taxonID	type	description	language	source
03B7D31BFF1BFC2BFF7EFC67EBFBFC65.taxon	type_taxon	Type species. Clubiona dubia O. Pickard-Cambridge, 1870 from Saint Helena Island, by monotypy.	en	Zhang, Jianshuang, Marusik, Yuri M., Oketch, Ambata D., Kioko, Esther N., Yu, Hao, Li, Shuqiang (2021): Resurrection of the spider genus Bucliona Benoit, 1977, with a description of a new species from Kenya (Araneae, Clubionidae). Zootaxa 5006 (1): 195-207, DOI: https://doi.org/10.11646/zootaxa.5006.1.21
03B7D31BFF1BFC2BFF7EFC67EBFBFC65.taxon	diagnosis	Diagnosis. Bucliona differs from Clubiona sensu stricto (type species Araneus pallidulus Clerck, 1757) by: having a distinct mesal ridge on the male chelicerae (Fig. 2 A, C) (vs. lacking in Clubiona); the presence of a dorsal abdominal scutum in males (Figs 4 E, F, 6 A, C) (vs. absent); the undivided, short tibial apophysis of the male palp (Figs 3 B, 5 B, 7 B – D) (vs. divided in many species); the presence of a tegular (= median) apophysis and the lack of a meandering sperm duct (Figs 3 B – E, 5 A – E) (vs. lack of a tegular apophysis and presence of a meandering sperm duct); the subtegulum with a prolateral extension (Fig. 8 A) (vs. without, Figs 8 B – F); the tip of the embolus directed anteriorly or antero-mesally (Figs 3 D, 5 D, 7 A, B) (vs. retrolaterally or posteriorly). Females of Bucliona differ from those of Clubiona sensu stricto in having an epigyne with a distinct, undivided epigynal fovea with an anterior hood and a pair of posterior hoods (Figs 1 C, D, 4 A, C) (vs. fovea and hoods lacking) and longer copulatory ducts (Fig. 4 D). Both the males and females of Bucliona can be recognized by the anterior tibiae with a prolateral “ scopula ”, which is reduced to a single row (Fig. 2 D – F) (vs. dense scopula ventrally on the anterior tibiae).	en	Zhang, Jianshuang, Marusik, Yuri M., Oketch, Ambata D., Kioko, Esther N., Yu, Hao, Li, Shuqiang (2021): Resurrection of the spider genus Bucliona Benoit, 1977, with a description of a new species from Kenya (Araneae, Clubionidae). Zootaxa 5006 (1): 195-207, DOI: https://doi.org/10.11646/zootaxa.5006.1.21
03B7D31BFF1BFC2BFF7EFC67EBFBFC65.taxon	description	Description. Medium to large size, with the body length of males 4.90 – 7.75 and of females 6.50 – 9.50. Male. Body yellow-brown, legs uniformly coloured as carapace (Figs 4 E, F, 6 A – C). Carapace (Figs 4 E, F, 6 A, C): elongate-oval, widest at midpoint, pars cephalica slightly elevated above thorax, pars thoracica distinctly wider and slightly higher than pars cephalica; integument smooth; distinctly darker in ocular region, without distinct pattern; cervical groove and radial grooves distinct; fovea a short slit on posterior half of carapace, longitudinal and reddish. Clypeus height distinctly narrower than AME (Figs 2 A, C). Chelicerae (Figs 2 A, C, 4 E, F, 6 A – C): robust and red brownish, consisting of a coniform paturon and claw-shaped fang; cheliceral paturon with distinct mesal ridge, distally with several setae; fang furrow with 3 promarginal and 2 retromarginal teeth. Both endites and labium longer than wide; endites depressed posteriorly, slightly convergent anteriorly, with dense scopulae on inner margin; labium nearly trapezoidal and