taxonID	type	description	language	source
038B87913142FFCA7B7E1463F42627B8.taxon	type_taxon	Type species Pomatias maculatum var. villae Strobel, 1851, by original designation.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B87913142FFCA7B7E1463F42627B8.taxon	discussion	Remarks The classification follows MolluscaBase (2021) except for the subgeneric name. Contrary to ICZN Art. 10.4, Neglecta Wagner, 1897 is used in MolluscaBase as a senior synonym of Clessiniella. Wagner (1897: 6 [570]) introduced Neglecta as “ Formenkreis Neglecta ”. It cannot be denied that the German term Formenkreis is “ .. a term such as “ superspecies ””. As a consequence, according to the ICZN ruling, Neglecta “ .. is not deemed to be a genus-group name ” (ICZN Art. 10.4, see also Zallot et al. 2015). Strobel (1851) and De Betta & Martinati (1855) named this entity Pomatias maculatum var. Villae – but referring to Pomatias maculatus Draparnaud, 1805 (= Co. (Co.) septemspirale).	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B87913142FFCA7B7E1463F42627B8.taxon	diagnosis	Diagnosis of the subgenus SHELL. Ribbed teleoconch with straight ribs, variable in strength and shape. The columellar lobe is curved backwards, covering the umbilicus. The teleoconch spots (if present) form 2 rows on the whorls (often, a 3 rd row of spots is visible on the body whorl): the upper one close to the suture with rectangular spots abapically elongated; the lower one roughly ⅓ up on the whorls (Fig. 12). In some populations the 2 rows of spots merge and there is a continuous vertical darker band on the whorls. The protoconch is dull and robust with fine riblets starting after 1.2 whorls or more from the apex; it is transparent and fragile or the ribbing starts close to the apex in other subgenera.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B87913142FFCA7B7E1463F42627B8.taxon	description	FEMALE GENITALIA. There is an apical connection of the pedunculus to the bursa copulatrix, a long seminal receptacle (normally club-shaped), which runs parallel to the pedunculus. The junction of the uterus gland is situated close to the connection between the pedunculus and the distal oviduct. This combination of characters in the female genitalia is most diagnostic to distinguish Clessiniella from the other subgenera. MALE GENITALIA. Apart from its apical part, the penial spermiduct is tortuous. The body spermiduct is protruding (often almost detached from the body) more than in the other subgenera and either straight or more or less tortuous. The posterior side of the penis is wrinkled.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B87913142FFCA7B7E1463F42627B8.taxon	discussion	Identification key for the species	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B8791315FFFC17BAB1477F36A252B.taxon	description	Figs 1, 3 C, 14 D, 15 D, 16 D, 17 – 21	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B8791315FFFC17BAB1477F36A252B.taxon	diagnosis	Diagnosis The last whorl, approaching the aperture, hardly enlarges (Fig. 4 A: FF). Very prominent and widely spaced teleoconch ribs, barely inclined in respect to the vertical axis. Spots almost always visible. The protoconch is relatively small. The apertural lip poorly developed. Visceral oviduct with convoluted loops; thin seminal receptacle. Straight body spermiduct.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B8791315FFFC17BAB1477F36A252B.taxon	materials_examined	Type material Syntypes ITALY • 1 ♀; “ Pomatias villae Spinel. Mizzole. Villa 49 ” [label of the original material] [Veneto, Verona, Mizzole] [thus, it could be interpreted as collected in 1849 by Villa]; PU 1610 - C 1 (Fig. 19) • 2 ♂♂; same label data as for preceding; PU 1610 - C 2, PU 1610 - C 3 (Fig. 20). Other material (fresh specimens from the type locality have also been analyzed) ITALY • Altissimo; 45.61511 ° N, 11.25225 ° E; 2009; Zallot leg.; EZ 0883 • Arqua’ Petrarca, surroundings; 45.26194 ° N, 11.69623 ° E; 1999; Zallot leg.; EZ 0895 • Bolca; 45.59389 ° N, 11.20972 ° E; 2010; Zallot leg.; EZ 0882 • Ca’Gottolo; 45.58921 ° N, 10.90787 ° E; Jun. 2010; Scarlassara leg.; EZ 1015 • Cornedo Vicentino-Cereda; 