taxonID	type	description	language	source
133A87A08148CF5BFF56FD44042EF80E.taxon	materials_examined	Type species: Cryptococcus ulmi Tang & Hao, 1995.	en	Nan, Nan, Deng, Jun, Wu, San’An (2013): A new felt scale genus Macroporicoccus gen. n. (Hemiptera: Coccoidea: Eriococcidae) from China, with a redescription of Macroporicoccus ulmi (Tang & Hao) comb. n. Zootaxa 3722 (2): 170-182, DOI: http://dx.doi.org/10.11646/zootaxa.3722.2.3
133A87A08148CF5BFF56FD44042EF80E.taxon	description	Description. Adult female: Body in life subglobular, orange-red, covered by a white ovisac. On slide, body circular or ovate. Antennae reduced, 6 - segmented. Mouthparts developed, labium 3 - segmented. Thoracic spiracles each with a group of disc pores, each with 3 or 5 loculi, near opening. Legs much reduced, only distal part of tarsus and complete claw visible; each claw with a denticle, hind legs each located on a pore plate. Anal ring circular or oval, with 6 short setae and no pores. Anal lobes indistinct. Macrotubular ducts with a cup-shaped invagination, present on both sides of body except ventral middle area. Microtubular ducts present on dorsum and margin of venter. Quinquelocular pores mainly present ventrally. Macrodisc pores present on dorsum and margin of venter. Setae small, sparsely distributed on venter. First-instar nymph: In life, body bright yellow. Body on slide oval. Antennae 6 - segmented. Eyes present. Mouthparts and legs well developed. Anal ring circular, with 6 setae and without pores. Anal lobes each with one long and one short flagellate seta. Macrotubular and microtubular ducts absent. Quinquelocular disc pores present on ventral abdominal segments and near spiracles. Macrodisc pores distributed on dorsal surface. Setae present, forming transverse or longitudinal rows on body surface.	en	Nan, Nan, Deng, Jun, Wu, San’An (2013): A new felt scale genus Macroporicoccus gen. n. (Hemiptera: Coccoidea: Eriococcidae) from China, with a redescription of Macroporicoccus ulmi (Tang & Hao) comb. n. Zootaxa 3722 (2): 170-182, DOI: http://dx.doi.org/10.11646/zootaxa.3722.2.3
133A87A08148CF5BFF56FD44042EF80E.taxon	discussion	Remarks. Adult females of Macroporicoccus Nan & Wu gen. n. can be easily distinguished from other eriococcid genera due to the presence of macrodisc pores, which occur throughout the dorsum and ventral margin of C. ulmi. Tang and Hao (1995) described this type of pore as a macroduct, but they noticed also that it was a rather large disc pore; Wu (2000) referred to it as a macrodisc pore and described it as a kind of large invaginated pore. As this pore is markedly larger than the simple pores in other Coccoidea (usually 2 – 3 µm in diameter), we consider the term macrodisc pore to be most appropriate. However, after SEM study, we can confirm that it is a simple closed pore (Foldi, 1997), about 10 µm in diameter, with a narrow rim and a weakly convex surface (Fig. 20, 21); it secretes long solid wax filaments, which make up the outline of the ovisac (Fig. 3) (Wu, unpublished data). Cryptococcus, Kuwanina Cockerell and the new genus Macroporicoccus live in crevices in the trunks and branches, and have many similar morphological characters. The shared features of the adult female are: (1) body subglobular covered by a white ovisac; (2) antennae reduced; (3) legs absent or reduced to tubercles, hind legs always replaced by pore-plates or vestigial flaps; and (4) anal lobes absent (Williams, 1985; Tang & Hao, 1995; Wu, 2000; Henderson, 2007; Wu & Liu, 2009). However, adult female Kuwanina apparently differ from those of other two genera in the absence of macrotubular ducts (C. nudata Brittin is an exception) and in the presence of unique invaginated 5 - locular disc pores (Williams, 1985; Henderson, 2007). As indicated above, adult female Macroporicoccus differ from Cryptococcus in the presence of macrodisc pores. In addition, Macroporicoccus has 6 - segmented antennae whereas Cryptococcus has 1 – 4 - segmented antennae (Kosztarab & Hale, 1968; Tang & Hao, 1995).	en	Nan, Nan, Deng, Jun, Wu, San’An (2013): A new felt scale genus Macroporicoccus gen. n. (Hemiptera: Coccoidea: Eriococcidae) from China, with a redescription of Macroporicoccus ulmi (Tang & Hao) comb. n. Zootaxa 3722 (2): 170-182, DOI: http://dx.doi.org/10.11646/zootaxa.3722.2.3
133A87A08148CF5BFF56FD44042EF80E.taxon	etymology	Etymology. The new genus name means those scale insects with macrodisc pores.	en	Nan, Nan, Deng, Jun, Wu, San’An (2013): A new felt scale genus Macroporicoccus gen. n. (Hemiptera: Coccoidea: Eriococcidae) from China, with a redescription of Macroporicoccus ulmi (Tang & Hao) comb. n. Zootaxa 3722 (2): 170-182, DOI: http://dx.doi.org/10.11646/zootaxa.3722.2.3
133A87A0814FCF5FFF56FF36029AF81C.taxon	description	(Figs 3 – 22)	en	Nan, Nan, Deng, Jun, Wu, San’An (2013): A new felt scale genus Macroporicoccus gen. n. (Hemiptera: Coccoidea: Eriococcidae) from China, with a redescription of Macroporicoccus ulmi (Tang & Hao) comb. n. Zootaxa 3722 (2): 170-182, DOI: http://dx.doi.org/10.11646/zootaxa.3722.2.3
