taxonID	type	description	language	source
B4615401FF842C1EFD71AAB3FDA3FABF.taxon	description	(Figs. 1 – 5)	en	Smith, Andrew B. T., Paulsen, M. J. (2017): Two New Species of Podolasia Harold (Coleoptera: Scarabaeidae: Melolonthinae: Podolasiini) from Coahuila, Mexico. The Coleopterists Bulletin 71 (3): 532-536, DOI: 10.1649/0010-065X-71.3.532, URL: http://dx.doi.org/10.1649/0010-065x-71.3.532
B4615401FF842C1EFD71AAB3FDA3FABF.taxon	materials_examined	Type Material. Holotype male (UNMC) labeled: a) “ Mexico: Coahuila: Cuatro / Cienegas Prot. Area / 16. V. 2010 / 26.9717 ° N 102.1224 ° W / Colr. K. R. Wetherill ”; b) “ Site A 1 / ~ 6 km W. Cuatrocienegas / lower bajada / general collecting ”; c) on red paper, “ HOLOTYPE / Podolasia diabla / Smith & Paulsen ”.	en	Smith, Andrew B. T., Paulsen, M. J. (2017): Two New Species of Podolasia Harold (Coleoptera: Scarabaeidae: Melolonthinae: Podolasiini) from Coahuila, Mexico. The Coleopterists Bulletin 71 (3): 532-536, DOI: 10.1649/0010-065X-71.3.532, URL: http://dx.doi.org/10.1649/0010-065x-71.3.532
B4615401FF842C1EFD71AAB3FDA3FABF.taxon	description	Description. Male, length 4.1 mm, greatest width 1.7 mm. Color dorsally and ventrally light reddish brown with frons and pronotum slightly darker (Figs. 1 – 2). Head subcircular with clypeal and genal margins nearly continuous. Clypeus (Fig. 3) narrower than 2 X length at middle, sides reflexed, more strongly so anteriorly, anterior margin rounded, not emarginate; posterior margin coplanar with frons, frontoclypeal suture visible laterally, absent medially; clypeal disc with scattered, moderate-size punctures; surface shiny with erect setae. Labrum not visible, presumably fused with clypeus. Genal margin arcuate, almost obsolete, curved, raised edge distinct. Frons lacking secondary transverse carina, with strong triangular horn above each eye (Fig. 3); surface concave, shiny. Mouthparts reduced, except for labial and maxillary palpomeres. Antenna with 9 antennomeres. Pronotum strongly convex (Figs. 1 – 2), declivous anteriorly, widest at middle; marginal bead complete, thin laterally, crenate near obtuse anterior angles; just posterior to wide anterior margin with distinct, setose transverse groove extending to anterior angles; lateral margin with sparse, long setae, posterior margin with short setae; anterior angles slightly produced near eyes, obtuse; posterior angles obsolete; pronotal disc except near anterior and posterior angles with sparse, large punctures, punctures closer laterally than on disc, surface between punctures shiny. Scutellum as long as wide, longitudinally depressed, tip bluntly rounded. Elytron lacking impressed striae, 4 striae indicated by irregular rows of large punctures, some punctures with fine, semi-erect seta; intervals between rows of punctures with irregular, sparse punctures lacking setae; interval punctures mixed fine to large; surface shiny; apex of elytron rounded near suture. Pygidium relatively short and broad, surface with scattered punctures, each with a long seta; surface shiny, feebly reticulate basally. Abdomen with ventral sutures mostly effaced (connate), only apical and basal segments with defined sutures. Protibia with 2 teeth (Figs. 1 – 2). Metafemur oval-elongate, approximately 2 X long as wide. Metatibia elongate, distinctly 3 X longer than wide (Fig. 1); slanted carina extending less than 1 / 2 of outer face near apical third of tibia, outer surface without distinct tubercles. Tarsal claws simple. Each paramere of genitalia (Fig. 5) sinuate with distinctive triangular expansion apically. Wings fully developed. Female. Unknown.	en	Smith, Andrew B. T., Paulsen, M. J. (2017): Two New Species of Podolasia Harold (Coleoptera: Scarabaeidae: Melolonthinae: Podolasiini) from Coahuila, Mexico. The Coleopterists Bulletin 71 (3): 532-536, DOI: 10.1649/0010-065X-71.3.532, URL: http://dx.doi.org/10.1649/0010-065x-71.3.532
