taxonID	type	description	language	source
507C87C0CD6E1C4FEEEA3BD9FA9BFB3D.taxon	diagnosis	Diagnosis. Small to medium size (1.9 – 4.3 mm). Post-pedicel with acuminate apex. Pterostigma and vein M 2 absent; cross vein r-m touching cell dm near middle. Mid coxa with sclerotized plate, setose; male hind trochanter sometimes modified, with protuberance or spine. Abdomen shorter than wing length, usually widest at middle. Apex of phallic guide simple. Phallus trifid. Ejaculatory apodeme usually dorsoventrally truncated, generally sandal-shaped.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD6E1C49EEEA3C15FB86FF69.taxon	description	Figs 1 – 10, 132	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD6E1C49EEEA3C15FB86FF69.taxon	diagnosis	Diagnosis. Male. Postcranium brown, dorsally pruinose, ventrally grayish-pruinose (Fig. 1). Hind trochanter with yellow-setose protuberance (Fig. 2). Tergites and sternites 6 and 7 as in Fig. 3. Surstyli similar in shape, with hooked-tip, having dorso-basal protuberance, right surstylus slightly shorter (Figs 4 – 5), both surstyli with tips downward-directed, mainly the left one (Figs 6 – 7). Phallic guide with apex simple, angled at about 45 degrees (Fig. 8). Phallus with one ejaculatory duct with two spines, one duct with a single spine and one duct without spine. Ejaculatory apodeme somewhat sandal-shaped, darker near base, rounded on both ends (Fig. 9). Gonopods subsymmetrical (Fig. 10).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD6E1C49EEEA3C15FB86FF69.taxon	materials_examined	Material examined. COLOMBIA: Boyacá, SFF [Santuario de Fauna y Flora] Iguaque, Cabaña Chaina, 5 º 25 ’ N – 73 º 27 ’ W, 2600 m [eters], Malaise, 31. viii. – 16. ix. 2001, A. Roberto, “ M 2066 ” (1 ♂, LEUA) (photographed specimen); “ idem, 01 – 14. feb [ii]. 2001 ”, “ M 1271 ” (1 ♂, IAvH).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD6E1C49EEEA3C15FB86FF69.taxon	distribution	Geographical distribution. Ecuador, Galapagos islands; Mexico, Baja California (Rodriguez & Rafael 2012); Colombia (new record) (Fig. 132).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD6E1C49EEEA3C15FB86FF69.taxon	biology_ecology	Habitat. The specimens were collected with Malaise trap at the ground level at the Santuario de Fauna y Flora Iguaque reserve, where the vegetation is composed of forest typical of the mountain range of the Northeast region of Colombia.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD6E1C49EEEA3C15FB86FF69.taxon	discussion	Comments. Specimens collected are similar to the holotype illustrated by Ale-Rocha (1992), differing only by the left surstylus, which has the apex more truncated when examined in lateral view.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD681C4BEEEA3829FCD0FA21.taxon	description	Figs 11 – 20, 133	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD681C4BEEEA3829FCD0FA21.taxon	diagnosis	Diagnosis. Male. Postcranium dorsally brownish-pruinose, ventrally grayish-pruinose, (Fig. 11). Hind trochanter with white protuberance, setose (Fig. 12). Tergites and sternites 6 and 7 as in Fig. 13. Surstyli similar in shape, with hooked-tip, right surstylus slightly shorter (Figs 14 – 15), tip of both surstyli downward-directed when seen in lateral view (Figs 16 – 17). Phallic guide with apex lined with the main body of the structure, bearing one fine dorsal spine (Fig. 18). Phallus with one ejaculatory duct with small spine (Fig. 18). Ejaculatory apodeme somewhat sandalshaped, darker near base, rounded on both ends (Fig. 9). Gonopods subsymmetric, right gonopod wider and bifid distally (Fig. 20).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD681C4BEEEA3829FCD0FA21.taxon	materials_examined	Material examined. COLOMBIA: Caquetá, San Vicente del Caguan, Vda. [Vereda] Alto Quebradón, Fca. [Finca] Rancho Veracruz, 02 º 17 ’ 52.7 ” N / 74 º 44 ’ 15.5 ” W, 374 m [eters], 04 – 18. i. 2017, trampa Malaise en cultivo de caña (Sacharum officinarum), Y. Ramos-Pastrana (1 ♂, LEUA – 1148); “ idem, 29. iii. – 12. iv. 2017, (1 ♂, LEUA – 1149) ”; “ idem, (1 ♂, LEUA – 1150) ”; “ idem, (1 ♂, LEUA – 1151) ”; “ idem, (1 ♂, LEUA – 1152) ”; “ idem, (1 ♂, LEUA – 1153) ”; “ idem, (1 ♂, LEUA – 1154) ”; “ idem, (1 ♂, LEUA – 1155) ”; “ idem, (1 ♂, LEUA – 1156) ”; “ idem, (1 ♂, LEUA – 1157) ”; “ idem, (1 ♂, LEUA – 1158) ”; “ idem, (1 ♂, INPA) ”; “ idem, (1 ♂, LEUA – 1160) ”; “ idem, (1 ♂, LEUA – 1161) ”; “ idem, (1 ♂, LEUA – 1162) ”; “ idem, (1 ♂, LEUA – 1163) ”; “ idem, (1 ♂, LEUA – 1164) ”; “ idem, 15 – 29. iii. 2017, (1 ♂, LEUA – 1165) ”; “ idem, (1 ♂, LEUA – 1166) ”; “ idem, (1 ♂, LEUA – 1167) ”; “ idem, (1 ♂, LEUA – 1168) ”; “ idem, 01 – 15. ii. 2017, (1 ♂, LEUA – 1169) ”; “ idem, 18. i. – 01. ii. 2017, (1 ♂, LEUA – 1170) ”; “ idem, 21. xii. 2016. – 04. i. 2017, (1 ♂, INPA) ”; “ idem, San José del Fragua, Vda. [Vereda] Bellavista, Fca. [Finca] Mi Ranchito, 01 º 18 ’ 3 ” N / 76 º 00 ’ 33 ” W, 265 m [eters], 04 – 18. i. 2017, trampa Malaise en cultivo de caña (Sacharum officinarum), Y. Ramos-Pastrana (1 ♂, LEUA – 1171) ”; “ idem, (1 ♂, INPA) ”; “ idem, 18. i. – 01. ii. 2017, (1 ♂, LEUA – 1173) ”; “ idem, (1 ♂, LEUA – 1174) ”; “ idem, 15. ii. – 01. iii. 2017, (1 ♂, LEUA – 1175) ”; “ idem, 15 – 29. iii. 2017, (1 ♂, LEUA – 1176) ”; “ idem, 26. x. – 09. xi. 2016, (1 ♂, LEUA – 1177) ”; “ idem, (1 ♂, LEUA – 1178) ”; “ idem, (1 ♂, LEUA – 1179) ”; “ idem, 23. xi. – 07. xii. 2016, (1 ♂, LEUA – 1180) ”; “ idem, (1 ♂, LEUA – 1181) ”; “ idem, La Montañita, Vda. [Vereda] Morros, Fca. [Finca] Las Dalias, 01 º 29´21.5 ” N / 75 º 24´17 ” W, 290 m [eters], 04 – 18. i. 2017, trampa Malaise en cultivo de caña (Sacharum officinarum), Y. Ramos-Pastrana (1 ♂, LEUA – 1182) ”; “ idem, (1 ♂, LEUA – 1183) ”; “ idem, (1 ♂, INPA) ”; “ idem, (1 ♂, LEUA – 1185) ”; “ idem, 29. iii. – 12. iv. 2017, (1 ♂ LEUA – 1186) ”; “ idem, (1 ♂, LEUA – 1187) ”; “ idem, (1 ♂, LEUA – 1188) ”; “ idem, (1 ♂, LEUA – 1189) ”; “ idem, (1 ♂, LEUA – 1190) ”; “ idem, (1 ♂, LEUA – 1191) ”; “ idem, 15. ii. – 01. iii. 2017, (1 ♂, LEUA – 1192) ”; “ idem, (1 ♂, LEUA – 1193) ”; “ idem, 15 – 29. iii. 2017, (1 ♂, LEUA – 1194) ”; “ idem, 23. xi. – 07. xii. 2016, (1 ♂, LEUA – 1195) ”; “ idem, 26. x. – 09. xi. 2016, (1 ♂, INPA) ”; “ idem, Florencia, Vda. [Vereda] La Viciosa, Fca. [Finca] Macagual, 01 º 30 ’ 32 ’’ N / 75 º 40 ’ 28.8 ’’ W, 253 m [eters], 15. ii. – 01. iii. 2017, trampa Malaise en cultivo de caña (Sacharum officinarum), Y. Ramos-Pastrana (1 ♂, LEUA – 1197) ”; “ idem, (1 ♂, INPA) ”; “ idem, 23. xi. – 07. xii. 2016, (1 ♂, LEUA – 1199) ”; “ idem, Vda. [Vereda] Paraiso, Fca. [Finca] Paraiso, 01 º 44 ’ 51.4 ’’ N / 75 º 37 ’ 46.8 ’’ W, 663 m [eters], 09 – 23. xi. 2016, trampa Malaise en cultivo de caña (Sacharum officinarum), Y. Ramos- Pastrana (1 ♂, LEUA – 1200) ”; “ idem, 04 – 18. i. 2017, (1 ♂, LEUA – 1201) ”; “ idem, 09 – 23. xi. 2016, (1 ♂, LEUA- 1202) ”; “ idem, 01 – 15. ii. 2017, (1 ♂, LEUA – 1203) ”; “ idem, Vda. [Vereda] San Francisco, Fca. [Finca] El Recreo, 01 º 42 ’ 24.3 ’’ N / 75 º 36 ’ 36.6 ’’ W, 643 m [eters], 15. ii. – 01. iii. 2017, trampa Malaise en cultivo de caña (Sacharum officinarum), Y. Ramos- Pastrana (1 ♂, LEUA – 1204) ” (photographed specimen); “ idem, 01 – 15. ii. 2017 ”, “ idem, 26. x. – 09. xi. 2016, (1 ♂, LEUA – 1205) ”; “ idem, Albania, Vda. [Vereda] Florida 1, Fca. [Finca] El Jardín, 01 º 15 ’ 08.9 ’’ N / 75 º 53 ’ 05.2 ’ W, 283 m [eters], 09 – 23. xi. 2016, trampa Malaise en cultivo de caña (Sacharum officinarum), Y. Ramos-Pastrana (1 ♂, LEUA – 1206) ”; “ idem, 23. xi. – 07. xii. 2016, (1 ♂, LEUA – 1207) ”; “ idem, 07 – 21. xii. 2016, (1 ♂, LEUA – 1208) ”; “ idem, 04 – 18. i. 2016, (1 ♂, INPA) ”; “ idem, (1 ♂, LEUA – 1210) ”; “ idem, 01 – 15. ii. 2017, (1 ♂, LEUA – 1211) ”; “ idem, 01 – 15. iii. 2017, (1 ♂, LEUA – 1212) ”; “ idem, Fca. [Finca] San Isidro, 01 º 14.5 ’ 50.1 ’’ N / 75 º 52 ’ 34.7 ’’ W, 295 m [eters], 18. i. 2017. – 01. ii. 2017, trampa Malaise en cultivo de caña (Sacharum officinarum), Y. Ramos-Pastrana (1 ♂, LEUA – 1213) ”; “ idem, Belén de los Andaquíes, Vda. [Vereda] Aletones, Fca. [Finca] La Cabaña, 01 º 29 ’ 31.8 ’’ N / 75 º 52 ’ 20.8 ’’ W, 369 m [eters], 21. xii. 2016. – 04. i. 2017, trampa Malaise en cultivo de caña (Sacharum officinarum), Y. Ramos-Pastrana (1 ♂, LEUA – 1214) ”; “ idem, (1 ♂, LEUA – 1215) ”; “ idem, 04 – 18. i. 2017, (1 ♂, LEUA – 1216) ”; “ idem, 18. i. – 01. ii. 2017, (1 ♂, LEUA – 1217) ”; “ idem, 01 – 15. ii. 2017, (1 ♂, LEUA – 1218) ”; “ idem, Morelia, Vda. [Vereda] Albano, Fca. [Finca] Guacarí, 01 º 25 ’ 41 ’’ N / 75 º 44 ’ 56.4 ’’ W, 251 m [eters], 09 – 23. xi. 2016, trampa Malaise en cultivo de caña (Sacharum officinarum), Y. Ramos-Pastrana (1 ♂, LEUA – 1219) ”; “ idem, Solano, Vda. [Vereda] Las Mercedes, Fca. [Finca] La Ceiba, 00 º 47 ’ 07 ’’ N / 75 º 19 ’ 00.5 ’’ W, 211 m [eters], 09 – 23. xi. 2016, trampa Malaise en cultivo de caña (Sacharum officinarum), Y. Ramos-Pastrana (1 ♂, LEUA – 1220) ”; “ idem, Valparaiso, Vda. [Vereda] La Florida, Fca. [Finca] La Florida, 01 º 10 ’ 18.0 ’’ N / 75 º 38 ’ 40.2 ’’ W, 235 m [eters], 23. xi. – 07. xii. 2016, trampa Malaise en cultivo de caña (Sacharum officinarum), Y. Ramos-Pastrana (1 ♂, LEUA – 1221) ”; “ idem, Doncello, Vda. [Vereda] La Arenosa, Fca. [Finca] El Carmen, 01 º 40 ’ 30.8 ’’ N / 75 º 16 ’ 03.5 ’’ W, 322 m [eters], 15. ii. – 01. iii. 2017, trampa Malaise en cultivo de caña (Sacharum officinarum), Y. Ramos-Pastrana (1 ♂, LEUA – 1299) ”; “ idem, (1 ♂, LEUA – 1300) ”; “ idem, Puerto Rico, Vda. [Vereda] La Soledad, Fca. [Finca] Borinquen, 01 º 55 ’ 30 ’’ N / 75 º 08 ’ 33.2 ’’ W, 308 m [eters], 15 – 29. iii. 2017, trampa Malaise en piso del bosque, Y. Ramos-Pastrana, (1 ♂ LEUA – 1302) ”; “ idem, Milan, Vda. [Vereda] San Rafael, Fca. [Finca] Bellavista, 01 º 09 ’ 56 ’’ N / 75 º 26 ’ 24.3 ’’ W, 231 m [eters], 04 – 18. i. 2017, trampa Malaise en cultivo de caña (Sacharum officinarum), Y. Ramos-Pastrana (1 ♂, LEUA – 1303) ”; “ idem, Cartagena del Chaira, Vda. [Vereda] Tigrera Alta, Fca. [Finca] Las Palmeras, 01 º 17 ’ 5.2 ’’ N / 74 º 49 ’ 1.2 ’’ W, 235 m [eters], 21. xii. 2016. – 04. i. 2017, trampa Malaise en cultivo de caña (Sacharum officinarum), Y. Ramos-Pastrana (1 ♂, LEUA – 1304) ”; “ idem, Meta, PNN [Parque Nacional Natural S [Serrania] La Macarena, Caño Curia, Sendero Cachicamos, 3 º 21 ’ N / 73 º 56 ’ W, 460 m [eters], Malaise, 01 – 10. iii. 2003, W. Villalba leg. “ M 3527 ” (4 ♂, IAvH) ”; “ idem, 17. i. – 09. ii. 2003, “ M 3529 ” (4 ♂, IAvH) ”; “ idem, 09 – 24. ii. 2003, “ M 3524 ” (2 ♂, IAvH) ”; “ idem, 10. xi. – 21. xii. 2002, “ M 2985 ” (1 ♂, INPA) ”; “ idem, Boliva r, SFF [Santuario de Fauna y Flora] Los Colorados, Villa Roca, 9 º 54 ’ N – 75 º 07 ’ W, 180 m [eters], Malaise, 06 – 21. xiii [sic]. 2001, E. Deulufuet leg. “ M 2630 ” (1 ♂, INPA) ”; “ idem, 14 – 30. xi. 2001, “ M 2627 ” (1 ♂, IAvH) ”; “ idem, Boyacá, SFF [Santuario de Fauna y Flora] Iguaque, Cabaña Chaina, 5 º 25 ’ N / 73 º 27 ’ W, 2600 m [eters], Malaise, 14. ii – 05. iii. 2001, P. Reina leg. “ M 1359 ” (1 ♂, IAvH) ”; “ idem, Putumayo, PNN [Parque Nacional Natural] La Paya, Fca. [Finca] Charapa, 0 º 8 ’ S / 74 º 57 ’ W, 330 m [eters], Malaise, 26. iii –. 10. iv. 2002, R. Cobete Leg. “ M 3135 ” (1 ♂, IAvH) ”.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD681C4BEEEA3829FCD0FA21.taxon	distribution	Geographical distribution. Argentina; Bahamas; Brazil; Costa Rica; Cuba; Dominica; USA.; Haiti; Jamaica; Mexico; Nicaragua; Peru; (Ale-Rocha & Souza 2011; Rodriguez & Rafael 2012); Colombia (new record) (Fig. 133).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD681C4BEEEA3829FCD0FA21.taxon	biology_ecology	Habitat. The specimens were collected using Malaise traps placed at the ground level in the following areas: a sugar cane plantation (Sacharum officinarum); a tropical humid forest reserve in the Colombian Amazon; Departament of Caquetá, Parque Nacional Natural La Paya with savanna vegetation; floodplain forests of the Orinoco region at Parque Nacional Natural La Macarena; the dry tropical forest of the Andean region in the Santuario de Fauna y Flora Iguaque and the Caribbean region at Santuario de Fauna y Flora Los Colorados.