taxonID	type	description	language	source
A97EBF375E3D5129BE6AB6BFF72F09CB.taxon	description	Figs 12 - 16, 17 - 18	en	Kasparyan, Dmitri R., Khalaim, Andrey I. (2021): New Darwin wasps (Hymenoptera, Ichneumonidae, Tersilochinae, Tryphoninae) from the Eastern Palaearctic region. Journal of Hymenoptera Research 84: 115-126, DOI: http://dx.doi.org/10.3897/jhr.84.68719, URL: http://dx.doi.org/10.3897/jhr.84.68719
A97EBF375E3D5129BE6AB6BFF72F09CB.taxon	description	Description. Female. Fore wing length 6.0 mm. Antenna with 29 flagellomeres; first flagellomere 4.0 times and second flagellomere 2.5 times as long as broad; combined length of flagellomeres 1 and 2 about 0.9 maximum eye diameter. Apical flagellomeres thin, about 0.6 times as wide as basal flagellomeres. Sensillae starting on flagellomere 4 and continuing on subsequent flagellomeres. Head (Figs 13, 14) shining, not narrowed behind eyes, temples slightly convex. Face centrally convex and coarsely punctate. Clypeus weakly and evenly convex, polished, finely and sparsely punctate; clypeal foveae small, without setae. Frons with rather dense, even and moderately fine punctures. Temples with fine and sparse punctures. Margins of antennal sockets on frons not raised. Malar space about half as long as basal mandibular width. Oral carina slightly higher than occipital carina, and 0.6 times as long as basal mandibular width. Thorax smooth, with even, distinct, moderately large and dense punctures. Pronotum with rugosity on transverse impression and lateral lower corner. Mesoscutum without notaulus. Scutellum with lateral carinae on its anterior 0.4. Mesopleuron in upper half and posteriorly smooth, shining, with moderately large punctures which are separated by about 1.5 diameters of puncture (Fig. 15); speculum large and polished. Metapleuron with punctures denser (distance between punctures about 0.5 - 1.0 diameter of puncture); lower margin of metapleuron with deep impression covered with rugae. Propodeum with superficial rugosity, impunctate except for irregular punctures on smooth background on first lateral areas; costulae present, extending from hind part of areola; areola confluent with basal area, its lateral sides parallel. Apical area 0.55 times as high as length of propodeum (measured at midline), with median longitudinal carina distinct. Fore wing with an areolet (Fig. 12); nervulus strongly postfurcal. Hind wing with nervellus intercepted at lower 0.23 (Fig. 15). Fore femur thick, 2.2 times as long as wide. Fore and mid tarsi shortened. Fore leg with first tarsomere 2.7 times as long as wide, tarsomeres 2 - 4 transverse. Mid tarsus with first tarsomere 3.0 times as long as wide and 1.4 times shorter than tarsomeres 2 - 5 combined. Hind femur 3.5 times as long as wide; length ratio of tarsomeres 1 - 5 of hind tarsus 4.8: 2.5: 1.7: 1.2: 2.2. Tarsal claws apically strongly curved and tapered to a sharp point; claws with relatively short and dense teeth (Fig. 16); fore tarsus with 6 teeth, mid and hind tarsi with 7 - 9 teeth. Metasomal tergites smooth, with dense and moderately large punctures; punctures denser on tergites 1 - 4 (Fig. 17), somewhat finer and sparser on posterior margins of tergites, on posterior half of tergite 4, and on tergite 5 (subsequent tergites retracted under tergite 5, see Fig. 18). First tergite about 0.9 times as long as wide, its longitudinal median dorsal carinae strong and extending to superficial transverse depression in posterior 0.2 of tergite; interspace between dorsal