depressed laterally (Fig. 6 B). Sternum (Fig. 6 B): shield shaped and brown, longer than wide, anteriorly straight; posterior region protruding strongly between coxae IV. Eyes: in dorsal view, AER slightly recurved, PER almost straight (Figs 4 E, 6 A); in anterior view, AER almost straight, PER slightly recurved (Figs 2 A, C); AME very slightly smaller than ALE, or equal in diameter, AME closer to ALE than to each other; in dorsal view, PME and PLE nearly equal in diameter, well-separated, PME located about 2 – 3 diameters apart, PME – PLE distance ca. 2 / 3 PME – PME distance. Legs: leg formula 4213; anterior legs with conspicuous scopulae disto-prolaterally on tibiae, and entire metatarsi and tarsi (Figs 2 D, F); anterior femora with 3 ‒ 5 dorsal spines, posterior femora with 5 ‒ 7 dorsal spines; all patellae with 0 ‒ 1 dorsal or retrolateral spine; tibiae I ‒ II with 2 or 3 pairs of ventral spines, 1 retrolateral, and 1 prolateral spine; metatarsi I ‒ II with 1 or 2 pair of spines, 1 retrolateral, and 1 prolateral spine; tibiae and metatarsi of posterior legs with more spines than anterior legs but spination varies among different individuals. Abdomen (Figs 4 E, F 6 A – C): elongate-oval, tapering posteriorly, dorsal scutum over 3 / 4 of length of abdomen; without distinct colour patterns in the type species and B. jucunda (with median band and chevrons in B. kirilli sp. n., see description below). Palp: femur and patella unmodified, tibia cylindrical with single, unbranched retrolateral apophysis (RTA), apophysis shorter than diameter of tibia; cymbium unmodified, lacking spines; bulb elongate-oval; subtegulum (ST) with characteristic prolateral extension (Figs 3 A, C, 5 A, C, 8 A); sperm duct not meandering, U-shaped in ventral view of tegulum; tegulum with well-developed hook-shaped distal apophysis (TA); embolus helical, originating prolatero-distally, either filamentous and wrapping around dorsal side of tegulum (B. kirilli sp. n.) or shorter, tip directed either anteriorly (B. dubia) or antero-mesally (B. jucunda). Female. General characters as in males, from which the females differ by (1) the cheliceral base without a mesal ridge (Fig. 2 B); (2) the abdomen without a scutum (Figs 1 A, 4 G); and (3) a denser “ scopula ” on anterior legs (Fig. 2 E). Epigyne: epigynal plate with large fovea, located posteriorly, with 1 anterior (AH) and 2 posterior hoods (PH); copulatory openings (CO) small and indistinct, located in central portion of fovea; copulatory ducts (CD) hyaline, strongly convoluted; receptacles (R) small and located posteriorly; bursae (BS) situated anteriorly, balloon shaped, larger than receptacles. Composition. Bucliona dubia (Saint Helena Island), B. kirilli sp. n. (Kenya), and B. jucunda (Far East Russia, China, Korea, Japan).	en	Zhang, Jianshuang, Marusik, Yuri M., Oketch, Ambata D., Kioko, Esther N., Yu, Hao, Li, Shuqiang (2021): Resurrection of the spider genus Bucliona Benoit, 1977, with a description of a new species from Kenya (Araneae, Clubionidae). Zootaxa 5006 (1): 195-207, DOI: https://doi.org/10.11646/zootaxa.5006.1.21