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD681C4BEEEA3829FCD0FA21.taxon	discussion	Comments. The specimens observed in this study are smaller and have a surstylus basally narrower compared to the holotype illustrated by Ale-Rocha & Rafael (1995).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD6A1C44EEEA3CECFBE6FD4E.taxon	description	Figs 21 – 36, 133 http: // zoobank. org: act: A 62 E 864 A- 73 F 9 - 46 A 5 - BCF 8 - 2 DF 274 A 08 CC 5	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD6A1C44EEEA3CECFBE6FD4E.taxon	diagnosis	Diagnosis. Male. Hind trochanter with semi-transverse truncated long protuberance, apically with yellow setae. Surstyli subsymmetrical, slender, as long as epandrium, with hooked-tip, right surstylus slightly shorter, both surstyli somewhat straight in lateral view. Cerci elongated. Phallic guide short, with apex simple, bearing one small seta and tip angled at about 30 degrees. Phallus with one ejaculatory duct with two spines, one duct with a single spine and one duct without spine.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD6A1C44EEEA3CECFBE6FD4E.taxon	description	Description. MALE (holotype). Body length 3 mm; wing length 3 mm. Head (Figs 21 – 22). Holoptic, eyes contiguous for seven facets. F, EM, V (mm) = 0.2, 0.1, 0.1. Frons grayish-pruinose. Postcranium black, dorsally brownish-pruinose, ventrally grayish-pruinose. Antenna brown; pedicel with three dorsal and one ventral setae; postpedicel dorsally clearer, with apex filiform, LPP / WPP = 2.4; Labellum brown. Thorax (Fig. 22 – 23). Postpronotal lobe yellow. Scutum dark-brown, shiny, with dense brownish-pruinosity medially. Notopleuron grayish-pruinose. Scutellum dark-brown, grayish-pruinose, with inconspicuous band of setae. Mesopleuron light-brown. Mediotergite dark-brown to black, grayish-pruinose. Wing (Fig. 24). LW / MWW = 4. LTC / LFC = 0.5. Membrane hyaline, almost completely microtrichose except as follow: cell bc bare; cell c bare on proximal quarter; cell sc bare on proximal three quarters; cell r 1 bare on proximal quarter; cell bm bare on proximal half; cell cup bare on proximal anal quarter; lobe bare on distal three quarters. Halter with stem and knob yellow, base brown. Legs (Fig. 25). Coxae dark-brown; trochanters light-brown; femora dark-brown except for base and apex yellow; tibiae dark-brown, with base and apex yellow; tarsomeres 1 - 3 yellow, 4 and 5 brown. Hind trochanter with long semi-transverse, truncated, apically with yellow setae protuberance (Fig. 25 – 26). Abdomen (Figs 22, 27 – 28). Dark-brown to black, shiny, dorsally brownish pruinose. Tergite 1 with six lateral setae. Tergites and sternites 6 and 7 as in Fig. 29. Syntergosternite 8 brown, brownish-pruinose, shorter than tergite 5, with membranous area occupying less than half of syntergosternite in dorsal view (Figs 27 – 28). Terminalia (Figs 29 – 36). Epandrium brown and surstyli light brown. Surstyli (Figs 30 – 33) subsymmetrical, slender, as long as epandrium, with hooked-tips; right surstylus slightly shorter, both surstyli somewhat straight in lateral view (Figs 32 – 33). Cerci elongated. Phallic guide short with apex simple, bearing one small reclinate seta and tip angled at about 30 degrees (Fig. 34). Phallus with one ejaculatory duct with two spines, one duct with a single spine and one duct without spine (Fig. 34). Ejaculatory apodeme somewhat sandal-shaped, darker near base (Fig. 35). Gonopods same length, left with acuter apex (Fig. 36). FEMALE. Unknown.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD6A1C44EEEA3CECFBE6FD4E.taxon	materials_examined	Type material. HOLOTYPE ♂: COLOMBIA, Meta, PNN [Parque Nacional Natural] S [errania] Macarena, Caño Curia, Sendero Cachicamos, 3 º 21 ’ N / 73 º 56 ’ W, 460 m [eters], Malaise, 10. xi. – 21. xii. 2002, M. Duarte leg, “ M 2985 ” (1 ♂, IAvH) (photographed specimen); PARATYPES ♂: “ idem, 09 – 24. ii. 2003, W. Villalba leg, “ M 3524 ” (1 ♂, LEUA) ”; “ idem, 17. i. – 09. ii. 2003, “ M 3529 ” (1 ♂, INPA) ”. The holotype has the left hind leg glued to a triangle and pinned along specimen, left wing mounted in Canada balsam on a microslide with balsam, and terminalia placed in a microvial with glycerin, both pinned along the specimen.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD6A1C44EEEA3CECFBE6FD4E.taxon	distribution	Geographical distribution. Colombia (Fig. 133).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD6A1C44EEEA3CECFBE6FD4E.taxon	etymology	Etymology. The specific epithet refers to the type locality, Macarena, National Natural Park, Colombia.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD6A1C44EEEA3CECFBE6FD4E.taxon	biology_ecology	Habitat. The specimens were collected with Malaise traps at ground level at the Parque Nacional Natural Serrania La Macarena, within savanna vegetation and in the flooded forests of the Orinoco region of Colombia.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD6A1C44EEEA3CECFBE6FD4E.taxon	discussion	Taxonomic notes. Tomosvaryella macarenensis sp. nov. runs to T. longiseta Ale-Rocha 1996 in the couplet 13 of the key presented by Ale-Rocha & Souza (2011) based on hind trochanter with a short semi-transverse truncated protuberance. It differs from T. longiseta in having slender surstyli and apexes curved inward (Figs 30 – 31) (versus surstyli short with truncated apex in T. longiseta, figure 69 of Ale-Rocha 1996); apex of phallic guide short, with a small reclinate seta, phallus with two spines in central branch (Fig. 34) (versus apex of phallic guide long with stout projection, phallus with a fine spine in one branch, figure 73 of Ale-Rocha 1996).