taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
C7BE87A15935594A9F4F197B961E885C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579480	https://doi.org/10.3897/jhr.84.68709.figures1-6	Figures 1 - 6. Arkaditilla spp., ♂, habitus 1, 3, 5 dorsal view 2, 4, 6 lateral view 1, 2 A. bagrada (Cameron) 3, 4 A. depressicornis (Mickel) 5, 6 A. frim sp. nov., holotype. Scale bars: 3 mm.	Figures 1 - 6. Arkaditilla spp., ♂, habitus 1, 3, 5 dorsal view 2, 4, 6 lateral view 1, 2 A. bagrada (Cameron) 3, 4 A. depressicornis (Mickel) 5, 6 A. frim sp. nov., holotype. Scale bars: 3 mm.	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
C7BE87A15935594A9F4F197B961E885C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579482	https://doi.org/10.3897/jhr.84.68709.figures13-18	Figures 13 - 18. Arkaditilla spp., ♂, face 13 A. bagrada (Cameron) 14 A. depressicornis (Mickel) 15 A. frim sp. nov., holotype 16 A. gibba sp. nov., paratype 17 A. leleji sp. nov., holotype 18 A. nallinia (Zavattari).	Figures 13 - 18. Arkaditilla spp., ♂, face 13 A. bagrada (Cameron) 14 A. depressicornis (Mickel) 15 A. frim sp. nov., holotype 16 A. gibba sp. nov., paratype 17 A. leleji sp. nov., holotype 18 A. nallinia (Zavattari).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
C7BE87A15935594A9F4F197B961E885C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579483	https://doi.org/10.3897/jhr.84.68709.figures19-36	Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari).	Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
C7BE87A15935594A9F4F197B961E885C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579484	https://doi.org/10.3897/jhr.84.68709.figures37-48	Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari).	Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
C7BE87A15935594A9F4F197B961E885C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579485	https://doi.org/10.3897/jhr.84.68709.figures49-66	Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari).	Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
C7BE87A15935594A9F4F197B961E885C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579486	https://doi.org/10.3897/jhr.84.68709.figures67-72	Figures 67 - 72. Arkaditilla spp., ♂, genital volsella, ventro-inner view 67 A. bagrada (Cameron) 68 A. depressicornis (Mickel) 69 A. frim sp. nov., holotype 70 A. gibba sp. nov., holotype 71 A. leleji sp. nov., holotype 72 A. nallinia (Zavattari). White arrows indicate the paracuspis.	Figures 67 - 72. Arkaditilla spp., ♂, genital volsella, ventro-inner view 67 A. bagrada (Cameron) 68 A. depressicornis (Mickel) 69 A. frim sp. nov., holotype 70 A. gibba sp. nov., holotype 71 A. leleji sp. nov., holotype 72 A. nallinia (Zavattari). White arrows indicate the paracuspis.	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
C7BE87A15935594A9F4F197B961E885C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579487	https://doi.org/10.3897/jhr.84.68709.figure73-76	Figure 73 - 76. Distributional map of Arkaditilla spp. based on specimens examined herein and the literature (Cameron 1902; Lelej 1996 b; Mickel 1935, 1937; Zavattari 1914) 73 A. bagrada (Cameron) (red circle) 74 A. depressicornis (Mickel) (blue circle) 75 A. frim sp. nov. (yellow circle), A. gibba sp. nov. (purple triangle), and A. leleji sp. nov. (light blue square) 76 A. nallinia (Zavattari) (orange circle).	Figure 73 - 76. Distributional map of Arkaditilla spp. based on specimens examined herein and the literature (Cameron 1902; Lelej 1996 b; Mickel 1935, 1937; Zavattari 1914) 73 A. bagrada (Cameron) (red circle) 74 A. depressicornis (Mickel) (blue circle) 75 A. frim sp. nov. (yellow circle), A. gibba sp. nov. (purple triangle), and A. leleji sp. nov. (light blue square) 76 A. nallinia (Zavattari) (orange circle).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
