taxonID	type	description	language	source
03BA87A5897C2A2FBD7A2D55FAACFA18.taxon	description	Figs 2 – 7	en	Zubair, R. M., Maqbool, Amir, Wachkoo, Aijaz Ahmad, Biffi, Gabriel (2021): A review of the Himalayan genus Trypheridium Brancucci (Coleoptera: Cantharidae: Chauliognathinae with description of a new species. European Journal of Taxonomy 764: 18-36, DOI: https://doi.org/10.5852/ejt.2021.764.1467, URL: http://dx.doi.org/10.5852/ejt.2021.764.1467
03BA87A5897C2A2FBD7A2D55FAACFA18.taxon	type_taxon	Type species Trypherus nuristanicus Wittmer, 1956, by original designation.	en	Zubair, R. M., Maqbool, Amir, Wachkoo, Aijaz Ahmad, Biffi, Gabriel (2021): A review of the Himalayan genus Trypheridium Brancucci (Coleoptera: Cantharidae: Chauliognathinae with description of a new species. European Journal of Taxonomy 764: 18-36, DOI: https://doi.org/10.5852/ejt.2021.764.1467, URL: http://dx.doi.org/10.5852/ejt.2021.764.1467
03BA87A5897C2A2FBD7A2D55FAACFA18.taxon	diagnosis	Diagnosis Trypheridium is similar to Trypherus, and distinguished by sexual characters: tergite IX with dorsal and ventral blades (“ lames tergales et sternales ”) very long and asymmetrical (Figs 3 A – D, 6 E – F); aedeagus without left paramere, median lobe membranous (Figs 3 F – J, 6 G – I), with orifice apically (not preapically as in Trypherus); female with sternite VII remarkably broad, partly or completely concealing most of sternite VIII (Figs 4 G – H, 7 B); bursa copulatrix elongate, with two tubular spermathecae and one accessory gland connecting the bursa in its postero-dorsal part (Figs 5 F, 7 D).	en	Zubair, R. M., Maqbool, Amir, Wachkoo, Aijaz Ahmad, Biffi, Gabriel (2021): A review of the Himalayan genus Trypheridium Brancucci (Coleoptera: Cantharidae: Chauliognathinae with description of a new species. European Journal of Taxonomy 764: 18-36, DOI: https://doi.org/10.5852/ejt.2021.764.1467, URL: http://dx.doi.org/10.5852/ejt.2021.764.1467
03BA87A5897C2A2FBD7A2D55FAACFA18.taxon	description	Re-description (Figs 6 – 7) Coloration General coloration pale to sulphur yellow with dark brown to black patches; head yellow anteriorly at frons and clypeus and infuscate at vertex and occipital region; antennae dark brown to black, the three or four basal antennomeres yellow; pronotum with a transversal black band at posterior half; elytra with dark brown to black triangular patches, from the base of elytra (not reaching the humeri) and extending posteriorly; wings dark brown; legs dark brown or black, usually fore and mid legs light brown with dark spots; abdominal tergites and ventrites black in the centre and yellow at the margins. Coloration in females usually darker. Male HABITUS. Length 6.5 – 8.5 mm. HEAD. Head (including the eyes) wider than pronotum; vertex flat, frons concave between the eyes, occipital region convex, sides rounded and regularly narrowing behind the eyes. Eyes large, oblong, bulging, longer than wide, broadly separated. Fronto-clypeal suture distinct laterally, indistinct in the middle; anterior margin of clypeus slightly sinuous, nearly straight; labrum membranous. Mandibles arcuate, apex acute, blunt, retinaculum in the same plan as incisor, apex rounded, with a small supplementary tooth at the base of retinaculum. Maxillary palp with fourth palpomere elongate, narrowing apicad. Antennae not juxtaposed, separated by at least the width of antennal sockets; with 11 antennomeres, filiform, without modifications; antennomere I slender, without swellings. PRONOTUM. Transverse, wider than long, convex; anterior margin broadly arched, lateral margins nearly straight, slightly narrowing posteriorly; posterior margin projected posteriorly; anterior and posterior angles rounded, barely distinct. Elytra short, 1.2 – 1.4 times longer than combined width, slightly narrowed laterally, suture dehiscent posteriorly, apex rounded. WINGS. With