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD651C41EEEA3A4DFB87FD39.taxon	description	Figs 37 – 52, 132 http: // zoobank. org: act: CDB 3843 B- 0 BE 4 - 484 D-BE 13 - 4 EC 760 E 75780	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD651C41EEEA3A4DFB87FD39.taxon	diagnosis	Diagnosis. Male. Hind trochanter ventrally with two bare spines, basal larger than apical. Surstyli subsymmetrical, shorter than epandrium, with hooked-tips, right surstylus slightly shorter, both surstyli somewhat straight in lateral view. Cerci robust and elongated. Phallic guide with apex simple, having one small inconspicuous lateral left seta and tip angled at about 25 degrees. Phallus with one ejaculatory duct with three long and reclined spines, one duct with a single and short reclined spine and one duct without spine. Ejaculatory apodeme sandal-shaped with truncated apex. Gonopods subsymmetrical, left gonopod slightly shorter than right (Fig. 52).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD651C41EEEA3A4DFB87FD39.taxon	description	Description. MALE (holotype). Body length 3 mm; wing length 3 mm. Head (Figs 37 – 38). Holoptic, eyes touching each other for a distance of eight facets. F, EM, V (mm) = 0.2, 0.1, 0.2. Frons grayish-pruinose. Postcranium black, dorsally brownish-pruinose, ventrally grayish-pruinose. Antenna with postpedicel basally dark-brown, apically light-yellow; pedicel with three dorsal and one ventral setae. LPP / WPP = 2.6. Thorax (Fig. 38 – 39). Postpronotal lobe yellow. Scutum dark-brown, shiny, with brownish-pruinuosity medially denser. Notopleuron grayishpruinose. Scutellum dark-brown, grayish-pruinose, with inconspicuous marginal setae. Mesopleuron and mediotergite dark-brown, grayish-pruinose. Wing (Fig. 40). LW / MWW = 3. LTC / LFC = 0.3. Membrane hyaline, almost completely microtrichose except as follow: cell c bare on proximal three quarters; cell br; cell bm, cell cup and anal lobe bare. Halter stem and knob yellow, base brown. Legs (Fig. 41). Coxae and trochanters dark-brown, except hind trochanter light-brown at apex of outer surface; femora dark-brown, with yellow apex, grayish-pruinose; tibiae dark brown, with base and apex yellow; tarsomeres dorsally brown and ventrally yellow; pulvilli yellow (Fig. 41). Hind trochanter ventrally with two bare spines, basal larger than apical (Fig. 41 – 42). Abdomen (Figs 38, 43 – 44). Elongate, dark-brown to black, shiny, dorsally and laterally grayish-pruinose. Tergite 1 with four setae; tergites and sternites 6 and 7 as in Fig. 45. Syntergosternite 8 brown, grayish-pruinose, shorter than tergite 5, slightly inclined to right with membranous area occupying less than half of syntergosternite in dorsal view (Figs 43 – 44). Terminalia (Figs 45 – 52). Epandrium and surstyli brown. Surstyli (Figs 46 - 49) subsymmetrical, shorter than epandrium, with hooked tips; right surstylus slightly shorter, both surstyli somewhat straight in lateral view (Figs 48 – 49). Cerci robust and elongated. Phallic guide with simple apex, with one small inconspicuous lateral left setae and tip angled at about 25 degrees (Fig. 50). Phallus with one ejaculatory duct with three long and reclined spines, one duct with a single and short reclined spine and one duct without spine. Ejaculatory apodeme sandal-shaped with apex truncated. Gonopods subsymmetrical, left gonopod slightly shorter than right one (Fig. 52). FEMALE. Unknown.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD651C41EEEA3A4DFB87FD39.taxon	materials_examined	Type material. HOLOTYPE ♂: COLOMBIA: Caquetá, San Vicente del Caguan, Vda. [Vereda] Alto Quebradon, Fca. [Finca] Rancho Veracruz, 02 º 17 ’ 52.7 ’’ N / 74 º 44 ’ 15.5 ’’ W, 374 m [eters], 15 – 29. iii. 2017, trampa Malaise en cultivo de caña (Sacharum officinarum), Y. Ramos-Pastrana (1 ♂, LEUA – 1236) (photographed specimen); PARATYPES ♂: “ idem, 15. ii. – 01. iii. 2017, (1 ♂, LEUA – 1237) ”; “ idem, 29. iii. – 12. iv. 2017, (1 ♂, LEUA – 1238) ”; “ idem, 18. i. – 01. ii. 2017, (1 ♂, INPA) ”; “ idem, 29. iii. – 12. iv. 2017, (1 ♂, LEUA – 1240) ”; “ idem, Meta, PNN [Parque Nacional Natrual] S [Serrania] La Macarena, Caño Curia, Sendero Cachicamos, 3 º 21 ’ N / 73 º 56 ’ W, 460 m [eters], Malaise, 01 – 10. iii. 2003, W. Villalba leg. “ M 3527 ” (1 ♂, IAvH) ”; “ idem, Bolivar, SFF Los Colorados, Alto el Mirador, 9 º 54 ’ N / 75 º 7 ’ W, 400 m [eters], Malaise, 7 – 22. v. 2001, E. Doulufeut Leg, “ M 1724 ” (1 ♂, IAvH) ”. The holotype has the left hind leg glued to a triangle and pinned with specimen, left wing mounted in Canada balsam on a microslide with balsam, and terminalia placed in a microvial with glycerin, both pinned along the specimen.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD651C41EEEA3A4DFB87FD39.taxon	distribution	Geographical distribution. Colombia (Fig. 132).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD651C41EEEA3A4DFB87FD39.taxon	etymology	Etymology. The name is a tribute to Dr. Marta Wolff, for her valuable contribution to the study of the dipterofauna of Colombia.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD651C41EEEA3A4DFB87FD39.taxon	biology_ecology	Habitat. The specimens were collected with Malaise traps at ground level in sugar cane (Sacharum officinarum) cultivation areas, at preserved areas of tropical rain forest at the Colombian Amazon Departament of Caquetá; savanna vegetation, floodplain forests of the Orinoco region of Parque Nacional Natural La Macarena and areas of tropical dry forest in the Caribbean region in the Santuario de Fauna y Flora Los Colorados, Colombia.