taxonID	type	description	language	source
03E687D65B46DE3AFF10EE53A243FF49.taxon	type_taxon	Type species: Cardiodactylus novaeguineae (Haan, 1842)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B46DE3AFF10EE53A243FF49.taxon	diagnosis	Diagnosis. Among Eneopterinae and Lebinthini genera, Cardiodactylus is characterized by its average to large size and its long wings in both sexes, while the other genera of the tribe are brachypterous (Robillard 2011 b) (Fig. 1). Eyes large and prominent, fastigium narrower than in Lebinthus and Agnotecous, as in Centuriarus. Face slightly higher than wide. Dorsal disk of pronotum trapezoidal, posterior margin straight or slightly bisinuated, especially in Efordi group. Legs long. TI with 2 tympana; inner tympanum covered by a sclerotized expansion, its membrane visible along a small longitudinal slit only; outer tympanum ellipsoidal, transversally plicate. TI with 2 inner and 1 outer apical spurs. TII with 2 inner and 2 outer spurs. FIII strong basally, filiform at apex. TIII serrulated on their whole length, furrowed longitudinally on their dorsal side and with 4 pairs of subapical spurs and 3 pairs of apical spurs; inner spurs long and curved, outer spurs short and straight. Tarsomeres III- 1 with 2 dorso-apical spines and a row of spines on outer dorsal edge; rarely with lateral outer spines. Wings longer than abdomen, hind wings longer than forewings, forming a tail of variable length. Cerci thin, longer than FIII. Male. Metanotal glands absent. Stridulum complete (Fig. 2): Harp longer than wide, with 1 – 2 w-shaped harp veins (obliques), delimiting a raised triangular area; posterior area of harp raised along the diagonal vein. In Novaeguineae group, FW coloration pattern usually more contrasted than in Efordi group, posterior angle of harp whitish and anterior region translucent, with different yellow or whitish areas around chords and posterior to mirror. Mirror not rounded but differentiated from other cells of D alignment; d 2 cell wide. CuP generally absent. Apical field involving 3 or more cell alignments posterior to the mirror. FW lateral field: Sc with several projections along its length; with variable number of more ventral veins. Subgenital plate clog-shaped; inner side with paired subapical swellings.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B46DE3AFF10EE53A243FF49.taxon	discussion	Male genitalia. (Fig. 3) Pseudepiphallus [epiphallus according to Randell’s (1964) terminology] setose laterally and ventrally, narrowed at mid-length, forming lateral shoulders delimiting the anterior part, which is generally wider than the posterior part, characterized by the presence of a median process of variable size and shape, with pair longitudinal dorsal ridges developed in Novaeguineae species group, generally independent from each other and sometimes asymmetrical, but fused together in C. novaeguineae and a few closely related species. Pseudepiphallic median process most often terminated posteriorly by a spoon-shaped apex more or less rounded, with a rounded translucent region. Structure including the dorsal ridges and the spoon-like apex always present in the genus, also present, although less developed in most species of Agnotecous (except a few species in Yahoue species group), but lacking in most Lebinthus species and in Centuriarus. Pseudepiphallic parameres [ectoparameres] with characteristic trilobate shape, the posterior lobe narrow and oriented externally. Anterior region of pseudepiphallic sclerite terminated with latero-anterior expansions usually triangular, the membrane between them setose in Novaeguineae species group, except in C. novaeguineae where these expansions delimit a membranous sac (Robillard & Ichikawa 2009). In Efordi group, the whole anterior region of pseudepiphallus is sclerotized and constitutes a wide dorsal concavity. Rami generally strong, with a wide preapical plate and a thin apical stem. Ectophallic apodemes [endoparameral apodemes] parallel and long, their base sometimes with anterior and posterior expansions. Ectophallic arc [endoparameres] well sclerotized, generally curved. Ectophallic fold [rachis] short, with thin lateral sclerites, apex trilobate, the median lobe sponge-like. Endophallic sclerite [formula] comprising a short median sclerotization and 2 lateral arms of variable size; endophallic apodeme [apodeme of formula] most often with a dorsal crest and lateral lamellas. Female. FWs with 7 – 10 strong longitudinal veins. Sc with several projections along its length; with variable number of more ventral veins. FW coloration variable across species, usually brown with patterns of yellow patches. Ovipositor laterally flattened, apex lanceolate (see Fig. 13), often denticulate on dorsal edge as in Agnotecous and Lebinthus species; denticulation also on ventral edge of ovipositor apex in most species of Novaeguineae group.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B46DE3AFF10EE53A243FF49.taxon	description	Female genitalia (see Fig. 14). Copulatory papilla with a wide ovoid basal part with pair basal sclerites and a narrowed apex generally rounded. Classification Species group Novaeguineae Otte, 2007 a	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B46DE3AFF10EE53A243FF49.taxon	diagnosis	Diagnosis. Species of average to large sizes, usually with more contrasted and vivid coloration patterns on FWs and pronotum, and shorter hind wings in both sexes than in Efordi group. Male genitalia with dorsal pseudepiphallic longitudinal ridges variable amongst species, but generally forming a wide gutter with a rounded spoon-shaped posterior apex; anterior apex of pseudepiphallus generally with pair triangular expansions surrounding a setose membrane. Species group Efordi Otte, 2007 a Diagnosis. Species of small to average sizes, more slender and usually with little contrasted brownish coloration, and longer hind wings in both sexes than in Novaeguineae group. Male genitalia with a very narrowed posterior part variable amongst species; anterior apex of pseudepiphallus sclerotized and constitutes a wide dorsal concavity.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B46DE3AFF10EE53A243FF49.taxon	discussion	List and distribution of Cardiodactylus species per island [country] in the region under study (Fig. 4) All the species in the region around the Wallace line (west of New Guinea Island) belong to the Novaeguineae species group. Species in the studied area but not directly considered here are mentioned in brackets, see corresponding references for more details.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B4ADE3DFF10EDC0A181FBE9.taxon	description	(Figs 4, 5 A, 7, 8, 9 A, 11 A, 13 A, 14 A)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B4ADE3DFF10EDC0A181FBE9.taxon	materials_examined	Type material. Male holotype: Indonesia. Central Kalimantan Province, Midden O. Borneo Exp. Marah [river], 10 – 28. XI. 1925, # 208 [?], H. C. Siebers (MNHN-EO-ENSIF 3497). Female allotype: same information as HT, # 200 (MNHN-EO-ENSIF 3498). Paratypes (12 ♂, 12 ♀): 2 ♂, same information as HT, # 204, # 208 (MNHN-EO-ENSIF 3499 - 3500). Midden O. Borneo Exp., no precision, H. C. Siebers: 1 ♂, 17. XI. 1925, # 200 (MNHN-EO-ENSIF 3502); 1 ♂, 22. IX. 1925, # 117 (MNHN-EO-ENSIF 3503); 1 ♂, 29. IX. 1925, # 135 (MNHN-EO-ENSIF 3501); 1 ♂, 23. XI. 1925, # 206 (MNHN-EO-ENSIF 3504); 1 ♀, 10. XI. 1925, # 105 (MNHN-EO-ENSIF 3505); 1 ♂, 23. IX. 1925, # 121; 1 ♀, 22. XI. 1925, # 205 (ZIN). Midden O. Borneo Exp., Long Petak, 450 m, 1 ♀, IX. 1925, H. C. Siebers (MNHN-EO-ENSIF 3506). East Kalimantan Province, Kec. Pujungan, Kayan-Mentarang Nat. Reserve, IIS 930549, lowland diptero forest, WWF station, Lalut Birai, base camp, Kebun, 355 m, 2 ° 52 ’ N 115 ° 49 ’ E, X. 1993, malaise trap head, DC. D. Rosichon U.: 1 ♂ (MZB-ORTH 12194); 1 ♀ (MZB-ORTH 12238). Bukit Soeharto Expt. For. 60 km S. Samarinda IIS 930613, 60 m, 0 ° 42 ’ S 117 ° 0 ’ E, lowland dipterocarp forest, primary rainforest, malaise trap head, XII. 1993, DC. D. Rosichon U.: 1 ♀ (MZB-ORTH 12195). Malaysia. South East Sabah, nr. Danum Valley Field C., c. 150 m, W 0, mal. trap 5, C. v. Achterberg & D. Kennedy: 26. V – 20. VI. 1987, 1 ♀; 20. VI – 12. VII. 1987, 1 ♂; 2 – 23. VIII. 1987, 1 ♀; 26. X – 22. XI. 1987, 1 ♂, 1 ♀; 5. XII. 1987 - 20 - I. 1988, 1 ♂; 22. XI- 4. XII. 1987, 1 ♀ (RMNH). nr. Danum Valley Field C., c. 150 m, W 0, mal. trap 11, C. v. Achterberg: 14 – 20. III. 1987, 1 ♀ (RMNH). North Sabah, Mangalum Island, 35 m, NW Jesselton, 8 – 15. VII. 1928, 1 ♂, identified Cardiodactylus novaeguineae Haan by L. Chopard, C. Boden Kloss (MNHN-EO-ENSIF 3507). Borneo, no precise locality, 1891, 2 ♀ (# 1269, -), Chaper (MNHN-EO-ENSIF 3508 - 3509). Malaysia (Borneo), Sabah, Crocker Range, 80 km S of Kota-Kinabalu City, environs of Ulu Kumanis Vill., 800 m, 5 – 10. V. 2009, A. Sotshivko, 1 ♂ (ZIN); Crocker Range National Park not far from Keningau Town, 1000 – 1300 m, secondary-primary forest, 2 – 6. V. 2013, A. Gorochov, M. Berezin, V. Gorochova, E. Tkatsheva, 1 ♀ (ZIN); ~ 5 km NWW of Kota-Kinabalu City, Manukan I., sea level, secondary forest, 24 – 25. IV. 2013, A. Gorochov, M. Berezin, V. Gorochova, E. Tkatsheva, 1 ♂, 1 ♀ (ZIN); Sandakan Division, environs of Sukau Vill. on Kinabatangan River (~ 35 km from its mouth), ~ sea level, secondary-primary forest, 8 – 13. V. 2013, A. Gorochov, M. Berezin, E. Tkatsheva, 2 ♂ (ZIN). Type locality. Indonesia, Borneo, Central Kalimantan, Marah [river]. Other material examined. Malaysia. Pahang, Tioman I. not far from Mersing City in Johor State (Malacca Peninsula), environs of Juara Vill. on eastern coast of island, disturbed primary forest, on leaf of bush, at night, 6 – 14. IV. 2010, A. Gorochov, M. Berezin, E. Tkatsheva, 3 ♂ (ZIN). Borneo, no presicion, C. Pictet, 1 ♀, identified Cardiodactylus novae-guineae (MHNG).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B4ADE3DFF10EDC0A181FBE9.taxon	etymology	Etymology. Named after the type locality.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B4ADE3DFF10EDC0A181FBE9.taxon	distribution	Distribution. Eastern, northern and central Borneo; Tioman Island.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B4ADE3DFF10EDC0A181FBE9.taxon	diagnosis	Diagnosis. Species of average size, general coloration brown and yellow brown, little contrasted, characterized by male FW venation with mirror separated in 2 sub-equal parts, and by shape of pseudepiphallic sclerite in male genitalia. Species very close to C. thailandia and C. admirabilis by shape of male genitalia (asymmetrical dorsal ridges of pseudepiphallus) and male FW venation, but differing by smaller size and coloration less contrasted (brown and yellow in C. borneoe, black and yellow in C. thailandia; diagonal vein including a black area in C. thailandia, absent in C. borneoe).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B4ADE3DFF10EDC0A181FBE9.taxon	description	Description. Average size for the species group. General coloration brown, little contrasted (Fig. 7 A – D). Head dorsum yellow brown with 4 wide dark brown bands; lateral bands complete except for a separate spot anteriorly; median bands punctuated; 2 short dark brown lines posterior to eyes (Fig. 5 A). Fastigium black or dark brown. Scapes yellow brown with a dark brown ring; antennae brown with yellow brown rings. Face variable, most often yellow brown with a yellow transverse band below eyes, 2 parallel stripes on front head and 2 small black dots bellow antennae. Mouthparts mostly yellow brown. Lateral sides of head brown. Maxillary palpi yellow brown, apex dark brown. Pronotum: Dorsal disk lateral margins yellow to whitish, with dark spots; median area yellow brown, with brown and black symmetrical patterns; posterior margin dark brown. Lateral lobes dark brown to black, ventral margins yellow. Legs I – II yellow brown, with dark brown spots on femora and rings on tibiae; tarsomeres I – III yellow brown, their ends dark brown. FIII yellow brown mottled with brown, knees black; TIII with yellow brown and dark brown rings. Hind wing tail black, 1.5 times longer than pronotum. Cerci yellow brown with dark brown spots basally and dark brown rings apically. Posterior tergites dark brown with lateral discontinuous yellow lines; posterior sternites yellow brown. Male: FW coloration mostly dark brown, some areas with yellowish or whitish venation (Fig. 9 A). Orange brown sclerotized area anterior to chords. Cell e 1 dark brown. Yellow areas include all veins anterior to harp, i. e. bases of anal veins, harp veins and anterior half of the diagonal vein; large oval area posterior to mirror with translucent cells. M / R / Sc area orange brown to dark brown; posterior part of Sc whitish; projections of Sc and ventral veins whitish, cells between them dark brown. FW venation (Fig. 9 A): 1 A slightly bisinuated. CuP missing. Diagonal vein homogeneously yellow brown (with a black median area in C. thailandia). Stridulatory file with 200 – 250 teeth on the transverse and longitudinal parts of 1 A, without discontinuity at the level of the angle, and ca. 30 more teeth on a bump near 1 A base (see Table 1, n = 3). Harp with a strong w-shaped vein and a faint anterior one. Mirror area: mirror (d 1) longer than wide, oval, separated in two sub-equal parts by a transverse vein; d 2 wide, prolonging mirror shape; e 1 variable, crossed by several accessory veins in HT. Apical field with 3 – 5 cell alignments posterior to mirror. Lateral field with 7 – 9 (m = 7.8, n = 5) projections of Sc and 4 – 5 (m = 4.6, n = 5) ventral veins. Subgenital plate yellow brown. Male genitalia (Fig. 7 E – G): Pseudepiphallus elongate, clearly narrowed twice, at mid-length and preapically. Dorsal ridges divergent posteriorly, their dorsal edges bean-like, slightly asymmetrical, the left one curved, the right one straight; pseudepiphallus bisinuated laterally. Pseudepiphallic sclerite with wide latero-anterior expansions pointed at apex, membrane between them with thin setae. Posterior region rounded, narrower than preapical part, apex with a small translucent area longer than wide. Rami with large preapical plates, indented on dorsal margin; apex slightly convergent. Ectophallic arc complete, wide and curved, with a short posterior expansion. Ectophallic apodemes as long as the rami, straight and parallel, their bases with a posterior membranous lobe and a long anterior sclerotization along the membrane. Ectophallic fold membranous, with weak lateral sclerites; apex trilobate, membranous, median lobe oval. Endophallic sclerite little sclerotized except lateral arms; with a wide posterior expansion. Endophallic apodeme with wide lateral lamellas and a strong medio-dorsal crest. Membrane of endophallic cavity smooth. Female: FW coloration (Fig. 11 A): mostly dark brown, veins light brown in anterior region, bases of anal veins whitish to yellow brown; dark brown area near base of CuA, CuP and 1 A. Anterior half of CuA / M area whitish, including transverse veins and associated sclerotization; posteror part dark brown. R / Sc area orange brown including veins. Projections of Sc and more ventral veins yellow brown, the cells between them dark brown. FW venation: 10 (n = 5) strong longitudinal veins on dorsal field; lateral field with 8 – 11 (n = 5) longitudinal veins including 4 – 7 Sc projections and 4 – 5 ventral veins. Ovipositor: slightly shorter than FIII; apex with both dorsal (slightly) and ventral edges denticulate (Fig. 13 A). Female genitalia (Fig. 14 A): Copulatory papilla trapezoidal with baso-lateral sclerites; apex rounded, slightly sclerotized and folded ventrally. Juvenile: Unknown. Variation: The species shows an important array of variation in details of coloration (specimens more or less dark or light) and in male genitalia: the length of the posterior part of the pseudepiphallus is slightly variable; in profile view, the dorsal edge of the pseudepiphallic ridges are variably indented, rounded or flat (Fig. 8). In particular, males from Tioman Island show a little indented genitalic profile with a flat dorsal edge (Fig. 8 C). Further studies including acoustic and molecular analyses will be necessary to test whether some of these populations may constitute separate species. Measurements. See Table 1. Habitat and life history traits. Unknown. Behavior. Unknown.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B59DE2AFF10EE2CA181F99C.taxon	description	(Figs 4, 5 B, 13 B, 14 B, 16)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B59DE2AFF10EE2CA181F99C.taxon	materials_examined	Type material. Male holotype: Indonesia. Maluku Province, Boeroe [Buru I.], 1921, Estrin & Smotritskaya (ZIN). Paratypes (5 ♂, 7 ♀): same information as HT, 1 ♂, 1 ♀ (ZIN). Buru, L. J. Toxopeus: Station 1 A, 1921, 1 ♂ (MZB-ORTH 10623), 1 ♂, MZB-ORTH 1826 (MNHN-EO-ENSIF 3153), 3 ♀ (MZB-ORTH 1827, 10622, 10586), 1 ♀, MZB-ORTH 10621 (MNHN-EO-ENSIF 3152); Station 4, 860 m, 29 – 31. I. 1922, 1 ♀ (MZB-ORTH 1836); Leksoela [?], V-VII- 1921, 1 ♀ (MZB-ORTH 1813); Station, IX. 1921, 1 ♂, E. Estrin (MZB-ORTH 1800); Station 1, IV-IX. 1921, 1 ♂ (MZB-ORTH 1811). Type locality. Indonesia, Buru Island. Other material examined. Indonesia. Maluku Province, Aboina [Ambon I.], 1859, Dr. Doleschal: 1 ♂, identified Eneoptera varia Br. (NHMW); 1 ♂, identified Pl. [Platydactylus] varius Br. (NHMW).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B59DE2AFF10EE2CA181F99C.taxon	etymology	Etymology. Named after type locality.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B59DE2AFF10EE2CA181F99C.taxon	distribution	Distribution. Indonesia, Buru Island and Ambon Island.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B59DE2AFF10EE2CA181F99C.taxon	diagnosis	Diagnosis. The new species is very close to C. muiri Otte, 2007 a but differs in distinctly more rounded subbasal projections of the dorsal epiphallic ridges and lateral field of tegmina brown (yellow in C. muiri); C. buru differs from C. halmahera by epiphallic ridges practically symmetrical; and from all the other species consired here in the subbasal projections of dorsal epiphallic ridges clearly curved aside.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B59DE2AFF10EE2CA181F99C.taxon	description	Description. Male (holotype): Species of average size. Coloration yellowish with following marks: face of head with light brown spot under each antennal cavity and a pair of small brown spots on rostral apex; head dorsum brown with a pair of slight lighter longitudinal marks between eyes and narrow yellowish stripe outlined anterior part of dorsum (Fig. 16 A – B); scape with brown medial spot; antennal flagellum light brown; pronotum brown with narrow light brown stripes along ventral edge of each lateral lobe and along lateral edges of disk (latter stripes interrupted at middle) (Fig 16 C); tegmina brown with 4 yellowish white spots on dorsal field, a few veins in region of chords, branches of Sc and crossveins between them, and with yellowish some oblique veins (membranes between Sc branches almost light brown, and spots at base of chords and near distal part of chords almost reddish brown) (Figs 16 D – E); exposed part of hind wings brownish gray with venation almost same color; legs uniformly light brown with only hardly darker apical part of hind femora and base of hind tibiae; abdomen with weakly spotted light brown tergites and anal plate as well as slightly darkened distal part of genital plate. Apical area of tegmina long, mirror slightly narrower than in C. empagatao (almost as in C. halmahera n. sp.), comb of tegminal Sc with 9 – 10 branches, and male genitalia with subbasal projections of dorsal epiphallic ridges distinctly more rounded and less strongly curved laterally (Figs 16 H – K). Female: General appearance similar to that of holotype, but face with light brown area between light brown spots under antennal cavities; tegmina with 2 yellowish white spots in dorsal field (basal spot and lateral one), connected with each other by narrow light humeral stripe, and light venation (excepting brown R, stem of Sc, and crossveins between them), as well as with 12 – 13 longitudinal veins in dorsal field, 7 – 8 branches of Sc, and much more numerous crossveins (especially in lateral field) (Figs 16 F – G). Ovipositor quite short (hind femur almost 1.2 times as long as ovipositor), its apex smooth on both dorsal and ventral edges (Fig. 13 B). Female genitalia (Fig. 14 B): Copulatory papilla triangular, with thin baso-lateral sclerites; apex rounded and slightly sclerotized, slightly folded ventrally. Juvenile: Unknown. Variation. Some male paratypes with slightly darker most part of head dorsum and almost without crossveins between Sc branches. Anterior part of the epiphallic ridges also variable in shape, slightly asymmetrical in some males. Measurements. Length in mm. Body: male 22 – 24, female 25; body with wings: male 29 – 31, female 33.5; pronotum: male 3.1 – 3.5, female 3.9; tegmina: male 17 – 18, female 20; hind femora: male 17 – 19.5, female 21; ovipositor 18. Habitat and life history traits. Unknown. Behavior. Unknown.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B5BDE2CFF10EE18A181FBE9.taxon	description	(Figs 4, 7, 8, 9 B, 17)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B5BDE2CFF10EE18A181FBE9.taxon	materials_examined	Type material. Male holotype: Indonesia. Celebes [Sulawesi], Pala ?? [Palapa or Palaga?], identified Cardiodactylus novae guineae Haan by L. Chopard (MZB-ORTH 10450). Type locality. Indonesia, Sulawesi.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B5BDE2CFF10EE18A181FBE9.taxon	etymology	Etymology. Named after the type locality.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B5BDE2CFF10EE18A181FBE9.taxon	distribution	Distribution. Indonesia, Sulawesi.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B5BDE2CFF10EE18A181FBE9.taxon	diagnosis	Diagnosis. Species of average size, coloration brownish with yellow brown patterns, characterized by male FW venation and coloration including a thin yellow line posterior to mirror, and by shape of pseudepiphallic and endophallic sclerites in male genitalia. Species close to C. fruhstorferi n. sp. from which it differs by the length of pseudepiphallus.