B4615401FF842C1EFD71AAB3FDA3FABF.taxon	discussion	Remarks. Podolasia diabla is the only species in the genus to possess horns on the head. Despite the small size of the only known specimen, the horns are well developed. Podolasia diabla keys to Podolasia pilosa Howden, 1954 in Howden (1997), but it is easily distinguished from both P. pilosa and Podolasia parapilosa Smith and Paulsen, new species by the head horns and the subcircular head with nearly continuous clypeal and genal margins.	en	Smith, Andrew B. T., Paulsen, M. J. (2017): Two New Species of Podolasia Harold (Coleoptera: Scarabaeidae: Melolonthinae: Podolasiini) from Coahuila, Mexico. The Coleopterists Bulletin 71 (3): 532-536, DOI: 10.1649/0010-065X-71.3.532, URL: http://dx.doi.org/10.1649/0010-065x-71.3.532
B4615401FF842C1EFD71AAB3FDA3FABF.taxon	etymology	Etymology. We name this species diabla, the feminine form of the Spanish word for “ devil ”, in regards to the small horns that are unique in the tribe and rarely found in the subfamily Melolonthinae. This name should be treated as a noun in apposition.	en	Smith, Andrew B. T., Paulsen, M. J. (2017): Two New Species of Podolasia Harold (Coleoptera: Scarabaeidae: Melolonthinae: Podolasiini) from Coahuila, Mexico. The Coleopterists Bulletin 71 (3): 532-536, DOI: 10.1649/0010-065X-71.3.532, URL: http://dx.doi.org/10.1649/0010-065x-71.3.532
B4615401FF862C18FF67AA66FF35FC2B.taxon	description	(Figs. 6 – 11)	en	Smith, Andrew B. T., Paulsen, M. J. (2017): Two New Species of Podolasia Harold (Coleoptera: Scarabaeidae: Melolonthinae: Podolasiini) from Coahuila, Mexico. The Coleopterists Bulletin 71 (3): 532-536, DOI: 10.1649/0010-065X-71.3.532, URL: http://dx.doi.org/10.1649/0010-065x-71.3.532
B4615401FF862C18FF67AA66FF35FC2B.taxon	materials_examined	Type Material. Holotype male (UNMC) labeled: a) “ Mexico: Coahuila: Cuatro / Cienegas Nat’ l. Preserve / 21. VI. 2011 / 26.84484 ° N 102.17866 ° W / D. C. Lightfoot et al., colrs ”; b) “ Site B 3 / ~ 19.43 km SW of Cuatro / gypsum dunes, pedestals / general collecting ”; c) on red paper, “ HOLOTYPE / Podolasia parapilosa / Smith & Paulsen ”. Eight male paratypes (3 CMNC, 1 MJPC, 1 UNAM, 2 UNMC, 1 UNSM) labeled as holotype except for c) on yellow paper “ PARATYPE / Podolasia parapilosa / Smith & Paulsen ”. Twenty male paratypes (4 CMNC, 2 MJPC, 1 UNAM, 10 UNMC, 3 UNSM) labeled: a) “ Mexico: Coahuila: Cuatro / Cienegas Prot. Area / 18. V. 2010 / 26.8336 ° N 102.1620 ° W / D. Lightfoot, K. R. Wetherill, et al. ”; b) “ Site B 1 / ~ 19 km SW Cuatrocienegas / gypsum dune / light trap, oatmeal line / general collecting ”; c) on yellow paper “ PARATYPE / Podolasia parapilosa / Smith & Paulsen ”. Twelve male paratypes (2 CMNC, 8 UNMC, 2 UNSM) labeled: a) “ Mexico: Coahuila: Cuatro- / Cienegas Prot. Area 18. V. 2010 / 26.833 ° N 102.162 ° W, ~ 20 km / SW Cuatrocienegas, colr. D. C. / Lightfoot, K. R. Wetherill, et al. ”; b) “ Site B 1 light trap / Gypsum Dunes ”; c) on yellow paper “ PARATYPE / Podolasia parapilosa / Smith & Paulsen ”.	en	Smith, Andrew B. T., Paulsen, M. J. (2017): Two New Species of Podolasia Harold (Coleoptera: Scarabaeidae: Melolonthinae: Podolasiini) from Coahuila, Mexico. The Coleopterists Bulletin 71 (3): 532-536, DOI: 10.1649/0010-065X-71.3.532, URL: http://dx.doi.org/10.1649/0010-065x-71.3.532
B4615401FF862C18FF67AA66FF35FC2B.taxon	description	Description. Male, length 6.5 mm, greatest width 2.5 mm. Color dorsally and ventrally reddish brown with head and pronotum slightly darker. Head apically arcuate (not subcircular), clypeal and genal margins not continuous. Clypeal length and width subequal (Fig. 10), sides reflexed, anteriorly thickened either side of weak median emargination, posteriorly not coplanar with frons; frontoclypeal suture carinate; clypeal disc with scattered large punctures, each with a long seta, surface between punctures roughened with micropunctures. Labrum not visible, presumably fused with clypeus. Genal margin distinctly produced, rounded; raised edge thickened anteriorly. Frons lacking secondary transverse carina, instead with oblique irregular carinae parallel to eyes; surface rugopunctate. Mouthparts reduced, except for labial and maxillary palpomeres. Antenna