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD651C41EEEA3A4DFB87FD39.taxon	discussion	Taxonomic notes. Tomosvaryella martae sp. nov. runs to couplet 5 [T. bidens Cresson 1911 and T. amazonensis (Ale-Rocha 1992 a)] in the key presented by Ale-Rocha & Souza (2011) based on hind trochanter with two ventral spiniform protuberances. It differs from T. bidens by apex of phallic guide elongated and forward-directed (Fig. 50) (versus short and lined with base of phallic guide in T. bidens, figure 11 of Ale-Rocha & Rafael 1995); central branch of phallus with three apical, stout, reclined spines and one of the lateral branches with reclined short spine (Fig. 50) (versus without stout reclined spines, Fig. 12 of Ale-Rocha & Rafael 1995); from the T. amazonensis it differs by having apex of phallic guide elongated and forward-directed (Fig. 50) (versus apex of phallic guide short and ventrally with a small lobe in T. amazonensis (as in figure 8 of Ale-Rocha 1992 a); central branch of phallus with three long, reclined spines, one of the lateral branches with reclined short spine (Fig. 50) (versus central branch of phallus with one apical stout spine in T. amazonensis, figure 8 of Ale-Rocha 1992 a).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD601C41EEEA3BD9FBCAF9FB.taxon	description	Figs 53 – 62, 132	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD601C41EEEA3BD9FBCAF9FB.taxon	diagnosis	Diagnosis. Male. Postcranium black, dorsally brownish-pruinose, ventrally grayish-pruinose (Fig. 53). Frontal triangle pale-yellowish pruinose. Hind trochanter with somewhat acute protuberance (Figs 56 – 57). Tergites and sternites 6 and 7 as in Fig. 55. Surstyli similar in shape, with hooked-tip, right surstylus slightly shorter (Figs 58 – 59), both surstyli with tips downward-directed when examined in lateral view (Figs 58 – 59). Apex of phallic guide short with stout dorsal spine, angled at about 25 degrees (Fig. 60). Phallus branches with central branch bearing row of small spines, one of the lateral branches with one small spine (Fig. 60). Ejaculatory apodeme sandal-shaped with apex rounded (Fig. 61). Gonopods subsymmetrical, apexes acute, right gonopod slightly longer (Fig. 62).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD601C41EEEA3BD9FBCAF9FB.taxon	materials_examined	Material examined. COLOMBIA, Magdalena, PNN [Parque Nacional Natural] Tayrona, Neguanje, 11 º 20 ’ N / 74 º 02 ’ W, 10 m [eters], Malaise, 17 – 27. ix. 2001, R. Henriquez leg, “ M 2136 ” (1 ♂, IAvH); “ idem, (1 ♂, LEUA) ”; “ idem, (1 ♂, INPA) ”; “ idem, 28. vii – 18. viii. 2001, “ M 2019 ” (2 ♂, IAvH) ” (photographed specimen); “ idem, (1 ♂, LEUA) ”; “ idem, (1 ♂, INPA) ”; “ idem, 01 – 16. xii. 2001, “ M 2754 ” (1 ♂, IAvH) ”; “ idem, 1 – 14. vii. 2001, “ M 1928 ” (1 ♂, IAvH) ”.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD601C41EEEA3BD9FBCAF9FB.taxon	distribution	Geographical distribution. Bahamas; Brazil; Costa Rica; Cuba; Haiti; Jamaica; Mexico; Nicaragua; Peru; Dominican Republic (Rodriguez & Rafael 2012); Colombia (new record) (Fig. 132).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD601C41EEEA3BD9FBCAF9FB.taxon	biology_ecology	Habitat. The specimens were collected with Malaise traps at ground level at the reserve Parque Nacional Natrual Tayrona. This area has mainly thorny bushes and dry tropical forest typical of the Caribbean region of Colombia.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD601C41EEEA3BD9FBCAF9FB.taxon	discussion	Comments. The analyzed specimens are larger and have the protuberance of the hind trochanter acuter apically compared to the holotype specimen illustrated by Ale-Rocha & Rafael (1995).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD601C40EEEA3F98FDEAFC75.taxon	description	Figs 63 – 72, 132	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD601C40EEEA3F98FDEAFC75.taxon	diagnosis	Diagnosis. Male. Postcranium with lower half grayish-pruinose, upper half brownish-pruinose (Fig. 63). Hind trochanter with long brown protuberance, usually with one fine lateral seta and setulose at apex (Fig. 64). Tergites and sternites 6 and 7 as in Fig. 65. Surstyli somewhat similar in shape, with distinct basal lobe, hooked-tips, right surstylus slightly shorter (Figs 66 – 67), both surstyli with tips slightly downward-directed when seen in lateral view (Figs 68 – 69). Apex of phallic guide conical, ventrally serrated, angled at about 25 degrees (Fig. 70). Phallus with one ejaculatory duct with three stout spines, the other two ducts without spines (Fig. 70). Ejaculatory apodeme sandal-shaped, darker near base (Fig. 71). Gonopods subsymmetrical, apexes somewhat rounded (Fig. 72).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD601C40EEEA3F98FDEAFC75.taxon	materials_examined	Material examined. COLOMBIA, Boyacá, SFF [Santuario de Fauna y Flora] Iguaque, 5 º 25 ’ 12 ’’ N / 73 º 27 ’ 24 ’’ W, 2855 m [eters], Malaise 4, 01 – 19. iv. 2000, P. Reina, (1 ♂, IAvH); “ idem, 25. Jun [vi] – 13. Jul [vii]. 2000. (2 ♂, IAvH) ”; “ idem, (1 ♂, LEUA) ”; “ idem, (1 ♂, INPA) ”; “ idem, Cab. [Cabaña] Chaina, 5 º 25 ’ N / 73 º 27 ’ W, 2550 m [eters], 01 – 14. feb [ii]. 2001, A. Roberto, “ M 1271 ” (4 ♂, IAvH) ”; “ idem, (1 ♂, LEUA) ”; “ idem, 2600 m [eters], 16 – 30. xi. 2001, P. Reina leg, “ M 2583 ” (1 ♂, IAvH) ”; “ idem, 31. viii – 16. ix. 2001, A. Roberto leg, “ M 2066 ” (1 ♂, IAvH) ”; “ idem, La Planada, 5 º 25 ’ 12 ’’ N / 73 º 27 ’ 24 ’’ W, 2975 m [eters], Malaise 8, 09 – 24. ii. 2003, W. Villalba leg, “ M 3524 ” (1 ♂, IAvH) ”; “ idem, 01 – 19. iv. 2000, P. Reina (1 ♂, INPA) ”; “ idem, Cab. [Cabaña] Mamaramos, 05 º 25 ’ N / 73 º 25 ’ W, 2855 m [eters], Sep [ix] 23 – Oct [x] 11.2000, P. Reina leg, (3 ♂, IAvH) ”; “ idem, Aug [xii] 01 – 17.2000, (1 ♂, IAvH) ” (photographed specimen); “ idem, 17. viii. – 01. ix. 2000, (2 ♂, IAvH) ”; “ idem, Norte de Santander, PNN [Parque Nacional Natural] Tamá, Mun. [Municipio] Toledo, Vda. [Vereda] La Camacha, Pozo Negro, 7 º 21 ’ N / 72 º 28 ’ W, 2203 m [eters], Malaise, 23. x. – 2. xi. 2003, C. Leal leg, “ M 4030 ” (2 ♂, IAvH) ”; “ idem, (1 ♂, LEUA) ”; “ idem, (1 ♂, INPA) ”; “ idem, 2213 m [eters], “ M 4029 ” (1 ♂, IAvH) ”; “ idem, 2 – 5. xi. 2003, “ M 4031 ” (1 ♂, IAvH) ”.