08E714DB16BD5B91AF5EBE4F2C2E111A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579480	https://doi.org/10.3897/jhr.84.68709.figures1-6	Figures 1 - 6. Arkaditilla spp., ♂, habitus 1, 3, 5 dorsal view 2, 4, 6 lateral view 1, 2 A. bagrada (Cameron) 3, 4 A. depressicornis (Mickel) 5, 6 A. frim sp. nov., holotype. Scale bars: 3 mm.	Figures 1 - 6. Arkaditilla spp., ♂, habitus 1, 3, 5 dorsal view 2, 4, 6 lateral view 1, 2 A. bagrada (Cameron) 3, 4 A. depressicornis (Mickel) 5, 6 A. frim sp. nov., holotype. Scale bars: 3 mm.	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
08E714DB16BD5B91AF5EBE4F2C2E111A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579482	https://doi.org/10.3897/jhr.84.68709.figures13-18	Figures 13 - 18. Arkaditilla spp., ♂, face 13 A. bagrada (Cameron) 14 A. depressicornis (Mickel) 15 A. frim sp. nov., holotype 16 A. gibba sp. nov., paratype 17 A. leleji sp. nov., holotype 18 A. nallinia (Zavattari).	Figures 13 - 18. Arkaditilla spp., ♂, face 13 A. bagrada (Cameron) 14 A. depressicornis (Mickel) 15 A. frim sp. nov., holotype 16 A. gibba sp. nov., paratype 17 A. leleji sp. nov., holotype 18 A. nallinia (Zavattari).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
08E714DB16BD5B91AF5EBE4F2C2E111A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579483	https://doi.org/10.3897/jhr.84.68709.figures19-36	Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari).	Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
08E714DB16BD5B91AF5EBE4F2C2E111A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579484	https://doi.org/10.3897/jhr.84.68709.figures37-48	Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari).	Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
08E714DB16BD5B91AF5EBE4F2C2E111A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579485	https://doi.org/10.3897/jhr.84.68709.figures49-66	Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari).	Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
08E714DB16BD5B91AF5EBE4F2C2E111A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579486	https://doi.org/10.3897/jhr.84.68709.figures67-72	Figures 67 - 72. Arkaditilla spp., ♂, genital volsella, ventro-inner view 67 A. bagrada (Cameron) 68 A. depressicornis (Mickel) 69 A. frim sp. nov., holotype 70 A. gibba sp. nov., holotype 71 A. leleji sp. nov., holotype 72 A. nallinia (Zavattari). White arrows indicate the paracuspis.	Figures 67 - 72. Arkaditilla spp., ♂, genital volsella, ventro-inner view 67 A. bagrada (Cameron) 68 A. depressicornis (Mickel) 69 A. frim sp. nov., holotype 70 A. gibba sp. nov., holotype 71 A. leleji sp. nov., holotype 72 A. nallinia (Zavattari). White arrows indicate the paracuspis.	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
08E714DB16BD5B91AF5EBE4F2C2E111A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579487	https://doi.org/10.3897/jhr.84.68709.figure73-76	Figure 73 - 76. Distributional map of Arkaditilla spp. based on specimens examined herein and the literature (Cameron 1902; Lelej 1996 b; Mickel 1935, 1937; Zavattari 1914) 73 A. bagrada (Cameron) (red circle) 74 A. depressicornis (Mickel) (blue circle) 75 A. frim sp. nov. (yellow circle), A. gibba sp. nov. (purple triangle), and A. leleji sp. nov. (light blue square) 76 A. nallinia (Zavattari) (orange circle).	Figure 73 - 76. Distributional map of Arkaditilla spp. based on specimens examined herein and the literature (Cameron 1902; Lelej 1996 b; Mickel 1935, 1937; Zavattari 1914) 73 A. bagrada (Cameron) (red circle) 74 A. depressicornis (Mickel) (blue circle) 75 A. frim sp. nov. (yellow circle), A. gibba sp. nov. (purple triangle), and A. leleji sp. nov. (light blue square) 76 A. nallinia (Zavattari) (orange circle).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
