taxonID	type	description	language	source
C7BE87A15935594A9F4F197B961E885C.taxon	description	Figures 1, 2, 13, 19, 25, 26, 37, 38, 49 - 51, 67, 73	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
C7BE87A15935594A9F4F197B961E885C.taxon	diagnosis	Diagnosis. Male. Clypeus dorsally elevated, sloping ventrally. Mandible with sharp subapical and large inner subbasal teeth (Fig. 13), ventrally deeply excised with large basal tooth (Fig. 19). Scape ventral carinae subparallel, mostly smooth between them (Fig. 25); F 1 apical 1 / 3 and F 2 basal 1 / 2 yellow, F 2 apical 1 / 2 and F 3 - F 11 with ventral reddish brown patch; F 1 cylindrical (Fig. 26). T 3 orange (Fig. 2); cuspis strongly widened on posterior 1 / 3 with rounded apex (Figs 49, 67); digitus posteriorly slender stick-like (Fig. 37); paracuspis stick-like (Fig. 67); penis valve weakly expanded ventrally (Figs 50, 51). Female. Unknown.	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
C7BE87A15935594A9F4F197B961E885C.taxon	materials_examined	Material examined. 24 ♂. Malaysia: Negeri Sembilan: Pasoh Forest Reserve, 25. III- 2. IV. 1993, K. Konishi & K. Maeto leg., coarse Malaise trap, plot 2 [2 ♂ FRIM, NARO], 2 - 8. IV. 1993 [1 ♂ FRIM], 8 - 15. IV. 1993 [1 ♂ NARO]; Pasoh Forest Reserve, 25. III- 2. IV. 1993, K. Konishi & K. Maeto leg., fine Malaise trap, plot 2 [2 ♂ FRIM, NARO]; Pasoh Forest Reserve, 26. III- 3. IV. 1993, K. Konishi & K. Maeto leg., coarse Malaise trap, plot 4 [1 ♂ FRIM], 9 - 16. IV. 1993 [1 ♂ NARO]; Pasoh Forest Reserve, 3 - 9. IV. 1993, K. Konishi & K. Maeto leg., fine Malaise trap, plot 4 [1 ♂ FRIM]; Pasoh Forest Reserve, 26. XI- 3. XII. 1993, K. Maeto leg., coarse Malaise trap, plot 1 [2 ♂ FRIM, NARO]; Pasoh Forest Reserve, 26. XI- 3. XII. 1993, K. Maeto leg., coarse Malaise trap, plot 4 [2 ♂ FRIM, NARO]; Pasoh Forest Reserve, 26. XI- 3. XII. 1993, K. Maeto leg., fine Malaise trap, plot 1 [2 ♂ FRIM, NARO], 10 - 17. XII. 1993 [1 ♂ FRIM]; Pasoh Forest Reserve, 26. XI- 3. XII. 1993, K. Maeto leg., fine Malaise trap, plot 3 [1 ♂ NARO], 10 - 17. XII. 1993 [1 ♂ FRIM]; Pasoh Forest Reserve, 26. XI- 3. XII. 1993, K. Maeto leg., fine Malaise trap, plot 4 [1 ♂ NARO], 3 - 10. XII. 1993 [1 ♂ NARO]; Pasoh Forest Reserve, 3 - 10. XII. 1993, K. Maeto leg., coarse Malaise trap, plot 6 [1 ♂ NARO]; Pasoh Forest Reserve, 10 - 17. XII. 1993, K. Maeto leg., coarse Malaise trap, plot 2 [1 ♂ FRIM]; Pasoh Forest Reserve, 10 - 17. XII. 1993, K. Maeto leg., fine Malaise trap, plot 2 [2 ♂ FRIM, NARO].	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
C7BE87A15935594A9F4F197B961E885C.taxon	distribution	Distribution. Indonesia: Kalimantan Timur (Lelej 1996 b). Malaysia: Negeri Sembilan *, Sarawak (Cameron 1902; Mickel 1937).	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
08E714DB16BD5B91AF5EBE4F2C2E111A.taxon	description	Figures 3, 4, 14, 20, 27, 28, 39, 40, 52 - 54, 68, 74	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
08E714DB16BD5B91AF5EBE4F2C2E111A.taxon	diagnosis	Diagnosis. Male. Clypeus dorsally elevated, sloping ventrally. Mandible with sharp subapical and large inner subbasal teeth (Fig. 14), ventrally deeply excised with large basal tooth (Fig. 20). Scape ventral carinae divergent apically (Fig. 27), densely punctate between them; F 1 apical 1 / 3, F 2 - F 3, F 2 - F 4, or F 2 - F 5 whitish yellow on ventral half, F 5 or F 5 - F 6 with ventral whitish yellow patch, F 6 - F 11 or F 7 - F 11 with ventral dark brown patch; F 1 strongly depressed, F 1 height ~ 0.8 x pedicel diameter (Fig. 28). T 3 orange (Fig. 4); cuspis weakly widened on posterior 1 / 3 with rounded apex (Figs 52, 68); digitus posteriorly slender stick-like (Fig. 39); paracuspis absent (Fig. 68); penis valve weakly expanded ventrally (Figs 53, 54). Female. Unknown.	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
08E714DB16BD5B91AF5EBE4F2C2E111A.taxon	materials_examined	Material examined. 3 ♂. Malaysia: Negeri Sembilan: Pasoh Forest Reserve, 31. III. 1993, K. Konishi leg. [1 ♂ NARO]; Pasoh Forest Reserve, 26. XI- 3. XII. 1993, K. Maeto leg., fine Malaise trap, plot 6 [1 ♂ FRIM]; Pahang: Cameron Highland, 19 mile, 15. III. 2006, K. Takasuka leg. [1 ♂ EUM].	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
08E714DB16BD5B91AF5EBE4F2C2E111A.taxon	distribution	Distribution. Malaysia: Negeri Sembilan *, Pahang *, Sabah, Sarawak (Mickel 1935, 1937; Lelej 1996 b).	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
2F91B07729FE5D91ABB3599BFDB59905.taxon	description	Figures 5, 6, 15, 21, 29, 30, 41, 42, 55 - 57, 69, 75	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