carinae broad, entirely densely punctate from transverse depression to base; transverse depression with short longitudinal rugae. Second and third tergites strongly transverse, about 0.5 times as long as broad; both tergites with shallow transverse depression just behind middle. Hypopygium large, with large oval desclerotized area medially in basal 0.7 (this area somewhat paler than its light yellow apical and lateral periphery); hypopygium with rather dense, erected and moderately long setae which are shorter than diameters of hind tarsomeres and extending to hind margin of hypopygium (Fig. 18). Ovipositor sheath 2.5 times as long as its median width, with dorsoapical emargination in its apical 0.4 (Fig. 18). Coloration (Fig. 12). Antenna blackish brown, scape and pedicel light yellow ventrally and flagellomeres 1 - 3 brownish yellow ventrally. Head black; face, clypeus, lower 0.3 of frontal orbits, palpi, mandible (except for reddish brown teeth), malar space, lower parts of gena and postgena yellow (Fig. 13). Mesosoma black; lower margin of propleuron (above fore coxa), tegula and plates at base of fore wing yellow; lower part of mesopleuron with obscure reddish marking ventrally (Fig. 15). Legs predominantly yellowish red. Fore and mid legs with coxae and trochanters entirely light yellow, and femora, tibiae and tarsi reddish yellow. Hind leg with coxa, trochanters and femur red; femur slightly darkened dorsally at apical 0.1; tibia brownish yellow in basal 0.6 and dark brown in apical 0.4; spurs yellow; hind tarsus brown, tarsomeres 1 - 3 pale at extreme base. Pterostigma dark brown with proximal 0.25 whitish. First metasomal tergite black except for yellowish red hind margin; second tergite reddish with two subbasal blackish spots (Fig. 17); subsequent tergites black except for tergites 3 and 4 which are brownish black in posterior 1 / 3 and with reddish tinge on hind margins (Figs 17, 18). Epipleura and sternites yellow, segments 2 - 4 each with brownish large sclerotized spot. Ovipositor sheath dark brown, its extreme apex and basal 0.6 latero-ventrally reddish yellow. Male. Unknown.	en	Kasparyan, Dmitri R., Khalaim, Andrey I. (2021): New Darwin wasps (Hymenoptera, Ichneumonidae, Tersilochinae, Tryphoninae) from the Eastern Palaearctic region. Journal of Hymenoptera Research 84: 115-126, DOI: http://dx.doi.org/10.3897/jhr.84.68719, URL: http://dx.doi.org/10.3897/jhr.84.68719
A97EBF375E3D5129BE6AB6BFF72F09CB.taxon	etymology	Etymology. We are happy to name this new species in honour of Prof. Arkady Lelej, after his 75 th birthday and in recognition of his enormous contribution to the taxonomy of Aculeata and his hard work organizing extensive insect studies in the Russian Far East.	en	Kasparyan, Dmitri R., Khalaim, Andrey I. (2021): New Darwin wasps (Hymenoptera, Ichneumonidae, Tersilochinae, Tryphoninae) from the Eastern Palaearctic region. Journal of Hymenoptera Research 84: 115-126, DOI: http://dx.doi.org/10.3897/jhr.84.68719, URL: http://dx.doi.org/10.3897/jhr.84.68719
A97EBF375E3D5129BE6AB6BFF72F09CB.taxon	materials_examined	Material examined. Holotype female (ZISP), Russia, Transbaikal Territory, railway station Karymskaya, ravine in right bank of Ingoda river, taiga (Pinus, Larix), 28. VI. 1975, coll. D. R. Kasparyan.	en	Kasparyan, Dmitri R., Khalaim, Andrey I. (2021): New Darwin wasps (Hymenoptera, Ichneumonidae, Tersilochinae, Tryphoninae) from the Eastern Palaearctic region. Journal of Hymenoptera Research 84: 115-126, DOI: http://dx.doi.org/10.3897/jhr.84.68719, URL: http://dx.doi.org/10.3897/jhr.84.68719