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD601C40EEEA3F98FDEAFC75.taxon	distribution	Geographical distribution. Colombia, Boyacá, Santuario de Fauna y Flora Iguaque; Cundinamarca (Rafael & Rodriguez 2016); Norte de Santander, Parque Nacional Natural Tamá (Fig. 132).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD601C40EEEA3F98FDEAFC75.taxon	biology_ecology	Habitat. Specimens were collected with Malaise traps at ground level at preserve areas that mainly have dry forests of the Andean region mountain range and temperate zones of the Northeast region of Colombia.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD601C40EEEA3F98FDEAFC75.taxon	discussion	Comments. The studied specimens are similar to the holotype illustrated by Ale-Rocha (1996), differing only by being smaller than the holotype by about 0.66 mm. The ejaculatory apodeme, lost in the holotype, is here illustrated based on non-type specimen.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD611C40EEEA3D15FC06F815.taxon	description	Figs 73 – 82, 133	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD611C40EEEA3D15FC06F815.taxon	diagnosis	Diagnosis. Male. Postcranium with lower half grayish-pruinose, upper half brownish-pruinose (Fig. 73). Hind trochanter with small triangular protuberance (Fig. 74). Tergites and sternites 6 and 7 as in Fig. 75. Surstyli asymmetrical, robust with laterobasal lobe setulose; apex somewhat apically truncated in dorsal view, right surstylus slightly shorter (Figs 76 – 77), both surstyli with tips slightly downward-directed when seen in lateral view (Figs 78 – 79). Apex of phallic guide short, angled at about 30 degrees (Fig. 80). Phallus with one ejaculatory duct with three spines reclined, the other two ducts without spine (Fig. 80). Ejaculatory apodeme shortened, distally truncated (Fig. 81). Gonopods subsymmetrical, right gonopod slightly longer (Fig. 82).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD611C40EEEA3D15FC06F815.taxon	materials_examined	Material examined. COLOMBIA, Bolivar, SFF [Santuario de Fauna y Flora] Los Colorados, Villa Roca, 9 º 54 ’ N / 75 º 07 ’ W, 180 m [eters], Malaise, 24. ix – 9. xi. 2001, E. Deulufeut leg, “ M 2419 ”, (1 ♂, IAvH) ” (photographed specimen); “ idem, Vichada, PNN [Parque Nacional Natural] Tuparro, Pie Cerro Tomas, 5 º 21 ’ N / 67 º 51 ’ W, 250 m [eters], 22. v. – 3. vi. 2001, W. Villalba leg “ M 1792 ” (1 ♂, LEUA) ”.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD611C40EEEA3D15FC06F815.taxon	distribution	Geographical distribution. Argentina; Cuba; Dominica; Mexico; Nicaragua; (Rodriguez & Rafael 2012); Colombia, Bolivar, Santuario de Fauna y Flora Los Colorados; Vichada, Parque Nacional Natural Tuparro (Fig. 133).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD611C40EEEA3D15FC06F815.taxon	biology_ecology	Habitat. The specimens were collected with Malaise traps at ground level in areas of tropical dry forest in the Caribbean region, at the Santuario de Fauna y Flora Los Colorados and Parque Nacional Natural Los Colorados, where the vegetation is composed of gallery forest with low elevation of the canopy from the ground in the Orinoco region of Colombia.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD611C40EEEA3D15FC06F815.taxon	discussion	Comments. The specimens collected are similar to the holotype illustrated by Ale-Rocha & Rafael (1995), differing only in the length, which exceeds in 1.16 mm that of the holotype.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD7D1C5CEEEA3998FC62FBE4.taxon	description	Figs 83 – 92, 133	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD7D1C5CEEEA3998FC62FBE4.taxon	diagnosis	Diagnosis. Male. Body shiny, robust (Fig. 83). Postcranium with lower three quarters grayish-pruinose, upper quarter brownish-pruinose (Fig. 83). Scutum mainly brownish-pruinose; thoracic setae long. Legs brownish-pruinose, except hind femur posteriorly and ventrally shiny. Hind trochanter with bulging, drop-shaped, protuberance, usually with a single ventral and a single dorsal seta (Fig. 84). Tergites and sternites 6 and 7 as in Fig. 85. Surstyli subsymmetrical, with external and internal margins somewhat straight, having slightly hooked-tips, right surstylus slightly shorter (Fig. 86 – 87), both surstyli with tips slightly downward-directed when seen in lateral view (Figs 88 – 89). Apex of phallic guide short, with inconspicuous spine, angled at about 20 degrees (Fig. 90). Phallus with ejaculatory ejaculary ducts elongated slender branches; one ejaculatory duct with three basal spines, the other two ducts without spines (Fig. 90). Ejaculatory apodeme sandal-shaped, darker near base (Fig. 91). Gonopods asymmetrical, left gonopod slender than right one (Fig. 92).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD7D1C5CEEEA3998FC62FBE4.taxon	materials_examined	Material examined. COLOMBIA, Boyacá, SFF [Santuario de Fauna y Flora] Iguaque, 5 º 25 ’ N / 73 º 27 ’ W, 2600 m [eters], Malaise, 16 – 30. xi. 2001, P. Reina leg, “ M 2583 ” (1 ♂, LEUA) (photographed specimen); “ idem, Vichada, PNN [Parque Nacional Natural] Tuparro, Centro Administrativo, 05 º 21 ’ N / 67 º 51 ’ W, 140 m [eters], Malaise 17, 12 – 29. Jun [vi]. 2000, V. Villalba (1 ♂, IAvH) ”.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD7D1C5CEEEA3998FC62FBE4.taxon	distribution	Geographical distribution. Bahamas; USA.; Mexico; Nicaragua (Rodriguez & Rafael 2012); Colombia (new record), Boyacá, Santuario de Fauna y Flora Iguaque; Vichada, Parque Nacional Natural Tuparro (Fig. 133).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD7D1C5CEEEA3998FC62FBE4.taxon	biology_ecology	Habitat. The specimens were collected with Malaise traps at ground level in areas of tropical dry forest, in the Northeast region in the Santuario de Fauna y Flora Iguaque, and in gallery forest with low elevation of the canopy from the ground in Parque Nacional Natural Tuparro in the Orinoco region of Colombia.