identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
DCA6ABECEBF05A7C805731A161B965CE.text	DCA6ABECEBF05A7C805731A161B965CE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Toxeuma acilius (Walker 1848)	<div><p>Toxeuma acilius (Walker, 1848)</p> <p>Figs 1-6</p> <p>Lamprotatus acilius Walker, 1848: 169. Lectotype male, (NHMUK, examined) designated by Graham 1959: 104.</p> <p>Toxeuma acilius (Walker, 1848) new combination in Graham, 1959: 104.</p> <p>Material examined.</p> <p>Lectotype male (NHMUK): United Kingdom: “LECTOTYPE”, “Type”, “England”, " Lamprotatus Acilius Walker ♂ / Stood under this name in old B.M. Coll. C. Waterhouse", " Acilius,", " Toxeuma ericae (Delucchi)", " Toxeuma acilius (Walk.) det. M.W.R. de V. Graham", "B.M. TYPE HYM. 5.1928", " NHMUK 013456778". Other material: United Kingdom: England, 1 female, Berkshire, Wytham Wood, 3.VIII.1954, coll. M.W. Graham. Russia: Bryansk Prov., 2 females, 15 km E Pochep Town, Uruch'e Vill., 52°55.00"N, 33°57.50"E, 20.VIII.2020, coll. O. Kosheleva and E. Tselikh; Kamchatka Reg., 1 female, 80 km S Lazo Vill., Kronotsky Reserve, Ipuin, 19.VII.2013, coll. D. Rachin and E. Tselikh; Sakhalinskaya Prov., 1 female, Kunashir Island, Dubovoe Vill., 22.VII.1973, coll. D. Kasparyan; 1 female, Mendeleevo Vill., 3.VIII.2011, coll. D. Rachin and E. Tselikh; Smolensk Prov., 1 female, Smolensk, 54°49.103"N, 32°5.09"E, 23.VIII.2020, coll. O.V. Kosheleva); Tyumen’ Prov., 1 female, 2 km SW Tyumen’, Bereznyakovsky Vill., 3.VII.2018, coll. Tselikh.</p> <p>Distribution.</p> <p>Czech Republic, Germany, Netherlands, Romania, Russia (Russian Far East; Western and Eastern Siberia, European part of Russia - new records), Sweden, United Kingdom (Walker 1848; Graham 1959; Tselikh 2014, 2019; Noyes 2019).</p></div> 	http://treatment.plazi.org/id/DCA6ABECEBF05A7C805731A161B965CE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tselikh, Ekaterina V.;Dale-Skey, Natalie	Tselikh, Ekaterina V., Dale-Skey, Natalie (2021): Review of the genus Toxeuma Walker, 1833 (Hymenoptera, Pteromalidae) from Russia, with a key to Palaearctic species. Journal of Hymenoptera Research 84: 391-403, DOI: http://dx.doi.org/10.3897/jhr.84.68627, URL: http://dx.doi.org/10.3897/jhr.84.68627
E094B7B426D7565784C4A3C662A53939.text	E094B7B426D7565784C4A3C662A53939.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Toxeuma discretum Graham 1984	<div><p>Toxeuma discretum Graham, 1984</p> <p>Figs 7-9</p> <p>Toxeuma discretum Graham, 1984: 501. Holotype female (NHMUK, examined).</p> <p>Material examined.</p> <p>Holotype female (NHMUK): United Kingdom: “Holotype”, "England: Oxon Bald Hill, nr. Lewknor, 18.6.1958", " Toxeuma discretum sp.n. HOLOTYPE ♀ M. de V. Graham det. 1983", "B.M. TYPE HYM 5.3020", " NHMUK 013456774".</p> <p>Distribution.</p> <p>Belgium, France, Sweden, United Kingdom (Graham 1984; Noyes 2019).</p></div> 	http://treatment.plazi.org/id/E094B7B426D7565784C4A3C662A53939	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tselikh, Ekaterina V.;Dale-Skey, Natalie	Tselikh, Ekaterina V., Dale-Skey, Natalie (2021): Review of the genus Toxeuma Walker, 1833 (Hymenoptera, Pteromalidae) from Russia, with a key to Palaearctic species. Journal of Hymenoptera Research 84: 391-403, DOI: http://dx.doi.org/10.3897/jhr.84.68627, URL: http://dx.doi.org/10.3897/jhr.84.68627
