taxonID	type	description	language	source
03F3EC13FFEAFF89FF0F0BDEBCA7FF37.taxon	description	(Figs 1 − 59)	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFEAFF89FF0F0BDEBCA7FF37.taxon	type_taxon	Type species: Haemolaelaps ulysses Domrow, 1961: 63.	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFEAFF89FF0F0BDEBCA7FF37.taxon	diagnosis	Diagnosis. Female. Medium-sized to large mesostigmatan mites. Dorsum. Mid-podonotal region of dorsal shield smooth, or almost so. Lateral margins and opisthonotal region of dorsal shield usually with reticulations. Many species with 39 paired dorsal setae as defined for holotrichous Laelapidae by Evans & Till (1965). Some species exhibit loss of podonotal z 3 and px 2 – 3, and U. sisyphus exhibits further hypotrichy as noted in the key. Dorsal setae smooth, except U. laertes has Z 5 with a few incipient barbs. Dorsal setae sometimes reduced to microsetae (U. calypso and U. telemachus). Anterior hexagon of podonotum lacks accessory setae. Accessory Jx setae absent in U. calypso, U. sisyphus and U. telemachus. Other species have 2 – 8 accessory Jx setae, which are borne in a posteriorly-positioned field in vicinity of J 4, some may appear paired, none anteriad level of mid J 3 – J 4. Gnathosoma. Lateral arms of internal malae have fimbriae bearing highly charactertistic expanded tips (except in U. laertes, U. telemachus and U. theoclymenus). These modified fimbriae are distally inflated, up to 2 µm wide, and may be densely clustered. Pilus dentilis short and uninflated. Cheliceral digits show great variation in size and dentition (Table 3). Fixed digit dentition varying in number and development from edentate to four strong teeth (count excludes apically-divided tip). Subcheliceral shelf usually with trident apex, but with three exceptions: U. calypso with 4 − 6 tines, U. species B with four tines, U. laertes with two tines and U. euryclea has an undivided pointed tip. Inner palp trochanteral setae, either setiform or blade-like (2 – 13 wide). Deutosternal groove with six rows of denticles; maximum number of denticles per row varies greatly, 2 – 19. Venter. Tritosternal base smooth. Genito-ventral shield bearing st 5, sometimes Zv 1 and rarely also Jv 1. Genitoventral shield extensive reaching close to, or touching, anal shield. Anterior portion of shield narrowly bordered by longitudinal striae extending posteriorly only to level of Jv 1 or less. Shield bearing 6 – 8 serially-regular, broadly transverse striae (except with obscure striae in U. ulixes, apparently smooth in unnamed species A and with reticulate striae in U. sisyphus). Anal shield broad being at least as broad as long or very broad (e. g. ca 1. 6 times broader in U. penelope). Opisthogaster mildly to strongly hypertrichous, or hypotrichous in U. sisyphus. Legs. Ulyxes spp are readily distinguished by certain dorsal leg setae being apically bifid on trochantera and femora (Figs 9, 18, 41). Apically bifid setae may be short or long, stout or slender, and have parallel, non-tapering sides. They are flattened (strap-like). They are never borne ventrally (with one possible exception, see Fig. 42). These differ in both number and position between species, providing setal signatures that are largely characteristic of, and consistent within, a species. There are 12 different setal positions known to bear apically bifid setae of which 5 – 10 are bifid on any one species (Table 4). Barbed setae absent. Tarsus I with apical stalk short, at most 16 long, or absent. Leg chaetotaxy holotrichous as defined by Evans (1963) for Laelapidae except genu IV variable with 8 – 10 setae, and U. sisyphus hypotrichous on various other segments (Domrow, 1981). Male. The six known males are the previously described U. telemachus, U. ulixes and U. sisyphus (Domrow, 1964, 1972 a, 1981); and described here are males of U. euryclea, U. laertes (Domrow) and U. penelope (Domrow). Cheliceral digits variable. Fixed digit varies from not reduced at all in U. laertes (Fig. 57) or U. euryclea (Fig. 56) to strongly reduced in U. telemachus (Fig. 52). All species have fixed digits with apical hooks except U. telemachus. In all species the pilus dentilis emerges from the distal half, including at the apex in U. telemachus. Spermatodactyl varies from being mostly free of movable digit from ca. 10 % of length fused, to 80 % of length fused. The spermatodactyl is moderately long at most and upturns abruptly with a distinct bend. Spermatodactyl tip usually blunt and rounded but sometimes with a small triangular projection providing a sharp-tip (U. euryclea, Fig. 24). Genital opening wider than deep, set anteriorly with posterior edge at least level with st 1 (Figs 26, 40, 46). Posterior holoventral shield with circumanal setae plus five setal pairs, or only four pairs with Jv 3 off shield (U. euryclea, U. ulixes) or just three pairs with Jv 3 and Zv 2 off shield (U. penelope, U. telemachus). Tarsus II av 2 distinctly modified in U. penelope (Fig. 42).	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFEAFF89FF0F0BDEBCA7FF37.taxon	etymology	Etymology. Ulyxes is a variant spelling of Ulysses.	