identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
2F5287828C6AFFA6FF71839DFCE4F996.text	2F5287828C6AFFA6FF71839DFCE4F996.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hedinomorpha Verhoeff 1934	<div><p>Genus Hedinomorpha Verhoeff, 1934</p> <p>Type species: Hedinomorpha hummelii Verhoeff, 1934, by monotypy.</p> <p>The following 24 species are currently included in Hedinomorpha, arranged combining both geographic and alphabetic orders:</p> <p>H. affinis Golovatch, 2014, H. altiterga Golovatch, 2019, H. flavobulbus Golovatch, 2019 and H. hummelii Verhoeff, 1934, all four from Gansu Province, China; H. bucharensis (Lohmander, 1933), from Tajikistan, Central Asia; H. biramipedicula Zhang &amp; Tang, 1985, from Shanxi Province, China; H. circofera Golovatch, 2013a and H. martensorum spec. nov., both from Qinghai Province; H. circularis (Takakuwa, in Takakuwa &amp; Takashima, 1949), H. jaegeri spec. nov., H. jeekeli (Golovatch, 2009) and H. sympatrica spec. nov., all four from Shaanxi Province; H. barbata Golovatch, 2020a, H. bifida Golovatch, 2019, H. crassiterga Golovatch, 2019, H. distincta Golovatch, 2020b, H. liuae Golovatch, 2020b, H. martensi Golovatch, 2014, H. nigra Golovatch, 2013b and H. reducta Golovatch, 2012, all from Sichuan Province; H. montana Golovatch, 2016b, H. proxima Golovatch, 2016b, H. subnigra Golovatch, 2013b and H. yunnanensis Golovatch, 2016b, all from Yunnan Province (Verhoeff 1934; Zhang &amp; Tang 1985; Jeekel 1988; Golovatch 2009, 2012, 2013a, 2013b, 2014, 2016b, 2019, 2020a, 2020b).</p> <p>The characters distinguishing the genus have recently been outlined by Golovatch (2019), but a brief description and an updated diagnosis seem warranted so as to properly assess the presently known species diversity and to discuss the closest affinities of Hedinomorpha.</p> <p>Brief description. A typical member of the tribe Sulciferini including only epigean, pigmented and mostly medium-sized to smaller species (10–40 mm long, 1.0–4.0 mm wide) with 20 body segments/rings and a normal pore formula (5, 7, 9, 10, 12, 13, 15–19). Paraterga from small, poorly developed and low (largely set at about half to upper third of midbody metaterga) to relatively strong and high keels (set at about upper quarter of midbody metaterga, rarely even elevated above dorsum), always showing calluses visible at least dorsally. Transverse sulci usually present on metaterga 5–17(18), but rarely up to 3–19. Pleurosternal carinae present, sometimes with small caudal teeth or lobes. Epiproct only sometimes conspicuously hirsute, enlarged and bulbous, but more usually typical, flattened dorsoventrally, its apex subtruncate, always devoid of terminal hooks. Legs relatively long and slender, male tarsal brushes often present, male prefemora not bulged laterad, adenostyles absent.</p> <p>Gonopods relatively simple, with a long coxite, subcylindrical and sparsely setose distoventrally, supplied with a small apicomesal cannula, devoid of modifications. Telopodites in situ crossing each other distally, each directed laterad and ca 2–3.5 times as long as coxite; prefemoral (= densely setose) part as usual, without modifications; femorite (fe) usually rather stout, but longer than either coxite or prefemoral part, suberect, slightly broadening distad, typically with a shallow gutter on mesal face to support a fully mesal seminal groove (sg) running all along, mostly (but not always) with a subtransverse distolateral sulcus (rarely two sulci) to delimit a small apical lobe (l); postfemoral region clearly bipartite, consisting of a long, free and flagelliform solenomere (sl) sheathed by and largely squeezed between a particularly strongly developed lamina lateralis (q) and a much smaller lamina medialis (j) of solenophore (sph), the latter a longer and stronger branch, loosely coiled, starting at base of l and never tightly spiraling, often barbed distally and acuminate apically, only rarely with 1–2 additional postfemoral outgrowths located immediately distal to l, but instead usually with 1–2 simple outgrowths (spines, lobes or ridges) near midway and/or distally.