2F91B07729FE5D91ABB3599BFDB59905.taxon	diagnosis	Diagnosis. Male. Clypeus dorsally elevated, sloping ventrally. Mandible with weak subapical and subbasal teeth (Fig. 15), ventrally deeply excised with large basal tooth (Fig. 21). Scape ventral carinae divergent apically (Fig. 29), densely punctate between them; F 1 apical 1 / 3, F 2, F 3 basal half whitish yellow on ventral half, F 5 - F 11 with ventral yellow to reddish brown patch; F 1 strongly depressed, F 1 height ~ 0.8 x pedicel diameter (Fig. 30). T 3 orange (Fig. 6); cuspis weakly widened on posterior 1 / 3 with rounded apex (Figs 55, 69); digitus posteriorly expanded, forming triangular projection (Fig. 41); paracuspis slightly tuberculate (Fig. 69); penis valve weakly expanded ventrally (Figs 56, 57). Female. Unknown.	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
2F91B07729FE5D91ABB3599BFDB59905.taxon	description	Description. Male. Body length: 11.2 - 15.0 mm (holotype: 15.0 mm; mean: 13.1 mm); forewing length: 8.09 - 10.7 mm (holotype: 10.7 mm; mean: 9.37 mm). Color and setae. Frons, vertex, gena, clypeus, pedicel, flagellum mostly, prementum, stipes, mesosoma, T 4 - T 7, S 4 - S 8 black; mandible brown to black, with apex always black and medial 1 / 5 - 1 / 2 yellowish brown; labrum, protibial spur yellow; scape, tegula brownish black; F 1 apical 1 / 3, F 2, F 3 basal half whitish yellow on ventral half; F 5 - F 11 yellow to reddish brown ventrally; trochanters brown to dark brown; coxae, femora, tibiae, tarsi dark brown to brownish black; meso- and metatibial spurs pale yellow; T 1 - T 3, S 1 - S 3 orange; wings pale brown on proximal 1 / 3, brown on distal 2 / 3, with veins dark brown. Upper frons, vertex, trochanters, T 1, S 1 with sparse long erect pale golden setae; lower frons, dorsal propodeal face, dorsal half of lateral propodeal face with dense short appressed and sparse long erect pale golden setae; eye dorsal margin with long erect black setae; gena, pronotal dorsum, metascutellum with sparse short recumbent and sparse long erect pale golden setae; clypeus with dense long recumbent pale golden setae; mandible, tibiae, tarsi, T 2 - T 5 with sparse long recumbent pale golden setae; scape, femora, S 2 with sparse long recumbent and sparse long erect pale golden setae; pedicel, F 1, tegula anterior portion with sparse short recumbent pale golden setae; F 2 - F 11 with sparse erect pale golden microsetae; pronotal collar anteriorly, T 2 disc, T 3 - T 5 medially, S 3 - S 6 with sparse short erect pale golden setae; pronotal collar posteriorly glabrous; pronotal lateral face meso- and metacoxae with sparse long appressed pale golden setae; propleuron with sparse long appressed and sparse long erect pale golden setae; mesoscutum with sparse short recumbent black and sparse short erect pale golden to black setae; tegula inner margin with sparse short recumbent pale golden and sparse short erect pale golden to black setae; mesoscutellum with sparse short recumbent pale golden, sparse long erect black, sparse long erect pale golden setae; anterior and posterior parts of mesopleuron, metapleuron, ventral half of lateral propodeal face with sparse short appressed pale golden setae; axilla, metanotal trough, medial and ventral parts of mesopleuron, procoxa with dense long appressed and sparse long erect pale golden setae; T 6 - T 7, S 7 - S 8 with sparse long recumbent and sparse long erect black setae; T 2 with lateral golden felt line; T 1 - T 5, S 2 - S 5 with posterior fringe of sparse pale golden setae; S 6 - S 8 with posterior fringe of sparse black setae. Structure. Head 1.82 - 2.00 x as wide as long (holotype: 2.00); lateral margins strongly convergent behind eye; vertex rounded posteriorly; POD / OOD = 0.56 - 0.58 (holotype: 0.56); OD: MOD: LOD = 1.00: 1.08 - 1.12: 0.94 - 1.00 (holotype: 1.00: 1.12: 0.94); ocellar region slightly convex, ocelli without posterior carina; antennal scrobe with weak inverted V-shaped dorsal carina not reaching inner eye margin; genal carina absent; postgenal bridge flattened; hypostomal carina weakly developed; clypeus dorsally elevated, sloping ventrally, with subventral transverse groove laterally not defined by carina; anterior clypeal margin projecting medially; mandible robust, narrowed to apex with inner margin expanded forming small subbasal tooth; mandible apex bidentate; mandible ventral margin excised into large basal tooth; mandible dorsal carina sharp; prementum flattened; scape curved medially, with two longitudinal ventral carinae divergent apically; F 1 depressed, F 1 height 0.67 - 0.80 