2871627491375E65B4EB7930B67EC2CA.text	2871627491375E65B4EB7930B67EC2CA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Toxeuma fuscicorne Walker 1833	<div><p>Toxeuma fuscicorne Walker, 1833</p> <p>Figs 10-14</p> <p>Toxeuma fuscicornis Walker, 1833: 378; lectotype female (NHMUK, not examined) designated by Graham (1969: 147).</p> <p>Miscogaster lugubris Walker, 1833: 462. Type specimens possibly lost (Graham 1969: 147). Synonymy by Walker (1846: 28) under Toxeuma ericae Walker, 1833.</p> <p>Toxeuma ericae Walker, 1833: 379; lectotype female (NHMUK, not examined) designated by Graham (1969: 147). Synonymy by Graham (1959: 101).</p> <p>Gastrancistrus accia Walker, 1848: 156; lectotype female, (NHMUK, not examined) designated by Graham (1969: 147). Synonymy by Graham (1959: 101).</p> <p>Material examined.</p> <p>Other material: Russia: Belgorod Prov., 12 females, 3 males, Borisovka Vill., “Belogor’e” Reserve, "Les na Vorskle", 11.VI.2018, coll. Tselikh.</p> <p>Distribution.</p> <p>Belgium, Croatia, Czech Republic, France, Germany, Hungary, Iran, Netherlands, Romania, Russia (new record), Spain, Sweden, Switzerland, United Kingdom (Walker 1833; Graham 1959, 1969; Noyes 2019).</p> <p>Biology.</p> <p>Primary parasitoid of Agromyza schineri Giraud, 1861 (Diptera, Agromyzidae) - Bouček (1977).</p> </div>	http://treatment.plazi.org/id/2871627491375E65B4EB7930B67EC2CA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tselikh, Ekaterina V.;Dale-Skey, Natalie	Tselikh, Ekaterina V., Dale-Skey, Natalie (2021): Review of the genus Toxeuma Walker, 1833 (Hymenoptera, Pteromalidae) from Russia, with a key to Palaearctic species. Journal of Hymenoptera Research 84: 391-403, DOI: http://dx.doi.org/10.3897/jhr.84.68627, URL: http://dx.doi.org/10.3897/jhr.84.68627
AC4F99ED3DB85ADFA0697A8B9BCC1109.text	AC4F99ED3DB85ADFA0697A8B9BCC1109.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Toxeuma leleji Tselikh 2021	<div><p>Toxeuma leleji Tselikh sp. nov.</p> <p>Figs 15-21</p> <p>Comparison.</p> <p>Toxeuma leleji is similar to T. acilius (Walker) and T. paludum Graham; the differences between these species are given in the key.</p> <p>Females of Toxeuma leleji are also similar to those of the Nearctic species Toxeuma inopinum Heydon because they have an antennal clava with needle-like spicula, F1-F4 elongate, fore wing with pilose basal vein, but Toxeuma leleji females have the gena not distinctly hollowed at mouth corner, petiole longer than wide, Mt2 0.85-0.9 times median length of metasoma.</p> <p>Description.</p> <p>Female. Body length 1.5 mm. Fore wing length 1.5 mm.</p> <p>Head, mesosoma and metasoma, metallic bluish-green with diffuse coppery lustre; antenna with scape yellow, pedicel yellowish-brown, flagellum brown; all coxae metallic bluish-green with diffuse coppery lustre; all femora, tibiae and tarsi yellow except last segment yellowish-brown; fore wing hyaline, venation yellowish-brown; ovipositor sheath black.</p> <p>Head in dorsal view 1.93-2.07 times as broad as long and 1.36-1.38 times as broad as mesoscutum; in frontal view 1.28-1.3 times broader than high. POL 1.46-1.57 times OOL. Eye height 1.21-1.26 times eye length and 2.90-2.92 times as long as malar space. Distance between antennal toruli and lower margin of clypeus 0.75 times distance between antennal toruli and median ocellus. Clypeus alutaceous, its lower margin slightly arched. Antenna with scape 0.92-0.96 times as long as eye height and 1.12-1.22 times as long as eye length; pedicel 1.61-1.66 times as long as broad and 1.40-1.42 times as long as F1; combined length of pedicel and flagellum 1.10-1.14 times breadth of head; flagellum almost filiform; F1-F3 longer than broad, F4-F5 subquadrate, F6 transverse (on dry material) and F1 longer than broad, F2-F3 subquadrate, F4-F6 transverse (on slide); clava 2.83-2.86 times as long as broad (on dry material) and 2.58 times (on slide).