reduced venation, radial cell 2 R 1 closed; vein r-m barely visible, vein Cu straight, vein Mr fading; transversal vein cu-a absent; Rr extends beyond the joining of vein r; vein A x 2 well visible, not joining Ax 1 + 2 and ending near the joining of vein A and Ax 1 + 2. LEGS. Slender, without swellings, teeth or remarkable modifications; apex of fore tibia with a pair of tiny spurs; all tarsal claws with broad internal teeth. ABDOMEN. Weakly sclerotised; tergites with a pair of small glandular pores on the distal corners; pores of tergite VIII in the apex of projections. Tergite VIII long and broad, with distal corners projected posteriorly, distal margin arcuate, concave. Tergite IX (paraproct) strongly modified in a capsule with a pair of long and asymmetrical lamellar projections directed ventrally; left blade (“ lame sternale ” of Brancucci) longer, flattened; right blade (“ lame tergale ” of Brancucci) shorter, more strongly curved ventrally. Tergite X (proctiger) divided posteriorly into two projecting lobes. Sternite VIII (ventrite VII) asymmetrical, distal margin sinuous and deeply notched, sides projected posteriorly. Sternite IX concealed underneath sternite VIII, elongate, lamellar, weakly sclerotised. AEDEAGUS. Elongated and weakly sclerotised; left paramere absent, right paramere and the prolongation of tegmen elongate and directed posteriorly; left prolongation of tegmen broad and lamellar, margins sinuous. Median lobe membranous, short, not twisted, with its broad opening located apically. Female Similar to males, except for eyes slightly smaller and more separated; sternite VII long and broad, mostly concealing sternite VIII, distal margin with elongate lobes projecting posteriorly. Sternite VIII membranous, broadly rounded, distal margin with a broad rounded lobe projecting posteriorly; coxites small, not fused medially, anterior arms very long and slender, apparently fused with valvifers, styles elongate, digitiform. Genitalia membranous, consisting of a long and broad vagina; oviduct anteriorly in the vagina; bursa copulatrix elongate, wrinkled, spiralling; spermatheca formed by two tubular projections; accessory gland large, connected by a long winding duct to the postero-dorsal part of bursa copulatrix.	en	Zubair, R. M., Maqbool, Amir, Wachkoo, Aijaz Ahmad, Biffi, Gabriel (2021): A review of the Himalayan genus Trypheridium Brancucci (Coleoptera: Cantharidae: Chauliognathinae with description of a new species. European Journal of Taxonomy 764: 18-36, DOI: https://doi.org/10.5852/ejt.2021.764.1467, URL: http://dx.doi.org/10.5852/ejt.2021.764.1467
03BA87A5897C2A2FBD7A2D55FAACFA18.taxon	distribution	Distribution Trypheridium seems to be endemic to the Hindu Kush Himalayan Region. The only species described prior to this work has been reported from Tajikistan, Afghanistan (Nuristan) and Pakistan (Fig. 8 A). Herein, we report a new species of this genus from India (Fig. 8 B).	en	Zubair, R. M., Maqbool, Amir, Wachkoo, Aijaz Ahmad, Biffi, Gabriel (2021): A review of the Himalayan genus Trypheridium Brancucci (Coleoptera: Cantharidae: Chauliognathinae with description of a new species. European Journal of Taxonomy 764: 18-36, DOI: https://doi.org/10.5852/ejt.2021.764.1467, URL: http://dx.doi.org/10.5852/ejt.2021.764.1467
03BA87A5897C2A2FBD7A2D55FAACFA18.taxon	discussion	Key to the species of Trypheridium Brancucci, 1985	en	Zubair, R. M., Maqbool, Amir, Wachkoo, Aijaz Ahmad, Biffi, Gabriel (2021): A review of the Himalayan genus Trypheridium Brancucci (Coleoptera: Cantharidae: Chauliognathinae with description of a new species. European Journal of Taxonomy 764: 18-36, DOI: https://doi.org/10.5852/ejt.2021.764.1467, URL: http://dx.doi.org/10.5852/ejt.2021.764.1467