x pedicel diameter (holotype: 0.67); relative length of pedicel, F 1 and F 2 = 1.00: 2.85 - 3.06: 1.92 - 2.29 (holotype: 1.00: 3.06: 2.29); F 2 - F 11 almost equal in width; F 11 as long as F 2, conical. Mesosoma widest at mesonotum including tegulae; head width: humeral width: pronotal width: mesonotal width = 88: 62: 91: 100; humeral angle rounded, with sharp humeral carina on ventral half; mesoscutum strongly convex; mesoscutal width 1.12 x its length; tegula short rounded, posteriorly reaching mesoscuto-scutellar articulation, 1.44 - 1.53 x as long as wide (holotype: 1.53); notaulus evenly wide, reaching anterior 1 / 2 of mesoscutum, subparallel; parapsidal line length 0.25 x mesoscutal length; parascutal carina forming rounded, weakly elevated process on postero-lateral angle of mesoscutum; mesoscutellum convex without medial carina, tubercle or impunctate line; mesopleuron divided into dorsal and ventral convexities by distinct transverse mesopleural groove, ventrally without transverse precoxal ridge, longitudinal impunctate line, or precoxal swelling; dorsal propodeal face without distinct dorsum and sublateral carina, with anterior inverted triangular cell and semicircular sublateral cell; lateral margins of propodeum gradually convergent posteriorly; dorsal and lateral propodeal faces not separated by carina. Wings well developed; distance between origin of RS on vein SC and base of stigmatic cell equal to stigmatic cell length and first abscissa of RS length; cell 2 RS present. Meso- and metacoxae evenly convex; metacoxa with weak complete inner longitudinal carina; protibial spur slender; tibiae without outer spines. T 1 without distinct dorsum; T 1 spiracle weakly tuberculate; T 2 evenly convex, 1.28 x as wide as long; S 1 with straight medial longitudinal carina with its ventral margin slightly elevated posteriorly; S 2 swollen anteriorly; T 7 medially with longitudinal impunctate convex line, posteriorly not forming tubercle; S 6 - S 8 without lateral carina or tubercle; S 7 entirely exposed, 0.75 x as long as S 8; S 8 mostly sclerotized with posterior membranous area evenly convex posteriorly. Paramere down-curved posteriorly, with weak inner protuberance on posterior 1 / 2 of dorso-inner margin; inner margin of paramere dorsally with short erect golden setae; ventral lobe posteriorly projecting, pointed; parapenial lobe large, with posterior margin slightly concave; cuspis long slender, weakly widened on posterior 1 / 3 with rounded apex, slightly surpassing apex of penis valve, with dense long erect setae ventrally and posteriorly; digitus straight, posteriorly expanded, forming triangular projection, with dense erect golden setae posteriorly; paracuspis slightly tuberculate with few erect golden setae; penis valves symmetrical slender, weakly expanded ventrally, with posterior ventral hook, without setae. Frons, vertex, gena, pronotal dorsum, mesoscutum, medial part of mesopleuron with large dense punctures; postgenal bridge, clypeus lateral portion, mandible outer face, scape between ventral carinae, flagellum, pronotal lateral face, propleuron, axilla, metanotal trough, anterior and posterior parts of mesopleuron, metapleuron, ventral half of lateral propodeal face, procoxa, tarsi with minute dense punctures; clypeus medial convexity with large sparse punctures; mandible mostly, pronotal collar posteriorly smooth; scape mostly, trochanters, femora, tibiae, T 1 medially, T 2 disc, T 3 - T 6, S 1 - S 2 with small sparse punctures; pedicel, pronotal collar anteriorly, meso- and metacoxa with minute sparse punctures; mesoscutellum with large confluent punctures; metascutellum with small confluent punctures; dorsal propodeal face, dorsal half of lateral propodeal face reticulate; T 1 - T 2 laterally, T 7, S 3 - S 8 with small dense punctures; tegula mostly microreticulate with few small punctures anteriorly, postero-inner portion with minute sparse punctures.	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
2F91B07729FE5D91ABB3599BFDB59905.taxon	distribution	Distribution. Malaysia: Negeri Sembilan.	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
2F91B07729FE5D91ABB3599BFDB59905.taxon	etymology	Etymology. This new species is named after FRIM, Forest Research Institute Malaysia, which was the Malaysian counterpart of the Japan-Malaysia joint research project in 1993 funded by the Ministry of the Environment of Japan. Treat as a noun in apposition.	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
BBE895FEA6A754F0BA61029FFD1C65EB.taxon	description	Figures 7, 8, 16, 22, 31, 32, 43, 44, 58 - 60, 70, 75	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
