taxonID	type	description	language	source
2F1787E8FFD88274FEBEFE8E8F4DF3D5.taxon	diagnosis	Diagnosis. During the evolution toward a canopy domain and away from a likely understory sistergroup (Erwin 1979, 1985, 1998), Agra adults acquired numerous generic-level autapotypic features: head elongate (cranium extended behind eyes), with prognathous mandibles, securiform ultimate labial palpomeres (Erwin 1982), and constricted neck; prothorax elongate and tubular, pleural sutures effaced; tarsomeres dilated with setiferous pads beneath (Erwin 1982), claws explanate and pectinate; elytron with latero-basal sinus and latero-apical callus, apex medially and laterally toothed or somewhat produced, apical margin truncate, sinuate, or medially lobed; male venter variously adorned with setal or pubescent patches; female reproductive system adapted to egg-laying telescopically deep in substrate (Erwin 2002), stylomere 2 stout (usually and with apical ensiform setae. Defense system very large (Erwin 1982). Size: ABL = 6.0 – 29.0 mm; TW = 1.5 – 6.0 mm. Note. For descriptions of species-groups and subgroups previously recognized, refer to Erwin (1996, 1998, 2002).	en	Erwin, Terry L. (2017): Agra Fabricius (Coleoptera: Carabidae: Harpalinae: Lebiini: Agrina), Arboreal Beetles of Neotropical Forests: The Rare, Non-Neotropical Texas Species at the Generic Northern Limit, with Notes on Their Way of Life. The Coleopterists Bulletin 71 (4): 639-651, DOI: 10.1649/0010-065X-71.4.639, URL: http://dx.doi.org/10.1649/0010-065x-71.4.639
2F1787E8FFD88274FEABFAC68DE5F5C7.taxon	description	Beetles of this rather uniform group are of medium to moderately large size and brown to black; some have aeneous to submetallic elytra in Costa Rica. A diagnostic combination of an oblongopunctata species-group adult includes the following: Labrum flat; male antenna with antennomeres more or less subequal in length (except short pedicel); female antenna with antennomere 8 short, about half the length of 7, and 9 – 11 somewhat shorter than 3 – 7; antenna of normal length reaching level of middle coxa, not markedly short and robust, nor long and markedly thin; head behind eyes elongate, evenly rounded posteriorly, more tapered to neck in the male; elytral interneurs each a series of medium to large variously separated punctures; elytral apex obliquely truncated, laterally with a small tooth, suturally obtuse; legs and tarsi normal, not markedly modified; male metasternum sparsely setigerous, abdominal sterna III – V bilaterally setigerous, setae in a patch; females simply sparsely setiferous.	en	Erwin, Terry L. (2017): Agra Fabricius (Coleoptera: Carabidae: Harpalinae: Lebiini: Agrina), Arboreal Beetles of Neotropical Forests: The Rare, Non-Neotropical Texas Species at the Generic Northern Limit, with Notes on Their Way of Life. The Coleopterists Bulletin 71 (4): 639-651, DOI: 10.1649/0010-065X-71.4.639, URL: http://dx.doi.org/10.1649/0010-065x-71.4.639
2F1787E8FFD88274FEABFAC68DE5F5C7.taxon	description	Eleven species are currently recognized in the species-group:	en	Erwin, Terry L. (2017): Agra Fabricius (Coleoptera: Carabidae: Harpalinae: Lebiini: Agrina), Arboreal Beetles of Neotropical Forests: The Rare, Non-Neotropical Texas Species at the Generic Northern Limit, with Notes on Their Way of Life. The Coleopterists Bulletin 71 (4): 639-651, DOI: 10.1649/0010-065X-71.4.639, URL: http://dx.doi.org/10.1649/0010-065x-71.4.639
2F1787E8FFD8827EFCB4FD308AB9F623.taxon	description	Riley’ s elegant canopy beetle	en	Erwin, Terry L. (2017): Agra Fabricius (Coleoptera: Carabidae: Harpalinae: Lebiini: Agrina), Arboreal Beetles of Neotropical Forests: The Rare, Non-Neotropical Texas Species at the Generic Northern Limit, with Notes on Their Way of Life. The Coleopterists Bulletin 71 (4): 639-651, DOI: 10.1649/0010-065X-71.4.639, URL: http://dx.doi.org/10.1649/0010-065x-71.4.639