</p> <p>Diagnosis. Like most of the typical genera of Sulciferini, Hedinomorpha is characterized by two main branches arising distal to the gonopodal femorite: a solenophore and a solenomere (Jeekel 1968). Moreover, both main branches of the distal part of the gonopodal telopodite in Hedinomorpha are long and slender, directed laterad, and the femorite usually shows a subtransverse distolateral sulcus to delimit a small postfemoral lobe (l). However, Hedinomorpha differs from the most similar genera by a shallow to occasionally undeveloped mesal gutter of the femorite, coupled with a slender, relatively simple and loosely coiled (= never spiraling) solenophore which is typically supplied with a particularly strongly developed lamina lateralis (q) and a much smaller lamina medialis (j), as well as a single midway outgrowth (k) and/or a parabasal process (p), only occasionally with two parabasal processes. The solenomere is flagelliform, similar in length to solenophore, tightly squeezed between and sheathed by both laminae q and j, being exposed only distally. The solenophore is ribbon-shaped, often barbed distally and acuminate apically (see also Affinities and Key below).</p> </div>	http://treatment.plazi.org/id/2F5287828C6AFFA6FF71839DFCE4F996	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Golovatch, Sergei I.	Golovatch, Sergei I. (2021): Review of the millipede genus Hedinomorpha Verhoeff, 1934, with descriptions of three new species collected in China by Professor Jochen Martens (Diplopoda Polydesmida, Paradoxosomatidae). Zootaxa 4984 (1): 368-384, DOI: https://doi.org/10.11646/zootaxa.4984.1.27
2F5287828C68FFA1FF7182BDFE8EFC39.text	2F5287828C68FFA1FF7182BDFE8EFC39.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hedinomorpha martensorum Golovatch 2021	<div><p>Hedinomorpha martensorum spec. nov.</p> <p>Figs 1–15</p> <p>Type material. Holotype male and 1 male paratype (SMF), China, Qinghai Prov., Lake Koko Nor, southern shore, near Heimahe, 3400 m a.s.l., 22.VI.1996, J. Martens leg.</p> <p>Name. This is the second species of the genus devoted to Jochen Martens, this time to honour the entire Martens family: Jochen, the collector, his wife Beate, their daughters Charlotte and Friederike, and their son Felix. I have been privileged to enjoy their warm home, company and hospitality many times during my stays at and visits to Mainz.</p> <p>Diagnosis. Differs from other species of the genus by the following combination of characters: colouration brown, devoid of a vivid colour pattern; paraterga poorly-developed, their lateral calluses smooth and delimited by a complete and distinct sulcus only dorsally; pleurosternal carinae present on segments 2–8, traceable as bulges until segment 18; and the peculiar shapes of the solenophore and its outgrowths (see also Key below).</p> <p>Description (male). Length of both holo- and paratype ca 17–18 mm, width of midbody pro- and metazonae 1.6 and 1.8 mm, respectively. Colouration in alcohol mainly brown (Figs 1–4), devoid of a vivid colour pattern; most of head, paraterga, venter, legs and tip of epiproct lighter, yellow- or light brown; antennomeres 6 and 7 infuscate, dark brown, but tip of antennae pallid. Paratype generally a little lighter brown than holotype.