BBE895FEA6A754F0BA61029FFD1C65EB.taxon	diagnosis	Diagnosis. Male. Clypeus medially strongly elevated even ventrally. Mandible with sharp subapical and large inner subbasal teeth (Fig. 16), ventrally deeply excised with large basal tooth (Fig. 22). Scape ventral carinae divergent apically (Fig. 31), densely punctate between them; F 1 apical 1 / 3 and F 2 whitish yellow on ventral half; F 1 strongly depressed, F 1 height ~ 0.8 x pedicel diameter (Fig. 32). T 3 orange (Fig. 8); cuspis widened near apex, forming oblique posterior margin (Figs 58, 70); digitus posteriorly expanded, forming triangular projection (Fig. 43); paracuspis slightly tuberculate (Fig. 70); penis valve weakly expanded ventrally (Figs 59, 60). Female. Unknown.	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
BBE895FEA6A754F0BA61029FFD1C65EB.taxon	description	Description. Male. Body length: 12.4 - 15.1 mm (holotype: 12.4 mm; mean: 13.8 mm); forewing length: 10.6 - 11.0 mm (holotype: 10.6 mm; mean: 10.8 mm). Color and setae. Frons, vertex, gena, clypeus, mandible mostly, scape, pedicel, flagellum mostly, prementum, stipes, mesosoma, coxae, femora, tibiae, tarsi, T 4 - T 7, S 1 medial carina, S 4 - S 8 black; mandible medial 1 / 4 reddish brown; F 1 apical 1 / 3, F 2 whitish yellow on ventral half; labrum, tegula brownish black; trochanters, protibial spur dark brown; meso- and metatibial spurs pale yellow; T 1 - T 3, S 1 except medial carina, S 2 - S 3 orange; wings pale brown, with veins dark brown. Upper frons, vertex, T 1, S 1 with sparse long erect pale golden setae; lower frons, pronotal dorsum, dorsal propodeal face, dorsal half of lateral propodeal face with dense short appressed and sparse long erect pale golden setae; gena, femora, tibiae with sparse short recumbent and sparse long erect pale golden setae; eye dorsal margin with long erect black setae; clypeus with dense long recumbent pale golden setae; mandible, trochanters, T 2 lateral margin with sparse long recumbent pale golden setae; scape with sparse long recumbent and sparse long erect pale golden setae; pedicel, F 1 with sparse short recumbent pale golden setae; F 2 - F 11 with sparse erect pale golden microsetae; pronotal collar anteriorly, T 2 disc, T 3 - T 5, S 3 - S 6 with sparse short erect pale golden setae; pronotal collar posteriorly glabrous; pronotal lateral face, coxae with sparse long appressed pale golden setae; propleuron with sparse long appressed and sparse long erect pale golden setae; mesoscutum with sparse short recumbent and sparse short erect goldish black setae; mesoscutellum, metascutellum with sparse short recumbent pale golden and sparse long erect pale golden to blackish golden setae; axilla, metanotal trough with dense long appressed pale golden setae; anterior and posterior parts of mesopleuron, metapleuron, ventral half of lateral propodeal face with sparse short appressed pale golden setae; medial and ventral parts of mesopleuron with dense long appressed and sparse long erect pale golden setae; tegula anterior portion, tarsi with sparse long recumbent pale golden setae; tegula inner margin with sparse short recumbent and sparse short erect golden setae; S 2 with sparse short recumbent and sparse short erect pale golden setae; T 6 - T 7, S 7 - S 8 with sparse short erect black setae; T 2 with lateral golden felt line; T 1 - T 5, S 2 - S 5 with posterior fringe of sparse pale golden setae; S 6 - S 8 with posterior fringe of sparse black setae. Structure. Head 1.89 - 1.92 x as wide as long (holotype: 1.92); lateral margins strongly convergent behind eye; vertex rounded posteriorly; POD / OOD = 0.56 - 0.77 (holotype: 0.77); OD: MOD: LOD = 1.00: 1.15 - 1.58: 0.92 - 1.08 (holotype: 1.00: 1.58: 0.92); ocellar region slightly convex, ocelli without posterior carina; antennal scrobe with weak inverted V-shaped dorsal carina not reaching inner eye margin; genal carina absent; postgenal bridge flattened; hypostomal carina weakly developed; clypeus medially strongly elevated even ventrally, without subventral transverse groove; anterior clypeal margin projecting medially; mandible robust, narrowed to apex with inner margin expanded forming large subbasal tooth; mandible apex bidentate; mandible ventral margin excised into large basal tooth; mandible dorsal carina sharp; prementum flattened; scape curved medially, with two longitudinal ventral carinae divergent apically; F 1 depressed, F 1 height 0.75 x pedicel diameter; relative length of pedicel, F 1 and F 2 = 1.00: 3.00 - 3.05: 