taxonID	type	description	language	source
039487A4FFE7FFE4FD84F13EFAA2FDE3.taxon	type_taxon	Type species Nisitra vittatus (Haan, 1844).	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFE7FFE4FD84F13EFAA2FDE3.taxon	discussion	Remarks on type species Kirby (1906) and Chopard (1968) listed N. vittata as the type species of Nisitra and treated the original type species N. marginata Walker, 1869 as a synonym of N. vittata. Subsequent authors listed N. vittatus as the type species of the replacement name Nisitrus, although Otte (1994) reinstated N. marginatus as a valid species (Braun 2013 in Cigliano et al. 2020). The present revision revalidates the synonymy of N. marginatus under N. vittatus, making N. vittatus the valid type species of the genus.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFE7FFE4FD84F13EFAA2FDE3.taxon	diagnosis	Diagnosis (emended from Robillard & Tan 2013) Among genera of Eneopterinae, Nisitrus is characterised by an elegant, wasp-like, slender and colourful body (Figs 7 – 9). Size average for subfamily. Colouration vivid, dominantly yellow, whitish and black, with species-specific patterns, particularly in the head (Figs 10 – 11), pronotum (Figs 11 – 12) and FWs (Figs 12 – 14). Characteristic head shape with narrow fastigium and large prominent eyes with thin dorso-ventral black stripes. Pronotum setose, square-shaped to slightly trapezoidal, usually bisinuated posteriorly. Metanotum with large glandular structures in males, made of a wide pit on scutum and a pair of posterior pits on scutellum, with glandular pores mostly organised by groups of 2 or 3 (Fig. 15). FWs more or less as long as abdomen. Male Nisitrus show modified FW venation and characteristic glossy transparency. Hindwings longer than FWs, bicoloured in most species, their base transparent with a wide dark posterior margin. Legs long and thin, FIII with a characteristic narrowed area before knee. Other general traits: TI with two tympana, inner one slit-like, covered by a swollen cuticular expansion; outer tympanum oval, its membrane transversally plicate in dorsal half. TI with three apical spurs; outer dorsal spur missing. TII with four apical spurs, inner longest. TIII with three inner and three outer apical spurs, median longest on each side; four pairs of subapical spurs, inner and outer spurs almost straight, their apex hook-like; TIII serrulate over whole length, slightly furrowed dorsally. Tarsomeres III- 1 with one row of dorsal spines on external edge in addition to apical spines. Apical claws of legs slightly indented. Both males and females show characteristic genitalia.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFE7FFE4FD84F13EFAA2FDE3.taxon	description	Male genitalia (Fig. 16) with membranous and setose lophi well individualised; pseudepiphallic parameres including two membranous posterior lobes and a sclerotized anterior lobe oriented anteriorly toward median axis of genitalia and carrying a secondary preapical sclerotized lobe; ectophallic arc transverse, incomplete due to a little-sclerotized median area; ectophallic fold narrowed preapically, its apex narrow, located between anterior apex of pseudepiphallic parameres; dorsal part of fold sclerotized, forming two elongated sclerites connected to the median area of arc; ventral part of ectophallic fold membranous, with two strong rounded lateral sclerites of variable shapes. Membrane of endophallic cavity smooth. Female copulatory papilla (Fig. 17) conical, apex generally sclerotized and pointed. Ovipositor typically longer than FIII; apex thin with both dorsal and ventral edges smooth. Phylogenetic relationships Based on previous phylogenetic studies (Robillard & Desutter-Grandcolas 2004 c; Nattier et al. 2011; Vicente et al. 2017), Nisitrus is the sister genus of Paranisitra Chopard, 1925 (revision in Gorochov 2009 and Baroga-Barbecho et al. 2019). The two genera form the Nisitrini tribe (Robillard & Desutter-Grandcolas 2008) and are very close morphologically; Paranisitra is mostly characterised by a lack of wings in both sexes, but except for this, it shares with Nisitrus its general body shape, long thin legs with indented claws, colouration with yellow and black, head shape with large prominent eyes and narrow fastigium, and the general structure of male and female genitalia. Nisitrini are less easy to relate to other clades of Eneopterinae.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFE7FFE4FD84F13EFAA2FDE3.taxon	biology_ecology	Natural history Species tend to be found at forest edges and gaps in the forest where the vegetation is more exposed. They are also found among shrubs and herbaceous plants (but not grasses). Nisitrus are observed to be generally diurnal, during which the males can be heard calling and individuals are found foraging in the vegetation. It seems as Nisitrus are mostly herbivorous, feeding on leaves, but also on flowers (Tan et al. 2017). Nisitrus are also observed to congregate with one another on the same plant, sometimes making up of more than 20 individuals of both adults and nymphs. They can be considered a pest in some parts of Southeast Asia (Willemse 2001).	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFE7FFE4FD84F13EFAA2FDE3.taxon	distribution	Distribution Borneo (Sabah and Sarawak States of East Malaysia, Brunei Darussalam, and Kalimantan State of Indonesia), Java, Malay Peninsula (Peninsular Malaysia and Singapore) and Sumatra (Indonesia) (Fig. 5).	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFE7FFE4FD84F13EFAA2FDE3.taxon	discussion	Key to the species of Nisitrus Saussure, 1878	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFF4FFE7FDD6F13DFEECF9EF.taxon	description	Figs 5, 7 A, 8 A – B, 10 A, 11 A, 12 A, 13 A, 14 A, 15 B, 16 A, 17 A – B, 18; Tables 1 – 3	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFF4FFE7FDD6F13DFEECF9EF.taxon	diagnosis	Diagnosis This species differs from all known congeners by the following characters: head nearly completely black (sometimes with yellow patch beneath scapes and eyes). Pronotum lateral lobe either black with small brown spot on postero-ventral corner, or black in dorsal half and yellow in ventral half. Male FW with basal area brown with tint of black, otherwise mostly hyaline; veins mostly yellow. Female FW with dorsal field mostly infumated black, with veins dark brown and lighten towards lateral field and apical areas. Legs red brown; FIII black dorso-basally without bands or stripes on external surface; with a distinct black ring before hind knee; hind knee black. Abdomen with tergites black, sternites mostly black. This species is similar to N. crucius sp. nov. and N. rindu sp. nov. but differs by the combination of these characters: frons nearly completely black (without black cross marking as in N. crucius sp. nov.), dorsum of head completely black (vs with yellow margins in N. crucius sp. nov. or pale with dark patterns in N. rindu sp. nov.) and lateral lobe either with ventral half yellow or with an oval yellow spot at the anterior end (vs with a characteristic dorsal yellow band that extends ventrally in N. rindu sp. nov.).	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFF4FFE7FDD6F13DFEECF9EF.taxon	materials_examined	Material examined Lectotype (here designated) NO COUNTRY • ♀; “ Boncard ded. ”; Brunner von Wattenwyl leg.; NMW. Paralectotype (here designated) NO COUNTRY • 1 ♀; same collection data as for lectotype; NMW. Additional specimens INDONESIA – Kalimantan • 1 ♀; [East Kalimantan] “ West Borneo, Marah ”, [near border of Koetai and Beraoe]; 1 ° 42 ′ 28 ″ N, 116 ° 15 ′ 40 ″ E; 10 Aug. 1925; H. C. Siebers leg.; MNHN • 1 ♂; [East Kalimantan] “ West Borneo, Marah ”; 0 ° 58 ′ 31 ″ N, 116 ° 33 ′ 14.8 ″ E; 10 – 28 Nov. 1925; H. C. Siebers leg.; MNHN-EO- ENSIF 1734 • 7 ♀♀; same collection data as for preceding; MNHN • 2 ♀♀; “ West Borneo, Midden ”, [near border of Koetai and Beraoe]; 1 ° 47 ′ 35 ″ N, 116 ° 12 ′ 22 ″ E; 12 Sep. 1925; H. C. Siebers leg.; MNHN • 1 ♂; “ West Borneo, Midden ”, [near border of Koetai and Beraoe]; 1 ° 50 ′ 14 ″ N, 116 ° 12 ′ 34 ″ E; 26 Sep. 1925; H. C. Siebers leg.; MNHN-EO-ENSIF 1726 • 1 ♂; “ West Borneo, Midden ”, [near Beraoe?]; 1 ° 58 ′ 53 ″ N, 116 ° 7 ′ 56 ″ E; 1 Oct. 1925; H. C. Siebers leg.; MNHN-ENSIF 1752 • 1 ♂; same collection data as for preceding; MNHN • 1 ♂; “ West Borneo, Midden ”, [near border of Koetai and Beraoe]; 1 ° 54 ′ 44 ″ N, 116 ° 11 ′ 27 ″ E; 6 Nov. 1925; H. C. Siebers leg.; MNHN-ENSIF 1715 • 2 ♀♀; same collection data as for preceding; MNHN • 1 ♀; [Melawi Regency] “ West Borneo, Melawi ”; 0 ° 35 ′ 43 ″ S, 0 ° 0 ′ 46 ″ E; Nov. – Dec. 1924; Blanchemanche leg.; MNHN • 1 ♀; “ West Borneo ”; 1924; Blanchemanche leg.; MNHN • 1 ♀; “ Lebang Hara ”; 25 Nov. – 5 Dec. 1924; H. Winkler leg.; ZMH • 1 ♀; “ Samarinda ”; 0 ° 29 ′ 47 ″ S, 117 ° 8 ′ 23 ″ E; M. - E. Walsh leg.; ZMLU • 1 ♀; Kayan Basin, Bahau; 2 ° 55 ′ 23.2 ″ N, 117 ° 33 ′ 13.1 ″ E; 13 Nov. 1999; H. H. Tan and D. Wowor leg.; ZRC • 1 ♂; same collection data as for preceding; ZRC • 1 ♀; “ O. Borneo Pelawan ”; May 1937; M. - E. Walsh leg.; ZMLU • 4 ♀♀; Penda Siroen, Soengai Maroewai [Sungai Maruwai]; 0 ° 34 ′ 1.9 ″ S, 114 ° 44 ′ 0.5 ″ E; 13 May 1932; Prince Leopold leg.; RBINS • 1 ♀; same collection data as for preceding; RBINS • 1 ♀; Kumai, West Kotawaringin Regency, Central Kalimantan; 2 ° 55 ′ 17.9 ″ S, 111 ° 54 ′ 16.4 ″ E; 24 Jun. 2014; G. Cahyadi leg.; iNaturalist. org / observations / 41219515. – Java • 2 ♂♂; “ Bibidjilan Djampang, Tengah VI. ”; 7 ° 20 ′ 8.2 ″ S, 107 ° 38 ′ 21.2 ″ E; M. - E. Walsh leg.; ZMLU. MALAYSIA – Sarawak • 1 ♀; Balleh River, Kapit jungle; 1 ° 45 ′ 14.51 ″ N, 113 ° 45 ′ 59.00 ″ E; 15 Apr. 1979; PII: N 53; MNHN-EO-ENSIF 11260 • 2 ♀♀; Balleh River, Kapit jungle; 1 ° 45 ′ 14.51 ″ N, 113 ° 45 ′ 59.00 ″ E; 15 Apr. 1979; MNHN • 1 ♀; 10 miles south of Kuching; 1 ° 15 ′ 12.94 ″ N, 110 ° 19 ′ 36.71 ″ E; 14 Jun. 1910; Beebe leg.; MZB • 1 juvenile; “ Gunung Mulu NP, Camp 4, Summit Trail ”; 4 ° 2 ′ 40.6 ″ N, 114 ° 54 ′ 50.8 ″ E; 5950 ft a. s. l.; 19 Jan. 2009; G. J. Svenson leg.; MNHN • 1 ♀; Sarawak; 1909; C. J. Brooks leg.; NHMUK 012497567 • 1 ♀; [Semongoh] “ Semongok, on oil palm ”; 1 ° 23 ′ 51 ″ N, 110 ° 18 ′ 43 ″ E; 19 Dec. 1966; “ Salin Iuses ” leg.; NHMUK 012497569. NO COUNTRY • 1 ♀; “ Borneo ”; [19] 32; “ Weir ” leg.; AMS-K 393086 • 1 ♀; “ Borneo ”; 1891; Chopard leg.; MNHN • 1 ♀; “ Federal Malay States ”; 1909; C. J. Brooks leg.; NHMUK 012497568.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFF4FFE7FDD6F13DFEECF9EF.taxon	description	Redescription Relatively large and slender (Figs 7 A, 18). Vertex completely black (Fig. 11 A). Fastigium velvety black, with anterior margins yellow. Scapes black, posterior end red brown. Antennae red brown with some segments black and with wide whitish rings. Eyes dark green when alive and red brown when dried. Frons generally black, sometimes with yellow to cream-coloured bands above and beneath antennal cavity and along ventral margins of eyes (Fig. 10 A). Mouthparts dark red brown to black. Maxillary palpi creamcoloured; apical segment black apically (Fig. 10 A). Head lateral side black (Fig. 12 A). Pronotum generally black, covered with white setae; dorsal disk trapezoidal, posterior margin bisinuated (Fig. 11 A). Lateral lobes of pronotum with two variants: (1) in lectotype, completely black except for a cream-coloured oblong spot near anterior margin (Fig. 12 A); (2) dorsal third black, otherwise completely yellow. Legs red brown. FIIIs dark red brown, sometimes with a dorsal black longitudinal stripe; knees dark brown to black; TIIIs brown with black spines and spurs, dark brown to black near distal end; tarsomeres dark brown to black. Hindwings hyaline brown, with a rounded transparent window near internal-basal area; longer than FWs, the dark brown tail exceeding FWs more than twice as long as pronotum. Tergites black, in some segments with cream-coloured transverse stripe at anterior end and / or with pale band laterally; sternites pale, their median area red brown. Cerci cream-coloured, short and conical. Male FOREWINGS. FW colouration (Fig. 13 A): dorsal field cells mostly transparent to pale yellow, veins mostly dark brown to black. Basal area brown with tint of black. R brown, Sc projections brown basally, transparent apically. Lateral field yellow brown, apex more transparent. FW venation (Fig. 13 A): 1 A somewhat straight. Harp slightly longer than wide, with 3 oblique veins, posterior-most vein bifurcated at base. c 1 long and wide, c 2 diamond-shaped; mirror (d 1) longer than wide, not rounded, generally separated into two parts by a distinct, roundly bent transverse vein, the posterior part hexagonal, shorter than anterior part. Cell d 2 narrower than d 1. Apical field short and rounded, with 2 wide cell alignments posterior to mirror and a narrow apical alignment; veins yellow. Lateral field with 5 – 6 projections of Sc. Epiproct black. Subgenital plate pale cream, its median area and anterior and posterior margins black. GENITALIA (Fig. 16 A). Pseudepiphallus sclerotized, relatively large and broad compared to congeners, anterior margin somewhat broad and straight, posterior margin also straight, lateral margins faintly converging posteriorly. Posterior apex with paired lophi longer than wide (at base), subacute at apex, sclerotized laterally only and covered with strong short setae; spaced apart from one another. Rami swollen preapically, anterior apex somewhat truncated. Pseudepiphallic parameres narrow, divergent posteriorly, their basis membranous, with a sclerotized lobe on anterior apex. Ectophallic arc transverse, complete, with anterior and posterior margins straight. Ectophallic fold strong rounded lateral sclerites appearing bean-shaped. Ectophallic apodemes long and thin, straight and slightly divergent basally. Endophallic sclerite large and sclerotized, its posterior apex with divergent lateral arms and with a short median expansion curved dorsally. Endophallic apodeme with lateral lamellae stout. Female FOREWINGS. FW colouration (Fig. 14 A): cells on dorsal field mostly infumated black, gradually becoming yellow near CuA, M and R veins; veins generally black or dark brown, also gradually becoming yellow towards CuA, M and R veins; veins near apical area also yellow. Sc projections cream-coloured basally, black apically. Lateral field dark. FW venation: 6 – 7 strong longitudinal veins on dorsal field, sometimes not smooth; lateral field with 6 projections of Sc. GENITALIA. Ovipositor: on average longer than FIII; apex thin with both dorsal and ventral edges smooth. Copulatory papilla conical, slender, more elongated than congeners; apex folded ventrally, long and slender, pointed; dorsal face with a sclerotized area; ventro-anterior end forming a somewhat triangular rim (Fig. 17 A – B). Juvenile Unknown. Measurements See Table 3.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFF4FFE7FDD6F13DFEECF9EF.taxon	biology_ecology	Ecology Unknown.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFF4FFE7FDD6F13DFEECF9EF.taxon	distribution	Distribution Borneo (Kalimantan, Sarawak), West Java.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFF4FFE7FDD6F13DFEECF9EF.taxon	description	Calling song Unknown.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFF7FFE2FDC5F533FEECFE3A.taxon	description	urn: lsid: zoobank. org: act: 3140 F 3 A 4 - 21 FB- 4 BB 4 - ACC 6 - 57 BD 749 A 6 C 3 B Figs 5, 7 B, 8 C, 10 B, 11 B, 12 B, 13 B, 14 B, 16 B, 17 C – D; Tables 2, 4	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFF7FFE2FDC5F533FEECFE3A.taxon	diagnosis	Diagnosis This species differs from all known congeners by the following combination of characters: frons yellow, with a cross-shaped black pattern above mouthparts; face part of fastigium between eyes black. Male FWs with a basal area generally paler; female ovipositor only about as long as FIII. Nisitrus crucius sp. nov. is similar to N. brunnerianus, from which it differs by smaller size and shorter ovipositor. It is also similar in size to N. vittatus and N. malaya sp. nov. but differs by female FW with cells on dorsal field mostly infumated black, gradually becoming yellow near CuA, M and R veins.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFF7FFE2FDC5F533FEECFE3A.taxon	etymology	Etymology The species name refers to the dark cross marking on the frons (ʻ crucis ʼ = ʻcrossʼ, in Latin).	