45.60667 ° N, 11.36833 ° E; 1999; Zallot leg.; EZ 0887 • Covoli di Velo; 45.60889 ° N, 11.12083 ° E; 2010; Zallot leg.; EZ 0879 • Ferrazza; 45.62137 ° N, 11.19538 ° E; Apr. 2010; Zallot leg.; EZ 0881 • Fumane-Molina; 45.56942 ° N, 10.89856 ° E; Jun. 2010; Scarlassara leg.; EZ 0936 • Lago di Fimon; 45.46972 ° N, 11.54 ° E; 2009; Zallot leg.; EZ 0892 • Lumignano; 45.45607 ° N, 11.57914 ° E; 2010; Zallot leg.; EZ 0893 • Mizzole; 45.48662 ° N, 11.05524 ° E; Apr. 2010; Zallot leg.; EZ 0878 • Monte delle Piume; 45.35477 ° N, 11.45641 ° E; May 2010; G. Zallot leg.; EZ 0889 • Mt Rusta; 45.28735 ° N, 11.68563 ° E; 2002; Zallot leg.; EZ 0894 • Novale-Valdagno; 45.66528 ° N, 11.29528 ° E; 2000; Zallot leg.; EZ 0884 • Perarolo dei Monti Berici, 45.47694 ° N, 11.50056 ° E; 1999, Zallot leg.; EZ 0891 • Priabona; 45.63389 ° N, 11.37417 ° E; 2000; Zallot leg.; EZ 0888 • San Giovanni di Zovencedo; 45.42833 ° N, 11.49417 ° E; 1999; Zallot leg.; EZ 0890 • Tregnago-Finetti; 45.52866 ° N, 11.1806 ° E; 2005; Zallot leg.; EZ 0880 • Valdagno; 45.64991 ° N, 11.30079 ° E; 2000; Zallot leg.; EZ 0885.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B8791315FFFC17BAB1477F36A252B.taxon	description	Description SHELL. The teleoconch whorls have whitish, very prominent, rather straight and widely spaced ribs on the upper whorls (see Fig. 11), but smooth specimens are found occasionally (see https: // www. naturamediterraneo. com / forum / topic. asp? TOPIC _ ID = 43816). LIP. Moderately developed, and more or less flattened by the internal callus of the aperture. It gently curves backwards at the columellar side, starting rather far from the upper part of the body whorl. On lateral view, the body whorl, approaching the aperture, in most of the specimens barely enlarging (Fig. 4 A: FF, Table 3). FEMALE GENITALIA. There is a long and relatively thin (as compared to the other species of Clessiniella; see Table 5) seminal receptacle with a curved apex. MALE GENITALIA. The penis is long and inflated. The penial spermiduct is slightly twisted and occupies only the central part of the penis. There is a clearly protruding but straight body spermiduct.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B8791315FFFC17BAB1477F36A252B.taxon	distribution	Distribution The species inhabits a relatively small area at the foothills of the eastern Alps. It can also be found on the hills stretching down the Venetian flatland of the “ pianura Veneta ” (Colli Berici and Colli Euganei). It has been found sympatric with Cochlostoma (Cochlostoma) septemspirale and Cochlostoma (Eupomatias) philippianum in some localities of the province of Verona (Fernando Scarlassara, personal communication).	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B8791315FFFC17BAB1477F36A252B.taxon	description	Habitat preference The snails are found on rocky cliffs as well as on smaller boulders and dry stone walls in woody places.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B8791315FFFC17BAB1477F36A252B.taxon	discussion	Remarks The range of this species is far from the almost continuous area in the east where the other Clessiniella species are known from. The species has been reported (and still is in the checklist of the Italian fauna – www. faunaitalia. it) as Cochlostoma villae (De Betta & Martinati, 1855) but Bank (1988) found that Strobel (1851) described it as Pomatias maculatum var. villae Spinelli, four years before De Betta & Martinati (1855). The description provided by Strobel (1851) is poor: “ Varieta’ generalmente maggiore, coi giri della spira piu’ gonfi e colle coste trasversali piu’ regolari, e saglienti ” = “ variety generally bigger, with more rounded whorls and more regular and strong ribs ” and follows “ P. Villae Spinelli in esemplari raccolti a Mizzole presso Verona. ” = “ P. villae Spinelli on specimens collected in Mizzole near Verona. ” A few years later, De Betta & Martinati (1855) gave almost the same poor description: “ c. elongatior, costulis anfract. super. distinctioribus ” = “ taller shell, ribs in the upper whorls spaced ” and follows “ Prov. Verona, Mizzole. Spin. ex spec. ” = “ Verona Province, Mizzole, specimens