</p> <p>Mesosoma 2.20-2.45 times as long as broad. Scutellum finely reticulate, 1.0-1.1 times as long as broad. Propodeum medially as long as scutellum; median carina and plicae present; sculpture reticulate with irregular rugae; nucha short and weakly reticulate. Sculptured part of petiole 1.17-1.25 times as long as broad, with irregular sculpture. Fore wing 2.34-2.89 times as long as maximum width; basal cell bare; basal vein pilose; speculum partly open; M 1.16-1.22 times as long as P and 1.95-2.00 times as long as S.</p> <p>Metasoma ovate, 1.50-1.64 times as long as broad and 0.40-0.45 times as long as mesosoma and head together; Mt2 0.85-0.90 times median length of metasoma. Ovipositor sheath projecting slightly beyond apex of metasoma.</p> <p>Etymology.</p> <p>The species is named in honour of the prominent entomologist, an expert on Mutillidae (Hymenoptera), Prof. Arkady S. Lelej.</p> <p>Material examined.</p> <p>Holotype female (ZISP): Russia: Sakhalin Prov., Kunashir Island, Golovnina Volcano, 27.VII.1981, coll. E. Sugonyaev.</p> <p>Paratype female (ZISP): same data as holotype.</p> <p>Distribution.</p> <p>Russian Far East (Sakhalinskaya Prov.).</p></div> 	http://treatment.plazi.org/id/AC4F99ED3DB85ADFA0697A8B9BCC1109	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tselikh, Ekaterina V.;Dale-Skey, Natalie	Tselikh, Ekaterina V., Dale-Skey, Natalie (2021): Review of the genus Toxeuma Walker, 1833 (Hymenoptera, Pteromalidae) from Russia, with a key to Palaearctic species. Journal of Hymenoptera Research 84: 391-403, DOI: http://dx.doi.org/10.3897/jhr.84.68627, URL: http://dx.doi.org/10.3897/jhr.84.68627
8A514512221F5F10A503721189B5580A.text	8A514512221F5F10A503721189B5580A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Toxeuma paludum Graham 1959	<div><p>Toxeuma paludum Graham, 1959</p> <p>Figs 22-27</p> <p>Toxeuma paludum Graham, 1959: 102. Holotype female (OUMNH, examined), paratypes (NHMUK, examined).</p> <p>Material examined.</p> <p>Holotype female (OUMNH): "Otmoor OX. MWG. (1) 6.7.1956", " Toxeuma paludum TYPE ♀", " Toxeuma paludum sp.n. M. de V. Graham det. 1957 HOLOTYPE [female symbol]", "TYPE HYM: 1219 Toxeuma paludum GRAHAM HOPE DEPT. OXFORD". Paratype female (NHMUK): United Kingdom: “PARATYPE”, "OXON. Otmoor. (3) 10.7.1956 M. de V. Graham", " Toxeuma paludum Paratype sp.n. M. de V. Graham", " NHMUK 013456775". Other material: Russia: Irkutsk Prov., 6 females, 3 males, Sludyanka Vill., "Shamansky Cape", 05.VIII.2019, coll. E. Tselikh; Kamchatka Terr., 6 females, 3 males, 80 km S Lazo Vill, Kronotsky Reserve, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=159.96587&amp;materialsCitation.latitude=55.11615" title="Search Plazi for locations around (long 159.96587/lat 55.11615)">Ipuin</a>, 55°06.969'N, 159°57.952'E, 18.VII.2013, coll. D. Rachin and E. Tselikh; Sakhalinskaya Prov., 1 female, Iturup Island, Goryachie Kluchi Vill., 30.VII.2002, coll. A. Lelej; 1 female, Kunashir Island, Mendeleevo Vill., 3.VIII.2011, coll. D. Rachin and E. Tselikh; 4 females, 3 males, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.49535&amp;materialsCitation.latitude=49.77465" title="Search Plazi for locations around (long 145.49535/lat 49.77465)">Dubovoe Vill.</a>, 49°46.479'N, 145°29.721'E, 21.VII.2011, coll. D. Rachin and E. Tselikh; 8 females, 3 males, Sakhalin Island, Sokol Vill., 4.VIII.2011, coll. D. Rachin and E. Tselikh; 1 female, Shikotan Island, Gorobets Bay, 18.VIII.1998, coll. A. Lelej; Tyumen’ Prov., 2 females, 5 males, 35 km NW Tyumen’ Sity, Salairka Vill., 5.VII.2018, coll. Tselikh; 5 km N Tyumen’ Sity, Lipovoe Lake, 7.VII.2018, coll. Tselikh.</p> <p>Distribution.