03BA87A5897A2A23BD012FADFD7AF84C.taxon	description	urn: lsid: zoobank. org: act: DF 4 DAB 14 - 589 A- 4 B 50 - A 37 C- 9 EFC 31438 D 86 Figs 2 – 5	en	Zubair, R. M., Maqbool, Amir, Wachkoo, Aijaz Ahmad, Biffi, Gabriel (2021): A review of the Himalayan genus Trypheridium Brancucci (Coleoptera: Cantharidae: Chauliognathinae with description of a new species. European Journal of Taxonomy 764: 18-36, DOI: https://doi.org/10.5852/ejt.2021.764.1467, URL: http://dx.doi.org/10.5852/ejt.2021.764.1467
03BA87A5897A2A23BD012FADFD7AF84C.taxon	diagnosis	Diagnosis This species resembles T. nuristanicum in general morphology and coloration, but differs in the structure of male and female terminal abdominal segments. In males, the right blade of tergite IX bifurcates terminally into two unequal lobes and tergite X divides into two very long asymmetrical projections. Sternite VII in females projects beyond the tip of abdomen, mostly concealing sternite VIII, and forms three projections apically.	en	Zubair, R. M., Maqbool, Amir, Wachkoo, Aijaz Ahmad, Biffi, Gabriel (2021): A review of the Himalayan genus Trypheridium Brancucci (Coleoptera: Cantharidae: Chauliognathinae with description of a new species. European Journal of Taxonomy 764: 18-36, DOI: https://doi.org/10.5852/ejt.2021.764.1467, URL: http://dx.doi.org/10.5852/ejt.2021.764.1467
03BA87A5897A2A23BD012FADFD7AF84C.taxon	etymology	Etymology The species epithet is in reference to the type locality.	en	Zubair, R. M., Maqbool, Amir, Wachkoo, Aijaz Ahmad, Biffi, Gabriel (2021): A review of the Himalayan genus Trypheridium Brancucci (Coleoptera: Cantharidae: Chauliognathinae with description of a new species. European Journal of Taxonomy 764: 18-36, DOI: https://doi.org/10.5852/ejt.2021.764.1467, URL: http://dx.doi.org/10.5852/ejt.2021.764.1467
03BA87A5897A2A23BD012FADFD7AF84C.taxon	materials_examined	Type material Holotype INDIA • ♂; Jammu and Kashmir, Baramulla, Tangmarg; 34 º 05 ′ 30 ″ N, 74 ° 33 ′ 31 ″ E; 1639 m a. s. l.; 10 Aug. 2018; Zubair R. M. leg.; KUIC 0013. Paratypes INDIA • 1 ♂, 2 ♀♀; same collection data as for holotype; KUIC 0014 to KUIC 0016. Other material INDIA • 1 ♂; Jammu and Kashmir, Baramulla, Tangmarg; 34 º 05 ′ 30 ″ N, 74 ° 33 ′ 31 ″ E; 1639 m a. s. l.; 10 Aug. 2018; Zubair R. M. leg.; KUIC 0017.	en	Zubair, R. M., Maqbool, Amir, Wachkoo, Aijaz Ahmad, Biffi, Gabriel (2021): A review of the Himalayan genus Trypheridium Brancucci (Coleoptera: Cantharidae: Chauliognathinae with description of a new species. European Journal of Taxonomy 764: 18-36, DOI: https://doi.org/10.5852/ejt.2021.764.1467, URL: http://dx.doi.org/10.5852/ejt.2021.764.1467
03BA87A5897A2A23BD012FADFD7AF84C.taxon	description	Description Coloration Head black in vertex and occipital region, yellow in frons and clypeus. Antennae dark brown, except the three and part of fourth basal antennomeres and sometimes underside of latter, yellow; sometimes antennomeres II and III infuscate. Mandibles yellowish with dark-brown apex. Palpi light brown, infuscate at distal palpomeres. Pronotum partly translucent, yellow with transversal black band at posterior half, not reaching lateral and posterior pronotal margins. Elytra sulphur yellow with triangular dark brown patch from base of elytra extending posteriorly along suture up to about posterior third of elytra; integument shinning, covered with fine yellow pubescence. Wings, meso- and metaventrite dark brown to black. Legs: fore and mid coxae yellow, hind coxa partly yellow and partly brown; fore and mid femora yellowish (black in females) except having black patch towards dorsal apex; fore tibia and tarsus yellow; mid tibia testaceous brown and tarsus dark brown; hind legs completely dark brown except yellowish brown basal margin; darker specimens have legs more broadly infuscate. Abdominal ventrites and tergites dark brown to black at the centre and yellow at margins; last ventrites and tergites light to dark brown (especially in females). Male (Figs 2 – 3) HABITUS. Length 7.8 mm in holotype (Fig. 2 A – C), 7.6 mm in paratype. HEAD. Head (including eyes) wider than pronotum (Fig. 2 D – E), 1.3 times as wide as long (across middle of eyes); vertex flat, frons slightly concave between eyes, with pair of longitudinal furrows running between antennae sockets; area around antennal sockets depressed and with carinated margin, forming ridge between antennae that reaches up to vertex; integument smooth and lustrous, with very small punctures and densely covered with short and fine setae. Fronto-clypeal suture defined laterally, indistinct medially. Eyes large, bulging, longer than wide, glabrous; interocular space 1.6 times wider than widest width of eye. Antennae filiform (Fig. 2 F), slender, 0.56 times body length, covered with short and fine setae; antennomere I 1.68 times length of II, 0.88 times length of antennomere III, 0.84 times length of antennomere IV, antennomeres IV – VI longest, subequal, latter decreasing apicad, XI pointed at apex. PRONOTUM. Transverse (Fig. 2 G) 1.3 times as wide as long, sides narrowed posteriorly, rounded, anterior margin arcuate, anterior angles rounded, not defined, posterior margin projected posteriorly, sinuous; disc slightly arched, integument partly translucent, smooth and lustrous, coarsely punctured and densely covered with fine pubescence. Scutellum with sides parallel, apex truncate. Scutellum trapezoidal, with pair of shallow and broad concavities anteriorly, posterior margin with rounded notch. ELYTRA. Short, 0.28 times as long as body length, about 1.89 times length of pronotum, 1.2 times longer than width of both elytra; sides slightly narrowed at posterior third, sutures parallel anteriorly and dehiscent from posterior half, apex of each elytron rounded and flattened; integument slightly rugose, almost smooth, with fine punctures and densely covered with short and fine setae. WINGS. With reduced venation (Fig. 2 H), radial cell 2 R 1 closed; vein r-m barely visible, vein Cu straight; transversal vein cu-a absent; Rr extends beyond the joining of vein r; vein A x 2 well visible, not joining Ax 1 + 2 and ending near the joining of vein A and Ax 1 + 2. LEGS. Slender, increasing in length from fore to hind, without strong modifications. Apex of fore tibia with a pair of very tiny spurs (Fig. 2 I). ABDOMEN. Weakly sclerotised; tergite VIII globose, longer and wider than preceding ones, distal margin arcuate, concave, posterior angles projected, with conspicuous glandular openings; tergite IX with two elongated, asymmetrical and protruding lamellae (blades) that converge towards the apex, giving it a shape of ‘ forceps’ (Fig. 2 J); right blade curves ventrally and bifurcates terminally into two unequal lobes (Fig. 3 A); left blade broad, apex rounded (Fig. 3 C), less sclerotised, almost parallel to abdominal axis; dorsally, tergite IX with pair of glandular openings in apex of short projections, lateral margins rounded, posterior margin slightly arcuate; tergite X large and produced into two unequal, asymmetrical, narrow and elongated lobes (Fig. 3 B – C). Sternite VIII wider than long, ‘ U’ shaped, asymmetrical, forming two narrow and very long asymmetrical lobes (Fig. 3 E), sternite IX internalised, weakly sclerotised, elongate, flanking the aedeagus laterally to left inside abdomen. AEDEAGUS. Elongated and weakly sclerotised (Fig. 3 F – G); left paramere absent, right paramere and prolongation of tegmen directed posteriorly. Right paramere elongated (Fig. 3 H), ventral face with row of setae, apex flattened, forming rounded lobe; left prolongation of tegmen broad and lamellar (Fig. 3 J), margins sinuous, apex rounded with tip pointing laterally. Median lobe membranous, short, not twisted, with its broad opening located apically. Female (Fig. 4 – 5) HABITUS. Length 7.4 – 7.6 mm (n = 2) (Fig. 4 A – C). Similar to male, except for eyes slightly smaller (Fig. 4 D – E), interocular distance across base of antennae 1.16 times wider than in male; elytra 0.32 times body length, 1.88 times the pronotum length; sternite VII variable (Fig. 4 G – H), long and projecting posteriorly into 3 flattened lobes: 2 shorter pointed lobes laterally and medial lobe broad, parallel-sided, distal margin, with rounded notch (Fig. 5 A); sometimes lateral lobes longer, almost as long as medial lobe, and distal margin of medial lobe truncate, not notched. Sternite VIII (Fig. 5 C) almost completely concealed by sternite VII, membranous, broadly rounded, distal margin with broad rounded lobe projecting posteriorly; tergite VIII rounded on sides basally with wide and deep notch apically forming two apical lobes; coxites small (Fig. 5 E), not fused medially, anterior arms very long and slender, apparently fused with valvifers, styles elongate, digitiform. Genitalia membranous (Fig. 5 F); vagina long and broad; bursa copulatrix elongate, wrinkled, spiralling; spermatheca formed by two tubular projections; accessory gland large, connected by long winding duct to the postero-dorsal part of bursa copulatrix. Biological notes	en	Zubair, R. M., Maqbool, Amir, Wachkoo, Aijaz Ahmad, Biffi, Gabriel (2021): A review of the Himalayan genus Trypheridium Brancucci (Coleoptera: Cantharidae: Chauliognathinae with description of a new species. European Journal of Taxonomy 764: 18-36, DOI: https://doi.org/10.5852/ejt.2021.764.1467, URL: http://dx.doi.org/10.5852/ejt.2021.764.1467
03BA87A5897A2A23BD012FADFD7AF84C.taxon	discussion	The specimens were collected on walnut tree leaves (Juglans regia L.) infested with walnut aphids, Chromaphis juglandicola (Kaltenbach, 1843) and Panaphis juglandis (Goeze, 1778).	en	Zubair, R. M., Maqbool, Amir, Wachkoo, Aijaz Ahmad, Biffi, Gabriel (2021): A review of the Himalayan genus Trypheridium Brancucci (Coleoptera: Cantharidae: Chauliognathinae with description of a new species. European Journal of Taxonomy 764: 18-36, DOI: https://doi.org/10.5852/ejt.2021.764.1467, URL: http://dx.doi.org/10.5852/ejt.2021.764.1467
03BA87A5897A2A23BD012FADFD7AF84C.taxon	distribution	Distribution India (Baramulla, Kashmir) (Fig. 8 A – B).	en	Zubair, R. M., Maqbool, Amir, Wachkoo, Aijaz Ahmad, Biffi, Gabriel (2021): A review of the Himalayan genus Trypheridium Brancucci (Coleoptera: Cantharidae: Chauliognathinae with description of a new species. European Journal of Taxonomy 764: 18-36, DOI: https://doi.org/10.5852/ejt.2021.764.1467, URL: http://dx.doi.org/10.5852/ejt.2021.764.1467
03BA87A589752A24BEFC2B59FC4BFE13.taxon	description	Figs 6 – 7	en	Zubair, R. M., Maqbool, Amir, Wachkoo, Aijaz Ahmad, Biffi, Gabriel (2021): A review of the Himalayan genus Trypheridium Brancucci (Coleoptera: Cantharidae: Chauliognathinae with description of a new species. European Journal of Taxonomy 764: 18-36, DOI: https://doi.org/10.5852/ejt.2021.764.1467, URL: http://dx.doi.org/10.5852/ejt.2021.764.1467
03BA87A589752A24BEFC2B59FC4BFE13.taxon	diagnosis	Diagnosis Trypheridium nuristanicum differs from T. kashmiricum sp. nov. in the structure of the male and female terminal abdominal segments. In males, the right blade of tergite IX is undivided terminally and tergite X is divided into two short asymmetrical lobes. Sternite VII in females is short, partly exposing sternite VIII, with 2 projections at the distal margin.	en	Zubair, R. M., Maqbool, Amir, Wachkoo, Aijaz Ahmad, Biffi, Gabriel (2021): A review of the Himalayan genus Trypheridium Brancucci (Coleoptera: Cantharidae: Chauliognathinae with description of a new species. European Journal of Taxonomy 764: 18-36, DOI: https://doi.org/10.5852/ejt.2021.764.1467, URL: http://dx.doi.org/10.5852/ejt.2021.764.1467