BCC4A660A2A951D0A314430B04C24CC7.taxon	description	Figs 19, 20	en	Kasparyan, Dmitri R., Khalaim, Andrey I. (2021): New Darwin wasps (Hymenoptera, Ichneumonidae, Tersilochinae, Tryphoninae) from the Eastern Palaearctic region. Journal of Hymenoptera Research 84: 115-126, DOI: http://dx.doi.org/10.3897/jhr.84.68719, URL: http://dx.doi.org/10.3897/jhr.84.68719
BCC4A660A2A951D0A314430B04C24CC7.taxon	description	Description. Male. Fore wing length 4.7 - 6.0 mm. Antenna with 28 - 30 flagellomeres, about ten basal flagellomeres thicker than subsequent ones. Head smooth, shining. Face weakly convex in profile, distinctly and rather densely punctate. Frons with distinct moderately dense punctures. Vertex and temple finely and sparsely punctate. Clypeus convex in upper 0.3, flat in lower part, almost impunctate. Malar space and oral carina behind mandible about 0.5 times as long as basal mandibular width. Pronotum laterally with large polished area, with subparallel striation near collar and at lower corner. Epomia weak. Mesopleuron finely and sparsely punctate, with large polished speculum and almost impunctate below subtegular ridge and next to mesopleural fovea along mesepimeron. Mesoscutum with uniform small and dense punctures. Notaulus very weak or absent. Scutellum with sparse moderately large punctures and distinct lateral carina on its anterior 0.4 - 0.5. Tarsal claw of fore leg with 4 (or 5) small teeth in basal 0.8; claw of mid leg with 3 small teeth in basal half; claw of hind leg with 3 teeth in basal 0.4. Metasoma shining, impunctate, tergites 1 and 2 partly with longitudinal striation in posterior half, especially on postmedian transverse impressions; posterior half of tergite 3 and subsequent tergites smooth, with very fine punctures and pubescence. Coloration. Antenna light brownish, scape and pedicel yellowish white ventrally, two or three basal flagellomeres brownish yellow ventrally. Head black; face, clypeus, malar space, palps, mandible (except for teeth) and part of gena from level of lower 0.3 of eye to mandible whitish. Mesosoma black; small spot in hind corner of pronotum whitish, tegula and plates at base of fore wing white. Fore and mid legs and hind coxa light reddish yellow, all trochanters yellow. Hind femur light brown, pale reddish in basal 0.2; hind tibia dull yellowish with apical 0.2 light brown. Metasomal tergites black, tergites 2 and 3 reddish, and tergite 4 in one specimen with small reddish marking at base. Sternites brown with yellow median longitudinal fold on sternites 1 - 5 (or 6).	en	Kasparyan, Dmitri R., Khalaim, Andrey I. (2021): New Darwin wasps (Hymenoptera, Ichneumonidae, Tersilochinae, Tryphoninae) from the Eastern Palaearctic region. Journal of Hymenoptera Research 84: 115-126, DOI: http://dx.doi.org/10.3897/jhr.84.68719, URL: http://dx.doi.org/10.3897/jhr.84.68719
BCC4A660A2A951D0A314430B04C24CC7.taxon	materials_examined	Material examined. 1 female, 2 males (ZISP), Russia, 24 km S of Khabarovsk, Khekhzir Ridge, Levaya River, 11. IV. 1983, coll. Kasparyan.	en	Kasparyan, Dmitri R., Khalaim, Andrey I. (2021): New Darwin wasps (Hymenoptera, Ichneumonidae, Tersilochinae, Tryphoninae) from the Eastern Palaearctic region. Journal of Hymenoptera Research 84: 115-126, DOI: http://dx.doi.org/10.3897/jhr.84.68719, URL: http://dx.doi.org/10.3897/jhr.84.68719
