identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
038B87913142FFCA7B7E1463F42627B8.text	038B87913142FFCA7B7E1463F42627B8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cochlostoma (Clessiniella) Zallot, Groenenberg, De Mattia, Feher & Gittenberger 2015	<div><p>Subgenus Clessiniella Zallot, Groenenberg, De Mattia, Fehér &amp; Gittenberger, 2015</p> <p>Clessiniella Zallot, Groenenberg, De Mattia, Fehér &amp; Gittenberger, 2015: 80.</p> <p>Type species</p> <p>Pomatias maculatum var. villae Strobel, 1851, by original designation.</p> <p>Remarks</p> <p>The classification follows MolluscaBase (2021) except for the subgeneric name. Contrary to ICZN Art. 10.4, Neglecta Wagner, 1897 is used in MolluscaBase as a senior synonym of Clessiniella. Wagner (1897: 6 [570]) introduced Neglecta as “Formenkreis Neglecta ”. It cannot be denied that the German term Formenkreis is “.. a term such as “superspecies””. As a consequence, according to the ICZN ruling, Neglecta “.. is not deemed to be a genus-group name” (ICZN Art. 10.4, see also Zallot et al. 2015). Strobel (1851) and De Betta &amp; Martinati (1855) named this entity Pomatias maculatum var. Villae – but referring to Pomatias maculatus Draparnaud, 1805 (= Co. (Co.) septemspirale).</p> <p>Diagnosis of the subgenus</p> <p>SHELL. Ribbed teleoconch with straight ribs, variable in strength and shape. The columellar lobe is curved backwards, covering the umbilicus. The teleoconch spots (if present) form 2 rows on the whorls (often, a 3 rd row of spots is visible on the body whorl): the upper one close to the suture with rectangular spots abapically elongated; the lower one roughly ⅓ up on the whorls (Fig. 12). In some populations the 2 rows of spots merge and there is a continuous vertical darker band on the whorls. The protoconch is dull and robust with fine riblets starting after 1.2 whorls or more from the apex; it is transparent and fragile or the ribbing starts close to the apex in other subgenera.</p> <p>FEMALE GENITALIA. There is an apical connection of the pedunculus to the bursa copulatrix, a long seminal receptacle (normally club-shaped), which runs parallel to the pedunculus. The junction of the uterus gland is situated close to the connection between the pedunculus and the distal oviduct. This</p> <p>combination of characters in the female genitalia is most diagnostic to distinguish Clessiniella from the other subgenera.</p> <p>MALE GENITALIA. Apart from its apical part, the penial spermiduct is tortuous. The body spermiduct is protruding (often almost detached from the body) more than in the other subgenera and either straight or more or less tortuous. The posterior side of the penis is wrinkled.</p> <p>Identification key for the species</p> <p>1. Body whorl inflated shortly before the lip (Fig. 4A: OB). In the female genitalia, simplified loop path with 1–2 loops............................................................................ Co. (Cl.) anomphale Boeckel, 1939</p> <p>– Body whorl not inflated before the lip (Fig. 4A: FF, AF, II)............................................................. 2</p> <p>2. In the female genitalia, the loops of the visceral oviduct are 1–2........ Co. (Cl.) stelucarum sp. nov.</p> <p>– In the female genitalia, the loops of the visceral oviduct are more than 2....................................... 3</p> <p>3. In the male genitalia, the body spermiduct is twisted............... Co. (Cl.) waldemari (Wagner, 1897)</p> <p>– In the male genitalia, the body spermiduct is straight...................................................................... 4</p> <p>4. Ribs of the shell widely spaced and barely inclined, protoconch relatively small (Fig. 13). Slim, club-shaped seminal receptacle in the female genitalia..................... Co. (Cl.) villae (Strobel, 1851)</p> <p>– Ribs less widely spaced and more inclined, relatively larger protoconch. Less slim seminal receptacle.......................................................................... Co. (Cl.) tergestinum (Westerlund, 1878)</p></div> 	http://treatment.plazi.org/id/038B87913142FFCA7B7E1463F42627B8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Zallot, Enrico;Mattia, Willy De;Fehér, Zoltán;Gittenberger, Edmund	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B8791315FFFC17BAB1477F36A252B.text	038B8791315FFFC17BAB1477F36A252B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cochlostoma (Clessiniella) villae (Strobel 1851)	<div><p>Cochlostoma (Clessiniella) villae (Strobel, 1851)</p> <p>Figs 1, 3C, 14D, 15D, 16D, 17–21</p> <p>Pomatias maculatum var. villae Strobel, 1851: 17 (“Mizzole presso Verona”).</p> <p>Pomatias maculatum var. villae De Betta &amp; Martinati, 1855: 74 (“Prov. Verona, Mizzole”).</p> <p>Cochlostoma (Auritus) villae – Alzona 1971: 22.</p> <p>Cochlostoma villae − Welter-Schultes 2012: 102.</p> <p>Diagnosis</p> <p>The last whorl, approaching the aperture, hardly enlarges (Fig. 4A: FF). Very prominent and widely spaced teleoconch ribs, barely inclined in respect to the vertical axis. Spots almost always visible. The protoconch is relatively small. The apertural lip poorly developed. Visceral oviduct with convoluted loops; thin seminal receptacle. Straight body spermiduct.</p> <p>Type material</p> <p>Syntypes</p> <p>ITALY • 1 ♀; “ Pomatias villae Spinel. Mizzole. Villa 49” [label of the original material] [Veneto, Verona, Mizzole] [thus, it could be interpreted as collected in 1849 by Villa]; PU1610-C1 (Fig. 19) • 2 ♂♂; same label data as for preceding; PU1610-C2, PU1610-C3 (Fig. 20).</p> <p>Other material (fresh specimens from the type locality have also been analyzed)</p> <p>ITALY • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=11.25225&amp;materialsCitation.latitude=45.61511" title="Search Plazi for locations around (long 11.25225/lat 45.61511)">Altissimo</a>; 45.61511° N, 11.25225° E; 2009; Zallot leg.; EZ0883 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=11.69623&amp;materialsCitation.latitude=45.26194" title="Search Plazi for locations around (long 11.69623/lat 45.26194)">Arqua’ Petrarca</a>, surroundings; 45.26194° N, 11.69623° E; 1999; Zallot leg.; EZ0895 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=11.20972&amp;materialsCitation.latitude=45.59389" title="Search Plazi for locations around (long 11.20972/lat 45.59389)">Bolca</a>; 45.59389° N, 11.20972° E; 2010; Zallot leg.; EZ0882 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=10.90787&amp;materialsCitation.latitude=45.58921" title="Search Plazi for locations around (long 10.90787/lat 45.58921)">Ca’Gottolo</a>; 45.58921° N, 10.90787° E; Jun. 2010; Scarlassara leg.; EZ1015 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=11.36833&amp;materialsCitation.latitude=45.60667" title="Search Plazi for locations around (long 11.36833/lat 45.60667)">Cornedo Vicentino-Cereda</a>; 45.60667° N, 11.36833° E; 1999; Zallot leg.; EZ0887 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=11.12083&amp;materialsCitation.latitude=45.60889" title="Search Plazi for locations around (long 11.12083/lat 45.60889)">Covoli di Velo</a>; 