1.84 - 2.26 (holotype: 1.00: 3.05: 1.84); F 2 - F 11 almost equal in width; F 11 as long as F 2, conical. Mesosoma widest at mesonotum including tegulae; head width: humeral width: pronotal width: mesonotal width = 87: 60: 93: 100; humeral angle rounded, with sharp humeral carina not reaching pronotal dorsum; mesoscutum strongly convex; mesoscutal width 1.20 x its length; tegula short rounded, posteriorly reaching mesoscuto-scutellar articulation, 1.43 - 1.44 x as long as wide (holotype: 1.44); notaulus evenly wide, reaching anterior 1 / 2 of mesoscutum, subparallel; parapsidal line length 0.25 x mesoscutal length; parascutal carina forming rounded, weakly elevated process on postero-lateral angle of mesoscutum; mesoscutellum convex without medial carina, tubercle or impunctate line; mesopleuron divided into dorsal and ventral convexities by distinct transverse mesopleural groove, ventrally without transverse precoxal ridge, longitudinal impunctate line, or precoxal swelling; dorsal propodeal face without distinct dorsum and sublateral carina, with anterior inverted triangular cell and semicircular sublateral cell; lateral margins of propodeum gradually convergent posteriorly; dorsal and lateral propodeal faces not separated by carina. Wings well developed; distance between origin of RS on vein SC and base of stigmatic cell equal to stigmatic cell length and first abscissa of RS length; cell 2 RS present. Meso- and metacoxae evenly convex; metacoxa with weak complete inner longitudinal carina; protibial spur slender; tibiae without outer spines. T 1 without distinct dorsum; T 1 spiracle weakly tuberculate; T 2 evenly convex, 1.25 x as wide as long; S 1 with straight medial longitudinal carina with its ventral margin convex; S 2 swollen anteriorly; T 7 medially with longitudinal impunctate convex line, posteriorly not forming tubercle; S 6 - S 8 without lateral carina or tubercle; S 7 entirely exposed, 0.75 x as long as S 8; S 8 mostly sclerotized with posterior membranous area evenly convex posteriorly. Paramere down-curved posteriorly, with weak inner protuberance on posterior 1 / 2 of dorso-inner margin; inner margin of paramere dorsally with short erect golden setae; ventral lobe posteriorly projecting, pointed; parapenial lobe large, with posterior margin concave; cuspis long slender, widened near apex, forming oblique posterior margin, apparently surpassing apex of penis valve, with dense long erect setae ventrally and posteriorly; digitus straight, posteriorly expanded, forming triangular projection, with dense erect golden setae posteriorly; paracuspis slightly tuberculate with few erect golden setae; penis valves symmetrical slender, weakly expanded ventrally, with posterior ventral hook, without setae. Frons, vertex, gena, pronotal dorsum, mesoscutum, medial part of mesopleuron with large dense punctures; postgenal bridge, trochanters, femora, tibiae, T 1 medially, T 2 disc, S 2 with small sparse punctures; clypeus lateral portion, mandible outer and dorsal faces, scape between ventral carinae, flagellum, pronotal lateral face, propleuron, axilla, metanotal trough, anterior and posterior parts of mesopleuron, metapleuron, ventral half of lateral propodeal face, coxae, tarsi with minute dense punctures; clypeus medial convexity, scape mostly, pedicel, T 1 - T 2 laterally, T 3 - T 7, S 1, S 3 - S 8 with small dense punctures; mesoscutellum with large confluent punctures; metascutellum with small confluent punctures; dorsal propodeal face, dorsal half of lateral propodeal face reticulate; pronotal collar anteriorly with minute sparse punctures; pronotal collar posteriorly smooth; tegula mostly microreticulate with few small punctures anteriorly, postero-inner portion with minute sparse punctures.	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
BBE895FEA6A754F0BA61029FFD1C65EB.taxon	distribution	Distribution. Indonesia: East Java.	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
BBE895FEA6A754F0BA61029FFD1C65EB.taxon	etymology	Etymology. The specific name Arkaditilla gibba is a Latin noun meaning gibbosity. It refers to the strongly elevated clypeus of this new species.	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
1A048A79BD0E579B8F8A615E811A9FDB.taxon	description	Figures 9, 10, 17, 23, 33, 34, 45, 46, 61 - 63, 71, 75	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