provided by Spinelli. ” It could be, as was common at the time, that the specimens used to describe the species by both Strobel (1851) and De Betta & Martinati (1855) have had a tortuous path, after being collected by Villa in 1849 (as it appears from Strobel’s label), given to Spinelli later on, who gave the collector’s name to the species (as far as is known, however, Giovanni Battista Spinelli never published the specific epithet villae, probably using the name only in private correspondence). Spinelli eventually gave part of the material to Strobel and part to De Betta (the De Betta material is currently kept at the Museo di Storia Naturale di Verona, see Fig. 21).	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B87913157FFC67B9811F3F29020C5.taxon	description	Figs 1, 3 A, 3 A 1, 14 A, 15 A, 16 A, 22 – 24	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B87913157FFC67B9811F3F29020C5.taxon	diagnosis	Diagnosis Shell quite densely ribbed, spots on the teleoconch generally clearly visible, body whorl inflated shortly before the lip. Visceral oviduct simplified with 1 – 2 superficial loops at most; seminal receptacle always club-shaped.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B87913157FFC67B9811F3F29020C5.taxon	materials_examined	Type material Paratypes AUSTRIA • 2 specs; Trögern, way to Vellach; SMF 42967 / 2 (Fig. 24). Other material (we analyzed fresh material from the type locality) AUSTRIA • EisenKappel; 46.48583 ° N, 14.59111 ° E; 2010, Mildner leg.; EZ 0034 • Vellach; 46.42332 ° N, 14.53639 ° E; Aug. 2010; De Mattia leg.; EZ 0991. SLOVENIA • 210 road, N of Kokra; 46.34318 ° N, 14.48336 ° E; Aug. 2010; De Mattia leg.; EZ 0993 • Gornji Grad; 46.2934 ° N, 14.82796 ° E; Aug. 2010; De Mattia leg.; EZ 0996 • Kamniska Bistrika; 46.32722 ° N, 14.59 ° E; 2010; Zallot leg.; EZ 0033 • Kokra; 46.30356 ° N, 14.4771 ° E; Aug. 2010; De Mattia leg.; EZ 0995 • Logarska Dolina; 46.41959 ° N, 14.65074 ° E; Aug. 2010; De Mattia leg.; EZ 0994 • Logarska Dolina; 46.42334 ° N, 14.66301 ° E; Aug. 2010; De Mattia leg.; EZ 0992 • Podvolovliek- Stahovica; 46.28787 ° N, 14.69765 ° E; Aug. 2005; Zallot leg.; EZ 0036 • Savinjska Dolina; 46.36596 ° N, 14.7318 ° E; 2010; De Mattia leg.; WdM 1579 • Solčava; 46.42002 ° N, 14.19172 ° E; Jul. 2010; De Mattia leg.; WdM 7010 • Stahovika; 46.26528 ° N, 14.60361 ° E; 2005; Zallot leg.; EZ 0035 • SV Anton; 46.28787 ° N, 14.69765 ° E; 2010; Zallot leg.; EZ 0032.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B87913157FFC67B9811F3F29020C5.taxon	discussion	Remarks The holotype and 23 paratypes should be in the Boeckel collection at the Naturkundemuseum Erfurt (Hartmann et al. 2018: 26). However, the shells could not be traced when this article was prepared (C. Unger in mail October 14, 2020). Type locality - Austria, Trögern, way to Vellach.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B87913157FFC67B9811F3F29020C5.taxon	description	Description SHELL. The outside of the body whorl is inflated shortly before the apertural lip (Fig. 4 A: OB). The lip is poorly developed; it is almost vertical at the palatal side, and curved backwards and covering the umbilicus at the columellar side. The shells are more densely sculptured and the ribs are weaker than in the other species of Clessiniella. Most of the shells have spots. FEMALE GENITALIA. There is a spindle-like seminal receptacle. The visceral oviduct has only 1 – 2, superficial, loops. MALE GENITALIA. The swollen penis is longer than the body and has a central spermiduct. There is a protruding, straight, body spermiduct.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B87913157FFC67B9811F3F29020C5.taxon	distribution	Distribution Cochlostoma (Clessiniella) anomphale inhabits the Karawanken in south-eastern Austria and northern Slovenia.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B87913157FFC67B9811F3F29020C5.taxon	description	Habitat preference The animals can be found under and on stones and boulders in shady habitats in forests.