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B5BDE2CFF10EE18A181FBE9.taxon	description	Description. Average size for the species group. General coloration little contrasted, brown with yellow areas. Head dorsum with 4 wide dark brown bands little distinct anteriorly, bordered by dark brown lines posterior to eyes (Fig. 17). Fastigium dark brown. Scapes orange brown; antennae with yellowish rings. Face and mouthparts almost homogeneously orange brown, slightly lighter ventral to eyes. Maxillary palpi brown, darker apically. Pronotum: Dorsal disk yellow brown laterally, median area mottled with dark brown. Lateral lobes dark brown, ventral margin yellow. Fore legs yellow brown, with faint dark spots on femur. Median legs and tarsomeres III missing in HT. Legs III homogeneously brown. Hind wing tail dark brown, 1.5 longer than pronotum. Cerci and apex of abdomen damaged in HT. Male: FW coloration contrasted (Fig. 9 B), mostly orange brown with yellowish and dark brown areas. Yellow areas include: bases of vein CuA, M and 2 A – 4 A, angle of 1 A, chords, harp veins, and a narrow transverse line posterior to mirror. Dark brown areas include region anterior to stridulatory file and external part of e 1. Lateral field mostly brown, including posterior part of M / R area; posterior part of R / Sc area yellowish. Projections of Sc yellow, cells between them brown. FW venation (Fig. 9 B): 1 A not clearly bisinuated. CuP missing. Stridulatory file with 166 teeth on the transverse part of the file, and 56 on the longitudinal part, without discontinuity at the level of 1 A angle. Harp with 2 w-shaped harp veins and several anterior accessory veins. Mirror area: mirror (d 1) longer than wide, not rounded but well defined, separated in two parts by a strong transverse vein, the posterior part small and triangular; d 2 wide, well defined, prolonging the mirror, without accessory vein; e 1 crossed by an accessory vein. Apical field with 4 cell alignments posterior to mirror. M / R fusion angle acute. Lateral field with 7 projections of Sc and 3 more ventral veins. Subgenital plate color not distinct in HT. Male genitalia (Fig. 17 C – E): Pseudepiphallus short, clearly narrowed twice, at mid-length and preapically. Dorsal ridges parallel, forming a narrow gutter. Pseudepiphallic sclerite with wide latero-anterior expansions, membrane between them with thick short setae. Posterior region rounded, widened laterally, with a small rounded translucent area at apex. Apex of rami convergent. Ectophallic arc complete, with a short posterior expansion. Ectophallic apodemes long and wide, their basis with long anterior expansions and short posterior ones. Ectophallic fold with lateral sclerites, apex trilobate, median lobe elongate. Endophallic sclerite narrow, its posterior arms fused with lateral sclerites of ectophalic fold. Endophallic apodeme with both lateral lamellas and a medio-dorsal crest. Membrane of endophallic cavity smooth. Female: Unknown. Juvenile: Unknown. Measurements. See Table 2. Habitat and life history traits. Unknown. Behavior. Unknown.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B5DDE2EFF10EE43A181FA54.taxon	description	(Figs 4, 5 C, 18)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B5DDE2EFF10EE43A181FA54.taxon	materials_examined	Type material. Male holotype: Indonesia. Sulawesi Tengah Province, Sulawesi I., Lore Lindu National Park, ~ 75 km SE of Palu City, environs of Wuasa Vill. (near eastern edge of park), ~ 1000 m, secondary forest, on large leaf of grassy plant, at night, 7 – 12. II. 2011, A. Gorochov (ZIN). Paratypes (2 ♀): same data as for holotype, 1 ♀ (ZIN), 1 ♀ (MNHN-EO-ENSIF 3653). Type locality. Indonesia, Sulawesi I., Lore Lindu National Park, environs of Wuasa village. Other material examined. Indonesia. Sulawesi Tengah Province, Sulawesi I., Lore Lindu National Park, ~ 45 km SSE of Palu City, environs of Tomado Vill. on Lindu Lake, ~ 1000 m, disturbed primary forest, on leaf of bush, at night, 13 – 17. II. 2011, A. Gorochov, 1 ♀ (ZIN).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B5DDE2EFF10EE43A181FA54.taxon	etymology	Etymology. The species is named in the connection with its contrasting (contrarius in Latin) coloration.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B5DDE2EFF10EE43A181FA54.taxon	distribution	Distribution. Indonesia, north-west of Sulawesi Island.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B5DDE2EFF10EE43A181FA54.taxon	diagnosis	Diagnosis. The new species is most similar to C. oeroe (also from Sulawesi), but it is distinguished from the latter species by the frons under rostral apex lighter, majority of darkened parts of male tegmina darker (including Sc-R area dark brown; in C. oeroe, this area is reddish), middle part of costal area dark with the proximal halves of Sc branches and crossveins between distal halves of Sc branches light (in C. oeroe, this part is with a wide light band along costal edge), tegminal mirror narrower, Sc-R area in female tegmina dark brown (not reddish) and with all crossveins or some of them clearly lighter, and epiphallus with the proximal corners of dorsal ridges widely rounded (not roundly angular) in profile.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B5DDE2EFF10EE43A181FA54.taxon	description	Description. Male (holotype): Coloration very characteristic: head light brown with labrum, clypeus and frons under rostal apex yellowish (Fig. 18 A), with dorsum brown and having a few lines yellowish (Fig. 18 B), with apex of apical segment of all palpi and middle and distal parts of antennal flagellum brown, and with a pair of small spots near dorsolateral corners of clypeus dark brown; pronotum with lateral lobes brown but having narrow stripe along each ventral edge yellowish (Fig. 18 C), and with disk yellowish and having rather numerous brown and light brown marks on its median 2 / 3; tegmina dark brown with several very large whitish spots (apical area of dorsal field almost completely whitish; Fig. 18 D), with R and stock of Sc as well as narrow areas between them and around diagonal vein light brown, with 3 – 4 whitish stripes along proximal half of some branches of Sc, and with a few whitish crossveins between distal halves of these branches (Fig. 18 E); exposed part of hind wings dark grayish brown with light brown venation; legs reddish brown but somewhat lighter than brown parts of head and of pronotum, with apical part of hind femora and middle segment of all tarsi dark brown, and with hind tibiae slightly darkened (grayish brown); rest of body slightly lighter than most part of legs (intermediate between brown and light brown) but with dark brown tergites having sparse lighter marks, and with uniformly light brown cerci. Body structure similar to that of C. reticulatus, however tegminal mirror 1.9 times as long as wide, Sc comb in tegmina with 7 branches (Fig. 18 E), and epiphallus of genitalia somewhat narrower in middle part and with proximal corners of dorsal ridges widely rounded in profile (Fig. 18 H – K). Female: General appearance similar to that of male, but with following differences: head in one specimen with narrow brown stripe along clypeal suture; tegminal dorsal field with 9 – 10 brown and moderately convex longitudinal veins, majority of membranes (excepting basal and lateral areas) light brown, large light spots less numerous, majority of longitudinal veins brown, and almost all crossveins whitish (Fig. 18 F); tegminal lateral field (Fig. 18 G) with Sc branches somewhat more straight, with comb of Sc having 4 – 5 branches, and with distal spot and rest of venation from whitish to light brown (including crossveins between Sc and R; but in one female, crossveins in distal half of Sc-R area brown); abdominal apex almost as in female of C. retuculatus but with ovipositor comparatively longer (hind femur 1.2 times as long as ovipositor). Variation. Female from Tomado village may correspond to another species. It is darker than the other specimens and C. oeroe, and ovipositor is slightly shorter (hind femur 1.3 times as long as ovipositor). Measurements. Length in mm. Body: male 16.5, female 18 – 18.5; body with wings: male 21, female 22 – 23; pronotum: male 2.4, female 2.6 – 2.7; tegmina: male 12, female 13 – 14; hind femora: male 13.5, female 14 – 15; ovipositor 12 – 12.5. Habitat and life history traits. Unknown. Behavior. Unknown.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B5FDE10FF10EE50A181FC31.taxon	description	(Figs 4, 19)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B5FDE10FF10EE50A181FC31.taxon	materials_examined	Type material. Male holotype: Indonesia. Sulawesi Utara Province, Wartabone National Park near Toraut Vill. not far from Doloduo Town (Minahassa Peninsula of Sulawesi I.), environs of Wallace Base Camp, primary forest near river, on leaf of bush, at night, 17 – 25. I. 2011, A. Gorochov (ZIN). Paratype (1 ♂): Indonesia. Sulawi Utara Province, Nord Celebes, Toli-toli, XI – XII. 1895, H. Fruhstorfer, identified Cardiodactylus novae-guineae by unknown person (MNHG). Type locality. Indonesia, Sulawesi I., Wartabone National Park near Toraut Vill. not far from Doloduo Town.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B5FDE10FF10EE50A181FC31.taxon	etymology	Etymology. Named after the type locality.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B5FDE10FF10EE50A181FC31.taxon	distribution	Distribution. Indonesia, north-east of Sulawesi Island.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B5FDE10FF10EE50A181FC31.taxon	diagnosis	Diagnosis. The new species is most similar to C. tangkoko in the shape of epiphallic dorsal ridges but differs in the presence of large whitish area in lateral field of tegmina and in the above-mentioned characters of male genitalia.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B5FDE10FF10EE50A181FC31.taxon	description	Description. Male (holotype): Coloration of body more or less similar to that of C. tangkoko (Fig. 19) but with following differences: spots on tegmina less contrast (general color of tegmen grayish brown with darker parts in basal area and grayish white or whitish gray marks); proximal half of tegminal lateral field (more lateral than Sc stem) and small apical part of this field rather light; most part of tegminal dorsal field (more distal than diagonal vein) almost light grayish brown with large slightly lighter spot in region of chords, with somewhat lighter (and distinctly wider than in C. tangkoko) spot at base of apical area, and with distal half of apical area somewhat lighter than proximal part of this area situated near (distad) light spot at base of this area (Fig. 19 D – E); exposed part of hind wings grayish brown with less distinct (slightly lighter) venation; legs with slightly lighter base of hind tibia; visible part of abdominal tergites less spotted. Some differences from C. tangkoko in structure of body parts also developed: tegmina with apical area slightly shorter, with mirror 1.6 times as long as wide, and with Sc comb having 6 – 7 branches; genitalia with distal half of epiphallus wider in proximal part and with high (proximal) part of dorsal epiphallic ridges shorter (compare Figs 19 F – I and 57 H – K). Female: unknown. Measurements. Length in mm. Body 17; body with wings 23; pronotum 2.7; tegmina 14.3; hind femora 16. Habitat and life history traits. C. duloduo has been found on leaf of bush, at night in primary forest near river. Behavior. Unknown.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B61DE10FF10EBB3A289FA6F.taxon	description	(Figs 4, 5 D)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B61DE10FF10EBB3A289FA6F.taxon	materials_examined	Type material. Male holotype: Philippines. Mindanao, Misamis Oriental, Mt. Empagatao, 28 km SW of Gingoog, 12. IV. 191, H. M. Torrevillas (BPBM) [not examined]. Type locality. Philippines: Mindanao, Misamis Oriental, Mt. Empagatao, 28 km SW of Gingoog. New signalization. Indonesia. Sulawesi Utara Province, small Bunaken I. very near Manado City on Sulawesi I., secondary forest near sea, on very large leaf of low plant, at night, 18 – 25. II. 2011, A. Gorochov (ZIN).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B61DE10FF10EBB3A289FA6F.taxon	distribution	Distribution. South of the Philippines, in islands of Mondanao and Leyte; north of Sulawesi.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B61DE10FF10EBB3A289FA6F.taxon	diagnosis	Diagnosis. Species of average size, contrasted coloration, face yellow with 3 dark spots, dorsal disk of pronotum pale yellow anteriorly; very similar in coloration and external morphology to C. riga, to which it differs by yellow Sc vein (brown in C. riga Otte, 2007 a) and male genitalia with only one level laterally.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B61DE10FF10EBB3A289FA6F.taxon	description	Redescription. See Robillard & Yap (2014).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B61DE13FF10EE41A181FD41.taxon	description	(Figs 4, 5 E, 9 C, 11 B, 13 C, 14 C, 20)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B61DE13FF10EE41A181FD41.taxon	materials_examined	Type material. Male holotype: Indonesia. West Java Province, Gn [Gunung = Mount in Indonesian] Halimun N [atural] P [ark], Res [ort] forest, Gn Kendeng, Cikaniki, canopy trail, 6 ° 44 ’ S 106 ° 32 ’ E, 900 – 1000 m, 8 – 15. V. 2001 (APR 01 - 2914), mal [a] ise trap 1 - A, GHNP 2001, S. Kahono, E. Cholik & Sarino (MZB). Female allotype: Indonesia. West Java Province, Gn. [Gunung = Mount] Halimun Nat [ional] Park; Hutan Resort Gn Kendeng, canopy trail, Cikaniki area, 6 ° 45 ’ S 106 ° 31 ’ E, 900 – 1000 m, Tgl 25. IV – 2. V. 2000, MTI T / GHNP 2000 (MAY 00 - 2405), E. Cholik & Sarino (MZB). Paratypes (11 ♂, 13 ♀): Indonesia. West Java Province, Gn [Gunung = Mount] Halimun N [atural] P [ark]: Resort forest, Gn Kendeng, Cikaniki, canopy trail, 6 ° 44 ’ S 106 ° 32 ’ E, 900 – 1000 m: 31. X – 7. XI. 2000, 1 ♂ (OKT- 3607), mal [a] ise trap 1 - A, GHNP 2000, E. Cholik & Sarino (molecular sample C 24) (MNHN-EO-ENSIF 3489); 8 – 15. V. 2001, 1 ♀ (APR 01 - 2915), mal [a] ise trap 1 - A, GHNP 2001, S. Kahono, E. Cholik & Sarino (ZIN); Canopy Walk (+ / - 25 m), 6 ° 44 ’ 91.1 ” S 106 ° 32 ’ 25.8 ” E, 950 m, Cikaniki: 9 – 16. V. 2000, 1 ♀ (MAY 00 - 2584), malaise trap 1 - A, E. Cholik, M. Rotik Sofyan (MNHN-EO-ENSIF 3490); 18 – 25. VII. 2000, 1 ♀ (JULY 00 - 3344), malaise trap 1 - B, M. Rotik & Sarino (MZB); Gunung Halimun [no presice locality], 1 ♂ (MNHN-EO-ENSIF 3491). Indonesia. West Java Province, Palaboean ratoe: I. 1921, 1 ♀, identified Cardiodactylus gaimardi Sauss. by L. Chopard (MNHN-EO-ENSIF 1201); II. 1921, 1 ♂, identified Cardiodactylus gaimardi Sauss. by L. Chopard (MNHN-EO-ENSIF 1212); III. 1921, 2 ♂, 2 ♀, identified Cardiodactylus gaimardi Sauss. by L. Chopard (MZB-ORTH 10488; ZIN; MNHN-EO-ENSIF 1200,1208). Indonesia. West Java Province, Java I., 20 – 25 km SE of Bogor City, Pangrango Mts, environs of Cemande Vill., 1000 m, secondary forest, on leaf of bush, at night, 27. XI – 7. XII. 1999, A. Gorochov: 5 ♂, 1 ♀ (ZIN), 1 ♂ (MNHN-EO-ENSIF 3492). Indonesia. West Java Province, Java I., environs of Sukabumi City, Gede-Pangrango Nature Reserve near Situ Gunung Lake, 11 – 12. IV. 2003, M. Berezin (ZIN), 1 ♀ (ZIN). Type locality. Indonesia, West Java, Mount Halimun Natural Park, Cikaniki area. Other material examined. Indonesia. West Java Province, Gn [Gunung = Mount] Halimun NP, Resort forest, Gn Kendeng, Cikaniki, canopy trail, 6 ° 45 ’ S 106 ° 31 ’ E, 900 – 1000 m: 4. IX- 7.2000, 1 juvenile (SEP- 3532), light trap B-III, GHNP 2000, E. Cholik & Sarino (MZB); Gunung [Mount] Halimun NP, Cikaniki, 6 ° 44 ’ 91 ” S 106 ° 32 ’ 25.8 ” E, 950 m, canopy trail, 13 – 20. III. 2001, 1 juvenile (FEB 01 - 2220), malaise trap 1 - A, Rofik & Sarino (MZB); Gn [Mount] Halimun National Park, Canopy Walk (+ / - 25 m), 6 ° 44 ’ 91.1 ” S 106 ° 32 ’ 25.8 ” E, 950 m, Cikaniki: 9 – 16. V. 2000, 1 juvenile (MAY 00 - 2953), malaise trap 1 - A, E. Cholik, M. Rotik Sofyan (MNHN).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B61DE13FF10EE41A181FD41.taxon	etymology	Etymology. Dedicated to Ms Erni Ernawati, curator of Orthoptera at MZB (Indonesia).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B61DE13FF10EE41A181FD41.taxon	distribution	Distribution. Indonesia, west of Java, Mount Halimun.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B61DE13FF10EE41A181FD41.taxon	diagnosis	Diagnosis. Species of average size, general coloration dark brown with yellow patterns. Male genitalia close to that of C. singapura and C. lampongsi, characterized by dorsal ridges of pseudepiphallus parallel or slightly divergent posteriorly, carinated laterally, with inner posterior expansions.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B61DE13FF10EE41A181FD41.taxon	description	Description. Average size for the species group. General coloration contrasted (Fig. 20 A – D), yellow brown laterally and dark brown to black dorsally. Head dorsum yellow brown with 4 dark brown bands, 2 wide lateral ones with an anterior notch fused to fastigium black coloration, 2 median bands punctuated; area posterior to eyes yellow brown with 2 short dark brown lines (Fig. 5 E). Scapes yellow brown with dark patterns on front side; antennae dark brown with yellowish rings; pedicel yellow brown with a dark brown spot on front side. Front part of fastigium yellow, sometimes with a pair of dark brown spots. Face and lateral side of head gray brown to orange brown, with a transverse yellow band ventral to eyes. Mouthparts and maxillary palpi mostly yellow brown with faint dark brown patterns. Pronotum: Dorsal disk lateral margins yellow brown to whitish; anterior area with dark brown symmetrical patterns; posterior area dark brown to black. Lateral lobes dark brown, ventral margins yellow brown. Legs I – II yellow brown to orange brown, femora with faint brown spots, tibiae with rings. Legs III yellow brown to orange brown including knees. Tarsomeres I – III (1 – 2) yellow brown, their apex dark brown. Hind wing tail dark brown, ca. 1.5 longer than pronotum. Cerci yellow brown, with dark brown spots basally then with dark brown rings. Abdomen mostly dark brown to black; dorsum with discontinuous longitudinal yellow lines. Male: FW coloration mostly dark brown. Anterior region and external part of e 1 cell black. Sclerotized area near base of chords orange brown. All veins anterior to mirror whitish, except diagonal, brown; anterior half of CuA whitish. Light area posterior to mirror almost not visible, with only lighter brown transverse veins; veins posterior to mirror brown. M / R area dark brown to orange brown, R / Sc area dark brown. Sc orange brown, posterior part yellow; projections of Sc and ventral veins of lateral field yellow, cells between them dark brown. FW venation (Fig. 9 C): 1 A slightly bisinuated; CuP missing but claval fold present in harp. Stridulatory file with ca. 200 teeth on the transverse and longitudinal parts of 1 A, without discontinuity at the level of the angle, and ca. 20 more teeth on a bump near 1 A base (see Table 3, n = 3). Harp with 2 w-shaped veins. Mirror area: mirror (d 1) longer than wide, separated near mid-length by a transverse vein, the posterior part shorter, triangular and variably subdivided by accessory veins; d 2 and next cells in D alignment wide until FW apex; E alignment widened posteriorly. Apical field with 3 – 4 cell alignments posterior to mirror (n = 4). Lateral field with 6 – 8 (m = 7.2, n = 5) projections of Sc and 4 – 5 (m = 4.2, n = 5) ventral veins. Subgenital plate yellow brown. Male genitalia (Fig. 20 E – G): Pseudepiphallic dorsal ridges parallel or slightly divergent posteriorly, carinated with inner posterior expansions. Pseudepiphallic sclerite with short flat and wide latero-anterior expansions, membrane between them with few setae. Posterior pseudepiphallic apex slightly curved toward left ventrally (n = 2), rounded, with a small rounded translucent area. Rami parallel, with large preapical plates. Ectophallic arc complete, curved. Bases of ectophallic apodemes with a posterior membranous lobe and a long anterior sclerotization. Ectophallic fold with weak lateral sclerites; apex trilobate, membranous, median lobe long and oval. Endophallic sclerite longer than wide, well sclerotized, with short posterior arms and a triangular posterior expansion. Endophallic apodeme with small lateral lamellas and a strong medio-dorsal crest. Membrane of endophallic cavity finely plicate. Female: FW coloration dark brown to black with brown veins, except bases of anal veins yellow brown. Anterior half of CuA, M and related accessory veins yellow. R / Sc area and veins dark brown. Sc projections and more ventral veins yellow brown, the cells between them dark brown. FW venation (Fig. 11 B): 10 – 11 (m = 10.2, n = 4) strong longitudinal veins on dorsal field; lateral field with 9 – 12 (m = 10.6, n = 5) longitudinal veins including 6 – 8 Sc projections and 3 – 5 ventral veins. Ovipositor of average length, shorter than FIII, apex with both dorsal and ventral edges denticulate (Fig. 13 C). Female genitalia (Fig. 14 C): Copulatory papilla trapezoidal with baso-lateral sclerites; apex rounded and sclerotized, folded ventrally. Juvenile: Light brown to yellow brown, mottled with dark brown. Measurements. See Table 3. Habitat and life history traits. Unknown. Behavior. Unknown.