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFF7FFE2FDC5F533FEECFE3A.taxon	materials_examined	Material examined Holotype INDONESIA – Kalimantan • ♂; “ Biv. Long Hoet ”; 1 ° 21 ′ 7.2 ″ N, 116 ° 31 ′ 4.1 ″ E; 16 – 20 Aug. 1925; H. C. Siebers leg.; MNHN-EO-ENSIF 11261. Paratypes INDONESIA – Kalimantan • 1 ♀; “ Long Navang ” [Long Nawang]; Mjöberg leg.; MNHN-EO- ENSIF 11262 • 1 ♀; “ Midden O. Borneo, Marah ”; 0 ° 58 ′ 31 ″ N, 116 ° 33 ′ 14.8 ″ E; 10 – 28 Nov. 1925; H. C. Siebers leg.; MNHN-EO-ENSIF 11263 • 1 ♀; “ Biv. Long Hoet ”; 1 ° 21 ′ 7.2 ″ N, 116 ° 31 ′ 4.1 ″ E; 16 – 20 Aug. 1925; H. C. Siebers leg.; MNHN-EO-ENSIF 11264 • 1 ♀; “ Melawi W Borneo ”; 0 ° 34 ′ 28 ″ S, 111 ° 45 ′ 5 ″ E; Nov. – Dec. 1924; Blanchemanche leg.; MNHN-EO-ENSIF 1732 • 1 ♂; same collection data as for preceding; MNHN-EO-ENSIF 11265 • 1 ♀; “ Samarinda ”; 0 ° 29 ′ 47 ″ S, 117 ° 8 ′ 23 ″ E; M. - E. Walsh leg.; ZMLU. MALAYSIA – Sarawak • 1 ♂; “ Kapit ”; 1 ° 46 ′ 0 ″ N, 113 ° 32 ′ 12 ″ E; 9 Jul. 1970; Baebe leg.; Ind. Mus.; PII: N 56; MNHN-EO-ENSIF 1750. – Sabah • 1 ♂; “ nr. Long Pa Sia (West) ”; 4 ° 24 ′ 21 ″ N, 115 ° 43 ′ 47 ″ E; “ c. 1200 m ”; 2 – 13 Apr. 1987; C. v. Achterberg leg.; RMNH. NO COUNTRY • 1 ♀; “ Borneo ”; 1891; Chaper leg.; MNHN-EO-ENSIF 11266.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFF7FFE2FDC5F533FEECFE3A.taxon	description	Description Average size (Fig. 7 B). Colour pattern somewhat intermediate between N. brunnerianus and N. vittatus. Vertex black with cream-coloured margin around eyes (Fig. 11 B). Fastigium velvety black with creamcoloured margins. Scapes black dorsally, sometimes reddish ventrally or along dorso-inner margins. Antennae black. Frons yellow, face part of fastigium black between eyes, with a cross-shaped black pattern above mouthparts; clypeus sometimes darkened, mouthparts dark red brown to black (Fig. 10 B). Maxillary palpi cream-coloured. Head lateral side generally black. Pronotal disk black with white setae (Fig. 11 B). Lateral lobes of pronotum with dorsal third black, cream-coloured ventrally (Fig. 12 B). Legs red brown, sometimes internal-basal area black. FIIIs dark red brown, knees dark brown to black; TIIIs brown with black spines and spurs, dark brown to black near distal end; tarsomeres dark brown to black. Hindwings dark hyaline, with a rounded transparent window near internal-basal area; longer than FWs, the dark brown tail exceeding the FWs about twice as long as pronotum. Tergites black, with a pale band laterally of variable thickness; sternites pale, with a black median area. Male FOREWINGS. FW colouration (Fig. 13 B): dorsal field cells mostly transparent; veins mostly creamcoloured, sometimes black. Basal area brown, basally cream-coloured, brown towards distal part. R and Sc projections cream-coloured. Lateral field hyaline, with dorsal parts infumated cream-coloured. FW venation (Fig. 13 B): 1 A somewhat straight. Harp slightly longer than wide, with 2 oblique veins, distal one bifurcate basally. Cell c 1 long and wide, c 2 diamond-shaped; mirror (d 1) longer than wide, not rounded, generally separated into two parts by a distinct straight transverse vein, the posterior part somewhat rectangular, much shorter than anterior part. Cell d 2 narrower than d 1, usually subdivided by accessory veins. Apical field short and rounded, with 3 wide cell alignments posterior to mirror and a narrow apical alignment; veins cream-coloured. Lateral field with 5 projections of Sc. Epiproct black. Subgenital plate pale cream-coloured with median area and anterior and posterior margins black. GENITALIA (Fig. 16 B). Pseudepiphallus sclerotized, stout (small but broad) compared to congeners, anterior margin somewhat broad and straight, posterior margin also straight, lateral margins faintly converging posteriorly. Posterior apex with paired lophi slightly longer than wide (at base), obtuse at apex, sclerotized laterally only and covered with short strong setae; narrowly spaced apart from one another. Rami swollen preapically, anterior apex somewhat truncated. Pseudepiphallic parameres narrow, divergent posteriorly, their basis membranous, with a sclerotized lobe on anterior apex. Ectophallic arc transverse, complete, with anterior and posterior margins straight. Ectophallic fold strong rounded lateral sclerites appearing bean-shaped. Ectophallic apodemes long, straight and slightly divergent basally. Endophallic sclerite large and sclerotized, its posterior apex with divergent lateral arms and with a short median expansion curved dorsally. Endophallic apodeme with lateral lamellae stout. Female FOREWINGS. FW with cells on dorsal field mostly infumated black, gradually becoming yellow near CuA, M and R veins; veins generally yellow (Fig. 14 B). Sc projections cream-coloured. Lateral field transparent, sometimes infumated black apically. GENITALIA. Ovipositor: only slightly longer than FIII. Copulatory papilla conical, smaller and stout; apex folded ventrally, short, pointed; dorsal face with a sclerotized area; ventro-anterior end forming an oval to pyriform rim (Fig. 17 C – D). Juvenile Unknown. Measurements See Table 4.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFF7FFE2FDC5F533FEECFE3A.taxon	biology_ecology	Ecology Unknown.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFF7FFE2FDC5F533FEECFE3A.taxon	distribution	Distribution Borneo (Kalimantan, Sabah, Sarawak).	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFF7FFE2FDC5F533FEECFE3A.taxon	materials_examined	Type locality Indonesia, Kalimantan.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFF7FFE2FDC5F533FEECFE3A.taxon	description	Calling song Unknown.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFF2FFDCFDC7F167FEECF96A.taxon	description	urn: lsid: zoobank. org: act: 0 DEB 5 BDB- 6 E 85 - 4691 - B 5 D 1 - E 6258239370 A Figs 5, 8 D, 10 C, 11 C, 12 C, 14 C, 17 E, 19; Tables 2, 5	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFF2FFDCFDC7F167FEECF96A.taxon	diagnosis	Diagnosis This new species differs from all known congeners by face part of fastigium with a black band across eyes and antennal cavity, frons otherwise yellow (reddish on preserved specimens); pronotal lateral lobe black dorsally and yellow ventrally; longitudinal white stripe only between M and R and lateral field matted black; Sc projections black. This new species is most similar to N. hughtani sp. nov. from Sumatra, but differs by the following characters: smaller in size, vertex black with yellow margin around eyes (instead of cream-coloured), frons with a black band across eyes and antennal cavity (yellow in N. hughtani sp. nov.); frons yellow (instead of black with a cream-coloured band ventral to eyes); mouthparts yellowish with some black spots; lateral part of head with a black band posterior of eye (instead of black with a cream-coloured ring around eye). This species is also similar to N. musicus from Kinabalu range, but differs by mouthparts yellow (instead of black), FIIIs not black dorsally, female papilla, and distribution: N. musicus is restricted to the highlands in the Kinabalu range, while N. danum sp. nov. occurs in the Danum Valley and Tawau Hill areas at the east of Sabah.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFF2FFDCFDC7F167FEECF96A.taxon	etymology	Etymology The specific name is a noun in apposition taken from the type locality, Danum Valley, Malaysia.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFF2FFDCFDC7F167FEECF96A.taxon	materials_examined	Material examined Holotype MALAYSIA – Sabah • ♀; Danum Valley Field Centre; 4 ° 57 ′ 52.95 ″ N, 117 ° 48 ′ 18.25 ″ E; ca 150 m a. s. l.; 26 May – 20 Jun. 1987; C. v. Achterberg and Kennedy leg.; PII: N 54; RMNH. Paratypes MALAYSIA – Sabah • 1 ♀; same collection data as for holotype; RMNH • 1 ♀; same collection data as for holotype; MNHN-EO-ENSIF 10945. Additional specimens MALAYSIA – Sabah • 1 ♀; Tawau Hill Park; 4 ° 24 ′ 0.0 ″ N, 117 ° 53 ′ 24.0 ″ E; 7 Mar. 2018; C. Elder leg.; iNaturalist. org / observations / 19511712 • 1 ♀; Danum Valley; 4 ° 57 ′ 42.6 ″ N, 117 ° 41 ′ 22.2 ″ E; 2 Oct. 2020; L. Day leg.; iNaturalist. org / observations / 21010925.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFF2FFDCFDC7F167FEECF96A.taxon	description	Description Slender and relatively large habitus (Figs 8 D, 19). Vertex black with yellow margin around eyes (Fig. 11 C). Fastigium velvety black with yellow margins. Scapes dark red brown to black (particularly on the dorsoinner margins). Antennae black with wide whitish rings. Face part of fastigium with a black band across eyes and antennal cavity, otherwise yellow (reddish in preserved specimens); frons almost entirely yellow, including mouthparts, with some black spots (Fig. 10 C). Maxillary palpi cream-coloured; apical segment black apically (Fig. 10 C). Lateral part of head with a black band posterior to eye (Fig. 12 C). Pronotum covered with white setae; dorsal disk dark brown to black (Fig. 11 C). Lateral lobes of pronotum dorsal third black, vivid yellow ventrally (cream-coloured in preserved specimens) (Fig. 12 C). Legs generally red brown, sometimes dorsally darker. FIIIs brown, knees dark brown to black; TIIIs brown with black spines and spurs, dark brown to black near distal end; tarsomeres dark brown to black. Tergites red brown, with a pale band laterally; sternites pale, with red brown median area. Cerci red brown, short and conical. Male Unknown. Female FOREWINGS. FW colouration (Fig. 14 C): veins generally yellow; cells infumated black. Longitudinal infumated white stripe only in M and R. Lateral field matted black, Sc projections black. FW venation: 6 – 7 strong longitudinal veins on dorsal field, cross-veins irregular and often sinuous; lateral field with 6 projections of Sc. GENITALIA. Ovipositor: distinctly longer than FIII; apex thin with both dorsal and ventral edges smooth. Copulatory papilla conical, smaller and stout; apex strongly folded (could be due to the preservation of specimen) ventrally, short, pointed; dorsal face with a sclerotized area (Fig. 17 E). Juvenile Unknown. Measurements See Table 5.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFF2FFDCFDC7F167FEECF96A.taxon	biology_ecology	Ecology Unknown.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFF2FFDCFDC7F167FEECF96A.taxon	distribution	Distribution Borneo (Sabah).	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFF2FFDCFDC7F167FEECF96A.taxon	materials_examined	Type locality Malaysia: Sabah.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFF2FFDCFDC7F167FEECF96A.taxon	description	Calling song Unknown.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFCCFFD8FDFAF5B4FEECF9E2.taxon	description	urn: lsid: zoobank. org: act: 8 E 5 BC 124 - 4 F 3 C- 432 E-BB 94 - A 19 C 32128 B 06 Figs 5, 7 C, 8 E, 10 D, 11 D, 12 D, 13 C, 14 D, 15 A, 16 C, 17 F – G, 20; Tables 1 – 2, 6	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFCCFFD8FDFAF5B4FEECF9E2.taxon	diagnosis	Diagnosis Large and slender species. Head mostly black, but with cream-coloured margin around eyes in both dorsal and lateral views; frons mostly black, but yellow ventral to eyes and antennal cavity; maxillary palpi white. Pronotum dorsal lobe black, lateral lobe mostly white or yellow, its dorsal margin black. Female FWs with dorsal field infumated black; lateral field ventrally black; M / R area forming a clearly defined white / yellow stripe. This species is most similar to N. danum sp. nov. from Borneo, but differs by the following characters: larger in size, vertex black with cream-coloured margin around eyes (instead of yellow); frons and mouthparts dark red brown to black; frons black with a cream-coloured band ventral to eyes (instead of completely yellow). This species also differs from the sympatric N. insignis by frons mostly black (instead of yellow); lateral field M / R area forming a clearly defined white stripe (instead of dark); female FW dorsal field with veins black (instead of yellow) and juvenile colouration.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFCCFFD8FDFAF5B4FEECF9E2.taxon	etymology	Etymology The new species is named after Prof. Hugh Tiang Wah Tan, a renowned botanist and conservation biologist from Singapore and the PhD supervisor of M. K. Tan.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFCCFFD8FDFAF5B4FEECF9E2.taxon	materials_examined	Material examined Holotype INDONESIA – Sumatra • ♀; “ Nambaru, Kec. Bundar Pasir Mandage, Kab. Asahan ”; 2 ° 44 ′ 24.17 ″ N, 99 ° 15 ′ 02.52 ″ E; 980 m a. s. l.; 5 Aug. 1983; Yusup leg.; MZB. ORTH. 12193. Paratypes INDONESIA – Sumatra • 1 ♀; same collection data as for holotype; (MZB. ORTH. 12192); MNHN- EO-ENSIF 10946 • 1 ♀; “ Sarapai ”; “ 8 July ” 1916; Edward Jacobson leg.; RMNH • 1 ♂; 20 km east of Sasak, environment of Harau Valley National Park; 0 ° 04 ′ 51.79 ″ S, 100 ° 39 ′ 08.13 ″ E; 24 – 26 Nov. 1999; A. V. Gorochov leg.; PIII: N 34 Nmb 1; ZIN • 1 ♀; Deli; 3 ° 19 ′ N, 98 ° 35 ′ E; 12 Oct. 1895; W. Burchard leg.; ZMH • 1 juvenile; Batu-Seri (Medan.); 3 ° 35 ′ 10 ″ N, 98 ° 40 ′ 19 ″ E; 10 Mar. 1900; B. Jachan leg.; ZMH • 1 juvenile; 20 km east of Sasak, environment of Harau Valley National Park; 0 ° 04 ′ 51.79 ″ S, 100 ° 39 ′ 08.13 ″ E; 24 – 26 Nov. 1999; A. V. Gorochov leg.; ZIN • 1 ♀; 1877 – 1878; RMNH. Additional specimens INDONESIA – Sumatra • 1 juvenile; Panti, Pasaman Regency, West Sumatra; 0 ° 20 ′ 45.7 ″ N, 100 ° 3 ′ 56.4 ″ E; 25 Jan. 2019; S. B. Shakya leg.; iNaturalist. org / observations / 20129372 • 1 ♀; Sungai Penuh City, Jambi; 2 ° 05 ′ 58.5 ″ S, 101 ° 15 ′ 57.8 ″ E; 18 Jun. 2019; S. Wollney leg.; iNaturalist. org / observations / 27481224 • 1 ♀; Lempur Tengah, Gunung Raya, Kerinci Regency, Jambi; 2 ° 19 ′ 24.2 ″ S, 101 ° 32 ′ 11.1 ″ E; 12 Nov. 2019; T. Sanders leg.; iNaturalist. org / observations / 36674976 • 1 juvenile; Kerinci, Jambi; 2 ° 15 ′ 12.6 ″ S, 101 ° 34 ′ 17.0 ″ E; 12 Aug. 2019; pmaaskant leg.; iNaturalist. org / observations / 37984028 • 1 juvenile; Aceh; 3 ° 36 ′ 47.6 ″ N, 97 ° 26 ′ 23.8 ″ E; 14 Apr. 2009; G. Bohne leg.; iNaturalist. org / observations / 27198231 • 1 juvenile; Aceh; 3 ° 36 ′ 55.2 ″ N, 97 ° 34 ′ 33.3 ″ E; 14 Apr. 2009; G. Bohne leg.; iNaturalist. org / observations / 27198588 • 1 juvenile; Aceh; 3 ° 46 ′ 11.5 ″ N, 97 ° 31 ′ 57.7 ″ E; 14 Apr. 2009; G. Bohne leg.; iNaturalist. org / observations / 27198509 • 1 ♀; Aceh, Gunung Leuser National Park, Gurah; 3 ° 46 ′ 15 ″ N, 97 ° 14 ′ 25.9 ″ E; 16 Feb. 2014; Michael Kesl leg.; https: // www. biolib. cz / en / image / id 334434 / • 1 ♀; Aceh, Gunung Leuser National Park, Gurah; 3 ° 46 ′ 15 ″ N, 97 ° 14 ′ 25.9 ″ E; 16 Feb. 2014; Michael Kesl leg.; https: // www. biolib. cz / en / image / id 334435 / • 1 juvenile; Aceh, Gunung Leuser National Park, Gurah; 3 ° 46 ′ 15 ″ N, 97 ° 14 ′ 25.9 ″ E; 16 Feb. 2014; M. Kesl leg.; https: // www. biolib. cz / en / image / id 334438 / • 2 juveniles; Aceh, Gunung Leuser National Park, Gurah; 3 ° 46 ′ 15 ″ N, 97 ° 14 ′ 25.9 ″ E; 16 Feb. 2014; M. Kesl leg.; https: // www. biolib. cz / en / image / id 334439 / • 1 ♀; West Sumatra, Gunung Talakmau camp 1; 0 ° 05 ′ 50.1 ″ N, 99 ° 56 ′ 50.8 ″ E; 15 Mar. 2018; O. Johnson leg.; https: // www. inaturalist. org / observations / 11085385.