45.60889° N, 11.12083° E; 2010; Zallot leg.; EZ0879 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=11.19538&amp;materialsCitation.latitude=45.62137" title="Search Plazi for locations around (long 11.19538/lat 45.62137)">Ferrazza</a>; 45.62137° N, 11.19538° E; Apr. 2010; Zallot leg.; EZ0881 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=10.89856&amp;materialsCitation.latitude=45.56942" title="Search Plazi for locations around (long 10.89856/lat 45.56942)">Fumane-Molina</a>; 45.56942° N, 10.89856° E; Jun. 2010; Scarlassara leg.; EZ0936 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=11.54&amp;materialsCitation.latitude=45.46972" title="Search Plazi for locations around (long 11.54/lat 45.46972)">Lago di Fimon</a>; 45.46972° N, 11.54° E; 2009; Zallot leg.; EZ0892 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=11.57914&amp;materialsCitation.latitude=45.45607" title="Search Plazi for locations around (long 11.57914/lat 45.45607)">Lumignano</a>; 45.45607° N, 11.57914° E; 2010; Zallot leg.; EZ0893 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=11.05524&amp;materialsCitation.latitude=45.48662" title="Search Plazi for locations around (long 11.05524/lat 45.48662)">Mizzole</a>; 45.48662° N, 11.05524° E; Apr. 2010; Zallot leg.; EZ0878 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=11.45641&amp;materialsCitation.latitude=45.35477" title="Search Plazi for locations around (long 11.45641/lat 45.35477)">Monte delle Piume</a>; 45.35477° N, 11.45641° E; May 2010; G. Zallot leg.; EZ0889 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=11.68563&amp;materialsCitation.latitude=45.28735" title="Search Plazi for locations around (long 11.68563/lat 45.28735)">Mt Rusta</a>; 45.28735° N, 11.68563° E; 2002; Zallot leg.; EZ0894 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=11.29528&amp;materialsCitation.latitude=45.66528" title="Search Plazi for locations around (long 11.29528/lat 45.66528)">Novale-Valdagno</a>; 45.66528° N, 11.29528° E; 2000; Zallot leg.; EZ0884 • Perarolo dei Monti Berici, 45.47694° N, 11.50056° E; 1999, Zallot leg.; EZ0891 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=11.37417&amp;materialsCitation.latitude=45.63389" title="Search Plazi for locations around (long 11.37417/lat 45.63389)">Priabona</a>; 45.63389° N, 11.37417° E; 2000; Zallot leg.; EZ0888 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=11.49417&amp;materialsCitation.latitude=45.42833" title="Search Plazi for locations around (long 11.49417/lat 45.42833)">San Giovanni di Zovencedo</a>; 45.42833° N, 11.49417° E; 1999; Zallot leg.; EZ0890 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=11.1806&amp;materialsCitation.latitude=45.52866" title="Search Plazi for locations around (long 11.1806/lat 45.52866)">Tregnago-Finetti</a>; 45.52866° N, 11.1806° E; 2005; Zallot leg.; EZ0880 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=11.30079&amp;materialsCitation.latitude=45.64991" title="Search Plazi for locations around (long 11.30079/lat 45.64991)">Valdagno</a>; 45.64991° N, 11.30079° E; 2000; Zallot leg.; EZ0885.</p> <p>Description</p> <p>SHELL. The teleoconch whorls have whitish, very prominent, rather straight and widely spaced ribs on the upper whorls (see Fig. 11), but smooth specimens are found occasionally (see https://www.naturamediterraneo.com/forum/topic.asp?TOPIC_ID=43816).</p> <p>LIP. Moderately developed, and more or less flattened by the internal callus of the aperture. It gently curves backwards at the columellar side, starting rather far from the upper part of the body whorl. On lateral view, the body whorl, approaching the aperture, in most of the specimens barely enlarging (Fig. 4A: FF, Table 3).</p> <p>FEMALE GENITALIA. There is a long and relatively thin (as compared to the other species of Clessiniella; see Table 5) seminal receptacle with a curved apex.</p> <p>MALE GENITALIA. The penis is long and inflated. The penial spermiduct is slightly twisted and occupies only the central part of the penis. There is a clearly protruding but straight body spermiduct.</p> <p>Distribution</p> <p>The species inhabits a relatively small area at the foothills of the eastern Alps. It can also be found on the hills stretching down the Venetian flatland of the “pianura Veneta” (Colli Berici and Colli Euganei). It has been found sympatric with Cochlostoma (Cochlostoma) septemspirale and Cochlostoma (Eupomatias) philippianum in some localities of the province of Verona (Fernando Scarlassara, personal communication).</p> <p>Habitat preference</p> <p>The snails are found on rocky cliffs as well as on smaller boulders and dry stone walls in woody places.</p> <p>Remarks</p> <p>The range of this species is far from the almost continuous area in the east where the other Clessiniella species are known from. The species has been reported (and still is in the checklist of the Italian fauna – www.faunaitalia.it) as Cochlostoma villae (De Betta &amp; Martinati, 1855) but Bank (1988) found that Strobel (1851) described it as Pomatias maculatum var. villae Spinelli, four years before De Betta &amp; Martinati (1855). The description provided by Strobel (1851) is poor: “Varieta’ generalmente maggiore, coi giri della spira piu’ gonfi e colle coste trasversali piu’ regolari, e saglienti” = “ variety generally bigger, with more rounded whorls and more regular and strong ribs” and follows “ P. Villae Spinelli in esemplari raccolti a Mizzole presso Verona.” = “ P. villae Spinelli on specimens collected in Mizzole near Verona.”</p> <p>A few years later, De Betta &amp; Martinati (1855) gave almost the same poor description: “c. elongatior, costulis anfract. super. distinctioribus” = “taller shell, ribs in the upper whorls spaced” and follows “ Prov. Verona, Mizzole. Spin. ex spec. ” = “Verona Province, Mizzole, specimens provided by Spinelli.”</p> <p>It could be, as was common at the time, that the specimens used to describe the species by both Strobel (1851) and De Betta &amp; Martinati (1855) have had a tortuous path, after being collected by Villa in 1849 (as it appears from Strobel’s label), given to Spinelli later on, who gave the collector’s name to the species (as far as is known, however, Giovanni Battista Spinelli never published the specific epithet villae, probably using the name only in private correspondence). Spinelli eventually gave part of the material to Strobel and part to De Betta (the De Betta material is currently kept at the Museo di Storia Naturale di Verona, see Fig. 21).</p> </div>	http://treatment.plazi.org/id/038B8791315FFFC17BAB1477F36A252B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Zallot, Enrico;Mattia, Willy De;Fehér, Zoltán;Gittenberger, Edmund	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B87913157FFC67B9811F3F29020C5.text	038B87913157FFC67B9811F3F29020C5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cochlostoma (Clessiniella) anomphale Boeckel 1939	<div><p>Cochlostoma (Clessiniella) anomphale Boeckel, 1939</p> <p>Figs 1, 3A, 3A 1, 14A, 15A, 16A, 22–24</p> <p>Cochlostoma septemspirale anomphale Boeckel, 1939: 41–50 (“Karawanken, Trögern. Weg nach Vellach”).