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B62DE17FF10ED8EA5BBFCEC.taxon	description	(Figs 4, 5 F, 9 D, 11 D, 13 D, 14 D, 21, 22, 23)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B62DE17FF10ED8EA5BBFCEC.taxon	materials_examined	Type material. Male holotype: Indonesia. East Nusa Tenggara Province, Flores, Ruteng district, Danau Ranamese Recreation Park, bord E du lac (GPS Flo 3 - 4), 08 ° 38 ' 22.1 " S 120 ° 33 ' 47.6 " E, 1225 m, 22. VI. 2010, nuit, sur plante basse, adulte en élevage, enregistrement appel, T. Robillard (MZB). Female allotype: same information as HT (MZB). Paratypes (3 ♂): Indonesia. East Nusa Tenggara Province, Flores, Ruteng district, Danau Ranamese Recreation Park, bord E du lac (GPS Flo 3 - 4), 08 ° 38 ' 22.1 " S, 120 ° 33 ' 47.6 " E, 1225 m, 22. VI. 2010, nuit, sur plante basse, adulte en élevage, T. Robillard: 1 ♂ (MZB); 2 ♂, enregistrement appel (MNHN-EO-ENSIF 3516 - 3517). Type locality. Indonesia, Flores, Ruteng district, Danau Ranamese Recreation Park. Other material examined. Indonesia. East Nusa Tenggara Province, Flores, Ruteng district, Danau Ranamese Recreation Park, bord E du lac (GPS Flo 3 - 4), 08 ° 38 ' 22.1 " S 120 ° 33 ' 47.6 " E, 1225 m, 22. VI. 2010, nuit, T. Robillard: 1 ♀, adulte en élevage (MNHN); 1 juvenile (TR 344), sur plante basse (MZB); 1 juvenile (TR 345), sur plante basse (MNHN).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B62DE17FF10ED8EA5BBFCEC.taxon	etymology	Etymology. Named after the type locality.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B62DE17FF10ED8EA5BBFCEC.taxon	distribution	Distribution. Indonesia, west of Flores Island.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B62DE17FF10ED8EA5BBFCEC.taxon	diagnosis	Diagnosis. Species of average to small size, general coloration brownish with yellow patterns. Characterized by male genitalia close to C. muria, with dorsal ridges slightly asymmetrical, carinated innerly and externally.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B62DE17FF10ED8EA5BBFCEC.taxon	description	Description. Average to small size for the species group. General coloration yellow brown (Fig. 21 A – D). Head dorsum yellow brown with 4 brown bands including 2 complete lateral ones fused to the dark brown fastigium without a clear anterior notch, and 2 median punctuated bands; areas posterior to eyes yellow brown with 2 brown triangles (Fig. 5 F). Scapes yellow brown, with a faint brown transverse band; rest of antennae orange brown. Front part of fastigium yellow brown 2 dark brown stripes extended below antennae. Face yellow brown, with a transverse yellow band extended bellow eyes. Lateral side of head variable, mostly gray brown mottled with dark brown. Mouthparts yellow brown more or less mottled with brown. Maxillary palpi yellow brown, apex dark brown. Pronotum: Dorsal disk almost completely yellow brown, more or less mottled with brown; posterior edge dark brown. Lateral lobes mostly brown, ventral quarter yellowish. Legs I – II yellow brown, femora with faint orange brown spots and tibiae with rings. FIII homogeneously orange brown, knees dark brown. TIII dark brown, with faint yellow brown rings. Tarsomeres III- 1 orange brown, their ends darker. Hind wing tail dark brown. Cerci yellow brown with faint dark brown rings, their bases dark brown. Abdomen mostly orange brown to dark brown. Subgenital plate yellow brown. Male: FW coloration dark brown with yellow areas (Fig. 9 D) including bases of anal veins and anterior half of CuA, angle of 1 A, harp veins, chords and a wide transverse band posterior to mirror. Black coloration on cell e 1 and area between bases of 1 A / 2 A and transverse part of 1 A. Diagonal vein orange brown. Apex of apical field grayish. M / R / Sc area orange brown; M dark brown; R and Sc orange brown, apex of Sc whitish; M / R posterior junction yellow. Sc projections and more ventral veins of lateral field yellow. FW venation (Fig. 9 D): 1 A clearly bisinuated. Stridulatory file with about 181 teeth (n = 1) on the transverse and angle parts of 1 A, and 14 more teeth on a bump near base of 1 A (Table 4). CuP absent. Harp with 2 w-shaped veins. Mirror area: mirror (d 1) distinct, longer than wide, separated at 2 / 3 by a transverse vein; d 2 as wide as mirror, made of 1 main cell. Cell e 1 not crossed by transverse veins. Apical field with 4 cell alignments posterior to mirror (n = 3). Lateral field with 7 projections of Sc (n = 3) and 4 – 5 ventral veins (m = 4, n = 3). Male genitalia (Fig. 21 E – G): Pseudepiphallus shaped as a wide gutter. Dorsal ridges divergent posteriorly, slightly asymmetrical in shape, with a postero-inner carination and a medio-outer one. Pseudepiphallic sclerite with long anterior expansions, blade-like, close in shape to the hook observed in C. rizali, the membrane between them with long setae. Posterior pseudepiphallic apex rounded, with a large square translucent area. Rami with short apical stems oriented anteriorly. Ectophallic arc complete, without posterior expansion. Ectophallic fold with lateral sclerites; apex trilobate, membranous, median lobe oval, smaller than lateral ones. Endophallic sclerite short its apodeme with a dorsal crest and narrow lateral lamellas. Membrane of endophallic cavity not clearly plicate. Female: FW coloration mostly dark brown, their base yellow brown, including bases of anal veins, and anterior half of CuA and M. Dark brown to black area including part of CuP and 1 A posterior to yellow basis. R / Sc area and veins orange brown, apex of Sc whitish. Sc projections and more ventral veins yellow brown, the cells between them orange brown. FW venation (Fig. 11 D): AT with 10 strong longitudinal veins on dorsal field; lateral field with 8 longitudinal veins including 5 projections of Sc and 3 more ventral veins. Ovipositor of average length, slightly shorter than FIII; apex with both dorsal and ventral edges denticulate (Fig. 13 D). Female genitalia (Fig. 14 D): Copulatory papilla triangular, with baso-lateral sclerites; apex rounded and slightly sclerotized, folded ventrally. Juvenile: Light brown, mottled with dark brown. Measurements. See Table 4. Habitat and life history traits. Cardiodactylus floresiensis is a nocturnal species living in forested areas. Only juveniles were observed in the field in Flores, on low branches of bushes or small trees (Fig. 22). Behavior. Calling song (Fig. 23): Cardiodactylus floresiensis has a mono-syllabic calling song. At 21 ° C, the calling songs show a very indented amplitude profile. Syllables have the following characteristics: syllable duration = 84.0 ± 3.6 ms; syllable period = 2.5 ± 0.1 s; syllable duty cycle = 3.4 %. The power spectrum shows a clear harmonic pattern; the dominant frequency is 15.43 ± 0.12 kHz and corresponds to the third frequency peak.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B66DE19FF10ED87A181F99C.taxon	description	(Figs 4, 5 G, 9 D, 11 C, 13 E, 14 E, 24)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B66DE19FF10ED87A181F99C.taxon	materials_examined	Type material. Male holotype: Indonesia. South Sulawesi Province, Samanga, S. Celebes [Sulawesi I., Buttu Samanga], XI. 1895, H. Fruhstorfer, # 61, 130 - 97, determined as C. novaeguineae by unknown person (ZIN). Paratypes (1 ♂, 3 ♀): same locality and collector as holotype, 1 ♀ (MHNG), 1 ♀ (MNHN-EO-ENSIF 3656). S. Celebes, [Sulawesi I.], Patunuang, I. 1896, H. Fruhstorfer: 1 ♂, # 61, 130 - 97, determined as C. novaeguineae by unknown person (ZIN); 1 ♀ (MHNG). Type locality. Indonesia, southern part of Sulawesi Island, Buttu Samanga. Other material examined. Indonesia. South Sulawesi Province, Mont Bulu Saraung, Pos 5. 31. VIII. 2007, forêt sur pente, espace dégagé en bord de piste, C. Villemant: 1 ♀, adulte en élevage (MNHN-EO-ENSIF 1223); 2 juveniles (MNHN).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B66DE19FF10ED87A181F99C.taxon	etymology	Etymology. Dedicated to the collector Dr. H. Fruhstorfer.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B66DE19FF10ED87A181F99C.taxon	distribution	Distribution. Indonesia, southern part of Sulawesi Island.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B66DE19FF10ED87A181F99C.taxon	diagnosis	Diagnosis. The new species is most similar to C. contrarius by the shape of male epiphallus, but it differs from the latter in the general coloration less contrasted, face of epicranium clearly darker, whitish spots on the male tegmina much smaller, and hind femora lacking any distinct dark areas at the apex. From C. oeroe, the new species differs in the same characters (excepting general coloration) and proximal corners of the dorsal epiphallic ridges widely rounded (not roundly angular) in profile, and from C. celebae (also from Sulawesi), in the whitish spots of male tegminal lateral field smaller, epiphallus distinctly longer, and its dorsal ridges slightly longer and in profile separated from the dorsoproximal epiphallic lobes by the notches clearly deeper.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B66DE19FF10ED87A181F99C.taxon	description	Description. Male (holotype): Coloration of body light brown with following marks: head brown with genae and most part of mandibles yellowish, small spots on rostral apex and on frons near clypeus as well as antennae and rest of mouthparts light brown (Figs 5 G, 24 A – B); pronotum light brown with most part of lateral lobes almost dark brown and median part of disk having rather small and weakly distinct brown marks (Fig. 24 B – C); tegminal dorsal field brown with semitransparent light brown areas in basal part and around oblique veins and chords, 2 rather large yellowish spots (one near mirror, and one at base of apical area), and oblique veins and part of chords almost whitish (Fig. 24 D – E); tegminal lateral field with M, R, stock of Sc, and areas and crossveins between these veins reddish brown, with two rather large whitish spots (one near mirror, and one in distal part), with Sc branches and crossveins between them also whitish, and with membranes between these crossveins grayish, semitransparent (Fig. 24 E); exposed part of hind wings grayish brown with venation weakly distinct; legs almost uniformly light brown but with middle part of tarsi darker (brown); rest of body brown with coxae, thoracic sternites, and genital plate light brown (coloration of anal plate and cerci intermediate between brown and light brown). Structure of body similar to that of C. reticulatus, however tegmina slightly not reaching abdominal apex, mirror 1.8 times as long as wide, Sc comb with 5 – 6 branches (Fig. 24 E), and epiphallus of genitalia distinguished from that of C. contrarius only by distal part slightly wider and proximal half of dorsal ridges slightly higher (Fig. 24 G – J). Female: FW brown (Fig. 24 F), its base whitish, longitidunal veins mostly orange brown, transverse veins variable, yellowish, whitish or brown; with a whitish sclerotizaion at FW mid-length on lateral external edge, not extended on dorsal field as in C. oeroe. R and Sc veins orange brown, the area between dark brown; projections of Sc yellow, the membrane between them dark brown; posterior corner of lateral field whitish. FW venation: dorsal field with 9 longitudinal veins, lateral field with 3 projections of Sc and 3 more ventral veins. Ovipositor short, its apex slightly denticulate dorsally and ventrally. Female genitalia: Copulatory papilla more or less trapezoidal, with wide baso-lateral sclerites, apex rounded, slightly sclerotized, folded ventrally. Juvenile: Contrasted coloration (Fig. 47 A), mostly light brown and yellow brown mottled with dark brown spots. Head coloration as in adults; pronotum yellow brown with few dark brown spots. Variation s. Paratype with lateral parts of frons under antennal cavities lighter (light brown), dorsum of head having narrow light longitudinal lines, and hind tibiae with weakly distinct (darkish) spots. Measurements. Length in mm. Body: male 19 – 20, female 21.6 – 23.7; body with wings: male 23 – 24, female 26.5 – 26.7; pronotum: male 2.7 – 2.8, female 3 – 3.4; tegmina: male 13 – 13.5, female 14.2 – 14.5; hind femora: male 14.5 – 15.5, female 16.2 – 17.9; ovipositor 13.3 – 14.4. Habitat and life history traits. Cardiodactylus fruhstorferi has been observed as juveniles only, at night on low plants and branches in open forested areas (Fig. 47). Behavior. Unknown.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B68DE1CFF10EE18A2F6FD16.taxon	description	(Figs 25 – 26)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B68DE1CFF10EE18A2F6FD16.taxon	materials_examined	Type material. Female holotype: Japan. Okinawa, Riukiu [Ryukyu] (K. Kuroiwa) (Hokaido University). Not examined. Type locality. Okinawa, Ryukyu Islands. New signalizations. Japan, Ryukyu, Okinawa Pref., Ishigaki I., 14 – 15. X. 1999, S. Belokobylsky, 4 ♂, 5 ♀ (ZIN); Iriomote I., 16 – 17. X. 1999, S. Belokobylsky, 1 ♂, 1 ♀ (ZIN); Okinawa-jima, Kunigami-cha, Yona, 14. VIII. 1991, A. Lelej, 1 ♂ (ZIN); Amami I., 11. IX. 2000, A. Bugrov (ZIN), 1 ♂, 1 ♀ (ZIN); Amami-Oshima, Komia near Sumno, 17. XII. 1926, P. Schmidt, 1 ♂ (ZIN). Vietnam (northern). Quang Tri Province, Huong Hoa Distr., Huong Lap Communa, XI. 2007, N. Orlov, 1 ♂ (ZIN). Quang Binh Province, Ba Ty Long Archipelago, Thanh Lan I., 18. X. 1990, V. Kuznetsov, 1 ♀ (ZIN). Vietnam (southern half). Gia Lai Province, 20 km N of Kannack Town, environs of Buon Luoi Vill., 24 – 30. IV. 1995, A. Gorochov, 1 ♂ (ZIN); same information, 3 – 11. XI. 1993 and 22. III- 10. V. 1995, A. Gorochov, 2 ♂, 2 ♀ (ZIN); same province, environs of Kannack Town, 8 – 16. XI. 1988, A. Gorochov, 1 ♀ (ZIN).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B68DE1CFF10EE18A2F6FD16.taxon	distribution	Distribution. Japan (Ryukuy Islands), coastal part of Vietnam (north to south), Hainan Island (China), Taiwan.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B68DE1CFF10EE18A2F6FD16.taxon	diagnosis	Diagnosis and redescription. See Robillard and Ichikawa (2009).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B68DE1CFF10EE18A2F6FD16.taxon	description	Variation. Specimens from southern Vietnam tend to vary in female coloration (median anterior witish spot on FWs nearly absent, Fig. 25 A), and one male has very angular peudepiphallic dorsal ridges in lateral view (Fig. 25 B). Such characteristics could be seen as sufficient to diagnose a new species or subspecies if they were stable within the sampled locality, which is not the case.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B6DDE1EFF10EA96A181FE61.taxon	description	(Figs 4, 5 H, 13 F, 14 F, 27)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B6DDE1EFF10EA96A181FE61.taxon	materials_examined	Type material. Male holotype: Indonesia. Maluku Utara Province, Halmahera I., environs of Subaim Vill. (to S from Lolobata Vill. but not far from it) near coast of Wasile Bay, low hills, disturbed forest, on leaf of bush, at night, 27. I – 1. II. 2011, A. Gorochov (ZIN). Paratypes (1 ♂, 3 ♀): same information as holotype, 1 ♂, 1 ♀ (ZIN); 1 ♀ (MNHN-EO-ENSIF 3655). Halmahera [I.] Noord-Oost [North-West], Bernstein, 1 ♀ (RMNH). Type locality. Indonesia, Halmahera Island, environs of Subaim Village (to S from Lolobata Village but not far from it) near coast of Wasile Bay.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B6DDE1EFF10EA96A181FE61.taxon	etymology	Etymology. Named after type locality.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B6DDE1EFF10EA96A181FE61.taxon	distribution	Distribution. Indonesia, Halmahera Island.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B6DDE1EFF10EA96A181FE61.taxon	diagnosis	Diagnosis. The new species is characterized by the male epiphallus with assymetrical dorsal ridges folded laterally, close to that of C. kotandora and with a rather deep notch at middle in profile view.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B6DDE1EFF10EA96A181FE61.taxon	description	Description. Male (holotype): General appearance similar to that of C. empagatao but with following differences: face of head with a pair of larger dark brown spots under antennal cavities connected with each other by reversed U-shaped brown narrow stripe almost reaching rostral apex, and with rather long dark stripe along clypeal suture; dorsum of head almost completely brown (including dorsal surface of rostrum) with narrow light brown stripe outlined anterior part of this dorsum, and with longitudinal yellowish stripe behind each eye (Fig. 27 A – C); pronotum with dark brown most part of lateral lobes and of disk (only 2 pairs of longitudinal light stripes along lateral edges of disk and along ventral edges of lateral lobes developed; Fig. 27 B – C); hind tibiae with somewhat more numerous darkish spots; tegmina with slightly darker most part of distal half of dorsal field and more contrast (white) light marks, but cell membranes in region of large light spot at base of apical area slightly darker, with grayish tinge; Fig. 27 D – E); hind wings with more distinct (light brown) venation; abdominal tergites somewhat darker (mainly grayish brown), and almost all sternites of almost same color (however with lateral parts light brown); wings somewhat longer, with narrower mirror (ratio length / width about 2) and 11 – 12 branches on Sc (Fig. 27 E). Genitalia distinguished by assymetrical epiphallic dorsal ridges, they dorsal edge folded laterally, and by presence of rather deep notch at middle in profile view (Fig. 27 H – K). Female: General appearance similar to that of holotype, but sometimes hind part of head dorsum slightly lighter and face without dark stripe along clypeal suture, tegmina in all females brown with less contrast light marks similar to those of female of C. empagatao and with additional yellowish spot between basal and lateral light spots on dorsal field but not along lateral edge of this field (area between this additional spot and basal spot almost dark brown) (Fig. 27 F – G). Sc comb with 8 – 9 branches. Ovipositor short (hind femur 1.2 – 1.3 times as long as ovipositor) (Fig. 13 F). Female genitalia (Fig. 14 F): Copulatory papilla more or less triangular, with small baso-lateral sclerites, its apex elongate, slightly sclerotized, folded ventrally. Juvenile: unknown. Variations. Second male somewhat lighter and with less contrasted coloration of head, pronotum and tegmina; its genitalia with some parts less sclerotized, and with notch on right epiphallic ridge less deep (this male possibly collected very shortly after moulting). Measurements. Length in mm. Body: male 20 – 22, female 22 – 23; body with wings: male 27 – 31, female 31 – 33; pronotum: male 3 – 3.3, female 3.5 – 3.7; tegmina: male 15.5 – 18, female 18.5 – 19.5; hind femora: male 18.5 – 20.5, female 20 – 21; ovipositor 16 – 17. Habitat and life history traits. Unknown. Behavior. Unknown.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B6FDE00FF10EA43A181FE1C.taxon	description	(Figs 4, 5 I, 9 E, 28)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B6FDE00FF10EA43A181FE1C.taxon	materials_examined	Type material. Male holotype: Indonesia. Riau Islands Province, Doerian [Durian Island], Riouw-Arch [Riau Islands], VI. 1923 (MNHN-EO-ENSIF 3496). Paratypes: Indonesia. No precise locality, 1877 – 1878, 1 ♂, Sumatra expedition, 272 (RMNH). Type locality. Indonesia, Riau Islands, Durian Island.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B6FDE00FF10EA43A181FE1C.taxon	etymology	Etymology. Named after the type locality.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B6FDE00FF10EA43A181FE1C.taxon	distribution	Distribution. Indonesia, Riau Islands, Durian Island.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B6FDE00FF10EA43A181FE1C.taxon	diagnosis	Diagnosis. Species of average to small size, rather slender for the species group, general coloration brown and yellow brown, characterized by male FW venation with only 1 harp vein and a light rounded area posterior to mirror. Male genitalia close to that of C. kondoi from which it differs by shape of pseudepiphallic dorsal ridges and apex and by curved ectophallic arc.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B6FDE00FF10EA43A181FE1C.taxon	description	Description. Average to small size, rather slender for the species group. General coloration brown with yellow areas. Head dorsum yellow brown with 4 dark brown bands, the lateral ones wide and complete, with an anterior notch fused to fastigium black coloration, the median ones narrower and punctuated; 2 thin dark brown lines posterior to eyes (Fig. 5 I). Scapes yellow brown with faint darker patterns; antennae orange brown. Face mottled with yellow brown and dark brown, with a yellow line below eyes. Mouthparts mostly yellow brown. Lateral sides of head brown. Maxillary palpi brown. Pronotum: Dorsal disk lateral margins yellow, median area yellow mottled with brown, posterior margin dark brown. Lateral lobes dark brown, ventral margin yellow. Legs I – II yellow brown, with faint dark brown spots on femora and rings on tibiae; tarsomeres yellow brown basally, their ends dark brown. Legs III orange brown, faintly mottled with yellow and dark brown, knees slightly darker. Hind wing tail dark brown, 1.5 times longer than pronotum. Cerci yellow brown with dark brown spots. Posterior tergites dark brown with yellow spots; posterior sternites yellow brown. Male: FW coloration contrasted (Figs 9 E, 28 A – B), mostly yellow and dark brown, with an orange brown sclerotized area anterior to chords. Yellow areas include: basal third of CuA, bases of 1 A – 3 A, angle of 1 A, base of chords, harp, anterior half of diagonal vein, small rounded area posterior to mirror and veins of apical field. M / R / Sc area orange brown, Sc projections yellow brown, cells between them dark brown. FW venation (Fig. 9 E): 1 A slightly bisinuated. CuP missing. Harp with 1 w-shaped vein; posterior margin only slightly raised along diagonal vein. Mirror area: mirror (d 1) longer than wide, not rounded but well defined, separated in two parts by a strong transverse vein, the posterior part smaller than anterior part and subdivided in 4 parts by accessory veins; d 2 little differentiated; e 1 separated in two parts by a transverse vein. Apical field with 4 cell alignments posterior to mirror. Lateral field with 7 projections of Sc and 4 more ventral veins. Subgenital plate yellow brown. Male genitalia (Fig. 28 C – E): Pseudepiphallus well sclerotized, clearly narrowed twice, at mid-length and preapically. Dorsal ridges diverging posteriorly, forming a wide gutter, their antero-dorsal edges slightly carinated. Pseudepiphallic sclerite with wide latero-anterior expansions cut straight at apex, membrane between them without setae. Posterior region rounded, narrower than preapical part of pseudepiphallus, apex with a small rounded translucent area, slightly denticulate. Rami strong, apex slightly convergent. Pseudepiphallic parameres median lobe very slecrotized, rectangular. Ectophallic arc complete, wide and curved, without posterior expansion. Ectophallic apodemes prolonging arc curvature, convergent, their bases with a posterior membranous lobe and a faint anterior sclerotization. Ectophallic fold laterally sclerotized, apex trilobate, membranous, median lobe oval. Endophallic sclerite well sclerotized, posterior arms disconnected from lateral sclerites of ectophalic fold. Endophallic apodeme with small lateral lamellas and a strong medio-dorsal crest. Membrane of endophallic cavity smooth. Female: Unknown. Juvenile: Unknown. Measurements. See Table 5. Habitat and life history traits. Unknown. Behavior. Unknown.