2F91B07729FE5D91ABB3599BFDB59905.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579480	https://doi.org/10.3897/jhr.84.68709.figures1-6	Figures 1 - 6. Arkaditilla spp., ♂, habitus 1, 3, 5 dorsal view 2, 4, 6 lateral view 1, 2 A. bagrada (Cameron) 3, 4 A. depressicornis (Mickel) 5, 6 A. frim sp. nov., holotype. Scale bars: 3 mm.	Figures 1 - 6. Arkaditilla spp., ♂, habitus 1, 3, 5 dorsal view 2, 4, 6 lateral view 1, 2 A. bagrada (Cameron) 3, 4 A. depressicornis (Mickel) 5, 6 A. frim sp. nov., holotype. Scale bars: 3 mm.	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
2F91B07729FE5D91ABB3599BFDB59905.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579482	https://doi.org/10.3897/jhr.84.68709.figures13-18	Figures 13 - 18. Arkaditilla spp., ♂, face 13 A. bagrada (Cameron) 14 A. depressicornis (Mickel) 15 A. frim sp. nov., holotype 16 A. gibba sp. nov., paratype 17 A. leleji sp. nov., holotype 18 A. nallinia (Zavattari).	Figures 13 - 18. Arkaditilla spp., ♂, face 13 A. bagrada (Cameron) 14 A. depressicornis (Mickel) 15 A. frim sp. nov., holotype 16 A. gibba sp. nov., paratype 17 A. leleji sp. nov., holotype 18 A. nallinia (Zavattari).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
2F91B07729FE5D91ABB3599BFDB59905.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579483	https://doi.org/10.3897/jhr.84.68709.figures19-36	Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari).	Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
2F91B07729FE5D91ABB3599BFDB59905.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579484	https://doi.org/10.3897/jhr.84.68709.figures37-48	Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari).	Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
2F91B07729FE5D91ABB3599BFDB59905.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579485	https://doi.org/10.3897/jhr.84.68709.figures49-66	Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari).	Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
2F91B07729FE5D91ABB3599BFDB59905.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579486	https://doi.org/10.3897/jhr.84.68709.figures67-72	Figures 67 - 72. Arkaditilla spp., ♂, genital volsella, ventro-inner view 67 A. bagrada (Cameron) 68 A. depressicornis (Mickel) 69 A. frim sp. nov., holotype 70 A. gibba sp. nov., holotype 71 A. leleji sp. nov., holotype 72 A. nallinia (Zavattari). White arrows indicate the paracuspis.	Figures 67 - 72. Arkaditilla spp., ♂, genital volsella, ventro-inner view 67 A. bagrada (Cameron) 68 A. depressicornis (Mickel) 69 A. frim sp. nov., holotype 70 A. gibba sp. nov., holotype 71 A. leleji sp. nov., holotype 72 A. nallinia (Zavattari). White arrows indicate the paracuspis.	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
2F91B07729FE5D91ABB3599BFDB59905.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579487	https://doi.org/10.3897/jhr.84.68709.figure73-76	Figure 73 - 76. Distributional map of Arkaditilla spp. based on specimens examined herein and the literature (Cameron 1902; Lelej 1996 b; Mickel 1935, 1937; Zavattari 1914) 73 A. bagrada (Cameron) (red circle) 74 A. depressicornis (Mickel) (blue circle) 75 A. frim sp. nov. (yellow circle), A. gibba sp. nov. (purple triangle), and A. leleji sp. nov. (light blue square) 76 A. nallinia (Zavattari) (orange circle).	Figure 73 - 76. Distributional map of Arkaditilla spp. based on specimens examined herein and the literature (Cameron 1902; Lelej 1996 b; Mickel 1935, 1937; Zavattari 1914) 73 A. bagrada (Cameron) (red circle) 74 A. depressicornis (Mickel) (blue circle) 75 A. frim sp. nov. (yellow circle), A. gibba sp. nov. (purple triangle), and A. leleji sp. nov. (light blue square) 76 A. nallinia (Zavattari) (orange circle).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
