallic blue with sparse to moderately dense gray-brown pollen. Legs: without major setae. Coxae concolorous with pleura. Coxa I with very sparse, short, white anterior hairs; 1 – 2 longer white hairs at apex (Fig. 2 A). Coxa III with minute, white lateral hair. Femora mostly dark brown; without preapical setae. Tibia I, II, and III dark yellow. Tarsi dark brown from apex of basal tarsomere. Ratios of tibia: tarsomeres for leg I: 9 – 5 – 3 – 2 – 2 – 2; for leg II: 12 – 7 – 3 – 2 – 2 – 2; for leg III: 12 – 4 – 3 – 2 – 2 – 2. Wing (Fig. 3): hyaline, usually with brownish clouding at very base, especially in costal cell. Vein R 2 + 3 short, fusing with costa about midway between termination of R 1 and R 4 + 5, still ending closer to termination of R 1. Halter knob yellow to yellow-brown; stem brown. Calypter very small, brown, without evident setae. Abdomen: with very short, rather sparse, stiff black setae (Fig. 4 A); dark metallic blue-green covered with moderately dense to dense ochre-brown pollen. Abdominal sternite 8 with setae, rather small and mostly hidden under tergite 7 (Fig. 4 B). Hypopygium (Figs. 4 - 5). Epandrium rather ovate with dorsoapical margin smooth, strongly sclerotized, projecting; with basal, long, narrow epandrial lobe bearing long seta near base and long seta at apex. Hypandrium with 2 broad, asymmetrical lobes that are ventroapically directed. Phallus abruptly arched and pointed ventrally between hypandrial lobes. Cercus roughly oval-shaped, covered with small hairs and several longer setae, especially along margin. Surstylus loosely attached to epandrium (at least ventrally), large, with many lobes, 2 large blade-like setae directed medially; with large, membranous, rugose, tapering, distolateral flange directed dorsomedially. Female: Similar to male.	en	Runyon, Justin B., Robinson, Harold (2010): Hurleyella, a new genus of Nearctic Dolichopodidae (Diptera). Zootaxa 2400 (1): 57-65, DOI: 10.11646/zootaxa.2400.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2400.1.6
5C28AA26060EFFFAFF7DFDB6008B85D5.taxon	etymology	Etymology: Specimens were collected in Cumberland Mountains physiographic region of southwestern Virginia.	en	Runyon, Justin B., Robinson, Harold (2010): Hurleyella, a new genus of Nearctic Dolichopodidae (Diptera). Zootaxa 2400 (1): 57-65, DOI: 10.11646/zootaxa.2400.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2400.1.6
5C28AA26060EFFFAFF7DFDB6008B85D5.taxon	materials_examined	Holotype: ♂, VIRGINIA: Wise Co., Phillips Creek, 3 mi. SW of Pound, 442 m (1450 ft), N 37 º 06.54 ’ W 82 º 40.25 ’, VI – 2 – 2008, J. B. Runyon (USNM). Paratypes: 6 ♂, 2 ♀, same data as holotype; 2 ♂, VIRGINIA: Buchanan Co., Hunts Creek, 1 mi. SW of Breaks, 320 m (1050 ft), N 37 º 18.37 ’ W 82 º 17.72 ’, VI – 12 – 2008, J. B. Runyon (USNM, CNC, MTEC). Two male paratypes from Phillips Creek were permanently slide mounted.	en	Runyon, Justin B., Robinson, Harold (2010): Hurleyella, a new genus of Nearctic Dolichopodidae (Diptera). Zootaxa 2400 (1): 57-65, DOI: 10.11646/zootaxa.2400.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2400.1.6
5C28AA26060EFFFAFF7DFDB6008B85D5.taxon	discussion	Remarks: The specimens at Phillips Creek were collected by sweeping open, dry, sandy soil with some very sparse vegetation at edge of an artificial beach where Phillips Creek flows into North Fork of Pound Lake. Those from Hunts Creek were collected near junction with Garden Creek from open, dry, bare areas of horizontal shale over which the creek flows. The two collection sites are in extreme southwest Virginia, near the Kentucky border, and about 40 km apart.	en	Runyon, Justin B., Robinson, Harold (2010): Hurleyella, a new genus of Nearctic Dolichopodidae (Diptera). Zootaxa 2400 (1): 57-65, DOI: 10.11646/zootaxa.2400.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2400.1.6
5C28AA26060BFFF8FF7DFF44008B81AD.taxon	description	Description: Very similar to H. cumberlandensis, showing only the following differences: Head: face with ventral half not as narrow (narrowest width approximately equal to width of first flagellomere), and more obviously widened toward palpus. Ochre pollen on frons and dorsal postcranium not as dense. Legs: coxae concolorous with pleura, but more yellowish in some specimens. Ratios of tibia: tarsomeres for leg I: 10 – 5 – 3 – 2 – 2 – 2; for leg II: 14 – 8 – 4 – 3 – 2 – 2; for leg III: 14 – 5 – 4 – 3 – 2 – 2. Abdomen: epandrial lobe with seta at apex only about one-third as long as seta near base (in H. cumberlandensis apical seta at least one-half length of basal seta). Surstylus (Fig. 5 C) with 3 large blade-like setae directed medially (H. cumberlandensis has 2 large setae - the third seta is very small); the flange of surstylus is very broad and rounded apically (Fig. 5 C). Female: Similar to male.	en	Runyon, Justin B., Robinson, Harold (2010): Hurleyella, a new genus of Nearctic Dolichopodidae (Diptera). Zootaxa 2400 (1): 57-65, DOI: 10.11646/zootaxa.2400.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2400.1.6
5C28AA26060BFFF8FF7DFF44008B81AD.taxon	etymology	Etymology: Named in appreciation of the dipterist Scott Brooks, who recognized and sent us the specimens of this species.	en	Runyon, Justin B., Robinson, Harold (2010): Hurleyella, a new genus of Nearctic Dolichopodidae (Diptera). Zootaxa 2400 (1): 57-65, DOI: 10.11646/zootaxa.2400.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2400.1.6
5C28AA26060BFFF8FF7DFF44008B81AD.taxon	materials_examined	Holotype: ♂, TEXAS: Brazos Co., College Station, Lick Creek Park, bottomland forest near creek, malaise trap, 28 – 30 March 2000, M. Buck (DEBU). Paratypes: 1 ♂, same data as holotype except 26 – 28 March 2000; 1 ♂, same data as holotype except 5 – 9 April 2000; 1 ♀, same data as holotype except “ post oak savanna by creek ”, 26 – 28 March 2000 (CNC, DEBU).	en	Runyon, Justin B., Robinson, Harold (2010): Hurleyella, a new genus of Nearctic Dolichopodidae (Diptera). Zootaxa 2400 (1): 57-65, DOI: 10.11646/zootaxa.2400.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2400.1.6
5C28AA26060BFFF8FF7DFF44008B81AD.taxon	discussion	Remarks: Hurleyella brooksi is very similar to H. cumberlandensis, but differs most noticeably in having a wider face and in the form of the surstylus.	en	Runyon, Justin B., Robinson, Harold (2010): Hurleyella, a new genus of Nearctic Dolichopodidae (Diptera). Zootaxa 2400 (1): 57-65, DOI: 10.11646/zootaxa.2400.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2400.1.6