</p> <p>Clypeolabral region moderately setose, vertex with a few setae; epicranial suture thin, but evident (Fig. 2). Antennae moderately long and only slightly clavate (Figs 1, 2), extending behind metatergum 2 when stretched dorsally (male). In length, antennomere 2&gt;3=4=5=6&gt;&gt;1=7. Interantennal isthmus about 1.5x as broad as diameter of antennal socket (Figs 1, 2). Tegument generally smooth and shining, only in places slightly rugulose above and below paraterga; surface below paraterga microgranulate; pro- and metazonae very delicately shagreened, strictures between them very faintly striolate; caudal margins of metaterga above a simple limbus densely and finely incised (Figs 3, 4). Tergal setae largely abraded or broken, few remaining ones thin and moderately long, ca 1/4–1/3 as long as metaterga; setation pattern traceable as 2+ 2 in a transverse fore (= pre-sulcus) row. In width, collum &lt;head &lt;segment 3=4 &lt;2 = 5–16, body gently and gradually tapering thereafter. Paraterga poorly developed, largely set at 1/3 of metatergal height, pore-bearing ones considerably thicker than poreless ones in lateral view, broadly rounded and delimited by a narrow sulcus on collum, clearly drawn both anteriad and caudad, and somewhat projecting beyond both tergal margins only on segment 2, caudal corner clearly rounded, never sharp and never extended behind rear tergal margin thereafter (Fig. 1). Lateral calluses subhorizontal, smooth and narrow, but evident, delimited by a distinct and complete sulcus only dorsally, with a fainter sulcus in caudal 1/3 also ventrally only on pore-bearing segments. Ozopores lateral, almost invisible from above, each lying inside an ovoid pit at ca 1/4–1/5 off caudal margin. Stricture between pro- and metazona thin and shallow, nearly smooth. Transverse metatergal sulci evident, thin, simple, slightly sinuate medially, far from reaching the bases of paraterga, present on segments 5–18, in places slightly punctured. Axial line missing. Pleurosternal carinae granulate ridges with rounded caudal flaps increasingly developed on segments 2–7, abruptly reduced to small flaps on segment 8, carinae being increasingly reduced and retained only as small bulges towards segment 18 (Fig. 1). Epiproct (Figs 4, 5) as usual, rather long, faintly concave to subtruncate at tip, lateral pre-apical papillae small. Hypoproct (Fig. 6) high, semi-circular, caudal margin with 1+1 long setae borne on flat rounded knobs. Sterna moderately setose, cross-impressions shallow, transverse sulci being slightly deeper than axial ones, without modifications except for a small, roundly subtrapeziform, poorly concave and setose lobe between coxae 4 (Figs 2, 8). No tubercles near gonopod aperture. Legs moderately long and slender, likely incrassate in male compared to a still unknown female, midbody ones ca 1.2–1.3x as long as body height, densely setose, neither adenostyles nor laterally swollen prefemora, ventral brushes on prefemora and tarsi until two last leg-pairs. In length, femur&gt; tarsus&gt; coxa = prefemur = tibia&gt; postfemur.</p> <p>Gonopods (Figs 1, 2, 9–15) largely suberect, in situ their distal parts coiled and in touch with each other; coxite subcylindrical, setose distoventrally, ca 2/3 as long as femorite (fe); prefemoral (= densely setose) part nearly half as long as fe; seminal groove (sg) usually running along fe at bottom of a shallow mesal groove/gutter (g) before moving onto a distinct flagelliform solenomere (sl) squeezed between a particularly strongly developed lamina lateralis (q) and a much smaller lamina medialis (j), and supported by a more complex postfemoral part; the latter forming a subtriangular, apical, postfemoral, lateral lobe (l) demarcated at base by a transverse lateral sulcus, and being largely represented by a long, slender, ribbon-shaped, loosely coiled, distally barbed and apically subacuminate solenophore branch (sph) supplied with a distinct spine (k) near midway and a rounded process/lobe (lo) in distal 2/3. No other outgrowths at base of l in addition to both main branches (sl and sph) except for a small tooth (t) distal to sl base.</p></div> 	http://treatment.plazi.org/id/2F5287828C68FFA1FF7182BDFE8EFC39	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Golovatch, Sergei I.	Golovatch, Sergei I. (2021): Review of the millipede genus Hedinomorpha Verhoeff, 1934, with descriptions of three new species collected in China by Professor Jochen Martens (Diplopoda Polydesmida, Paradoxosomatidae). Zootaxa 4984 (1): 368-384, DOI: https://doi.org/10.11646/zootaxa.4984.1.27