03B7D31BFF19FC2AFF7EFF7BE9C5FDAA.taxon	description	Fig. 1	en	Zhang, Jianshuang, Marusik, Yuri M., Oketch, Ambata D., Kioko, Esther N., Yu, Hao, Li, Shuqiang (2021): Resurrection of the spider genus Bucliona Benoit, 1977, with a description of a new species from Kenya (Araneae, Clubionidae). Zootaxa 5006 (1): 195-207, DOI: https://doi.org/10.11646/zootaxa.5006.1.21
03B7D31BFF19FC2AFF7EFF7BE9C5FDAA.taxon	materials_examined	Type examined. Photograph of the holotype female from OUMNH, Mr. Melliss’s Saint Helena Island collection, May 1869.	en	Zhang, Jianshuang, Marusik, Yuri M., Oketch, Ambata D., Kioko, Esther N., Yu, Hao, Li, Shuqiang (2021): Resurrection of the spider genus Bucliona Benoit, 1977, with a description of a new species from Kenya (Araneae, Clubionidae). Zootaxa 5006 (1): 195-207, DOI: https://doi.org/10.11646/zootaxa.5006.1.21
03B7D31BFF19FC2AFF7EFF7BE9C5FDAA.taxon	diagnosis	Diagnosis and description. See Benoit (1977) and Mikhailov (1997). Epigyne and habitus of the holotype female are as in Fig. 1 A – D.	en	Zhang, Jianshuang, Marusik, Yuri M., Oketch, Ambata D., Kioko, Esther N., Yu, Hao, Li, Shuqiang (2021): Resurrection of the spider genus Bucliona Benoit, 1977, with a description of a new species from Kenya (Araneae, Clubionidae). Zootaxa 5006 (1): 195-207, DOI: https://doi.org/10.11646/zootaxa.5006.1.21
03B7D31BFF19FC2AFF7EFF7BE9C5FDAA.taxon	discussion	Comments. The holotype female was thought to be lost (Benoit 1977), however Zoë Simmons, the curator of OUMNH has managed to find the vial with the holotype female.	en	Zhang, Jianshuang, Marusik, Yuri M., Oketch, Ambata D., Kioko, Esther N., Yu, Hao, Li, Shuqiang (2021): Resurrection of the spider genus Bucliona Benoit, 1977, with a description of a new species from Kenya (Araneae, Clubionidae). Zootaxa 5006 (1): 195-207, DOI: https://doi.org/10.11646/zootaxa.5006.1.21
03B7D31BFF19FC2AFF7EFF7BE9C5FDAA.taxon	distribution	Distribution. The species is known from Saint Helena Island only.	en	Zhang, Jianshuang, Marusik, Yuri M., Oketch, Ambata D., Kioko, Esther N., Yu, Hao, Li, Shuqiang (2021): Resurrection of the spider genus Bucliona Benoit, 1977, with a description of a new species from Kenya (Araneae, Clubionidae). Zootaxa 5006 (1): 195-207, DOI: https://doi.org/10.11646/zootaxa.5006.1.21
03B7D31BFF19FC21FF7EFD5DEF3CF9E3.taxon	description	Figs 2 A ‒ B, D ‒ E, 3, 4, 7 B, D	en	Zhang, Jianshuang, Marusik, Yuri M., Oketch, Ambata D., Kioko, Esther N., Yu, Hao, Li, Shuqiang (2021): Resurrection of the spider genus Bucliona Benoit, 1977, with a description of a new species from Kenya (Araneae, Clubionidae). Zootaxa 5006 (1): 195-207, DOI: https://doi.org/10.11646/zootaxa.5006.1.21
03B7D31BFF19FC21FF7EFD5DEF3CF9E3.taxon	materials_examined	Material examined. CHINA: Hubei: Xiantao City, Shahu Town, Shishangang Village, Shahu National Wetland Park (N 30.1588 º, E 117.7179 º, 15 m), 2 ♂ 5 ♀ (MGEU), February 2011, H. Yu. leg.	en	Zhang, Jianshuang, Marusik, Yuri M., Oketch, Ambata D., Kioko, Esther N., Yu, Hao, Li, Shuqiang (2021): Resurrection of the spider genus Bucliona Benoit, 1977, with a description of a new species from Kenya (Araneae, Clubionidae). Zootaxa 5006 (1): 195-207, DOI: https://doi.org/10.11646/zootaxa.5006.1.21