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B71DE04FF10EB8BA44DFDD1.taxon	description	(Figs 4, 5 J, 9 F, 12 A, 13 G, 14 G, 29, 30, 31)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B71DE04FF10EB8BA44DFDD1.taxon	materials_examined	Type material. Male holotype: Philippines: Sulu Province, Tarawakan NE, 12. II. 1957, Y. Kondo (BPBM) (not examined). Type locality. Philippines: Sulu Province, Tarawakan NE. Material examined. Philippines: Manille [Manila, no precise locality], identified Cardiodactylus novaeguineae, 1 ♂ (MHNG). Laguna Province, Luzon, Los Baňos, Mont Makiling, forêt de mousse vers sommet # 2, 14 ° 08 ' 15,2 " N 121 ° 11 ' 47,7 " E, 925 m (GPS MOSS), 1. VII. 2011, nuit, sur végétation, T. Robillard: 1 ♂, adultes en élevage, enregistrements appel (MNHN-EO-ENSIF 3512); 3 ♀, adultes en élevage (MNHN); 3 juveniles (MNHN); 3 ♀, adultes en élevage (UPLB-MNH); 3 juveniles (UPLB-MNH). Paete, University of Philippines Laguna Land Grant, forêt secondaire à 500 m de la station, 14 ° 23 ' 56,9 " N 121 ° 32 ' 47,2 " E, 376 m, 4 – 6. VII. 2011, nuit, sur végétation, T. Robillard: 2 ♂, adultes en élevage, enregistrements appel (MNHN-EO-ENSIF 3513 - 3514); 2 juveniles (MNHN); 1 juvenile (MNHN-EO-ENSIF 3154); 5 juveniles (UPLB-MNH). Mt Makiling, 22. II. [19] 07, 1908 - 228, 1 ♀, identified Cardiodactylus novaeguineae by Chopard, C. S. Banks (BMNH). Romblon Province, Sibuyan Island, 1 ♂, 20435, Baker (MNHN). Quezon Province, Pollilo Island, Puting Bato, forest near limestone cave, 2 - 11 - 2012, 1 ♀, Mark V. Yngente (MNHN-EO-ENSIF 3160). Negros Occidental Province, Negros Island, Mount Kanlaon, 2010, 1 ♂, S. A. Yap & O. L. Eusebio (UPLB-MNH). Oriental Mindoro Province, Mindoro I., environs of Puerto Galera Town (northern coast of island), on leaf of bush, at night, 11 – 13. III. 2004, A. Gorochov, 1 ♂ (ZIN).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B71DE04FF10EB8BA44DFDD1.taxon	distribution	Distribution. Philippines, islands of Luzon, Sulu, Negros, Sibuyan and Mindanao.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B71DE04FF10EB8BA44DFDD1.taxon	diagnosis	Diagnosis. Species of average size, coloration contrasted, dark brown with yellow patterns. Characterized by head dorsum coloration with one median punctuated brown band. Male genitalia close to that of C. guttulus, with bean-shaped dorsal ridges slightly diverging posteriorly.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B71DE04FF10EB8BA44DFDD1.taxon	description	Redescription. In addition to the characters given by Otte (2007): size average for the genus, coloration variable, with contrasted yellow and dark brown patterns (Fig. 29 A – D). Head dorsum yellow brown with 3 dark brown bands including 2 wide lateral ones with an anterior notch not related to black coloration of fastigium, and a median punctuated band made of the fusion of two thin parallel bands (Fig. 5 J); short triangular bands posterior to eyes slightly connected to lateral bands. Scapes yellow brown with a dark brown ring; antennae orange brown. Face and mouthparts yellow, with a dark brown area below eyes. Lateral part of head sometimes with a dark brown band posterior to eyes. Pronotum: Dorsal disk mostly brown, mottled with dark brown and yellow; anterior corners yellow. Lateral lobes dark brown, with a narrow yellowish ventral margin. Legs yellow to orange brown, femora faintly mottled with dark brown, tibiae with dark brown rings. Hind knees dark brown. Male: FW coloration mostly brown (cells and veins), anterior area dark brown to black; with yellow areas including bases of anal veins and CuA, harp veins, oval region posterior to mirror, part of the chords and apex of apical field. Bases of chords with an orange brown sclerotization. M / R / Sc area and veins dark orange brown. Sc projections and more ventral veins of lateral field yellow, their bases brown. FW venation (Fig. 9 F): 1 A slightly bisinuated; CuP absent. Harp with 2 w-shaped veins, the anterior one weaker. Mirror area: mirror (d 1) oval, separated in two parts near mid-length; d 2 as wide as mirror, not distinctive; e 1 as long as d 1. Apical field with 4 cell alignments posterior to mirror (n = 3). Lateral field with 6 – 7 projections of Sc (m = 7, n = 3) and 3 – 5 ventral veins (m = 4, n = 3). Male genitalia (Fig. 29 E – G): Pseudepiphallus forming a wide gutter not narrowed preapically, its posterior apex rounded. Dorsal ridges diverging posteriorly, slightly carinated externally, without lateral expansions. Pseudepiphallic sclerite with very short and wide latero-anterior expansions, membrane between them with short setae. Female: FW coloration mostly dark brown, anterior and latero-external regions black, with yellow areas including bases of anal veins, CuA and anterior quarter of M, a yellow area at mid-length including a part of CuP, CuA, M and R, and 2 yellow spots at ¼ of FW length (Fig. 12 A). R / Sc area and veins orange brown. Sc projections and more ventral veins yellow, cells between them dark brown. FW venation (Fig. 12 A): 9 – 10 (m = 9, n = 3) strong longitudinal veins on dorsal field; lateral field with 7 – 9 (m = 8, n = 3) longitudinal veins including 4 – 6 projections of Sc and 3 – 4 ventral veins. Ovipositor of average length, apex with both dorsal and ventral edges slightly denticulate (Fig. 13 G). Female genitalia (Fig. 14 G): Copulatory papilla rectangular, with lateral sclerites; apex rounded and slightly sclerotized, folded ventrally. Juvenile: Head dark red brown, body and legs yellow brown highly mottled with brown and black. Lateral lobes of pronotum black with a ventral yellow margin (Fig. 30 D). Habitat and life history traits. Cardiodactylus kondoi is a nocturnal species living in forested areas (Fig. 30 A). Only juveniles were observed in Luzon, on low branches of bushes or small trees. Behavior. Calling song (Fig. 31): Cardiodactylus kondoi has a mono-syllabic calling song. At 21 ° C, the syllables show a very indented amplitude profile with the following characteristics: syllable duration = 90.0 ± 14 ms; syllable period = 2.2 ± 0.6 s; syllable duty cycle = 4.1 %. The power spectrum shows a clear harmonic pattern; the dominant frequency is 12.81 ± 0.16 kHz and corresponds to the third frequency peak.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B75DE07FF10EACEA0E4FB2E.taxon	description	(Figs 4, 5 K, 9 G, 11 F, 13 H, 14 H, 32, 33, 34)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B75DE07FF10EACEA0E4FB2E.taxon	materials_examined	Type material. Male holotype: Indonesia. East Nusa Tenggara Province, Flores, Borong district, Mont Boco Nbegi, prox. village Kotandora, forêt secondaire sur pente (GPS Flo 5) 08 ° 49 ' 06.2 " S 120 ° 37 ' 55.9 " E, 245 m, 23 – 24. VI. 2010, jour, sur plante basse, adulte en élevage, enregistrement appel, T. Robillard (MZB). Female allotype: same information as HT (MZB). Paratypes (4 ♂, 2 ♀): same information as HT, 1 ♂ (MZB); 1 ♂, 1 ♀ (ZIN); 2 ♂, 1 ♀ (MNHN-EO-ENSIF 3518 - 3520). Type locality. Indonesia, Flores, Borong district, Mont Boco Nbegi, prox. village Kotandora. Other material examined. Indonesia. East Nusa Tenggara Province, Flores, Borong district, Mont Boco Nbegi, prox. village Kotandora, forêt secondaire sur pente (GPS Flo 5) 08 ° 49 ' 06.2 " S 120 ° 37 ' 55.9 " E, 245 m, T. Robillard: 23 – 24. VI. 2010, jour, 2 juveniles (TR 362, 451) (MNHN); 3 ♂, (MNHN-EO-ENSIF 3649 - 3651), 1 ♀, reared specimens (F 1 generation, TR- 2010 - 2011) (MNHN).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B75DE07FF10EACEA0E4FB2E.taxon	etymology	Etymology. Named after the type locality.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B75DE07FF10EACEA0E4FB2E.taxon	distribution	Distribution. Indonesia, south-west of Flores Island.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B75DE07FF10EACEA0E4FB2E.taxon	diagnosis	Diagnosis. Species of average size, general coloration contrasted, dark brown with yellow patterns. Characterized by male genitalia, close to C. sumba and C. muria, with dorsal ridges carinated innerly and externally and with asymmetrical triangular lateral expansions.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B75DE07FF10EACEA0E4FB2E.taxon	description	Description. Average size for the species group. General coloration contrasted. Head dorsum yellow brown with 4 faint brown bands including 2 wide complete lateral ones with an anterior notch, most often fused to the dark brown fastigium, and 2 median, narrower punctuated bands; areas posterior to eyes yellow brown with 2 dark brown triangles (Fig. 5 K). Scapes yellow brown, with a faint brown transverse band; rest of antennae orange brown. Front part of fastigium yellow with 2 dark brown spots. Face and mouthparts almost homogeneously yellow or light brown. Lateral side of head mostly gray brown or yellow brown, with a faint brown dorsal band. Maxillary palpi yellow brown, apex dark brown. Pronotum: Dorsal disk almost completely yellow brown, more or less mottled with brown; posterior edge dark brown. Lateral lobes mostly orange brown, ventral quarter yellowish. Legs I – II yellow brown, femora with faint orange brown spots and tibiae with rings. FIII homogeneously orange or yellow brown, knees dark brown. TIII dark brown. Tarsomeres III- 1 yellow brown. Hind wing tail dark gray brown. Cerci light brown mottled with dark brown. Abdomen mostly orange brown to dark brown with vivid yellow and orange patterns laterally. Subgenital plate yellow brown. Male: FW coloration with dark brown to black and vivid yellow areas (Fig. 9 G). Yellow areas include base of FWs (including bases of anal veins, CuA, M, R, Sc), anterior part of 3 A, 1 A excluding a small dark brown area near base), harp veins, most part of chordal area (veins yellow, cells translucent), and a wide band posterior to mirror with translucent cells and yellow veins, wider on inner region than externally. Black coloration variable, either on all non-yellow areas, or limited to region between FW base and transverse part of 1 A and region posterior to harp including mirror and apical field. Sclerotized area at base of chords dark brown or orange brown. Apex of apical field grayish. Diagonal brown. M / R / Sc area and veins orange brown before, M / R area and veins black posterior to their fusion; apex and projections of Sc and more ventral veins of lateral field yellow, the cells between them dark brown. Ventral margin of and posterior triangular apex of lateral field translucent. FW venation (Fig. 9 G): 1 A clearly bisinuated. Stridulatory file with 215 – 230 teeth (m = 223; n = 3) on the transverse and angle parts of 1 A, and 10 – 16 (m = 13; n = 3) teeth on a bump near base of 1 A, with no clear discontinuity with the rest of the file (Table 6). CuP absent. Harp with 2 w-shaped veins. Mirror area: mirror (d 1) distinct, usually as long as wide, diamond-shaped, crossed at mid-length by a transverse vein; d 2 not distinct. Cell e 1 not crossed by transverse veins. Apical field with 4 cell alignments posterior to mirror (n = 5). Lateral field with 8 – 11 projections of Sc, some bifurcated basally (m = 10; n = 5) and 3 – 4 more ventral veins (m = 4; n = 5). Male genitalia (Fig. 32 E – G): Pseudepiphallus shaped as a wide gutter. Apex of dorsal ridges with short setae, folded laterally, the lateral triangular expansions clearly asymmetrical, the left one larger and more posterior, the right one smaller and more anterior. Dorsal ridges almost convex in lateral view. Pseudepiphallic sclerite with wide triangular anterior expansions, the membrane between them setose. Posterior pseudepiphallic apex rounded, with a rounded translucent area. Rami with short apical stems oriented posteriorly. Ectophallic arc complete, without posterior expansion. Ectophallic fold with thin lateral sclerites; apex trilobate, membranous, median lobe oval. Endophallic sclerite small, with a triangular posterior expansion. Endophallic apodeme with a dorsal crest and lateral lamellas. Membrane of endophallic cavity smooth. Female: FWs mostly brown with orange brown veins, their base whitish, dorsal field with four whitish areas: a lateral one at FW mid-length, between CuP and Sc, followed by a black area; second whitish area median, at 1 / 4 of FW length; 2 fainter light areas more variable at 1 / 3 of FW length near inner margin. CuA and M whitish basally, then successively dark brown, whitish and dark brown again apically. M / R / Sc areas and veins orange brown; Sc apex whitish. Projections of Sc whitish, their base orange brown. FW venation (Fig. 11 F): 10 – 11 (m = 10, n = 3) strong longitudinal veins on dorsal field; lateral field with 11 (m = 11, n = 3) longitudinal veins including 7 – 8 projections of Sc and 3 – 4 ventral veins. Ovipositor of average length, slightly shorter than FIII; apex dorsal edge with only thinely denticulate; ventral edge more clearly denticulate (Fig. 13 H). Female genitalia (Fig. 14 H): Copulatory papilla conical, with strong and thick baso-lateral sclerites; apex rounded, slightly sclerotized, folded ventrally. Ventral membrane plicate longitudinally. Juvenile: brown, mottled with dark brown and yellow. Measurements. See Table 6. Habitat and life history traits. Cardiodactylus kotandora is a nocturnal species living in secondary forested areas. Only juveniles were observed in the field in Flores, on low branches of bushes or small trees (Fig. 33). Behavior. Calling song (Fig. 34): Cardiodactylus kotandora has a mono-syllabic calling song. At 21 ° C, the calling songs are emitted in irregular groups of 1 – 9 (m = 4.2 ± 2.0, n = 3). Each syllable shows a very indented amplitude profile and has the following characteristics: syllable duration = 129.4 ± 14 ms; syllable period = 1.48 ± 0.33 s; syllable duty cycle = 8.7 %. The power spectrum shows a clear harmonic pattern of the end of the syllable, but the spectrum has a rather wide band; the dominant frequency is 13.6 ± 0.9 kHz and corresponds to the third frequency peak. inued.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B79DE0CFF10EE0FA181FED4.taxon	description	(Figs 4, 5 L, 9 J, 12 E, 13 I, 14 I, 35, 36)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B79DE0CFF10EE0FA181FED4.taxon	materials_examined	Type material. Male holotype: Indonesia. Lampung Province, Wai Lima [estate] Z. [uid] Sum [atra], Lampongs [= Lampung], XI – XII. 1921, No # 222, identified Cardiodactylus novae guineae Haan by L. Chopard, [H. H.] Karny & [H. C.] Siebers (MZB-ORTH 10432). Female allotype: same information as HT, No # 376, identified Cardiodactylus novae guineae Haan by L. Chopard, [H. H.] Karny & [H. C.] Siebers (MZB-ORTH 10440). Paratypes (2 ♂, 6 ♀): same information as HT: 2 ♀, No # 435, 460, [H. H.] Karny & [H. C.] Siebers (MZB-ORTH 10444,10439); 1 ♂, No # 158, [H. H.] Karny & [H. C.] Siebers (MNHN-EO-ENSIF 3493); 1 ♀, No # 395 (MZB-ORTH 10442), identified Cardiodactylus novae guineae Haan by L. Chopard, [H. H.] Karny & [H. C.] Siebers (MNHN-EO-ENSIF 3494). Indonesia. Sumatera Selatan Province, Sumatra I., environs of Banding Agung Vill [age], on lake Ranau Danau, S 04 ° 48,695 ’ E 103 ° 55.289 ’, 600 – 700 m, secondary forest, on leaf of bush, at night, 19 – 22. IV. 2009, A. Gorochov, M. Berezin, E. Tkatsheva: 1 ♂, 1 ♀ (ZIN), 1 ♀ (MNHN-EO-ENSIF 3495). Indonesia. Bengkulu Province, Sumatra I., environs of Curup Town (not far from Bengkulu City), S 03 ° 28 – 29 ’, E 102 ° 31 – 38 ’, 1000 – 1500 m, secondary forest, on leaf of bush, at night, 24. IV – 2. V. 2009, A. Gorochov, M. Berezin, E. Tkatsheva (ZIN). Type locality. Indonesia, Lampung Province, South Sumatra, Wai Lima estate. Other material examined. Malaysia: Str. [Straight] Malacca, Eil. Berhala [Berhala I.], VIII. 1927, J. C. v. d. Meer Mohr, 1 ♀ (MNHN), 1 ♀ (MZB).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B79DE0CFF10EE0FA181FED4.taxon	etymology	Etymology. Named after the type locality.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B79DE0CFF10EE0FA181FED4.taxon	distribution	Distribution. Southern part of Sumatra and islands in Malacca Straight.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B79DE0CFF10EE0FA181FED4.taxon	diagnosis	Diagnosis. Species of average size, close in shape, coloration and male venation to C. jdoeria, but more similar to C. singapura and C. pelagus in terms of male genitalia, with parallel carinated dorsal ridges and apex of pseudepiphallic sclerite trapezoidal (instead of rounded). The species differs from C. reticulatus in the coloration of tegmina less contrasted (including crossveins between M and R not remarkably lighter), head and legs distinctly less spotted, pseudepiphallus wider and with proximal corners of high parts of dorsal ridges rounded (not angular).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B79DE0CFF10EE0FA181FED4.taxon	description	Description. Average size for the species group. General coloration little contrasted, yellowish brown with dark areas. Head dorsum yellow brown with 4 dark brown bands, 2 wide lateral ones with an anterior notch fused to fastigium dark brown coloration, 2 median bands punctuated; area posterior to eyes yellow brown with 2 short dark brown lines (Fig. 5 L). Scapes yellow brown, their base darker; antennae orange brown with yellowish rings. Face yellow brown; HT and paratypes from Sumatra Selatan Province with a light yellow transverse band ventral to eyes. Lateral side of head yellow brown with a dark brown band posterior to eyes. Mouthparts and maxillary palpi mostly yellow brown. Pronotum: Dorsal disk lateral margins yellow brown; anterior area yellow brown mottled with dark brown; posterior area dark brown. Lateral lobes dark brown, ventral margins yellow brown. Legs I – II yellow brown, femora with faint orange brown spots and tibiae with faint orange brown rings. Legs III completely orange brown including knees. Tarsomeres III- 1 yellow brown. Hind wing tail dark brown, 1.5 longer than pronotum. Cerci yellow brown, with dark brown spots basally then with dark brown rings. Abdomen: Sternites yellow brown, tergites mostly dark brown with discontinuous longitudinal yellow lines. Male: FW coloration little contrasted, mostly dark brown with orange brown veins (9 J). Anterior area external part of cell e 1 black. Sclerotized area near base of chords orange brown. Areas with yellow veins and translucent cells include bases of anal veins, anterior third of CuA, harp veins, part of chords and a small triangular area posterior to mirror. M / R / Sc area and veins orange brown. Sc projections and more ventral veins of lateral field yellow brown, cells between them dark brown. FW venation (Fig. 9 J): 1 A very slightly bisinuated; CuP missing but claval fold present in harp. Stridulatory file in HT with 161 teeth on the transverse part of 1 A and 51 teeth on the longitudinal part, without discontinuity at the level of the angle. Harp with 2 w-shaped veins. Mirror area: mirror (d 1) longer than wide, oval, separated near mid-length by a transverse vein, the posterior part slightly shorter than the anterior one, triangular with few accessory veins; d 2 as wide as d 1 with many accessory veins; cell e 1 long. Apical field with 4 – 5 cell alignments posterior to mirror (n = 2). Lateral field with 8 (n = 2) projections of Sc and 3 – 4 (n = 2) ventral veins. Male genitalia (Figs 35 E – G, 36 H – K): Pseudepiphallus little setose. Dorsal ridges almost parallel or slightly divergent posteriorly, their posterior inner margin carinated, with short inner expansions. Pseudepiphallic sclerite with very short latero-anterior expansions, membrane between them densely setose. Posterior pseudepiphallic apex narrowed at mid-length, trapezoidal. Rami parallel, little sclerotized, apex sliglthly convergent, ended by a thin expansion. Ectophallic arc complete, curved. Bases of ectophallic apodemes with a posterior membranous lobe and a long anterior sclerotization. Ectophallic fold with weak lateral sclerites. Endophallic sclerite longer than wide, well sclerotized, with short posterior arms and a triangular posterior expansion. Endophallic apodeme with small lateral lamellas and a strong medio-dorsal crest. Membrane of endophallic cavity finely plicate. Female: FW brown (cells and veins), except bases of anal veins yellow brown. Anterior half of CuA, M and related accessory veins yellow brown. R / Sc area and veins dark orange brown. Sc projections and more ventral veins yellow brown, the cells between them dark brown. FW venation (Fig. 12 E): 9 – 10 (m = 9.5, n = 4) strong longitudinal veins on dorsal field; lateral field with 8 (m = 8, n = 4) longitudinal veins including 5 Sc projections and 3 ventral veins. Ovipositor of average length, shorter than FIII, apex with both dorsal and ventral edges denticulate (Fig. 13 I). Female genitalia (Fig. 14 I): Copulatory papilla triangular with baso-lateral sclerites; apex rounded and sclerotized, folded ventrally. Juvenile: Unknown. Measurements. See Table 7. Habitat and life history traits. Unknown. Behavior. Unknown.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B7DDE0EFF10EC30A181FB84.taxon	description	(Figs 4, 5 M, 9 H, 11 E, 13 J, 14 J, 37)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B7DDE0EFF10EC30A181FB84.taxon	materials_examined	Type material. Male holotype: Indonesia. North Sulawesi Province. Sangi en Talaud [Sangihe and Talaud Islands, Karakelong Island], Lobo, sol. Doeata [?], Erie, 1926 (MZB-ORTH 10632). Female allotype: same information as HT (MZB-ORTH 10629). Paratype (1 ♀): same information as HT, MZB-ORTH 10630 (MNHN-EO-ENSIF 3652). Type locality. Indonesia, Karakelong Island (Talaud Islands).