BCC4A660A2A951D0A314430B04C24CC7.taxon	distribution	Distribution. Norway, Germany, Russia (south of Far East).	en	Kasparyan, Dmitri R., Khalaim, Andrey I. (2021): New Darwin wasps (Hymenoptera, Ichneumonidae, Tersilochinae, Tryphoninae) from the Eastern Palaearctic region. Journal of Hymenoptera Research 84: 115-126, DOI: http://dx.doi.org/10.3897/jhr.84.68719, URL: http://dx.doi.org/10.3897/jhr.84.68719
631055174F785212A9B979718711B29E.taxon	description	Figs 1 - 6, 7 - 11	en	Kasparyan, Dmitri R., Khalaim, Andrey I. (2021): New Darwin wasps (Hymenoptera, Ichneumonidae, Tersilochinae, Tryphoninae) from the Eastern Palaearctic region. Journal of Hymenoptera Research 84: 115-126, DOI: http://dx.doi.org/10.3897/jhr.84.68719, URL: http://dx.doi.org/10.3897/jhr.84.68719
631055174F785212A9B979718711B29E.taxon	description	Description. Female. Body length 3.2 mm. Fore wing length 2.2 mm. Head with gena distinctly and roundly constricted posterior to eyes (Fig. 5); gena in dorsal view 0.45 times as long as eye width. Clypeus unusually small, 2.1 times as broad as long and 0.6 times as broad as face (shortest distance between eye margins), smooth, with very fine and sparse punctures in upper half, separated from face by sharp furrow (Fig. 3). Mandible strongly constricted in basal 0.7; width of mandible at level of teeth about 0.4 times width at base; lower tooth almost 1.5 times longer than the upper (Fig. 4). Malar space about 1.1 times as long as basal mandibular width. Flagellum with 16 - 17 flagellomeres, filiform (Fig. 2); second flagellomere almost twice, flagellomeres 3 - 5 about 1.8 times, and subapical flagellomeres 1.2 times as long as broad; flagellomeres 4 to 6 (or more) bearing very small subapical finger-shaped structures on outer surface (these structures hardly discernible with a light microscope). Face with median swelling. Face, frons and vertex finely granulate, impunctate, dull. Gena shallowly granulate, impunctate, weakly shining. Occipital carina complete, evenly convex mediodorsally. Hypostomal carina absent (at least in lower part). Mesosoma almost entirely granulate, impunctate and dull, except for mesopleuron, which is widely smooth and shining centrally above foveate groove (Fig. 6). Notaulus with irregular wrinkles (Fig. 5). Scutellum with lateral longitudinal carinae developed on anterior 0.3 - 0.4. Upper end of epicnemial carina curved abruptly forward to anterior margin of mesopleuron (Fig. 4). Foveate groove of mesopleuron long, oblique, almost straight, deep and moderately broad, with distinct transverse wrinkles, extending over anterior 0.6 of mesopleuron, almost reaching epicnemial carina anteriorly (Fig. 6). Propodeum with narrow to moderately broad basal area which is 2.5 - 3.5 times as long as broad and about half as long as apical area (Fig. 8). Propodeal spiracle separated from pleural carina by about 2.0 diameters of spiracle. Apical area slightly impressed along midline, narrowly rounded anteriorly (Fig. 8); apical longitudinal carinae complete and reaching transverse carina anteriorly (Fig. 8). Fore wing (Fig. 7) with second recurrent vein (2 m-cu) postfurcal. First abscissa of radius (Rs + 2 r) straight, somewhat longer than width of pterostigma. First and second sections of radius (Rs + 2 r and Rs) meeting at slightly acute angle. Intercubitus (2 rs-m) slightly thickened, about as long as abscissa of cubitus between intercubitus and second recurrent vein (abscissa of M between 2 rs-m and 2 m-cu). Metacarpus (R 1) short, not reaching tip of the wing. Second abscissa of postnervulus (Cu & 2 cu-a) present, thus brachial cell is closed