03B7D31BFF19FC21FF7EFD5DEF3CF9E3.taxon	diagnosis	Diagnosis. Bicluona jucunda is similar to the generotype, B. dubia (Mikhailov 1997: 96, figs 1 ‒ 4). Males of the two species can be distinguished by the shape of the embolus and RTA. In B. jucunda, the embolus is filamentous, wraps around the tegulum, and the tip is directed antero-mesally, while in B. dubia, the embolus is shorter and stouter, Ƨ-shaped, and the tip is directed anteriorly (cf. Figs 3 C – E, 7 B and Mikhailov 1997: fig. 1). The RTA in B. jucunda has a blunt tip vs. pointed in B. dubia (cf. Figs 3 B, 7 D and Mikhailov 1997: fig. 2). Females of B. jucunda can be distinguished from those of B. dubia by the distinctly longer and strongly convoluted copulatory ducts (Fig. 4 B, D) (vs. relatively shorter, see Mikhailov 1997: fig. 4) and the smaller receptacles, less than 1 / 2 length and 1 / 3 width of epigynal fovea (Fig. 4 A – D) (vs. comparatively larger, length subequal to fovea; see Mikhailov 1997: figs 3, 4).	en	Zhang, Jianshuang, Marusik, Yuri M., Oketch, Ambata D., Kioko, Esther N., Yu, Hao, Li, Shuqiang (2021): Resurrection of the spider genus Bucliona Benoit, 1977, with a description of a new species from Kenya (Araneae, Clubionidae). Zootaxa 5006 (1): 195-207, DOI: https://doi.org/10.11646/zootaxa.5006.1.21
03B7D31BFF19FC21FF7EFD5DEF3CF9E3.taxon	description	Description. Male. Total length 6.70; carapace 3.05 long, 2.20 wide; abdomen 3.65 long, 1.97 wide. Carapace (Fig. 4 E, F) dark yellow with darkened eye region, without distinct pattern; cervical groove and radial grooves indistinct. Labium and endites dark reddish brown. Sternum yellow. Eyes: AER slightly recurved, PER wider than AER and straight in dorsal view. AME dark, other eyes light; with black rings. Eye sizes and interdistances: AME 0.12, ALE 0.15, PME 0.12, PLE 0.14, AME – AME 0.16, AME – ALE 0.11, PME – PME 0.36, PME – PLE 0.22, MOQL 0.41, MOQA 0.40, MOQP 0.63. Legs yellowish, without distinct markings. Leg measurements: I 5.84 (1.53, 2.60, 1.05, 0.67), II 6.29 (1.83, 2.62, 1.12, 0.71), III 4.95 (1.46, 1.70, 1.26, 0.53), IV 6.94 (1.96, 2.37, 1.91, 0.70). Abdomen (Figs 4 E, F), dorsally with a narrow scutum extending ca. 1 / 2 of abdomen length, gradually widening posteriorly, with two pairs of inconspicuous sigilla on either side; venter white with no distinct pattern. Palp (Figs 3 A – E, 7 B, D): Tibia comparatively short, ca. 1 / 2 of cymbium length, ca. 1.7 times longer than wide, retrolateral tibial apophysis (RTA) about 1 / 4 of tibia length, about 1.5 times longer than wide, thumb shaped. Embolus filamentous, wrapping around dorsal side of tegulum, tip directed antero-mesally, base (EB) slightly shorter than free (filamentous) part of (E). Tegular apophysis (TA) originating from antero-retrolateral side of tegulum, semicircular, directed antero-mesally, distally curved postero-retrolaterally. Female. (Fig. 4 G, H): total length 6.50; carapace 2.88 long, 2.16 wide; abdomen 3.57 long, 2.20 wide. Eye sizes and interdistances: AME 0.12, ALE 0.14, PME 0.12, PLE 0.010, AME – AME 0.13, AME – ALE 0.15, PME – PME 0.38, PME – PLE 0.26, MOQL 0.39, MOQA 0.37, MOQP 0.57. Sternum 1.49 long, 1.01 wide. Leg measurements: I 4.99 (1.44, 2.112, 0.87, 0.571), II 5.54 (1.51, 2.31, 1.08, 0.64), III 4.50 (1.40, 1.54, 1.048, 0.51), IV 6.64 (1.75, 2.32, 1.90, 0.67). Slightly smaller and distinctly