with 9 antennomeres (Fig. 10). Pronotum weakly convex (Figs. 6 – 7), not declivous anteriorly, widest at middle, with marginal bead complete, thin laterally, not distinctly crenate near anterior angles; lacking distinct transverse groove behind anterior margin; lateral margin with dense, long setae; setae becoming short and sparse on posterior margin. Anterior angles slightly produced near eyes, acute; posterior angles obsolete; pronotal disc except near anterior and posterior margins with sparse, large punctures, punctures closer laterally than on disc, surface between punctures shiny; some lateral punctures with setae. Scutellum longer than wide, with bluntly rounded apex, surface smooth, impunctate. Elytron lacking impressed striae, 4 striae indicated by irregular rows of large punctures, some punctures with a fine, semi-erect seta; intervals between rows of punctures with irregular, sparse punctures lacking setae; interval punctures mixed fine to large; surface shiny; apex of elytron rounded near suture. Pygidium relatively short and broad, surface with scattered punctures, each with a long seta; surface shiny, feebly reticulate basally. Abdomen with ventral sutures mostly effaced (connate), only apical and basal segments with defined sutures. Protibia with 2 teeth. Metafemur oval, less than 2 X long as wide. Metatibia elongate, about 2 X longer than width at apex; apex with basotarsal flange narrowly produced in lateral view; slanted carina extending across 1 / 2 of outer face near apical third of tibia, outer surface without distinct tubercles. Tarsal claws simple. Each paramere of genitalia (Fig. 11) straight externally with distinctive triangular expansion apically. Wings present. Female. Unknown. Variation. Length 4.5 – 6.3 mm, greatest width 1.5 – 2.6 mm. Smaller specimens lack a clypeal emargination.	en	Smith, Andrew B. T., Paulsen, M. J. (2017): Two New Species of Podolasia Harold (Coleoptera: Scarabaeidae: Melolonthinae: Podolasiini) from Coahuila, Mexico. The Coleopterists Bulletin 71 (3): 532-536, DOI: 10.1649/0010-065X-71.3.532, URL: http://dx.doi.org/10.1649/0010-065x-71.3.532
B4615401FF862C18FF67AA66FF35FC2B.taxon	discussion	Remarks. Podolasia parapilosa keys to P. pilosa in Howden (1997). Podolasia parapilosa is distinguished by the clypeus being longer than wide (wider than long in P. pilosa) and usually weakly emarginate (never emarginate in P. pilosa), the metatibiae being 2 – 3 X as thick at the apex compared to the base (at least 3 X thicker in P. pilosa), the straight outer edges of the parameres (slightly bowed medially in P. pilosa), and the distribution in Coahuila, Mexico (New Mexico, Chihuahua, and bordering areas of Texas in P. pilosa). When describing the tribe Podolasiini, Howden (1997) provided a list of other “ primitive ” melolonthine taxa that shared some characters. One additional character that we observed and warrants mention is the sclerotized male genital capsule (Fig. 9). This structure has two apical knobs with forward projecting setae, which is remarkable for a taxon in the phytophagous scarab clade (Smith et al. 2006). The vast majority of phytophagous scarabs have a soft genital capsule without hard sclerotized parts or setae.	en	Smith, Andrew B. T., Paulsen, M. J. (2017): Two New Species of Podolasia Harold (Coleoptera: Scarabaeidae: Melolonthinae: Podolasiini) from Coahuila, Mexico. The Coleopterists Bulletin 71 (3): 532-536, DOI: 10.1649/0010-065X-71.3.532, URL: http://dx.doi.org/10.1649/0010-065x-71.3.532
B4615401FF862C18FF67AA66FF35FC2B.taxon	etymology	Etymology. We name this species P. parapilosa, meaning “ close to P. pilosa ”, to indicate the many similarities between these two species. This name should be treated as an adjective in the nominative singular.	en	Smith, Andrew B. T., Paulsen, M. J. (2017): Two New Species of Podolasia Harold (Coleoptera: Scarabaeidae: Melolonthinae: Podolasiini) from Coahuila, Mexico. The Coleopterists Bulletin 71 (3): 532-536, DOI: 10.1649/0010-065X-71.3.532, URL: http://dx.doi.org/10.1649/0010-065x-71.3.532