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFEAFF89FF0F0BDEBCA7FF37.taxon	discussion	Remarks. Ulyxes spp comprises thirteen species, eight previously described and five undescribed: U. telemachus on two small congeneric dasyurid marsupials (Domrow, 1964); U. laertes from an Australian endemic murid rodent (Domrow, 1972 a); U. euryclea from the cavity nests of cockatoos and Leadbeater’s possum described herein; U. theoclymenus from a Papuan lory here described, and the remaining ten species on an diverse array of possums in the superfamilies Petauroidea (Acrobatidae, Petauridae, Pseudocheiriidae) and Phalangeroidea (Domrow, 1961, 1964, 1981). Host and area relationships of Ulyxes spp, described and undescribed, are given in Table 2. Haemolaelaps was resurrected from synonomy and rediagnosed in a separate paper (Shaw 2014), where it is redefined as equivalent to the Haemolaelaps marsupialis group (Domrow, 1988) and herein referred to as Haemolaelaps sensu stricto. Here I briefly contrast the diagnoses of Haemolaelaps ss and its former congener Ulyxes. The following five characters apply to Haemolaelaps sensu stricto, but not Ulyxes: there are never unpaired dorsal setae, the pilus dentilis is long, being at least as long as the movable digit (except shorter in H. hattenae), the fimbriae on the lateral arms of internal malae never have inflated tips, the second poststigmatal pore is enlarged, leg setae are always barbed. Also the pilus dentilis of the male always emerges from the apex of the fixed digit in Haemolaelaps ss. Haemolaelaps is one of seven described Australian genera that are related to Ulyxes, and I regard Ulyxes as more closely related to some of these (e. g. Nidilaelaps) than it is to Haemolaelaps ss.	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFEDFF89FF0F0E42BC83FD40.taxon	discussion	Remarks. Male unknown. This species is only reported from Leadbeater’s possum Gymnobelides leadbeateri from a highland site in Victoria (Tanjil Bren). It could easily be confused with the closely-related U. euryclea which also occurs in Leadbeater’s possum nests. See Remarks for U. euryclea. It has two teeth on the fixed digit which is slightly stronger dentition than some of its parasitic congeners (Table 3).	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFEDFF8BFF0F0975BD4EFAF7.taxon	description	(Figs 1 – 5) Specimens examined. 1 holotype female, 2 paratype females, Collin’s Sawmill, Mt Otto, Papua New Guinea, 7 Aug 1959, H. M. Van Deusen coll., ex “ Pseuocheirus cupreus ” = Pseudochirops cupreus Coppery Ringtail, 2215 metres, A. E. (Archbold Expedition) No 16229. Holotype and paratypes in BBM. Description of female (n = 3). Dorsum. Dorsal shield 850 (840 – 850) x 610 (610 – 660), with 39 paired setae and 2 – 3 unpaired Jx setae (between J 4 to mid J 3 – 4) (Fig. 1). Podonotal j 1 46 (41 – 48), j 2 55 (55 – 57), z 1 28 (28 – 33), z 3 45 (41 – 45). Discal setae shorter, opisthonotal J 2 27 (27 – 28). Z 5 49 (49 – 53). Marginal setae r 2 46 (46 – 50), r 3 48 (48 – 60), r 4 49 (49 – 52), r 5 57 (57 – 61), S 1 55 (54 – 55), S 2 53 (53 – 54), S 4 51 (51 – 57), S 5 49 (49 – 55). Fine reticulations over opisthonotal and lateral portions of shield. Discal portion of podonotum glabrous. Z 5 smooth. Distinctive gland pore posterolaterad px 2, only weakly encircled by striae. Gnathosoma. Epistome a somewhat triangular lobe with weakly – toothed edge (Fig. 4). Corniculi protrude 24. Hypostomal setae: h 1 43 (42 – 43), h 2 26 (23 – 26), h 3 60 (56 – 60), capitular seta 30 (30 – 32). Fixed digit edentate except for divided tip. Movable digit 52 (50 – 52) long with two weak teeth (Fig. 3). Cheliceral seta dorsal, 15 long. Second cheliceral segment 153 (148 – 153), first cheliceral segment 65 (62 – 65). Six denticulate deutosternal rows, Q 2 – 5 with 4 – 8 denticles and Q 6 – 7 with 12 or more very fine bead-like denticles (Fig. 2). Deutosternal groove flanked by 2 – 3 lateral lines. Inner palp trochanteral setae setiform. Palp genu al 1 short with blunt tip. Palp genu al 2 with subparallel sides and distally compressed. Lateral arms of malae short, fimbriae tips distinctly expanded (Fig. 2). Labrum projects to level of mid palp genu. Venter. (Fig. 5) Smooth tritosternal base, 41 (38 – 41) between origin and suture. Lacinae fuse 0 – 4 distad suture. Sternal shield wider 154 (137 – 154) than long 94 (90 – 94). Anterior edge of sternal shield is subcuticular, obscured, giving the appearance of st 1 being off shield. Moderately long cornua present. Sternal shield with usual two lyrifissures, with st 1 situated immediately external to level of first lyrifissure; third lyrifissure is free. St 1 52 (52 – 58), st 2 59 (59 – 63), st 3 (55 – 58). Sternal shield extending only to level of anterior edge coxa III, posterior edge a strong concave arch. Genito-ventral shield with 6 strongly transverse and serially-regular striae, 5 – 6 rows at or below level of Zv 1. Zv 1 missing from RHS holotype and LHS of a paratype. Genitoventral shield 221 (221 – 241) long x 150 (142 – 150) wide at st 5. Maximum width 253 (246 – 253) at Jv 1. Longitudinal striae extend only to level of Zv 1. Genito-ventral shield bearing only st 5, associated lyrifissure (iv 5) is off shield in soft cuticle. Peritreme extends past middle of coxa I. Principal (= external) metapodal platelet oval, 50 (39 – 50) x 14 (14 – 21). Inner metapodal a small broad oval 20 (13 – 20) x 12 (12 – 14). Paragenital platelet present. 12 – 14 pairs of setae on unarmed ventral opisthogaster (count excludes Jv 1 – 3 and Zv 1 – 2). Post-stigmatal plate abbreviated, with 3 porelike structures, very narrowly rebordered externally. Median and posterior pore-like structures are connected by short gutter. Genito-ventral shield extends to within 9 (5 – 9) of anal plate. Anal plate 108 (108 – 109) long x 126 (122 – 126) wide between cribral pores. Maximum width 142 (140 – 144). Cribrum of two apparent rows, in holotype a short third row appressed to second. Para-anal setae 37 (37 – 39), postanal 55 (55 – 65). Legs. Leg setation holotrichous as defined by Evans (1963), genu IV with a single pl seta, proximally positioned. Apically bifid setae on femur I ad 1, pd 2; femur II ad 1, pd 2; femur III ad 1, pd 1 and femur IV ad 1, ad 2 and trochanter IV pl. Leg segment lengths as in Table 4.	