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B7DDE0EFF10EC30A181FB84.taxon	etymology	Etymology. Named after the type locality.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B7DDE0EFF10EC30A181FB84.taxon	distribution	Distribution. Indonesia, Karakelong Island.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B7DDE0EFF10EC30A181FB84.taxon	diagnosis	Diagnosis. Species of large size, coloration brownish with yellow brown patterns, characterized by shape of pseudepiphallic sclerite in male genitalia with almost parallel dorsal ridges, slightly asymmetrical, close to C. gagnei Otte, 2007 a and C. chessmani Otte, 2007 a from Vanuatu.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B7DDE0EFF10EC30A181FB84.taxon	description	Description. Large size for the species group, especially for females. General coloration dark brown with yellow areas. Head dorsum almost homogeneously dark brown (Fig. 5 M). Fastigium dark brown to black. Scapes yellow brown, dark brown basally. Antennae brown. Face mostly brown, with 2 parallel dark brown stripes on front part of fastigium (not visible in HT) and a yellow transverse band below eyes. Mouthparts yellow brown. Maxillary palpi yellow brown, darker apically. Pronotum: Dorsal disk dark brown faintly mottled with yellow brown, the 4 corners yellow; lateral lobes almost entirely dark brown, ventral corners yellow brown. Legs homogeneously orange brown, hind knees dark brown. Hind wing tail dark brown. Male: FW coloration mostly brown with yellowish areas. Yellow areas include: bases of CuA, veins 2 A – 4 A, transverse part of 1 A, chord veins, anterior part of diagonal vein, harp veins, and oval area posterior to mirror (translucent cells). Orange brown sclerotized area near base of chords. Lateral field: M / R area dark brown, veins orange brown; Sc vein and projections yellow, cells between them dark brown. FW venation (Fig. 9 H): 1 A slightly bisinuated. CuP missing. Stridulatory file with 221 teeth on the transverse part of the file, and 13 on a ventral bump near base of 1 A. Harp with 2 w-shaped harp veins. Mirror area: mirror (d 1) longer than wide, oval, well defined, crossed by an accessory vein near mid-length, the posterior part crossed by several transverse veins; d 2 not distinctive; e 1 crossed by transverse accessory veins. Apical field with 5 cell alignments posterior to mirror (n = 1). Lateral field with 10 projections of Sc and 4 more ventral veins. Subgenital plate dark brown. Male genitalia (Fig. 37 EG): Pseudepiphallic dorsal ridges almost parallel, their dorsal edges setose, carinated on dorso-external edges and slightly asymmetrical, the left one thicker; forming a homogeneous curve in lateral view (Fig. 37 G). Pseudepiphallic sclerite with wide latero-anterior expansions, membrane between them not setose. Posterior region of pseudepiphallus rounded, with a small rounded translucent area. Posterior lobe of Pseudepiphallic parameres curved externally, their bases very sclerotized. Rami strong, apical expansion oriented posteriorly. Ectophallic arc complete, curved, without a posterior expansion. Ectophallic fold membranous, with weak lateral sclerites. Endophallic sclerite with short posterior arms and a wide posterior expansion. Endophallic apodeme with a large medio-dorsal crest and lateral lamellas. Membrane of endophallic cavity plicate. Female: Size large for the genus. FW coloration dark brown (cells and veins), their bases yellow brown, including bases of anal veins and CuA and first quarter of M. CuP orange brown, very strong. Yellow brown area near FW mid-length including part of CuA and M and surrounding cells and transverse veins. R / Sc area and veins dark orange brown, with many transverse veins; posterior part of Sc yellow brown. Projections of Sc and more ventral veins yellow, their bases dark brown, the cells between them dark brown. FW venation (Fig. 11 E): 13 – 14 (n = 2) strong longitudinal veins on dorsal field; lateral field with 10 – 11 (n = 2) longitudinal veins including 6 projections of Sc and 4 – 5 ventral veins. Ovipositor long, apex with both dorsal and ventral edges denticulate, ventral valves with two preapical notches (Fig. 13 J). Female genitalia (Fig. 14 J): Copulatory papilla trapezoidal, ventral part raised, apex slightly curved ventrally and sclerotized. Juvenile: Unknown. Measurements. See Table 8. Habitat and life history traits. Unknown. Behavior. Unknown.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B7FDE72FF10EEDFA217FDD1.taxon	description	(Figs 4, 6 A, 10 A, 11 G, 13 K, 15 A, 38, 39, 40)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B7FDE72FF10EEDFA217FDD1.taxon	materials_examined	Type material. Male holotype: Indonesia. West Nusa Tenggara Province, Lombok, Joben district, Rinjani National Park, forêt de Joben, 08 ° 31 ' 04.7 " S 116 ° 23 ' 33.4 " E, 870 m, 29. VI. 2010, TR 393, jour, litière, T. Robillard (MZB). Female allotype: Lombok, Tetebatu district, Rinjani National Park, 29. VI. 2010, morte en élevage, T. Robillard (MZB). Paratypes (5 ♂, 1 ♀): Lombok, Tetebatu district, Rinjani National Park, forêt de Jeruk Manis, 08 ° 30 ' 39.7 " S 116 ° 25 ' 23 " E, 848 m, 29. VI. 2010, T. Robillard: 1 ♂ (TR 403), jour (5 PM), sur plante, enregistrement appel, photo TR (MNHN-EO-ENSIF 3673); 1 ♂ (TR 404), nuit, sur plante en lisière (h = 80 cm), enregistrement appel (MNHN-EO-ENSIF 3672). Lombok, Tetebatu district, Rinjani National Park, 08 ° 30 ' S 116 ° 25 ' E, 29. VI. 2010, morts en élevage, T. Robillard: 1 ♂ (TR 446) (ZIN), 2 ♂ (TR 447,?) (MZB), 1 ♀ (MNHN-EO-ENSIF 3674). Type locality. Indonesia, Lombok, Rinjani National Park. Other material examined. Indonesia. West Nusa Tenggara Province, Lombok, Tetebatu district, Rinjani National Park, 08 ° 30 ' S 116 ° 25 ' E, 29. VI. 2010, 2 juveniles, morts en élevage, T. Robillard (MNHN, MZB). Lombok, Pusuk forest, 550 m, 9 – 13. X. 1991, 1 ♀, Krikken, Huijbregts & de Vries (RMNH). Sumbawa, Batu Dulang, forêt au S du village, sommet 1 au-dessus du village, 08 ° 36 ' 56.4 ’’ S 117 ° 15 ' 58.4 ’’ E, 26 – 27. VI. 2010 (GPS 2), zone ouverte herbacée, 26 – 27. VI. 2010, T. Robillard: 1 ♂ (TR 375), nuit, sur plante (h = 1.3 m), enregistrement appel (MNHN-EO-ENSIF 3675); 1 ♂ (TR 376), nuit, sur plante (h = 80 cm), enregistrement appel (MNHN-EO-ENSIF 3677). Batu Dulang, forêt au S du village, 08 ° 37 ' S 117 ° 15 ' E, 26 – 27. VI. 2010, mort en élevage, T. Robillard, morts en élevage: 2 ♂ (TR 390 - TR 391) (MZB); 1 ♂ (Male 1 - F 0), enregistrement appel en captivité (MNHN-EO-ENSIF 3676); 2 ♂ (Male 2 - F 0, Male 3 - F 0), enregistrement appel en captivité (MNHN-EO-ENSIF 3678 - 3679); 1 ♀ (MZB); 3 juveniles (MZB); 2 juveniles (MNHN).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B7FDE72FF10EEDFA217FDD1.taxon	etymology	Etymology. Named after the contraction of the type locality and island (Lombock and Rinjani National Park).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B7FDE72FF10EEDFA217FDD1.taxon	distribution	Distribution. Indonesia, Lombok and Sumbawa Islands.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B7FDE72FF10EEDFA217FDD1.taxon	diagnosis	Diagnosis. Species of average size, general coloration brown, little contrasted. Characterized by male pseudepiphallic dorsal ridges showing wide expansions folded innerly, close to that of C. vella Otte, 2007 a and C. riga.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B7FDE72FF10EEDFA217FDD1.taxon	description	Description. Average size for the species group. General coloration brown, little contrasted. Head dorsum yellow brown with 4 brown bands including 2 wide lateral ones, fused to the dark brown fastigium, and 2 median punctuated bands; area posterior to eyes with dark brown lines (Fig. 6 A). Scapes yellow brown, with a transverse dark brown band. Antennae orange brown. Front part of fastigium yellow, sometimes with 2 dark brown spots. Lateral side of head mostly light brown. Face yellow brown, with a faint whitish transverse band ventral to eyes. Mouthparts whitish; maxillary palpi yellow brown. Pronotum: Dorsal disk mostly yellow brown mottled with dark brown; posterior area dark brown. Lateral lobes dark brown dorsally, ventral part yellow brown to whitish. Legs I – II orange brown, femora with faint dark brown spots and tibiae with rings. FIII homogeneously yellow to orange brown, knees dark brown; TIII dark brown, with faint yellow brown rings. Tarsomeres III- 1 dark brown. Hind wing tail dark brown. Cerci yellow brown, faintly mottled with dark brown near base, ringed at apex. Abdomen mostly yellow brown dorsally; lateral and dorsal sides dark brown with discontinuous yellow lines on posterior tergites. Subgenital plate yellow brown. Male: FWs brown with yellow and dark brown patterns (Fig. 10 A). Yellow brown to whitish areas including bases of anal veins and of CuA, harp veins, part of chords and a wide band posterior to mirror. Black spot near base of 1 A and CuA. Bases of M, R and Sc yellow brown, then dark orange brown, including veins and areas between them; apex of Sc whitish. Projections of Sc and more ventral veins of lateral field yellow brown, their bases brown. FW venation (Fig. 10 A): 1 A clearly bisinuated. CuP absent. Stridulatory file with about 240 teeth on the transverse and curved parts of 1 A, and 21 more teeth on top of a bump near 1 A base (Table 9). Harp with 2 w-shaped veins. Mirror area: mirror (d 1) longer than wide, its posterior region variably crossed by longitudinal and transverse accessory veins; d 2 as wide as mirror, variable. Cell e 1 crossed at mid-length by an accessory vein. Apical field with 3 – 4 cell alignments posterior to mirror (m = 4, n = 4). Lateral field with 8 – 10 projections of Sc (m = 9; n = 4) and 4 – 5 ventral veins (m = 4; n = 4). Male genitalia (Fig. 38 E – H): Pseudepiphallus shaped as a wide gutter at the level of the dorsal ridges, almost closed dorsally by wide inner expansions of the ridges touching each other posteriorly (Fig. 38 H). Pseudepiphallic sclerite with large triangular anterior expansions, the membrane between them setose. Posterior pseudepiphallic apex rounded, with a long oval translucent area. Rami with short convergent apical stems. Ectophallic arc complete, without posterior expansion. Ectophallic fold with weak lateral sclerites; apex trilobate, membranous, median lobe oval. Endophallic sclerite long, with a rectangular posterior expansion and short lateral arms. Endophallic apodeme with a dorsal crest and wide lateral lamellas. Membrane of endophallic cavity finely plicate. Female: FW coloration dark brown (cells and veins), their bases yellow, and with a whitish area on external edge at mid-length. CuA / M area whitish on basal part. R orange brown. Sc dark brown, apex whitish, Sc projections and more ventral veins yellow, the cells between them dark brown. FW venation (Fig. 11 G): dorsal field with 12 (n = 2) strong longitudinal veins; lateral field with 10 – 11 (n = 2) longitudinal veins including 7 projections of Sc and 3 – 4 more ventral veins. Ovipositor of average length, apex with both dorsal and ventral edges denticulate (Fig. 13 K). Female genitalia (Fig. 15 A): Copulatory papilla triangular with baso-lateral sclerotizations; apex rounded, slightly sclerotized and folded ventrally. Juvenile: Brown, little contrasted, head and legs with same pattern of coloration as adults (Fig. 39 D). Variation. One female specimen from Pusuk forest may belong to another species according to its larger size (Table 9). The specimens from Sumbawa Island are slightly smaller than that from Lombok (type series), but they differ only by details of male genitalia. In particular the region posterior to the pseudepiphallic dorsal ridges seems smaller and the inner triangular expansions of the dorsal ridges are slightly stockier, and the endophallic sclerite tends to be smaller. However, these differences are not stable among the specimens of each island, so only one species is distinguished here. Measurements. See Table 9. Habitat and life history traits. Cardiodactylus lombrinjani is a nocturnal species living both in forested areas and in / or near secondary habitats such as bushes and hedges. Males produce calling songs at night, from tree branches and bushes (Fig. 39). During the day, couples have been observed mating under the bark of trees and several specimens have been found in blocks of aerial litter. Juveniles are found day and night on low vegetation. Behavior. Calling song (Fig. 40): Cardiodactylus lombrinjani has a mono-syllabic calling song. At 23 ° C, the calling songs show a very indented amplitude profile. Syllables have the following characteristics: syllable duration = 193 ± 12 ms; syllable period = 18.2 ± 11 s; syllable duty cycle = 1.6 %. The power spectrum consists of a rather broad band dominating at 13.94 ± 0.37 kHz which corresponds to the third frequency peak of the song despite harmonic pattern is unclear.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B02DE75FF10ED73A4F9FE61.taxon	description	(Figs. 4, 6 B, 10 B, 12 B, 13 L, 15 B, 41)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B02DE75FF10ED73A4F9FE61.taxon	materials_examined	Type material. Male holotype: Indonesia: Malaka Province, Ceram, Piroe [Seram I., Piru], II. 1909, F. Muir (BPBM) (not examined). Type locality. Indonesia, Seram Island, Piru. Material examined. Indonesia. Malaka Province, Ceram, Piroe [Seram I., Piru], 1 ♂ (MNHN); Ceram [Seram I.], Januar-Februar [I – II.] 1910, 1 ♀ (MNHN); Ceram I. [Seram I.], 1 ♀, determined Cardiodactylus novaeguineae by L. Chopard (MZB. ORTH 10448); Seram [I.], Ruhuwa, 30 km E. of Amahai, 16. VI. 1970, R. F. Ellen (BMNH 1972 - 282).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B02DE75FF10ED73A4F9FE61.taxon	distribution	Distribution. Indonesia, west and south of Seram Island.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B02DE75FF10ED73A4F9FE61.taxon	diagnosis	Diagnosis. Species of large size, close to C. buru n. sp., characterized by coloration of male FW with lateral field whitish (brown to dark brown in C. muiri) and with yellowish base and wide yellowish band posterior to mirror; characteristic male genitalia made of 3 levels in lateral view, the most anterior level of dorsal ridges, the highest, forming small triangular carinae in dorsal view.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B02DE75FF10ED73A4F9FE61.taxon	description	Redescription. In addition to the few characters given by Otte (2007 a): size large for the species group, coloration brown with contrasted yellowish patterns (Fig. 41). Head dorsum almost entirely dark brown, with 4 wide brown bands almost fused together (Fig. 6 B); fastigium dark brown; yellow lines posterior to eyes. Scapes dark brown. Face mostly brown, with sometimes a yellow transverse band (not visible in some old specimens). Pronotum: Dorsal disk yellow laterally, median area mottled symmetrical orange brown patterns, its posterior edge dark brown; lateral lobes mostly dark brown, their ventral margin yellowish, the anterior region with a wider yellow margin. Legs yellow brown, femora mottled with orange brown, tibiae with orange brown rings; FIII knees dark brown. Cerci yellow brown, faintly mottled with brown. Male: FW coloration brown with the following large yellowish areas (Fig. 10 B): base of FWs including bases of anal veins, CuA, M, R and Sc veins, region anterior to file near plectrum, part of chords, harp veins and a wide band posterior to mirror. Bases of chords with an orange brown sclerotization. Lateral field whitish, M / R / Sc area and cells orange brown; posterior region of Sc and its rojections and more ventral veins whitish, the membrane between them translucent. FW venation (Fig. 10 B): 1 A slightly bisinuated. CuP absent. Harp with 2 w-shaped veins (variably in shape, the male from south Seram having angular veins and the male from Piru having sinuated veins). Mirror area: mirror (d 1) oval, twice longer than wide, its posterior region made of a variable number of small cells; d 2 not distinctive from the rest of D aligment. Apical field with 4 or 5 (n = 2) cell alignments posterior to mirror. Lateral field with 8 – 9 projections of Sc (n = 2) and 5 more ventral veins (n = 2). Male genitalia: Pseudepiphallus narrowed at mid-length with clear lateral shoulders; posterior region widened preapically; apex rounded. In lateral view, pseudepiphallus made of 3 levels, the most anterior level of dorsal ridges, the highest, forming small triangular carinae in dorsal view; posterior part of dorsal ridges forming parallel crests slightly divergent and assymetrical. Female: FW coloration mostly brown with a whitish spot at mid-length. Base of FWs yellow brown, including bases of anal veins, CuA and anterior half of M. CuA / M / R / Sc areas brown, including veins; Sc apex and projections whitish. R / Sc area and veins orange brown. Projections of Sc and more ventral veins yellow, the cells between them translucent. FW venation (Fig. 12 B): 11 (n = 2) strong longitudinal veins on dorsal field; lateral field with 11 (n = 2) longitudinal veins including 6 – 7 projections of Sc and 4 – 5 more ventral veins. Ovipositor long, apex with both dorsal and ventral edges slightly denticulate (Fig. 13 L). Female genitalia (Fig. 15 B): Copulatory papilla trapezoidal, with large baso-lateral sclerites; apex rounded, slightly sclerotized and finely plicate, slightly folded ventrally. Juvenile: unknown. Measurements. See Table 10.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B02DE75FF10ED73A4F9FE61.taxon	discussion	Remark. The male specimen from the south of Seram (BMNH) is smaller than the males from Piru in western Seram (HT and male examined in MNHN), but it differs otherwise only by the more angular shape of the harp veins; both general coloration and male genitalia are consistent with the holotype of the species.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B04DE7AFF10EC5FA17DFE1C.taxon	description	(Figs 4, 6 C, 10 C, 11 H, 13 M, 15 C, 42, 43, 44)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B04DE7AFF10EC5FA17DFE1C.taxon	materials_examined	Type material. Male holotype: Indonesia. Central Java Province, Mt. Moeria [Mount Muria], # 9, identified Cardiodactylus novaeguineae by L. Chopard (MNHN-EO-ENSIF 1199). Female allotype: same information as HT, (MNHN-EO-ENSIF 1211). Paratypes (5 ♂, 9 ♀): same information as HT, 3 ♀, identified C. gaimardi by L. Chopard (MNHN-EO-ENSIF 1209, 1210, 1222); Mont Muria, NW village de Colo, forêt secondaire, 06 ° 39 ' 40 " S 110 ° 53 ' 47.9 " E, 1027 m, 9. VI. 2010, 1 ♀ (TR 200), nuit, copulation entre 2 feuilles sur arbuste (h = 1.2 m, 22 h 30) (molecualr sample Cmur) (MNHN-EO-ENSIF 3481); Mont Muria, NW village de Colo, sentier bordé de cultures, 06 ° 39 ' 36.1 " S 110 ° 53 ' 47.3 " E, 1021 m, 8. VI. 2010, 1 ♀ (TR 201), nuit, arbuste (h = 1.5 m) (MZB); Mont Muria, NW village de Colo, Mont Muria, forêt secondaire et chemin en bord de cultures, 06 ° 39 ' 18.3 " S 110 ° 53 ' 46.2 " E, 1115 m, 8 – 9. VI. 2010: 1 ♂, 3 ♀, adultes en élevage (MZB); 1 ♀, 1 ♂, adultes en élevage (ZIN); 1 ♀, adulte en élevage (MNHN-EO-ENSIF 3482); 3 ♂, adultes en élevage, enregistrement appel (MNHN-EO-ENSIF 3478 - 3480). Type locality. Indonesia, Java, Mount Muria. Other material examined. Indonesia. Central Java Province, Mont Muria, NW village de Colo, forêt secondaire, souche élevage, 4 ♂, F 1, enregistrement appel (MNHN-EO-ENSIF 3483 - 3487); 3 ♂, F 2 (MNHN-EO-ENSIF 3488 - 3490).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B04DE7AFF10EC5FA17DFE1C.taxon	etymology	Etymology. Named after the type locality.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B04DE7AFF10EC5FA17DFE1C.taxon	distribution	Distribution. Indonesia, northern part of Central Java.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B04DE7AFF10EC5FA17DFE1C.taxon	diagnosis	Diagnosis. Species of average size, general coloration dark brown with yellow patterns, with a large yellow crescent-shaped area posterior to mirror. Female with a light band on latero-anterior part of FW. Characterized by male genitalia with folded asymmetrical dorsal ridges and female copulatory papilla with a ventral crest.