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD7D1C5CEEEA3998FC62FBE4.taxon	discussion	Comments. The studied specimens are similar to the holotype illustrated by Ale-Rocha & Rafael (1995), differing only the length, which exceeds in 0.58 mm that of the holotype.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD7D1C59EEEA3D90FC45FE6E.taxon	description	Figs 93 – 102, 133	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD7D1C59EEEA3D90FC45FE6E.taxon	diagnosis	Diagnosis. Male. Body shiny (Fig. 95). Hind trochanter ventrally slightly elevated, white-setose (Fig. 94). Tergites and sternites 6 and 7 as in Fig. 95. Syntergosternite 8 symmetrical, sub-he mispherical (Figs 96 – 97). Surstyli asymmetrical, right surstylus wider than left, with distinct external sub-median lobe (Figs 96 – 97), when seen in lateral view both somewhat short and wide (Figs 98 – 99). Apex of phallic guide thickened, elongated, angled at about 45 degrees (Fig. 100). Phallus with ejaculatory ducts elongated, one ejaculatory duct somewhat coiled and noticable thicker than the other two (Fig. 100). Ejaculatory apodeme fan-shaped (Fig. 101). Gonopods subsymmetrical (Fig. 102).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD7D1C59EEEA3D90FC45FE6E.taxon	materials_examined	Material examined. COLOMBIA, Boyacá, SFF [Santuario de Fauna y Flora] Iguaque, 5 º 26 ’ N / 73 º 27 ’ W, 2600 m [eters], Malaise, 01 – 16. xii. 2001, A. Roberto leg, “ M 2582 ” (1 ♂, IAvH) (photographed specimen); “ idem, Antioquia, Envigado, Separador vial sector bodegas Éxito- Pirelli, Zona industrial, Jama, 6,16913 N / 75,60201 W, 15. ago [viii] – 01. sep [ix]. 2017, J. Torres-Toro (1 ♂, LEUA – 161730) ”.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD7D1C59EEEA3D90FC45FE6E.taxon	distribution	Geographical distribution. Species widely distributed in the Palearctic regions, Afrotropical, Indomalaya, Australian, Nearctic and Neotropical, Mexico Bahamas, Belize (Ale-Rocha & Rafael 1995), Argentina; Brazil; Chile; Costa Rica; Cuba; Dominica; Jamaica; Nicaragua; Puerto Rico; Colombia (Rodriguez & Rafael 2012), Antioquia, Envigado; Boyacá, Santuario de Fauna y Flora Iguaque (Fig. 133).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD7D1C59EEEA3D90FC45FE6E.taxon	biology_ecology	Habitat. The specimens were collected with Malaise traps at ground level at the Santuario de Fauna y Flora Iguaque reserve, in tropical dry forest of the Northeast region, and in urban areas at Envigado, Antioquia Andean region of Colombia.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD7D1C59EEEA3D90FC45FE6E.taxon	discussion	Comments. It is one of the most common species of this genus in the Neotropical Region. The specimens collected are identical to the holotype illustrated by Ale-Rocha (1996).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD781C5AEEEA3B18FF04FB52.taxon	description	Figs 103 – 119, 132	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD781C5AEEEA3B18FF04FB52.taxon	diagnosis	Diagnosis. Male. Postcranium with lower half grayish-pruinose, upper half dark-brownish pruinose (Fig. 103). Hind trochanter with variable stout spiniform protuberance (Fig. 104 – 108). Tergites and sternites 6 and 7 as in Fig. 109. Surstyli similar in shape, with hooked-tips, right surstylus slightly shorter (Figs 110 – 111), both surstyli dorsally lobated and tips slightly downward-directed when seen in lateral view (Figs 112 – 113). Phallic guide with short apex, bearing a small dorsal spine, not angled (Fig. 114). Phllus with ejaculatory ducts short, two ejaculatory ducts with one fine spine, the other ejaculatory duct with a spine (Fig. 114). Ejaculatory apodeme sandal-shaped, darker near base (Fig. 115). Gonopods asymmetrical (Fig. 116).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD781C5AEEEA3B18FF04FB52.taxon	description	Description of female. Body length 2 mm (Fig. 117). Wing 1.5 mm; frons medially grayish-pruinose, upper third sparser. Ocellar triangle dark-brown, shiny. Pedicel with two dorsal setae. Scutum dark-brown, with long band of setae. Scutellum brown, brownish-pruinose. Tegula yellow. Femora dark-brown except yellow apex. Abdomen (Fig. 117 – 118) brown, shiny metallic with blue tinge. Terminalia: Ovipositor OL: 0.50 mm, PL: 0.34 mm, B: 0.20 mm, base light-brown, in dorsal view longer than wide, piercer yellow, except basal part light-brown, apex shiny, straight (Fig. 119).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD781C5AEEEA3B18FF04FB52.taxon	materials_examined	Material examined. COLOMBIA, Caquetá, San José del Fragua, Vda. [Vereda] Bellavista, Fca. [Finca] Mi Ranchito, 01 º 18 ’ 23 ’’ N / 76 º 00 ’ 33 ’’ W, 265 m [eters], 26. x. – 09. xi. 2016, trampa Malaise en cultivo de caña (Sacharum officinarum), Y. Ramos-Pastrana (1 ♂, LEUA – 1222); “ idem, (1 ♂, LEUA – 1223) ”; “ idem, (1 ♂, LEUA – 1224) ”; “ idem, (1 ♂, LEUA – 1225) ”; “ idem, (1 ♂, LEUA – 1226) ”; “ idem, 09 – 23. xi. 2016, (1 ♂, LEUA – 1227) ”; “ idem, 23. xi. – 07. xii. 2016, (1 ♂, LEUA – 1228) ”; “ idem, 07 – 21. xii. 2016, (1 ♂, LEUA – 1229) ”; “ idem, 26. x. – 09. xi. 2016, (1 ♂, LEUA – 1230) ”; “ idem, (1 ♂, LEUA – 1231) ”; “ idem, 15 – 29. iii. 2017, (1 ♂, LEUA – 1232) ”; “ idem, 04 – 18. i. 2017, (1 ♂, LEUA – 1233) ”; “ idem, 26. x. – 09. xi. 2016, (1 ♂, LEUA – 1234) ”; “ idem, 15. ii. – 01. iii. 2017, (1 ♀, LEUA – 1287) ”; “ idem, Puerto Rico, Vda. [Vereda] La Soledad, Fca. [Finca] Borinquen, 01 º 55 ’ 26 ’’ N / 75 º 08 ’ 58.8 ’’ W, 270 m [eters], 07 – 21. xii. 2016, trampa Malaise en cultivo de caña (Sacharum officinarum), Y. Ramos-Pastrana (1 ♂, LEUA – 1235) ”; “ idem, Bolivar, SFF [Santuario de Fauna y Flora] Los Colorados, Villa Roca, 9 º 54 ’ N / 75 º 07 ’ W, 180 m [eters], Malaise, 24. ix – 9. xi. 2001, E. Deulufeut leg, “ M 2419 ”, (1 ♂, IAvH) ”; “ idem, 18 – 31. viii. 2001, “ M 2049 ” (1 ♂, IAvH); “ idem, 06 – 21. xiii [sic]. 