BBE895FEA6A754F0BA61029FFD1C65EB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579481	https://doi.org/10.3897/jhr.84.68709.figures7-12	Figures 7 - 12. Arkaditilla spp., ♂, habitus 7, 9, 11 dorsal view 8, 10, 12 lateral view 7, 8 A. gibba sp. nov., holotype 9, 10 A. leleji sp. nov., holotype 11, 12 A. nallinia (Zavattari). Scale bars: 3 mm.	Figures 7 - 12. Arkaditilla spp., ♂, habitus 7, 9, 11 dorsal view 8, 10, 12 lateral view 7, 8 A. gibba sp. nov., holotype 9, 10 A. leleji sp. nov., holotype 11, 12 A. nallinia (Zavattari). Scale bars: 3 mm.	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
BBE895FEA6A754F0BA61029FFD1C65EB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579482	https://doi.org/10.3897/jhr.84.68709.figures13-18	Figures 13 - 18. Arkaditilla spp., ♂, face 13 A. bagrada (Cameron) 14 A. depressicornis (Mickel) 15 A. frim sp. nov., holotype 16 A. gibba sp. nov., paratype 17 A. leleji sp. nov., holotype 18 A. nallinia (Zavattari).	Figures 13 - 18. Arkaditilla spp., ♂, face 13 A. bagrada (Cameron) 14 A. depressicornis (Mickel) 15 A. frim sp. nov., holotype 16 A. gibba sp. nov., paratype 17 A. leleji sp. nov., holotype 18 A. nallinia (Zavattari).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
BBE895FEA6A754F0BA61029FFD1C65EB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579483	https://doi.org/10.3897/jhr.84.68709.figures19-36	Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari).	Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
BBE895FEA6A754F0BA61029FFD1C65EB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579484	https://doi.org/10.3897/jhr.84.68709.figures37-48	Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari).	Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
BBE895FEA6A754F0BA61029FFD1C65EB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579485	https://doi.org/10.3897/jhr.84.68709.figures49-66	Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari).	Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
BBE895FEA6A754F0BA61029FFD1C65EB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579486	https://doi.org/10.3897/jhr.84.68709.figures67-72	Figures 67 - 72. Arkaditilla spp., ♂, genital volsella, ventro-inner view 67 A. bagrada (Cameron) 68 A. depressicornis (Mickel) 69 A. frim sp. nov., holotype 70 A. gibba sp. nov., holotype 71 A. leleji sp. nov., holotype 72 A. nallinia (Zavattari). White arrows indicate the paracuspis.	Figures 67 - 72. Arkaditilla spp., ♂, genital volsella, ventro-inner view 67 A. bagrada (Cameron) 68 A. depressicornis (Mickel) 69 A. frim sp. nov., holotype 70 A. gibba sp. nov., holotype 71 A. leleji sp. nov., holotype 72 A. nallinia (Zavattari). White arrows indicate the paracuspis.	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
BBE895FEA6A754F0BA61029FFD1C65EB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579487	https://doi.org/10.3897/jhr.84.68709.figure73-76	Figure 73 - 76. Distributional map of Arkaditilla spp. based on specimens examined herein and the literature (Cameron 1902; Lelej 1996 b; Mickel 1935, 1937; Zavattari 1914) 73 A. bagrada (Cameron) (red circle) 74 A. depressicornis (Mickel) (blue circle) 75 A. frim sp. nov. (yellow circle), A. gibba sp. nov. (purple triangle), and A. leleji sp. nov. (light blue square) 76 A. nallinia (Zavattari) (orange circle).	Figure 73 - 76. Distributional map of Arkaditilla spp. based on specimens examined herein and the literature (Cameron 1902; Lelej 1996 b; Mickel 1935, 1937; Zavattari 1914) 73 A. bagrada (Cameron) (red circle) 74 A. depressicornis (Mickel) (blue circle) 75 A. frim sp. nov. (yellow circle), A. gibba sp. nov. (purple triangle), and A. leleji sp. nov. (light blue square) 76 A. nallinia (Zavattari) (orange circle).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
1A048A79BD0E579B8F8A615E811A9FDB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579481	https://doi.org/10.3897/jhr.84.68709.figures7-12	Figures 7 - 12. Arkaditilla spp., ♂, habitus 7, 9, 11 dorsal view 8, 10, 12 lateral view 7, 8 A. gibba sp. nov., holotype 9, 10 A. leleji sp. nov., holotype 11, 12 A. nallinia (Zavattari). Scale bars: 3 mm.	Figures 7 - 12. Arkaditilla spp., ♂, habitus 7, 9, 11 dorsal view 8, 10, 12 lateral view 7, 8 A. gibba sp. nov., holotype 9, 10 A. leleji sp. nov., holotype 11, 12 A. nallinia (Zavattari). Scale bars: 3 mm.	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