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B04DE7AFF10EC5FA17DFE1C.taxon	description	Description. Average size for the species group. General coloration brown to dark brown with contrasted vivid yellow areas (Fig. 42 A – D). Head dorsum with a large brownish area comprising short dark brown lines posterior to eyes and 4 more or less distinct dark brown bands, faint posteriorly, the lateral bands complete, the median ones punctuated (Fig. 6 C). Fastigium dark brown to black. Scapes orange brown with a transverse whitish ring. Antennae brown. Face most often brown with a thin yellow transverse band ventral to eyes. Mouthparts mostly yellow brown. Lateral sides of head gray brown. Maxillary palpi yellow brown. Pronotum: Dorsal disk anterior and lateral areas yellow with variable dark brown symmetrical patterns and black spots; posterior area dark brown to black. Lateral lobes dark orange brown, ventral margins yellow brown. Legs I – II yellow brown to orange brown. FIII yellow brown to orange brown; knees dark brown; tibiae faintly ringed with dark brown at spur level; tarsomeres III- 1 yellow brown with both ends dark brown. Hind wing tail dark brown to black, 1.5 longer than pronotum. Cerci brown with faint dark brown rings. Abdomen mostly dark brown. Male: FW coloration mostly dark brown to black with yellow areas including bases of anal veins (4 A completely yellow), angle of 1 A, first third of CuA, harp veins, chords, veins within a crescent-shaped area posterior to mirror. Translucent areas include harp anterior region, chord area and area posterior to mirror. M / R area dark brown, R / Sc area orange brown. Sc orange brown, its posterior part yellow. Projections of Sc and more ventral veins of lateral field yellow, cells between them dark brown. FW venation (Fig. 10 C): 1 A slightly bisinuated; CuP missing but claval fold present in harp. Stridulatory file with about 200 teeth on the transverse and curved parts of 1 A without discontinuity, and 15 more teeth on top of a bump near 1 A base (Table 11, n = 4). Harp with 2 w-shaped veins. Mirror area: mirror (d 1) longer than wide, oval, crossed near mid-length by a transverse vein, the posterior part variably subdivided by an accessory veins; d 2 as wide as d 1, variably subdivided by accessory veins; e 1 crossed by several accessory veins. Apical field with 4 – 5 cell alignments posterior to mirror (n = 4). Lateral field with 7 – 9 (m = 8, n = 5) projections of Sc and 4 – 5 (m = 4.2, n = 5) ventral veins. Subgenital plate yellow brown. Male genitalia (Fig. 42 E – G): Region of pseudepiphallus posterior to mid-length narrowing almost rectangular. Dorsal ridges divergent posteriorly, their anterior part with dorsal asymmetrical expansions folded laterally, the left one larger and more anterior, the right one smaller and more posterior; pseudepiphallus slightly bisinuated laterally. Pseudepiphallic sclerite with flat and wide latero-anterior expansions, membrane between them with thick dense setae. Posterior pseudepiphallic apex rounded, with a small rounded translucent area. Rami parallel, with large preapical plates, apical stems slightly convergent. Pseudepiphallic parameres median lobe rectangular. Ectophallic arc complete, narrow and curved, without posterior expansion. Bases of ectophallic apodemes with a posterior membranous lobe but with no clear anterior sclerotization. Ectophallic fold with weak lateral sclerites; apex trilobate, membranous, median lobe oval. Endophallic sclerite well sclerotized, with long posterior arms and a triangular posterior expansion. Endophallic apodeme with wide lateral lamellas and a strong medio-dorsal crest. Membrane of endophallic cavity smooth. Female: FW coloration orange brown to dark brown, including veins and cells (Fig. 11 H). Base of FWs whitish, including bases of anal veins. Whitish lateral band on FW anterior part, including anterior half of CuA and M and associated transverse veins, their posterior half orange brown. R / Sc area orange brown to dark brown, veins orange brown. Projections of Sc and more ventral veins yellow brown, the cells between them dark brown. FW venation (Fig. 11 H): 10 – 11 (m = 10.2, n = 5) strong longitudinal veins on dorsal field; lateral field with 7 – 9 (m = 8.2, n = 5) longitudinal veins including 4 – 5 projections of Sc and 3 – 4 ventral veins. Ovipositor long, apex with both dorsal and ventral edges denticulate (Fig. 13 M). Female genitalia (Fig. 15 C): Copulatory papilla trapezoidal with baso-lateral sclerotizations; apex membranous, folded ventrally and related to a medio-ventral crest; apex with a posterior sclerotized area near base. Juvenile: Light brown mottled with dark brown. Measurements. See Table 11. Habitat and life history traits. Cardiodactylus muria is a nocturnal species living in or near secondary forested areas. Males produce calling songs at night, from tree branches and leaves (Fig. 43). Juveniles are found day and night on low vegetation. Behavior. Calling song (Fig. 44): High speed video observations at 1250 frames per second show that each calling song corresponds to only one FW closure (i. e. one syllable), the gaps within the syllables are made by the jerking closing movement of the FWs. At 20 ° C, these mono-syllabic calling songs are emitted in irregular groups of 7 – 12 (m = 10.6 ± 2.2, n = 5) that lasts for 77 ± 13 s with a period of 208 + / - 50 s. Each syllable shows an indented temporal profile due to the jerking FW closures, with the following characteristics: syllable duration = 48 ± 2 ms; syllable period (within the echemes / syllable groups) = 8.0 ± 2.3 s; syllable duty cycle = 0.65 %. The dominant frequency is 13.28 ± 0.22 kHz and corresponds to the third frequency peak of the song, with a clear harmonic series.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B0BDE7CFF10E998A181FD8C.taxon	description	(Figs 4, 6 D, 10 D, 12 C, 13 N, 15 D, 45)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B0BDE7CFF10E998A181FD8C.taxon	materials_examined	Type material. Male holotype: Indonesia. Maluku Utara Province, Obi Latoe [Obi I.], Snellius Exp., 23 – 27. IV. 1930 (RMNH). Paratypes (2 ♀): Indonesia. Maluku Utara Province, Obi [Obi I.], 1904, 1 ♀, Rolle (ZIN); [Obi I.], Doherty, 1 ♀, ex coll. Fruhstorfer (MHNG). Type locality. Indonesia, Obi Island.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B0BDE7CFF10E998A181FD8C.taxon	etymology	Etymology. Named after the type locality.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B0BDE7CFF10E998A181FD8C.taxon	distribution	Distribution. Indonesia, Obi Island.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B0BDE7CFF10E998A181FD8C.taxon	diagnosis	Diagnosis. Species of average to large size, close to C. manus Otte, 2007 a by the male genitalia but differing by coloration (male and female FWs, lateral lobes of pronotum mostly dark brown) and details of male genitalia.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B0BDE7CFF10E998A181FD8C.taxon	description	Description. Size average to large for the species group, shape stocky (Fig. 45 A – D). General coloration contrasted. Head dorsum almost entirely dark brown (Fig. 6 D), with 4 wide brown bands almost fused together separated by faint yellow lines; fastigium dark brown; yellow lines posterior to eyes. Scapes yellow brown, with a dark brown ring; rest of antennae orange brown. Front part of fastigium yellow with 2 dark brown spots. Face and mouthparts yellowish to brown, with a transverse dark brown band bellow antennae and a yellow band bellow eyes. Lateral side of head mostly dark brown dorsally, progressively lighter ventrally; eyes underlined by a yellow line ventrally. Pronotum: Dorsal disk almost homogeneously brown, anterior corners yellow, posterior edge dark brown. Lateral lobes mostly dark brown, ventral margin yellowish. Legs almost homogeneously orange brown, FIII knees darker. Hind wing tail dark brown with yellow transverse veins. Cerci light brown faintly mottled with brown. Abdomen mostly dark, with yellow stripes laterally. Subgenital plate yellow brown. Male: FW coloration dark brown with the following whitish areas (Fig. 10 D): base of FWs including bases of anal veins and CuA, wide transverve band posterior to mirror, part of chords, 1 A (file) and harp veins. Orange brown sclerotized area near base of chords. M and R orange brown, the area between dark brown; Sc orange brown, its projections whitish, the membrane between them dark brown. FW venation (Fig. 10 D): 1 A clearly bisinuated. CuP absent. Harp with 2 w-shaped veins, and a small straight anterior one. Mirror area: mirror (d 1) oval, distinct, crossed at 2 / 3 by a transverse vein; d 2 not distinct. Apical field with 4 cell alignments posterior to mirror (n = 1). Lateral field with 9 projections of Sc and 4 more ventral veins. Male genitalia (Fig. 45 E – G): Pseudepiphallus as in C. manus, shaped as a wide gutter. Apex of dorsal ridges with short setae, carinated innerly and folded laterally, the lateral triangular expansions clearly asymmetrical, the left one larger and more posterior, the right one smaller and more anterior. Dorsal ridges almost convex in lateral view. Pseudepiphallic sclerite with wide rectangular anterior expansions, slightly divergent, the membrane between them with thin setae. Rami with large preapical rectangular plates and with short globular convergent apical stems. Posterior pseudepiphallic apex rounded, with a triangular translucent area. Pseudepiphallic parameres with posterior arms thin and curved laterally. Ectophallic arc complete, curved posteriorly, without posterior expansion. Ectophallic fold with thin lateral sclerites; apex trilobate, membranous, median lobe oval. Endophallic sclerite small, with a wide posterior expansion. Endophallic apodeme with a short dorsal crest and lateral lamellas. Membrane of endophallic cavity smooth. Female: Base of tegmina yellow; dorsal field variable, one specimen with practically uniform brown coloration (ZIN) with only one lateral yellowish spot, the other female specimen with an additional yellowish elongate pattern in anterior part of tegmina (MHNG) (Fig. 12 C); lateral field of tegmina with proximal branches of Sc and crossveins between them yellowish white, distal branches of Sc and membranes between all Sc branches light brown (however R, stem of Sc, and area between them brown, as in female of C. buru n. sp.); tegminal venation with 11 – 12 longitudinal veins in dorsal field and with 8 – 9 branches on Sc and distinctly less numerous crossveins in lateral field. Ovipositor approximately equal or shorter to hind femur in length, its apex slightly denticulate on both dorsal and ventral edges (Fig. 13 N). Female genitalia (Fig. 15 D): Copulatory papilla longer than wide, trapezoidal, with elongate and basal sclerites mostly on dorsal side; apex elongate, not rounded, slightly sclerotized, folded ventrally. Juvenile: Unknown. Measurements. Length in mm. Body: male 27.2, female 21 – 32.2; body with wings: male 34.2, female 31 – 41.7; pronotum: male 3.5, female 3.3 – 4.2; tegmina: male 20, female 18 – 23.5; hind femora: male 22.8, female 19 – 24.2; ovipositor 17.5 – 18.5. Habitat and life history traits. Unknown. Behavior. Unknown.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B0DDE7FFF10EAE2A181FE1C.taxon	description	(Figs. 1, 4, 5 N, 9 I, 12 D, 13 O, 15 E, 46)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B0DDE7FFF10EAE2A181FE1C.taxon	materials_examined	Type material. Male holotype: Indonesia. South Sulawesi Province, Celebes [Sulawesi I.], VIII. 1930, Latimadjong Geb [irge], Oeroe [3 ° 25 ' S 120 ° 0 ' W] 800 m ([G.] Heinrich), identified Cardiodactylus novae guineae Haan by L. Chopard (1933) (MNHN-EO-ENSIF 1225). Female allotype: same information as HT (MNHN-EO-ENSIF 1224). Paratype: 1 ♀, same information as HT (MZB). Type locality. Indonesia, South Sulawesi, Latimadjong Gebirge, Oeroe [3 ° 25 ' S 120 ° 0 ' W], 800 m. Other material examined. Indones. South Sulawesi Province, no precision, VIII. 2007, 6 juveniles, T. Robillard (MZB), 6 juveniles, T. Robillard (MNHN). South East Sulawesi Province, Sulawesi Tengara, S. Sangona, foot Gn [Gunung = Mount] Matuwila, 150 m, disturbed forest, 3 ° 48 ° S 121 ° 39 ’ E, 30. X. 1989, 1 ♀, Van Tol (RMNH).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B0DDE7FFF10EAE2A181FE1C.taxon	etymology	Etymology. Species named after the type locality.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B0DDE7FFF10EAE2A181FE1C.taxon	distribution	Distribution. Indonesia, south and south-east of Sulawesi Island.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B0DDE7FFF10EAE2A181FE1C.taxon	diagnosis	Diagnosis. Species close to several species of the Novaeguineae species group, especially to Cardiodactylus frustorferi n. sp., C. guttulus and C. kondoi, but differing by its lighter coloration and male and female genitalia; the species is also close to C. contrarius n. sp. which has a lighter coloration of FWs and a longer pseudepiphallus.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B0DDE7FFF10EAE2A181FE1C.taxon	description	Description. Size average for the genus (Figs 1, 46 A – D). Head: Dorsum yellowish brown with four dark brown longitudinal bands, the median ones punctuated, and a brown triangle posterior to eye (5 N). Fastigium black, its margins yellow. Antennae, scapes and face almost homogeneously yellow brown. Lateral side of head gray brown. Maxillary palpi yellow brown, apex slightly darker. Pronotum: Dorsal disk yellow brown mottled with dark brown; lateral margins whitish, posterior margin dark brown. Lateral lobes dark brown, ventral margin yellow. Legs orange brown with few darker spots; hind knees dark brown. Tarsomeres III- 1 orange brown, darker at both ends, with 3 spines on dorso-external edges (n = 3). Hind wing tail gray brown, 1.5 times as long as the pronotum. Cerci yellow brown, with faint darker spots. Male: FW coloration (Figs 1, 9 I): Dorsal field mostly whitish and brown. Whitish areas include bases of 3 A – 4 A, transverse region of 1 A, harp veins, bases of chords, wide transverse band posterior to mirror and apex of anal region. Black areas between bases of 1 A and 3 A, e 1 cell and base of CuA. CuA / M area whitish and brown anteriorly. M and R orange brown anterior to their fusion, whitish posteriorly. Sc vein orange brown, its projections whitish; cells immediately ventral to Sc dark brown, more ventral part of lateral field whitish. FW venation (Fig. 9 I): 1 A slightly bisinuated. CuP absent. Harp with 2 w-shaped harp veins and 1 faint vein in anterior region. Mirror area: mirror (d 1) not rounded, subdivided by a transverse vein, posterior part small; d 2 little distinctive, separated into several longitudinal cells, longer than wide; e 1 cell not crossed by transverse vein. Apical field with 5 cell alignments. Lateral field with 9 projections of Sc (n = 1) and 4 more ventral longitudinal veins. Male genitalia (Fig. 46 E – G): Pseudepiphallic dorsal ridges parallel, their anterior edges forming a right angle with the rest of the sclerite. Pseudepiphallic sclerite with latero-anterior expansions, membrane between them with short setae. Posterior region rounded, widened laterally, with a small rounded translucent area. Rami wide. Ectophallic arc complete, with a short posterior expansion. Bases of ectophallic apodemes with long anterior expansions and short posterior ones. Apex of ectophallic fold trilobate, laterally sclerotized; median lobe rounded, sclerotized basally. Endophallic sclerite wider than long, with a short anterior expansion; lateral arms fused with lateral sclerites of ectophalic fold. Endophallic apodeme with lateral lamellas and a narrow medio-dorsal crest. Membrane of endophallic cavity plicate. Female: FW coloration (Fig. 12 D): mostly dark brown. External margin of dorsal field successively dark brown and whitish, starting from base of longitudinal veins whitish. Inner part of dorsal field dark brown, except a translucent area at one third of FW length. M / R area brown with whitish veins until FW mid-length, with a whitish sclerotization at mid-length, as in males, then dark brown until posterior margin. R / Sc area and veins orange brown, projections of Sc and more ventral veins whitish. Posterior angle white, including distal parts of R and Sc. FW venation (Fig. 12 D): dorsal field with 10 (n = 2) strong longitudinal veins; lateral field with 9 – 11 (n = 2) longitudinal veins including 5 – 6 projections of Sc and 4 – 5 more ventral veins. Ovipositor: Shorter than FIII; apex with both dorsal and ventral edges denticulate (Fig. 13 O). Female genitalia (Fig. 15 E): Copulatory papilla triangular, with baso-lateral sclerites; apex rounded, sclerotized and folded ventrally. Juvenile: Contrasted coloration (Fig. 47 A), mostly light brown and yellow brown mottled with dark brown spots. Head coloration as in adults; pronotum yellow brown with few dark brown spots. Measurements. See Table 12. Habitat and life history traits. Cardiodactylus oeroe has been observed as juveniles only, at night on low plants and branches in open forested areas (Fig. 47). Behavior. Unknown.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B0EDE61FF10E99AA181FC89.taxon	description	(Figs 4, 6 E, 48)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B0EDE61FF10E99AA181FC89.taxon	materials_examined	Type material. Male holotype: Philippines. Palawan Province, northern part of Palawan I., environs of Taytay Town on eastern coast, secondary forest, on leaf of bush, at night, 25 – 26. II. 2004, A. Gorochov (ZIN). Paratype: Philippines. Palawan Province, northern part of Palawan I., environs of Port Barton on western coast, 27 – 29. II. 2004, 1 ♀, secondary forest on hills, at night, A. Gorochov (ZIN). Type locality. Philippines, northern part of Palawan I., environs of Taytay Town on eastern coast, secondary forest. Other material examined: Philippines. Palawan, coll. Br. v. W. [Brunner von Wattenwyl], 1 ♀, # 18876 (MNHN-EO-ENSIF 1197).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B0EDE61FF10E99AA181FC89.taxon	etymology	Etymology. Named after the type locality.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B0EDE61FF10E99AA181FC89.taxon	distribution	Distribution. Philippines, northern part of Palawan Island.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B0EDE61FF10E99AA181FC89.taxon	diagnosis	Diagnosis. The new species differs from similar species by the following characters: from C. reticulatus, C. singapura, C. lampongsi, C. tello, C. erniae and C. floresiensis in the epiphallus clearly narrowed in middle part; additionally from C. reticulatus and C. tello, it differs in a more easily S-shaped dorsal edge of the dorsal epiphallic ridges in profile, and from the other species, in roundly-angular (not only rounded) proximal corners of the high parts of dorsal epiphallic ridges in profile. From C. pelagus, the new species differs in a wider subapical part of the epiphallus, more strongly S-shaped dorsal edges of the dorsal epiphallic ridges (this character also separates the new species from C. loboe), and less distinctly darkened apical parts of the hind femora, and from C quatei, C. oeroe, C. celebae and C. gagnei, in the notches between the proximal corners of dorsal epiphallic ridges and dorsoproximal lobes of epiphallus in profile clearly less deep (from the first species) or distinctly narrower (from the second, third and forth species). Female of C. palawan differs in the longitudinal veins of the dorsal tegminal field weakly convex, area between Sc and R with the crossveins and membranes of the same color, or distal part of the hind femora not contrastingly darker than the other parts of these femora.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B0EDE61FF10E99AA181FC89.taxon	description	Description. Male (holotype): Coloration of body light brown with following marks: face of epicranium with a few rather small brown spots (some of them weakly distinct; Fig. 6 E); dorsum of head brown with small light brown areas on rostrum and distinct light pattern from longitudinal and oblique lines; lateral part of head with only brown area behind each eye (Fig. 48 C); most part of labrum and of clypeus yellowish; scape and palpi with slightly darker (almost grayish brown) small marks (including darkened apical part of fifth segment of maxillary palpi); antennal flagellum with brown middle and distal parts; pronotum with most part of lateral lobes brown and with several darkish and weakly distinct spots on disk (Fig. 48 C); tegmina brown with reddish brown M, R, stock of Sc, areas (and crossveins) between them, and large spot situated slightly distad than plectrum, with light grayish brown membranes around chords and at apex of dorsal field, with whitish two spots on basal area of this field, one spot near mirror, one spot (much larger) on proximal half of this field, narrow and rather short humeral stripe in proximal part of tegmina, oblique veins, some of chords, all branches of Sc, and distal area of lateral field, as well as with yellowish diagonal vein and crossveins in apical area of dorsal field (Fig. 48 D – E); exposed part of hind wings brown with weakly lighter (weakly distinct) venation; legs with two slightly darkened spots on middle tibiae and on distal part of hind tibiae, and with partly brown (but not dark) fore and middle tarsi; coxae, all sternites, and genital plate yellowish; visible parts of abdominal tergites dorsally brown with light brown spots; anal plate with a few small brown marks; cerci yellowish with numerous small brown marks. Structure of body similar to that of C. reticulatus, but tegmina (Fig. 48 D – E) slightly not reaching abdominal apex, their mirror 2.1 times as long as wide; epiphallus of genitalia not long, clearly narrower in middle part and widened in subapical part (Fig. 48 H – K); proximal corners of high parts of epiphallic dorsal ridges roundly angular in profile and separated from dorsoproximal lobes of epiphallus by not deep notches; these dorsal ridges in profile roundly and easily (not roundly-angularly) S-shaped (Fig. 48 H – I). Female: General appearance as in male, however with following differences: face of epicranium with brown spots slightly larger; darkened area on rostral dorsum somewhat lighter; most part of pronotal lateral lobes darker, almost dark brown; pronotal disk with more distinct spots; tegminal dorsal field slightly lighter (intermediate between brown and light brown), with yellowish-whitish large basal spot in dorsal field and rather wide humeral stripe (fused with this spot proximally), with light brown majority of longitudinal veins in dorsal field, and with almost yellowish crossveins between latter veins (Fig. 48 F – G); femora somewhat more spotted (all they with slightly darker spots, but small spots along ventral ridges of hind femora more distinct, brown; hind tibiae missing); structure of tegmina and of abdominal apex similar to that in female of C. reticulatus, but genital plate with widely truncate apex, and ovipositor clearly longer (hind femur 1.25 times as long as ovipositor). Measurements. Length in mm. Body: male 19, female 16; body with wings: male 22, female 25; pronotum: male 2.5, female 3; tegmina: male 12.5, female 15.5; hind femora: male 13.5, female 15; ovipositor 12. Habitat and life history traits. Unknown. Behavior. Unknown.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B10DE63FF10EBEBA456FB84.taxon	description	(Figs. 4, 6 F, 10 E, 12 H, 13 P, 15 F, 49)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B10DE63FF10EBEBA456FB84.taxon	materials_examined	Type material. Male holotype: Malaysia. Sarawak, Nanga Pelagus near Kapit, 180 – 585 m, 7 – 14. VIII. 1958, Maa (BPBM) (not examined).