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFCCFFD8FDFAF5B4FEECF9E2.taxon	description	Description Habitus large and slender, colouration very contrasting with black, white and vivid yellow when alive (Figs 7 C, 20 A – B). Vertex black with cream-coloured margin around eyes (Fig. 11 D). Fastigium velvety black with cream-coloured margins. Scapes dark red brown to black (particularly on the dorso-inner margins). Antennae black with wide white rings. Frons and mouthparts dark red brown to black, frons cream-coloured (whitish when alive) ventral to eyes (Fig. 10 D). Maxillary palpi cream-coloured; apical segment black apically (Fig. 10 D). Head lateral side dark coloured, with broad cream-coloured band around eyes extending to area behind eyes (Fig. 12 D). Pronotum covered with white setae; dorsal disk velvety black (Fig. 11 D). Lateral lobes of pronotum with dorsal-third black, cream-coloured ventrally, vivid yellow when alive (Fig. 12 D). Legs dark brown. FIII brown, knees dark brown to black; TIIIs brown with black spines and spurs, dark brown to black near distal end; tarsomeres dark brown to black. Hindwings hyaline brown apically, longer than FWs, the dark brown tail exceeding FW more than twice as long as pronotum. Tergites red brown, with a pale band laterally; sternites pale, with a red brown median area. Cerci red brown. Male FOREWINGS. FW colouration (Fig. 13 C): dorsal field cells mostly transparent, veins mostly dark brown to black, cream-coloured in apical field. Basal area brown, basally cream-coloured, brown towards distal part. R cream-coloured, Sc projections cream-coloured basally, black apically. Lateral field brown. FW venation (Fig. 13 C): 1 A curved, slightly angulated. Harp slightly longer than wide, with 3 oblique veins, basal one bifurcate distally, distal one bifurcate basally. c 1 long and wide, c 2 diamond-shaped; mirror (d 1) longer than wide, not rounded, generally separated into two parts by a distinct transverse vein, the posterior part hexagonal, shorter than anterior part. Cell d 2 narrower than d 1, more or less rectangular. Apical field short and rounded, with 3 wide cell alignments posterior to mirror and a narrow apical alignment. Lateral field with 6 projections of Sc. Epiproct black. Subgenital plate pale cream-coloured with median area, anterior and posterior margins black. GENITALIA (Fig. 16 C). Pseudepiphallus sclerotized, large and slender, anterior margin narrow and straight, posterior margin also straight, lateral margins distinctly converging posteriorly. Posterior apex with paired lophi longer than wide (at base), subacute at apex, sclerotized laterally only and covered with short strong setae, narrowly spaced apart from each other. Rami swollen preapically, somewhat sinuous. Pseudepiphallic parameres narrow, divergent posteriorly sclerotized arm base to parameres distinctly curved laterally. Ectophallic arc transverse, complete, with anterior and posterior margins straight. Ectophallic fold strong rounded lateral sclerites appearing bean-shaped. Ectophallic apodemes long, straight and slightly divergent. Endophallic sclerite large and sclerotized, its posterior apex with divergent lateral arms and with a short median expansion curved dorsally. Endophallic apodeme with lateral lamellae and dorsal crest only slightly longer than wide. Female FOREWINGS. FW colouration (Fig. 14 D): cells black, veins generally black. Longitudinal white stripe around region of CuA, M and R with a distinct margin against black colouration on dorsal field; Sc projections cream-coloured basally, black apically. Lateral field dark. FW venation: 6 – 7 strong longitudinal veins on dorsal field; lateral field with 6 projections of Sc. GENITALIA. Ovipositor: distinctly longer than FIII. Copulatory papilla conical, large (similar to N. brunnerianus but less slender — possibly due to preservation of specimen); apex folded ventrally, elongated like in N. brunnerianus, pointed; dorsal face with a sclerotized area, but less sclerotized than in congeners; ventro-anterior end forming a somewhat triangular rim (Fig. 17 F – G). Juvenile (Fig. 20 C – E) Early instars with a pale colouration with numerous dark stripes all around body. Head generally reddish. Eyes with numerous dark stripes. Pronotum with three thick, dark longitudinal stripes on dorsal disk, one very thick and velvety black stripe between dorsal disk and lateral lobe; another curved stripe on lateral lobe ventral of the thick stripe; stripes never reaching the margins. Fore and middle legs with longitudinal dark stripes. FIII also with longitudinal dark stripes, two dorsally, one on each side; and two ventrally, one on each side. Thoracic (excluding pronotum) and abdominal tergites typically dark or greyish, anterior margin darkest, posterior margin whitish. Cerci whitish with a black stripe. Measurements See Table 6.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFCCFFD8FDFAF5B4FEECF9E2.taxon	biology_ecology	Ecology Unknown.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFCCFFD8FDFAF5B4FEECF9E2.taxon	distribution	Distribution Northern Sumatra.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFCCFFD8FDFAF5B4FEECF9E2.taxon	materials_examined	Type locality Indonesia, northern Sumatra.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFCCFFD8FDFAF5B4FEECF9E2.taxon	description	Calling song Unknown.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFC8FFDAFDB5F53BFEE5FE03.taxon	description	Fig. 21; Table 2	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFC8FFDAFDB5F53BFEE5FE03.taxon	materials_examined	Type locality Borneo: Kalimantan, “ Benjermassing ” [Banjarmasin]. Type details Syntypes, RMNH (missing).	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFC8FFDAFDB5F53BFEE5FE03.taxon	discussion	Remarks The syntypes in RMNH are missing. L. Chopardʼs records (Chopard 1929) of this species in Mentawai, North Pagi, Siberut and Padang (all part of or related to Sumatra, Indonesia) correspond to specimens of N. insignis, while N. hyalinus is known only from Borneo. We cannot revise this species owing to the absence of type material and of other specimens from the region of the type locality in the south of Borneo. Nonetheless, we have found images of specimens from Sungai Utik, Embaloh Hulu, Kapuas Hulu Regency, West Kalimantan, albeit not exactly near the type locality in Banjarmasin in South Kalimantan, which could possibly correspond to N. hyalinus. These photographs (Fig. 21) match the description by Saussure (1878): “ Les élytres latéralement hyalins dans leur partie marginale, dʼun noir opaque luisant dans leur partie médiastino-humérale; le champ dorsal brunâtre, réticulé en jaune par mailles plus grandes que chez le N. vittatus. Ailes assez courtes, hyalines, nébuleuses, surtout le long du limbe. Fémurs postérieurs très longs; leur moitié apicale linéaire, comme chez le N. insignis ”. [The elytra laterally hyaline in their marginal part, a shiny opaque black in their medial-humeral part; the dorsal field brownish, reticulated in yellow with larger meshes than in N. vittatus. Wings rather short, hyaline, nebulous, especially along the limbus. Very long FIII; their apical half linear, as in N. insignis.] We can also confirm that none of the Bornean specimens examined in this study match the description of N. hyalinus. Saussure (1878) repeatedly compared N. hyalinus with N. vittatus, but not with other Bornean species which differ distinctly from N. vittatus (i. e., N. brunnerianus). Nisitrus danum sp. nov. differs from N. hyalinus by dorsal field clearly not brownish; lateral field humeral stripe whitish, not shining black; lateral field is not hyaline either. Furthermore, the type locality of N. hyalinus from Banjarmasin in southern Kalimantan is quite far from Danum Valley and of a different forest type (Danum Valley is lowland forest, whereas Banjarmasin would have been peat swamp forest). Nisitrus rindu sp. nov. also differs by medial-humeral part not shining black opaque. Given the fact that very few specimens of Nisitrus are known from Southern Borneo, it is likely that N. hyalinus is a valid species but rarely collected.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFCAFFD6FDBAF15CFABDF811.taxon	description	Figs 2 A, 5, 7 D, 8 F, 10 E, 11 E, 12 E, 13 D, 14 E, 16 D, 17 H – I, 22; Tables 1 – 2, 7	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFCAFFD6FDBAF15CFABDF811.taxon	diagnosis	Diagnosis Of smaller size but stouter habitus than congeners. Vertex coloration ranging from black with creamcoloured margins around eyes and fastigium to cream-coloured with a strong longitudinal black band and one or two faint transverse black band (forming a T- or a cross-shaped black pattern). FIIIs brown, with a brown longitudinal stripe along external ventral half; sometimes dorsal margin with a paler longitudinal stripe. Dorsal field of male FW always with cream-coloured triangular (sometimes rounded) patch, with very distinctly different venation; harp very small, slightly longer than wide, with one distinct straight oblique vein. This species is similar to N. malaya sp. nov. and N. vittatus but differs by legs with lighter colouration and dorsum of head yellow with variable black pattern (instead of entirely black). This species also differs from the sympatric N. hughtani sp. nov. by smaller size, frons yellow (black in N. hughtani sp. nov.) and vertex colouration pattern; FW lateral field M / R area dark (instead of forming a clearly defined white / yellow stripe), and female FW dorsal field with veins yellow (instead of mostly black).	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFCAFFD6FDBAF15CFABDF811.taxon	materials_examined	Material examined Lectotype (here designated) INDONESIA – Sumatra • ♀; “ Neit Suma ” [North Sumatra]; 1853; Mus. Caes. Vindobon; NMW. Additional specimens INDONESIA – Sumatra • 1 ♂ (holotype of N. sumatrensis); R. Weber leg.; No. 20486; AMNH • 1 ♂; “ Mentawei ” [Siberut] “ Siberoet ”; 1 ° 23 ′ 17.3 ″ S, 98 ° 56 ′ 7.7 ″ E; 22 Sep. 1924; H. H. Karny leg.; MNHN • 2 ♂♂; same locality data as for preceding; 8 Sep. 1924; H. H. Karny leg.; MNHN • 2 ♀♀; same locality data as for preceding; 8 Sep. 1924; H. H. Karny leg.; identified as Nisitra hyalina by L. Chopard; MNHN • 3 ♀♀; same locality data as for preceding; 12 Sep. 1924; H. H. Karny leg.; MNHN • 3 ♀♀; same locality data as for preceding; Sep. 1924; C. B. K. and N. S. leg.; MNHN • 3 ♂♂; same locality data as for preceding; Sep. 1924; C. B. K. and N. S. leg.; MNHN; • 1 ♂; “ Mentawei ” Sipora Island; 2 ° 13 ′ 28.7 ″ S, 99 ° 40 ′ 9.4 ″ E; Oct. 1924; C. B. K. and N. S. leg.; identifed Nisitrus brunnerianus by T. Robillard, 2004; MNHN-EO-ENSIF 1748 • 1 ♀; same collection data as for preceding; MNHN • 1 ♀; “ Mentawei ” [Siberut] “ Siberoet ”; 1 ° 23 ′ 17.3 ″ S, 98 ° 56 ′ 7.7 ″ E; Sep. 1924; C. B. K. and N. S. leg.; ZRC • 1 ♂; same collection data as for preceding; PII: N 55; ZRC • 2 ♀♀; Sibolangit; 3 ° 18 ′ 2 ″ N, 98 ° 34 ′ 8 ″ E; Sep. – Oct. 1929; D. V. Leeuwen leg.; MNHN • 2 ♂♂; same collection data as for preceding; MNHN • 2 ♀♀; Sibolangit; 3 ° 18 ′ 2 ″ N, 98 ° 34 ′ 8 ″ E; 8 Oct. 1925; Fulmek and H. Karny leg.; • 2 ♀♀; 20 km east of Sasak environs of Harau Valley National Park; 0 ° 4 ′ 57 ″ S, 100 ° 39 ′ 12 ″ E; 600 m a. s. l.; 24 – 26 Nov. 1999; A. V. Gorochov leg.; ZIN • 3 ♂♂; same collection data as for preceding; ZIN • 1 ♀; Pematang Siantar “ Pem. Siantar ”, Research Centre? “ Prapat Hutan Penelitian ”; 10 – 1100 m a. s. l.; 10 Mar. 1993; S. Ingrisch leg.; SMTD • 1 ♂; same collection data as for preceding; SMTD • 2 ♂; Alas Valley, Belelutu; 3 ° 43 ′ N, 97 ° 38 ′ E; ca 320 m a. s. l.; 3 – 8 Aug. 1972; J. Krikken leg.; RMNH • 1 ♀; Alas Valley, vicinity of Gumpang; 3 ° 48 ′ N, 97 ° 29 ′ E; 13 Jun. 1972; J. Krikken leg.; RMNH • 1 ♀; Deli Sibolangit Nature Reserve; 3 ° 19 ′ N, 98 ° 35 ′ E; 500 m a. s. l.; 29 Jul. 1972; J. Krikken leg.; RMNH • 1 ♂; Soban Ajam; Jul. 1916; E. Jacobson leg.; RMNH • 2 ♀♀; Kuala Simpang, lowland forest; 4 ° 17 ′ 3.7 ″ N, 98 ° 3 ′ 29 ″ E; Jan. 1954; A. Sollaart leg.; RMNH • 1 ♀; Serdang, Tanjong Morawai; 1882; B. Hagen leg.; RMNH • 2 ♀♀; Sumatra Expedition 1877 – 1878 leg.; RMNH • 1 ♀; Ketambe, at Jeb Tenggara; 3 ° 41 ′ 6 ″ N, 97 ° 44 ′ 57 ″ E; 350 m a. s. l.; 18 Nov. 1973; H. D. Rijksen leg.; RMNH • 1 ♂; Medan; 3 ° 35 ′ 10 ″ N, 98 ° 40 ′ 19 ″ E; 20 Jun. 1949; Kalshoven leg.; RMNH • 1 ♂; 1908; E. Jacobson leg.; RMNH • 1 ♂; Padang Panjang, Singalang; 0 ° 54 ′ 9 ″ S, 100 ° 23 ′ 16 ″ E; 700 m a. s. l.; 15 Feb. 1995; H. Deumer and M. Schaarschmidt leg.; N 5; MNHN-EO-ENSIF 4180 • 2 ♀♀; 2 km from Padang Panjang (Taglang) [Talang?]; “ 3.9.1991 ”; Wolfram Guidatti leg.; MNHN • 2 ♂♂; [Lubu?] “ Lubu Sampin ”; 19 Jan. 1885; MHN • 1 ♀; [Lubu?] “ Lubu Sampin ”; 2 Dec. 1884; MHN • 2 ♀♀; Padang in Batangproepoe “ Pad. Bovenl ”; 0 ° 54 ′ 9 ″ S, 100 ° 23 ′ 16 ″ E; Nov. 1924; C. B. K. and N. S. leg.; ZRC • 1 ♂; Jambi “ Djambi ”; O. Posthumus leg.; MNHN • 1 ♂; “ Habinsaran Tangga ”; 2 Aug. 1928; J. C. v. d. MeerMohr leg.; MNHN • 1 ♀; same collection data as for preceding; MNHN • 1 ♂; Fort de Kock; 0 ° 18 ′ 1 ″ S, 100 ° 22 ′ 8 ″ E; 920 m a. s. l.; 1924; E. Jacobson leg.; MNHN • 1 juvenile; same collection data as for preceding; MNHN • 1 ♀; same collection data as for preceding; MNHN • 1 ♂; Soilak Daras, Korinchi Valley; 1 ° 41 ′ 12 ″ S, 101 ° 13 ′ 46 ″ E; 3100 ft a. s. l.; Mar. 1914; NHMUK • 1 ♀; Lut [“ Laut ″] Tawar; 4 ° 37 ′ 36 ″ N, 96 ° 51 ′ 4 ″ E; 1400 m a. s. l.; 23 Dec. 1929; H. T. Pagdan leg.; NHMUK • 1 ♂; Dohrn leg.; identified as Nisitrus hyalinus by L. Chopard; MNHN • 1 ♀; Dohrn leg.; MNHN • 1 ♀; Padang in Batangproepoe “ Pad. Bovenl ”; 0 ° 54 ′ 9 ″ S, 100 ° 23 ′ 16 ″ E; 9 Aug. 1920 “ 20 ”; MNHN • 1 ♂; same locality data as for preceding; MNHN • 3 ♀♀; same locality data as for preceding; MNHN • 1 ♂; Medan; 3 ° 35 ′ 10 ″ N, 98 ° 40 ′ 19 ″ E; Mjöberg leg.; GNME • 1 ♀; Dolok Baros; 1906; “ Maindon ” leg.; MNHN • 1 ♀; Medan Batu-Seri; 3 ° 35 ′ 10 ″ N, 98 ° 40 ′ 19 ″ E; 10 Mar. 1900; B. Jachan leg.; ZMH • 1 ♀; Deli; 3 ° 19 ′ N, 98 ° 35 ′ E; 3 Jan. 1896; Gebr. Siemssen leg.; ZMH • 3 ♂♂; Deli [Bindjai] “ Bindjey-Estate ”; 3 ° 36 ′ 44 ″ N, 98 ° 29 ′ 20 ″ E; 28 Apr. 1894; W. Burchard leg.; ZMH • 1 ♀; same collection data as for preceding; ZMH • 1 ♂; A. Grubauer leg.; ZMH • 1 ♂; “ Anei Kloof ”; 500 m a. s. l.; 1925; E. Jacobson leg.; ZMH • 1 ♀; Fort de Kock; 0 ° 18 ′ 1 ″ S, 100 ° 22 ′ 8 ″ E; 920 m a. s. l.; 1925 / 1926?; ZMH • 1 ♂; “ Deli ”; 3 ° 19 ′ N, 98 ° 35 ′ E; MNHN • 1 ♀; same locality data as for preceding; MNHN • 1 ♂; v. W. leg.; MNHN • 1 ♀; v. W. leg.; MNHN • 1 ♂; “ Sumatra ”; MNHN • 1 ♀; “ Sumatra ”; MNHN • 3 ♀♀; Haran Kloof; 0 ° 27 ′ 14.6 ″ S, 101 ° 24 ′ 53.192 ″ E; 23 Apr. 1929; Prince Leopold leg.; RBINS • 1 ♂; same collection data as for preceding; RBINS • 1 ♂; Singalang [Mount Singgalang]; 0 ° 24 ′ 36 ″ S, 100 ° 19 ′ 51 ″ E; 23 Apr. 1929; Prince Leopold leg.; RBINS • 1 ♂; Medan S. O. K. [Deli River]; 3 ° 35 ′ 42.7 ″ N, 98 ° 40 ′ 20.0 ″ E; Fulmek L. leg.; NHMW • 1 ♂; Labuhan Batu; 2 ° 0 ′ 4.8 ″ N, 99 ° 53 ′ 54.1 ″ E; 22 Sep. 2020; K. S. Zebua leg.; iNaturalist. org / observations / 60403336 • 1 ♂; Labuhan Batu; 2 ° 10 ′ 42.9 ″ N, 99 ° 57 ′ 25.3 ″ E; 6 Oct. 2020; K. S. Zebua leg.; iNaturalist. org / observations / 61849044 • 1 ♂; Labuhan Batu; 2 ° 08 ′ 7.3 ″ N, 99 ° 52 ′ 59.2 ″ E; 6 Oct. 2020; K. S. Zebua leg.; iNaturalist. org / observations / 61849074 • 1 ♂; Kong Paluh, Kutapanjang, Gayo Lues Regency, Aceh; 3 ° 48 ′ 16.0 ″ N, 97 ° 14 ′ 6.9 ″ E; 20 Jul. 2018; G. Jarasunas leg.; iNaturalist. org / observations / 20738568 • 1 ♀; Kerinci, Jambi; 2 ° 14 ′ 54.4 ″ S, 101 ° 32 ′ 21.3 ″ E; 12 Aug. 2018; pmaaskant leg.; iNaturalist. org / observations / 37307112 • 1 ♂; “ Ланкат, СевернаЯ Суматра, ИндонеЗИЯ ” [Langkat, North Sumatra, Indonesia]; 3 ° 33 ′ 6.3 ″ N, 98 ° 6 ′ 37.2 ″ E; 3 May 2019; reflectitur _ photons leg.; iNaturalist. org / observations / 25730568 • 1 ♀; Lima Puluh Koto, ID-SB, ID; 0 ° 6 ′ 39.0 ″ S, 100 ° 40 ′ 15.2 ″ E; 6 Jun. 2011; D. Nys and M. Vancauwenbergh leg.; iNaturalist. org / observations / 34083286 • 1 ♀; Lubuk Landua; 0 ° 6 ′ 19.5 ″ N, 99 ° 53 ′ 12.3 ″ E; 13 Mar. 2018; O. Johnson leg.; iNaturalist. org / observations / 11018930 • 1 ♂; Gayo Lues Regency, Aceh; 3 ° 52 ′ 51.7 ″ N, 97 ° 19 ′ 23.1 ″ E; 21 May 2010; ninjawil leg.; iNaturalist. org / observations / 9181395. MALAYSIA • 4 ♀♀; “ Camp Jor ”, border between Perak and Pahang; 4 ° 9 ′ 43 ″ N, 102 ° 22 ′ 20 ″ E; 30 Dec. 1902; Alb. Grubauer leg.; ZMH.