03BA87A589752A24BEFC2B59FC4BFE13.taxon	materials_examined	Material examined (from photographs) Paratypes AFGHANISTAN • 1 ♂; Nuristan, Baschgaltal; 1200 m a. s. l.; 15 Apr. 1953; Wittmer W. leg.; [Klapperich J., Afghanistan exped.]; “ Paratype; CANTHARIDAE, CANTH 00014269, [QR Code label] ”; NHMB • 1 ♂, 4 ♀♀, same collection data as for preceding; det. Wittmer W.; “ Paratype; Trypherus nuristanicus Wittm.; CANTH 00013839; [QR Code label] ”; NHMB • 1 ♂; Nuristan, Baschgaltal; 1100 m a. s. l.; 15 Apr. 1953; Wittmer W. leg.; “ Klapperich J., Afghanistan exped. ”; NHMB.	en	Zubair, R. M., Maqbool, Amir, Wachkoo, Aijaz Ahmad, Biffi, Gabriel (2021): A review of the Himalayan genus Trypheridium Brancucci (Coleoptera: Cantharidae: Chauliognathinae with description of a new species. European Journal of Taxonomy 764: 18-36, DOI: https://doi.org/10.5852/ejt.2021.764.1467, URL: http://dx.doi.org/10.5852/ejt.2021.764.1467
03BA87A589752A24BEFC2B59FC4BFE13.taxon	description	Re-description (partly adapted from Wittmer 1956 and Brancucci 1985 a) Coloration Head black in the vertex and occipital region, yellow in the frons and clypeus. Antennae dark brown, except the three or four first antennomeres and sometimes the underside of the later, yellow. Palpi light brown, infuscate at distal palpomeres. Pronotum yellow with a transversal black band at posterior half, nearly reaching the lateral and posterior pronotal margins. Elytra sulphur yellow, sometimes lighter at the apex; a triangular dark brown patch from the base of elytra extending posteriorly along the suture up to about posterior third of elytra. Wings, meso- and metaventrite dark brown to black. Legs usually black; fore tibia and base of femur sometimes testaceous to light brown, fore femur rarely completely testaceous; bases of mid femur and tibia sometimes lighter. Abdominal ventrites and tergites dark brown to black at the centre and yellow at the margins; last ventrites and tergites mostly to completely yellow. Coloration in females, especially of the legs, usually a little darker. Male (Fig. 6) HABITUS. Length 6.5 – 8.5 mm (Fig. 6 A). HEAD. Head (including the eyes) wider than pronotum; vertex flat, frons slightly concave between eyes, with fine longitudinal furrow in middle, which ends in semicircular impression between antennae sockets; integument smooth and lustrous, with very small punctures and densely covered with short and fine setae. Eyes large, bulging; interocular space slightly wider than widest width of eye. Antennae filiform, slender, reaching apex of elytra posteriorly, covered with short and fine setae; antennomere I slender, II more than a third shorter than III, antennomeres IV and V subequal and following decreasing in length. PRONOTUM. Transverse (Fig. 6 B) 1.1 times as wide as long, sides narrowed posteriorly, anterior margin regularly arched, anterior angles barely defined, posterior margin projected posteriorly, truncate at middle and concave laterally; disc slightly arched, integument smooth and lustrous, coarsely punctured and densely covered with fine pubescence. Scutellum with sides parallel, apex truncate. ELYTRA. Short, 1.4 times longer than width of both elytra; sides slightly narrowed at posterior third, sutures parallel anteriorly and dehiscent from posterior third, apex of each elytron rounded and flattened; integument slightly rugose, almost smooth, with fine punctures and densely covered with short and fine setae. LEGS. Slender, increasing in length from fore to hind, without strong modifications. Apex of fore tibia with pair of tiny spurs (Fig. 6 C). ABDOMEN. Weakly sclerotised; tergite VIII globose (Fig. 6 D), longer and wider than preceding ones, distal margin arcuate, concave, posterior angles with conspicuous glandular openings; tergite IX modified into capsule with two elongate, asymmetrical and protruding lamellae (blades) that converge towards apex (Fig. 6 E – F): left blade curved ventrally and posteriorly, long, flattened, weakly sclerotised, apex broadly rounded; right blade shorter, more