1A048A79BD0E579B8F8A615E811A9FDB.taxon	diagnosis	Diagnosis. Male. Clypeus dorsally elevated, sloping ventrally. Mandible with sharp subapical and large inner subbasal teeth lamellately expanded near apex (Fig. 17), ventrally slightly excised with small basal tooth (Fig. 23). Scape ventral carinae divergent apically (Fig. 33), densely punctate between them; F 1 apical half and F 2 - F 6 whitish yellow on ventral half, F 7 - F 11 with ventral dark brown patch; F 1 weakly depressed, F 1 height subequal to pedicel diameter (Fig. 34). T 3 black (Fig. 10); cuspis weakly widened on posterior 1 / 3 with rounded apex (Figs 61, 71); digitus posteriorly slender stick-like (Fig. 45); paracuspis slightly tuberculate (Fig. 71); penis valve strongly expanded ventrally on posterior half (Figs 62, 63). Female. Unknown.	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
1A048A79BD0E579B8F8A615E811A9FDB.taxon	description	Description. Male. Body length: 15.9 mm; forewing length: 13.0 mm. Color and setae. Frons, vertex, gena, clypeus, mandible mostly, pedicel, flagellum mostly, prementum, stipes, mesosoma, meso- and metacoxae, T 3 - T 7, S 1 medial carina, S 4 - S 8 black; mandible medial 1 / 3 brown; labrum, protibial spur yellow; scape, F 7 - F 11 ventrally, trochanters dark brown; F 1 apical half, F 2 - F 6 whitish yellow on ventral half; tegula, procoxa, femora, tibiae, tarsi, S 3 lateral portion brownish black; meso- and metatibial spurs pale yellow; T 1 - T 2, S 1 except medial carina, S 2 - S 3 mostly orange; wings brown, with veins dark brown. Upper frons, vertex, T 1, S 1 with sparse long erect pale golden setae; lower frons, gena, pronotal dorsum, dorsal propodeal face, dorsal half of lateral propodeal face, coxae with dense short appressed and sparse long erect pale golden setae; eye dorsal margin with long erect black setae; clypeus with dense long recumbent pale golden setae; mandible with sparse short recumbent and sparse long erect pale golden setae; scape with sparse long recumbent and sparse long erect pale golden setae; pedicel, F 1, trochanters, T 2 lateral margin with sparse short recumbent pale golden setae; F 2 - F 11 with sparse erect pale golden microsetae; pronotal collar anteriorly, T 2 disc, T 3 - T 5, S 3 - S 6 with sparse short erect pale golden setae; pronotal collar posteriorly glabrous; pronotal lateral face with sparse long appressed pale golden setae; propleuron with sparse long appressed and sparse long erect pale golden setae; mesoscutum with sparse short recumbent and sparse short erect dark golden setae; meso- and metascutellum, femora with sparse short recumbent pale golden and sparse long erect golden setae; anterior and posterior parts of mesopleuron, metapleuron, ventral half of lateral propodeal face with sparse short appressed pale golden setae; axilla, metanotal trough, medial and ventral parts of mesopleuron with dense long appressed and sparse long erect pale golden setae; tegula anterior portion, tarsi with sparse long recumbent golden setae; tegula inner margin with sparse short recumbent and sparse short erect golden setae; tibiae with sparse short recumbent golden and sparse long erect golden setae; T 6 - T 7, S 7 - S 8 with sparse short erect black setae; S 2 with sparse short recumbent and sparse short erect pale golden setae; T 2 with lateral golden felt line; T 1 - T 5, S 2 - S 5 with posterior fringe of sparse pale golden setae; S 6 - S 8 with posterior fringe of sparse black setae. Structure. Head 2.00 x as wide as long; lateral margins strongly convergent behind eye; vertex rounded posteriorly; POD / OOD = 0.54; OD: MOD: LOD = 1.00: 1.29: 1.00; ocellar region slightly convex, ocelli without posterior carina; antennal scrobe with weak inverted V-shaped dorsal carina not reaching inner eye margin; genal carina absent; postgenal bridge flattened; hypostomal carina weakly developed; clypeus dorsally elevated, sloping ventrally, without subventral transverse groove; anterior clypeal margin projecting medially; mandible robust, narrowed to apex with inner margin expanded forming large subbasal tooth; mandible apex bidentate; mandible ventral margin slightly excised into small basal tooth; mandible dorsal carina sharp; prementum flattened; scape curved medially, with two longitudinal ventral carinae divergent apically; F 1 depressed, F 1 height 0.91 x pedicel diameter; relative length of pedicel, F 1 and F 2 = 1.00: 3.00: 2.37; F 2 - F 11 almost equal in width; F 11 as long as F 2, conical. Mesosoma widest at mesonotum including tegulae; head width: humeral width: pronotal width: mesonotal width = 83: 62: 85: 100; humeral angle rounded, with sharp humeral carina not reaching pronotal dorsum; mesoscutum strongly convex; mesoscutal width 1.11 x its length; tegula short rounded, posteriorly