1A048A79BD0E579B8F8A615E811A9FDB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579482	https://doi.org/10.3897/jhr.84.68709.figures13-18	Figures 13 - 18. Arkaditilla spp., ♂, face 13 A. bagrada (Cameron) 14 A. depressicornis (Mickel) 15 A. frim sp. nov., holotype 16 A. gibba sp. nov., paratype 17 A. leleji sp. nov., holotype 18 A. nallinia (Zavattari).	Figures 13 - 18. Arkaditilla spp., ♂, face 13 A. bagrada (Cameron) 14 A. depressicornis (Mickel) 15 A. frim sp. nov., holotype 16 A. gibba sp. nov., paratype 17 A. leleji sp. nov., holotype 18 A. nallinia (Zavattari).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
1A048A79BD0E579B8F8A615E811A9FDB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579483	https://doi.org/10.3897/jhr.84.68709.figures19-36	Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari).	Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
1A048A79BD0E579B8F8A615E811A9FDB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579484	https://doi.org/10.3897/jhr.84.68709.figures37-48	Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari).	Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
1A048A79BD0E579B8F8A615E811A9FDB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579485	https://doi.org/10.3897/jhr.84.68709.figures49-66	Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari).	Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
1A048A79BD0E579B8F8A615E811A9FDB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579486	https://doi.org/10.3897/jhr.84.68709.figures67-72	Figures 67 - 72. Arkaditilla spp., ♂, genital volsella, ventro-inner view 67 A. bagrada (Cameron) 68 A. depressicornis (Mickel) 69 A. frim sp. nov., holotype 70 A. gibba sp. nov., holotype 71 A. leleji sp. nov., holotype 72 A. nallinia (Zavattari). White arrows indicate the paracuspis.	Figures 67 - 72. Arkaditilla spp., ♂, genital volsella, ventro-inner view 67 A. bagrada (Cameron) 68 A. depressicornis (Mickel) 69 A. frim sp. nov., holotype 70 A. gibba sp. nov., holotype 71 A. leleji sp. nov., holotype 72 A. nallinia (Zavattari). White arrows indicate the paracuspis.	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
1A048A79BD0E579B8F8A615E811A9FDB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579487	https://doi.org/10.3897/jhr.84.68709.figure73-76	Figure 73 - 76. Distributional map of Arkaditilla spp. based on specimens examined herein and the literature (Cameron 1902; Lelej 1996 b; Mickel 1935, 1937; Zavattari 1914) 73 A. bagrada (Cameron) (red circle) 74 A. depressicornis (Mickel) (blue circle) 75 A. frim sp. nov. (yellow circle), A. gibba sp. nov. (purple triangle), and A. leleji sp. nov. (light blue square) 76 A. nallinia (Zavattari) (orange circle).	Figure 73 - 76. Distributional map of Arkaditilla spp. based on specimens examined herein and the literature (Cameron 1902; Lelej 1996 b; Mickel 1935, 1937; Zavattari 1914) 73 A. bagrada (Cameron) (red circle) 74 A. depressicornis (Mickel) (blue circle) 75 A. frim sp. nov. (yellow circle), A. gibba sp. nov. (purple triangle), and A. leleji sp. nov. (light blue square) 76 A. nallinia (Zavattari) (orange circle).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
7603CBDB1C6855F4B10DBD3ED5EBBA9B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579481	https://doi.org/10.3897/jhr.84.68709.figures7-12	Figures 7 - 12. Arkaditilla spp., ♂, habitus 7, 9, 11 dorsal view 8, 10, 12 lateral view 7, 8 A. gibba sp. nov., holotype 9, 10 A. leleji sp. nov., holotype 11, 12 A. nallinia (Zavattari). Scale bars: 3 mm.	Figures 7 - 12. Arkaditilla spp., ♂, habitus 7, 9, 11 dorsal view 8, 10, 12 lateral view 7, 8 A. gibba sp. nov., holotype 9, 10 A. leleji sp. nov., holotype 11, 12 A. nallinia (Zavattari). Scale bars: 3 mm.	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