133A87A0814FCF5FFF56FF36029AF81C.taxon	materials_examined	Material examined. 12 ad ♀♀, China, Shanxi Province, Taigu County, campus of Shanxi Agricultural University in crevices in the trunks of Ulmus pumila, 10 December 1997, Coll. San-an Wu (the measurements and illustration are based on this slide); 4 ad ♀♀, China, Tianjin City on U. pumila, 15 June 1999, Coll. San-an Wu; 7 ad ♀♀, China, Beijing City, Songshan Mountain in the crevices of trunks of U. japonica, 29 May 2012, Coll. Nan Nan.	en	Nan, Nan, Deng, Jun, Wu, San’An (2013): A new felt scale genus Macroporicoccus gen. n. (Hemiptera: Coccoidea: Eriococcidae) from China, with a redescription of Macroporicoccus ulmi (Tang & Hao) comb. n. Zootaxa 3722 (2): 170-182, DOI: http://dx.doi.org/10.11646/zootaxa.3722.2.3
133A87A0814FCF5FFF56FF36029AF81C.taxon	description	Redescription of adult female. Body in life sub-globular, orange-red, covered by white waxy ovisac. Body on slide circular or ovate, 0.90 – 1.45 mm long and 0.70 – 1.33 mm wide. Antennae (Fig. 7) reduced, 6 - segmented, length of each segment (µm): I 14 – 18, II 8 – 11, III 6 – 7, IV 4 – 6, V 5 – 6, VI 12 – 16; segment II with one campaniform sensillum, segment IV and V each bearing a fleshy seta, segment VI with 3 fleshy and 4 long flagellate setae and a pair of coeloconic sensilla at apex. Mouthparts developed, clypeolabral shield (Fig. 8) with 2 small setae in middle of surface; labium (Fig. 9) 3 - segmented, 56 – 62 µm long, with 5 pairs of short setae on apical segment. Thoracic spiracles each 34 – 40 µm long and atrium 19 – 21 µm wide, each with a group of quinquelocular (rarely trilocular) (Fig. 12) pores near opening, with 6 – 21 near each anterior spiracle (Fig. 10) and 6 – 23 near each posterior spiracle (Fig. 11). Legs (Figs 13 – 15) much reduced, only distal part of tarsus and complete claw visible; each claw with a denticle near apex; fore and middle claw 9 – 10 µm long, hind claw stouter, 6 – 8 µm long; each leg with one pair of tarsal digitules and one pair of claw digitules, all slightly knobbed, tarsal digitule longer than claw, claw digitule nearly equal to length of claw; each leg with several setae around base; hind legs each located on a 51 – 70 µm long and 42 – 50 µm wide pore plate (Fig. 16). Anal ring (Fig. 17) circular to oval, 26 – 28 µm long and 18 – 26 µm wide, with 6 short setae, each 17 – 20 µm long, and without pores. Anal lobes absent, each represented by a long flagellate seta, 48 – 70 µm long at posterior end of body. Vulva (Fig. 18) distinct, located on ventromedial area between abdominal segments VI and VII. Dorsum (Fig. 5). Setae absent. Macrotubular ducts (Fig. 19) with a cup-shaped invagination, each 20 – 26 µm long and about 3 µm wide, with an inner ductule 17 – 19 µm long; numerous, distributed over all surface. Microtubular ducts each with an outer ductule about 2 µm long and a dermal orifice about 2 µm wide; sparse on dorsum. Macrodisc pores (Figs 20, 21) with rim slightly protruding, each about 10 µm in diameter, forming transverse rows across dorsum. Quinquelocular disc pores, each about 5 µm in diameter; few on margin. Venter (Fig. 6). Setae (Fig. 22) spine- or hair-like, 4 – 16 µm long, sparsely distributed on venter but with a small group around each leg. Macroducts, microducts and macrodisc pores similar to those on dorsum, present on marginal and submarginal areas. Quinquelocular pores also similar to those on dorsum, but with a group near each spiracle and mouthparts; also distributed on abdominal venter and in marginal and submarginal areas, sometimes with a few near spiracles replaced by trilocular pores.	en	Nan, Nan, Deng, Jun, Wu, San’An (2013): A new felt scale genus Macroporicoccus gen. n. (Hemiptera: Coccoidea: Eriococcidae) from China, with a redescription of Macroporicoccus ulmi (Tang & Hao) comb. n. Zootaxa 3722 (2): 170-182, DOI: http://dx.doi.org/10.11646/zootaxa.3722.2.3
133A87A0814FCF5FFF56FF36029AF81C.taxon	biology_ecology	Host plants. Ulmus japonica, U. pumila, U. pumila ‘ Tenue’ (Ulmaceae), Syringa oblata (Oleaceae).	en	Nan, Nan, Deng, Jun, Wu, San’An (2013): A new felt scale genus Macroporicoccus gen. n. (Hemiptera: Coccoidea: Eriococcidae) from China, with a redescription of Macroporicoccus ulmi (Tang & Hao) comb. n. Zootaxa 3722 (2): 170-182, DOI: http://dx.doi.org/10.11646/zootaxa.3722.2.3
133A87A0814FCF5FFF56FF36029AF81C.taxon	distribution	Distribution. China (Beijing, Shanxi, Tianjin).	en	Nan, Nan, Deng, Jun, Wu, San’An (2013): A new felt scale genus Macroporicoccus gen. n. (Hemiptera: Coccoidea: Eriococcidae) from China, with a redescription of Macroporicoccus ulmi (Tang & Hao) comb. n. Zootaxa 3722 (2): 170-182, DOI: http://dx.doi.org/10.11646/zootaxa.3722.2.3