reaching mesoscuto-scutellar articulation, 1.54 x as long as wide; notaulus evenly wide, reaching anterior 1 / 2 of mesoscutum, subparallel; parapsidal line length 0.3 x mesoscutal length; parascutal carina forming rounded, weakly elevated process on postero-lateral angle of mesoscutum; mesoscutellum convex without medial carina, tubercle or impunctate line; mesopleuron divided into dorsal and ventral convexities by distinct transverse mesopleural groove, ventrally with transverse precoxal ridge, without longitudinal impunctate line, or precoxal swelling; dorsal propodeal face without distinct dorsum and sublateral carina, with anterior inverted triangular cell and semicircular sublateral cell; lateral margins of propodeum gradually convergent posteriorly; dorsal and lateral propodeal faces not separated by carina. Wings well developed; distance between origin of RS on vein SC and base of stigmatic cell equal to stigmatic cell length and first abscissa of RS length; cell 2 RS present. Meso- and metacoxae evenly convex; metacoxa with weak complete inner longitudinal carina; protibial spur slender; tibiae without outer spines. T 1 without distinct dorsum; T 1 spiracle weakly tuberculate; T 2 evenly convex, 1.15 x as wide as long; S 1 with straight medial longitudinal carina with its ventral margin convex; S 2 swollen anteriorly; T 7 medially with longitudinal impunctate convex line, posteriorly not forming tubercle; S 6 - S 8 without lateral carina or tubercle; S 7 entirely exposed, 0.75 x as long as S 8; S 8 mostly sclerotized with posterior membranous area evenly convex posteriorly. Paramere down-curved posteriorly, without inner protuberance on dorso-inner margin; inner margin of paramere dorsally with short erect golden setae; ventral lobe posteriorly projecting, pointed; parapenial lobe large, with posterior margin slightly concave; cuspis long slender, weakly widened on posterior 1 / 3 with rounded apex, apparently surpassing apex of penis valve, with dense long erect setae ventrally and posteriorly; digitus straight, posteriorly not expanded, with dense erect golden setae posteriorly; paracuspis slightly tuberculate with few erect golden setae; penis valves symmetrical slender, strongly expanded ventrally on posterior half, with posterior ventral hook, without setae. Frons, vertex, gena, pronotal dorsum, mesoscutum, medial part of mesopleuron with large dense punctures; postgenal bridge, trochanters, femora, tibiae, T 1 medially, T 2 disc with small sparse punctures; clypeus mostly, mandible outer and dorsal faces, scape between ventral carinae, flagellum, pronotal lateral face, propleuron, axilla, metanotal trough, anterior and posterior parts of mesopleuron, metapleuron, ventral half of lateral propodeal face, coxae, tarsi with minute dense punctures; clypeus anterior margin with few large punctures; scape mostly, pedicel, T 1 - T 2 laterally, T 3 - T 7, S 1 - S 8 with small dense punctures; pronotal collar anteriorly with minute sparse punctures; pronotal collar posteriorly smooth; mesoscutellum with large confluent punctures; metascutellum with small confluent punctures; dorsal propodeal face, dorsal half of lateral propodeal face reticulate; tegula mostly microreticulate with few small punctures anteriorly, postero-inner portion with minute sparse punctures.	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
1A048A79BD0E579B8F8A615E811A9FDB.taxon	distribution	Distribution. Indonesia: Kalimantan Timur.	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
1A048A79BD0E579B8F8A615E811A9FDB.taxon	etymology	Etymology. This new species is dedicated to Arkady Lelej as is the genus itself.	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
7603CBDB1C6855F4B10DBD3ED5EBBA9B.taxon	description	Figures 11, 12, 18, 24, 35, 36, 47, 48, 64 - 66, 72, 76	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
7603CBDB1C6855F4B10DBD3ED5EBBA9B.taxon	diagnosis	Diagnosis. Male. Clypeus dorsally elevated, sloping ventrally. Mandible with sharp subapical and large inner subbasal teeth (Fig. 18), ventrally deeply excised with large basal tooth (Fig. 24). Scape ventral carinae divergent apically (Fig. 35), densely punctate between them; F 1 apical 1 / 3, F 2 and F 3 basal 1 / 3 whitish yellow on ventral half; F 1 strongly depressed, F 1 height ~ 0.8 x pedicel diameter (Fig. 36). T 3 orange (Fig. 12); cuspis strongly widened on posterior 1 / 3 with rounded apex (Figs 64, 72); digitus posteriorly expanded, forming triangular projection (Fig. 47); paracuspis slightly tuberculate (Fig. 72); penis valve weakly expanded ventrally (Figs 65, 66). Female. Unknown.	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