7603CBDB1C6855F4B10DBD3ED5EBBA9B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579482	https://doi.org/10.3897/jhr.84.68709.figures13-18	Figures 13 - 18. Arkaditilla spp., ♂, face 13 A. bagrada (Cameron) 14 A. depressicornis (Mickel) 15 A. frim sp. nov., holotype 16 A. gibba sp. nov., paratype 17 A. leleji sp. nov., holotype 18 A. nallinia (Zavattari).	Figures 13 - 18. Arkaditilla spp., ♂, face 13 A. bagrada (Cameron) 14 A. depressicornis (Mickel) 15 A. frim sp. nov., holotype 16 A. gibba sp. nov., paratype 17 A. leleji sp. nov., holotype 18 A. nallinia (Zavattari).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
7603CBDB1C6855F4B10DBD3ED5EBBA9B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579483	https://doi.org/10.3897/jhr.84.68709.figures19-36	Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari).	Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
7603CBDB1C6855F4B10DBD3ED5EBBA9B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579484	https://doi.org/10.3897/jhr.84.68709.figures37-48	Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari).	Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
7603CBDB1C6855F4B10DBD3ED5EBBA9B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579485	https://doi.org/10.3897/jhr.84.68709.figures49-66	Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari).	Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
7603CBDB1C6855F4B10DBD3ED5EBBA9B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579486	https://doi.org/10.3897/jhr.84.68709.figures67-72	Figures 67 - 72. Arkaditilla spp., ♂, genital volsella, ventro-inner view 67 A. bagrada (Cameron) 68 A. depressicornis (Mickel) 69 A. frim sp. nov., holotype 70 A. gibba sp. nov., holotype 71 A. leleji sp. nov., holotype 72 A. nallinia (Zavattari). White arrows indicate the paracuspis.	Figures 67 - 72. Arkaditilla spp., ♂, genital volsella, ventro-inner view 67 A. bagrada (Cameron) 68 A. depressicornis (Mickel) 69 A. frim sp. nov., holotype 70 A. gibba sp. nov., holotype 71 A. leleji sp. nov., holotype 72 A. nallinia (Zavattari). White arrows indicate the paracuspis.	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
7603CBDB1C6855F4B10DBD3ED5EBBA9B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579487	https://doi.org/10.3897/jhr.84.68709.figure73-76	Figure 73 - 76. Distributional map of Arkaditilla spp. based on specimens examined herein and the literature (Cameron 1902; Lelej 1996 b; Mickel 1935, 1937; Zavattari 1914) 73 A. bagrada (Cameron) (red circle) 74 A. depressicornis (Mickel) (blue circle) 75 A. frim sp. nov. (yellow circle), A. gibba sp. nov. (purple triangle), and A. leleji sp. nov. (light blue square) 76 A. nallinia (Zavattari) (orange circle).	Figure 73 - 76. Distributional map of Arkaditilla spp. based on specimens examined herein and the literature (Cameron 1902; Lelej 1996 b; Mickel 1935, 1937; Zavattari 1914) 73 A. bagrada (Cameron) (red circle) 74 A. depressicornis (Mickel) (blue circle) 75 A. frim sp. nov. (yellow circle), A. gibba sp. nov. (purple triangle), and A. leleji sp. nov. (light blue square) 76 A. nallinia (Zavattari) (orange circle).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
6A19D29C19245A50A33A7EB56087CE48.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579480	https://doi.org/10.3897/jhr.84.68709.figures1-6	Figures 1 - 6. Arkaditilla spp., ♂, habitus 1, 3, 5 dorsal view 2, 4, 6 lateral view 1, 2 A. bagrada (Cameron) 3, 4 A. depressicornis (Mickel) 5, 6 A. frim sp. nov., holotype. Scale bars: 3 mm.	Figures 1 - 6. Arkaditilla spp., ♂, habitus 1, 3, 5 dorsal view 2, 4, 6 lateral view 1, 2 A. bagrada (Cameron) 3, 4 A. depressicornis (Mickel) 5, 6 A. frim sp. nov., holotype. Scale bars: 3 mm.	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