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFEDFF8BFF0F0975BD4EFAF7.taxon	etymology	Etymology. All members of this genus are named after characters in Homer’s Odyssey. Autolycus was a thief and a relative of Ulysses.	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFEDFF8BFF0F0975BD4EFAF7.taxon	discussion	Remarks. This is the first pseudocheriid-associated Ulyxes species to have podonotal z 3 present. This disrupts the formerly neat morphological divide between petaurid- and pseudocheriid-associated species, and confounds the implication of cophylogeny made by Domrow (1981). Based on its small edentate chelicerae this species is assumed to be parasitic (Table 3).	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFEFFF8BFF0F0AFABE06F8D0.taxon	description	(Figs 6 – 9, 58)	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFEFFF8BFF0F0AFABE06F8D0.taxon	discussion	Remarks. Males are unknown. Nesting material from 29 nestboxes used by Petaurus breviceps or P. norfolcensis in Victoria and SE Queensland were extracted and four nests were found to contain female U. calypso but no males were found. Based mainly on its small, weakly-dentate chelicerae this species can be assumed to be parasitic (Table 3). Its strong, long sternal setae and thickened coxal setae are likely to be adaptations for resisting grooming while in its hosts’ fur.	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFE0FF82FF0F09D2BD48F881.taxon	description	(Figs 10 – 31, 56) Specimens examined. 1 holotype female, 1 paratype male, 3 paratype females, 9 females, Candlebark Park, Templestowe, Victoria, 37 ° 31 ’ 56 ” S, 145 ° 06 ’ 39 ” E, 15 Nov 1998, I. Temby coll., ex nestbox recently occupied by Cacatua longirostris Long-billed Corella, collected four days after last chick fledged, nest 87; 4 females, Stokes Bay, Kangaroo Is, South Australia, 35 ° 37 ’ S 137 ° 12 ’ E, 1 Jun 1999, L. Pedler coll., ex tree hollow occupied by Calyptorhynchus lathamihalmaturinus Glossy Black Cockatoo, “ Tragic ”, ca. 10 week old nestling present, nest 245; 1 female, Cygnet River, Kangaroo Is, South Australia, 35 ° 42 ’ S 137 ° 12 ’ E, 20 Mar 1999, L. Pedler coll., ex tree hollow occupied by incubating female Calyptorhynchus lathami halmaturinus Glossy Black Cockatoo, “ Sunglasses ”, nest 239; 2 females, 1 male, Yellingbo, Victoria, 37 º 49 ’ S 145 º 30 ’ E, 13 May 2002, D. Harley coll., nestbox occupied by Gymnobelideus leadbeateri Leadbeater’s Possum, L 9, nest 486. Holotype and paratypes in QM. Description of females (n = 8). Dorsum. 775 (710 – 785) long x 545 (500 – 550) wide, margins and posterior of shield faintly reticulate, becoming smooth anteromedially. Standard 39 setae plus 5 – 7 medial accessory Jx setae, posterior level J 3 (Fig. 10). At least 2 of these accessory setae are placed almost symmetrically. Mediodorsal setae of relatively uniform length, j 1 43 – 54, j 2 43 – 60, z 1 25 – 32, z 3 46 – 68, J 2 40 – 50. J 5 30 – 35, Z 5 35 – 47. Marginal setae slightly longer r 2 48 – 60, r 3 52 – 56, r 4 54 – 62, r 5 58 – 59, S 1 55 – 63, S 2 57 – 68, S 3 57 – 67, S 4 55 – 65, S 5 47 – 58. Dorsal shield covers entire dorsum. Z 5 smooth. Distinct gland pore posterolaterad px 2, within subcircular lacuna 11 – 13 in diameter. Gnathosoma. Epistome with essentially smooth edge, some specimens with sparse denticles (Fig. 11). Hypostomal setae h 1 41 (33 – 47), h 2 28 (19 – 29), h 3 70 (59 – 71), capitular seta 45 (46 – 49). Deutosternal groove parallel-sided, bearing six deutosternal rows with 8 – 13 denticles per row. Fringed internal malae aligned medially, with short lateral lobes. Tips of fimbriae have distinct swellings (Fig. 14). Corniculi 22 (20 – 22) long, and somewhat broad. Palp trochanter-tibia holotrichous (2: 5: 6: 14). Palp genu al 1 shorter than al 2, both distally compressed. Four teeth on fixed digit, with one small tooth distal of uninflated pilus dentilis and three welldeveloped teeth proximal. Tip of fixed digit divided (Figs 12, 21). Cheliceral seta dorsal, 8 long. Movable digit 63 (59 – 63) with two well-developed teeth borne on antiaxial edge (Fig. 16). Second cheliceral segment 151 – 174, first segment 65 – 71. Labrum projects strongly to level of distal edge of palp femur. Venter (Fig. 15). Smooth tritosternal base, 35 (35 – 40) between origin and suture, 17 – 18 wide at origin, 10 – 11 at suture. Lacinae are fused 18 (9 – 18) above level of suture and free for 93 (85 – 105) (Fig. 13). Sternal shield wider 132 (130 – 136) than long 116 (114 – 125). Anterior edge of sternal shield is distinctly recessed or stepped-down immediately internal of st 1. Long thin cornua abruptly meet, but do not fuse, with exopodal I. Usual two lyrifissures on sternal shield, with st 1 external of first lyrifissure, third lyrifissure is free. St 1 62 (60 – 63), St 2 63 (63 – 72), St 3 64 (60 – 64). Sternal shield extending to level mid-coxa III, concave posteriorly. Genito-ventral shield (Fig. 15, 19) with 7 – 8 serially-regular, broadly transverse striae, 204 (190 – 225) long x 150 (132 – 150) wide at st 5. Maximum width 214 (193 – 215) at Jv 1. Lateral edges of genito-ventral shield subparallel. Longitudinal striae extend to just past Zv 1. Genito-ventral shield bearing only st 5, associated lyrifissure (iv 5) is off shield in soft cuticle. In ovigerous females, the anterior pole of the large egg (405 – 439 x 305 – 365), has a distinctive sclerotised plate, maximum width ca. 56 (Fig. 20; see Remarks). Peritreme extends past middle of coxa 1. Principal or external metapodal platelet oval, 43 (39 – 48) x 15 (14 – 17). Inner metapodal small oval, 10 – 22 x 6 – 10. 