7603CBDB1C6855F4B10DBD3ED5EBBA9B.taxon	materials_examined	Material examined. 2 ♂. Indonesia: Bali: Tabanan, Bengkel, Pura Batu Salahan, 08 ° 23 ' 31.8 " S, 115 ° 05 ' E, 682 m alt., 12. VIII. 2006, K. Watanabe leg. [1 ♂ TUA].	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
7603CBDB1C6855F4B10DBD3ED5EBBA9B.taxon	distribution	Distribution. Indonesia: Bali, Java (Zavattari 1914; Mickel 1935; Lelej 1996 b).	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
6A19D29C19245A50A33A7EB56087CE48.taxon	description	Figures 1 - 6, 7 - 12, 13 - 18, 19 - 36, 37 - 48, 49 - 66, 67 - 72, 73 - 76	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
6A19D29C19245A50A33A7EB56087CE48.taxon	diagnosis	Diagnosis. Male. Frons punctate; vertex roundly sloping posteriorly; postgenal bridge flattened; hypostomal carina simple, without denticle; clypeus dorsally elevated; prementum without tubercle. Mandible apically bidentate, ventrally excised into basal tooth (Figs 19 - 24); inner margin expanded, forming subbasal tooth (Figs 13 - 18); dorso-lateral carina sharp, not crooked. Scape ventrally with two longitudinal carinae, densely punctate or mostly smooth with sparse punctures apically between carinae (Figs 25, 27, 29, 31, 33, 35); flagellum with yellow or reddish brown ventral markings; F 1 longer than F 2, depressed (Figs 28, 30, 32, 34, 36) except A. bagrada with F 1 cylindrical (Fig. 26). Mesosoma black; notaulus present, reaching anterior 1 / 3 to 1 / 2 of mesoscutum; parascutal carina forming weakly elevated, rounded tubercle posteriorly; tegula posterior margin slightly protruding over transscutal articulation, not recurved; mesoscutellum evenly convex, puncto-reticulate; mesopleuron divided into dorsal and ventral convexities by transverse groove; propodeum gradually sloping posteriorly, without sublateral longitudinal carina; propodeal dorso-lateral margin without denticles. Pterostigma length subequal to distance between origin of RS on vein SC and proximal base of pterostigma. Protibial spur slender; mesocoxa evenly convex; metacoxa evenly convex with long sparse recumbent setae, without tubercle on inner margin. Metasoma black with segments 1 - 2 or 3 orange; S 2 swollen anteriorly, without lateral felt line (Figs 2, 4, 6, 8, 10, 12); T 7 medially with slender longitudinal smooth convex line; S 6 - S 8 without lateral carina; S 7 entirely exposed, 0.75 x as long as S 8; S 8 mostly sclerotized with posterior membranous area evenly convex posteriorly; left and right basiparameres ventro-medial margins connected to form postero-medial tongue-like tubercle (Figs 38, 40, 42, 46, 48); parapenial lobe spatulate, not flexed; ventral portion of basiparamere strongly projecting posteriorly to connect volsella (Figs 49, 52, 55, 58, 61, 64); paramere antero-ventrally serrate (Figs 49, 52, 55, 58, 61, 64); digitus straight, dorsal margin lamellate (Figs 67 - 72); paracuspis absent, tuberculate or stick-like; penis valves symmetrical, with posterior hook (Figs 50, 51, 53, 54, 56, 57, 59, 60, 62, 63, 65, 66). Female. Unknown.	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
6A19D29C19245A50A33A7EB56087CE48.taxon	distribution	Distribution. Widespread in Sundaland, the Oriental Region (Figs 73 - 76).	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
6A19D29C19245A50A33A7EB56087CE48.taxon	etymology	Etymology. I am delighted to name this new genus in honor of Arkady Stepanovich Lelej, a fellow mutillidologist, pioneer of Old World mutillid systematics, and friend, to celebrate his 75 th birthday and 50 th anniversary of his career since the publication of Lelej (1971), with a common suffix for mutillid genera. Gender feminine.	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
6A19D29C19245A50A33A7EB56087CE48.taxon	biology_ecology	Biology. This genus is rare in collections; only twelve specimens of A. bagrada, nine specimens of A. depressicornis, and seven specimens of A. nallinia have been reported by previous authors (Cameron 1902; Zavattari 1914; Mickel 1935, 1937; Lelej 1996 b). In this study, however, a series of 28 specimens collected in Malaise traps placed in a lowland rain forest at Pasoh Forest Reserve was examined, suggesting greater abundance in forested habitats. See Maeto et al. (1999) for sampling sites.	en	Okayasu, Juriya (2021): A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species. Journal of Hymenoptera Research 84: 145-168, DOI: http://dx.doi.org/10.3897/jhr.84.68709, URL: http://dx.doi.org/10.3897/jhr.84.68709