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFCAFFD6FDBAF15CFABDF811.taxon	description	Redescription Slightly smaller than other species (Fig. 7 D). Vivid yellow, brown and black when alive (Fig. 22 A – C). Vertex variable: ranging from black with yellow margins around eyes (mostly in individuals from Sumatra) to cream-coloured with a strong longitudinal black band and one or two faint transverse black bands (forming a T- or cross-shaped black pattern; mostly in individuals from Siberut Islands and Malaysia) (Fig. 11 E); dark pattern variable between individuals, some have a narrow pattern appearing more like a longitudinal band. Scapes red brown to black. Antennae red brown (individuals from Siberut) to black (individuals from Sumatra), with wide whitish rings. Frons yellow, cream-coloured to yellow, face part of fastigium brown to dark brown or black between eyes, lighter for individuals from Siberut than from Sumatra (Fig. 10 E). Mouthparts dark brown to black in specimens from Sumatra, lighter or sometimes yellow in specimens from Siberut (Fig. 10 E). Maxillary palpi cream-coloured; apical segment black apically (Fig. 10 E). Head lateral side yellow to cream-coloured, including gena, with a black band posterior to eyes (Fig. 12 E). Pronotum dorsal disk black, covered with white setae, rectangular with lateral margin barely widening posteriorly, posterior margin substraight to faintly bisinuous (Fig. 11 E). Lateral lobes of pronotum dorsal half black, vivid yellow ventrally (cream-coloured in preserved specimens) (Fig. 12 E). Legs generally light grey brown, sometimes with some small dark spots. FIIIs brown, with a brown longitudinal stripe along ventral half (more distinct and darker in individuals from Sumatra); sometimes dorsal margin with a paler longitudinal stripe; knees dark brown to black. TIIIs brown with black spines and spurs, dark brown to black near distal end; tarsomeres dark brown to black. Hindwings hyaline brown apically; longer than FWs and surpassing cerci, forming a black tail with yellow veins exceeding FWs for less than twice the length of the pronotum. Tergites red brown, with a pale band laterally; sternites pale, sometimes with a dark brown median area. Subgenital plate with a dark median area. Male FOREWINGS. FW colouration (Fig. 13 D): dorsal field narrow, cells mostly transparent, veins varying from pale yellow to black. Basal area either infumated brown to black (individuals from Sumatra) or almost transparent (individuals from Siberut), but always with a cream-coloured triangular patch (sometimes more rounded). Chords most often with a white patch, most readily observed in individuals from Sumatra. R black, Sc projections black basally, cream-coloured or transparent apically. Lateral field transparent. FW venation (Fig. 13 D): 1 A curved, slightly angulated. Harp very small, slightly longer than wide, with one distinct straight oblique vein. c 1 long and wide, broader basally; c 2 diamond-shaped; mirror (d 1) small, longer than wide, not rounded, generally separated into two parts by a transverse vein, the anterior part triangular, about as long as posterior part, posterior part rectangular. Cell d 2 about as wide as d 1, usually subdivided by accessory veins. Apical field short, including one cell alignment posterior to mirror and a narrow apical alignment; its veins cream-coloured. Lateral field transparent, with 5 or 6 projections of Sc. Epiproct black. Subgenital plate with a dark median area. Cerci red brown. GENITALIA (Fig. 16 D). Pseudepiphallus sclerotized, stout (medium-sized but broad) compared to congeners, anterior margin somewhat broad and straight, posterior margin also straight, lateral margins faintly converging posteriorly. Posterior apex with paired lophi slightly longer than wide (at base), obtuse at apex, moderately spaced apart from one another. Rami swollen preapically, anterior apex somewhat truncated. Pseudepiphallic parameres narrow. Ectophallic fold rounded lateral sclerites appearing beanshaped to rectangular. Endophallic sclerite wide and rounded laterally, with relatively long lateral arms. Female FOREWINGS. FW colouration (Fig. 14 E): base of dorsal field with a yellow area; cells black, veins mostly vivid yellow (cream-coloured in preserved specimens). Region of CuA, M and R black, including the veins. Sc projections yellow basally, black apically. Lateral field transparent. FW venation: 6 – 7 strong longitudinal veins on dorsal field; lateral field with 6 projections of Sc. GENITALIA. Ovipositor slightly longer than FIII. Copulatory papilla conical, smaller and stout; apex folded ventrally, stout, broader than congeners, pointed; dorsal face with a sclerotized area, forming a somewhat triangular rim (Fig. 17 H – I). Juvenile (Fig. 22 D – E) Young instar with head nearly completely cream-coloured. Pronotum disk velvety black, lateral lobe with dorsal half velvety black and ventral half cream-coloured. Legs generally cream-coloured with small dark spots; tarsi darker. Abdomen with tergites velvety black dorsally, with sternites cream-coloured ventrally. Epiproct cream-coloured. Later instar resembles adult colouration. Measurements See Table 7.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFCAFFD6FDBAF15CFABDF811.taxon	biology_ecology	Ecology Habitat and mating behaviour, including multiple mating, male guarding behaviour, female eating spermatophores and male-male competition for females, were described in detail by Preston-Mahfam (2000).	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFCAFFD6FDBAF15CFABDF811.taxon	distribution	Distribution Sumatra, Malay Peninsula (Malaysia).	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFCAFFD6FDBAF15CFABDF811.taxon	materials_examined	Type locality Indonesia: Sumatra.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFCAFFD6FDBAF15CFABDF811.taxon	description	Calling song Unknown.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFCAFFD6FDBAF15CFABDF811.taxon	discussion	Remarks One syntype of Saussure from Northern Sumatra found in NMW is designated here as the lectotype of the species. Based on comparisons with the type of N. sumatrensis, the latter name is considered as a junior synonym of N. insignis (new synonymy, see below).	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFCAFFD6FDBAF15CFABDF811.taxon	description	The observed specimen series vary greatly in size and colouration, which made us hesitate between defining one or more species, in particular for the lighter specimens from the Mentawai Islands. To remain conservative, we considered these differences as species variation only, but more information about molecular divergence and acoustic parameters may lead to revise this decision in future studies.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFC7FFC4FDC6F26CFAB7FE1B.taxon	description	urn: lsid: zoobank. org: act: AD 6647 DB- 81 AE- 421 B- 8433 - C 4 E 3 D 9 ACC 314 Figs 4 – 5, 7 E, 9 A, 10 F, 11 F, 12 F, 13 E, 14 F, 16 E, 17 J – K, 23 A – D, 24 A – C, 25 A – C, 26; Tables 1 – 2, 8	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFC7FFC4FDC6F26CFAB7FE1B.taxon	diagnosis	Diagnosis Average size, colouration contrasting with black and yellow, characterised by black vertex, yellow frons and orange brown legs. This new species is very close to N. vittatus from which it differs by adult and juvenile colouration, male and female genitalia and calling song. General colouration darker, particularly among FW cells and veins beneath R; R black (instead of cream-coloured as in N. vittatus); in male venation, anal-most and middle chords narrow and nearly parallel with one another (instead of diverging apically), and basal-most chord extend from basal half of middle chord (instead of at / after midpoint). It differs from N. vittatus by the male genitalia with pseudepiphallus more robust, broad, anterior margin more strongly concave. Female copulatory papilla with truncated lobule at the ventro-anterior end. Juvenile differs strikingly in the body colour pattern. Male calling song also differs by longer echeme period. This species is also similar to N. crucius sp. nov. in size and habitus, but differs by the absence of cross-shaped black pattern on frons and does not occur in Borneo.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFC7FFC4FDC6F26CFAB7FE1B.taxon	etymology	Etymology This specific name is a noun in apposition referring to the speciesʼ distribution in the Malay Peninsula (including Singapore and Peninsular Malaysia).	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFC7FFC4FDC6F26CFAB7FE1B.taxon	materials_examined	Material examined Holotype SINGAPORE • ♂; Dairy Farm Nature Park; 1 ° 21 ′ 44 ″ N, 103 ° 46 ′ 9 ″ E; 17 Jul. 2020; M. K. Tan leg.; ZRC. Paratypes MALAYSIA – Selangor • 1 ♂; Petaling Jaya, Mount Kiara; 3 ° 10 ′ 17.44 ″ N, 101 ° 38 ′ 42.70 ″ E; 7 Sep. 2002; T. Robillard leg.; MNHN-EO-ENSIF 3132 • 1 ♂; same collection data as for preceding; call recording; MNHN-EO-ENSIF 3133 • 1 ♂; same collection data as for preceding; MNHN-EO-ENSIF 3134 • 1 ♀; same collection data as for preceding; MNHN-EO-ENSIF 11140 • 1 ♀; same locality data as for preceding; Aug. 2002; T. Robillard leg.; MNHN-EO-ENSIF 11331 • 1 ♀; same locality data as for preceding; T. Robillard leg.; PIII: Nvith 2; MNHN-EO-ENSIF 1708 • 1 ♂; 1909; C. J. Brooks leg.; NHMUK. SINGAPORE • 1 ♂; Bukit Timah, “ près de l’entrée de la réserve ” [near entrance]; 1 ° 21 ′ 6.7 ″ N, 103 ° 46 ′ 45.2 ″ E; 92 m a. s. l.; 10 Jun. 2011; T. Robillard leg.; call recording; MNHN-EO-ENSIF 3136 • 1 ♂; Bukit Timah, “ sommet ” [summit]; 1 ° 21 ′ 16.2 ″ N, 103 ° 46 ′ 35.9 ″ E; 170 m a. s. l.; 29 Jun. 2009; T. Robillard leg.; call recording MNHN-SO- 2016 - 41; TR 9; MNHN-EO-ENSIF 3102 • 1 ♂; same locality data as for preceding; 10 Jun. 2011; T. Robillard leg.; TR 1; MNHN-EO-ENSIF 10948 • 1 ♂; Bukit Timah Nature Reserve, Hindhede Trail; 1 ° 20 ′ 57.1 ″ N, 103 ° 46 ′ 45.2 ″ E; 68 m a. s. l.; 20 Jun. 2011; T. Robillard leg.; TR 26; MNHN-EO-ENSIF 10949 • 1 ♂; MacRitchie Reserve; 1 ° 21 ′ 10.2 ″ N, 103 ° 46 ′ 33.6 ″ E; 68 m a. s. l.; 2 Jul. 2009; T. Robillard leg.; call recording; MNHN-EO-ENSIF 3135 • 1 ♂; MacRitchie Reserve; 1 ° 21 ′ 10.2 ″ N, 103 ° 46 ′ 33.6 ″ E; 68 m a. s. l.; 2 Jul. 2009; T. Robillard leg.; call recording; MNHN-EO-ENSIF 2742 • 1 ♀; Mandai; 1 ° 24 ′ 52.7 ″ N, 103 ° 47 ′ 22.2 ″ E; 27 Dec. 2015; M. K. Tan and H. Yeo leg.; C 2. F. 50; MNHN-EO- ENSIF 11136 • 1 ♀; same collection data as for preceding; C 2. F. 51; MNHN-EO-ENSIF 11138 • 1 ♀; same collection data as for preceding; C 2. F. 28; MNHN-EO-ENSIF 11137. Additional specimens MALAYSIA • 6 juveniles; Petaling Jaya, Mount Kiara; 3 ° 10 ′ 17.44 ″ N, 101 ° 38 ′ 42.70 ″ E; 7 Sep. 2002; T. Robillard leg.; MNHN • 1 juvenile; Kepong FRIM (Forest Research Insitute of Malaysia); 3 ° 13 ′ 60 ″ N, 101 ° 37 ′ 51 ″ E; 18 Aug. 2003; Svenson leg.; MNHN • 16 ♂♂; Petaling Jaya, Mount Kiara; T. Robillard leg.; élevage; MNHN • 11 ♀♀; same collection data as for preceding; MNHN • 1 ♀; PD Villa Country Resort, 11, Jln PDV 3, Taman Rakyat, 71050 Teluk Kemang, Negeri Sembilan; 2 ° 27 ′ 27.3 ″ N, 101 ° 52 ′ 5.9 ″ E; 17 Feb. 2019; J. Howes leg.; iNaturalist. org / observations / 20414064 • 1 ♀; Bestari Jaya, Selangor; 3 ° 16 ′ 7 ″ N, 101 ° 27 ′ 28.6 ″ E; 27 Apr. 2019; S. Siva Perumal leg.; iNaturalist. org / observations / 24568214 • 1 ♂; Kuala Lumpur; 3 ° 7 ′ 44.8 ″ N, 101 ° 39 ′ 3.20 ″ E; 5 Mar. 2019; msone leg.; iNaturalist. org / observations / 30373276 • 1 ♂; Kuala Lumpur; 3 ° 7 ′ 44.8 ″ N, 101 ° 39 ′ 3.20 ″ E; 5 Mar. 2019; msone leg.; iNaturalist. org / observations / 30373274 • 1 ♂; Kuala Lumpur; 3 ° 7 ′ 56.5 ″ N, 101 ° 39 ′ 16.3 ″ E; 9 Jan. 2019; msone leg.; iNaturalist. org / observations / 20052999 • 1 ♀; Kledong Saiong Forest Eco Park, Perak; 4 ° 40 ′ 59.7 ″ N, 101 ° 4 ′ 6.5 ″ E; 25 Jul. 2019; K. Wenyuh leg.; iNaturalist. org / observations / 29510836 • 1 ♀; Bukit Gasing, Petaling Jaya, Selangor; 3 ° 5 ′ 54.3 ″ N, 101 ° 39 ′ 29.1 ″ E; 17 Nov. 2006; J. Howes leg.; iNaturalist. org / observations / 35667227 • 1 ♂; Taman Tugu PT 88, Jalan Sultan Salahuddin, Kuala Lumpur, 50480 Kuala Lumpur Federal Territory of Kuala Lumpur; 3 ° 9 ′ 6.28 ″ N, 101 ° 41 ′ 0.900 ″ E; 17 Nov. 2019; T. Maul leg.; iNaturalist. org / observations / 35769550 • 1 ♀; Semenyih, Selangor; 2 ° 56 ′ 36.2 ″ N, 101 ° 54 ′ 4 ″ E; 3 Dec. 2019; B. C. A. Howes leg.; iNaturalist. org / observations / 36316633 • 1 ♂; Bukit Persekutuan, Kuala Lumpur, Federal Territory of Kuala Lumpur; 3 ° 8 ′ 21.3 ″ N, 101 ° 40 ′ 50.6 ″ E; 11 Jul. 2007; J. Howes leg.; iNaturalist. org / observations / 36663502 • 1 ♀; Lot 240, Jalan Raja Chulan, Kuala Lumpur, 50250 Kuala Lumpur, Wilayah Persekutuan Kuala Lumpur; 3 ° 9 ′ 3.28 ″ N, 101 ° 42 ′ 7.5 ″ E; 6 Jan. 2020; T. Maul leg.; iNaturalist. org / observations / 37332417 • 1 ♀; 31600, Perak; 4 ° 27 ′ 29.1 ″ N, 101 ° 12 ′ 30.5 ″ E; 4 Jan. 2020; M. Ng leg.; iNaturalist. org / observations / 37357322 • 1 ♀; Hutan Lipur Sungai Tua; 3 ° 20 ′ 1.2 ″ N, 101 ° 43 ′ 26.1 ″ E; 6 Oct. 2014; T. Kirschey leg.; iNaturalist. org / observations / 49430837 • 1 ♀; unnamed road, Kampung Padang, 43100 Hulu Langat, Selangor; 3 ° 11 ′ 55.2 ″ N, 101 ° 51 ′ 4 ″ E; 11 Jan. 2020; F. Cheong leg.; iNaturalist. org / observations / 37615383 • 1 ♂; Beranang, Selangor; 2 ° 56 ′ 17.8 ″ N, 101 ° 53 ′ 56.2 ″ E; 19 Jan. 2020; M. A. M. Nasir leg.; iNaturalist. org / observations / 37730365 • 1 ♀; Kampung Tanjung Jelai, Jerantut, Pahang; 4 ° 4 ′ 11.1 ″ N, 102 ° 19 ′ 0 ″ E; 20 Feb. 2019; djhiker leg.; iNaturalist. org / observations / 24363726 • 1 ♂; 5 ° 12 ′ 53.7 ″ N, 100 ° 30 ′ 53.2 ″ E; 19 Mar. 2018; R. Ong leg.; iNaturalist. org / observations / 29697626 • 1 ♂; 53100 Kuala Lumpur, Federal Territory of Kuala Lumpur; 3 ° 13 ′ 53.1 ″ N, 101 ° 44 ′ 25.8 ″ E; 15 Feb. 2020; T. Maul leg.; iNaturalist. org / observations / 38668098 • 1 ♂; Balai Cerap UTM; 1 ° 34 ′ 11.8 ″ N, 103 ° 38 ′ 41 ″ E; 21 Feb. 2020; S. Mierul leg.; iNaturalist. org / observations / 38906538 • 1 ♂; Selangor; 3 ° 35 ′ 39.6 ″ N, 101 ° 44 ′ 5.9 ″ E; 28 Jul. 2019; msone leg.; iNaturalist. org / observations / 39490018 • 1 ♀; Johor Baharu, MY-JH; 1 ° 32 ′ 28.1 ″ N, 103 ° 40 ′ 32.2 ″ E; 11 Mar. 2020; gonaturelam leg.; iNaturalist. org / observations / 39833380 • 1 ♂; 2 ° 50 ′ 40.7 ″ N, 101 ° 56 ′ 32.2 ″ E; 1 May 2019; anukma leg.; iNaturalist. org / observations / 24256308 • 1 juvenile; Taman Tugu PT 88, Jalan Sultan Salahuddin, Kuala Lumpur, 50480 Kuala Lumpur, Federal Territory of Kuala Lumpur; 3 ° 9 ′ 6.28 ″ N, 101 ° 41 ′ 0.900 ″ E; 17 Nov. 2019; T. Maul leg.; iNaturalist. org / observations / 35769588 • 1 ♀; Mersing, Johor; 2 ° 25 ′ 7.56 ″ N, 103 ° 51 ′ 22.8 ″ E; 7 Feb. 2020; msone leg.; iNaturalist. org / observations / 38427233 • 1 juvenile; Tamarind Springs Jalan 1, Taman Tun Abdul Razak, 68000 Ampang Jaya, Selangor; 3 ° 10 ′ 14.6 ″ N, 101 ° 46 ′ 14.3 ″ E; 21 Jun. 2020; T. Maul leg.; iNaturalist. org / observations / 50422192 • 1 juvenile; Selama; 5 ° 13 ′ 6.5 ″ N, 100 ° 41 ′ 36.5 ″ E; 5 Jul. 2020; M. Syarifudin leg.; iNaturalist. org / observations / 51984928 • 1 juvenile; Tamarind Springs Jalan 1, Taman Tun Abdul Razak, 68000 Ampang Jaya, Selangor; 3 ° 10 ′ 14.6 ″ N, 101 ° 46 ′ 14.3 ″ E; 5 Sep. 2020; T. Maul leg.; iNaturalist. org / observations / 58822053 • 1 ♂; Bentong, Pahang, MY; 3 ° 18 ′ 47.2 ″ N, 101 ° 52 ′ 49.6 ″ E; 12 Sep. 2020; J. Marriappan leg.; iNaturalist. org / observations / 59301449 • 1 juvenile; Tamarind Springs Jalan 1, Taman Tun Abdul Razak, 68000 Ampang Jaya, Selangor; 3 ° 10 ′ 14.6 ″ N, 101 ° 46 ′ 14.3 ″ E; 19 Sep. 2020; T. Maul leg.; iNaturalist. org / observations / 60070623 • 1 ♂; Jalan Melawati 5, Taman Zoo View, Selangor, MY; 3 ° 13 ′ 0.400 ″ N, 101 ° 45 ′ 2 ″ E; 2 Sep. 2020; K. J. M. Anbumani leg.; iNaturalist. org / observations / 60166965 • 1 juvenile; Taman Bukit Perdana, 83000 Batu Pahat, Johor; 1 ° 50 ′ 53.6 ″ N, 102 ° 57 ′ 37.3 ″ E; 26 Sep. 2020; J. Neo leg.; iNaturalist. org / observations / 60810954 • 1 ♂; Kulaijaya; 1 ° 34 ′ 12.6 ″ N, 103 ° 38 ′ 40.1 ″ E; 2 Mar. 2020; A. Zaidi leg.; iNaturalist. org / observations / 39477877 • 1 juvenile; Tamarind Springs Jalan 1, Taman Tun Abdul Razak, 68000 Ampang Jaya, Selangor; 3 ° 10 ′ 14.6 ″ N, 101 ° 46 ′ 14.3 ″ E; 8 Mar. 2020; T. Maul leg.; iNaturalist. org / observations / 39698484 • 1 ♂; UTM Skudai, Johor Bahru; 1 ° 33 ′ 26.3 ″ N, 103 ° 38 ′ 14.1 ″ E; 29 Apr. 2020; B. Kumpau leg.; iNaturalist. org / observations / 44237939 • 1 juvenile; Kuala Lumpur, 50480 Kuala Lumpur, Federal Territory of Kuala Lumpur; 3 ° 9 ′ 23.1 ″ N, 101 ° 41 ′ 5.9 ″ E; 26 May 2020; H. Shanmugam leg.; iNaturalist. org / observations / 47737528 • 1 ♀; 2 ° 48 ′ 23.6 ″ N, 101 ° 52 ′ 20.3 ″ E; 3 May 2020; anukma leg.; iNaturalist. org / observations / 47829023 • 1 ♂; Klinik Kesihatan Sungai Lembing, Jalan Sungai Lembing, Kampung Bayas, 26200 Sungai Lembing, Pahang; 3 ° 55 ′ 31 ″ N, 103 ° 4 ′ 17 ″ E; 7 Jun. 2020; A. Yusuf leg.; iNaturalist. org / observations / 48735148 • 1 ♀; Perdana Botanical Gardens, Kuala Lumpur, Federal Territory of Kuala Lumpur; 3 ° 8 ′ 41.7 ″ N, 101 ° 41 ′ 10.6 ″ E; 19 Jun. 2020; N. Rahman leg.; iNaturalist. org / observations / 50133159 • 1 ♂; Tamarind Springs Jalan 1, Taman Tun Abdul Razak, 68000 Ampang Jaya, Selangor; 3 ° 10 ′ 14.6 ″ N, 101 ° 46 ′ 14.3 ″ E; 20 Jun. 2020; T. Maul leg.; iNaturalist. org / observations / 50276251 • 1 ♂; Seberang Perai Selatan; 5 ° 17 ′ 13.6 ″ N, 100 ° 30 ′ 30 ″ E; 5 Jul. 2020; alexiuslzl leg.; iNaturalist. org / observations / 52006113 • 1 ♀; KJ 75 Pertubuhan Peladang, Pekan Batu 14 Hulu Langat, 43100 Hulu Langat, Selangor; 3 ° 6 ′ 50.1 ″ N, 101 ° 48 ′ 56.3 ″ E; 19 Jul. 2020; S. Saphian leg.; iNaturalist. org / observations / 53567997 • 1 juvenile; Batu Pahat, Johor; 1 ° 50 ′ 28 ″ N, 102 ° 57 ′ 36.8 ″ E; 31 Jul. 2020; diclim _ eschsbp leg.; iNaturalist. org / observations / 54914512 • 1 ♂; Taman Bukit Perdana, 83000 Batu Pahat, Johor; 1 ° 50 ′ 53.6 ″ N, 102 ° 57 ′ 37.3 ″ E; 31 Jul. 2020; ziki _ eschsbp leg.; iNaturalist. org / observations / 54932721 • 1 juvenile; NCP, Taman Tugu, Kuala Lumpur; 3 ° 9 ′ 4.4 ″ N, 101 ° 41 ′ 8.13 ″ E; 9 Aug. 2020; T. Maul leg.; iNaturalist. org / observations / 55895941 • 1 ♀; NCP, Taman Tugu, Kuala Lumpur; 3 ° 9 ′ 4.4 ″ N, 101 ° 41 ′ 8.13 ″ E; 9 Aug. 2020; T. Maul leg.; iNaturalist. org / observations / 55896227 • 1 juvenile; Jalan Perdana 17, Taman Bukit Perdana, 83000 Batu Pahat, Johor; 1 ° 50 ′ 53.7 ″ N, 102 ° 57 ′ 37.7 ″ E; 16 Aug. 2020; ziki _ eschsbp leg.; iNaturalist. org / observations / 56605014 • 1 ♂, 1 ♀; Jalan Perdana 17, Taman Bukit Perdana, 83000 Batu Pahat, Johor; 1 ° 50 ′ 53.7 ″ N, 102 ° 57 ′ 37.7 ″ E; 16 Aug. 2020; ziki _ eschsbp leg.; iNaturalist. org / observations / 56605016 • 1 juvenile; Taman Bukit Indah, 68000 Ampang Jaya, Selangor; 3 ° 9 ′ 34.7 ″ N, 101 ° 46 ′ 34.3 ″ E; 20 Aug. 2020; T. Maul leg.; iNaturalist. org / observations / 57008216 • 1 ♂; Taman Bukit Perdana, 83000 Batu Pahat, Johor; 1 ° 50 ′ 53.6 ″ N, 102 ° 57 ′ 37.3 ″ E; 20 Aug. 2020; ziki _ eschsbp leg.; iNaturalist. org / observations / 57014573 • 1 ♀; 83200 Senggarang, Johor; 1 ° 46 ′ 20 ″ N, 103 ° 1 ′ 33.1 ″ E; 22 Aug. 2020; ziki _ eschsbp leg.; iNaturalist. org / observations / 57193924 • 1 ♂; Taman Bukit Perdana 2, 83000 Batu Pahat, Johor; 1 ° 50 ′ 32.8 ″ N, 102 ° 57 ′ 15 ″ E; 20 Aug. 2020; tstan _ eschsbp