22 pairs of setae on unarmed opisthogaster excluding Jv 1 – 3 and Zv 1 – 2. Post-stigmatal plate narrow but retains narrow external rebordering, and usual three pore-like structures, median and posterior pores are connected by short gutter (Fig. 17). Mild exopodal IV lobe, 18 – 20 wide. Genito-ventral shield reaches to within 0 – 8 µm of anal plate. Anal plate 97 (95 – 118) long x 115 (98 – 115) wide between cribral pores. Maximum width 120 (113 – 123), cribrum of only two rows in most specimens, portion of third cribral row sometimes present. Para-anal setae 31 (28 – 36), postanal 46 (38 – 46). Legs. Leg setation holotrichous as defined by Evans (1963), genu IV with a single pl seta, proximally positioned. Apically bifid setae distributed between femora I – IV and trochanter IV as femur I ad 1, pd 2, ad 3, femur II ad 1, pd 2, femur III ad 1, trochanter IV pl and femur IV ad 1, ad 2 (Fig. 18). Ventral setae of tibiae and tarsi II – IV and genu III – IV are stouter than dorsal leg setae. Pretarsal opercula with ca. 7 – 9 tines. Leg segment lengths as in Table 4. Description of males (n = 2). As for female except as described below. Dorsum. Dorsal shield 640 – 650 x 410 – 425. Gnathosoma. Fixed digit with rounded tooth level with pilus dentilis; second small tooth distal. Movable digit, 45 – 49, not reduced, with usual single tooth and notched apex. Spermatodactyl free in distal half, broad bore, 4 wide at apex with a sharp triangular projection (Fig. 24). Lateral lobes of internal malae have fimbrae with tips expanded. Second cheliceral segment 137 – 138. Slender corniculi 23 – 24 long. Hypostomal setae h 1 35 – 41, h 2 17 – 26, h 3 35 – 37, capitular seta 49. Venter. Tritosternal base 25 – 30 long to suture, 10 from suture to fork of lacinae. Lacinae 83 – 88 long. Male genital opening oval 27 – 30 wide x 17 deep. Opening set anteriorly with posterior edge of opening level with st 1 (Fig. 26). Holoventral shield (Figs 22, 25) bearing st 1 – 5, circumanal setae and 4 other pairs of setae with Jv 3 off shield, 112 – 115 wide at st 2, st 1 39 – 45, st 2 48 – 46, st 3 47 – 51. Metapodal platelets subsumed by holoventral plate. Holoventral shield not broadened at level of anal opening, 88 – 97 between cribral pores. Para-anal setae 29, postanal 27 – 35. Legs. Femur II av 1 is strong spike, 33 long, shifted proximally to central point on segment making pv 1 - av 1 - av 2 essentially straight (Figs 23, 27). Leg segment measures in Table 4. No modified setae on tarsus II.	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFE0FF82FF0F09D2BD48F881.taxon	etymology	Etymology. From Euryclea, Ulysses’ nurse, who lived in the same house as Anticlea, Ulysses’ mother.	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFE0FF82FF0F09D2BD48F881.taxon	discussion	Remarks. Ulyxes euryclea was compared to five paratypes of the closely-related U. anticlea Domrow (QM S 58721 – 4, 58739), which also occurs with Leadbeater’s possums. Both species share similar body size and an identical arrangement of apically bifid setae. However Ulyxes euryclea females differ from those of U. anticlea in always having only st 5 on the genito-ventral shield, and this shield being relatively narrow with nearly parallel lateral edges in contrast to the broader, distinctly rounded shield of U. anticlea bearing st 5, often Zv 1 and sometimes also Jv 1. U. euryclea has a distinctive gland pore posterolaterad px 2 (as is in many Laelapidae), whereas this pore is anterolaterad px 2 in U. anticlea. There are other consistent differences between the two species (U. euryclea vs U. anticlea): the number of fixed digit teeth (4 vs 2), number of denticles in deutosternal rows (8 − 13 vs 3 − 8), basal width of tritosternum (17 – 18 wide at origin, 10 – 11 at suture vs 27 − 30 wide at origin, 12 − 17 at suture), maximum width of the genito-ventral shield (193 – 215 vs 249 – 259), and distance between genitoventral and anal shields (0 – 8 vs 14 − 17). Ulyxes euryclea has been recovered from natural and artificial nest cavities occupied by Glossy Black Cockatoos, Long-billed Corellas in Victoria and South Australia and also from nestboxes used solely by Leadbeater’s possums at Yellingbo, a lowland site. The other member of this genus recorded with Leadbeater’s possum is the closely-related U. anticlea (Domrow, 1972 b). The feeding biology of U. euryclea is unknown, but it is probably a nest-based predator of small invertebrates. Although U. euryclea has cheliceral digits the same length as the blood-feeding U. penelope Domrow, its dentition is stronger, with four teeth on the fixed digit (Table 3). Also it is probably relevant that males retain chelate-dentate dentition (cf U. laertes) and, in contrast to many putatively parasitic Ulyxes spp, it is found associated with a broad range of vertebrate species as listed above. The complex sclerotised plate-like structure seen on the anterior pole of many of the eggs has not been detected in other Ulyxes spp. This structure appears to bear the pair of ramus sacculi (rs), opening into a pair of circular areas which may be the bilobed sacculus foemineus (sf, Fig. 20). U. euryclea is superficially similar in its gracile setae, form of genito-ventral shield, and in general appearance to Nidilaelaps holdsworthi Shaw, which is only known from parrot nests in southern Australia. This species, along with Nidilaelaps annectans (Womersley) and U. euryclea, co-occur in Glossy Black cockatoo nests on Kangaroo Island, South Australia (Shaw, 2012). However U. euryclea is instantly distinguishable from these species by those characters which diagnose Ulyxes spp, e. g. presence of apically bifid leg setae. A female and two protonymphs in very poor, decayed condition were kerofloated from a nestbox housing Trichoglossus moluccanus Rainbow Lorikeet in Brisbane (nest 89). The female has similar dentition and anterior edge of sternal shield to U. euryclea, but it lacks a bifid seta on trochanter IV. More specimens are required before this population can be identified.	