2001, “ M 2630 ” (1 ♂, IAvH) ” (photographed specimen); “ idem, 14 – 30. xi. 2001, “ M 2627 ” (1 ♂, IAvH) ”; “ idem, Magdalena, PNN [Parque Nacional Natural] Tayrona, Zaino, 11 º 20 ’ N / 74 º 02 ’ W, 50 m [eters], Malaise, 04. Dic [xii] – 15. xii. 2000, R. Henriquez leg, “ M 966 ” (1 ♂, IAvH) ”; “ idem, (1 ♂, INPA) ”; “ idem, 15 – 31. dic [xii]. 2000, “ M 967 ” (1 ♂, INPA) ”; “ idem, 28. Jun [vi] – 17. Jul [vii]. 2000, “ M 301 ” (1 ♂, IAvH) ”; “ idem, Vaupes, Estación Biológica Mosiro-Itajura (Caparú) Centro Ambiental, 1 º 4 ’ S / 69 º 31 ’ W, 60 m [eters], Malaise, 17 – 24. xii. 2003, J. Pinzón leg, “ M 4436 ” (1 ♂, LEUA) ”; “ idem, 01 – 08. xii. 2003, “ M 4442 ” (1 ♂, IAvH) ”; “ idem, Vichada, PNN [Parque Nacional Natural] Tuparro, Cerro Tomas, 5 º 21 ’ N / 67 º 51 ’ W, 250 m [eters], Malaise, 22. v. – 3. vi. 2001, W. Villalba leg, “ M 1792 ” (1 ♂, LEUA) ”; “ idem, Amazonas, PNN [Parque Nacional Natural] Amacayacu, Mocagua, 03 º 23 ’ S / 70 º 06 ’ W, 300 m [eters], Malaise, 07 – 14. ago [viii]. 2000, A. Parente, “ M 677 ” (1 ♂, LEUA) ”; “ idem, Antioquia, Cimitarra, Central Termocentro Instalaciones, 6.44275 ºN / 74.88251 ºW, 118 m [eters], Trampa Malaise, 20 – 24. Ago [viii]. 2015, Y. Correa-C, (2 ♂, CEUA – 101594) ”; “ idem, 20 – 24. Ago [viii]. 2015, (1 ♀, CEUA – 101594) ” (photographed specimen).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD781C5AEEEA3B18FF04FB52.taxon	distribution	Geographical distribution. Bahamas; Brazil; Mexico; Costa Rica; Cuba; Dominica; Dominican Republic; Haiti; Jamaica; Santa Lucia (Rodriguez & Rafael 2012); Colombia (new record), Amazonas, Parque Nacional Natural Amacayacu; Antioquia, Cimitarra; Bolivar, Santuario de Fauna y Flora Los Colorados, Caquetá, San José del Fragua, Puerto Rico; Magdalena, Parque Nacional Natural Tayrona; Vaupes, Estación Biológica Mosiro-Itajúra (Caparú); Vichada, Parque Nacional Natural Tuparro (Fig. 132).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD781C5AEEEA3B18FF04FB52.taxon	biology_ecology	Habitat. The specimens were collected using Malaise traps at ground level in a number of different habitats including plantations, humid and dry tropical forests, gallery forest and urban araes. Plantations included sugar cane (Sacharum officinarum). Humid tropical forests were located in the Department of Caquetá, Colombia, in the Parque Nacional Natural Amacayacu and at the Estación Biológica Mosiro-Itajúra (Caparú) in the Department of Amazonas. Tropical dry forests were located in Parque Nacional Natural Tayrona and at Santuario de Fauna y Flora Los Colorados in the Northeat region of Colombia. Low elevation of the canopy from the ground gallery forests were located in Parque Nacional Natural Tuparro in the Orinoco region. Urban areas in the Department of Antioquia of the Andean region of Colombia were also sampled.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD781C5AEEEA3B18FF04FB52.taxon	discussion	Comments. Regarding the different shapes of the trochanter, there was no association of any particular shape with a specific locality. Usually different shapes of the trochanter are found in each locality. At this moment, all different morphos are being treated as a single species due to the fact that the shape of the genitalia is similar in all specimens. These variations in the hind trochanter had already been reported by Ale-Rocha & Rafael (1995). The male-female association made here is based on two males and one female that were collected together in the same trap.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD7B1C55EEEA3C30FEF6FEA0.taxon	description	Figs 120 – 131, 133	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD7B1C55EEEA3C30FEF6FEA0.taxon	diagnosis	Diagnosis. Male. Postcranium with lower half grayish-pruinose, upper half dark-brownish pruinose (Fig. 120). Tergite 1 entirely grayish-pruinose (Fig. 121). Hind trochanter with small protuberance (Figs 122 – 123). Tarsi brown (Fig. 123). Tergites and sternites 6 and 7 as in Fig. 124. Surstyli asymmetrical, nearly same length, having hookedtips, left surstylus with external margin somewhat straight, right surtylus with external margin curved (Figs 125 – 126), in lateral view left surstylus with dorsal margin straight (Fig. 127), right surstylus distinctly dorsally lobate (Fig. 128). Phallic guide with apex elongated, angled at about 55 degrees (Fig. 129). Phallus with distinct ventral projection before trifurcation, two branches with stout reclined spines (Fig. 129). Ejaculatory apodeme sandalshaped, darker near base (Fig. 130). Gonopods asymmetrical, left gonopod shorter (Fig. 131).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD7B1C55EEEA3C30FEF6FEA0.taxon	materials_examined	Material examined. COLOMBIA, Magdalena, PNN [Parque Nacional Natural] Tayrona, Neguanje, 11 º 20 ’ N / 74 º 02 ’ W, 10 m [eters], Malaise, 28. vii. – 18. viii. 2001, R. Henriquez leg, “ M 2019 ” (1 ♂, LEUA) (photographed specimen); “ idem, (1 ♂, INPA) ”; “ idem, 17 – 27. ix. 2001, “ M 2136 ” (1 ♂, IAvH) ”.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD7B1C55EEEA3C30FEF6FEA0.taxon	distribution	Geographical distribution. Venezuela; Colombia (new record), Magdalena, Parque Nacional Natural Tayrona (Fig. 133).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD7B1C55EEEA3C30FEF6FEA0.taxon	biology_ecology	Habitat. The specimens were collected with Malaise traps at ground level at the reserve Parque Nacional Natural Tayrona, where the vegetation is composed of thorny bushes, and in the tropical dry forests of the Caribbean region of Colombia.	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
507C87C0CD7B1C55EEEA3C30FEF6FEA0.taxon	discussion	Comments. The studied specimens are similar and longer in comparison with the holotype illustrated by Ale- Rocha (1993).	en	Ramos-Pastrana, Yardany, Rafael, José Albertino (2021): Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species. Zootaxa 4985 (1): 37-68, DOI: https://doi.org/10.11646/zootaxa.4985.1.2