strongly curved ventrally, apex rounded, undivided; tergite X strongly narrowed (Fig. 6 F), longer than wide, split at base into two asymmetrical lobes projected posteriorly, one of lobes slightly shorter than other. Sternite VIII asymmetrical, distal margin deeply notched, sinuous, with sides projected posteriorly; sternite IX lamellar (Fig. 6 F), weakly sclerotised, concealed underneath tergites VII and VIII. AEDEAGUS. Elongate, weakly sclerotised (Fig. 6 G – I); left paramere absent, right paramere and prolongation of tegmen directed posteriorly. Right paramere elongated, ventral face with row of setae, apex flattened, forming rounded lobe; left prolongation of tegmen broad and lamellar, margins sinuous, apex truncate with tip pointing laterally, outer margin with fringe of setae. Median lobe membranous, short, not twisted, opening broad, located apically. Female (Fig. 7) Similar to males, except for eyes smaller, less protruding, interocular space 1.4 times larger than widest diameter of eye; antennae shorter, not reaching apex of elytra posteriorly. Sternite VII strongly and variably modified, elongate, projected distally into pair of forked lamellar projections, parallel to slightly divergent, apices rounded, sides of projections angulate at base or near mid length (Fig. 7 B); sternite VIII subquadrate, broad and long, partly concealed by sternite VII, lateral margins arched, distal margin with rounded projecting lobe in middle tergite VIII convex, lateral margins slightly arched, distal margin broadly arched, concave, distal angles projected posteriorly, with glandular openings; tergite IX short and broad, distal margins slightly arched, concave (Fig. 7 C). Genitalia membranous (Fig. 7 D); vagina long and broad; bursa copulatrix elongate, wrinkled, spiralling; spermatheca formed by two tubular projections; accessory gland large, connected by long winding duct to the postero-dorsal part of bursa copulatrix.	en	Zubair, R. M., Maqbool, Amir, Wachkoo, Aijaz Ahmad, Biffi, Gabriel (2021): A review of the Himalayan genus Trypheridium Brancucci (Coleoptera: Cantharidae: Chauliognathinae with description of a new species. European Journal of Taxonomy 764: 18-36, DOI: https://doi.org/10.5852/ejt.2021.764.1467, URL: http://dx.doi.org/10.5852/ejt.2021.764.1467
03BA87A589752A24BEFC2B59FC4BFE13.taxon	discussion	Remarks Brancucci (1985 a) proposed Trypheridium nuristanicus (sic) as a new combination in his description of the neuter genus Trypheridium, of which it is the type species. He omitted to adjust the gender of the species name to agree with Trypheridium. The gender of nuristanicus (an adjective formed from the name of the type locality Nuristan in Afghanistan) is emended to nuristanicum to agree with the neuter Latin suffix - idium of Trypheridium (ICZN 1999, Art. 34.2). When proposing the genus, LeConte (1851) did not give an etymology for Trypherus, but Gemminger & Harold (1869) defined the name as meaning “ debilis ” (week, feeble, fragile, etc). “ Trypheridium ” is a diminutive, a ‘ little Trypherus ’.	en	Zubair, R. M., Maqbool, Amir, Wachkoo, Aijaz Ahmad, Biffi, Gabriel (2021): A review of the Himalayan genus Trypheridium Brancucci (Coleoptera: Cantharidae: Chauliognathinae with description of a new species. European Journal of Taxonomy 764: 18-36, DOI: https://doi.org/10.5852/ejt.2021.764.1467, URL: http://dx.doi.org/10.5852/ejt.2021.764.1467
03BA87A589752A24BEFC2B59FC4BFE13.taxon	distribution	Distribution Tajikistan; Afghanistan (Nuristan); Pakistan (Swat) (Fig. 8 A).	en	Zubair, R. M., Maqbool, Amir, Wachkoo, Aijaz Ahmad, Biffi, Gabriel (2021): A review of the Himalayan genus Trypheridium Brancucci (Coleoptera: Cantharidae: Chauliognathinae with description of a new species. European Journal of Taxonomy 764: 18-36, DOI: https://doi.org/10.5852/ejt.2021.764.1467, URL: http://dx.doi.org/10.5852/ejt.2021.764.1467