leg.; iNaturalist. org / observations / 57307974 • 1 ♀; NCP, Taman Tugu, Kuala Lumpur; 3 ° 9 ′ 4.4 ″ N, 101 ° 41 ′ 8.13 ″ E; 23 Aug. 2020; T. Maul leg.; iNaturalist. org / observations / 57311814 • 1 juvenile; Jalan Cenderasari, Kuala Lumpur; 3 ° 8 ′ 44.6 ″ N, 101 ° 41 ′ 18.5 ″ E; 30 Aug. 2020; mengarooo leg.; iNaturalist. org / observations / 58026678 • 1 juvenile; Tamarind Springs Jalan 1, Taman Tun Abdul Razak, 68000 Ampang Jaya, Selangor; 3 ° 10 ′ 14.6 ″ N, 101 ° 46 ′ 14.3 ″ E; 31 Aug. 2020; T. Maul leg.; iNaturalist. org / observations / 58233139 • 1 ♀; 83200 Senggarang, Johor; 1 ° 44 ′ 6.4 ″ N, 103 ° 4 ′ 1.2 ″ E; 22 Aug. 2020; J. Neo leg.; iNaturalist. org / observations / 58578339 • 1 ♂; Choa Chu Kang Park; 1 ° 23 ′ 11.9 ″ N, 103 ° 44 ′ 51 ″ E; 6 Sep. 2020; mingwei leg.; iNaturalist. org / observations / 58695507 • 1 ♂, 1 ♀; Tamarind Springs Jalan 1, Taman Tun Abdul Razak, 68000 Ampang Jaya, Selangor; 3 ° 10 ′ 14.6 ″ N, 101 ° 46 ′ 14.3 ″ E; 5 Sep. 2020; T. Maul leg.; iNaturalist. org / observations / 58826217 • 1 ♀; Ampang Jaya, Selangor; 3 ° 13 ′ 0.100 ″ N, 101 ° 45 ′ 3.5 ″ E; 20 Sep. 2020; B. Sg leg.; iNaturalist. org / observations / 60147373 • 1 ♂, 1 ♀; 83200 Senggarang, Johor; 1 ° 46 ′ 36.8 ″ N, 103 ° 1 ′ 24.6 ″ E; 4 Oct. 2020; ziki _ eschsbp leg.; iNaturalist. org / observations / 61669789 • 1 juvenile; 83200 Senggarang, Johor; 1 ° 46 ′ 36.8 ″ N, 103 ° 1 ′ 24.6 ″ E; 4 Oct. 2020; ziki _ eschsbp leg.; iNaturalist. org / observations / 61669797 • 1 ♂; Garden Avenue, 70300 Seremban, Negeri Sembilan; 2 ° 40 ′ 39.7 ″ N, 101 ° 54 ′ 2.70 ″ E; 7 Oct. 2020; O. J. Seng leg.; iNaturalist. org / observations / 61917637 • 1 ♂; Garden Avenue, 70300 Seremban, Negeri Sembilan; 2 ° 40 ′ 53 ″ N, 101 ° 53 ′ 56.2 ″ E; 7 Oct. 2020; O. J. Seng leg.; iNaturalist. org / observations / 61920273 • 1 juvenile; Bukit Kiara, 60000 Kuala Lumpur, Federal Territory of Kuala Lumpur; 3 ° 9 ′ 12 ″ N, 101 ° 38 ′ 11 ″ E; 8 Oct. 2020; V. Ganesh leg.; iNaturalist. org / observations / 62013556 • 1 juvenile; Kuala Lumpur, Federal Territory of Kuala Lumpur; 3 ° 8 ′ 20.3 ″ N, 101 ° 41 ′ 12.6 ″ E; 8 Oct. 2020; C. W. Chun leg.; iNaturalist. org / observations / 62037651 • 1 ♀; Perdana Botanical Gardens, Kuala Lumpur, Federal Territory of Kuala Lumpur; 3 ° 8 ′ 37.2 ″ N, 101 ° 41 ′ 9.9 ″ E; 8 Oct. 2020; C. W. Chun leg.; iNaturalist. org / observations / 62075913 • 1 ♀; 1 ° 50 ′ 33.5 ″ N, 102 ° 57 ′ 38.2 ″ E; 10 Oct. 2020; joyce _ tan leg.; iNaturalist. org / observations / 62628458 • 1 juvenile; Petaling Jaya, Selangor; 3 ° 10 ′ 33.6 ″ N, 101 ° 36 ′ 31.1 ″ E; 25 Oct. 2020; msone leg.; iNaturalist. org / observations / 63555073 • 1 juvenile; Petaling Jaya, Selangor; 3 ° 10 ′ 33.6 ″ N, 101 ° 36 ′ 31.1 ″ E; 25 Oct. 2020; msone leg.; iNaturalist. org / observations / 63555074 • 1 juvenile; Perdana Botanical Gardens, Kuala Lumpur, Federal Territory of Kuala Lumpur; 3 ° 8 ′ 37.2 ″ N, 101 ° 41 ′ 9.9 ″ E; 8 Oct. 2020; C. W. Chun leg.; iNaturalist. org / observations / 62078581 • 1 ♂, 1 ♀; Gombak; 3 ° 15 ′ 41.7 ″ N, 101 ° 38 ′ 7 ″ E; 28 Apr. 2019; Y. Joyce leg.; iNaturalist. org / observations / 23581051 • 1 juvenile; Gunung Lambak; 2 ° 1 ′ 39 ″ N, 103 ° 21 ′ 27 ″ E; 2 Aug. 2020; howe leg.; iNaturalist. org / observations / 55174550 • 1 ♀; Kuala Terengganu; 5 ° 16 ′ 55 ″ N, 103 ° 6 ′ 32 ″ E; 27 Dec. 2013; Lanzz leg.; https: // www. projectnoah. org / spottings / 130726139? fbclid = IwAR 1 CsvK 9 ownDvtCZtRiad 5 lj 9 dnh 3 aKasTEVLyyZGkwfXm 1 PNj dkKMrq-TM. SINGAPORE • 1 juvenile; Central Catchment Nature Reserve, “ sentier forestier ”; 1 ° 22 ′ 49 ″ N, 103 ° 49 ′ 6 ″ E; 79 m a. s. l.; 16 Jun. 2011; T. Robillard leg.; MNHN • 1 ♂; Elevage F 1; T. Robillard leg.; MNHN • 3 ♂♂; Elevage F 2; T. Robillard leg.; MNHN • 2 ♂♂; Elevage F 3; T. Robillard leg.; MNHN • 1 ♂; Elevage F 1; T. Robillard leg.; MNHN • 1 ♂; Elevage F 4; T. Robillard leg.; MNHN • 1 ♂; Elevage F 5; T. Robillard leg.; MNHN • 1 ♂; Elevage F 6; T. Robillard leg.; MNHN • 1 ♂; Elevage F 9; T. Robillard leg.; MNHN-EO-ENSIF 3102 • 4 ♂♂; Elevage; T. Robillard leg.; MNHN • 1 ♀; Elevage F 2; T. Robillard leg.; MNHN • 1 ♀; Elevage F 3; T. Robillard leg.; MNHN • 1 ♀; Elevage F 5; T. Robillard leg.; MNHN • 3 ♀♀; Elevage; T. Robillard leg.; MNHN • 1 ♀; Lim Chu Kang; 1 ° 26 ′ 46.1 ″ N, 103 ° 43 ′ 45.7 ″ E; 26 Dec. 2019; E. C. Maxwell leg.; iNaturalist. org / observations / 39013226 • 1 ♂; Blackmore Drive, Holland Village; 1 ° 19 ′ 59.8 ″ N, 103 ° 46 ′ 53.1 ″ E; 27 Oct. 2020; stefaniek leg.; iNaturalist. org / observations / 63613982 • 1 ♂; Central Water Catchment; 1 ° 20 ′ 48.2 ″ N, 103 ° 50 ′ 8.5 ″ E; 10 Apr. 2020; M. Finlay leg.; iNaturalist. org / observations / 41811938 • 1 ♂; Blk 70 A; 1 ° 16 ′ 35.6 ″ N, 103 ° 48 ′ 42 ″ E; 23 May 2020; lcwoo leg.; iNaturalist. org / observations / 46927932 • 1 ♀; Woodlands (SLE) Park Connector, Woodlands; 1 ° 25 ′ 36.7 ″ N, 103 ° 47 ′ 14.3 ″ E; 29 May 2020; looiec leg.; iNaturalist. org / observations / 47701690 • 1 ♀; Champions Way, Woodlands; 1 ° 25 ′ 40.1 ″ N, 103 ° 47 ′ 16.8 ″ E; 29 May 2020; looiec leg.; iNaturalist. org / observations / 47701715 • 1 juvenile; 36 C Lor Halwa 298637; 1 ° 20 ′ 10 ″ N, 103 ° 49 ′ 22.6 ″ E; 9 May 2020; flyingswallow 123 leg.; iNaturalist. org / observations / 48998890 • 1 ♀; Tanglin; 1 ° 18 ′ 2.5 ″ N, 103 ° 48 ′ 41.1 ″ E; 15 Jun. 2020; D. Rham leg.; iNaturalist. org / observations / 49656881 • 1 ♂; Yishun; 1 ° 24 ′ 51.7 ″ N, 103 ° 50 ′ 34.2 ″ E; 20 Jun. 2020; M. Ng leg.; iNaturalist. org / observations / 50287104 • 1 ♂, 1 ♀; Rambutan Trail, Thomson; 1 ° 23 ′ 6.28 ″ N, 103 ° 49 ′ 13.8 ″ E; 28 Jun. 2020; simchoonheok leg.; iNaturalist. org / observations / 51206798 • 1 juvenile; Tengah; 1 ° 21 ′ 58 ″ N, 103 ° 43 ′ 47.2 ″ E; 23 Jun. 2020; C. W. Gan leg.; iNaturalist. org / observations / 51352950 • 1 ♀; Bishan; 1 ° 20 ′ 57.1 ″ N, 103 ° 49 ′ 59.8 ″ E; 27 Jun. 2020; M. Finlay leg.; iNaturalist. org / observations / 51885537 • 1 ♂; Bukit Timah Nature Reserve; 1 ° 21 ′ 38.8 ″ N, 103 ° 46 ′ 38.1 ″ E; 5 Jul. 2020; ramesh-birding-butterflying leg.; iNaturalist. org / observations / 52003328 • 1 ♂; Kent Ridge Park, Pasir Panjang; 1 ° 17 ′ 7.28 ″ N, 103 ° 47 ′ 28.3 ″ E; 2 Jul. 2020; skm 3355 leg.; iNaturalist. org / observations / 52035813 • 1 ♀; Bef Dairy Farm Road; 1 ° 22 ′ 0 ″ N, 103 ° 46 ′ 32 ″ E; 10 Jul. 2020; N. Sanjai leg.; iNaturalist. org / observations / 52529111 • 1 ♀; Bukit Panjang; 1 ° 21 ′ 50.8 ″ N, 103 ° 46 ′ 34.2 ″ E; 10 Jul. 2020; N. Sanjai leg.; iNaturalist. org / observations / 52536392 • 1 ♀; Bef Old Upp Thomson Road; 1 ° 23 ′ 12.3 ″ N, 103 ° 49 ′ 12.9 ″ E; 10 Jul. 2020; Y. H. Koh leg.; iNaturalist. org / observations / 52542825 • 1 ♂; MacRitchie Reservoir; 1 ° 20 ′ 41.3 ″ N, 103 ° 49 ′ 20.5 ″ E; 10 Jul. 2020; falseamoebae leg.; iNaturalist. org / observations / 52574945 • 1 ♀; Novena; 1 ° 20 ′ 0.598 ″ N, 103 ° 49 ′ 39.3 ″ E; 10 Jul. 2020; M. Finlay leg.; iNaturalist. org / observations / 52644701 • 1 ♀; Upper Seletar Reservoir Park; 1 ° 23 ′ 58.7 ″ N, 103 ° 48 ′ 29.3 ″ E; 10 Jul. 2020; K. Well leg.; iNaturalist. org / observations / 52703749 • 1 juvenile; Bukit Panjang; 1 ° 20 ′ 52.7 ″ N, 103 ° 46 ′ 30.3 ″ E; 14 Jul. 2020; M. Nasry leg.; iNaturalist. org / observations / 53034819 • 1 ♀; Rifle Range Link; 1 ° 21 ′ 22.6 ″ N, 103 ° 48 ′ 6 ″ E; 12 Jul. 2020; K. Well leg.; iNaturalist. org / observations / 53036305 • 1 ♀; 100 Dairy Farm Road, 679057; 1 ° 21 ′ 42.7 ″ N, 103 ° 46 ′ 31.1 ″ E; 17 Jul. 2020; M. Ng leg.; iNaturalist. org / observations / 53356890 • 1 ♀; 100 Dairy Farm Road, 679057; 1 ° 21 ′ 42.7 ″ N, 103 ° 46 ′ 31.1 ″ E; 17 Jul. 2020; M. Ng leg.; iNaturalist. org / observations / 53356981 • 1 ♂; Central Water Catchment; 1 ° 20 ′ 48.7 ″ N, 103 ° 50 ′ 8 ″ E; 12 Jul. 2020; M. Finlay leg.; iNaturalist. org / observations / 53451611 • 1 ♂; Central Water Catchment; 1 ° 20 ′ 46.7 ″ N, 103 ° 50 ′ 9.3 ″ E; 12 Jul. 2020; M. Finlay leg.; iNaturalist. org / observations / 53451612 • 1 ♀; Bishan; 1 ° 20 ′ 51.5 ″ N, 103 ° 50 ′ 7.70 ″ E; 12 Jul. 2020; M. Finlay leg.; iNaturalist. org / observations / 53452345 • 1 ♂; Novena; 1 ° 20 ′ 8.13 ″ N, 103 ° 49 ′ 40.6 ″ E; 18 Jul. 2020; M. Finlay leg.; iNaturalist. org / observations / 53454664 • 1 ♀; Upper Thomson Road; 1 ° 23 ′ 7.4 ″ N, 103 ° 49 ′ 15.6 ″ E; 19 Jul. 2020; M. Ng leg.; iNaturalist. org / observations / 53596352 • 1 ♀; Wallace Centre, Dairy Farm Nature Park; 1 ° 21 ′ 35.3 ″ N, 103 ° 46 ′ 39 ″ E; 19 Jul. 2020; K. Well leg.; iNaturalist. org / observations / 53622733 • 1 ♂, 1 ♀; Bukit Panjang; 1 ° 21 ′ 42.7 ″ N, 103 ° 46 ′ 31.1 ″ E; 19 Jul. 2020; lcwoo leg.; iNaturalist. org / observations / 53668007 • 1 juvenile; Dairy Farm Nature Park; 1 ° 21 ′ 42.7 ″ N, 103 ° 46 ′ 31.1 ″ E; 19 Jul. 2020; lcwoo leg.; iNaturalist. org / observations / 53673593 • 1 ♂; Bukit Timah Nature Reserve, Bukit Timah; 1 ° 21 ′ 22.1 ″ N, 103 ° 46 ′ 8.56 ″ E; 18 Jul. 2020; gcynature leg.; iNaturalist. org / observations / 53775496 • 1 ♂; 1 ° 22 ′ 22.5 ″ N, 103 ° 49 ′ 26.6 ″ E; 24 Jul. 2020; noahartpradid leg.; iNaturalist. org / observations / 54131093 • 1 juvenile; Windsor Park; 1 ° 21 ′ 41.3 ″ N, 103 ° 49 ′ 35.5 ″ E; 22 Jul. 2020; C. W. Gan leg.; iNaturalist. org / observations / 54154877 • 1 ♂; Neo Tiew Lane 2 at Kranji Marsh; 1 ° 25 ′ 0.80 ″ N, 103 ° 43 ′ 34.2 ″ E; 25 Jul. 2020; K. Well leg.; iNaturalist. org / observations / 54259065 • 1 ♀; Central Water Catchment; 1 ° 22 ′ 26.3 ″ N, 103 ° 49 ′ 13.6 ″ E; 26 Jul. 2020; gracelsy leg.; iNaturalist. org / observations / 54379190 • 1 ♂; Thomson Nature Park; 1 ° 22 ′ 56 ″ N, 103 ° 49 ′ 19.5 ″ E; 30 Jul. 2020; K. Well leg.; iNaturalist. org / observations / 54847762 • 1 ♂, 1 ♀; Thomson Nature Park; 1 ° 23 ′ 7.4 ″ N, 103 ° 49 ′ 15.6 ″ E; 31 Jul. 2020; S. Chan leg.; iNaturalist. org / observations / 54945933 • 1 ♂; Thomson Nature Park; 1 ° 23 ′ 7.4 ″ N, 103 ° 49 ′ 15.6 ″ E; 31 Jul. 2020; S. Chan leg.; iNaturalist. org / observations / 54946444 • 1 juvenile; Lorong Pelita, Thomson; 1 ° 22 ′ 58.8 ″ N, 103 ° 49 ′ 17.3 ″ E; 31 Jul. 2020; yeehwa leg.; iNaturalist. org / observations / 55005657 • 1 juvenile; North; 1 ° 22 ′ 5.9 ″ N, 103 ° 46 ′ 52.1 ″ E; 1 Aug. 2020; S. G. Wen leg.; iNaturalist. org / observations / 55005954 • 1 ♂, 1 ♀; Central Water Catchment; 1 ° 23 ′ 49.1 ″ N, 103 ° 48 ′ 13 ″ E; 2 Aug. 2020; S. K. Yung leg.; iNaturalist. org / observations / 55122889 • 1 juvenile; Sungei Buloh Wetland Reserve; 1 ° 26 ′ 46.2 ″ N, 103 ° 43 ′ 39.7 ″ E; 2 Aug. 2020; K. Well leg.; iNaturalist. org / observations / 55167429 • 1 ♂; Botanic Gardens, Holland Village; 1 ° 18 ′ 28.8 ″ N, 103 ° 49 ′ 3 ″ E; 3 Aug. 2020; pb 1785 leg.; iNaturalist. org / observations / 55235146 • 1 ♂; Aft Tagore Drive; 1 ° 23 ′ 5.78 ″ N, 103 ° 49 ′ 14.9 ″ E; 8 Aug. 2020; lcwoo leg.; iNaturalist. org / observations / 55762965 • 1 juvenile; Bukit Drive, Bukit Timah; 1 ° 20 ′ 54.5 ″ N, 103 ° 46 ′ 29.8 ″ E; 2 Aug. 2020; gcynature leg.; iNaturalist. org / observations / 56017697 • 1 ♂; Central Water Catchment; 1 ° 20 ′ 44.7 ″ N, 103 ° 50 ′ 10.1 ″ E; 8 Aug. 2020; T. Z. Wan leg.; iNaturalist. org / observations / 56103224 • 1 juvenile; Andrews Avenue, Sembawang; 1 ° 27 ′ 26.8 ″ N, 103 ° 50 ′ 21.1 ″ E; 11 Aug. 2020; K. Well leg.; iNaturalist. org / observations / 56116568 • 1 ♀; Mandai; 1 ° 25 ′ 3.5 ″ N, 103 ° 47 ′ 27.8 ″ E; 16 Aug. 2020; S. K. Yung leg.; iNaturalist. org / observations / 56602046 • 1 ♀; Central Water Catchment; 1 ° 23 ′ 50.7 ″ N, 103 ° 48 ′ 20.3 ″ E; 16 Aug. 2020; kangmin leg.; iNaturalist. org / observations / 56625952 • 1 juvenile; Bukit Panjang; 1 ° 21 ′ 2.5 ″ N, 103 ° 46 ′ 44.2 ″ E; 3 Aug. 2020; T. Z. Wan leg.; iNaturalist. org / observations / 56827799 • 1 juvenile; Admiralty West Park Connector, Woodlands; 1 ° 27 ′ 4.4 ″ N, 103 ° 46 ′ 48 ″ E; 18 Aug. 2020; M. Farook leg.; iNaturalist. org / observations / 56946793 • 1 ♀; Lower Peirce Reservoir; 1 ° 22 ′ 12.3 ″ N, 103 ° 49 ′ 23.1 ″ E; 21 Aug. 2020; G. Lim leg.; iNaturalist. org / observations / 57110204 • 1 ♀; Bukit Batok East Avenue 2, Bukit Batok Nature Park Playground; 1 ° 20 ′ 59.3 ″ N, 103 ° 45 ′ 49 ″ E; 22 Aug. 2020; V. Lim leg.; iNaturalist. org / observations / 57189865 • 1 juvenile; Bukit Batok; 1 ° 20 ′ 58.6 ″ N, 103 ° 45 ′ 51.6 ″ E; 22 Aug. 2020; V. Lim leg.; iNaturalist. org / observations / 57191688 • 1 ♂; Lim Chu Kang; 1 ° 25 ′ 7.5 ″ N, 103 ° 43 ′ 15.6 ″ E; 23 Aug. 2020; S. K. Yung leg.; iNaturalist. org / observations / 57315540 • 1 ♀; Lower Peirce Reservoir Park; 1 ° 22 ′ 23 ″ N, 103 ° 49 ′ 26.8 ″ E; 23 Aug. 2020; darkendaltrax leg.; iNaturalist. org / observations / 57315659 • 1 ♀; Upper Seletar Reservoir Park; 1 ° 24 ′ 3.20 ″ N, 103 ° 48 ′ 25.3 ″ E; 23 Aug. 2020; S. Chan leg.; iNaturalist. org / observations / 57317179 • 1 juvenile; 1 ° 20 ′ 46.7 ″ N, 103 ° 53 ′ 9.63 ″ E; 23 Aug. 2020; lowch leg.; iNaturalist. org / observations / 57318298 • 1 ♀; Bukit Timah Nature Reserve; 1 ° 21 ′ 51.1 ″ N, 103 ° 46 ′ 33.2 ″ E; 24 Aug. 2020; noahartpradid leg.; iNaturalist. org / observations / 57429535 • 1 ♂; Lower Peirce Reservoir; 1 ° 22 ′ 12.3 ″ N, 103 ° 49 ′ 23.1 ″ E; 29 Aug. 2020; G. Lim leg.; iNaturalist. org / observations / 57916937 • 1 ♀; Lim Chu Kang; 1 ° 25 ′ 1.2 ″ N, 103 ° 43 ′ 32.3 ″ E; 3 Aug. 2020; S. K. Yung leg.; iNaturalist. org / observations / 58007762 • 1 ♂, 1 ♀; Upper Thomson Road; 1 ° 23 ′ 7.4 ″ N, 103 ° 49 ′ 15.6 ″ E; 10 Jul. 2020; C. Tong leg.; iNaturalist. org / observations / 58179315 • 1 ♂; Sungei Kadut; 1 ° 25 ′ 24.8 ″ N, 103 ° 45 ′ 19.8 ″ E; 18 Aug. 2020; S. Hui leg.; iNaturalist. org / observations / 58230167 • 1 ♂, 1 ♀; Champions Golf Course, Bukit Timah; 1 ° 20 ′ 24.1 ″ N, 103 ° 47 ′ 54.6 ″ E; 28 Aug. 2020; noahartpradid leg.; iNaturalist. org / observations / 58407499 • 1 ♀; Ang Mo Kio; 1 ° 22 ′ 52.5 ″ N, 103 ° 49 ′ 16.8 ″ E; 6 Sep. 2020; lowch leg.; iNaturalist. org / observations / 58680372 • 1 juvenile; Zhenghua Park; 1 ° 22 ′ 36.2 ″ N, 103 ° 46 ′ 34.8 ″ E; 6 Sep. 2020; S. Chan leg.; iNaturalist. org / observations / 58696289 • 1 ♀; 1 ° 21 ′ 49.2 ″ N, 103 ° 46 ′ 19.8 ″ E; 7 Sep. 2020; kei _ y leg.; iNaturalist. org / observations / 58931194 • 1 ♀; 1 ° 21 ′ 50.2 ″ N, 103 ° 46 ′ 36.1 ″ E; 1 Sep. 2020; kei _ y leg.; iNaturalist. org / observations / 58931653 • 1 juvenile; Bef Mandai Road; 1 ° 24 ′ 37.2 ″ N, 103 ° 46 ′ 41.8 ″ E; 4 Sep. 2020; T. Z. Wan leg.; iNaturalist. org / observations / 58957627 • 1 ♂, 1 ♀; Dairy Farm Nature Park; 1 ° 21 ′ 42.7 ″ N, 103 ° 46 ′ 31.1 ″ E; 9 Sep. 2020; S. Chan leg.; iNaturalist. org / observations / 59036868 • 1 ♀; Central Water Catchment; 1 ° 22 ′ 16.5 ″ N, 103 ° 46 ′ 46.2 ″ E; 9 Sep. 2020; S. Chan leg.; iNaturalist. org / observations / 59038833 • 1 juvenile; Upper Seletar Reservoir Park; 1 ° 24 ′ 3.20 ″ N, 103 ° 48 ′ 25.3 ″ E; 11 Sep. 2020; S. Chan leg.; iNaturalist. org / observations / 59209460 • 1 ♀; Novena; 1 ° 20 ′ 13.9 ″ N, 103 ° 49 ′ 36.8 ″ E; 29 Aug. 2020; M. Finlay leg.; iNaturalist. org / observations / 59402675 • 1 ♂; Central Water Catchment; 1 ° 23 ′ 47.7 ″ N, 103 ° 48 ′ 12.1 ″ E; 13 Sep. 2020; S. K. Yung leg.; iNaturalist. org / observations / 59419512 • 1 ♀; Central Water Catchment; 1 ° 23 ′ 48.8 ″ N, 103 ° 48 ′ 13.1 ″ E; 13 Sep. 2020; S. K. Yung leg.; iNaturalist. org / observations / 59419728 • 1 ♀; 1 ° 23 ′ 5.28 ″ N, 103 ° 49 ′ 21.3 ″ E; 15 Sep. 2020; kei _ y leg.; iNaturalist. org / observations / 59666269 • 1 ♂; 1 ° 22 ′ 56.7 ″ N, 103 ° 49 ′ 16.8 ″ E; 15 Sep. 2020; kei _ y leg.; iNaturalist. org / observations / 59666275 • 1 ♀; Champions Golf Course, Bukit Timah; 1 ° 20 ′ 28.3 ″ N, 103 ° 47 ′ 53.7 ″ E; 16 Sep. 2020; noahartpradid leg.; iNaturalist. org / observations / 59759002 • 1 ♀; Bukit Panjang; 1 ° 21 ′ 44.3 ″ N, 103 ° 46 ′ 36.1 ″ E; 20 Sep. 2020; M. Nasry leg.; iNaturalist. org / observations / 60171530 • 1 ♂; Upper Seletar Reservoir Park; 1 ° 24 ′ 3.20 ″ N, 103 ° 48 ′ 25.3 ″ E; 19 Sep. 2020; S. Chan leg.; iNaturalist. org / observations / 60189103 • 1 ♀; Kranji Marshes, Neo Tiew Lane 2; 1 ° 25 ′ 0.80 ″ N, 103 ° 43 ′ 35.7 ″ E; 20 Sep. 2020; mingwei leg.; iNaturalist. org / observations / 60196972 • 1 ♀; 1 ° 19 ′ 14.3 ″ N, 103 ° 40 ′ 42.2 ″ E; 20 Sep. 2020; prakashbr leg.; iNaturalist. org / observations / 60402098 • 1 ♂; Dairy Farm Nature Park; 1 ° 21 ′ 42.7 ″ N, 103 ° 46 ′ 31.1 ″ E; 24 Sep. 2020; S. Chan leg.; iNaturalist. org / observations / 60596693 • 1 ♂; Block 265; 1 ° 20 ′ 59.7 ″ N, 103 ° 45 ′ 35.1 ″ E; 25 Sep. 2020; yq _ 1810 leg.; iNaturalist. org / observations / 60696154 • 1 ♂; Tanglin; 1 ° 19 ′ 11.3 ″ N, 103 ° 48 ′ 54.3 ″ E; 26 Sep. 2020; scarlett 26 leg.; iNaturalist. org / observations / 60807200 • 1 juvenile; Bukit Panjang; 1 ° 21 ′ 22.6 ″ N, 103 ° 46 ′ 56.7 ″ E; 26 Sep. 2020; S. Chan leg.; iNaturalist. org / observations / 60811750 • 1 ♀; Dairy Farm Nature Park; 1 ° 21 ′ 42.7 ″ N, 103 ° 46 ′ 31.1 ″ E; 26 Sep. 2020; S. Chan leg.; iNaturalist. org / observations / 60817045 • 1 juvenile; Bukit Panjang; 1 ° 21 ′ 14.6 ″ N, 103 ° 46 ′ 37 ″ E; 27 Sep. 2020; S. Chan leg.; iNaturalist. org / observations / 60972069 • 1 ♀; Thomson Nature Park; 1 ° 23 ′ 7.5 ″ N, 103 ° 49 ′ 15.6 ″ E; 1 Oct. 2020; G. Lim leg.; iNaturalist. org / observations / 61387731 • 1 ♂; Queenstown; 1 ° 17 ′ 6.70 ″ N, 103 ° 47 ′ 29 ″ E; 6 Oct. 2020; S. K. Yung leg.; iNaturalist. org / observations / 61843628 • 1 ♀; Thomson Nature Park; 1 ° 22 ′ 53.3 ″ N, 103 ° 49 ′ 20.1 ″ E; 4 Oct. 2020; patricialorenz leg.; iNaturalist. org / observations / 61869845 • 1 ♂; 100 Dairy Farm Road, 679057; 1 ° 21 ′ 42.7 ″ N, 103 ° 46 ′ 31.1 ″ E; 9 Oct. 2020; C. Tong leg.; iNaturalist. org / observations / 62099010 • 1 ♂; Tanglin, Swan Lake; 1 ° 18 ′ 28.5 ″ N, 103 ° 48 ′ 55.2 ″ E; 11 Oct. 2020; L. J. Yi leg.; iNaturalist. org / observations / 62263904 • 1 ♂, 1 ♀; Botanic Gardens; 1 ° 18 ′ 49.7 ″ N, 103 ° 48 ′ 57.2 ″ E; 11 Oct. 2020; lcwoo leg.; iNaturalist. org / observations / 62384745 • 1 ♂; Holland Village; 1 ° 19 ′ 50.5 ″ N, 103 ° 46 ′ 52.7 ″ E; 13 Oct. 2020; geledek leg.; iNaturalist. org / observations / 62453318 • 1 juvenile; Unnamed Road; 1 ° 26 ′ 31.8 ″ N, 103 ° 44 ′ 9 ″ E; 13 Oct. 2020; S. K. Yung leg.; iNaturalist. org / observations / 62472505 • 1 ♂; Central Water Catchment; 1 ° 22 ′ 24.8 ″ N, 103 ° 49 ′ 27.8 ″ E; 16 Oct. 2020; Y. Jun leg.; iNaturalist. org / observations / 62719526 • 1 ♂; Yio Chu Kang; 1 ° 24 ′ 7.06 ″ N, 103 ° 49 ′ 4.78 ″ E; 17 Oct. 2020; M. Ng leg.; iNaturalist. org / observations / 62797963 • 1 ♀; Central Water Catchment; 1 ° 23 ′ 58.7 ″ N, 103 ° 48 ′ 28.6 ″ E; 18 Oct. 2020; S. K. Yung leg.; iNaturalist. org / observations / 62877098 • 1 ♂; HortPark; 1 ° 16 ′ 43.5 ″ N, 103 ° 48 ′ 6.56 ″ E; 18 Oct. 2020; S. Chan leg.; iNaturalist. org / observations / 62896486 • 1 ♀; Kent Ridge Park; 1 ° 17 ′ 2 ″ N, 103 ° 47 ′ 28.6 ″ E; 18 Oct. 2020; S. Chan leg.; iNaturalist. org / observations / 62898852 • 1 ♂; Woodlands; 1 ° 26 ′ 59.6 ″ N, 103 ° 46 ′ 43.2 ″ E; 19 Oct. 2020; Y. Jun leg.; iNaturalist. org / observations / 63007200 • 1 ♀; Holland Village; 1 ° 19 ′ 49.8 ″ N, 103 ° 46 ′ 53 ″ E; 20 Oct. 2020; irwin 88 leg.; iNaturalist. org / observations / 63062077 • 1 ♀; Holland Village; 1 ° 20 ′ 2.78 ″ N, 103 ° 46 ′ 53.7 ″ E; 20 Oct. 2020; rocherussel leg.; iNaturalist. org / observations / 63062082 • 1 ♀; Holland Village; 1 ° 19 ′ 50.3 ″ N, 103 ° 46 ′ 52.8 ″ E; 20 Oct. 2020; khim 99 leg.; iNaturalist. org / observations / 63062122 • 1 ♀; Bukit Timah Road, Holland Village; 1 ° 20 ′ 8 ″ N, 103 ° 46 ′ 53.2 ″ E; 20 Oct. 2020; i 328026 leg.; iNaturalist. org / observations / 63062126 • 1 ♀; BukitTimah; 1 ° 19 ′ 33.5 ″ N, 103 ° 46 ′ 51.5 ″ E; 20 Oct. 2020; kangmin leg.; iNaturalist. org / observations / 63076249 • 1 ♂; Central Water Catchment; 1 ° 21 ′ 18.5 ″ N, 103 ° 47 ′ 49.8 ″ E; 23 Oct. 2020; C. Tong leg.; iNaturalist. org / observations / 63332336 • 1 ♀; Central Water Catchment; 1 ° 21 ′ 18.5 ″ N, 103 ° 47 ′ 49.8 ″ E; 23 Oct. 2020; C. Tong leg.; iNaturalist. org / observations / 63332337 • 1 ♀; Novena; 1 ° 20 ′ 7.20 ″ N, 103 ° 49 ′ 43.2 ″ E; 22 Oct. 2020; M. Finlay leg.; iNaturalist. org / observations / 63354764 • 1 ♂, 2 ♀♀; Novena; 1 ° 20 ′ 7.78 ″ N, 103 ° 49 ′ 43.2 ″ E; 22 Oct. 2020; M. Finlay leg.; iNaturalist. org / observations / 63354765 • 1 ♀; Novena; 1 ° 20 ′ 7.70 ″ N, 103 ° 49 ′ 41.1 ″ E; 18 Oct. 2020; M. Finlay leg.; iNaturalist. org / observations / 63357376 • 1 ♂; Novena; 1 ° 20 ′ 8.8 ″ N, 103 ° 49 ′ 44.2 ″ E; 18 Oct. 2020; M. Finlay leg.; iNaturalist. org / observations / 63357384 • 1 ♂, 1 ♀; Central Water Catchment; 1 ° 23 ′ 55.5 ″ N, 103 ° 46 ′ 40.1 ″ E; 23 Oct. 2020; M. Finlay leg.; iNaturalist. org / observations / 63371589 • 1 juvenile; Kent Ridge Park; 1 ° 17 ′ 2 ″ N, 103 ° 47 ′ 28.6 ″ E; 24 Oct. 2020; S. Chan leg.; iNaturalist. org / observations / 63383652 • 1 ♂; Kent Ridge Park, Pasir Panjang; 1 ° 16 ′ 54.7 ″ N, 103 ° 47 ′ 33.7 ″ E; 14 Oct. 2020; skm 3355 leg.; iNaturalist. org / observations / 63400835 • 1 juvenile; Hindhede Nature Park; 1 ° 20 ′ 56.1 ″ N, 103 ° 46 ′ 32.6 ″ E; 25 Oct. 2020; lcwoo leg.; iNaturalist. org / observations / 63466017 • 1 ♂; Aft WʼLands Waterfront Park; 1 ° 27 ′ 2.89 ″ N, 103 ° 46 ′ 47 ″ E; 29 Oct. 2020; S. G. Wen leg.; iNaturalist. org / observations / 63750085 • 1?; Bukit Panjang; 1 ° 21 ′ 37.7 ″ N, 103 ° 46 ′ 37.3 ″ E; 30 Oct. 2020; M. Nasry leg.; iNaturalist. org / observations / 63878925 • 1 ♂, 1 ♀; Thomson Nature Park; 1 ° 23 ′ 7.5 ″ N, 103 ° 49 ′ 15.6 ″ E; 12 Jul. 2020; G. Lim leg.; iNaturalist. org / observations / 52933922 • 1 ♂; Sungei Buloh Wetland Reserve; 1 ° 26 ′ 53.1 ″ N, 103 ° 43 ′ 28.5 ″ E; 28 Dec. 2013; T. K. Hui leg.; iNaturalist. org / observations / 19011702.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFC7FFC4FDC6F26CFAB7FE1B.taxon	description	Description Habitus similar to N. vittatus (Figs 7 E, 23 A – D): vertex black with cream-coloured margin around eyes (Fig. 11 F). Fastigium velvety black with cream-coloured margins. Scapes dark red brown to black (particularly on dorso-inner margins). Antennae black with wide whitish rings. Frons and mouthparts most often yellow with a few dark spots (Fig. 10 F). Maxillary palpi cream-coloured, apical segment black apically. Head lateral side yellow, with a wide black band posterior to eye (Fig. 12 F). Pronotum covered with white setae; dorsal disk dark red brown to black (Fig. 11 F); lateral lobes of pronotum dorsal third black, vivid yellow ventrally (cream-coloured in preserved specimens) (Fig. 12 F). Legs dark orange brown. FIIIs brown, knees dark brown to black; TIIIs brown with black spines and spurs, dark brown to black near distal end; tarsomeres dark brown to black. Hindwings hyaline brown apically, transparent basally; longer than FWs, forming a dark brown tail exceeding FWs for more than twice the length of pronotum. Tergites red brown, with a pale band laterally; sternites pale, with a red brown median area. Cerci red brown, lighter dorsally. Male FOREWINGS. FW colouration (Fig. 13 E): dorsal field cells mostly transparent, veins mostly dark brown to black. Basal FW area black (instead of cream-coloured or brown as in N. vittatus), with a relatively small cream-coloured patch basally. R black (instead of cream-coloured as in N. vittatus), Sc projections cream-coloured basally, black apically. Apex of dorsal field somewhat truncated. Lateral field dark. FW venation (Fig. 13 E): 1 A curved, slightly angulated. Harp slightly longer than wide, with 3 oblique veins, basal one bifurcated distally, distal one bifurcated basally. c 1 long and wide, c 2 diamond-shaped; mirror (d 1) longer than wide, not rounded, generally separated into two parts by a distinct transverse vein, the posterior part nearly rectangular, shorter than anterior part. Cell d 2 narrower than d 1. Anal-most and middle chords narrow and nearly parallel; basal-most chord extending from basal half of middle chord. Apical field short and rounded, with 3 wide cell alignments posterior to mirror and a narrow apical alignment; veins cream-coloured. Lateral field with 5 projections of Sc. Epiproct black. Subgenital plate pale cream-coloured with median area and anterior and posterior margins black. GENITALIA. Very similar to N. vittatus (Fig. 16 E) but differ by pseudepiphallus faintly less robust and broad, anterior margin more strongly concave. Rami more strongly curved. Female FOREWINGS. FW colouration (Fig. 14 F): cells black, veins generally cream-coloured. Without distinct longitudinal white stripe around region of CuA, M and R with distinct margin from black colouration on dorsal field; Sc projections cream-coloured basally, black apically. Lateral field dark. FW venation: 6 – 7 strong longitudinal veins on dorsal field; lateral field with 6 projections of Sc. GENITALIA. Ovipositor about as long as FIII. Copulatory papilla conical, small and stout; apex folded ventrally, short, pointed; dorsal face with a sclerotized area; ventro-anterior end forming a truncated lobule (Fig. 17 J – K). Juvenile (Fig. 24 A – C) Differing from that of N. vittatus by darker colouration, usually grey, never as white as N. vittatus. Stripes on head similar to N. vittatus but sometimes very faint; pronotum dorsal disk homogeneously dark grey, without longitudinal stripes; sometimes with a large velvety black spot. Legs grey with dark stripes, but stripes not as distinct as in N. vittatus. Abdominal tergites dark grey, their posterior margins thinly lined with yellow, not forming a pattern of transverse stripes. In some individuals, first and second abdominal tergites with a particularly dark (black), velvety band. Measurements See Table 8.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFC7FFC4FDC6F26CFAB7FE1B.taxon	biology_ecology	Ecology This species can be found among shrubs and herbaceous plants (but not grasses) (Fig. 27). The adults and nymphs from Singapore and Malaysia (previously reported as N. vittatus) have been reported to feed on the flower parts of a varieties of native (e. g., Dillenia suffruticosa (Griff.) Martelli) and exotic weedy (e. g., Lantana camara L.) plant species (Tan et al. 2017). They also feed on the foliages of understory plants such as Piper sarmentosum Roxb. (Fig. 27). In Singapore, N. malaya sp. nov. can be found abundantly in the understory where Piper sarmentosum dominates.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFC7FFC4FDC6F26CFAB7FE1B.taxon	distribution	Distribution Malay Peninsula (Singapore and Peninsular Malaysia [northern limit at the state of Penang]) (Fig. 5).	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFC7FFC4FDC6F26CFAB7FE1B.taxon	materials_examined	Type locality Singapore.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFC7FFC4FDC6F26CFAB7FE1B.taxon	description	Calling song (Singapore, n = 4 males, 20 echemes; MNHN-SO- 2851 – 2854) (Fig. 25 A – C) At 27.3 ° C, the calling song consists of bouts of highly variable length, ranging from a few seconds to several minutes. Each bout is composed of short echemes made of triplets of syllables; echeme duration is 32.3 ± 2.8 ms (29.0 – 37.56 ms); echeme period is 64.5 ± 4.3 ms (59.5 – 75.5 ms); downtime between consecutive echemes is 32.1 ± 4.4 ms (24.3 – 41.3 ms). Durations for the first to third syllables of the triplet are as follows: 6.1 ± 1.5 ms (3.7 – 8.0 ms), 6.7 ± 2.7 ms (2.5 – 10.6 ms) and 5.8 ± 2.9 ms (3.0 – 11.2 ms). Call spectrum consists of a series of harmonics with three distinct peaks in the first three harmonics: first harmonic (fundamental frequency) at 7.35 ± 1.28 kHz (6.201 – 9.39 kHz), which is also the dominant frequency; second harmonic at 13.33 ± 1.01 kHz (12.23 – 15.07 kHz); third harmonic at 18.88 ± 0.77 kHz (17.23 – 20.07 kHz).	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFC7FFC4FDC6F26CFAB7FE1B.taxon	discussion	Remarks We split the species previously recognised as N. vittatus into two distinct species after integrating evidence from general and nymphal colouration, female genitalia morphology, acoustics, DNA (12 S sequence) and distribution. In terms of adult morphology, the new species is very similar to N. vittatus in habitus, other than being darker in colouration. The male genitalia are not always distinguishable between these species; but the sclerotization of the female genitalia can be differentiated. However, the colour patterns in the nymphs are drastically different. Likewise, a multivariate analysis of the call properties also showed distinct calling songs. The 12 S gene tree also demonstrates that the two species form distinct clades. Lastly, the new species occurs in Malay Peninsula whereas N. vittatus occurs in Borneo and Sumatra. Integrating these pieces of evidence indicates that N. vittatus and N. malaya sp. nov. are indeed separate species having probably diverged recently, despite the similarities in adult habitus and male genitalia.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD4FFC4FE49F146FDFEFAF7.taxon	materials_examined	Type locality Indonesia: Maluka, Seram Island.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD4FFC4FE49F146FDFEFAF7.taxon	description	Type details Male, not located.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD4FFC4FE49F146FDFEFAF7.taxon	discussion	Remarks According to the uninformative original description, the type of the species is from Seram Island, while no species of Nisitrus has ever been recorded in the region of Maluku Archipelago (Indonesia). In addition, the description mentions that the eyes are little prominent, and that the FWs extend beyond the abdomen, which is not compatible with any other species of Nisitrus. With the type missing, we consider this species as nomen dubium (new status).	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD4FFC4FF2FF62BFAD2F810.taxon	materials_examined	Material examined Syntype INDONESIA – Sumatra • 1 ♀; Stamford Raffles leg.; NHMUK 012497570.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD4FFC4FF2FF62BFAD2F810.taxon	discussion	Remarks It is unclear why Otte (1994) reinstated N. marginatus as a valid species, even though earlier literature had already synonymised this species under N. vittatus. Based on the observation of a female syntype showing no difference with N. vittatus, we synonymise N. marginatus again under N. vittatus.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD5FFC0FDBCF26CFD86FB63.taxon	description	Figs 5, 7 F, 9 B, 10 G, 11 G, 12 G, 13 F, 14 G, 16 F, 17 L – M, 26, 28; Tables 1 – 2, 9	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD5FFC0FDBCF26CFD86FB63.taxon	diagnosis	Diagnosis (revised from Ingrisch, 1987) Frons whitish with a black stripe between eyes and black mouthparts; femora reddish brown with a black stripe dorsally; male genitalia with pseudepiphallus slenderer than in congeners. This species is most similar to N. danum sp. nov., also from Borneo, from which it differs by mouthparts black (instead of yellow), FIIIs black dorsally and female papilla. While this species is restricted to the highlands in the Kinabalu range, N. danum sp. nov. occurs in the Danum Valley and Tawau Hill areas at the east of Sabah. It is also similar to N. vittatus, N. crucius sp. nov. and N. malaya sp. nov., from which it differs by slightly larger and slenderer habitus, frons whitish with black front head and mouthparts, FIIIs with a black pattern dorsally, and slenderer pseudepiphallus.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD5FFC0FDBCF26CFD86FB63.taxon	materials_examined	Material examined Holotype MALAYSIA – Sabah • ♂; Mount Kinabalu; 6 ° 07 ′ 16.09 ″ N, 116 ° 35 ′ 46.86 ″ E; 1000 – 1800 m a. s. l.; 5 – 7 Aug. 1984; S. Ingrisch leg.; SMFD. Paratype MALAYSIA – Sabah • 1 ♀; same collection data as for holotype; SMFD. Additional specimens MALAYSIA – Sabah • 6 ♂♂; Mount Kinabalu Mesilau Trail; 6 ° 02 ′ 37.74 ″ N, 116 ° 35 ′ 46.30 ″ E; 5000 ft a. s. l.; 13 – 15 Mar. 1964; S. Kueh (Royal Society Expedition) leg.; NHMUK • 1 ♀; Mount Kinabalu Mesilau Trail; 6 ° 02 ′ 37.74 ″ N, 116 ° 35 ′ 46.30 ″ E; 15 Feb. 1964; Royal Society Expedition leg.; NHMUK 012497572 • 3 ♂♂; Mount Kinabalu; 6 ° 02 ′ 37.74 ″ N, 116 ° 35 ′ 46.30 ″ E; 20 Mar. 1964; S. Kueh (Royal Society Expedition) leg.; NHMUK • 1 ♂; west of 3.1 km on Road Tenom Keningau, depleted lowland forest; 5 ° 15 ′ 39.17 ″ N, 115 ° 58 ′ 30.22 ″ E; 240 m a. s. l.; 25 Jun. 1986; J. Huisman leg.; RMNH • 2 ♀♀; Tenompok 30 miles east of Jesselton; 6 ° 00 ′ 10.40 ″ N, 116 ° 31 ′ 45.20 ″ E; 1460 m a. s. l.; 17 – 21 Oct. 1958; T. C. Maa leg.; BPBM • 2 ♂♂; same collection data as for preceding; BPBM • 2 ♂♂; west coast Residency, Kundasan; 5 ° 57 ′ 37.51 ″ N, 116 ° 33 ′ 18.03 ″ E; 1240 m a. s. l.; 15 Oct. 1958; T. C. Maa leg.; BPBM • 4 ♂♂; southern part of Kinabalu National Park; 5 ° 59 ′ 50.78 ″ N, 116 ° 33 ′ 39.77 ″ E; ~ 1500 – 2000 m a. s. l.; 26 Apr. – 1 May 2013; A. Gorochov, M. Berezin, V. Gorochova and E. Tkatsheva leg.; PII: N 52; ZIN • 3 ♀♀; same collection data as for preceding; ZIN • 5 ♂♂; Mount Trus Madi; 5 ° 33 ′ 53.94 ″ N, 116 ° 30 ′ 11.31 ″ E; ~ 1000 m a. s. l.; 13 – 25 May 2007; A. V. Gorochov leg.; PIII: N 37 Nmu 3; ZIN • 2 ♀♀; same collection data as for preceding; ZIN • 2 ♂♂; Mount Kinabalu; 6 ° 07 ′ 16.09 ″ N, 116 ° 35 ′ 46.86 ″ E; J. Zhang leg.; MNHN • 1 ♂; same locality data as for preceding; Dec. 2018; J. Zhang leg.; PII: N 48; MNHN-EO- ENSIF 11070 • 3 ♀♀; same locality data as for preceding; J. Zhang leg.; MNHN • 2 ♀♀; Tenompok near Kinabalu; 6 ° 00 ′ 10.40 ″ N, 116 ° 31 ′ 45.20 ″ E; 4500 ft a. s. l.; 18 Mar. 1929; H. M. Pendlebury leg.; identified as Nisitra vittata by L. Chopard; MNHN-EO-ENSIF 1738, MNHN-EO-ENSIF 1743 • 1 ♀; “ Crocker Range National Park not far from Keningau Town, secondary-primary forest ”; 5 ° 28 ′ 07.79 ″ N, 116 ° 04 ′ 22.95 ″ E; 1000 – 1300 m a. s. l.; 6 May 2013; A. Gorochov, M. Berezin, V. Gorochova and E. Tkatsheva leg.; N 29 - NboG 2; ZIN • 1 ♀; Kinabalu Park, Ranau; 6 ° 0 ′ 22.0 ″ N, 116 ° 32 ′ 33.5 ″ E; 3 Aug. 2018; lhurteau leg.; iNaturalist. org / observations / 15440494 • 1 ♀; Ranau, Kinabalu Park World Heritage Site; 6 ° 10 ′ 18.9 ″ N, 116 ° 43 ′ 45.9 ″ E; 10 Mar. 2018; rbeunen leg.; iNaturalist. org / observations / 18843361 • 1 ♂; Kinabalu Park, Kota Belud; 6 ° 3 ′ 24.8 ″ N, 116 ° 33 ′ 58.7 ″ E; 1 Aug. 2019; a 1 virginia leg.; iNaturalist. org / observations / 30690023 • 1 ♀; Kota Belud District; 6 ° 0 ′ 49.0 ″ N, 116 ° 30 ′ 35.3 ″ E; 28 Jun. 2019; paulsullivan 25 leg.; iNaturalist. org / observations / 27783166 • 1 juvenile; Tambunan; 5 ° 45 ′ 11.5 ″ N, 116 ° 30 ′ 0.6 ″ E; 24 Jun. 2019; paulsullivan 25 leg.; iNaturalist. org / observations / 28232546 • 1 ♂; Crocker Range, 89300 Tambunan; 5 ° 45 ′ 0 ″ N, 116 ° 30 ′ 0 ″ E; 24 Jun. 2019; paulsullivan 25 leg.; iNaturalist. org / observations / 28232543 • 1 ♀; Tambunan; 5 ° 45 ′ 10.9 ″ N, 116 ° 30 ′ 1.9 ″ E; 24 Jun. 2019; paulsullivan 25 leg.; iNaturalist. org / observations / 28232551 • 1 juvenile; 89300 Ranau; 6 ° 12 ′ 26.5 ″ N, 116 ° 38 ′ 27.2 ″ E; 2 Aug. 2019; a 1 virginia leg.; iNaturalist. org / observations / 30808860 • 1 ♀; Kinabalu Park, Ranau; 6 ° 0 ′ 19.4 ″ N, 116 ° 32 ′ 36.4 ″ E; 25 Oct. 2019; twan 3253 leg.; iNaturalist. org / observations / 35365957 • 1 juvenile; Tambunan; 5 ° 45 ′ 58.3 ″ N, 116 ° 20 ′ 32.9 ″ E; 31 Dec. 2019; F. Jarvis leg.; iNaturalist. org / observations / 37398325 • 1 ♂; Kinabalu Park, Ranau; 6 ° 0 ′ 31.1 ″ N, 116 ° 32 ′ 34.2 ″ E; 23 Nov. 2016; K. O’Brien leg.; iNaturalist. org / observations / 42450618 • around 20 individuals; Kinabalu Park, Ranau; 6 ° 0 ′ 22.9 ″ N, 116 ° 32 ′ 38.6 ″ E; 23 Nov. 2016; K. O’Brien leg.; iNaturalist. org / observations / 42450617 • 2 ♀♀; Kundasang; 6 ° 0 ′ 27.0 ″ N, 116 ° 32 ′ 1. 