darker than male, other characters as in male. Epigyne (Figs 4 A – D): Epigynal plate distinctly longer than wide, anterior and lateral margin not rebordered, part of bursae and receptacles distinctly visible through plate. Epigynal fovea triangular, with rebordered margins, ca. 1 / 2 of epigyne length and 2 / 3 of epigyne width; both anterior (AH) and posterior hoods (PH) heavily sclerotized. Copulatory openings (CO) small and indistinct, on posterior part of fovea. Copulatory ducts (CD) hyaline and strongly convoluted. Receptacles (R) small, bean shaped, ca. 1.9 times longer than wide. Bursae (BS) situated anteriorly, balloon shaped, relatively large, ca. 1.3 times longer than wide, anterior surface wrinkled; two bursae close together. Fertilisation ducts (FD) acicular and membranous, curved, on posterior surface of receptacles, separated from epigastric furrow by about 1 / 3 epigyne length.	en	Zhang, Jianshuang, Marusik, Yuri M., Oketch, Ambata D., Kioko, Esther N., Yu, Hao, Li, Shuqiang (2021): Resurrection of the spider genus Bucliona Benoit, 1977, with a description of a new species from Kenya (Araneae, Clubionidae). Zootaxa 5006 (1): 195-207, DOI: https://doi.org/10.11646/zootaxa.5006.1.21
03B7D31BFF19FC21FF7EFD5DEF3CF9E3.taxon	distribution	Distribution. Russia (south part of the Far East), China (from Heilongjiang to Guizhou), Korea, Japan.	en	Zhang, Jianshuang, Marusik, Yuri M., Oketch, Ambata D., Kioko, Esther N., Yu, Hao, Li, Shuqiang (2021): Resurrection of the spider genus Bucliona Benoit, 1977, with a description of a new species from Kenya (Araneae, Clubionidae). Zootaxa 5006 (1): 195-207, DOI: https://doi.org/10.11646/zootaxa.5006.1.21
03B7D31BFF12FC22FF7EF998E881FC40.taxon	description	Figs 2 E, F, 5, 6, 7 A, C	en	Zhang, Jianshuang, Marusik, Yuri M., Oketch, Ambata D., Kioko, Esther N., Yu, Hao, Li, Shuqiang (2021): Resurrection of the spider genus Bucliona Benoit, 1977, with a description of a new species from Kenya (Araneae, Clubionidae). Zootaxa 5006 (1): 195-207, DOI: https://doi.org/10.11646/zootaxa.5006.1.21
03B7D31BFF12FC22FF7EF998E881FC40.taxon	materials_examined	Type material. Holotype ♂ (NMK), KENYA: Nyeri County, Naro Moru Town, Mount Kenya National Park, Metrological Station (S 0.1702 º, E 37.2140 º, 3000 m), 16. VIII. 2018, Kioko G. leg. Other material examined. KENYA: Laikipia County, Nanyuki Town, Mount Kenya National Park, Sirimon Gate, Bamboo Forest (S 0.2675 º, E 37.28276 º, 3176 m), 1 ♂ (MGEU), 17. VIII. 2018, Kioko G. leg.	en	Zhang, Jianshuang, Marusik, Yuri M., Oketch, Ambata D., Kioko, Esther N., Yu, Hao, Li, Shuqiang (2021): Resurrection of the spider genus Bucliona Benoit, 1977, with a description of a new species from Kenya (Araneae, Clubionidae). Zootaxa 5006 (1): 195-207, DOI: https://doi.org/10.11646/zootaxa.5006.1.21
03B7D31BFF12FC22FF7EF998E881FC40.taxon	etymology	Etymology. The species is named in honour of Kirill G. Mikhailov from Moscow State University for his contributions to the systematics of clubionid spiders and on the occasion of his 60 th birthday; noun (name) in genitive case.	en	Zhang, Jianshuang, Marusik, Yuri M., Oketch, Ambata D., Kioko, Esther N., Yu, Hao, Li, Shuqiang (2021): Resurrection of the spider genus Bucliona Benoit, 1977, with a description of a new species from Kenya (Araneae, Clubionidae). Zootaxa 5006 (1): 195-207, DOI: https://doi.org/10.11646/zootaxa.5006.1.21