6A19D29C19245A50A33A7EB56087CE48.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579481	https://doi.org/10.3897/jhr.84.68709.figures7-12	Figures 7 - 12. Arkaditilla spp., ♂, habitus 7, 9, 11 dorsal view 8, 10, 12 lateral view 7, 8 A. gibba sp. nov., holotype 9, 10 A. leleji sp. nov., holotype 11, 12 A. nallinia (Zavattari). Scale bars: 3 mm.	Figures 7 - 12. Arkaditilla spp., ♂, habitus 7, 9, 11 dorsal view 8, 10, 12 lateral view 7, 8 A. gibba sp. nov., holotype 9, 10 A. leleji sp. nov., holotype 11, 12 A. nallinia (Zavattari). Scale bars: 3 mm.	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
6A19D29C19245A50A33A7EB56087CE48.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579482	https://doi.org/10.3897/jhr.84.68709.figures13-18	Figures 13 - 18. Arkaditilla spp., ♂, face 13 A. bagrada (Cameron) 14 A. depressicornis (Mickel) 15 A. frim sp. nov., holotype 16 A. gibba sp. nov., paratype 17 A. leleji sp. nov., holotype 18 A. nallinia (Zavattari).	Figures 13 - 18. Arkaditilla spp., ♂, face 13 A. bagrada (Cameron) 14 A. depressicornis (Mickel) 15 A. frim sp. nov., holotype 16 A. gibba sp. nov., paratype 17 A. leleji sp. nov., holotype 18 A. nallinia (Zavattari).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
6A19D29C19245A50A33A7EB56087CE48.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579483	https://doi.org/10.3897/jhr.84.68709.figures19-36	Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari).	Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
6A19D29C19245A50A33A7EB56087CE48.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579484	https://doi.org/10.3897/jhr.84.68709.figures37-48	Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari).	Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
6A19D29C19245A50A33A7EB56087CE48.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579485	https://doi.org/10.3897/jhr.84.68709.figures49-66	Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari).	Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
6A19D29C19245A50A33A7EB56087CE48.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579486	https://doi.org/10.3897/jhr.84.68709.figures67-72	Figures 67 - 72. Arkaditilla spp., ♂, genital volsella, ventro-inner view 67 A. bagrada (Cameron) 68 A. depressicornis (Mickel) 69 A. frim sp. nov., holotype 70 A. gibba sp. nov., holotype 71 A. leleji sp. nov., holotype 72 A. nallinia (Zavattari). White arrows indicate the paracuspis.	Figures 67 - 72. Arkaditilla spp., ♂, genital volsella, ventro-inner view 67 A. bagrada (Cameron) 68 A. depressicornis (Mickel) 69 A. frim sp. nov., holotype 70 A. gibba sp. nov., holotype 71 A. leleji sp. nov., holotype 72 A. nallinia (Zavattari). White arrows indicate the paracuspis.	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
6A19D29C19245A50A33A7EB56087CE48.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/579487	https://doi.org/10.3897/jhr.84.68709.figure73-76	Figure 73 - 76. Distributional map of Arkaditilla spp. based on specimens examined herein and the literature (Cameron 1902; Lelej 1996 b; Mickel 1935, 1937; Zavattari 1914) 73 A. bagrada (Cameron) (red circle) 74 A. depressicornis (Mickel) (blue circle) 75 A. frim sp. nov. (yellow circle), A. gibba sp. nov. (purple triangle), and A. leleji sp. nov. (light blue square) 76 A. nallinia (Zavattari) (orange circle).	Figure 73 - 76. Distributional map of Arkaditilla spp. based on specimens examined herein and the literature (Cameron 1902; Lelej 1996 b; Mickel 1935, 1937; Zavattari 1914) 73 A. bagrada (Cameron) (red circle) 74 A. depressicornis (Mickel) (blue circle) 75 A. frim sp. nov. (yellow circle), A. gibba sp. nov. (purple triangle), and A. leleji sp. nov. (light blue square) 76 A. nallinia (Zavattari) (orange circle).	2021-08-24	Okayasu, Juriya		Zenodo	biologists	Okayasu, Juriya			