</p> <p>Cochlostoma septemspirale anomphale – Zilch 1958: 57.</p> <p>Cochlostoma (Turritus) anomphale – Gittenberger 1971: 115–118.</p> <p>Cochlostoma anomphale – Welter-Schultes 2012: 91.</p> <p>Diagnosis</p> <p>Shell quite densely ribbed, spots on the teleoconch generally clearly visible, body whorl inflated shortly before the lip. Visceral oviduct simplified with 1–2 superficial loops at most; seminal receptacle always club-shaped.</p> <p>Type material</p> <p>Paratypes AUSTRIA • 2 specs; Trögern, way to Vellach; SMF 42967/2 (Fig. 24).</p> <p>Other material (we analyzed fresh material from the type locality)</p> <p>AUSTRIA • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.59111&amp;materialsCitation.latitude=46.48583" title="Search Plazi for locations around (long 14.59111/lat 46.48583)">EisenKappel</a>; 46.48583° N, 14.59111° E; 2010, Mildner leg.; EZ0034 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.53639&amp;materialsCitation.latitude=46.42332" title="Search Plazi for locations around (long 14.53639/lat 46.42332)">Vellach</a>; 46.42332° N, 14.53639° E; Aug. 2010; De Mattia leg.; EZ0991.</p> <p>SLOVENIA • 210 road, N of Kokra; 46.34318° N, 14.48336° E; Aug. 2010; De Mattia leg.; EZ0993 • Gornji Grad; 46.2934° N, 14.82796° E; Aug. 2010; De Mattia leg.; EZ0996 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.59&amp;materialsCitation.latitude=46.32722" title="Search Plazi for locations around (long 14.59/lat 46.32722)">Kamniska Bistrika</a>; 46.32722° N, 14.59° E; 2010; Zallot leg.; EZ0033 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.4771&amp;materialsCitation.latitude=46.30356" title="Search Plazi for locations around (long 14.4771/lat 46.30356)">Kokra</a>; 46.30356° N, 14.4771° E; Aug. 2010; De Mattia leg.; EZ0995 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.65074&amp;materialsCitation.latitude=46.41959" title="Search Plazi for locations around (long 14.65074/lat 46.41959)">Logarska Dolina</a>; 46.41959° N, 14.65074° E; Aug. 2010; De Mattia leg.; EZ0994 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.66301&amp;materialsCitation.latitude=46.42334" title="Search Plazi for locations around (long 14.66301/lat 46.42334)">Logarska Dolina</a>; 46.42334° N, 14.66301° E; Aug. 2010; De Mattia leg.; EZ0992 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.69765&amp;materialsCitation.latitude=46.28787" title="Search Plazi for locations around (long 14.69765/lat 46.28787)">Podvolovliek- Stahovica</a>; 46.28787° N, 14.69765° E; Aug. 2005; Zallot leg.; EZ0036 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.7318&amp;materialsCitation.latitude=46.36596" title="Search Plazi for locations around (long 14.7318/lat 46.36596)">Savinjska Dolina</a>; 46.36596° N, 14.7318° E; 2010; De Mattia leg.; WdM1579 • Solčava; 46.42002° N, 14.19172° E; Jul. 2010; De Mattia leg.; WdM7010 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.60361&amp;materialsCitation.latitude=46.26528" title="Search Plazi for locations around (long 14.60361/lat 46.26528)">Stahovika</a>; 46.26528° N, 14.60361° E; 2005; Zallot leg.; EZ0035 • SV <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.69765&amp;materialsCitation.latitude=46.28787" title="Search Plazi for locations around (long 14.69765/lat 46.28787)">Anton</a>; 46.28787° N, 14.69765° E; 2010; Zallot leg.; EZ0032.</p> <p>Remarks</p> <p>The holotype and 23 paratypes should be in the Boeckel collection at the Naturkundemuseum Erfurt (Hartmann et al. 2018: 26). However, the shells could not be traced when this article was prepared (C. Unger in mail October 14, 2020).Type locality - Austria, Trögern, way to Vellach.</p> <p>Description</p> <p>SHELL. The outside of the body whorl is inflated shortly before the apertural lip (Fig. 4A: OB). The lip is poorly developed; it is almost vertical at the palatal side, and curved backwards and covering the umbilicus at the columellar side. The shells are more densely sculptured and the ribs are weaker than in the other species of Clessiniella. Most of the shells have spots.</p> <p>FEMALE GENITALIA. There is a spindle-like seminal receptacle. The visceral oviduct has only 1–2, superficial, loops.</p> <p>MALE GENITALIA. The swollen penis is longer than the body and has a central spermiduct. There is a protruding, straight, body spermiduct.</p> <p>Distribution</p> <p>Cochlostoma (Clessiniella) anomphale inhabits the Karawanken in south-eastern Austria and northern Slovenia.</p> <p>Habitat preference</p> <p>The animals can be found under and on stones and boulders in shady habitats in forests.</p> <p>Remarks</p> <p>This species was initially described as a subspecies of Cochlostoma (Cochlostoma) septemspirale. Gittenberger (1971) considered it a separate species because of the sympatry with Co. (Co.) semptemspirale heydenianum and the lack of conchologically intermediate specimens. This is an epitome of quite common Cochlostoma taxonomical issues: based on shell morphology it was initially described as subspecies of a taxon which is only distantly related.</p> </div>	http://treatment.plazi.org/id/038B87913157FFC67B9811F3F29020C5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Zallot, Enrico;Mattia, Willy De;Fehér, Zoltán;Gittenberger, Edmund	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B87913150FFFB7BB014C8F26D2664.text	038B87913150FFFB7BB014C8F26D2664.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cochlostoma (Clessiniella) stelucarum Zallot & Mattia & Fehér & Gittenberger 2021	<div><p>Cochlostoma (Clessiniella) stelucarum sp. nov.</p> <p>urn:lsid:zoobank.org:act: 4609B41C-3611-4F8A-9048-5F97EA0446E6</p> <p>Figs 1, 3E, 14B, 15B, 16B, 15–27</p> <p>Diagnosis</p> <p>Slender shell with 8½–10½ rather widely ribbed whorls. Spots always visible. Poorly developed lip with FF or AF external lobe. Visceral oviduct simplified and seminal receptacle sometimes oval.</p> <p>Etymology</p> <p>The epithet ‘ steluca ’ is an acronym noun formed from the initial syllables of the names of the wife and two daughters (Stefania, Ludovica and Camilla Zallot) of the senior author.</p> <p>Type material (Fig. 27)</p> <p>Holotype ITALY • ♀ (in ethanol); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.025&amp;materialsCitation.latitude=46.2297" title="Search Plazi for locations around (long 13.025/lat 46.2297)">Friuli Venezia Giulia</a>, Udine, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.025&amp;materialsCitation.latitude=46.2297" title="Search Plazi for locations around (long 13.025/lat 46.2297)">Cornino</a>; 46.2297° N, 13.0250° E; 180 m a.s.l.; 5 Jun. 2002; Zallot leg.; rocks surrounding the lake; RMNH.MOL.347617.</p> <p>Paratypes ITALY • 1 ♂ (in ethanol); same collection data as for holotype; RMNH.MOL.347618 • 1 ♀ (empty shell); same collection data as for holotype; GenBank voucher H3: KX120830; RMNH.MOL.347619.