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B87913157FFC67B9811F3F29020C5.taxon	discussion	Remarks This species was initially described as a subspecies of Cochlostoma (Cochlostoma) septemspirale. Gittenberger (1971) considered it a separate species because of the sympatry with Co. (Co.) semptemspirale heydenianum and the lack of conchologically intermediate specimens. This is an epitome of quite common Cochlostoma taxonomical issues: based on shell morphology it was initially described as subspecies of a taxon which is only distantly related.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B87913150FFFB7BB014C8F26D2664.taxon	description	urn: lsid: zoobank. org: act: 4609 B 41 C- 3611 - 4 F 8 A- 9048 - 5 F 97 EA 0446 E 6 Figs 1, 3 E, 14 B, 15 B, 16 B, 15 – 27	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B87913150FFFB7BB014C8F26D2664.taxon	diagnosis	Diagnosis Slender shell with 8 ½ – 10 ½ rather widely ribbed whorls. Spots always visible. Poorly developed lip with FF or AF external lobe. Visceral oviduct simplified and seminal receptacle sometimes oval.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B87913150FFFB7BB014C8F26D2664.taxon	etymology	Etymology The epithet ‘ steluca ’ is an acronym noun formed from the initial syllables of the names of the wife and two daughters (Stefania, Ludovica and Camilla Zallot) of the senior author.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B87913150FFFB7BB014C8F26D2664.taxon	materials_examined	Type material (Fig. 27) Holotype ITALY • ♀ (in ethanol); Friuli Venezia Giulia, Udine, Cornino; 46.2297 ° N, 13.0250 ° E; 180 m a. s. l.; 5 Jun. 2002; Zallot leg.; rocks surrounding the lake; RMNH. MOL. 347617. Paratypes ITALY • 1 ♂ (in ethanol); same collection data as for holotype; RMNH. MOL. 347618 • 1 ♀ (empty shell); same collection data as for holotype; GenBank voucher H 3: KX 120830; RMNH. MOL. 347619. Other material ITALY • Anduins-cliffs; 46.23755 ° N, 12.97106 ° E; 1997; Zallot leg.; EZ 0102 • Campone; 46.26194 ° N, 12.83333 ° E; 2010; Zallot leg.; EZ 0098 • Castelmonte; 46.094 ° N, 13.52095 ° E; Oct. 2010; Margelli leg.; EZ 1020 • Cjanet-Peonis; 46.25917 ° N, 13.04111 ° E; 2002; Zallot leg.; EZ 0105 • Cornino; 46.22972 ° N, 13.025 ° E; 2002; Zallot leg.; EZ 0104 • F. Piccola Mt Valinins; 46.245 ° N, 12.80111 ° E; 2000; Zallot leg.; EZ 0095 • Mt Bernadia; 46.23361 ° N, 13.25556 ° E; 2005; Zallot leg.; EZ 0107 • Mt Ciaurlec; 46.21226 ° N, 12.88018 ° E; 2005; Zallot leg.; EZ 0099 • Mt Prat; 46.24577 ° N, 12.9907 ° E; 2002; Zallot leg.; EZ 0103 • Mt Valinis; 46.23417 ° N, 12.8075 ° E; 2002; Zallot leg.; EZ 0096 • San Vito D’asio; 46.2337 ° N, 12.94531 ° E; 1997; Zallot leg.; EZ 0101 • Tramonti-Campone; 46.27833 ° N, 12.81917 ° E; 2000; Zallot leg.; EZ 0097 • Val di Torre-all’imbocco; 46.23861 ° N, 13.23111 ° E; 1998; Zallot leg.; EZ 0106 • Valle D’Arzino; 46.30278 ° N, 12.92972 ° E; 2000; Zallot leg.; EZ 0100.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B87913150FFFB7BB014C8F26D2664.taxon	description	Description SHELL. The shell has prominent spots and strong and rounded ribs, which are regularly and widely spaced and coloured like the background. The ribs become less prominent on the body whorl. The body whorl slightly and gradually enlarges while approaching the aperture (Fig. 4 A: FF, AF). In comparison with the other species of the subgenus, the whorls increase more slowly in size so that, given the same shell height, there is a higher number of whorls (Fig. 9). FEMALE GENITALIA. The visceral oviduct has only 1 – 2 superficial loops, what makes it one of the simplest in the genus Cochlostoma. The seminal receptacle is thin and may be oval, thus not always club-shaped as in the other species. MALE GENITALIA. The penis is longer than the body and inflated, although less so than in the other species of Clessiniella. The penial spermiduct is gently twisted. The body spermiduct is straight. There is a long groove, ending in a well-formed sperm pocket, delimited at the frontal side by an ascending sperm funnel.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B87913150FFFB7BB014C8F26D2664.taxon	distribution	Distribution The species inhabits a restricted area in the eastern Southern Pre-Alps. Our sampling localities are located between Val Tramontina to the west and Val di Torre to the east (Friuli region, Italy).	