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFE7FF9EFF0F0A12BBF9FDDF.taxon	description	(Figs 28 − 37, 57)	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFE7FF9EFF0F0A12BBF9FDDF.taxon	description	Females were described by Domrow (1972 a). Description of males (n = 3). Dorsum. (Fig. 34) Very large mites. Dorsal shield 1150 – 1190 long x 780 – 800 wide. 39 standard setae plus 3 – 4 unpaired Jx setae at level of J 4. Podonotal j 1 85 – 92, z 1 ca. 42, other medial and discal setae short, e. g. j 5 ca. 42, j 6 ca. 47. J 2 50, J 5 50. Marginal setae heteromorphic in size, long and thick, increasing in length and becoming sabre-shaped posteriorly; s 1 52, s 2 60, r 2 68, r 3 90, r 4 85, r 5 87, S 1 90 – 104, S 2 95 – 98, S 3 92 – 108, S 4 90 – 120, S 5 110 – 118. Dorsal shield covered in fine reticulations, fainter in mid-podonotal region. Gnathosoma. (Fig. 32) Palp genu al 1 broad, 8 µm maximum width, hyaline, lateral edges subparallel. Palp genu al 2 narrow with long spatulate tip. Row Q 1 bare, followed by usual six denticulate deutosternal rows (Q 2 – Q 7) and bare Q 8. Movable digit 188 – 205 long bearing single tooth. Spermatodactyl relatively small, shorter than movable digit with outline of movable digit completely encompassing that of spermatodactyl (Figs 30, 35). Most of spermatodactyl, 80 % of length, fused with movable digit. Fixed digit without divided tip, with three strong teeth with pilus dentilis adjacent to second tooth. Cheliceral seta dorsal. Cheliceral segment II 415 – 430, segment I ca. 152. Epistome denticulate, narrows abruptly, leaving a narrow-based gradually-tapering medial point (Figs 29, 36). Venter (Fig. 28). Tritosternal base 50 – 77 long to suture. Lacinae separate ca. 12 above suture and are free for ca. 212. Genital opening broad, very narrow oval, 65 wide x 22 deep. Opening set anteriorly, posterior edge level with st 1. St 1 65 – 80, st 2 52 – 60, st 3 52 – 53. Holoventral plate bears five pairs setae in addition to st 1 – 5 and circumanal setae. Para-anal setae 48 – 50; postanal seta 73 – 80. Cribral pores separated by 197 – 230. Cribrum a dense beard of fine spicules, not in discernible rows. Legs. Ten setae on trochanter I and femora I – IV modified as apically bifid (Fig. 37); trochanter 1 d, femur I pd 2, ad 3, femur II ad 1, ad 3, pd 2, femur III ad 1, pd 1 and femur IV ad 1, ad 2. Femur II ventral setae unmodified. No modified setae on tarsus II. Pretarsal opercula with 10 – 12 tines. No apical stalk on leg I. Leg segment lengths as in Table 4.	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFE7FF9EFF0F0A12BBF9FDDF.taxon	discussion	Remarks. Specimens of U. laertes were kept in culture for three months. Breeding occurred but no second generation adults were produced. Adult stages readily predated on adult Stratiolaelaps sp. Other prey that were accepted were a blood-fed endemic macronyssid protonymph and an anopluran louse. A female was also observed to grasp nematodes. The predatory habits of U. laertes match its unusually strong and large chelicerae. However, all postlarval stages also readily orientated towards sources of free blood and fed avidly from these. Judging by its chelicerae it seems unlikely that it would feed on unbroken skin of its host. Its attraction to, and ingestion of blood, could be an adaptation for utilising other sources of blood available within nests such as engorged mites, fed flea larvae, and the faeces of adult fleas. Engorged lice might also be available in the host’s fur. In culture U. laertes showed a strong cryptic tendency, invariably hiding under squares of Parafilm provided for them. This behaviour could be a thigmotactic response. The mites did not seem to tolerate conspecifics in close proximity and apparent intraspecific aggression was observed on many occasions when a U. laertes would venture underneath a square of Parafilm and move too close to an already resident mite. The resident would make a short advance towards the intruder, with the intruder then changing direction and allowing the resident to maintain its position underneath its Parafilm shelter. Ulyxes laertes has an enlarged capitulum (Figs 32, 33). The palp femora are internally concave (Domrow, 1972 a), and the epistome narrows abruptly creating a strong medial point (Figs 29, 36). These features appear to be accommodations for its enormously-enlarged chelicerae. Three cusps are found on the fixed digit of both females and males (Figs 30, 31) and in this respect, U. laertes appears unusually, though superficially, sexually monomorphic. Note that these dentitions between the sexes are not considered homologous as demonstrated by the different relative position of the pilus dentilis. The movable digits of females are three to seven times longer than other members of Ulyxes spp in absolute terms, and even as a ratio of body size they are two to more than four times longer (Table 3). The digits are much larger than in any morphologically similar mites too, even including Laelapsella humi Womersley, that similarly-sized inhabitant of ground nests, whose digits are still only half as long as those of U. laertes. This fascinating species is rare in collections. Prior to this study there were just three females in Queensland collections despite it being larger than most other symbiotic Laelapidae and despite it ocurring on small mammals that are frequently trapped by zoologists. I suggest its rarity in collections is because most collections of commensal arthropods are from hosts, and this mite is probably strongly nidicolous, rarely occupying its host’s fur. Ulyxes laertes was collected from two Crow’s Nest ferns at Tenison Woods Mountain. These ferns were used as nests by small mammals, judging by the co-occurrence of most life stages, including males and nymphs, of Mesolaelaps sp. in one nest, and the macronyssid Trichonyssus praedo (Domrow) in the other.	