</p> <p>Russia (Russian Far East; Western and Eastern Siberia - new records), Sweden, United Kingdom (Graham 1959; Tselikh 2014, 2019; Noyes 2019).</p> <p>Biology.</p> <p>Primary parasitoid of Coleophora sp. (Lepidoptera, Coleophoridae) (Askew 1968).</p> </div>	http://treatment.plazi.org/id/8A514512221F5F10A503721189B5580A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tselikh, Ekaterina V.;Dale-Skey, Natalie	Tselikh, Ekaterina V., Dale-Skey, Natalie (2021): Review of the genus Toxeuma Walker, 1833 (Hymenoptera, Pteromalidae) from Russia, with a key to Palaearctic species. Journal of Hymenoptera Research 84: 391-403, DOI: http://dx.doi.org/10.3897/jhr.84.68627, URL: http://dx.doi.org/10.3897/jhr.84.68627
9CF07E7674CB5B37872F5697CCD717BA.text	9CF07E7674CB5B37872F5697CCD717BA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Toxeuma styliclava (Hedqvist 1974)	<div><p>Toxeuma styliclava (Hedqvist, 1974)</p> <p>Figs 28-32</p> <p>Cirdania styliclava Hedqvist, 1974: 145-146. Holotype female (NHRS, not examined).</p> <p>Toxeuma styliclava (Hedqvist, 1974) new combination in Bouček (1991: 205).</p> <p>Toxeuma mucronatum Graham, 1984: 501-502. Holotype female (NHMUK, not examined). Synonymy by Bouček, 1991: 205.</p> <p>Material examined.</p> <p>Paratype female (NHMUK): Sweden: "Sk. Sandhammaren 24/7 1973 leg. K. J. Hedqvist", "PARATYPUS Cirdania gen.n. Cirdania styliclava sp.n. ♀ K-J Hedqvist det. 1973", "Hedqvist coll. BMNH(E) 2011-27", " NHMUK 013456781". Other material: Russia: Primorskii Reg., 1 female, Lazovsky Reserve, “Korpad”, 10-14.VIII.2010, coll. D. Rachin and E. Tselikh; 1 female, Ussuriysky Reserve, 3.VIII.1961, coll. V. Trjapitzin; Sakhalinskaya Prov., 1 female, Kunashir Island, Stolbchaty Cape, 3.VIII.2011, coll. D. Rachin and E. Tselikh; 1 female, Sernovodsk Vill., 22.VII.1981, coll. E. Sugonyaev; 1 female, Tret'yakovo Vill., 29-30.VII.2011, coll. D. Rachin and E. Tselikh; 1 female, Sakhalin Island, Sokol Vill., 4.VIII.2011, coll. D. Rachin and E. Tselikh.</p> <p>Distribution.</p> <p>Belgium, France, Russia (Russian Far East - new record), Sweden (Hedqvist 1974; Noyes 2019).</p></div> 	http://treatment.plazi.org/id/9CF07E7674CB5B37872F5697CCD717BA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tselikh, Ekaterina V.;Dale-Skey, Natalie	Tselikh, Ekaterina V., Dale-Skey, Natalie (2021): Review of the genus Toxeuma Walker, 1833 (Hymenoptera, Pteromalidae) from Russia, with a key to Palaearctic species. Journal of Hymenoptera Research 84: 391-403, DOI: http://dx.doi.org/10.3897/jhr.84.68627, URL: http://dx.doi.org/10.3897/jhr.84.68627
46DFEE0142B551AA8C3734E04AA9EBF9.text	46DFEE0142B551AA8C3734E04AA9EBF9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Toxeuma subtruncatum Graham 1959	<div><p>Toxeuma subtruncatum Graham, 1959</p> <p>Figs 33-38</p> <p>Toxeuma subtruncatum Graham, 1959: 105. Holotype male (OUMNH, not examined).</p> <p>Material examined.</p> <p>Paratype female (NHMUK): United Kingdom: "ENGLAND: Warw Ufton Wood 15.vi.1957", "M.W.R. de V. Graham coll. BMNH(E) 1995-489", " Toxeuma subtruncatum Paratype ♀", “PARATYPE”, " NHMUK 013456776"; paratype male (NHMUK): "Wytham Wood BERKS. (2) 16.5.1951 M.W. Graham", " Toxeuma subtruncatum M. de V. G. sp.n", " NHMUK 010749000". Other material: United Kingdom: "Wytham Wood BERKS 11.6.1951 M.W. Graham", " Toxeuma subtruncatum Graham", " NHMUK 013456773"; Russia: Primorskii Reg., 1 female, Ussuriysky Reserve, 30.VII.1975, coll. N. Storozheva; Sakhalinskaya Prov., 1 female, Kunashir Island, Dubovoe Vill., 31.VIII.1973, coll. D. Kasparyan.</p> <p>Distribution.