2F1787E8FFD8827EFCB4FD308AB9F623.taxon	description	(Figs. 1, 3, 4)	en	Erwin, Terry L. (2017): Agra Fabricius (Coleoptera: Carabidae: Harpalinae: Lebiini: Agrina), Arboreal Beetles of Neotropical Forests: The Rare, Non-Neotropical Texas Species at the Generic Northern Limit, with Notes on Their Way of Life. The Coleopterists Bulletin 71 (4): 639-651, DOI: 10.1649/0010-065X-71.4.639, URL: http://dx.doi.org/10.1649/0010-065x-71.4.639
2F1787E8FFD8827EFCB4FD308AB9F623.taxon	materials_examined	Holotype. USA, TEXAS: Cameron County, Paloma Blanca Road, near Sabal Palm Groove, 9 m, 25.8663 ° N, 97.4343 ° W, 26 October 1991 (EG Riley) (NMNH: ADP 109066, male). Derivation of Specific Epithet. The epithet “ rileyi ” is an eponym based on the family name of the Texas Coleopterist, Edward G. Riley, at Texas A & M University, who collected adults of this species near and at the type locality. Proposed English Vernacular Name. Riley’ s elegant canopy beetle.	en	Erwin, Terry L. (2017): Agra Fabricius (Coleoptera: Carabidae: Harpalinae: Lebiini: Agrina), Arboreal Beetles of Neotropical Forests: The Rare, Non-Neotropical Texas Species at the Generic Northern Limit, with Notes on Their Way of Life. The Coleopterists Bulletin 71 (4): 639-651, DOI: 10.1649/0010-065X-71.4.639, URL: http://dx.doi.org/10.1649/0010-065x-71.4.639
2F1787E8FFD8827EFCB4FD308AB9F623.taxon	diagnosis	Diagnosis. This species has all the attributes of the genus and species-group as described above and is small for the oblongopunctata species-group. Adults with brown integument, elytra slightly aeneous with a subtly greenish tint at bottom of punctures in interneurs; head behind eyes and prothorax dark brown. Frons and occiput slightly domed, without punctures, with scattered fine setae. Pronotal disc with 4 rows of longitudinal punctures, 2 rows of larger punctures adjacent to midline each side, and a row of smaller punctures along each lateral margin; a few randomly spaced, very small punctures on smooth areas. Elytral apex obliquely truncate, lateral corner slightly obtuse, not dentate, sutural corner rounded. Interneurs with rounded or slightly elongate punctures, without even spaces between puncture.	en	Erwin, Terry L. (2017): Agra Fabricius (Coleoptera: Carabidae: Harpalinae: Lebiini: Agrina), Arboreal Beetles of Neotropical Forests: The Rare, Non-Neotropical Texas Species at the Generic Northern Limit, with Notes on Their Way of Life. The Coleopterists Bulletin 71 (4): 639-651, DOI: 10.1649/0010-065X-71.4.639, URL: http://dx.doi.org/10.1649/0010-065x-71.4.639
2F1787E8FFD8827EFCB4FD308AB9F623.taxon	description	Description. Size: Small for species-group, ABL = 10.4 – 14.3 mm, SBL = 8.98 – 13.04 mm, TW = 2.65 – 3.89 mm (Table 1). Color: As described above and antennomeres 1 – 5 brownish, 6 – 11 dark testaceous with apical brown rings; mouthparts brownish, palpomere 4 with pale apex, and legs and tarsi brown. Luster: Head, pronotum, and legs shiny metallic, elytra brown aeneous. Head: As described above. Prothorax: Elongate, moderately constricted near base, devoid of hind angles, narrowed anteriorly to about width of neck; surface of disc as described above (Fig. 1). Pterothorax: Elytron moderately convex, narrow in anterior third, moderately flared from middle to apical third and rounded to lateral hind angle; intervals slightly convex, somewhat irregular. Legs: Normal in both sexes (see Erwin 2002). Abdomen: As described above. Male genitalia: Phallus (Fig. 3) with phalloshaft elongate and narrow apically, widened to phallobase which is at about 45 ° angle from phalloshaft, with ostium modestly elongate, about 1 / 4 length of phallus, apex of phalloshaft small and arrow-shaped with rounded lateral corners; endophallus without sclerotized features. Parameres small, left twice the size of the right, both broadly rounded. Female