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFFAFF98FF0F084CBE0DFA22.taxon	description	(Figs 38 – 42)	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFFAFF98FF0F084CBE0DFA22.taxon	description	Females were described by Domrow (1964). Description of male (n = 1). Medium-sized mites. Dorsum. Dorsal shield 476 long x 240 wide. Podonotal j 1 22, z 1 17, other medial and discal setae also short: j 5 15 – 16, j 6 15 – 16. J 2 17 – 18, J 5 23. Marginal setae slightly longer: s 1 22, s 2 21, r 2 20, r 3 27, r 4 22, r 5 27, S 1 27, S 2 25, S 3 31, S 4 32, S 5 33, Z 5 36. Dorsal shield podonotal region without reticulations except narrow margin, with 39 standard setae plus two unpaired Jx setae posteriad level of J 4. Gnathosoma. Corniculi slender. Deutosternal groove with six rows of 1 – 2 denticles each (Fig. 39). Movable digit 32 long bearing single weak tooth and a strong apical hook. Spermatodactyl longer than, and mostly separate from, movable digit. Fixed digit reduced, edentate with weak apical hook (Fig. 38). Cheliceral segment II 82, segment I 28. Epistome not discernible. Hypostomal setae h 1 30, h 2 14, h 3 30, capitular seta 17. Venter (Fig. 40). Tritosternal base short, 12 long to suture. Lacinae separate immediately above suture, 85 long. Genital opening broad, oval, 27 wide x 19 deep, set anteriorly, posterior edge level with st 1. Sternal shield 100 wide at st 2. St 1 22, st 2 21, st 3 21. Holoventral plate bears three pairs setae in addition to st 1 – 5 and circumanal setae, Jv 3 and Zv 2 off shield. Primary metapodal platelet 26 x 8. Para-anal setae 18; postanal seta 25. Cribral pores separated by 72. Cribrum two rows of fine spicules. Legs. A total of seven setae on femora I – IV are modified as apically bifid: femur I ad 1, pd 1; femur II ad 1, pd 2; femur III ad 1; femur IV ad 1, ad 2. No trochanteral setae modified. Femur II ventral setae av 1 not modified, nor shifted laterally (Fig. 41). Tarsus II av 2 modified as a stout spine, with a sharp accessory projection halfway along its length (Fig. 42). Leg I without apical stalk. Tarsus I forked sensory seta present (as in female). Leg segment lengths as in Table 4.	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFFAFF98FF0F084CBE0DFA22.taxon	discussion	Remarks. Some freshly caught female U. penelope contained blood in their caeca (nest 475). A bulk culture of 14 U. penelope from nest 475 (10 nymphs, 3 females, 1 male) was established. All life stages fed avidly on fresh blood supplied in short lengths of heparinsed micropipettes. While the initial population declined after 3.5 months, two mites remained active and healthy for more than five months. However none were observed to feed on, or show interest in, regularly-supplied cultured nematodes. In contrast to some other nidicolous parasites such as macronyssids and Mesolaelaps australiensis, the short-limbed U. penelope showed a tendency to remain on the substrate floor and not climb the walls of their container, hiding quickly under pieces of humus when disturbed. Ulyxes penelope appears to be restricted to the Short-eared possum Trichosurus caninus and the closely-related Trichosurus cunninghami (Lindenmayer et al., 2002). Despite various collection attempts, no U. penelope have been confirmed from Trichosurus vulpecula, a common possum that is sometimes sympatric with T. caninus. Ulyxes penelope was collected from nestboxes used by Trichosurus spp possums at Woodford, South-east Queensland. Both Trichosurus vulpecula and T. caninus use the same nestboxes at this locality so the host data is not known here.	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFFCFF98FF0F094EBE65F832.taxon	description	(Figs 55, 59)	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFFCFF98FF0F094EBE65F832.taxon	discussion	Remarks. The description of this species appears in a journal issue intended for 1979 and stated therein to be published 1980 but the actual publication date is 1981 (Domrow, 1988; Halliday, 1998). Ulyxes sisyphus is recorded from the intranasal passages of southeastern New South Wales populations of Trichosurus vulpecula, the Common Brushtail Possum. In one specimen peritematal and exopodal IV plates are narrowly fused (QMS 58771). Even though it has moderately strong chelicerae (Table 3), these are dissimilar to those of predatory forms, and it is assumed to be a parasite based on its highly specialised habitat and derived morphology.	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFFDFF9AFF0F0FEEBE84FE87.taxon	description	(Figs 43 – 46)	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFFDFF9AFF0F0FEEBE84FE87.taxon	discussion	Remarks. Ulyxes telemachus is found on the small carnivorous marsupials Antechinus stuartii sensu stricto and A. flavipes. These hosts are excellent climbers and they nest in cavities, typically in trees although they will use fallen logs too. U. telemachus appears to be strongly modified for host association having short legs, an almost glabrous dorsum, minute dorsal setae, and short leg setae. For feeding, its tiny cheliceral digits are probably used to take fluid directly from its host.	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFFEFF9AFF0F0EEABAC0F8A8.taxon	description	(Figs 47 – 51) Specimen examined. 