posteriorly. Hind wing with nervellus (cu 1 & cu-a) weakly reclivous. Legs slender, tarsal claws not pectinate. First tergite about 3.4 times as long as posteriorly broad, more or less trapeziform in central cross-section; lateral sides of petiole distinctly longitudinally striate before glymma (Fig. 10), petiole dorsally and postpetiole smooth; glymma deep, round, situated at posterior 0.55 of tergite and joining by thin and sharp furrow with ventral part of postpetiole (Fig. 10); in dorsal view, petiole with lateral margins subparallel, postpetiole distinctly widened anteriorly and almost twice broader than petiole (Fig. 9); upper margin of first tergite, in lateral view, nearly straight in anterior 0.7 and distinctly arcuate in posterior 0.3 (Fig. 10). Second tergite 1.5 times as long as anteriorly broad (Fig. 9). Thyridial depression deep and long, 2.5 - 3.0 times as long as broad, with posterior end rounded (Fig. 9). Ovipositor slender, weakly upcurved, with weak dorsal subapical depression (Fig. 11); sheath twice as long as first tergite (Fig. 1). Head and mesosoma dark reddish brown (Fig. 6); lower 0.7 of clypeus, mouthparts and mandibles (teeth reddish black) yellow to brownish yellow. Antenna with scape and pedicel yellow ventrally and brownish dorsally; flagellum brown with basal flagellomeres yellowish brown. Tegula yellow-brown. Pterostigma brown. Legs brownish yellow; hind coxa slightly darkened with brown. First metasomal tergite dark brown. Metasoma posterior to first tergite predominantly brown, yellowish to pale brown ventrally (Fig. 1). Male. Unknown.	en	Kasparyan, Dmitri R., Khalaim, Andrey I. (2021): New Darwin wasps (Hymenoptera, Ichneumonidae, Tersilochinae, Tryphoninae) from the Eastern Palaearctic region. Journal of Hymenoptera Research 84: 115-126, DOI: http://dx.doi.org/10.3897/jhr.84.68719, URL: http://dx.doi.org/10.3897/jhr.84.68719
631055174F785212A9B979718711B29E.taxon	etymology	Etymology. The species is named in honour of the Russian entomologist, an expert in Mutillidae, Prof. Arkady S. Lelej (Vladivostok, Russia).	en	Kasparyan, Dmitri R., Khalaim, Andrey I. (2021): New Darwin wasps (Hymenoptera, Ichneumonidae, Tersilochinae, Tryphoninae) from the Eastern Palaearctic region. Journal of Hymenoptera Research 84: 115-126, DOI: http://dx.doi.org/10.3897/jhr.84.68719, URL: http://dx.doi.org/10.3897/jhr.84.68719
631055174F785212A9B979718711B29E.taxon	materials_examined	Material examined. Holotype female (EUM), Japan, Hokkaido I., Sapporo, Hitsujigaoka, 43 ° 00 ' 27 " N, 141 ° 24 ' 53 " E, Malaise trap, 29. VI- 6. VII. 2010, coll. K. Konishi. Paratypes. Japan: 1 female (EUM), same data as holotype, but 6 - 13. VII. 2010. 1 female (ZISP), same data, but 27. VI- 4. VII. 2011. 1 female (ZISP), same data, but 4 - 11. VII. 2011. 1 female (EUM), same data, but 11 - 18. VII. 2011.	en	Kasparyan, Dmitri R., Khalaim, Andrey I. (2021): New Darwin wasps (Hymenoptera, Ichneumonidae, Tersilochinae, Tryphoninae) from the Eastern Palaearctic region. Journal of Hymenoptera Research 84: 115-126, DOI: http://dx.doi.org/10.3897/jhr.84.68719, URL: http://dx.doi.org/10.3897/jhr.84.68719
631055174F785212A9B979718711B29E.taxon	distribution	Distribution. Japan (Hokkaido I.).	en	Kasparyan, Dmitri R., Khalaim, Andrey I. (2021): New Darwin wasps (Hymenoptera, Ichneumonidae, Tersilochinae, Tryphoninae) from the Eastern Palaearctic region. Journal of Hymenoptera Research 84: 115-126, DOI: http://dx.doi.org/10.3897/jhr.84.68719, URL: http://dx.doi.org/10.3897/jhr.84.68719