ovipositor: Female internal parts not investigated (but see Erwin 1982). Stylomere 2 as in Fig. 3 C. Dispersal Potential. These beetles are macropterous and capable of flight; they are attracted to UV light. They are swift and agile runners. They ‘ sleep’ in the daytime under curled or dry leaves, aligned with the mid-rib of the leaf. Way of Life. Adults of other Agra species are found in the canopy of rainforest trees; larvae of species in this genus are found under the bark of these trees (Arndt et al. 2001), however, they must also roam on the surface during the night, as they have been collected by insecticidal fogging techniques in the very early morning before first light. Members of A. rileyi occur at lowland elevations along the Rio Grande in southeastern Texas where sabal palms and sugar hackberry grow. Adults are active in the spring, summer, and fall, mostly in the rainy season of September and October. Other Specimens Examined. USA: TEXAS, Cameron County, Brownsville, Esperanza Ranch, 8 m, 25.8906 ° N, 97.4502 ° W, 25 July 1899 (EC Van Dyke) (CAS: ADP 091220, male paratype), 19 August 1899 (BMC: ADP 091221, female paratype), 26 June 1899 (BMC: ADP 091222, male paratype), 10 June 1899 (BMC: ADP 091223, male paratype), 8 August 1899 BMC: ADP 091231, female paratype), 28 - 29 September 2004 (B Raber) (BTRC: ADP 109102, ADP 109104, male paratypes); Cameron County, Brownsville, southernmost sector, Sabal Palm Grove Sanctuary, 8 m, 25.8419 ° N, 97.4247 ° W, 28 - 29 September 2004 (B Raber) (BTRC: ADP 109100, male), 29 September 2004 (B Raber / B Smith) (CMNH: ADP 116793, female paratype, ADP 116281, male paratype), 28 September 2004 (B Raber / B Smith) (CMNH: ADP 116279, female paratype), 18 - 19 October 2002 (BT Raber) (BTRC: ADP 109106, male paratype), Brownsville, southmost sector, Sabal Palm Grove Sanctuary, 13 - 17 May 1977 (FT Hovore) (FTHC: ADP 124942, female paratype), 7 - 11 October 1975 (FT Hovore) (FTHC: ADP 054437, female paratype), 18 October 1985 (JE Wappes) (JEWC: ADP 109110, female paratype), 21 October 1993 (JE Wappes) (JEWC: ADP 109112, male paratype) (JE Wappes) (TAMU: ADP 109068, male paratype), Brownsville, 8 m, 25.8906 ° N, 97.4502 ° W, (JW Green) (CAS: ADP 091224, female paratype), 8 May 1904 (HS Barber) (NMNH: ADP 091225, male paratype), 5 - 8 October 1967 (N Rolien) (FSCA: ADP 113638, female paratype), 27 June 1899 (CHT Townsend) (NMNH: ADP 091226, male paratype, ADP 091227, female paratype), – July —— (HF Wickham) (MCZ: ADP 091228, male paratype), – July —— (HF Wickham) (FSCA: ADP 085333, male paratype), 20 July 1899 (HF Wickham) (FSCA: ADP 085312, male paratype), Cameron County, Sabal Palm Grove nr. Southmost, 29 September 1976 (JE Wappes) (JEWC: ADP 109116, male paratype), Cameron County, Sabal Palm Grove, 9 - 10 June 1978 (JE Wappes) (JEWC: ADP 004768, male paratype), 14 April 1979 (JE Wappes) (JEWC: ADP 062418, male paratype, ADP 058560, female paratype), 10 October 1981 (JE Wappes) (JEWC: ADP 058561, male paratype), 27 km W Brownsville, Hwy 281, Sabal Palm Grove Sanctuary, 8 m, 25.8419 ° N, 97.4247 ° W, 9 July 1986 (GH Nelson) (JEWC: ADP 116098, male paratype), Hidalgo County, Anzalduas County Park, 27 October 1980 (N Downie, JE Wappes) (NMNH: ADP 058533, female paratype), 10 October 1981 (R Turnbow) (JEWC: ADP 116086, male paratype), Cameron County vicinity of Sabal Palm Grove, 21 October 1989 (EG Riley) (EGRC: ADP 109062, male), Cameron County, Sabal Palm Grove Sanctuary, 16 - 17 October 1993 (EG Riley) (EGRC: ADP 109072, female paratype), Cameron County, Sabal Palm Grove Sanctuary, 21 - 22 April 1990 UV (DJ Heffern) (TAMU: 109064, female paratype), Cameron County, SE Brownsville, Las Palomas Road, 26 October 1991 (DJ Heffern) (TAMU: 109070, female paratype), Cameron County, 16.1 km W Boca Chica State Park, 29 May 1979 (JE Wappes) (JEWC: ADP 109108, male