</p> <p>Other material</p> <p>ITALY • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.97106&amp;materialsCitation.latitude=46.23755" title="Search Plazi for locations around (long 12.97106/lat 46.23755)">Anduins-cliffs</a>; 46.23755° N, 12.97106° E; 1997; Zallot leg.; EZ0102 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.83333&amp;materialsCitation.latitude=46.26194" title="Search Plazi for locations around (long 12.83333/lat 46.26194)">Campone</a>; 46.26194° N, 12.83333° E; 2010; Zallot leg.; EZ0098 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.52095&amp;materialsCitation.latitude=46.094" title="Search Plazi for locations around (long 13.52095/lat 46.094)">Castelmonte</a>; 46.094° N, 13.52095° E; Oct. 2010; Margelli leg.; EZ1020 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.04111&amp;materialsCitation.latitude=46.25917" title="Search Plazi for locations around (long 13.04111/lat 46.25917)">Cjanet-Peonis</a>; 46.25917° N, 13.04111° E; 2002; Zallot leg.; EZ0105 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.025&amp;materialsCitation.latitude=46.22972" title="Search Plazi for locations around (long 13.025/lat 46.22972)">Cornino</a>; 46.22972° N, 13.025° E; 2002; Zallot leg.; EZ0104 • F. Piccola <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.80111&amp;materialsCitation.latitude=46.245" title="Search Plazi for locations around (long 12.80111/lat 46.245)">Mt Valinins</a>; 46.245° N, 12.80111° E; 2000; Zallot leg.; EZ0095 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.25556&amp;materialsCitation.latitude=46.23361" title="Search Plazi for locations around (long 13.25556/lat 46.23361)">Mt Bernadia</a>; 46.23361° N, 13.25556° E; 2005; Zallot leg.; EZ0107 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.88018&amp;materialsCitation.latitude=46.21226" title="Search Plazi for locations around (long 12.88018/lat 46.21226)">Mt Ciaurlec</a>; 46.21226° N, 12.88018° E; 2005; Zallot leg.; EZ0099 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.9907&amp;materialsCitation.latitude=46.24577" title="Search Plazi for locations around (long 12.9907/lat 46.24577)">Mt Prat</a>; 46.24577° N, 12.9907° E; 2002; Zallot leg.; EZ0103 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.8075&amp;materialsCitation.latitude=46.23417" title="Search Plazi for locations around (long 12.8075/lat 46.23417)">Mt Valinis</a>; 46.23417° N, 12.8075° E; 2002; Zallot leg.; EZ0096 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.94531&amp;materialsCitation.latitude=46.2337" title="Search Plazi for locations around (long 12.94531/lat 46.2337)">San Vito D’asio</a>; 46.2337° N, 12.94531° E; 1997; Zallot leg.; EZ0101 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.81917&amp;materialsCitation.latitude=46.27833" title="Search Plazi for locations around (long 12.81917/lat 46.27833)">Tramonti-Campone</a>; 46.27833° N, 12.81917° E; 2000; Zallot leg.; EZ0097 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.23111&amp;materialsCitation.latitude=46.23861" title="Search Plazi for locations around (long 13.23111/lat 46.23861)">Val di Torre-all’imbocco</a>; 46.23861° N, 13.23111° E; 1998; Zallot leg.; EZ0106 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=12.92972&amp;materialsCitation.latitude=46.30278" title="Search Plazi for locations around (long 12.92972/lat 46.30278)">Valle D’Arzino</a>; 46.30278° N, 12.92972° E; 2000; Zallot leg.; EZ0100.</p> <p>Description</p> <p>SHELL. The shell has prominent spots and strong and rounded ribs, which are regularly and widely spaced and coloured like the background. The ribs become less prominent on the body whorl. The body whorl slightly and gradually enlarges while approaching the aperture (Fig. 4A: FF, AF). In comparison with the other species of the subgenus, the whorls increase more slowly in size so that, given the same shell height, there is a higher number of whorls (Fig. 9).</p> <p>FEMALE GENITALIA. The visceral oviduct has only 1–2 superficial loops, what makes it one of the simplest in the genus Cochlostoma. The seminal receptacle is thin and may be oval, thus not always club-shaped as in the other species.</p> <p>MALE GENITALIA. The penis is longer than the body and inflated, although less so than in the other species of Clessiniella. The penial spermiduct is gently twisted. The body spermiduct is straight. There is a long groove, ending in a well-formed sperm pocket, delimited at the frontal side by an ascending sperm funnel.</p> <p>Distribution</p> <p>The species inhabits a restricted area in the eastern Southern Pre-Alps. Our sampling localities are located between Val Tramontina to the west and Val di Torre to the east (Friuli region, Italy).</p> <p>Habitat preference</p> <p>Like in most species of Cochlostoma, these are obligate rock-dwelling snails but, rather than on the surface of limestone cliffs, their main habitat is on smaller surfaces of stones and boulders which are normally found in large numbers at the foot of the cliffs.</p> <p>Remarks</p> <p>Cochlostoma (Clessiniella) stelucarum sp. nov. was introduced as Cochlostoma sp. by Zallot (2002: 98). At most of the localities, it lives syntopically with either Co. (Eupomatias) philippianum (Gredler, 1853) or Co. (Eupomatias) henricae (Strobel, 1851). Near the village of Anduins it has been found a few centimeters apart from both Co. (Eu.) henricae and Co. (Cochlostoma) septemspirale on stones and boulders. This is one of the rare cases where 3 species of Cochlostoma, belonging to different subgenera, live syntopically. The range of the species is situated in between the range of Co. (Cl.) villae in the west and that of the other species of Clessiniella in the east.</p> </div>	http://treatment.plazi.org/id/038B87913150FFFB7BB014C8F26D2664	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Zallot, Enrico;Mattia, Willy De;Fehér, Zoltán;Gittenberger, Edmund	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B8791316CFFFD7BF8116BF3D126E0.text	038B8791316CFFFD7BF8116BF3D126E0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cochlostoma (Clessiniella) tergestinum (Westerlund 1878)	<div><p>Cochlostoma (Clessiniella) tergestinum (Westerlund, 1878)</p> <p>Figs 1, 3B, 5, 14C, 15C, 16C, 28–30, 34 (bottom), 35 (bottom), 36 (top)</p> <p>Pomatias tergestinus Westerlund, 1878: 108–109 (“Triest”).</p> <p>Pomatias tortivus Westerlund, 1885: 124 (“Fiume, San Daniel u. Karstgebirge”, which should be Rijeka in Croatia and Štanjel in Slovenia).</p> <p>Pomatias tergestinus var. grahovana A.J. Wagner, 1897: 616, pl. 9 fig. 89 (“Grahovo bei Fiume”, a locality ca 10 km north of Rijeka in Croatia).</p> <p>Pomatias tergestinus var. tortiva – A.J. Wagner 1897: 616, pl. 9 fig. 88.</p> <p>Cochlostoma tergestinum − Welter-Schultes 2012: 102.</p> <p>Diagnosis</p> <p>Shell very variable, lip curved backwards more or less abruptly at the columellar side. Strong ribs on the teleoconch, in most of the populations widely spaced and, on the upper whorls, whitish and contrasting with the corneous, dark brown to light yellow, shell background. Visceral oviduct with convoluted loops path; body spermiduct straight.</p> <p>Type material</p> <p>Syntypes ITALY • “Triest” [Friuli Venezia Giulia, Trieste]; Westerlund leg.; SMF160677/1 • same label data as for preceding; SMF 160676/7 (Fig. 30).