3 ″ E; 1 Feb. 2017; V. Volkova leg.; iNaturalist. org / observations / 51690441 • 1 ♀; Kinabalu Park, Ranau; 6 ° 1 ′ 45.5 ″ N, 116 ° 32 ′ 49.1 ″ E; 4 Jan. 2020; F. Jarvis leg.; iNaturalist. org / observations / 37711505 • 1 ♀; Ranau; 6 ° 1 ′ 36.8 ″ N, 116 ° 36 ′ 11.1 ″ E; 4 Jul. 2020; M. Haffiszul Mohd Said leg.; iNaturalist. org / observations / 51878361 • 1 ♀; Kinabalu Park, Ranau; 6 ° 1 ′ 38.3 ″ N, 116 ° 36 ′ 11.2 ″ E; 4 Jul. 2020; tanuki 18 leg.; iNaturalist. org / observations / 52011854 • 1 ♀; Kinabalu Park, Kota Belud; 6 ° 1 ′ 36.7 ″ N, 116 ° 32 ′ 51.8 ″ E; 13 Mar. 2019; D. Shaw leg.; iNaturalist. org / observations / 23278330.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD5FFC0FDBCF26CFD86FB63.taxon	description	Additional description (after Ingrisch 1987) Male GENITALIA (Fig. 16 F). Pseudepiphallus very slender and elongated, distinctly longer than wide, anterior and anterior margins gently concave. Pseudepiphallic lophi slightly longer than wide (at base), subacute and slightly folded dorsally at apex, sclerotized laterally only, widely spaced apart from each other. Rami straight, swollen preapically, slightly diverging. Pseudepiphallic parameres narrow, diverging posteriorly, their basal sclerite short and narrow. Ectophallic lateral sclerites appearing rounded. Ectophallic apodemes straight and parallel. Endophallic sclerite elongated, its posterior lateral arms short. Female GENITALIA. Copulatory papilla conical, smaller and stout; apex folded ventrally, elongated, pointed; dorsal face with a weakly sclerotized area; at ventro-anterior end a narrow oblong rim (Fig. 17 L – M). Juvenile (Fig. 28 D – E) Similar to adults in colour patterns, but younger instars differ by pronotal disk grey with dark longitudinal stripes, lateral lobes with a grey longitudinal stripe slightly ventral of the dorsal-third, sometimes in femora with white dorsal stripe, FIII with a white dorsal stripe at basal half. Legs darker in older instars, similar to adults. Measurements See Table 9.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD5FFC0FDBCF26CFD86FB63.taxon	biology_ecology	Ecology Adult and nymphal individuals have been observed to aggregate in large number (~ 20 individuals) on the same plant (Fig. 28 E).	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD5FFC0FDBCF26CFD86FB63.taxon	distribution	Distribution Borneo (Mount Kinabalu, Trus Madi, Tenompok, Tenom Keningau in Sabah) (Fig. 5).	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD5FFC0FDBCF26CFD86FB63.taxon	materials_examined	Type locality Borneo: Sabah: Mount Kinabalu.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD5FFC0FDBCF26CFD86FB63.taxon	description	Calling song (n = 1 male, 9 echemes, MNHN-SO- 2020 - 2857) The calling song consists of long homogeneous bouts formed of very short echemes consisting of doublets of syllables; echeme duration is 29.7 ± 1.1 ms (28.4 – 31.3 ms); echeme period is 72.2 ± 2.1 ms (69.8 – 76.8 ms); downtime between consecutive echemes is 42.5 ± 2.2 ms (39.4 – 45.9 ms). Duration for first and second syllables of the doublets are as follows: 7.5 ± 0.6 ms (6.6 – 8.1 ms) and 18.7 ± 1.5 ms (15.8 – 20.5 ms); with second syllable usually with greater amplitude than the first. The song spectrum consists of a series of harmonics, with two main peaks in the first two harmonics: first harmonic (fundamental frequency) which is also the dominant frequency peaks at 5.46 ± 0.03 (5.42 – 5.51) kHz; second harmonic peaking at 10.93 ± 0.04 (10.88 – 11.02) kHz.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD0FFC2FDD0F7BCFEECFB7A.taxon	description	urn: lsid: zoobank. org: act: A 3 AE 87 F 7 - 8196 - 4 C 9 B-BDEF- 8 E 9 D 21 B 92 BE 5 Figs 5, 9 C – D, 10 H, 11 H, 12 H, 14 H, 17 N – O; Tables 2, 10	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD0FFC2FDD0F7BCFEECFB7A.taxon	diagnosis	Diagnosis This species differs from all known congeners by the combination of these characters: dorsum of head cream-coloured with a black band or T-shaped marking; frons completely black; pronotal lateral lobes yellow, their anterior ventral corner black. Female FW dorsal field with cells almost completely black; veins yellow; longitudinal veins on dorsal field with many more cross-veins, creating a net-like mosaic; lateral field with M, R cream-coloured. This species is similar to N. insignis by pattern on head dorsum, but differs by frons completely black (instead of yellow), unique pronotum colouration and female FW lateral field with M and R cream-coloured (instead of dark coloured). This species is also similar to N. brunnerianus, which probably occurs in sympatry, and N. hughtani sp. nov. by dark frons, but differs by the characteristic pattern on dorsum of head and pronotal lateral lobe, and by female FW lateral field hyaline in ventral half. It also differs from the sympatric N. vittatus and N. crucius sp. nov. by the black frons, pronotum lateral lobe and FW colouration.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD0FFC2FDD0F7BCFEECFB7A.taxon	etymology	Etymology This species name refers to the word ʻrinduʼ, which means ʻloveʼ [verb] in Iban [a group of indigenous people from Borneo] language and ʻhome-sicknessʼ = ʻmissʼ [verb] in Bahasa Melayu (Malay language) and Indonesian. This new species is dedicated to the front-liners fighting against the COVID- 19 pandemic who were far from their homes and loved ones during the early phase of the global pandemic (when the speciesʼ name was chosen).	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD0FFC2FDD0F7BCFEECFB7A.taxon	materials_examined	Material examined Holotype MALAYSIA – Sarawak • ♀; Mount Murud; 6500 ft a. s. l.; 3 ° 54 ′ 28.44 ″ N, 115 ° 29 ′ 12.68 ″ E; 15 Apr. 1979; identified as Nisitra brunneriana by L. Chopard; MNHN-EO-ENSIF 10939. Paratypes INDONESIA – Kalimantan • 3 ♀; “ West Borneo, Midden ” [borders of Apokajan, Boeloengan and Beraoe]; 1 ° 58 ′ 43 ″ N, 116 ° 6 ′ 43 ″ E; 15 – 20 Oct. 1925; H. C. Siebers leg.; identified as Nisitrus brunnerianus by T. Robillard 2004; MNHN-EO-ENSIF 10940 to ENSIF 10942 • 1 ♀; Long Petak; 1 ° 42 ′ 6 ″ N, 116 ° 12 ′ 25 ″ E; 450 m a. s. l.; Sep. 1925; Brugs leg.; identified as Nisitrus brunnerianus by T. Robillard 2004; MNHN-EO-ENSIF 10943. MALAYSIA – Sabah • 1 ♀; near Long Pasia “ Long Pa Sia ”; 4 ° 24 ′ 22.05 ″ N, 115 ° 43 ′ 40.97 ″ E; 1020 m a. s. l.; 1 – 14 Apr. 1987; C. v. Achterberg leg.; RMNH • 1 ♀; same collection data as for preceding; PII: N 57; RMNH. – Sarawak • 1 ♀; Mount Tibang; 1 ° 39 ′ N, 114 ° 34 ′ E; 1700 m a. s. l.; E. G. Mjöberg leg.; MNHN-EO-ENSIF 10944.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD0FFC2FDD0F7BCFEECFB7A.taxon	description	Description Relatively large for the genus, colouration contrasted (Fig. 9 C – D). Vertex cream-coloured with a variable longitudinal black band and one or two faint transverse black bands forming a T- or ʻ + ʼ-shaped black pattern (Fig. 11 H). Scapes black, posterior end red brown. Antennae red brown with some segments black. Frons and face part of fastigium completely black, without yellow spots or bands (Fig. 10 H). Mouthparts dark red brown to black. Maxillary palpi cream-coloured (Fig. 10 H). Head lateral side black, sometimes with a small yellow spot at the ventro-posterior margin of eye (Fig. 12 H). Pronotal disk generally black with lateral and posterior margins yellow (Fig. 11 H); pronotal lateral lobe yellow, its ventro-anterior corner black (Fig. 12 H). Legs red brown. FIIIs dark red brown, their inner basal area black; knees dark brown to black; TIIIs brown with black spines and spurs, dark brown to black near distal end; tarsomeres dark brown to black. Hindwings hyaline brown, with a longitudinal transparent window near external-basal area; longer than FWs, forming a dark brown tail exceeding FWs more than twice as long as pronotum. Abdomen generally black. Male Unknown. Female FOREWINGS. FW dorsal field with cells almost completely black; veins almost always yellow (Fig. 14 H). Lateral field with M and R cream-coloured, sometimes with a cream-coloured patch at base; otherwise transparent or dark. FW venation: longitudinal veins on dorsal field with many cross-veins, creating a thin net-like mosaic. GENITALIA. Ovipositor distinctly longer than FIII; apex thin. Copulatory papilla conical, smaller and stout; apex folded ventrally, elongated, pointed; dorsal face with a sclerotized area; ventro-anterior end an oblong rim (Fig. 17 N – O). Measurements See Table 10.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD0FFC2FDD0F7BCFEECFB7A.taxon	biology_ecology	Ecology Unknown.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD0FFC2FDD0F7BCFEECFB7A.taxon	distribution	Distribution Borneo: Sabah, Sarawak and Kalimantan.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD0FFC2FDD0F7BCFEECFB7A.taxon	materials_examined	Type locality Malaysia: Sabah.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD0FFC2FDD0F7BCFEECFB7A.taxon	description	Calling song Unknown.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD2FFC2FF1FF7A6FCA7F811.taxon	materials_examined	Material examined Holotype INDONESIA – Sumatra • ♂; R. Weber leg.; AMNH. Type locality Indonesia: Sumatra.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD2FFC2FF1FF7A6FCA7F811.taxon	discussion	Remarks Based on the study of the type from Sumatra, we consider N. sumatrensis as a junior synonym of N. insignis, even though the colour pattern of the types of both species is slightly different. After the examination of the material from Sumatra identified “ N. sumatrensis ” and “ N. insignis ”, we observed that the colour patterns form a continuum between a lighter version in N. insignis and the darker, more contrasting specimens previously identified as in the type of N. sumatrensis. There are also little diagnostic differences in the male and female genitalia for both series.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD3FFB8FD89F26CFD84FCA3.taxon	description	Figs 1, 3, 5 – 6, 7 G, 9 E, 10 I, 11 I, 12 I, 13 G, 14 I, 16 G, 17 P – Q, 23 E – H, 24 D – F, 25 D – F, 26; Tables 1 – 2, 11	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD3FFB8FD89F26CFD84FCA3.taxon	diagnosis	Revised diagnosis Species of average size, colouration contrasting with black and yellow, characterised by black vertex, frons yellow, orange brown legs. Male area between M and R yellow, veins black or yellow; cells beneath R partly yellow, otherwise transparent. Anal-most and middle chords distinctly diverging apically, basalmost chord extend from midpoint / after midpoint of middle chord. Female copulatory papilla without truncated lobule at ventro-anterior end. Species very similar to N. malaya sp. nov., from which it differs by male genitalia with pseudepiphallus faintly less robust, narrower, anterior margin less concave; female copulatory papilla without truncated lobule at ventro-anterior end. Juvenile differs strikingly in colour patterns. Male calling song also differs by shorter echeme period.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD3FFB8FD89F26CFD84FCA3.taxon	materials_examined	Material examined Neotype INDONESIA – Sumatra • ♂; Wai Lima, “ Z. Sum. ” [South Sumatra], Lampongs; 5 ° 25 ′ 38.06 ″ S, 105 ° 01 ′ 22.37 ″ E; Nov. – Dec. 1921; Karny and Siebers leg.; RMNH. Additional specimens BRUNEI DARUSSALAM – Temburong • 1 ♂; Kuala Belalong Field Studies Centre; 4 ° 32 ′ 47 ″ N, 115 ° 9 ′ 27 ″ E; 8 Jan. 2017; M. K. Tan leg.; KB. 17.34; N 37; MNHN-EO-ENSIF 11069 • 1 ♀; Kuala Belalong Field Studies Centre; 4 ° 32 ′ 28.57 ″ N, 115 ° 09 ′ 28.94 ″ E; 23 Sep. 2016; M. K. Tan leg.; KB. 16.10; UBDM • 1 ♀; Kuala Belalong Field Studies Centre; 4 ° 32 ′ 28.57 ″ N, 115 ° 09 ′ 28.94 ″ E; 23 Sep. 2016; M. K. Tan leg.; KB. 16.11; UBDM • 1 ♀; same locality data as for preceding; 25 Jul. 2017; M. K. Tan leg.; KB. 17.53; UBDM • 1 juvenile; 4 ° 32 ′ 49.2 ″ N, 115 ° 9 ′ 20.3 ″ E; 7 Jan. 2014; J. Hodecek leg.; iNaturalist. org / observations / 46113125 • 1 ♂; 4 ° 32 ′ 49.2 ″ N, 115 ° 9 ′ 20.3 ″ E; 16 Jan. 2014; J. Hodecek leg.; iNaturalist. org / observations / 46113093. INDONESIA – Sumatra • 1 ♂; same collection data as for neotype; Karny leg.; MNHN-ENSIF 1742 • 1 ♀; same collection data as for neotype; MNHN-EO-ENSIF 3200 • 1 ♀; same collection data as for neotype; MNHN-ENSIF 3198 • 1 ♀; Pahoe; 22 Oct. [1925]; O. Posthumus leg.; MNHN-EO- ENSIF 3199 • 1 ♀; same collection data as for preceding; MNHN • 1 ♀; S Sumatra, SW Lampongs, Mt Tanggaamues; 1 ° 37 ′ 15.25 ″ S, 103 ° 36 ′ 23.79 ″ E; 600 m a. s. l.; Dec. 1934; Giesting, Lieftinck and Toxopeus leg.; MNHN-EO-ENSIF 3201 • 1 ♀; S Sumatra, Lalembang, Soeban Djerigi; 15 Jun. 1933; Soekarno leg.; MNHN-EO-ENSIF 3202 • 1 ♀; Damm [?] Muntok Banka; 2 ° 03 ′ 16.62 ″ S, 105 ° 10 ′ 43.11 ″ E; 25 Nov. [19] 23; MNHN-EO-ENSIF 3203 • 1 ♀; [Pidada] “ Pedada-B Lampongs ”; 5 ° 27 ′ 42.68 ″ S, 105 ° 19 ′ 20.91 ″ E; MNHN-EO-ENSIF 3204 • 6 ♂♂; “ southern Sumatra, Bengkulu, environs of town Curup (not far from city Bengkulu) ”; 3 ° 28 – 29 ′ S, 102 ° 31 – 38 ′ E; 24 Apr. – 2 May 2009; A. Gorochov, M. Berezin and E. Tkatsheva leg.; ZIN • 1 ♀; same collection data as for preceding; ZIN • 2 ♀♀; “ southern Sumatra, Lampung, National Park Way Kambas, ~ 60 km NE of city Tanjung Karang, camp Way Kenam ”; 5 ° 1.678 ′ S, 105 ° 46572 ′ E; 20 – 30 m a. s. l.; 13 – 14 Apr. 2009; A. Gorochov, M. Berezin and E. Tkatsheva leg.; ZIN • 1 ♂; same collection data as for preceding; ZIN • 7 ♂♂; “ Sumatra, southern Sumatra, environs of village Banding Agung on Lake Ranau Danau ”; 4 ° 48.695 ′ S, 103 ° 55.289 ′ E; 600 – 700 m a. s. l.; 19 – 29 Apr. 2009; A. Gorochov, M. Berezin and E. Tkatsheva leg.; ZIN • 3 ♀♀; “ Southern Sumtra, prov Lampung, National Park Way Kambas, ~ 60 km NE of city Tanjung Karang, camp Way Kenam ”; 5 ° 1.678 ′ S, 105 ° 46.572 ′ E; 20 – 30 m a. s. l.; 13 – 14 Apr. 2009; A. Gorochov, M. Berezin and E. Tkatsheva leg.; PII N 31; ZIN • 1 juvenile; “ Sumatra, southern Sumatra, environs of village Banding Agung on Lake Ranau Danau ”; 4 ° 48.695 ′ S, 103 ° 55.289 ′ E; 600 – 700 m a. s. l.; 19 – 29 Apr. 2009; A. Gorochov, M. Berezin and E. Tkatsheva leg.; ZIN • 1 ♂; “ southern Sumatra, Lampong, National Park Bukit Barisan Selatan, 15 – 20 km NW of town Krui, camp Kubu Prahu ”; 5 ° 4.341 ′ S, 104 ° 3.093 ′ E; ~ 600 m a. s. l.; 18 – 19 Apr. 2009; A. Gorochov, M. Berezin and E. Tkatsheva leg.; N 33; ZIN. – Java • 1 ♂; “ Java ”; H. D. Saussure leg.; MHN • 1 ♂; Padang; 0 ° 57 ′ 22.4 ″ S, 100 ° 25 ′ 5.36 ″ E; 21 Apr. 1929; Prince Leopold leg.; RBINS • 1 ♂; Tandjong-Karang; 0 ° 32 ′ 12.4 ″ S, 105 ° 15 ′ 29.88 ″ E; 21 Apr. 1929; Prince Leopold leg.; RBINS • 1 ♀; Sungai Penuh City, Jambi; 2 ° 7 ′ 53.7 ″ S, 101 ° 15 ′ 53.8 ″ E; 18 Jun. 2019; S. Wollney leg.; iNaturalist. org / observations / 27481225 • 1 ♂; Pesawaran Regency, Lampung; 5 ° 26 ′ 2.5 ″ S, 105 ° 9 ′ 57.6 ″ E; 9 Apr. 2019; J. - S. Wang leg.; iNaturalist. org / observations / 36723277 • 1 ♂; East Lampung Regency, Lampung; 5 ° 7 ′ 3.2 ″ N, 105 ° 44 ′ 16.6 ″ E; 7 Aug. 2019; pmaaskant leg.; iNaturalist. org / observations / 37027091 • 1 ♀; Sungai Penuh City, Jambi; 2 ° 7 ′ 56.8 ″ S, 101 ° 15 ′ 53.2 ″ E; 20 Jun. 2019; S. Wollney leg.; iNaturalist. org / observations / 27481235 • 1 ♀; Pesawaran Regency, Lampung; 5 ° 26 ′ 4.5 ″ S, 105 ° 10 ′ 29.3 ″ E; 9 Apr. 2019; J. - S. Wang leg.; iNaturalist. org / observations / 36723272 • 1 ♀; Eka Jaya, Kec. Jambi Sel., Kota Jambi, Jambi 36132; 1 ° 37 ′ 1.2 ″ S, 103 ° 40 ′ 16.8 ″ E; 7 Aug. 2020; suhendra leg.; iNaturalist. org / observations / 55695657 • 1 ♂; Banyu Asin, Riau Silip, Bangka Regency, Bangka Belitung Islands; 1 ° 53 ′ 3 ″ S, 105 ° 44 ′ 55 ″ E; 20 Oct. 2020; R. Maharani leg.; iNaturalist. org / observations / 63092557. – Kalimantan • 6 ♂; “ Marah ”; 0 ° 58 ′ 31.3 ″ N, 116 ° 33 ′ 14.8 ″ E; 10 – 28 Nov. 1925; H. C. Siebers leg.; MNHN • 3 ♂; “ Midden ” [near border of Koetai and Beraoe]; 1 ° 50 ′ 14 ″ N, 116 ° 12 ′ 34 ″ E; 26 Sep. 1925; H. C. Siebers leg.; MNHN • 1 ♂; “ Midden ” [near border of Koetai and Beraoe]; 1 ° 42 ′ 28 ″ N, 116 ° 15 ′ 40 ″ E; 4 Aug. 1925; H. C. Siebers leg.; MNHN • 1 ♂; “ Midden ”; 1925; H. C. Siebers leg.; MNHN 1730 • 1 ♂; “ Midden ” [near border of Koetai and Beraoe]; 1 ° 50 ′ 14 ″ N, 116 ° 12 ′ 34 ″ E; 27 Sep. 1925; H. C. Siebers leg.; MNHN • 1 ♀; “ Midden ” [near border of Koetai and Beraoe]; 1 ° 47 ′ 35 ″ N, 116 ° 12 ′ 22 ″ E; 12 Sep. 1925; H. C. Siebers leg.; MNHN • 1 juvenile; “ Midden ” [near border of Koetai and Beraoe]; 1 ° 50 ′ 14 ″ N, 116 ° 12 ′ 34 ″ E; 30 Sep. 1925; H. C. Siebers leg.; MNHN • 1 ♀; “ Kac. Pujungan Kayan Mentarang Nat. Reserve, lowland dipterocarp forest, WWF station, Lalut Birai Base Camp ”; 2 ° 52 ′ N, 115 ° 49 ′ E; 355 m a. s. l.; Jun. 1993; D. C. Darling and U. Rosichon leg.; MZB • 1 ♀; “ Karimata Island Nat. Keptapang Res., primary rainforest, canopy cover ”; 1 ° 25 ′ N, 106 ° 40 ′ E; 350 m a. s. l.; Feb. 1991; Sutrisno Djenal leg.; MZB • 1 ♂; Samarinda; 0 ° 31 ′ 54.17 ″ S, 117 ° 7 ′ 35.31 ″ E; 9 Feb. 1929; Prince Leopold leg.; RBINS • 2 ♀♀; “ Melawi ”; 0 ° 37 ′ 11.5 ″ S, 111 ° 45 ′ 4 ″ E; Nov. – Dec. 1924; Blanchmanche leg.; MNHN • 1 ♂; “ Hollandska Ostindien ”; Mrs M. - E. Walsh leg.; ZMLU • 2 ♀♀; same collection data as for preceding; ZMLU • 4 ♂♂; “ Samarinda ”; 0 ° 28 ′ 59 ″ S, 117 ° 8 ′ 33 ″ E; Mrs M. - E. Walsh leg.; ZMLU • 3 ♀♀; same collection data as for preceding; ZMLU • 1 ♂; [Tanggarong] “ Tanggarang ”; 0 ° 25 ′ 39 ″ S, 116 ° 59 ′ 16 ″ E; 5 Jul. 1957; Mrs M. - E. Walsh leg.; ZMLU • 1 ♂; same locality data as for preceding; 4 Jul. 1957; Mrs M. - E. Walsh leg.; ZMLU • 1 ♀; [Kaliorang] “ Kariorang ”; 0 ° 52 ′ 29 ″ N, 117 ° 56 ′ 31 ″ E; Apr. 1937; Mrs M. - E. Walsh leg.; ZMLU. MALAYSIA – Sabah • 1 ♂; N R Sandakan; 5 ° 50 ′ 38 ″ N, 117 ° 57 ′ 19 ″ E; 22 Jul. 1927; H. M. Pendlebury leg.; MNHN • 1 ♂; same collection data as for preceding; MNHN • 1 ♂; Mount Kinabalu, Kiau; 6 ° 1 ′ 40 ″ N, 116 ° 29 ′ 26 ″ E; 3000 ft a. s. l.; H. M. Pendlebury leg.; MNHN • 1 ♀; “ Crocker Range National Park not far from Keningau Town, secondary – primary forest ”; 5 ° 28 ′ 07.79 ″ N, 116 ° 04 ′ 22.95 ″ E; 1000 – 1300 m a. s. l.; 6 May 2013; A. Gorochov, M. Berezin, V. Gorochova and E. Tkatsheva leg.; N 32 - Nbo 5; ZIN • 1 ♂; “ Mulu National Park, not far from borders with Brunei and Indonesia ”; 4 ° 01 ′ 58.72 ″ N, 114 ° 49 ′ 00.56 ″ E; 100 – 300 m a. s. l.; 24 – 27 Mar. 2012; A. V. Gorochov, M. Berezin, E. Tkatsheva and I. Kamskov leg.; N 27 - Nbo 3; ZIN • 9 ♂♂; Mount Trus Madi, “ partly primary / partly secondary forest ”; 5 ° 33 ′ 53.94 ″ N, 116 ° 30 ′ 11.31 ″ E; ~ 1000 m a. s. l.; 13 – 25 May 2007; A. V. Gorochov leg.; ZIN • 10 ♀♀; Mount Trus Madi, “ partly primary / partly secondary forest ”; 5 ° 33 ′ 53.94 ″ N, 116 ° 30 ′ 11.31 ″ E; ~ 1000 m a. s. l.; 13 – 25 May 2007; A. V. Gorochov leg.; ZIN • 1 ♂; “ southern part of Kinabalu National Park, primary forest ”; 5 ° 59 ′ 50.78 ″ N, 116 ° 33 ′ 39.77 ″ E; ~ 1500 – 2000 m a. s. l.; 26 Apr. – 1 May 2013; A. Gorochov, M. Berezin, V. Gorochova and E. Tkatsheva leg.; PII N 25 - Nbo 1; ZIN • 1 ♂; “ Sandakan, environs of Sukau Village on Kinabatangan River (~ 35 km from sea), secondary – primary forest ”; 5 ° 51 ′ 21.27 ″ N, 117 ° 51 ′ 49.17 ″ E; 0 m a. s. l.; 8 – 13 May 2013; A. Gorochov, M. Berezin and E. Tkatsheva leg.; PII N 28 - Nbo 4; ZIN • 1 ♂; “ Tawau Hills National Park near Tawau City, secondary-primary forest ”; 4 ° 23 ′ 55.38 ″ N, 117 ° 53 ′ 20.60 ″ E; 200 – 400 m a. s. l.; 14 – 20 Apr. 2013; A. V. Gorochov, M. Berezin, E. Tkatsheva and I. Kamskov leg.; N 26 - Nbo 2; ZIN • 4 ♂♂; [Long Pasia] “ Long Pa Sia ”; 4 ° 24 ′ 22.05 ″ N, 115 ° 43 ′ 40.97 ″ E; ca 1025 m a. s. l.; 1 – 13 Apr. 1987; C. v. Achterberg leg.; RMNH • 2 ♀♀; same collection data as for preceding; RMNH • 2 ♀♀; Danum Valley Field Studies Centre; 4 ° 57 ′ 52.95 ″ N, 117 ° 48 ′ 18.25 ″ E; 14 – 20 Mar. 1987; C. v. Achterberg leg.; RMNH • 1 ♀; “ West of 3.1 km on Road Tenom – Keningau Depleted lowland forest; 5 ° 19 ′ 15.2 ″ N, 116 ° 13 ′ 2 ″ E; 240 m a. s. l.; 25 Jun. 1986; C. v. Achterberg leg.; RMNH • 2 ♂♂; Ranau “ west coast Residency, Ranau, 8 miles north Paring Hot Springs ”; 5 ° 50 ′ 47.95 ″ N, 116 ° 44 ′ 02.41 ″ E; 500 m a. s. l.; 8 – 11 Oct. 1958; L. W. Quate leg.; BPBM • 1 ♂; Tawau, Quoin Hill; 4 ° 23 ′ 55.38 ″ N, 117 ° 53 ′ 20.60 ″ E; 8 – 14 Jul. 1962; H. Holtmann leg.; BPBM • 1 ♂; Sandakan “ Sandakan Bay (NW), Sepilok Fore. Res. ”; 5 ° 50 ′ 38 ″ N, 117 ° 57 ′ 19 ″ E; 1 – 10 m a. s. l.; 27 Oct 1957; J. L. Gressitt leg.; BPBM • 1 ♂; [Crocker Range National Park] “ Croker Range n. p. ”; 5 ° 28 ′ 07.79 ″ N, 116 ° 04 ′ 22.95 ″ E; 11 – 17 Aug. 2003; “ Whiting, Svenson, Bybee ” leg.; PII: NviSA; MNHN-EO-ENSIF 1709 • 2 ♀♀; same collection data as for preceding; MNHN • 1 juvenile; same collection data as for preceding; MNHN • 1 ♂; “ Sipitang ”, “ Mendolong T 3 / W 5 ”; 4 ° 54 ′ 58 ″ N, 115 ° 45 ′ 60 ″ E; 25 Apr. 1988; S. Adebratt leg.; ZMLU • 1 ♂; “ Sipitang ”, “ Mendolong T 1 B / W 4 ”; 4 ° 54 ′ 58 ″ N, 115 ° 45 ′ 60 ″ E; 14 Mar. 1989; S. Adebratt leg.; ZMLU • 4 ♀♀; “ Sipitang ”, “ Mendolong T 1 B / W 4 ”; 4 ° 54 ′ 58 ″ N, 115 ° 45 ′ 60 ″ E; 14 Mar. 1989; S. Adebratt leg.; ZMLU • 1 ♀; “ Sipitang ”, “ Mendolong T 3 / W 5 ”; 4 ° 54 ′ 58 ″ N, 115 ° 45 ′ 60 ″ E; 31 Mar. 1989; S. Adebratt leg.; ZMLU • 2 ♀♀; “ Sipitang ”, “ Mendolong T 3 / W 5 ”; 4 ° 54 ′ 58 ″ N, 115 ° 45 ′ 60 ″ E; 9 Apr. 1988; S. Adebratt leg.; ZMLU • 1 ♀; “ Sipitang ”, “ Mendolong T 3 / W 5 ”; 4 ° 54 ′ 58 ″ N, 115 ° 45 ′ 60 ″ E; 24 Feb. 1989; S. Adebratt leg.; ZMLU • 2 ♀♀; “ Sipitang ”, “ Mendolong T 1 B / W 4 ”; 4 ° 54 ′ 58 ″ N, 115 ° 45 ′ 60 ″ E; 17 Mar. 1989; S. Adebratt leg.; ZMLU • 7 ♀♀; “ Sipitang ”, “ Mendolong T 2 B / W 5 ”; 4 ° 54 ′ 58 ″ N, 115 ° 45 ′ 60 ″ E; 19 Apr. 1988; S. Adebratt leg.; ZMLU • 1 juvenile; “ Sipitang ”, “ Mendolong T 5 / R ”; 4 ° 54 ′ 58 ″ N, 115 ° 45 ′ 60 ″ E; 28 Apr. 1988; S. Adebratt leg.; ZMLU • 1 ♂; “ Sipitang ”, “ Mendolong T 5 / R ”; 4 ° 54 ′ 58 ″ N, 115 ° 45 ′ 60 ″ E; 28 Apr. 1988; S. Adebratt; ZMLU • 1 juvenile; “ Batu Niah 18. mbbbb ”; 3 ° 48 ′ 14 ″ N, 113 ° 45 ′ 15 ″ E; Ulf Gärdenfors leg.; ZMLU • 1 ♀; “ Sipitang ” Mendolong; 4 ° 54 ′ 58 ″ N, 115 ° 45 ′ 60 ″ E; S. Adebratt leg.; ZMLU • 1 ♀; Kinabatangan; 5 ° 29 ′ 33.2 ″ N, 118 ° 12 ′ 23 ″ E; 30 Jul. 2018; lhurteau leg.; iNaturalist. org / observations / 15437369 • 1 ♂; Kota Kinabatangan; 5 ° 28 ′ 20.6 ″ N, 118 ° 15 ′ 28.8 ″ E; 7 Mar. 2013; rbeunen leg.; iNaturalist. org / observations / 18843166 • 1 ♂; Jalan Poring, 89300 Ranau; 6 ° 2 ′ 36.6 ″ N, 116 ° 42 ′ 10.6 ″ E; 6 Aug. 2013; doggy 0406 leg.; iNaturalist. org / observations / 24398754 • 1 ♂; Lahad Datu District; 4 ° 57 ′ 48.7 ″ N, 117 ° 40 ′ 20.3 ″ E; 17 Oct. 2019; N. K. Thangasamy leg.; iNaturalist. org / observations / 35226617 • 1 ♀; Lahad Datu; 4 ° 57 ′ 42.6 ″ N, 117 ° 41 ′ 21.1 ″ E; 27 Sep. 2016; S. Myers leg.; iNaturalist. org / observations / 17100119 • 1 ♂; Lahad Datu, Danum Valley Field Centre trails; 4 ° 58 ′ 15.8 ″ N, 117 ° 48 ′ 8.63 ″ E; 17 Jun. 2014; B. Crins leg.; iNaturalist. org / observations / 40612983 • 1 ♂; Kinabatangan; 5 ° 29 ′ 46.2 ″ N, 118 ° 16 ′ 39.7 ″ E; 26 Sep. 2018; z _ mcgrath leg.; iNaturalist. org / observations / 27490441 • 1 ♀; Bahagian Sandakan; 5 ° 22 ′ 0.8 ″ N, 117 ° 25 ′ 47.2 ″ E; 24 May 2018; K. Squires leg.; iNaturalist. org / observations / 38619342 • 1 ♀; Penampang; 5 ° 50 ′ 50.7 ″ N, 116 ° 5 ′ 27.8 ″ E; 28 Apr. 2020; tanuki 18 leg.; iNaturalist. org / observations / 44101645 • 1 juvenile; Penampang; 5 ° 50 ′ 50.6 ″ N, 116 ° 5 ′ 28 ″ E; 23 May 2020; tanuki 18 leg.; iNaturalist. org / observations / 46949300 • 1 juvenile; Tuaran; 5 ° 49 ′ 17.3 ″ N, 116 ° 20 ′ 28.8 ″ E; 6 Jun. 2020; tanuki 18 leg.; iNaturalist. org / observations / 48631961 • 1 ♀; Tuaran; 6 ° 0 ′ 34.1 ″ N, 116 ° 14 ′ 48.6 ″ E; 14 Jun. 2020; tanuki 18 leg.; iNaturalist. org / observations / 49562737 • 1 ♂; 90000 Kinabatangan; 5 ° 24 ′ 49.3 ″ N, 118 ° 2 ′ 15.5 ″ E; 7 Mar. 2020; M. de Groot leg.; iNaturalist. org / observations / 40232424 • 1 ♀, 1 ♂; 5 ° 13 ′ 0.40 ″ N, 118 ° 43 ′ 5.7 ″ E; 24 Jul. 2003; C. Shu leg.; iNaturalist. org / observations / 39450135. – Sarawak • 1 ♀; Mount Dulit, moss forest; 3 ° 19 ′ 58 ″ N, 114 ° 9 ′ E; 4000 ft a. s. l.; 22 Oct. 1932; B. M. Hobby and A. W. Moore leg.; MNHN • 1 ♂; Batu Niah; 3 ° 48 ′ N, 113 ° 45 ′ E; 4 – 8 Mar. 1984; S. Ingrisch leg.; SMTD • 2 ♀; “ Gunung Gading National Park, 80 – 90 km northwest of Kuching City ”; 1 ° 41 ′ 25.88 ″ N, 109 ° 50 ′ 44.68 ″ E; 100 – 300 m a. s. l.; 8 – 9 Mar. 2012; A. Gorochov, M. Berezin, E. Tkatsheva and I. Kamskov leg.; ZIN • 3 ♂♂; “ Kuching, environs of Kubah National Park, (situated on Matang Mountain and partly near it) ”; 1 ° 33 ′ 05.74 ″ N, 110 ° 18 ′ 54.26 ″ E; 200 – 500 m a. s. l.; 10 – 17 Mar. 2012; A. V. Gorochov, M. Berezin, E. Tkatsheva and I. Kamskov leg.; PII: N 24 Nku 1; ZIN • 1 ♀; “ Kuching, environs of Kubah National Park, (situated on Matang Mountain and partly near it) ”; 1 ° 33 ′ 05.74 ″ N, 110 ° 18 ′ 54.26 ″ E; 200 – 500 m a. s. l.; 10 – 17 Mar. 2012; A. V. Gorochov, M. Berezin, E. Tkatsheva and I. Kamskov leg.; ZIN • 1 ♂; Nanga Pelagus near Kapit; 1 ° 42 ′ 03.32 ″ N, 113 ° 32 ′ 32.38 ″ E; 180 – 585 m a. s. l.; 7 – 14 Oct. 1958; T. Maa leg.; BPBM • 1 ♂; Kapit District, Merirai Valley; 1 ° 53 ′ 52.61 ″ N, 113 ° 46 ′ 13.62 ″ E; 30 – 300 m a. s. l.; 1 – 6 Aug. 1958; T. Maa leg.; BPBM • 2 ♂♂; Tuba; 1 ° 16 ′ 15 ″ N, 111 ° 5 ′ 13 ″ E; 16 – 22 Jan. 1979; Gärdenfors, Hall, Hansson and Samuelsson leg.; ZMLU • 2 ♀♀; same collection data as for preceding; ZMLU • 1 ♀; “ 20 km SE Sebuyau, Tuba Forest clearing at longhouse ”; 1 ° 16 ′ 15 ″ N, 111 ° 5 ′ 13 ″ E; 15 – 22 Jan. 1979; Gärdenfors, Hall, Hansson and Samuelsson leg.; ZMLU • 1 ♀; Mount Matang, clearing; 1 ° 35 ′ 34 ″ N, 110 ° 11 ′ 25 ″ E; 1000 ft a. s. l.; 1 Jun. 1939; N. C. E. Miller leg.; NHMUK 012497573; • 1 ♀; Mount Dulit, moss forest; 3 ° 19 ′ 58 ″ N, 114 ° 9 ′ E; 4000 ft a. s. l.; 22 Oct. 1932; B. M. Hobby and A. W. Moore leg.; NHMUK 012497571 • 1 ♂; 1 ° 45 ′ 24.8 ″ N, 110 ° 19 ′ 11.9 ″ E; 15 May 2017; N. Vladimirov leg.; iNaturalist. org / observations / 8840806 • 1 ♂; Siburan; 1 ° 7 ′ 19.3 ″ N, 110 ° 12 ′ 56.8 ″ E; 21 Jun. 2013; flosker leg.; iNaturalist. org / observations / 19469334 • 1 ♀; Bau; 1 ° 22 ′ 52.2 ″ N, 110 ° 13 ′ 15.3 ″ E; 17 Sep. 2013; ml _ ls leg.; iNaturalist. org / observations / 28583992 • 1 ♂; Long Lellang Airport, 98008 Marudi; 3 ° 25 ′ 17.5 ″ N, 115 ° 9 ′ 9.5 ″ E; 2 Oct. 2016; Q. Lu leg.; iNaturalist. org / observations / 48562286 • 1 ♂; Gunung Serapi Kuching, 93050, Petra Jaya, 93050 Kuching; 1 ° 36 ′ 16.3 ″ N, 110 ° 11 ′ 19.6 ″ E; 19 Jul. 2019; plea 84 leg.; iNaturalist. org / observations / 29106017 • 1 ♀; Kuching; 1 ° 34 ′ 33.2 ″ N, 110 ° 17 ′ 54.1 ″ E; 18 Jul. 2017; C. Shu leg.; iNaturalist. org / observations / 39208711 • 1 ♀; Bahagian Miri; 3 ° 25 ′ 16.8 ″ N, 115 ° 9 ′ 15.8 ″ E; 30 Sep. 2018; K. Lin leg.; iNaturalist. org / observations / 20053192. NO COUNTRY • 1 ♂; “ South Borneo ”; H. Fruhstorfer leg.; MHN • 1 ♀; “ South Borneo ”; H. Fruhstorfer leg.; MHN • 1 ♀; “ South Borneo ”; H. Fruhstorfer leg.; MHN • 1 ♀; MHN • 1 ♀; MHN • 1 ♂; MNHN • 1 ♂; “ Borneo ”; Blanchmanche leg.; MNHN • 1 ♂; “ Borneo ”; MNHN • 1 ♀; “ Borneo ”; 1891; Chaper leg.; MNHN • 1 ♀; “ Borneo ”; 1891; Chaper leg.; MNHN • 1 ♂; “ Borneo ”; 1891; Chaper leg.; MNHN • 1 ♀; “ Borneo ”; Bruner leg.; MNHN • 1 ♂; “ Borneo ”; Bruner leg.; MNHN.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD3FFB8FD89F26CFD84FCA3.taxon	description	Redescription Size average (Figs 2, 23 E – H), similar to N. malaya sp. nov. but differs by slightly lighter colouration, male genitalia, female copulatory papilla and juvenile colouration. Male GENITALIA (Fig. 16 G). Pseudepiphallus relatively slender, slightly longer than broad, anterior margin straight, posterior margin indented in median area to faintly concave, lateral margins with basal half converge posteriorly, apical half somewhat parallel. Pseudepiphallic lophi slightly longer than wide (at base), sclerotized laterally only and covered with strong short setae; their apex slightly folded dorsally and sub-obtuse; moderately spaced apart from each other. Similar to N. malaya sp. nov. but differs by pseudepiphallus faintly more robust and broad, anterior margin more straight. Rami more straight. Female GENITALIA. Copulatory papilla conical, smaller and stout; apex folded ventrally, elongated, pointed; dorsal face with a sclerotized area; ventro-anterior end with an oblong rim, without truncated lobule (Fig. 17 P – Q). Juvenile (Fig. 24 D – F) Characterised by pale white or yellow colouration with numerous dark stripes all around body. Dorsum of head with three transverse black stripes that extends to gena. Eyes with numerous dark stripes. Fastigium with margin black; antennal black with pale rings; frons with a few black spots; palpal segments white with apices black. Pronotum with three thick, dark longitudinal stripes on dorsal disk, one very thick velvety black stripe between dorsal disk and lateral lobe; another curved stripe on lateral lobe ventral of the thick stripe; these stripes not reaching margins. Fore and middle legs with longitudinal dark stripes, two on each face, usually one dorsal and one ventral. FIII with longitudinal dark stripes, two dorsally, and two ventrally. Thoracic (excluding pronotum) and abdominal tergites typically dark or greyish, anterior margin dark, their posterior margin whitish forming a homogeneous light / dark striped pattern. Cerci whitish with a black stripe. Measurements See Table 11.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD3FFB8FD89F26CFD84FCA3.taxon	biology_ecology	Ecology This species can be found among shrubs and herbaceous plants (but not grasses) at the forest edge (Fig. 27 A). Similar to N. malaya sp. nov., the adults and nymphs of this species have been observed in Brunei Darussalam to feed on the flower parts of a varieties of native and exotic plant species (Tan et al. 2017).	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD3FFB8FD89F26CFD84FCA3.taxon	distribution	Distribution Sumatra, Borneo.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD3FFB8FD89F26CFD84FCA3.taxon	materials_examined	Type locality Indonesia: Sumatra: Lampongs, Wai Lima.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD3FFB8FD89F26CFD84FCA3.taxon	description	Calling song (Kuala Belalong, n = 2 males, 20 echemes; MNHN-SO- 2020 - 2859 to MNHN-SO- 2020 - 2860) (Fig. 25 D – F) At 27.0 ° C, the calling song consists of long homogeneous bouts made of short echemes consisting of triplets of syllables; echeme duration is 32.47 ± 3.79 ms (27.00 – 38.65 ms); echeme period is 55.20 ± 5.90 ms (50.52 – 74.27 ms); downtime between consecutive echemes is 22.73 ± 7.26 ms (14.16 – 41.82 ms). Durations for the first to third syllables of the triplet are as follows: 6.93 ± 1.73 ms (4.57 – 9.95 ms), 7.51 ± 2.03 ms (4.80 – 11.67 ms) and 7.22 ± 2.82 ms (3.24 – 12.44 ms). The call spectrum consists of a series of harmonics with three distinct peaks corresponding to the first three harmonics: first harmonic (fundamental frequency) at 6.86 ± 0.28 kHz (6.46 – 7.15 kHz), is usually the dominant frequency; second harmonic peaking at 14.08 ± 0.39 kHz (13.35 – 14.64 kHz); third harmonic peaking at 19.96 ± 0.35 kHz (19.12 – 20.67 kHz).	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
039487A4FFD3FFB8FD89F26CFD84FCA3.taxon	discussion	Remarks See N. malaya sp. nov. for arguments about considering two distinct species. About synonymy of Nisitrus marginatus (Walker, 1869) under N. vittatus, considering that N. vittatus is variable in colouration and widely distributed in Borneo and Sumatra, and that the only known syntype of N. marginatus from Sumatra resembles N. vittatus, we consider the species described by Walker (1869) as a junior synonym of N. vittatus.	en	Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C., Robillard, Tony (2021): Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species. European Journal of Taxonomy 761 (1): 1-75, DOI: 10.5852/ejt.2021.761.1449, URL: http://dx.doi.org/10.5852/ejt.2021.761.1449