2F5287828C6EFFACFF7186ACFEE1FBE1.text	2F5287828C6EFFACFF7186ACFEE1FBE1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hedinomorpha jaegeri Golovatch 2021	<div><p>Hedinomorpha jaegeri spec. nov.</p> <p>Figs 16–28</p> <p>Material examined. Holotype male and 1 male paratype (SMF), China, Shaanxi Prov., Taibai Shan Mts, S flanks, above Houshenzi, 1700 m a.s.l., primary broadleaved forest, 20.VII.1997, P. Jäger &amp; J. Martens leg.</p> <p>Name. This species is dedicated to Peter Jäger, the main collector.</p> <p>Diagnosis. Differs from other species of the genus by the following combination of characters: colouration brown to nearly pallid, devoid of a vivid colour pattern; paraterga poorly-developed, their pore-bearing and poreless calluses with two or three lateral incisions, respectively, mostly sharp caudally, delimited by a distinct sulcus only dorsally and a very narrow sulcus ventrally, both sulci being complete; pleurosternal carinae present only on segment 2, and the peculiar shapes of the solenophore and its outgrowths (see also Key below).</p> <p>Description (male). All characters as in H. martensorum spec. nov., except as follows.</p> <p>Length of holo- and paratype ca 14 or 10 mm, width of midbody pro- and metazonae 0.95 and 1.1 or 0.85 and 1.0 mm, respectively. Colouration in alcohol mainly brown (holotype) (Figs 16–19) or pallid, devoid of a vivid colour pattern; in holotype, legs and tip of epiproct lighter, nearly pallid; antennomeres 6 and 7 slightly infuscate, light brown, but tip of antennae pallid.</p> <p>Clypeolabral region rather densely setose. In width, collum &lt;segment 3=4 &lt;2 = 5–16 &lt;head, body gently and gradually tapering on segments 17–20. Tegument generally shining and finely shagreened, most metaterga slightly tuberculate, in places rugulose to rugose above and below paraterga; surface below paraterga very delicately microgranulate; strictures between pro- and metazonae densely ribbed (Figs 16, 18, 19). Tergal setae often abraded, but remaining ones thin and moderately long, mostly ca 1/3 as long as metaterga; setation pattern traceable as 2+ 2 in a transverse pre-sulcus row, setae being borne on evident, but small bosses/tubercles; about 3+3 similar tuberculations arranged in two transverse rows, one each in pre-sulcus (with 2+2 setigerous) and post-sulcus halves of metaterga, but rear row always devoid of setae or their insertion points. Dorsal surface of collum and segment 2 smooth, that of metaterga 3–18 uneven, slightly tuberculate, with a mostly unusually deep, microgranulate and wide transverse sulcus, and an axial line. Paraterga poorly developed, largely set at upper 1/3 of metatergal height, clearly drawn only anteriad and somewhat projecting beyond fore tergal margin only on segment 2; caudal corner drawn into a small and sharp tooth, but never extended behind rear tergal margin thereafter (Figs 16–19). Lateral calluses subhorizontal, narrow, with two (pore-bearing segments) or three (poreless ones) small setigerous incisions at lateral margin, delimited by complete sulci both dorsally and ventrally, but ventral sulcus very narrow and indistinct. Ozopores lateral, almost invisible from above, each lying inside an ovoid pit at ca 1/4–1/5 off caudal margin. Transverse metatergal sulci exceptionally evident, deep and wide, microgranulate at bottom, slightly sinuate medially, reaching the bases of