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B10DE63FF10EBEBA456FB84.taxon	distribution	Distribution. Malaysia, western part of Borneo.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B10DE63FF10EBEBA456FB84.taxon	materials_examined	Material examined. Malaysia. Sarawak State, foot of Mt. Dulit, junction of rivers Tinjar & Lejok, 10 – 24. IX. 1932, Oxford Univ. Exp., B. M. Hobby & A. W. Moore, B. M. 1933 - 254: 1 ♂, fish trap 1, identified Cardiodactylus gaimardi by L. Chopard (MNHN-EO-ENSIF 3511); 1 ♀, # 28, traps 1, identified Cardiodactylus gaimardi by L. Chopard (BMNH); 1 ♂, traps, identified Cardiodactylus gaimardi by L. Chopard (BMNH). East Malaysia, 4 th Division, Baram District, Gunong [Gunung = Mount] Mulu National Park, 4 ° 03 ’ N 114 ° 56 ’ E, 150 m, 12 – 18. V. 1978, 1 ♀, malaise in mixed dipterocarp forest, N. M. Collins (BMNH. 1978 - 11). Sarawak, Mulu National Park not far from borders with Brunei and Indonesia, 100 – 300 m, primary forest, on leaf of bush, at night, 24 – 27. III. 2012, A. Gorochov, M. Berezin, E. Tkatsheva, I. Kamskov, 1 ♂, 2 ♀ (ZIN). Borneo I., environs of Kuching City, Kubah National park on Matang Mt., 200 – 500 m, primary forest, on leaf of tree not far from soil, at night, 10 – 17. III. 2012, A. Gorochov, M. Berezin, E. Tkatsheva, I. Kamskov, 1 ♀ (ZIN).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B10DE63FF10EBEBA456FB84.taxon	diagnosis	Diagnosis. Species of small size, coloration contrasted, dark brown to black, with yellowish patterns. Male genitalia close to that of C. singapura with parallel carinated dorsal ridges and apex of pseudepiphallic sclerite not rounded. Species close to C. variegatus n. sp. but differing by coloration of pronotum (dorsal disk laterally yellow but median area mostly dark brown; lateral lobe yellow ventrally), and by coloration of FIII orange brown, less contrasted.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B10DE63FF10EBEBA456FB84.taxon	description	Redescription. In addition to the characters given by Otte (2007): size small for the species group, coloration dark brown to black with contrasted yellow brown patterns (Fig. 49 A – D). Head dorsum yellow brown with 4 dark brown bands including 2 complete lateral ones and 2 thin median punctuated bands; area posterior to eyes yellow brown with 2 short dark brown lines (Fig. 6 F). Pronotum: Dorsal disk yellow brown laterally, median area mottled with dark brown and with black spots. Lateral lobes dark brown dorsally, ventral half yellow. Legs yellow brown, femora mottled with dark brown, tibiae with dark brown rings. FIII orange brown, almost homogeneous, knees slightly darker than rest of femora. Tarsomeres 1 – 2 yellow brown, their apex dark brown. Hind knees black or dark brown. Cerci yellow brown, base faintly mottled with dark brown, apex with dark brown rings. Male: FW coloration (Fig. 10 E) mostly dark brown to black (cells and veins), with yellow to whitish areas including bases of anal veins and CuA, harp veins, oval region posterior to mirror and transverse veins in apical field. Bases of chords orange brown. M / R / Sc area and cells dark orange brown. Projections of Sc and more ventral veins of lateral field yellow. FW venation (Fig. 10 E): 1 A slightly bisinuated. CuP absent. Harp with 2 w-shaped veins. Mirror area: mirror (d 1) oval, separated in two parts near mid-length; d 2 as wide as mirror. Apical field with 3 cell alignments posterior to mirror (n = 3). Lateral field with 4 – 7 projections of Sc (m = 5; n = 3) and 3 more ventral veins (n = 3). Male genitalia (Fig. 49 E – F): Pseudepiphallus short, narrowed twice, at mid-length and preapically. Lateral ridges parallel, thick, wider anteriorly, their dorsal edges carinated. Pseudepiphallic sclerite with short triangular anterior expansions. Posterior pseudepiphallic apex trapezoidal, with a small rounded translucent area. Rami parallel, with very short apical stems. Ectophallic arc complete, v-shaped. Ectophallic fold with strong lateral sclerites; apex trilobate, membranous, median lobe elongate. Endophallic sclerite long, with a triangular posterior expansion and short lateral arms. Endophallic apodeme with a dorsal crest and wide lateral lamellas. Membrane of endophallic cavity finely plicate. Female: FW coloration (Fig. 12 H) mostly dark brown with contrasted yellow cells. Base of FWs yellow brown, including bases of anal veins, CuA and anterior half of M. CuA / M area yellow brown basally, then dark brown, with a yellowish spot at mid-length, extended on M / R area. R / Sc area and veins orange brown. Projections of Sc and more ventral veins yellow, the cells between them dark brown. FW venation (Fig. 12 H): 10 (n = 2) strong longitudinal veins on dorsal field; lateral field with 7 – 8 (n = 2) longitudinal veins including 4 – 5 projections of Sc and 3 – 4 more ventral veins. Ovipositor of average length, apex with both dorsal and ventral edges slightly denticulate (Fig. 13 P). Female genitalia (Fig. 15 F): Copulatory papilla triangular, with baso-lateral sclerites; apex rounded and slightly sclerotized, slightly folded ventrally. Juvenile: unknown.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B10DE63FF10EBEBA456FB84.taxon	discussion	Remark. The female specimen photographed by Otte (2007 a) probably does not belong to C. pelageu, but to C. borneoe n. sp., according to the pattern of coloration of the material examined here.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B12DE65FF10ECE0A181FC5C.taxon	description	(Figs 4, 6 G, 50)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B12DE65FF10ECE0A181FC5C.taxon	materials_examined	Type material. Male holotype: Malaysia. Pahang State, Tioman I. not far from Mersing City in Johor State (Malacca Peninsula), environs of Juara Vill. on eastern coast of island, disturbed primary forest, on leaf of bush, at night, 6 – 14. IV. 2010, A. Gorochov, M. Berezin, E. Tkatsheva (ZIN). Paratypes (2 ♀): same data as HT, 1 ♀ (ZIN), 1 ♀ (MNHN-EO-ENSIF 1133). Type locality. Malaysia, Tioman Island, not far from Mersing City in Johor State (Malacca Peninsula), environs of Juara Vill. on eastern coast of island.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B12DE65FF10ECE0A181FC5C.taxon	etymology	Etymology. Named after the reticular coloration of tegmina with partly whitish venation, but without large white spots.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B12DE65FF10ECE0A181FC5C.taxon	distribution	Distribution. Malaysia, Tioman Island.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B12DE65FF10ECE0A181FC5C.taxon	diagnosis	Diagnosis. The new species is most similar to C. pelagus from Borneo and C. quatei Otte, 2007 a from New Guinea by the shape of epiphallus but differs from these species in the presence of one (not three or two) large light spot in the distal half of male tegmen and epiphallic ridges in profile slightly less rounded (more straight).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B12DE65FF10ECE0A181FC5C.taxon	description	Description. Male (holotype): Coloration of body yellowish with following ornament: head with a few small dark brown spots on face, dark brown area behind each eye, rather numerous dark brown spots on dorsum (Fig. 6 G), small brown spots on scapes and palpi (including almost dark brown apex of apical segment of maxillary palpus), light brown proximal part of antennal flagellum, and brown middle and distal parts of this flagellum; pronotum with dark brown most part of lateral lobes and with numerous diverse brown and dark brown spots on dorsal disk (Fig. 50 B – C); FWs brown (almost dark brown) with rather large spot near mirror and majority of veins and crossveins contrastingly whitish, M and R as well as stock of Sc reddish (R branches and crossveins between Sc and R whitish, and areas between these crossveins brown), a few veins in dorsal field brown and light brown, and small areas near stridulatory vein and at base of dorsal field as well as short humeral stripe in proximal part of this field from whitish to light brown (Fig. 50 D – E); exposed part of hind wings dark brown with contrastingly yellowish and whitish venation; legs with rather numerous small dark brown spots on femora and coxae (and on pleurites near them), less numerous and larger dark brown spots on tibiae and tarsi (hind tibiae also with brown ventral part), and dark brown apical part of hind femora; abdomen with numerous dark brown and brown spots on tergites and supra-anal plate, small dark brown spots on cerci, light brown median band on subgenital plate, and brown lateral edges of this plate. Structure of body typical of Cardiodactylus; rostrum between antennal cavities about 1.2 times as wide as scape; tegmina reaching abdominal apex, with mirror almost 2.2 times as long as wide and with Sc having 8 – 9 branches (Fig. 50 E); hind wings significantly longer than FWs; fore tibiae with moderately large and oval outer tympanum and with rather short and slit-like inner tympanum. Male genitalia: Epiphallus moderately long, rather narrow, slightly asymmetrical, with almost parallel lateral edges (Fig. 50 H – K), with high parts of dorsal ridges rather long (approximately equal to more distal part of epiphallus in length) as well as not contacted to each other and not curved aside, and with proximal corners of these parts angular in profile and separated from dorsoproximal lobes of epiphallus by rather deep notches. Female: General appearance similar to that of male, but FW dorsal field dark brown with 11 almost parallel light brown longitudinal veins moderately convex, and numerous (but not very dense) whitish crossveins (Fig. 50 F – G); lateral field with Sc having 5 – 6 straight branches, sometimes with stock of R and M yellowish. Subgenital plate uniformly light. Ovipositor rather short (hind femur 1.45 times as long as ovipositor). Juvenile: Unknown. Measurements. Length in mm. Body: male 17, female 18 – 19; body with wings: male 23, female 23 – 25; pronL: male 2.4, female 2.5 – 2.7; FWL: male 12.5, female 13 – 14; FIIIL: male 14, female 14.5 – 16; ovipositor: 10 – 11. Habitat and life history traits. Cardiodactylus reticulatus was found in disturbed primary forest, on leaf or bush, at night. Behavior. Unknown.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B14DE65FF10EC58A285F9FB.taxon	description	(Figs 4, 6 H) Cardiodactylus riga 2007 a: 348; 2007 b: 30 (confirmation of depository); Robillard & Yap 2014: submitted.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B14DE65FF10EC58A285F9FB.taxon	materials_examined	Type material. Male holotype: Philippines. Luzon, Camarines Sur, Mt. Iriga, 500 – 600 m, coll. H. M. Torrevillas, 3 Apr. 1962 (BPBM), [not examined]. Type locality. Philippines: Luzon, Camarines Sur, Mt. Iriga, 500 – 600 m. New signalization. Philipppines, North Surigao Province, Surigao, Mindanao, Baker: 1 ♂, 14108 (MNHN-EO-ENSIF 1196); 1 ♀, 20433 (MNHN-EO-ENSIF 1195).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B14DE65FF10EC58A285F9FB.taxon	distribution	Distribution. Philippines, from to Luzon to Mindanao, including Leyte Island.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B14DE65FF10EC58A285F9FB.taxon	diagnosis	Diagnosis. Species of average size, colouration contrasted, face yellow with 3 black spots, very similar in colouration and external morphology to C. empagatao, to which it differs by brown Sc vein (yellow in C. empagatao), female venation yellow with dark brown cells and male genitalia with three levels in lateral view.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B14DE65FF10EC58A285F9FB.taxon	description	Redescription. see Robillard & Yap (2014).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B14DE66FF10EEFAA181FA54.taxon	description	(Figs. 4, 6 I, 10 F, 51)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B14DE66FF10EEFAA181FA54.taxon	materials_examined	Type material. Male holotype: Philippines. North Zamboanga Province, Mindanao, Dapitan, Baker, 7523 (MNHN-EO-ENSIF 1192). Type locality. Philippines, Mindanao Island, Dapitan.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B14DE66FF10EEFAA181FA54.taxon	etymology	Etymology. Named after Jose Rizal, the national hero of the Philippines who was exiled in the city of Dapitan.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B14DE66FF10EEFAA181FA54.taxon	distribution	Distribution. Philippines, north of Mindanao Island.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B14DE66FF10EEFAA181FA54.taxon	diagnosis	Diagnosis. Species of average to large size, coloration brownish with yellow brown patterns, characterized by head coloration and shape of pseudepiphallic sclerite in male genitalia with hook-like anterior expansions and slightly asymmetrical lateral expansions of dorsal ridges close to C. muria. Male genitalia close to that of C. empagatao Otte, 2007 a (shape of pseudepiphallic dorsal ridges and rami little sclerotized) but the species differ strikingly in terms of coloration.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B14DE66FF10EEFAA181FA54.taxon	description	Description. Average size for the species group. General coloration orange brown with yellow and dark areas (Fig. 51 A – B). Head dorsum with 4 dark brown bands, 2 wide lateral ones with an anterior notch separated from black coloration of fastigium, and 2 thin median parallel and punctuated bands; short triangular bands posterior to eyes slightly connected to lateral bands. Scapes yellow brown with a dark brown ring. Antennae orange brown. Face yellow brown, with a transverse dark brown band below antennae; front part of fastigium with faint parallel dark stripes. Mouthparts yellow brown. Maxillary palpi brown, apex darker. Lateral part of head dark brown dorsally, lateral margin gray brown. Pronotum: Dorsal disk 4 corners yellow; median area yellow brown with dark brown patterns; posterior area dark brown. Lateral lobes dark brown, ventral margin yellow. Legs I – II orange brown, with faint dark spots on femora and rings on tibiae. FIII orange brown, knees and tibiae dark brown. Hind wing tail dark brown, transverse veins yellow. Cerci orange brown. Apex of abdomen dark brown, sternites yellow brown. Male: FW coloration mostly dark brown (Fig. 10 F); area anterior to stridulatory file black; orange brown sclerotization near base of chords. Venation mostly yellowish, except part of 2 A, posterior half of diagonal and apical field, orange brown. Small oval area posterior to mirror with translucent cells and yellow veins. Lateral field brown. M / R area dark red brown. Sc vein orange brown, apex yellow and bifurcations whitish, including their basis. FW venation (Fig. 10 F): 1 A slightly bisinuated. CuP faint but present until the file level. Harp with 1 wshaped harp vein and a faint anterior vein. Mirror area: mirror (d 1) longer than wide, oval, well defined, separated in two parts by a strong transverse vein, the posterior part smaller and triangular; d 2 crossed by several accessory veins; e 1 not crossed by accessory veins. Apical field with 4 cell alignments posterior to mirror. Lateral field with 8 projections of Sc and 3 more ventral veins. Male genitalia (Fig. 51 C – E): Pseudepiphallus forming a narrow gutter, very sclerotized and narrowed preapically. Dorsal ridges parallel, slightly carinated innerly, with antero-lateral expansions folded externally, slightly asymmetrical as in C. muria. Anterior edge of pseudepiphallic sclerite with hook-like convergent expansions, membrane between them with thin setae. Posterior region rounded, with a small rounded translucent area at apex. Rami strong, with convergent apical stems. Ectophallic arc strong, with a short triangular posterior expansion. Bases of ectophallic apodemes with posterior lobes sclerotized at apex, without anterior sclerotization. Ectophallic fold laterally sclerotized, apex trilobate, median lobe conical and sponge-like. Endophallic sclerite with short lateral arms, with a triangular posterior expansion. Endophallic apodeme with lateral lamellas and a mediodorsal crest. Membrane of endophallic cavity finely plicate. Female: Unknown. Juvenile: Unknown. Measurements. See Table 13. Habitat and life history traits. Unknown. Behavior. Unknown.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B19DE6CFF10E8F0A2D9FD34.taxon	description	(Figs 4, 6 J, 10 G, 12 F, 13 Q, 15 G, 52, 53, 54)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B19DE6CFF10E8F0A2D9FD34.taxon	materials_examined	Type material. Male holotype: Indonesia. East Nusa Tenggara Province, Sumba, Laiwangi-Wanggameti National Park, près du village de Praingkareha, 10 ° 01 ' 11.7 " S 120 ° 03 ' 22.9 " E, 340 m, 15 – 18. VI. 2010, mort en élevage (TR 265), T. Robillard (MZB). Female allotype: same information as HT (TR 303) (MZB). Paratypes (7 ♂, 7 ♀): same information as HT, 15 – 18. VI. 2010, mort en élevage, T. Robillard: 1 ♂ (TR 298) (MZB); 3 ♀, (TR 264 - 266) (MZB); 1 ♀, souche élevage (MNHN-EO-ENSIF 3657); 2 ♂, souche élevage, enregistrement appel (MNHN-EO-ENSIF 3658 - 3659); 1 ♂ (TR 297), 1 ♀ (TR 303) (ZIN). Sumba, Laiwangi-Wanggameti National Park, près du village de Praingkareha, forêt sur pente à l'E du camp, 10 ° 00 ' 59,8 " S 120 ° 03 ' 36,5 " E, 425 m (GPS 6), T. Robillard: 15 – 18. VI. 2010, nuit, 2 ♂ (TR 246 - 247), zone ouverte, sur plante (h = 80 cm), enregistrement appel (MNHN-EO-ENSIF 3660 - 3661); 17. VI. 2010, nuit, 1 ♀ (TR 276), sur arbuste (h = 1 m) (MNHN-EO-ENSIF 3662). Sumba, Laiwangi-Wanggameti National Park, près du village de Praingkareha, forêt karstique au S du camp, 10 ° 02 ' 02,9 " S 120 ° 03 ' 28,3 " E, 539 m (GPS 7), T. Robillard: 16. VI. 2010, nuit, 1 ♂ (TR 257), sur arbuste épineux en bord de sentier (h = 1.1 m), enregistrement appel (MNHN-EO-ENSIF 3663); 1 ♀ (TR 251), sur branche (h = 2 m) (MZB). Type locality. Indonesia, Sumba, Laiwangi-Wanggameti National Park. Other material examined. Indonesia. East Nusa Tenggara Province, O. Soemba [West Sumba], Kananggar [village], 700 m, V. 1925, 1 ♂ (MNHN). N. W. Soemba [Sumba], Laora [village], 100 m, IV. 1925, 1 ♀ (MNHN). Sumba, Laiwangi-Wanggameti National Park, près du village de Praingkareha, 10 ° 01 ' 11.7 " S 120 ° 03 ' 22.9 " E, 340 m (Sumba camp), 15 – 18. VI. 2010, mort en élevage, T. Robillard: 4 juveniles (TR 346, 347, 449, 450) (MZB); 4 juveniles (TR 299 - 301, TR 448) (MNHN).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B19DE6CFF10E8F0A2D9FD34.taxon	etymology	Etymology. Named after the type locality.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B19DE6CFF10E8F0A2D9FD34.taxon	distribution	Distribution. Indonesia, south of Sumba Island.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B19DE6CFF10E8F0A2D9FD34.taxon	diagnosis	Diagnosis. Species of average to large size, general coloration yellow brown with lighter patterns, with a wide yellow band posterior to mirror, characterized by male genitalia with folded asymetrical dorsal ridges close to C. kotandora n. sp. Distinctive mono-syllabic calling song with a very indented amplitude pattern.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B19DE6CFF10E8F0A2D9FD34.taxon	description	Description. Average to large size for the species group, stocky shape (Fig. 52 A – D). General coloration yellow brown, little contrasted. Head dorsum yellow brown with 4 faint brown bands including 2 complete lateral ones fused to the dark brown coloration of the fastigium, and 2 median punctuated bands; area posterior to eyes yellow brown with 2 short dark brown lines (Fig. 6 J). Scapes yellow brown, with a transverse dark brown band. Antennae orange brown to dark brown. Front part of fastigium yellow, sometimes with 2 dark brown spots. Lateral side of head mostly yellow brown, with orange brown areas posterior to eyes. Face homogeneously gray brown. Mouthparts yellow brown to whitish; insertion of mandibles black. Maxillary palpi yellow brown or orange brown, apex dark brown. Pronotum: Dorsal disk almost completely yellow brown with few dark brown spots; posterior edge mottled with dark brown. Lateral lobes mostly dark brown, ventral margin yellow brown. Legs I – II yellow brown, femora with faint orange brown spots and tibiae with rings. Tarsomeres I – II- 1 yellow brown, apex brown. FIII homogeneously orange brown; knees dark brown; TIII dark brown with faint yellow brown rings. Tarsomeres III- 1 orange brown, their apex slightly darker. Hind wing tail short, gray brown. Cerci orange brown, faintly mottled with dark brown. Abdomen mostly orange brown to dark brown. Subgenital plate yellow brown. Male: FW coloration with yellow brown areas including bases of anal veins and anterior half of CuA (Fig. 10 G). Whitish areas include harp veins, part of chords and a thin transverse band posterior to mirror. Two black spots near bases of 1 A and 2 A. Bases of chords and posterior part of diagonal orange brown. Apex of apical field grayish brown. M / R / Sc area and cells dark brown, apex of Sc whitish. Projections of Sc and more ventral veins of lateral field whitish. FW venation (Fig. 10 G): 1 A slightly bisinuated. Stridulatory file with about 220 teeth on the transverse and longitudinal parts of 1 A, without discontinuity at the level of the angle, and 20 – 30 more teeth on top of a bump near 1 A base (see Table 14, n = 2). CuP present basally. Harp with 2 w-shaped veins. Mirror area: mirror (d 1) variable, generally oval, crossed by longitudinal and transverse veins; d 2 as wide as mirror. Cell e 1 crossed at mid-length by an accessory vein. Apical field with 5 cell alignments posterior to mirror (n = 5). Lateral field with 8 projections of Sc (n = 5) and 4 – 5 ventral veins (m = 4; n = 5). Male genitalia (Fig. 52 E – G): Pseudepiphallus shaped as a wide gutter. Apex of dorsal ridges setose, folded laterally, slightly asymmetrical, the left one larger than the right. Dorsal ridges almost convex in lateral view. Pseudepiphallic sclerite with wide triangular anterior expansions, the membrane between them setose. Posterior pseudepiphallic apex rounded, with a small rounded translucent area with a median sclerotization. Rami with short convergent apical stems. Ectophallic arc complete, without posterior expansion. Ectophallic fold with strong lateral sclerites; apex trilobate, membranous, median lobe oval. Endophallic sclerite long, with a triangular posterior expansion and long lateral arms. Endophallic apodeme with a dorsal crest and wide lateral lamellas. Membrane of endophallic cavity finely plicate. Female: FW coloration dark brown with orange brown to gray brown veins (Fig. 12 F). Base of FWs yellow brown, including bases of anal veins, CuA and M. CuP strong, orange brown. CuA / M area dark brown, with a yellowish area near FW mid-length. R / Sc area and veins dark brown, apex of Sc whitish, its projections and more ventral veins whitish, the cells between them dark brown. Variation: dark brown area with whitish veins sometimes present in the third quarter of FW. FW venation (Fig. 12 F): 11 – 14 (m = 12.2, n = 5) strong longitudinal veins on dorsal field; lateral field with 9 – 11 (m = 10.4, n = 5) longitudinal veins including 5 – 7 projections of Sc and 4 – 5 ventral veins. Ovipositor of average length, apex with both dorsal and ventral edges denticulate (Fig. 13 Q). Female genitalia (Fig. 15 G): Copulatory papilla elongated with baso-lateral sclerites; apex rounded and slightly sclerotized, folded ventrally. Juvenile: Light grayish brown, head with same patterns of coloration as adults. Measurements. See Table 14. Habitat and life history traits. Cardiodactylus sumba is a nocturnal species living both in forested areas and in or near secondary habitats such as bushes and hedges. Males produce calling songs at night, from tree branches and leaves (Fig. 53). During the day, couples have been observed mating under the bark of trees. Juveniles are found day and night on low vegetation. Behavior. Calling song (Fig. 54): High speed video observations at 1250 frames per second show that each calling song of C. sumba corresponds to only one FW closure (1 syllable), the gaps within the syllables being made by the jerking movement of the FW closure. At 20.5 ° C, these mono-syllabic calling songs have the following characteristics: syllable duration = 319 ± 29 ms; syllable period = 3.2 ± 0.8 s; syllable duty cycle = 9.9 %. The power spectrum consists of a broad band dominating at 11.74 ± 0.20 kHz with no harmonic structure, suggesting that the system is not sustained as usually in crickets.