1 holotype female, May River, West Sepik province, 3 Jun 1963, P. Temple coll., ex Lorius lory det A. C. Zeigler, BBM-NG 22634. Holotype in BBM. Description of female (n = 1). Dorsum. Dorsal shield 890 x 630, with 39 paired setae and eight unpaired Jx setae at ca. level of J 4, spread longitudinally between J 5 and mid J 3 – 4 (Fig. 47). Podonotal j 1 58, j 2 70, z 1 38, z 3 72. Discal setae shorter e. g. j 6 36. Marginal setae r 2 65, r 3 75, r 4 68, r 5, S 1 80, S 2 77, S 3 76, S 4 82, S 5 79, J 5 51 Z 5 77. Fine reticulations over opisthonotal and lateral portions of shield. Discal portion of podonotum glabrous. Z 5 smooth. Distinctive gland pore posterolaterad px 2, set in subcircular lacuna ca. 20 wide. Gnathosoma. Epistome a shallow, smoothly-rounded lobe (Fig. 49). Corniculi protrude 28. Hypostomal setae: h 1 61, h 2 41, h 3 83, capitular seta 54. Fixed and movable digits are scoop-shaped, with teeth borne on both lateral edges of movable digit (Fig. 48). Movable digit 70 long with two strong teeth borne on antiaxial edge of digit which do not oppose any teeth in fixed digit, although they shear against undulating antiaxial edge of fixed digit. In addition another strong tooth borne on paraxial edge of movable digit shears against three triangular teeth on paraxial edge of fixed digit. Cheliceral seta not discernible. Second cheliceral segment 188, first cheliceral segment 85. Six deutosternal rows of 8 – 14 denticles, flanked by single pair of lateral lines at ca. level of Q 7 (Fig. 50). Palp genu al 1 with rounded, unexpanded tip. Palp genu al 2 strongly spatulate. Lateral arms of malae lengthened with fimbriae extending past tip of corniculi. Fimbriae tips not expanded. Venter. Tritosternum with base 51 long to suture. Lacinae separate 10 above suture. Lacinae free for 118. Sternal shield 155 wide, 120 deep, anterior edge straight, lacking excavations, posterior edge straight. St 1 68, st 2 72, st 3 75. Shield 240 long, extends posteriorly to within 1 µm of anal shield. Genito-ventral shield broad, 62 between st 5, maximum width 285, between level of Zv 1 and Jv 1 (Fig. 51). Longitudinal striae narrowly border genito-ventral shield, falling short of Jv 1. Serially-regular, broadly transverse striae in 7 – 8 rows, 5 – 6 rows at or below level of Zv 1. Post-stigmatal plate with median and posterior pore-like structures connected by narrow gutter. Exopodal IV a slender rim, 10 wide, without ridges. Primary metapodal platelet oval 62 x 13, secondary (inner) metapodal platelet 28 x 8. Paragenital platelet present. Opisthogaster hypertrichous with ca. 24 pairs of setae (excluding Zv 1, Jv 1 – 2). Anal shield 118 long by 137 wide at cribral pores. Maximum width 148. Para-anal setae 46. Cribrum in 3 closely-appressed rows. Legs. Leg setation holotrichous as defined by Evans (1963), genu IV with a single pl seta, proximally positioned. Eight setae are modified as apically bifid; femur I ad 1, femur I pd 2, femur I ad 3, femur II ad 1, femur II pd 2, femur III ad 1, femur IV ad 1 and femur IV ad 2. Pretarsal opercula concealed in this unique specimen. Ambulacra I borne on apical stalk, 16 long. Leg segment lengths as in Table 4.	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFFEFF9AFF0F0EEABAC0F8A8.taxon	etymology	Etymology. Theoclymenus was a diviner who read the auspices of birds.	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFFEFF9AFF0F0EEABAC0F8A8.taxon	discussion	Remarks. The dentition of the movable digit is highly conservative throughout the Laelapidae so U. theoclymenus is remarkable in having a third tooth that has arisen de novo, creating an additional, paraxial, shearing plane. This tooth contacts complementary teeth that have developed, or shifted, on the fixed digit. Other Ulyxes spp are like most other Laelapidae in that adjacent teeth on a given digit are spaced closely along a proximal-distal axis. I tentatively suggest this species may be a predator merely because of the relatively strong dentition and the relative size of the digits (Table 2). Ulyxes theoclymenus is similar to U. euryclea however it differs in its unique dentition, broader genitoventral shield, its moderately well-developed apical stalk on tarsus I, and lacking an apically bifid seta on trochanter IV.	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFF0FF94FF0F0FEEBA5CFDEC.taxon	description	(Fig. 54)	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFF0FF94FF0F0FEEBA5CFDEC.taxon	materials_examined	Additional material: 1 female, Bauple State Forest, 8 Dec 1999, M. Shaw, K. Wormington, ex tree hollow in stag ca. 10 metres high occupied by regularly resident Petauroides volans Greater Glider, material vacuumed from floor of hollow one hour after glider left for nightly feeding, nest 279.	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFF0FF94FF0F0FEEBA5CFDEC.taxon	discussion	Remarks. This species is only known from Petauroides volans, the Greater Glider. Based on its small chelicerae, and the single weak tooth in the fixed digit, this species is assumed to be parasitic (Table 3).	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFF0FF94FF0F0D0ABCF8FC5C.taxon	discussion	Remarks. The species is type for the genus. It is only known from Pseudocheirus peregrinus, the Common Ringtail Possum. Original collections were from ears of its host. Based on its small chelicerae, and the single weak tooth in the fixed digit, this species is assumed to be parasitic (Table 3).	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFF0FF94FF0F0C97BDF1F831.taxon	materials_examined	Specimens examined. 1 female, 1 male, 2 deutonymphs (see Remarks), Spargo Creek, Wombat State Forest, Victoria, 29 Jan 1999, S. Ward coll., ex nestbox occupied by 19 adult and several juvenile Acrobates pygmaeus Feathertail Glider, nest 143.	