paraterga, present on segments 3–19, but less distinct on segments 3, 4 and 19. Axial line sulciform, present on both halves of metazonae. Pleurosternal carina a small squarish lappet only on segment 2, carinae totally missing thereafter (Fig. 17). Epiproct (Figs 19, 20) as usual, rather long, faintly concave to subtruncate at tip, lateral pre-apical papillae small. Hypoproct (Fig. 21) roundly subtrapeziform, caudal margin with 1+1 long setae borne on flat rounded knobs. Sterna without modifications except for a lower, roundly subtrapeziform, slightly concave and setose lobe between coxae 4 (Figs 17, 22). Legs moderately long and slender, obviously incrassate in male compared to a still unknown female, midbody ones ca 1.3–1.4x as long as body height, mostly less densely setose, ventral brushes on tarsi present only until leg-pair 9 or 10, gradually thinning out thereafter (Fig. 17).</p> <p>Gonopods (Figs 17, 23–28) largely suberect, in situ their distal parts coiled and in touch with each other; coxite subcylindrical, setose distoventrally, ca 2/3 as long as femorite (fe); prefemoral (= densely setose) part nearly half as long as fe; seminal groove (sg) running along fe at bottom of a shallow mesal groove/gutter (g) before moving onto a distinct flagelliform solenomere (sl) squeezed between a particularly strongly developed lamina lateralis (q) and a much smaller lamina medialis, and supported by a more complex postfemoral part; the latter forming a subtriangular, apical, postfemoral, lateral lobe (l) demarcated at base by a transverse sulcus, and being largely represented by a long, ribbon-shaped, loosely coiled, distally barbed and apically subacuminate solenophore branch (sph) supplied with a distinct spine (k) near midway and a small rounded lobule (l) in distal 2/3. A large, ribbon-shaped, curved and apically acuminate process (p) set at base of l in addition to both main branches (sl and sph) and a small subapical tooth (d).</p></div> 	http://treatment.plazi.org/id/2F5287828C6EFFACFF7186ACFEE1FBE1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Golovatch, Sergei I.	Golovatch, Sergei I. (2021): Review of the millipede genus Hedinomorpha Verhoeff, 1934, with descriptions of three new species collected in China by Professor Jochen Martens (Diplopoda Polydesmida, Paradoxosomatidae). Zootaxa 4984 (1): 368-384, DOI: https://doi.org/10.11646/zootaxa.4984.1.27
2F5287828C63FFA8FF718548FA74F921.text	2F5287828C63FFA8FF718548FA74F921.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hedinomorpha sympatrica Golovatch 2021	<div><p>Hedinomorpha sympatrica spec. nov.</p> <p>Figs 29–43</p> <p>Material examined. Holotype male, 1 male (without gonopods) and 3 female paratypes (SMF), China, Shaanxi Prov., Taibai Shan Mts, S flanks, above Houshenzi, 1700 m a.s.l., primary broadleaved forest, 20.VII.1997, P. Jäger &amp; J. Martens leg.</p> <p>Name. To emphasize strict sympatry, and even syntopy, with H. jaegeri spec. nov.; adjective.</p> <p>Diagnosis. Differs from other species of the genus by the following combination of characters: colouration light brown to brown, devoid of a vivid colour pattern; paraterga poorly-developed, their pore-bearing and poreless calluses smooth lasterally, mostly blunt and subrectangular teeth caudally, delimited by a distinct and complete sulcus only dorsally; pleurosternal carinae present on segments 2–15(16), and the peculiar shapes of the solenophore and its outgrowths (see also Key below).</p> <p>Description. All characters as in H. martensorum spec. nov., except as follows.