</p> <p>Other material</p> <p>AUSTRIA • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.6825&amp;materialsCitation.latitude=46.59222" title="Search Plazi for locations around (long 13.6825/lat 46.59222)">Dobratsch</a>; 46.59222° N, 13.6825° E; 2009; Mildner and Giovannelli leg.; EZ0799 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.75002&amp;materialsCitation.latitude=46.5888" title="Search Plazi for locations around (long 13.75002/lat 46.5888)">Kranzwand</a>; 46.588799° N, 13.75002° E; Jul. 2015; AL leg.; NHMW109000.</p> <p>CROATIA • Sveta Nedelja near Labin; 45.13741° N, 14.14488° E; 2000; De Mattia leg.; WdM6961 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.00625&amp;materialsCitation.latitude=45.45444" title="Search Plazi for locations around (long 14.00625/lat 45.45444)">Brest</a>, Cicaria; 45.45444° N, 14.00625° E; Apr. 2008; De Mattia leg.; WdM6715 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.64828&amp;materialsCitation.latitude=45.24147" title="Search Plazi for locations around (long 14.64828/lat 45.24147)">Drivenik</a> spring; 45.24147° N, 14.64828° E; Mar. 2009; De Mattia leg.; WdM6714 • from A6 to 501 down Bakar; 45.30543° N, 14.58579° E; Jul. 2009; De Mattia and Zallot leg.; EZ0938 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.004384&amp;materialsCitation.latitude=45.179493" title="Search Plazi for locations around (long 14.004384/lat 45.179493)">Jakačici</a>; 45.179494° N, 14.004384° E; 2009; De Mattia leg.; WdM6699 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.68499&amp;materialsCitation.latitude=45.43558" title="Search Plazi for locations around (long 13.68499/lat 45.43558)">Kremenje-Buje</a>; 45.43558° N, 13.68499° E; Jan. 2008; De Mattia leg.; WdM6400 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.07216&amp;materialsCitation.latitude=45.19369" title="Search Plazi for locations around (long 14.07216/lat 45.19369)">Kukurini</a>; 45.19369° N, 14.07216° E; Feb. 2009; De Mattia leg.; WdM6698 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.74039&amp;materialsCitation.latitude=45.12957" title="Search Plazi for locations around (long 13.74039/lat 45.12957)">Limski Kanal</a>; 45.12957° N, 13.74039° E; 2010; De Mattia leg.; WdM6693 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.74039&amp;materialsCitation.latitude=45.12957" title="Search Plazi for locations around (long 13.74039/lat 45.12957)">Limski Kanal</a>; 45.12957° N, 13.74039° E; Jun. 2010; De Mattia leg.; WdM6998 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.19715&amp;materialsCitation.latitude=45.27605" title="Search Plazi for locations around (long 14.19715/lat 45.27605)">Mala Ucka</a>; 45.27605° N, 14.19715° E; 2009; De Mattia leg.; WdM6768 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.58348&amp;materialsCitation.latitude=45.3037" title="Search Plazi for locations around (long 14.58348/lat 45.3037)">Meja</a>, Bakarac; 45.3037° N, 14.58348° E; 2009; De Mattia leg.; WdM6951 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.71384&amp;materialsCitation.latitude=45.442" title="Search Plazi for locations around (long 13.71384/lat 45.442)">Momjan</a>; 45.442° N, 13.71384° E; 2009; De Mattia leg.; WdM6854 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.20865&amp;materialsCitation.latitude=45.30636" title="Search Plazi for locations around (long 14.20865/lat 45.30636)">Mt Učka</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.20865&amp;materialsCitation.latitude=45.30636" title="Search Plazi for locations around (long 14.20865/lat 45.30636)">beech forest</a>; 45.30636° N, 14.20865° E; 2009; De Mattia leg.; WdM6794 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.20341&amp;materialsCitation.latitude=45.28967" title="Search Plazi for locations around (long 14.20341/lat 45.28967)">Mt Učka</a>, top; 45.28967° N, 14.20341° E; 2009; De Mattia leg.; WdM6794 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.93099&amp;materialsCitation.latitude=45.24134" title="Search Plazi for locations around (long 13.93099/lat 45.24134)">Pazin</a>; 45.24134° N, 13.93099° E; Jun. 1986; Drimmer leg.; HNHM97227 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.28905&amp;materialsCitation.latitude=45.43742" title="Search Plazi for locations around (long 14.28905/lat 45.43742)">Permani</a>; 45.43742° N, 14.28905° E; 2009; De Mattia leg.; WdM6708 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.08854&amp;materialsCitation.latitude=45.0872" title="Search Plazi for locations around (long 14.08854/lat 45.0872)">Raša</a>; 45.0872° N, 14.08854° E; 2010; De Mattia leg.; WdM6968 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.0538&amp;materialsCitation.latitude=45.15638" title="Search Plazi for locations around (long 14.0538/lat 45.15638)">Rasko Polje</a>; 45.15638° N, 14.0538° E; 2010; De Mattia leg. • EZ0813 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.0538&amp;materialsCitation.latitude=45.15638" title="Search Plazi for locations around (long 14.0538/lat 45.15638)">Rasko Polje</a>; 45.15638° N, 14.0538° E; 1900; De Mattia leg.; EZ1027 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.88873&amp;materialsCitation.latitude=45.13991" title="Search Plazi for locations around (long 13.88873/lat 45.13991)">Zminj</a>; 45.13991° N, 13.88873° E; 2000; De Mattia leg.; WdM6700.</p> <p>ITALY • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.87139&amp;materialsCitation.latitude=45.62" title="Search Plazi for locations around (long 13.87139/lat 45.62)">Val Rosandra</a>; 45.62° N, 13.87139° E; 2000; Zallot leg.; EZ0840 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.80594&amp;materialsCitation.latitude=45.71878" title="Search Plazi for locations around (long 13.80594/lat 45.71878)">Torri di Monrupino</a>; 45.71878° N, 13.80594° E; 1998; Zallot leg.; EZ0839.</p> <p>SLOVENIA • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.01433&amp;materialsCitation.latitude=45.82055" title="Search Plazi for locations around (long 14.01433/lat 45.82055)">Abram-Nanos Mt</a>; 45.82055° N, 14.01433° E; 2008; De Mattia leg.; WdM6731 • Brestovica pri <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.65016&amp;materialsCitation.latitude=45.80996" title="Search Plazi for locations around (long 13.65016/lat 45.80996)">Komnu</a>; 45.80996° N, 13.65016° E; 2009; De Mattia leg.; WdM6948 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.09732&amp;materialsCitation.latitude=45.67071" title="Search Plazi for locations around (long 14.09732/lat 45.67071)">Čepno</a>; 45.67071° N, 14.09732° E; Jun. 2009; De Mattia leg.; WdM6866 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.88633&amp;materialsCitation.latitude=45.55276" title="Search Plazi for locations around (long 13.88633/lat 45.55276)">Črnotiče-Črni Kal</a>; 45.55276° N, 13.88633° E; Mar. 2009; De Mattia leg.; EZ1028 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.06699&amp;materialsCitation.latitude=45.79813" title="Search Plazi for locations around (long 14.06699/lat 45.79813)">Dolina Nanos-Abran-Podkraj</a>; 45.79813° N, 14.06699° E; Apr. 2009; De Mattia leg.; WdM6733 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.95442&amp;materialsCitation.latitude=45.92429" title="Search