paratype); Starr County, Salineno Park, 29 May 1979 (JE Wappes) (JEWC: ADP 062420, male paratype); Hidalgo County, 17 km SE McAllen, 27 m, 26.1674 ° N, 98.0160 ° W, 28 - 30 July 1976 (no collector label) (UTIC: ADP 148392, female paratype). Geographic Distribution. This species is currently known from the type locality area in Cameron County, and the nearby Texas counties of Hidalgo and Starr (Fig. 4). Notes. This species, one of many in the oblongopunctata pecies-s group, represents the northern limit of the entire Agra distribution pattern. At the southern end, a few species occur in northernmost Argentina. The greater species richness of the genus straddles the equator, decreasing with distance from the equator.	en	Erwin, Terry L. (2017): Agra Fabricius (Coleoptera: Carabidae: Harpalinae: Lebiini: Agrina), Arboreal Beetles of Neotropical Forests: The Rare, Non-Neotropical Texas Species at the Generic Northern Limit, with Notes on Their Way of Life. The Coleopterists Bulletin 71 (4): 639-651, DOI: 10.1649/0010-065X-71.4.639, URL: http://dx.doi.org/10.1649/0010-065x-71.4.639
2F1787E8FFD2827CFCE4FE868FE9F54F.taxon	description	Members of this rather uniform group are of small to medium size and blackish; the nominate species has a subtle bluish tint to the elytra. A diagnostic combination of the adults of the truquii speciesgroup includes the following: labrum slightly depressed medially with rounded anterior lateral margin and slightly emarginate apex; antennae of males with antennomeres more or less subequal in length (except short pedicel); that of females with antennomere 8 short, slightly less than half the length of 7, antennomeres 9 – 11 half the length of 3 - 7. Antenna of normal length, reaching level of middle coxa, not markedly short and robust nor long and markedly thin. Head behind eyes moderately elongate, broadly rounded posteriorly in female, slightly tapered to neck in male; elytral interneurs each a series of large, coarse, variously separated punctures; elytral apex nearly perfectly truncated, laterally with a small, broadly acute tooth, sutural corner rounded; legs and tarsi normal, not markedly modified; male metasternum moderately setigerous, abdominal sterna III – V bilaterally setigerous, setae long and numerous, simply sparsely setiferous on female; male phallus (Fig. 3 B) broadly arrowshaped, tip rounded.	en	Erwin, Terry L. (2017): Agra Fabricius (Coleoptera: Carabidae: Harpalinae: Lebiini: Agrina), Arboreal Beetles of Neotropical Forests: The Rare, Non-Neotropical Texas Species at the Generic Northern Limit, with Notes on Their Way of Life. The Coleopterists Bulletin 71 (4): 639-651, DOI: 10.1649/0010-065X-71.4.639, URL: http://dx.doi.org/10.1649/0010-065x-71.4.639
2F1787E8FFD2827CFCE4FE868FE9F54F.taxon	description	Three species are currently recognized in the species-group:	en	Erwin, Terry L. (2017): Agra Fabricius (Coleoptera: Carabidae: Harpalinae: Lebiini: Agrina), Arboreal Beetles of Neotropical Forests: The Rare, Non-Neotropical Texas Species at the Generic Northern Limit, with Notes on Their Way of Life. The Coleopterists Bulletin 71 (4): 639-651, DOI: 10.1649/0010-065X-71.4.639, URL: http://dx.doi.org/10.1649/0010-065x-71.4.639
2F1787E8FFD2827CFCE4FE868FE9F54F.taxon	description	Agra wickhami Erwin, new species Wickham’ s elegant canopy beetle (Figs. 2, 3, 5)	en	Erwin, Terry L. (2017): Agra Fabricius (Coleoptera: Carabidae: Harpalinae: Lebiini: Agrina), Arboreal Beetles of Neotropical Forests: The Rare, Non-Neotropical Texas Species at the Generic Northern Limit, with Notes on Their Way of Life. The Coleopterists Bulletin 71 (4): 639-651, DOI: 10.1649/0010-065X-71.4.639, URL: http://dx.doi.org/10.1649/0010-065x-71.4.639