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B87913150FFFB7BB014C8F26D2664.taxon	description	Habitat preference Like in most species of Cochlostoma, these are obligate rock-dwelling snails but, rather than on the surface of limestone cliffs, their main habitat is on smaller surfaces of stones and boulders which are normally found in large numbers at the foot of the cliffs.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B87913150FFFB7BB014C8F26D2664.taxon	discussion	Remarks Cochlostoma (Clessiniella) stelucarum sp. nov. was introduced as Cochlostoma sp. by Zallot (2002: 98). At most of the localities, it lives syntopically with either Co. (Eupomatias) philippianum (Gredler, 1853) or Co. (Eupomatias) henricae (Strobel, 1851). Near the village of Anduins it has been found a few centimeters apart from both Co. (Eu.) henricae and Co. (Cochlostoma) septemspirale on stones and boulders. This is one of the rare cases where 3 species of Cochlostoma, belonging to different subgenera, live syntopically. The range of the species is situated in between the range of Co. (Cl.) villae in the west and that of the other species of Clessiniella in the east.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B8791316CFFFD7BF8116BF3D126E0.taxon	description	Figs 1, 3 B, 5, 14 C, 15 C, 16 C, 28 – 30, 34 (bottom), 35 (bottom), 36 (top)	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B8791316CFFFD7BF8116BF3D126E0.taxon	diagnosis	Diagnosis Shell very variable, lip curved backwards more or less abruptly at the columellar side. Strong ribs on the teleoconch, in most of the populations widely spaced and, on the upper whorls, whitish and contrasting with the corneous, dark brown to light yellow, shell background. Visceral oviduct with convoluted loops path; body spermiduct straight.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B8791316CFFFD7BF8116BF3D126E0.taxon	materials_examined	Type material Syntypes ITALY • “ Triest ” [Friuli Venezia Giulia, Trieste]; Westerlund leg.; SMF 160677 / 1 • same label data as for preceding; SMF 160676 / 7 (Fig. 30). Other material AUSTRIA • Dobratsch; 46.59222 ° N, 13.6825 ° E; 2009; Mildner and Giovannelli leg.; EZ 0799 • Kranzwand; 46.588799 ° N, 13.75002 ° E; Jul. 2015; AL leg.; NHMW 109000. CROATIA • Sveta Nedelja near Labin; 45.13741 ° N, 14.14488 ° E; 2000; De Mattia leg.; WdM 6961 • Brest, Cicaria; 45.45444 ° N, 14.00625 ° E; Apr. 2008; De Mattia leg.; WdM 6715 • Drivenik spring; 45.24147 ° N, 14.64828 ° E; Mar. 2009; De Mattia leg.; WdM 6714 • from A 6 to 501 down Bakar; 45.30543 ° N, 14.58579 ° E; Jul. 2009; De Mattia and Zallot leg.; EZ 0938 • Jakačici; 45.179494 ° N, 14.004384 ° E; 2009; De Mattia leg.; WdM 6699 • Kremenje-Buje; 45.43558 ° N, 13.68499 ° E; Jan. 2008; De Mattia leg.; WdM 6400 • Kukurini; 45.19369 ° N, 14.07216 ° E; Feb. 2009; De Mattia leg.; WdM 6698 • Limski Kanal; 45.12957 ° N, 13.74039 ° E; 2010; De Mattia leg.; WdM 6693 • Limski Kanal; 45.12957 ° N, 13.74039 ° E; Jun. 2010; De Mattia leg.; WdM 6998 • Mala Ucka; 45.27605 ° N, 14.19715 ° E; 2009; De Mattia leg.; WdM 6768 • Meja, Bakarac; 45.3037 ° N, 14.58348 ° E; 2009; De Mattia leg.; WdM 6951 • Momjan; 45.442 ° N, 13.71384 ° E; 2009; De Mattia leg.; WdM 6854 • Mt Učka, beech forest; 45.30636 ° N, 14.20865 ° E; 2009; De Mattia leg.; WdM 6794 • Mt Učka, top; 45.28967 ° N, 14.20341 ° E; 2009; De Mattia leg.; WdM 6794 • Pazin; 45.24134 ° N, 13.93099 ° E; Jun. 1986; Drimmer leg.; HNHM 97227 • Permani; 45.43742 ° N, 14.28905 ° E; 2009; De Mattia leg.; WdM 6708 • Raša; 45.0872 ° N, 14.08854 ° E; 2010; De Mattia leg.; WdM 6968 • Rasko Polje; 45.15638 ° N, 14.0538 ° E; 2010; De Mattia leg. • EZ 0813 • Rasko Polje; 45.15638 ° N, 14.0538 ° E; 1900; De Mattia leg.; EZ 1027 • Zminj; 45.13991 ° N, 13.88873 ° E; 2000; De Mattia leg.; WdM 6700. ITALY • Val Rosandra; 45.62 ° N, 13.87139 ° E; 2000; Zallot leg.; EZ 0840 • Torri di Monrupino; 45.71878 ° N, 13.80594 ° E; 1998; Zallot leg.; EZ 0839. SLOVENIA • Abram-Nanos Mt; 45.82055 ° N, 14.01433 ° E; 2008; De Mattia leg.; WdM 6731 • Brestovica pri Komnu; 45.80996 ° N, 13.65016 ° E; 2009; De Mattia leg.; WdM 6948 • Čepno; 45.67071 ° N, 14.09732 ° E; Jun. 2009; De Mattia leg.; WdM 6866 • Črnotiče-Črni Kal; 45.55276 ° N, 13.88633 ° E; Mar. 2009; De Mattia leg.; EZ 1028 • Dolina Nanos-Abran-Podkraj; 45.79813 ° N, 14.06699 ° E; Apr. 2009; De Mattia leg.; WdM 6733 • Farjev Vrh; 45.92429 ° N, 13.95442 ° E; 2009; De Mattia leg.; WdM 6740 • Golac, Vela Vrata; 45.51315 ° N, 14.02969 ° E; 2009; De Mattia leg.; WdM 6762 • Gorjansko; 45.78485 ° N, 13.70068 ° E; 2000; De Mattia leg.; WdM 6950 • Hruščica; 45.85832 ° N, 14.1117 ° E; 2000; De Mattia leg.; WdM 2409 • Ilirska Bistrika; 45.56944 ° N, 14.23917 ° E; 2009; Zallot leg.; EZ 0821 • Kovk; 45.94614 ° N, 13.95058 ° E; 2015 • De Mattia leg.; WdM 6737 • Križna Jama; 45.82003 ° N, 14.24579 ° E; Jun. 2009; Jochum leg.; HNHM 97142 • Markovščina-Dimnice cave; 45.58357 ° N, 14.03994 ° E; 2009; De Mattia leg.; WdM 6765 • Mt Kobilica; 46.06528 ° N, 13.80907 ° E; 2005; De Mattia leg.; WdM 6849 • Mt Nanos, top, loc. Plesa; 45.79813 ° N, 14.06699 ° E; Jun. 2005; De Mattia leg.; WdM 0000 • Mt Snežnik 1161; 45.56816 ° N, 14.37049 ° E; Jun. 2010; De Mattia leg.; WdM 7012 • Mt Snežnik 1390; 45.57179 ° N, 14.41044 ° E; Jun. 2010; De Mattia leg.; EZ 0965 • Mt Snežnik 1447; 45.57877 ° N, 14.43839 ° E; Jun. 2010; De Mattia leg.; EZ 0964 • Mt Snežnik 1450; 45.58195 ° N, 14.43129 ° E; Jun. 2010; De Mattia leg.; EZ 0962 • Mt Snežnik car parking; 45.58235 ° N, 14.43117 ° E; Jun. 2002; De Mattia leg.; WdM 7006 • Mt Vremščica; 45.69898 ° N, 14.03388 ° E; 2009; De Mattia leg.; WdM 6758 • near Gorenje; 45.82976 ° N, 14.10755 ° E; 2009; Zallot leg.; EZ 0817 • near Logatec; 45.88076 ° N, 14.27448 ° E; 2009; Zallot leg.; EZ 0828 • Osp, 45.57071 ° N, 13.86168 ° E; 2010; De Mattia leg.; WdM 6756 • Ospo, small cliff; 45.57119 ° N, 13.8582 ° E; May 2010; De Mattia leg.; WdM 6993 • Otlica; 45.92822 ° N, 13.90918 ° E; 1986; De Mattia leg.; WdM 6752 • Planina; 45.82405 ° N, 14.24708 ° E; 2000; Zallot leg.; EZ 0823 • Planina; 45.82972 ° N, 14.25056 ° E; 2000; Zallot leg.; EZ 0824 • Planina, Postojna; 45.83 ° N, 14.26056 ° E; 2009; Zallot leg.; EZ 0826 • Podgorje; 45.53872 ° N, 13.9449 ° E; 2000; De Mattia leg.; WdM 6735 • Podgorje; 45.5364 ° N, 13.9463 ° E; 2009; De Mattia leg.; WdM 6735 • Podlešče-Bainsizza; 46.05738 ° N, 13.72709 ° E; May 2009; De Mattia leg.; WdM 6814 • Prešnica; 45.56587 ° N, 13.94191 ° E; Mar 2009; De Mattia leg.; WdM 6734 • Pri Cerkvi- Bainsizza; 46.04133 ° N, 13.77584 ° E; 2009; De Mattia leg.; WdM 6834 • Rakow Skocjan; 45.79467 ° N, 14.28948 ° E; 2010; Zallot leg.; EZ 0838 • San Canzian (Škocjan); 45.66532 ° N, 13.99356 ° E; 2009; De Mattia leg.; WdM 6754 • Senožeče; 45.71421 ° N, 14.07856 ° E; 2009; De Mattia leg.; WdM 6865 • 10 km S of Vrhnika; 45.88549 ° N, 14.26091 ° E; 2000; Zallot leg.; EZ 0827 • Vrhnika, Močilnik; 45.95591 ° N, 14.29268 ° E; 2009; Zallot leg.; EZ 0829.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B8791316CFFFD7BF8116BF3D126E0.taxon	discussion	Remarks The original indication of the type locality is rather vague, since only a city in the Friuli-Venezia Giulia region, in the easternmost part of Italy is indicated. The closest to Trieste where we found the species is at “ Val Rosandra ”, 8 km north-east of the city. The fresh specimens collected there have shell features similar to those of the syntypes. We therefore consider them as topotypical.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B8791316CFFFD7BF8116BF3D126E0.taxon	description	Description SHELL. The size of the shells is extremely variable (height 6.1 – 9.6 mm in the females) as well as their general morphology (see Fig. 28). Several populations are spotless whereas others have clear spots on the teleoconch. Depending on the population, the lip can be either poorly or even hugely developed. In lateral view, the body whorl, while approaching the aperture, may widen in various ways, depending on the population (Fig. 4 A: FF, AF, II). The H / W ratio is very variable and the slenderness of the shells in different populations varies accordingly. FEMALE GENITALIA. There is a long, club-shaped seminal receptacle. In some cases there is a change of colour at its base, presumably indicating the starting point of the rather long distal oviduct. As shown in Table 5, there is a considerable variability in the part of the seminal receptacle that is curved down and in the number of loops. MALE GENITALIA. The inflated penis is longer than the body. The penial spermiduct is confined to the central part of the penis and gently twisted. The body spermiduct is straight and well defined.