Plazi for locations around (long 13.95442/lat 45.92429)">Farjev Vrh</a>; 45.92429° N, 13.95442° E; 2009; De Mattia leg.; WdM6740 • Golac, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.02969&amp;materialsCitation.latitude=45.51315" title="Search Plazi for locations around (long 14.02969/lat 45.51315)">Vela Vrata</a>; 45.51315° N, 14.02969° E; 2009; De Mattia leg.; WdM6762 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.70068&amp;materialsCitation.latitude=45.78485" title="Search Plazi for locations around (long 13.70068/lat 45.78485)">Gorjansko</a>; 45.78485° N, 13.70068° E; 2000; De Mattia leg.; WdM6950 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.1117&amp;materialsCitation.latitude=45.85832" title="Search Plazi for locations around (long 14.1117/lat 45.85832)">Hruščica</a>; 45.85832° N, 14.1117° E; 2000; De Mattia leg.; WdM2409 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.23917&amp;materialsCitation.latitude=45.56944" title="Search Plazi for locations around (long 14.23917/lat 45.56944)">Ilirska Bistrika</a>; 45.56944° N, 14.23917° E; 2009; Zallot leg.; EZ0821 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.95058&amp;materialsCitation.latitude=45.94614" title="Search Plazi for locations around (long 13.95058/lat 45.94614)">Kovk</a>; 45.94614° N, 13.95058° E; 2015 • De Mattia leg.; WdM6737 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.24579&amp;materialsCitation.latitude=45.82003" title="Search Plazi for locations around (long 14.24579/lat 45.82003)">Križna Jama</a>; 45.82003° N, 14.24579° E; Jun. 2009; Jochum leg.; HNHM97142 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.03994&amp;materialsCitation.latitude=45.58357" title="Search Plazi for locations around (long 14.03994/lat 45.58357)">Markovščina-Dimnice cave</a>; 45.58357° N, 14.03994° E; 2009; De Mattia leg.; WdM6765 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.80907&amp;materialsCitation.latitude=46.06528" title="Search Plazi for locations around (long 13.80907/lat 46.06528)">Mt Kobilica</a>; 46.06528° N, 13.80907° E; 2005; De Mattia leg.; WdM6849 • Mt Nanos, top, loc. <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.06699&amp;materialsCitation.latitude=45.79813" title="Search Plazi for locations around (long 14.06699/lat 45.79813)">Plesa</a>; 45.79813° N, 14.06699° E; Jun. 2005; De Mattia leg.; WdM0000 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.37049&amp;materialsCitation.latitude=45.56816" title="Search Plazi for locations around (long 14.37049/lat 45.56816)">Mt Snežnik</a> 1161; 45.56816° N, 14.37049° E; Jun. 2010; De Mattia leg.; WdM7012 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.41044&amp;materialsCitation.latitude=45.57179" title="Search Plazi for locations around (long 14.41044/lat 45.57179)">Mt Snežnik</a> 1390; 45.57179° N, 14.41044° E; Jun. 2010; De Mattia leg.; EZ0965 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.43839&amp;materialsCitation.latitude=45.57877" title="Search Plazi for locations around (long 14.43839/lat 45.57877)">Mt Snežnik</a> 1447; 45.57877° N, 14.43839° E; Jun. 2010; De Mattia leg.; EZ0964 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.43129&amp;materialsCitation.latitude=45.58195" title="Search Plazi for locations around (long 14.43129/lat 45.58195)">Mt Snežnik</a> 1450; 45.58195° N, 14.43129° E; Jun. 2010; De Mattia leg.; EZ0962 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.43117&amp;materialsCitation.latitude=45.58235" title="Search Plazi for locations around (long 14.43117/lat 45.58235)">Mt Snežnik car parking</a>; 45.58235° N, 14.43117° E; Jun. 2002; De Mattia leg.; WdM7006 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.03388&amp;materialsCitation.latitude=45.69898" title="Search Plazi for locations around (long 14.03388/lat 45.69898)">Mt Vremščica</a>; 45.69898° N, 14.03388° E; 2009; De Mattia leg.; WdM6758 • near Gorenje; 45.82976° N, 14.10755° E; 2009; Zallot leg.; EZ0817 • near Logatec; 45.88076° N, 14.27448° E; 2009; Zallot leg.; EZ0828 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.86168&amp;materialsCitation.latitude=45.57071" title="Search Plazi for locations around (long 13.86168/lat 45.57071)">Osp</a>, 45.57071° N, 13.86168° E; 2010; De Mattia leg.; WdM6756 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.8582&amp;materialsCitation.latitude=45.57119" title="Search Plazi for locations around (long 13.8582/lat 45.57119)">Ospo</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.8582&amp;materialsCitation.latitude=45.57119" title="Search Plazi for locations around (long 13.8582/lat 45.57119)">small cliff</a>; 45.57119° N, 13.8582° E; May 2010; De Mattia leg.; WdM6993 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.90918&amp;materialsCitation.latitude=45.92822" title="Search Plazi for locations around (long 13.90918/lat 45.92822)">Otlica</a>; 45.92822° N, 13.90918° E; 1986; De Mattia leg.; WdM6752 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.24708&amp;materialsCitation.latitude=45.82405" title="Search Plazi for locations around (long 14.24708/lat 45.82405)">Planina</a>; 45.82405° N, 14.24708° E; 2000; Zallot leg.; EZ0823 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.25056&amp;materialsCitation.latitude=45.82972" title="Search Plazi for locations around (long 14.25056/lat 45.82972)">Planina</a>; 45.82972° N, 14.25056° E; 2000; Zallot leg.; EZ0824 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.26056&amp;materialsCitation.latitude=45.83" title="Search Plazi for locations around (long 14.26056/lat 45.83)">Planina</a>, Postojna; 45.83° N, 14.26056° E; 2009; Zallot leg.; EZ0826 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.9449&amp;materialsCitation.latitude=45.53872" title="Search Plazi for locations around (long 13.9449/lat 45.53872)">Podgorje</a>; 45.53872° N, 13.9449° E; 2000; De Mattia leg.; WdM6735 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.9463&amp;materialsCitation.latitude=45.5364" title="Search Plazi for locations around (long 13.9463/lat 45.5364)">Podgorje</a>; 45.5364° N, 13.9463° E; 2009; De Mattia leg.; WdM6735 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.72709&amp;materialsCitation.latitude=46.05738" title="Search Plazi for locations around (long 13.72709/lat 46.05738)">Podlešče-Bainsizza</a>; 46.05738° N, 13.72709° E; May 2009; De Mattia leg.; WdM6814 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.94191&amp;materialsCitation.latitude=45.56587" title="Search Plazi for locations around (long 13.94191/lat 45.56587)">Prešnica</a>; 45.56587° N, 13.94191° E; Mar 2009; De Mattia leg.; WdM6734 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.77584&amp;materialsCitation.latitude=46.04133" title="Search Plazi for locations around (long 13.77584/lat 46.04133)">Pri Cerkvi- Bainsizza</a>; 46.04133° N, 13.77584° E; 2009; De Mattia leg.; WdM6834 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.28948&amp;materialsCitation.latitude=45.79467" title="Search Plazi for locations around (long 14.28948/lat 45.79467)">Rakow</a> Skocjan; 45.79467° N, 14.28948° E; 2010; Zallot leg.; EZ0838 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.99356&amp;materialsCitation.latitude=45.66532" title="Search Plazi for locations around (long 13.99356/lat 45.66532)">San Canzian</a> (Škocjan); 45.66532° N, 13.99356° E; 2009; De Mattia leg.; WdM6754 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.07856&amp;materialsCitation.latitude=45.71421" title="Search Plazi for locations around (long 14.07856/lat 45.71421)">Senožeče</a>; 45.71421° N, 14.07856° E; 2009; De Mattia leg.; WdM6865 • 10 km S of Vrhnika; 45.88549° N, 14.26091° E; 2000; Zallot leg.; EZ0827 • Vrhnika, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.29268&amp;materialsCitation.latitude=45.95591" title="Search Plazi for locations around (long 14.29268/lat 45.95591)">Močilnik</a>; 45.95591° N, 14.29268° E; 2009; Zallot leg.; EZ0829.