</p> <p>Length of male holo- and paratype ca 12 mm, width of midbody pro- and metazonae 0.85 and 1.0 mm, respectively. Length of female paratypes ca 13 mm, width of midbody pro- and metazonae 1.3–1.35 and 1.5–1.55 mm, respectively. Colouration in alcohol mainly light brown (Figs 29–32) to brown, devoid of a vivid colour pattern, but with a slightly lighter, broad, dorsal stripe divided by a darker axial line; sides of head, venter, legs and tip of epiproct lighter, nearly pallid; antennomeres 6 and 7 slightly infuscate, light brown to brown, but tip of antennae pallid.</p> <p>Clypeolabral region moderately setose, vertex bare (Figs 29, 30). Antennae shorter and only slightly clavate (Figs 29, 30), extending behind until about midway of segment 2 (male) or midway of collum (female) when stretched dorsally. Interantennal isthmus about 1.3x as broad as diameter of antennal socket. Tegument generally smooth and shining, metaterga slightly rugulose above and below paraterga; strictures between pro- and metazonae finely striolate. Tergal setae often abraded, but remaining ones thin and moderately long, mostly ca 1/3 as long as metaterga; setation pattern traceable as 2+ 2 in a transverse pre-sulcus row, setae not being borne on bosses. Paraterga poorly developed, largely set at 1/2 of metatergal height, slightly drawn both anteriad and caudad only on segment 2, and only caudad on segment 3; caudal corner drawn into a small, mostly subrectangular and blunt tooth, but never extended behind rear tergal margin thereafter (Figs 29–32). Lateral calluses subhorizontal, in lateral view narrow (poreless segments) or thicker (pore-bearing segments), smooth at lateral margin, delimited by complete sulci only dorsally. Ozopores lateral, invisible from above, each lying inside an ovoid pit a little in front of or at caudal margin (Fig. 29). Transverse metatergal sulci very simple, shallow, smooth at bottom, slightly sinuate medially, not reaching the bases of paraterga, present on segments 5–18, but far less distinct on segment 18. Axial line missing. Pleurosternal carinae microgranulate ridges, in male increasingly developed on segments 2–7, abruptly reduced to a small ridge on segment 8, with small, rounded, caudal flaps only on segments 4–7; in female smaller, thin ridges devoid of caudal flaps and visible until segment 9; in both sexes being increasingly reduced and retained only as small bulges towards segment 15 or 16 (Fig. 29). Epiproct (Figs 29, 32) bulbous and hirsute. Hypoproct (Fig. 40) semi-circular, caudal margin with 1+1 long setae borne on minute knobs. Sterna without modifications except for a low, setose, roundly subtrapeziform and medially slightly concave lobe between male coxae 4 (Fig. 41). Legs (Figs 29–33) moderately long and slender, obviously incrassate in male compared to female, midbody ones ca 1.3–1.4x (male) or 0.9–1.0x (female) as long as body height, densely setose, ventral brushes on tarsi present until two last leg-pairs in male (Fig. 33).</p> <p>Gonopods (Figs 34–39, 42, 43) much like in H. martensi spec. nov., but prefemoral (= densely setose) part almost as long as femorite (fe); solenomere (sl) also squeezed between a particularly strongly developed lamina lateralis (q) and a much smaller lamina medialis (j), and supported by a similarly complex postfemoral part; the latter forming a rounded, apical, postfemoral, lateral lobe (l) demarcated at base by a subtransverse sulcus, and being largely represented by a long, ribbon-shaped, loosely coiled, distally barbed and apically subacuminate solenophore branch (sph) supplied with a stronger spine (k) near midway and a similarly small rounded lobule (l) in distal 2/3.</p> </div>	http://treatment.plazi.org/id/2F5287828C63FFA8FF718548FA74F921	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Golovatch, Sergei I.	Golovatch, Sergei I. (2021): Review of the millipede genus Hedinomorpha Verhoeff, 1934, with descriptions of three new species collected in China by Professor Jochen Martens (Diplopoda Polydesmida, Paradoxosomatidae). Zootaxa 4984 (1): 368-384, DOI: https://doi.org/10.11646/zootaxa.4984.1.27