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B8791316CFFFD7BF8116BF3D126E0.taxon	distribution	Distribution This species inhabits the eastern side of the province of Trieste in Italy, the southwestern part of Slovenia and the Istrian peninsula. It has also been found in several Croatian localities in the surroundings of Bakar and, quite far from its main range, on the Villacher Alps, in Austria (see Mildner 1997). The species has been erroneously reported for Albania (see Fehér & Erőss 2009).	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B8791316CFFFD7BF8116BF3D126E0.taxon	description	Habitat preference It is an obligate rock-dwelling species. Co. (Cl.) tergestinum is found in a variety of habitats, from open, exposed limestone cliffs to shady spots in deciduous and evergreen forests.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B8791316CFFFD7BF8116BF3D126E0.taxon	discussion	Remarks According to Zilch (1958) and Alzona (1971) there are 3 subspecies of Co. (Cl.) tergestinum, viz. the nominate subspecies Co. (Clessiniella) tergestinum tergestinum (Westerlund, 1878), Co. (Clessiniella) tergestinum grahovanum (A. J. Wagner, 1897) and Co. (Clessiniella) tergestinum tortivum (Westerlund, 1885). For the time being, we here synonymize these nominal taxa because the conchological features which supposedly characterize them, like measures, density and prominence of the ribs, and the structure of the lip, are population features without an obvious geographical pattern. There is no significant correlation between latitude and longitude of the locality of the sample and the character states that are mentioned in the original description of the so-called subspecies. Moreover, in the surroundings of Trieste, the type locality of the nominate subspecies, only populations with some ‘ tortivum ’ shell features, like the thin lip, can be found (De Mattia & Prodan 2006). Some of the populations have a peculiar shell morphology (see for instance Fig. 28 E – F); their taxonomical status should be investigated more thoroughly especially since the available molecular H 3 analysis indicates that population “ EZ 0813 ” from Rasko Polje, corresponding to the specimen in Fig. 28 E, stands apart from the others.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B8791316BFFF37B9A12ABF5F92792.taxon	description	Figs 1, 3 D, 14 E, 15 E, 16 E, 31 – 33, 34 (top), 35 (top), 36 (bottom)	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B8791316BFFF37B9A12ABF5F92792.taxon	diagnosis	Diagnosis Spotless shell. Moderately strong, more or less widely spaced ribs. Columellar lobe gently curved backwards. External lobe of type “ II ” in almost all the populations. The body spermiduct is huge and twisted instead of straight as in all the other species of Clessiniella.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B8791316BFFF37B9A12ABF5F92792.taxon	materials_examined	Type material Syntypes CROATIA • ♀; Ogulin; A. J. Wagner leg.; MIZPAS-W- 6979 a (Fig. 33) • 2 ♀♀, 3 ♂♂; same collection data as for preceding; MIZPAS-W- 6979 / 5. Other material CROATIA • Kapela pass, Ogulin; 45.07534 ° N, 15.20837 ° E; 2009; De Mattia leg.; EZ 0909 • Klek Mt, close to the top; 45.25904 ° N, 15.14044 ° E; 2009; Zallot leg.; EZ 0904 • Klek Mt, cliffs halfway; 45.25438 ° N, 15.14561 ° E; 1996; Zallot leg.; EZ 0905 • Mt Risnjak; 45.43095 ° N, 14.6188 ° E; 1400 m a. s. l.; 2009; De Mattia leg.; EZ 0845 • Mt Risnjak; 45.39072 ° N, 14.61748 ° E; 1000 m a. s. l.; 2010; Zallot leg.; EZ 0907 • Ročko Polje; 45.3607 ° N, 14.09534 ° E; 2009; De Mattia leg.; WdM 6697 • Rudnica; 45.23402 ° N, 15.33827 ° E; 2009; De Mattia leg.; EZ 0906 • Spilja Vrelo; 45.31777 ° N, 14.71065 ° E; 