</p> <p>Remarks</p> <p>The original indication of the type locality is rather vague, since only a city in the Friuli-Venezia Giulia region, in the easternmost part of Italy is indicated. The closest to Trieste where we found the species is at “Val Rosandra”, 8 km north-east of the city. The fresh specimens collected there have shell features similar to those of the syntypes. We therefore consider them as topotypical.</p> <p>Description</p> <p>SHELL. The size of the shells is extremely variable (height 6.1–9.6 mm in the females) as well as their general morphology (see Fig. 28). Several populations are spotless whereas others have clear spots on the teleoconch. Depending on the population, the lip can be either poorly or even hugely developed. In lateral view, the body whorl, while approaching the aperture, may widen in various ways, depending on the population (Fig. 4A: FF, AF, II). The H/W ratio is very variable and the slenderness of the shells in different populations varies accordingly.</p> <p>FEMALE GENITALIA. There is a long, club-shaped seminal receptacle. In some cases there is a change of colour at its base, presumably indicating the starting point of the rather long distal oviduct. As shown in Table 5, there is a considerable variability in the part of the seminal receptacle that is curved down and in the number of loops.</p> <p>MALE GENITALIA. The inflated penis is longer than the body. The penial spermiduct is confined to the central part of the penis and gently twisted. The body spermiduct is straight and well defined.</p> <p>Distribution</p> <p>This species inhabits the eastern side of the province of Trieste in Italy, the southwestern part of Slovenia and the Istrian peninsula. It has also been found in several Croatian localities in the surroundings of Bakar and, quite far from its main range, on the Villacher Alps, in Austria (see Mildner 1997). The species has been erroneously reported for Albania (see Fehér &amp; Erőss 2009).</p> <p>Habitat preference</p> <p>It is an obligate rock-dwelling species. Co. (Cl.) tergestinum is found in a variety of habitats, from open, exposed limestone cliffs to shady spots in deciduous and evergreen forests.</p> <p>Remarks</p> <p>According to Zilch (1958) and Alzona (1971) there are 3 subspecies of Co. (Cl.) tergestinum, viz. the nominate subspecies Co. (Clessiniella) tergestinum tergestinum (Westerlund, 1878), Co. (Clessiniella) tergestinum grahovanum (A.J. Wagner, 1897) and Co. (Clessiniella) tergestinum tortivum (Westerlund, 1885).</p> <p>For the time being, we here synonymize these nominal taxa because the conchological features which supposedly characterize them, like measures, density and prominence of the ribs, and the structure of the lip, are population features without an obvious geographical pattern. There is no significant correlation between latitude and longitude of the locality of the sample and the character states that are mentioned in the original description of the so-called subspecies. Moreover, in the surroundings of Trieste, the type locality of the nominate subspecies, only populations with some ‘ tortivum ’ shell features, like the thin lip, can be found (De Mattia &amp; Prodan 2006).</p> <p>Some of the populations have a peculiar shell morphology (see for instance Fig. 28E–F); their taxonomical status should be investigated more thoroughly especially since the available molecular H3 analysis indicates that population “EZ0813” from Rasko Polje, corresponding to the specimen in Fig. 28E, stands apart from the others.</p></div> 	http://treatment.plazi.org/id/038B8791316CFFFD7BF8116BF3D126E0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Zallot, Enrico;Mattia, Willy De;Fehér, Zoltán;Gittenberger, Edmund	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B8791316BFFF37B9A12ABF5F92792.text	038B8791316BFFF37B9A12ABF5F92792.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cochlostoma (Clessiniella) waldemari (A. J. Wagner 1897)	<div><p>Cochlostoma (Clessiniella) waldemari (Wagner, 1897)</p> <p>Figs 1, 3D, 14E, 15E, 16E, 31–33, 34 (top), 35 (top), 36 (bottom)</p> <p>Pomatias (Auritus) waldemari A.J. Wagner, 1897: 616, pl. 9, 90 (“Ogulin”).</p> <p>Cochlostoma waldemari – Welter-Schultes 2012: 103.</p> <p>Diagnosis</p> <p>Spotless shell. Moderately strong, more or less widely spaced ribs. Columellar lobe gently curved backwards. External lobe of type “II” in almost all the populations. The body spermiduct is huge and twisted instead of straight as in all the other species of Clessiniella.</p> <p>Type material</p> <p>Syntypes CROATIA • ♀; Ogulin; A.J. Wagner leg.; MIZPAS-W-6979a (Fig. 33) • 2 ♀♀, 3 ♂♂; same collection data as for preceding; MIZPAS-W-6979/5.</p> <p>Other material</p> <p>CROATIA • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.20837&amp;materialsCitation.latitude=45.07534" title="Search Plazi for locations around (long 15.20837/lat 45.07534)">Kapela pass</a>, Ogulin; 45.07534° N, 15.20837° E; 2009; De Mattia leg.; EZ0909 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.14044&amp;materialsCitation.latitude=45.25904" title="Search Plazi for locations around (long 15.14044/lat 45.25904)">Klek Mt</a>, close to the top; 45.25904° N, 15.14044° E; 2009; Zallot leg.; EZ0904 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.14561&amp;materialsCitation.latitude=45.25438" title="Search Plazi for locations around (long 15.14561/lat 45.25438)">Klek Mt</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.14561&amp;materialsCitation.latitude=45.25438" title="Search Plazi for locations around (long 15.14561/lat 45.25438)">cliffs halfway</a>; 45.25438° N, 15.14561° E; 1996; Zallot leg.; EZ0905 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.6188&amp;materialsCitation.latitude=45.43095" title="Search Plazi for locations around (long 14.6188/lat 45.43095)">Mt Risnjak</a>; 45.43095° N, 14.6188° E; 1400 m a.s.l.; 2009; De Mattia leg.; EZ0845 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.61748&amp;materialsCitation.latitude=45.39072" title="Search Plazi for locations around (long 14.61748/lat 45.39072)">Mt Risnjak</a>; 45.39072° N, 14.61748° E; 1000 m a.s.l.; 2010; Zallot leg.; EZ0907 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.09534&amp;materialsCitation.latitude=45.3607" title="Search Plazi for locations around (long 14.09534/lat 45.3607)">Ročko Polje</a>; 45.3607° N, 14.09534° E; 2009; De Mattia leg.; WdM6697 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.33827&amp;materialsCitation.latitude=45.23402" title="Search Plazi for locations around (long 15.33827/lat 45.23402)">Rudnica</a>; 45.23402° N, 15.33827° E; 2009; De Mattia leg.; EZ0906 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.71065&amp;materialsCitation.latitude=45.31777" title="Search Plazi for locations around (long 14.71065/lat 45.31777)">Spilja Vrelo</a>; 45.31777° N, 14.71065° E; 2009; Zallot leg.; EZ0899 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.14041&amp;materialsCitation.latitude=45.29114" title="Search Plazi for locations around (long 15.14041/lat 45.29114)">Vitunj-Ogulin</a>; 45.29114° N, 15.14041° E; 2005; Zallot leg.; EZ0903.