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFF0FF94FF0F0C97BDF1F831.taxon	description	Description of female. Dorsum. Large mite, 900 long. Dorsal shield almost textureless centrally, margins with fine reticulations, bearing 37 pairs of setae, z 3 and px 3 absent. Unpaired setae below J 3 present. Discal setae minute. Z 5 smooth, distinctly thickened, 5 wide. Gnathosoma. Epistome a smooth lobe. Fimbriae on lateral arms of internal malae with distinctly expanded tips. Fixed digit with single tooth. Palp genu al 1 not broadened or spatulate. Palp trochanter av seta distinctly broadened, sabre-shaped, 5 wide. Deutosternal groove with 6 rows of denticles, flanked by two lateral lines. Venter. Sternal shield anterior edge subcuticular, posterior edge straight. Exopodal IV well-developed, 21 wide. Post-stigmatal shield abbreviated, with usual three pore-like structures. Genito-ventral shield extensive, reaching to within 4 of anal shield; without visible striae, possibly smooth. Zv 1 thickened, distinctly peg-like. Opisthogaster bearing 29 pairs of setae excluding Jv 1 – 3 and Zv 1 – 2. Jv 5 elongate, 93 µm (cf U. ulixes). Legs. Chaetotaxy holotrichous for Laelapidae as defined by Evans (1963). Tarsus I lacks a forked sensory seta. Genu IV with single seta on pl face, distally placed thus pl 2 appears absent or has migrated (cf. U. calypso, U. sisyphus, Figs 58, 59). Femur I ad 1 long (50) and thick (6). Other d setae on femur II – III are slightly strengthened but none longer than 32 or thicker than 4. The following setae are apically bifid: femur I ad 1, femur I pd 2, femur II ad 1, femur II ad 2, femur III ad 1, trochanterIV pl, femur IV ad 1 and femur IV ad 2. Trochanter I v seta thickened, peg-like. Trochanter IV ventral seta slightly thickened, not peg-like. Coxa II anterior seta thickened, peg-like; posterior seta slightly thickened, not peg-like. Coxa III anterior and posterior seta slightly thickened, not peg-like. Description of male. Dorsal shield with long marginal setae (cf. U. ulixes). Fixed digit slightly reduced, no teeth in fixed digit. Genital opening subcircular, 20 wide, 18 deep. Zv 1 setiform, not peg-like.	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFF0FF94FF0F0C97BDF1F831.taxon	discussion	Remarks. The examined specimens cannot be figured or formally named because they were destroyed during an attempt to remount them. These notes are offered in case they help interpret future material. This is the first record of a Ulyxes species from a possum in the family Acrobatidae. In the same locality from which these specimens were found, five other nestboxes used by Acrobates did not contain Ulyxes sp. Nor did six other nestboxes used by Acrobates in SE Queensland.	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFF1FF95FF0F0FEEBA45FB21.taxon	materials_examined	Specimen examined. 1 female (QM S 99209), Massey Creek, Queensland 17 ˚ 37 ’ 38 ” S 145 ˚ 33 ’ 45 ” E, 1 Dec 1998, coll. A. Johnson, M. Shaw, ex humus from floor of tree hollow “ Helen’s Den ” in rainforest regularly occupied by Trichosurus vulpecula johnstoni Northern Brushtail Possum,	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFF1FF95FF0F0FEEBA45FB21.taxon	description	Description of female. Dorsum. Large mite, 860 long. Dorsal shield becoming glabrous centrally, margins and opisthonotal portion with reticulations, bearing 38 pairs of setae, z 3 absent. Unpaired setae present below J 3. Z 5 smooth. Gnathosoma. Epistome appears to have smooth margin although view obscured. Fimbriae on lateral arms of internal malae with distinctly expanded tips. Fixed digit with a single tooth. Cheliceral setae dorsal. Palp genu al 1 blunt-ended with hyaline lateral edges. Palp trochanter av seta slightly broadened 4 wide. Deutosternal groove with six rows of denticles with up to six denticles per row, flanked by only two lateral lines. Subcheliceral shelf with two apical tines and two subapical irregular tines. Venter. Sternal shield anterior edge not subcuticular, excavated immediately lateral of st 1 (cf U. ulysses and U. calypso), posterior edge mildly concave. Exopodal IV well-developed, 28 wide. Post-stigmatal shield abbreviated, with usual three pore-like structures. Genito-ventral shield broad and long, reaching to within 2 of anal shield; with six broad serially-regular transverse striae. Primary metapodal shield a narrow oval, 58 long, 24 wide; secondary metapodal shields present. Anal shield very broad 190 wide at mid opening, 160 deep, 225 maximum width. Cribrum with four rows of denticles. Opisthogaster hypertrichous bearing twenty-seven pairs of setae excluding Jv 1 – 3 and Zv 1 – 2. Legs. Chaetotaxy holotrichous for Laelapidae as defined by Evans (1963). Genu IV with single seta on pl face, proximally placed. Only slight thickening of some femoral setae: Femur I ad 1 30 long and 4 wide; Femur II ad 1 ad 1 36 long and 4 wide. The following setae are apically bifid: femur 1 ad 1 and pd 2, femur II ad 1 and pd 2, femur III ad 1, femur IV ad 1 and ad 2, trochanter IV ad 1 and trochanter IV pl. Coxa II and coxa III anterior setae strong and stout.	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
03F3EC13FFF1FF95FF0F0FEEBA45FB21.taxon	discussion	Remarks. In contrast to the new Papuan species U. theoclymenus above, there are good prospects for recovering more specimens of this species in NE Queensland so I refrain from formally naming this singleton specimen here. The brief notes are given to allow initial identification and comparison of material that may be collected in future. The tree hollow this mite was collected from had been carefully monitored for several years and the resident possum and its den use were well known, but note that the area where this specimen was collected has a remarkably high diversity of arboreal marsupials. Thus host association needs to inferred with care.	en	Shaw, Matthew D. (2014): Ulyxes, a new Australopapuan mite genus associated with arboreal nests (Acari: Laelapidae). Zootaxa 3878 (3): 261-290, DOI: http://dx.doi.org/10.11646/zootaxa.3878.3.3