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B1DDE6EFF10ED19A181FC14.taxon	description	(Figs 4, 6 K, 10 H, 55)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B1DDE6EFF10ED19A181FC14.taxon	materials_examined	Type material. Male holotype: Indonesia. North Sulawesi Province, Beo (Talaud) eilanden [Sangihe and Talaud Islands, Karakelong Island], IV. 1924, B. Rufmans [?] (MZB-ORTH 10632). Type locality. Indonesia, Karakelong Island (Talaud Islands).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B1DDE6EFF10ED19A181FC14.taxon	etymology	Etymology. Named after the type locality.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B1DDE6EFF10ED19A181FC14.taxon	distribution	Distribution. Indonesia, Karakelong Island.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B1DDE6EFF10ED19A181FC14.taxon	diagnosis	Diagnosis. Species of average size, coloration brownish with yellow brown patterns, characterized by head coloration and shape of pseudepiphallic sclerite in male genitalia. Male genitalia close to that of C. empagatao (shape of pseudepiphallic dorsal ridges and rami little sclerotized) but the two species differ strikingly in terms of coloration.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B1DDE6EFF10ED19A181FC14.taxon	description	Description. Average size for the species group. General coloration brown with yellow areas (Fig. 55 A – B). Head dorsum yellow brown with 3 wide dark brown bands, the lateral ones very wide, connected to black coloration of fastigium; the ventral band thin and punctuated, resulting from the partial fusion of 2 median lines (Fig. 6 K); area posterior to eyes without dark brown lines. Scapes yellow brown, with a transverse dark ring. Antennae orange brown. Face and mouthparts almost homogeneously yellow, with a transverse dark brown band below antennae, extended laterally posterior to eyes; small yellow area ventral to eyes. Pronotum: Dorsal disk dark brown with black spots, the 4 corners yellow. Lateral lobes almost black, ventral margin yellowish. Legs orange brown, with faint dark spots on femora. Hind knees dark brown. Hind wing tail dark brown. Abdomen yellow brown ventrally. Cerci orange brown with dark brown rings. Male: FW coloration dark brown, yellow basally (Fig. 10 H). Yellow areas include base and transverse part of 1 A, harp veins, a pale transverse band posterior to mirror, small part of chords. Orange sclerotized area near base of chords absent. Lateral field: M / R veins dark brown, area orange brown; Sc orange brown, its posterior part yellowish. Projections of Sc yellow, cells between them brown. FW venation (Fig. 10 H): 1 A not bisinuated. CuP missing. Harp with 2 w-shaped veins. Mirror area: mirror (d 1) longer than wide, oval and well defined, separated in two parts by a strong transverse vein, the posterior part triangular; d 2 not distinctive; e 1 crossed by transverse veins in posterior part. Apical field with 4 cell alignments posterior to mirror. Lateral field with 8 projections of Sc and 6 more ventral veins (n = 1). Male genitalia (Fig. 55 C – E): Pseudepiphallus forming a wide gutter not narrowed preapically, posterior region rounded. Dorsal ridges diverging posteriorly, slightly carinated innerly, with lateral expansions folded externally. Pseudepiphallic sclerite with wide latero-anterior expansions almost rectangular, membrane between them slightly setose. Rami median part little sclerotized. Ectophallic arc and apodemes little sclerotized. Ectophallic fold little sclerotized laterally. Endophallus little sclerotized. Female: Unknown. Juvenile: Unknown. Measurements. See Table 15. Habitat and life history traits. Unknown. Behavior. Unknown.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B1FDE50FF10EDE0A181FBCC.taxon	description	(Figs 4, 6 L, 56)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B1FDE50FF10EDE0A181FBCC.taxon	materials_examined	Type material. Male holotype: Indonesia. Sulawesi Utara Province, ~ 40 km NE of Manado City (Sulawesi I.), Tangkoko National Park on eastern coast of Minahassa Peninsula, environs of Tangkoko Lodge, disturbed primary forest near sea, on leaf of bush, at night, 3 – 6. II. 2011, A. Gorochov (ZIN). Paratypes (1 ♂, 2 ♀): same data as for holotype, 1 ♀ (ZIN), 1 ♂, 1 ♀ (MNHN-EO-ENSIF 3654). Type locality. Indonesia, ~ 40 km NE of Manado City (Sulawesi I.), Tangkoko National Park on eastern coast of Minahassa Peninsula, environs of Tangkoko Lodge.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B1FDE50FF10EDE0A181FBCC.taxon	etymology	Etymology. Named after the type locality.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B1FDE50FF10EDE0A181FBCC.taxon	distribution	Distribution. Indonesia, north of Sulawesi Island.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B1FDE50FF10EDE0A181FBCC.taxon	diagnosis	Diagnosis. The new species is most similar to C. borneoe but distinguished by general coloration and by higher proximal part of dorsal epiphallic ridges and deeper notch between proximal part of these ridges and more proximal part of epiphallus in profile.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B1FDE50FF10EDE0A181FBCC.taxon	description	Description. Male (holotype): Coloration of body light brown with following pattern: head with a pair of very small dark brown spots at anterolateral corners of clypeus, with brown most part of genae, and with dorsum having six narrow light brown stripes (Fig. 6 L); pronotum with several dark brown spots on disk partly fused with each other (Fig. 56 B) and dark brown most part of lateral lobes (Fig. 56 C); tegmina dark brown with yellowish white branches of Sc, narrow areas around proximal halves of these branches, two rather large spots (basal and lateral) on dorsal field, narrow transverse stripe at base of apical area, small parts of venation along lateral and medial edges of apical area, and venation of three other parts of dorsal field (as in Fig. 56 D), as well as with whitish gray membranes in region of oblique veins and almost transparent membranes in region of chords; exposed part of hind wings almost dark grayish brown with distinct whitish gray venation; legs with rather numerous (but not very distinct) darkish dots, small spots and oblique stripes (these stripes situated only on hind femora), as well as with grayish brown apical part of hind femora, basal part of hind tibiae and areas on dorsal surface of these tibiae; abdomen with dark brown (almost blackish) visible part of tergites having rather numerous lighter spots. Structure of body more or less similar to that of C. borneoe but length of tegmina and of hind wings as in long-winged representatives of this species or slightly longer (but not somewhat shortened as in C. thailandia), tegminal mirror 1.8 times as long as wide, comb of tegminal Sc with 7 – 10 branches (Fig. 56 E), and genitalia (Fig. 56 H – K) with somewhat higher proximal part of dorsal epiphallic ridges and deeper notch between proximal part of these ridges and more proximal part of epiphallus in profile. Female: General appearance as in male, but tegmina with 10 – 11 longitudinal branches in dorsal field, 8 – 9 branches on Sc, and characteristic coloration of dorsal field: dark brown with yellowish white two rather large spots (basal and lateral), and venation in middle part of this field as well as along medial edge of this field and lateral edge of apical area (Fig. 56 F – G); ovipositor rather long (hind femur 1.2 times as long as ovipositor). Juvenile: Unknown. Variation. Male paratype with light median stripes on head dorsum almost indistinct, with disk almost completely dark brown but having light lateral margins and a few small and almost indistinct lightish marks on median part. Measurements. Length in mm. Body: male 21 – 21.5, female 23 – 24; body with wings: male 24.5 – 25.5, female 29 – 30.5; pronotum: male 2.7 – 2.9, female 3.4 – 3.6; tegmina: male 19 – 19.5, female 17 – 18; hind femora: male 16 – 17, female 18.5 – 19.5; ovipositor 15.5 – 16. Habitat and life history traits. C. tangkoko was found in disturbed primary forest near sea, on leaf of bush, at night. Behavior. Unknown.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B21DE53FF10ECA8A198FE39.taxon	description	(Figs. 4, 6 M, 10 I, 12 G, 13 R, 15 H, 57)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B21DE53FF10ECA8A198FE39.taxon	materials_examined	Type material. Male holotype: Indonesia. West Sumatra Province, P [ulau] Tello [Telo Island], XI. 1924, C. B. K. & N. S. (MNHN-EO-ENSIF 3670). Female allotype: same information as HT (MNHN-EO-ENSIF 3671). Type locality. Indonesia, West Sumatra, Telo Island. Other material examined. Indonesia. West Sumatra Province, Mentawei [Mentawai Islands], Siberoet [Siberut Island], 30. IX. 1924, 1 ♀ # 135, H. H. Karny (MNHN). Lampung Province, Wai Lima, Z. Sum. [South Sumatra], Lampongs [Lampung], XI. XII. 1921, 1 ♂, No 14, identified Cardiodactylus novaeguineae Haan by L. Chopard, [H. H.] Karny (MZB-ORTH 10426).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B21DE53FF10ECA8A198FE39.taxon	etymology	Etymology. Named after the type locality.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B21DE53FF10ECA8A198FE39.taxon	distribution	Distribution. Indonesia, wester Sumatra, Telo Island, Siberut Island.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B21DE53FF10ECA8A198FE39.taxon	diagnosis	Diagnosis. Species of average size, coloration brownish with yellow brown patterns, characterized by male FW coloration including absence of yellow area posterior to mirror, and by the following features of male genitalia: posterior region of pseudepiphallus wider than anterior region, bases of rami membranous as in C. empagatao and C. talaudae n. sp.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B21DE53FF10ECA8A198FE39.taxon	description	Description. Average size for the species group. General coloration mostly light brown and yellow brown, with dark brown patterns on head and lateral lobes of pronotum (Fig. 57 A – D). Head dorsum yellow brown with 4 dark brown bands including 2 complete lateral ones with an anterior notch sometimes discontinuous and 2 median punctuated bands; area posterior to eyes yellow brown with 2 short dark brown lines (Figs 6 M). Scapes yellow brown, their bases darker; pedicel dark brown; rest of antennae light brown. Dorsal part of fastigium dark brown, front part yellow. Lateral side of head dark brown, face gray brown to dark brown, with a transverse yellow brown band ventral to eyes. Mouthparts dark brown mottled with yellow. Maxillary palpi yellow brown, apex dark brown. Pronotum: Dorsal disk lateral margins yellow brown to whitish; median area with dark brown patterns and black spots. Lateral lobes dorsal half dark brown, ventral part yellow brown. Legs I – II yellow brown, femora with brown spots and tibiae with brown rings; tarsomeres I – II- 1 yellow brown, apex brown. Legs III yellow brown including knees. Tarsomeres III- 1 yellow brown, their apex dark brown. Hind wing tail dark brown, 1.5 longer than pronotum. Cerci yellow brown, with dark brown spots basally then with dark brown rings. Abdomen mostly dark brown to black with dorsal longitudinal discontinuous yellow lines. Male: FW coloration brown, little contrasted (Fig. 10 I), with whitish or yellow areas including bases of anal veins, anterior half of CuA, harp veins and part of chords. Diagonal orange brown. Light area posterior to mirror almost not visible, except for a few translucent transverse veins. External part of e 1 dark brown. M / R area orange brown, R / Sc area orange brown, basal part dark brown. Projections of Sc whitish, more ventral veins and cells of lateral field brown. FW venation (Fig. 10 I): 1 A slightly bisinuated. CuP missing. Harp with 2 w-shaped veins. Mirror area: mirror (d 1) oval, crossed near mid-length by a transverse vein; d 2 as wide as mirror, crossed by several accessory veins. Cell e 1 crossed at mid-length by an accessory vein. Apical field with 3 cell alignments posterior to mirror (HT). Lateral field with 3 projections of Sc and 3 more ventral veins. Subgenital plate yellow brown. Male genitalia (Fig. 57 E – G): Posterior region of pseudepiphallus wider than anterior region. Dorsal ridges parallel or slightly divergent posteriorly, little carinated laterally, without inner expansion as in C. erniae n. sp. Pseudepiphallic sclerite with short antero-lateral expansions, the membrane between them with few setae. Posterior pseudepiphallic apex rounded, with a small rounded translucent area. Rami characterized by their membranous bases; parallel, with large preapical plates and thin apical stems. Ectophallic arc membranous, sclerotized on median area only with a short posterior expansion. Ectophallic fold with weak lateral sclerites; apex trilobate, membranous, median lobe oval. Endophallic sclerite wider than long, with a triangular posterior expansion. Endophallic apodeme absent or very small, with no visible lateral lamellas and dorsal crest. Membrane of endophallic cavity finely plicate, dorsal cavity longer than in most Cardiodactylus species. Female: FW coloration brown with yellow brown veins (Fig. 12 G). Bases of anal veins whitish. CuP strong, orange brown. Anterior half of CuA, M and related accessory veins yellow. R / Sc area and veins orange brown. Projections of Sc and more ventral veins whitish, the cells between them brown. FW venation (Fig. 12 G): 11 – 12 (n = 2) strong longitudinal veins on dorsal field; lateral field with 10 (n = 2) longitudinal veins including 7 projections of Sc and 3 more ventral veins. Ovipositor of average length, apex with both dorsal and ventral edges denticulate (Fig. 13 R). Female genitalia (Fig. 15 H): Copulatory papilla triangular with baso-lateral sclerites; apex rounded and sclerotized, folded ventrally. Juvenile: Unknown. Measurements. See Table 16. Habitat and life history traits. Unknown. Behavior. Unknown.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B21DE53FF10ECA8A198FE39.taxon	discussion	Remark. The male specimen from Lampung Province (Indonesia, South Sumatra) shows differences with C. tello in terms of coloration and male FW venation which barely allow distinguishing it from other species from the same region (C. lampongsi and C. jdoeria). However the male genitalia are very similar to that of C. tello with small differences suggesting that it may be another species, to be assessed with additional material from Sumatra and surrounding islands.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B22DE53FF10E9BBA0CDFBBB.taxon	materials_examined	Type material. Male holotype: Thailand. Khaoyai [Khao Yai] National Park, about 14. VII. 1986 (M. Takeda) (OMNH) [examined]. Type locality. Thailand, Khao Yai National Park. New signalizations: Thailand. Trat Province, eastern Thailand, Chang I. in Siam Bay, low mountains near sea, primary forest, 5 – 20. XI. 2000, night, A. Gorochov and L. Anisyutkin: 1 ♂, 1 ♂ juvenile (ZIN). Cambodia. 10 – 15 km NEE of Sihanoukville (City near Siam Bay), environs of waterfall, 200 m, secondary forest, 11 – 12. IX. 2003, night, A. Gorochov: 1 ♂ (adult in captivity), 1 ♀ juvenile (ZIN).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B22DE53FF10E9BBA0CDFBBB.taxon	distribution	Distribution. Thailand, Cambodia.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B22DE53FF10E9BBA0CDFBBB.taxon	diagnosis	Diagnosis. Species of average size for the genus, with very contrasted black and vivid yellow coloration, close to C. borneoe n. sp. from Borneo, C. admirabilis from Singapore, C. guttulus from Ryukyu Islands (Japan), and C. kondoi from the Philippines, with differences on color pattern and the shape of the pseudepiphallic sclerite in male genitalia.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B22DE55FF10EC35A181FD34.taxon	description	(Figs 4, 6 N, 12 I, 13 S, 15 I, 58)	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B22DE55FF10EC35A181FD34.taxon	materials_examined	Type material. Female holotype: Malaysia. Sabah State, Borneo I., Trus Madi Mt, ~ 1000 m, partly primary / partly secondary forest, on leaf of bush, at night, 13 – 25. V. 2007, A. Gorochov (ZIN). Paratypes (3 ♀): Malaysia, Sabah State, [Borneo I.], Sabah, nr. Long Pa Sia (West), c. 1020 m, 25. XI. 1987, Mal [aise] trap 2, RMNH 87, C. v. Achterberg: 1 ♀ (RMNH); 1 ♀ (MNHN-EO-ENSIF 3510). Type locality. Malaysia, Borneo, Trus Madi Mount. Other material examined. Malaysia. Sabah State, [Borneo I.], Sabah, nr. Long Pa Sia (West), c. 1020 m, 25. XI. 1987, Mal [aise] trap 2, RMNH 87, C. v. Achterberg: 2 juveniles (RMNH); 1 juvenile (MNHN).	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B22DE55FF10EC35A181FD34.taxon	etymology	Etymology. Named after very spotted (variegate) colouration.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B22DE55FF10EC35A181FD34.taxon	distribution	Distribution. Malaysia, northern part of Borneo.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B22DE55FF10EC35A181FD34.taxon	diagnosis	Diagnosis. The new species is very close to C. pelagus. It differs from all the other similar congeners in the characteristic coloration: face almost uniformly yellow, pronotum with two dark bands on each lateral lobe, legs strongly spotted, and hind femora with very dark apical part.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
03E687D65B22DE55FF10EC35A181FD34.taxon	description	Description. Female: Coloration of body yellowish (Fig. 58 A – B) with following ornament: head (Figs 6 N, 58 C – E) with yellow face, light brown narrow stripe along clypeal suture, several brown spots on dorsum and behind eyes, light brown scape and palpi, slightly darker proximal part of antennal flagellum having several brown spots, and brown middle and distal parts of this flagellum; pronotum with two dark brown longitudinal bands on lateral lobes (wider upper band, and narrower and shorter lower band situated along anterior 2 / 3 of ventral edge of these lobes) and with several brown spots on median part of disk (Fig. 58 D); tegmina (Fig. 12 I) brown (almost dark brown) with three whitish spots (basal spot, proximal humeral stripe fused with previous spot, and distal humeral spot), one of longitudinal vein in dorsal field also whitish, crossveins in this field light brown, some branches of Sc and crossveins between them whitish, and R and stock of Sc as well as crossveins between them reddish brown; exposed part of hind wings dark brown with light brown venation; legs with numerous distinct brown spots, very dark brown apical part of hind femora, dark brown base and ventral surface of hind tibiae as well as very distinct dark brown spots on rest of these tibiae and on hind tarsi; abdomen with dark brown tergites having lighter spots and with small dark spots on cerci. Structure of body similar to that of female of C. reticulatus, dorsal tegminal field with 10 moderately convex longitudinal veins, genital plate intermediate between those of C. reticulatus and C. borneoe in shape of its apical part, and hind femur 1.3 times as long as ovipositor. Apex of ovipositor with both dorsal and ventral edges denticulate (Fig. 13 S). Female genitalia (Fig. 15 I): Copulatory papilla diamond-shaped with wide baso-lateral sclerites; apex barely rounded and sclerotized, only slightly folded ventrally. Male: Unknown. Juvenile: Coloration pattern of head and legs close to that of adults; body light brown mottled with dark brown. Measurements. Length in mm. Body 16.5; body with wings 21; pronotum 2.5; tegmina 12.5; hind femora 13.5; ovipositor 10.5. Habitat and life history traits. Unknown. Behavior. Unknown.	en	Robillard, Tony, Gorochov, Andrej V., Poulain, Simon, Suhardjono, Yayuk R. (2014): Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species. Zootaxa 3854 (1): 1-104, DOI: http://dx.doi.org/10.11646/zootaxa.3854.1.1