</p> <p>SLOVENIA • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.56752&amp;materialsCitation.latitude=45.76192" title="Search Plazi for locations around (long 14.56752/lat 45.76192)">Benete</a>, 45.76192°N, 14.56752°E;2009; De Mattia leg.; WdM6805 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.43533&amp;materialsCitation.latitude=45.95607" title="Search Plazi for locations around (long 14.43533/lat 45.95607)">Bresovica</a>; 45.95607°N, 14.43533° E; 2006; De Mattia leg.; EZ0897 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.0437&amp;materialsCitation.latitude=45.55164" title="Search Plazi for locations around (long 15.0437/lat 45.55164)">Bresovica pri Predgradu</a>; 45.55164° N, 15.0437° E; 2009; De Mattia leg.; WdM6612 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.0277&amp;materialsCitation.latitude=45.9827" title="Search Plazi for locations around (long 14.0277/lat 45.9827)">Divje Jezero</a>; 45.9827° N, 14.0277° E; 2009; De Mattia leg.; WdM6840 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.81189&amp;materialsCitation.latitude=46.06445" title="Search Plazi for locations around (long 13.81189/lat 46.06445)">Dol-E of Čepovan</a>; 46.06445° N, 13.81189° E; 2009; De Mattia leg.; WdM6837 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.05346&amp;materialsCitation.latitude=45.5115" title="Search Plazi for locations around (long 15.05346/lat 45.5115)">Jelenja Vas</a>; 45.5115° N, 15.05346° E; 2009; De Mattia leg.; WdM6804 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.09528&amp;materialsCitation.latitude=45.53417" title="Search Plazi for locations around (long 15.09528/lat 45.53417)">Krempa Kolpa</a>; 45.53417° N, 15.09528° E; 1996; Mildner leg.; EZ0902 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.09389&amp;materialsCitation.latitude=45.53444" title="Search Plazi for locations around (long 15.09389/lat 45.53444)">Marija Ceplje Kolpa</a>; 45.53444° N, 15.09389° E; 2006; Mildner leg.; EZ0901 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.321&amp;materialsCitation.latitude=45.56913" title="Search Plazi for locations around (long 14.321/lat 45.56913)">Mt Snežnik</a> 1150; 45.56913° N, 14.321° E; Jun. 2010; De Mattia leg.; WdM7014 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.44209&amp;materialsCitation.latitude=45.58845" title="Search Plazi for locations around (long 14.44209/lat 45.58845)">Mt Snežnik</a> 1620; 45.58845° N, 14.44209° E; Jun. 2010; De Mattia leg.; EZ0963 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.43117&amp;materialsCitation.latitude=45.58235" title="Search Plazi for locations around (long 14.43117/lat 45.58235)">Mt Snežnik car parking</a>; 45.58235° N, 14.43117° E; Jun. 2010; De Mattia leg.; WdM7006 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.81275&amp;materialsCitation.latitude=46.06387" title="Search Plazi for locations around (long 13.81275/lat 46.06387)">Polog-E of Čepovan</a>; 46.06387° N, 13.81275° E; 2009; De Mattia leg.; WdM6842 • SW Pescena Glava; 46.05156° N, 13.73844° E; 843 m a.s.l.; 2009; De Mattia leg.; WdM6829 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.76859&amp;materialsCitation.latitude=45.81809" title="Search Plazi for locations around (long 14.76859/lat 45.81809)">Tisovec</a>; 45.81809° N, 14.76859° E; 2009; De Mattia leg.; WdM6749 • same collection data as for preceding; WdM6750 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.76417&amp;materialsCitation.latitude=46.44278" title="Search Plazi for locations around (long 13.76417/lat 46.44278)">Vrsic Pass</a>; 46.44278° N, 13.76417° E; 1985; Zallot leg.; EZ0896 • <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=14.83417&amp;materialsCitation.latitude=45.86028" title="Search Plazi for locations around (long 14.83417/lat 45.86028)">Zagradec- Polyane</a>; 45.86028° N, 14.83417° E; Bodon leg.; EZ0900.</p> <p>Description</p> <p>SHELL. The shell is more or less light corneous brown and spotless. The size of the aperture varies considerably among the populations; shells with a large aperture and a correspondingly large lip occur next to shells with a poorly developed lip and a relatively small aperture. The lip is gently curved backwards on the columellar side.</p> <p>FEMALE GENITALIA. The female genitalia are hardly distinguishable from those of Co. (Cl.) tergestinum if not for the more relaxed loops.</p> <p>MALE GENITALIA. There is a long, inflated penis. The penial spermiduct is slightly twisted and occupies only the central part of the penis. The body spermiduct is twisted and conspicuously protruding (see Figs 34–35).</p> <p>Distribution</p> <p>The species inhabits the north-western and south-eastern part of Slovenia and an adjacent area in Croatia. It is reported from Austria for the Karawanken and the Gailtaler Alpen (Edlinger &amp; Mildner 1979). Its range largely surrounds that of Co. (Cl.) tergestinum (see Fig. 1).</p> <p>Habitat preference</p> <p>The snails are found on rocky cliffs as well as on boulders in woody areas.</p> <p>Remarks</p> <p>To assign a given population to Co. (Cl.) waldemari rather than to Co. (Cl.) tergestinum on the basis of the morphology of the shell and the female genitalia alone could be challenging if not impossible (see Figs 34–35). Only the structure of the male genitalia allows a convincing identification of the species. The H3 analysis (Fig. 7) confirms that these sibling species belong to separate clades within Clessiniella.</p> <p>Boeckel (1939) considered Co. (Cl.) waldemari a subspecies of Co. (Cl.) tergestinum. This view was contradicted by Bole (1994). In some areas, like on Mount Snežnik in southern Slovenia, the two species live not far from each other and here they can be distinguished also conchologically while differing in the presence or absence of the spots on the teleoconch (with Co. (Cl.) waldemari having a spotless shell) and in the shape of the body whorl approaching the aperture, in lateral view (Fig. 36).</p> </div>	http://treatment.plazi.org/id/038B8791316BFFF37B9A12ABF5F92792	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Zallot, Enrico;Mattia, Willy De;Fehér, Zoltán;Gittenberger, Edmund	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