2F1787E8FFD2827CFCE4FE868FE9F54F.taxon	materials_examined	Holotype: USA: TEXAS: Cameron County, Brownsville, 9 m, 25.86 ° N, 97.43 ° W, no date (HF Wickham) (CAS: ADP 091173, female). Derivation of Specific Epithet. The epithet “ wickhami ” is an eponym based on the family name of H. F. Wickham, Coleopterist and Paleoentomologist of the late 19 th and early 20 th centuries, who collected the adult holotype of this species somewhere in or near Brownsville, Texas. Proposed English Vernacular Name. Wickham’ s elegant canopy beetle.	en	Erwin, Terry L. (2017): Agra Fabricius (Coleoptera: Carabidae: Harpalinae: Lebiini: Agrina), Arboreal Beetles of Neotropical Forests: The Rare, Non-Neotropical Texas Species at the Generic Northern Limit, with Notes on Their Way of Life. The Coleopterists Bulletin 71 (4): 639-651, DOI: 10.1649/0010-065X-71.4.639, URL: http://dx.doi.org/10.1649/0010-065x-71.4.639
2F1787E8FFD2827CFCE4FE868FE9F54F.taxon	diagnosis	Diagnosis. Agra wickhami has all the attributes of the genus and species-group as described above and is large for the truquii species-group. Adults with black integument; appendages darkly infuscated; antennomeres 6 – 11 bicolored. Frons and occiput moderately domed, smooth, mostly glabrous, with less than 10 scattered setae in addition to fixed supraorbital setae. Pronotum with linear rows of large, coarse punctures from apex to base. Elytral interneurs each with large coarse puncture; apex truncate with small, obtuse dent apicolaterally.	en	Erwin, Terry L. (2017): Agra Fabricius (Coleoptera: Carabidae: Harpalinae: Lebiini: Agrina), Arboreal Beetles of Neotropical Forests: The Rare, Non-Neotropical Texas Species at the Generic Northern Limit, with Notes on Their Way of Life. The Coleopterists Bulletin 71 (4): 639-651, DOI: 10.1649/0010-065X-71.4.639, URL: http://dx.doi.org/10.1649/0010-065x-71.4.639
2F1787E8FFD2827CFCE4FE868FE9F54F.taxon	description	Description. Size: Large for species group, ABL = 15.5 – 16.3 mm, SBL = 14.92 – 17.47 mm, TW = 4.34 – 4.45 mm (Table 2). Color: As described above and antennomers 1 – 5 infuscated, 6 – 11 bicolored with lateral black stripes; mouthparts infuscated, especially maxillary palpomeres, legs and tarsi dark brownish. Luster: Head, pronotum, and legs shiny, elytra matte black. Head: As described above. Prothorax: Short, about the length of head, moderately constricted near base, devoid of hind angles, narrowed anteriorly to about width of neck; surface of disc as described above (Fig. 2). Pterothorax: Elytron moderately convex, narrow in anterior third, slightly flared from middle to apical third and rounded to lateral hind angle; intervals moderately convex, quite regular in width. Legs: Simple in females. Abdomen: As described above. Male genitalia: Unknown. Female ovipositor: Female internal parts not investigated. Female stylomere 2 as in Fig. 3. Dispersal Potential. These beetles are macropterous and probably capable of flight; they are swift and agile runners. Way of Life. See A. rileyi above for general information. Specimens of A. wickhami were taken at lowland elevations along the Rio Grande Valley in southeastern Texas, where sabal palms and sugar hackberry grow. Adults are active in the late rainy season in the Yucatán of Mexico. See Wickham (1897) for descriptions of the vegetation at Brownsville at the time of his collecting the holotype. Other Specimens Examined. Mexico: YUCATÁN, Arco, 16 m, 21.0918 ° N, 88.6654 ° W, 23 September 1965 (AMNH: ADP 056075, female paratype). Geographic Distribution. This species is currently known from the type locality and the Yucatán peninsula of Mexico (Fig. 5).	en	Erwin, Terry L. (2017): Agra Fabricius (Coleoptera: Carabidae: Harpalinae: Lebiini: Agrina), Arboreal Beetles of Neotropical Forests: The Rare, Non-Neotropical Texas Species at the Generic Northern Limit, with Notes on Their Way of Life. The Coleopterists Bulletin 71 (4): 639-651, DOI: 10.1649/0010-065X-71.4.639, URL: http://dx.doi.org/10.1649/0010-065x-71.4.639