</p> <p>Belgium, Czech Republic, Netherlands, Russia (Russian Far East; including Primorskii Reg. - new record), Sweden, United Kingdom (Graham 1959; Tselikh 2014, 2019; Noyes 2019).</p></div> 	http://treatment.plazi.org/id/46DFEE0142B551AA8C3734E04AA9EBF9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tselikh, Ekaterina V.;Dale-Skey, Natalie	Tselikh, Ekaterina V., Dale-Skey, Natalie (2021): Review of the genus Toxeuma Walker, 1833 (Hymenoptera, Pteromalidae) from Russia, with a key to Palaearctic species. Journal of Hymenoptera Research 84: 391-403, DOI: http://dx.doi.org/10.3897/jhr.84.68627, URL: http://dx.doi.org/10.3897/jhr.84.68627
29588486EEC151C39B3E8D8DB3200A9E.text	29588486EEC151C39B3E8D8DB3200A9E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Toxeuma Walker 1833	<div><p>Toxeuma Walker, 1833.</p> <p>Toxeuma fuscicornis Type species: Toxeuma fuscicornis Walker, 1833, by subsequent designation of Westwood (1839: 68).</p> <p>Cirdania Hedqvist, 1974: 145. Type species: Cirdania styliclava Hedqvist, 1974, by original designation. Synonymy by Bouček (1991: 205).</p> <p>Diagnosis.</p> <p>Clypeal margin entire, truncate or slightly arched; antennal clava with or without short or long spicula; antennal formula 11263; pronotum with collar margin carinate; notauli complete; prepectus reticulate, without carinae; sculpture of propodeum at least partly irregular, petiole from transverse to slightly elongate; fore wing with slender M, metasoma ovate or lanceolate (Graham 1959; Hedqvist 1974; Heydon and Grissell 1988; Bouček and Rasplus 1991).</p> <p>Distribution.</p> <p>Palaearctic, Nearctic and Neotropics.</p> <p>Comments.</p> <p>Unfortunately, the biology of most Toxeuma species is unknown. The only biological records exist for T. fuscicorne, which was reared from the primary host Agromyza schineri Giraud, 1861 (Diptera, Agromyzidae) (Bouček 1977), and for T. paludum, reared from the primary host Coleophora sp. (Lepidoptera, Coleophoridae) (Askew 1968).</p> <p>The genus can be distinguished from other Pteromalidae genera using the keys of Graham (1969) and Bouček and Rasplus (1991).</p> <p>In this genus, the antennal clava can have three variations: acute (Fig. 24), with a spicula (e.g., Figs 4, 17, 27, 30) or without spicula and rounded apically (e.g., Figs 13, 35). This is an important feature to identify species of this genus. Unfortunately, in the original descriptions of T. acilius (Walker, 1848) and T. paludum (Graham, 1959) as well as in previously published keys (Graham 1959, 1969) this feature was neither mentioned nor shown on the figures. However, after studying the types and additional material, the presence of a spicula was found in almost all individuals of these species. Interestingly, T. paludum Graham is characterized by variation of antennal clava from acute (Fig. 24) to that with very short and straight spicula (e.g., Figs 23, 27).</p> <p>The discovery of a previously undescribed spicula in some Palaearctic species suggests that some members of Toxeuma from Nearctic Region described by Heydon and Grissell (1988) might be synonyms of the Palaearctic species, but additional research is needed to confirm this assumption.</p> </div>	http://treatment.plazi.org/id/29588486EEC151C39B3E8D8DB3200A9E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tselikh, Ekaterina V.;Dale-Skey, Natalie	Tselikh, Ekaterina V., Dale-Skey, Natalie (2021): Review of the genus Toxeuma Walker, 1833 (Hymenoptera, Pteromalidae) from Russia, with a key to Palaearctic species. Journal of Hymenoptera Research 84: 391-403, DOI: http://dx.doi.org/10.3897/jhr.84.68627, URL: http://dx.doi.org/10.3897/jhr.84.68627