2009; Zallot leg.; EZ 0899 • Vitunj-Ogulin; 45.29114 ° N, 15.14041 ° E; 2005; Zallot leg.; EZ 0903. SLOVENIA • Benete, 45.76192 ° N, 14.56752 ° E; 2009; De Mattia leg.; WdM 6805 • Bresovica; 45.95607 ° N, 14.43533 ° E; 2006; De Mattia leg.; EZ 0897 • Bresovica pri Predgradu; 45.55164 ° N, 15.0437 ° E; 2009; De Mattia leg.; WdM 6612 • Divje Jezero; 45.9827 ° N, 14.0277 ° E; 2009; De Mattia leg.; WdM 6840 • Dol-E of Čepovan; 46.06445 ° N, 13.81189 ° E; 2009; De Mattia leg.; WdM 6837 • Jelenja Vas; 45.5115 ° N, 15.05346 ° E; 2009; De Mattia leg.; WdM 6804 • Krempa Kolpa; 45.53417 ° N, 15.09528 ° E; 1996; Mildner leg.; EZ 0902 • Marija Ceplje Kolpa; 45.53444 ° N, 15.09389 ° E; 2006; Mildner leg.; EZ 0901 • Mt Snežnik 1150; 45.56913 ° N, 14.321 ° E; Jun. 2010; De Mattia leg.; WdM 7014 • Mt Snežnik 1620; 45.58845 ° N, 14.44209 ° E; Jun. 2010; De Mattia leg.; EZ 0963 • Mt Snežnik car parking; 45.58235 ° N, 14.43117 ° E; Jun. 2010; De Mattia leg.; WdM 7006 • Polog-E of Čepovan; 46.06387 ° N, 13.81275 ° E; 2009; De Mattia leg.; WdM 6842 • SW Pescena Glava; 46.05156 ° N, 13.73844 ° E; 843 m a. s. l.; 2009; De Mattia leg.; WdM 6829 • Tisovec; 45.81809 ° N, 14.76859 ° E; 2009; De Mattia leg.; WdM 6749 • same collection data as for preceding; WdM 6750 • Vrsic Pass; 46.44278 ° N, 13.76417 ° E; 1985; Zallot leg.; EZ 0896 • Zagradec- Polyane; 45.86028 ° N, 14.83417 ° E; Bodon leg.; EZ 0900.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B8791316BFFF37B9A12ABF5F92792.taxon	description	Description SHELL. The shell is more or less light corneous brown and spotless. The size of the aperture varies considerably among the populations; shells with a large aperture and a correspondingly large lip occur next to shells with a poorly developed lip and a relatively small aperture. The lip is gently curved backwards on the columellar side. FEMALE GENITALIA. The female genitalia are hardly distinguishable from those of Co. (Cl.) tergestinum if not for the more relaxed loops. MALE GENITALIA. There is a long, inflated penis. The penial spermiduct is slightly twisted and occupies only the central part of the penis. The body spermiduct is twisted and conspicuously protruding (see Figs 34 – 35).	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B8791316BFFF37B9A12ABF5F92792.taxon	distribution	Distribution The species inhabits the north-western and south-eastern part of Slovenia and an adjacent area in Croatia. It is reported from Austria for the Karawanken and the Gailtaler Alpen (Edlinger & Mildner 1979). Its range largely surrounds that of Co. (Cl.) tergestinum (see Fig. 1).	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B8791316BFFF37B9A12ABF5F92792.taxon	description	Habitat preference The snails are found on rocky cliffs as well as on boulders in woody areas.	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
038B8791316BFFF37B9A12ABF5F92792.taxon	discussion	Remarks To assign a given population to Co. (Cl.) waldemari rather than to Co. (Cl.) tergestinum on the basis of the morphology of the shell and the female genitalia alone could be challenging if not impossible (see Figs 34 – 35). Only the structure of the male genitalia allows a convincing identification of the species. The H 3 analysis (Fig. 7) confirms that these sibling species belong to separate clades within Clessiniella. Boeckel (1939) considered Co. (Cl.) waldemari a subspecies of Co. (Cl.) tergestinum. This view was contradicted by Bole (1994). In some areas, like on Mount Snežnik in southern Slovenia, the two species live not far from each other and here they can be distinguished also conchologically while differing in the presence or absence of the spots on the teleoconch (with Co. (Cl.) waldemari having a spotless shell) and in the shape of the body whorl approaching the aperture, in lateral view (Fig. 36).	en	Zallot, Enrico, Mattia, Willy De, Fehér, Zoltán, Gittenberger, Edmund (2021): Cochlostoma revised: the subgenus Clessiniella Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae). European Journal of Taxonomy 762 (1): 49-95, DOI: https://doi.org/10.5852/ejt.2021.762.1453, URL: http://dx.doi.org/10.5852/ejt.2021.762.1453