03B7D31BFF12FC22FF7EF998E881FC40.taxon	diagnosis	Diagnosis. Males of B. kirilli sp. n. resemble those of B. dubia but differ by the parallel sides and a triangular tip of the RTA (vs. wide base and claw-like tip), as well as by the tip of the embolus which is directed antero-mesally (vs. anteriorly).	en	Zhang, Jianshuang, Marusik, Yuri M., Oketch, Ambata D., Kioko, Esther N., Yu, Hao, Li, Shuqiang (2021): Resurrection of the spider genus Bucliona Benoit, 1977, with a description of a new species from Kenya (Araneae, Clubionidae). Zootaxa 5006 (1): 195-207, DOI: https://doi.org/10.11646/zootaxa.5006.1.21
03B7D31BFF12FC22FF7EF998E881FC40.taxon	description	Description. Male. Total length 7.66; carapace 3.65 long, 2.54 wide; abdomen 4.01 long, 2.14 wide. Carapace (Figs 6 A – C) uniformly yellow-brown. Labium and endites dark reddish brown. Sternum coloured as carapace. Eyes: AER slightly recurved, PER wider than AER and straight in dorsal view. AME dark, other eyes light; with black rings. Eye sizes and interdistances: AME 0.17, ALE 0.20, PME 0.17, PLE 0.18, AME – AME 0.14, AME – ALE 0.15, PME – PME 0.39, PME – PLE 0.28, MOQL 0.47, MOQA 0.44, MOQP 0.71. Legs yellowish, without distinct pattern. Leg measurements: I and II missing, III (2.50, —, —, —), IV 11.45 (3.20, 3.95, 3.33, 0.97). Abdomen (Figs 6 A – C) with wide dorsal scutum covering almost entire abdomen; dorsum anteriorly with triangular, dark, median stripe, reaching 1 / 3 of abdomen length, centrally without distinct patterns (faded in the preserved holotype but with 2 transverse chevrons in the second female), posteriorly with 3 transverse chevrons; venter anteriorly without any markings, posteriorly with 2 broken longitudinal stripes. Palp (Figs 5 A – E, 7 A, C): Tibia relatively long, over 1 / 2 cymbium length, ca. 1.9 times longer than wide, retrolateral tibial apophysis (RTA) about 1 / 2 of tibia length, about 5 times longer than wide, slightly wider at base, gradually tapering towards apex, tip triangular, sharply pointed. Embolus Ƨ-shaped in ventral view, bent at almost a right angle, tip directed antero-mesally, base (EB) slightly longer than free part (E), embolus 6.6 times longer than wide. Tegular apophysis (TA) heavily sclerotized, arising retrolatero-distally from tegulum, finger shaped, directed proximally, distally curved retrolaterally. Female. Unknown.	en	Zhang, Jianshuang, Marusik, Yuri M., Oketch, Ambata D., Kioko, Esther N., Yu, Hao, Li, Shuqiang (2021): Resurrection of the spider genus Bucliona Benoit, 1977, with a description of a new species from Kenya (Araneae, Clubionidae). Zootaxa 5006 (1): 195-207, DOI: https://doi.org/10.11646/zootaxa.5006.1.21
03B7D31BFF12FC22FF7EF998E881FC40.taxon	distribution	Distribution. Known only from the type locality.	en	Zhang, Jianshuang, Marusik, Yuri M., Oketch, Ambata D., Kioko, Esther N., Yu, Hao, Li, Shuqiang (2021): Resurrection of the spider genus Bucliona Benoit, 1977, with a description of a new species from Kenya (Araneae, Clubionidae). Zootaxa 5006 (1): 195-207, DOI: https://doi.org/10.11646/zootaxa.5006.1.21
