identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03FB6C167440FFF15A9BFE37E485FA57.text	03FB6C167440FFF15A9BFE37E485FA57.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Fenestrobates Balogh and Mahunka 1969	<div><p>Fenestrobates Balogh and Mahunka 1969</p> <p>Type species: Fenestrobates capucinus Balogh and Mahunka, 1969, p.62</p> <p>Oribatella (Fenestrobates) capucinus (Balogh and Mahunka, 1969) Subías, 2004, p. 172.</p> <p>Diagnosis. Adult. Species comprising this genus are unique among Oribatellidae (Grandjean 1953 b, Bernini 1978) in having the following combination of character states. Integument micropunctate/microfoveolate. Lamella and pedotectum I with fine, long longitudinal striae. Rostrum medially with longitudinal ridge; rostral tectum with small u-shaped indentation medially. Notogaster flat to slightly convex anteriorly. Pteromorph with small dens ventrodistally; without ridges anteriorly on notogaster lateral of bothridium. Lenticulus absent. Octotaxic system as four pairs of porose areas. Notogaster with 10 pairs of short to long setae. Lamellae long, broad, with large, medially converging cusps; cusps with large medial and lateral dentes; translamella very short or absent; interlamellar pocket present or absent. Interlamellar setae positioned medially or laterally on posterior of lamella. Bothridium flask-shaped with indentation laterally. Genal tooth broad, subtriangular in shape, with dens ventrodistally, without longitudinal ridge ventrally. Tutorium curved, lamelliform; cusp rectangular or subrectangular, with teeth distally and dorsodistally, lying parallel to dorsal contour of prodorsum in lateral aspect, extending well anterior to insertion of rostral seta. Epimeral setal formula 2(3)–1–3–2, 2 or 3 pairs of setae on epimere I; seta 4c absent. Setae 3b, 3c and 4b, or 3b and 3c only, thick, barbed, very long. Ventral depression between pedotectum I and the lateral body wall distinctly flask-shaped, extending into podosoma (Figs. 2, 16), such that in lateral aspect invagination extending well ventral of level of pedotectum II. Pedotectum II with or without blunt tooth posterolaterally. Custodium present, with short free distal point. Chelicera chelate-dentate. Mentum with narrow tectum, overlaying insertion of seta m, without recurved ridge distally. Palp setal formula 0–2–1–3–9(+ω); eupathidium acm subequal in length to solenidion, forming double horn with solenidion along length. Axillary saccule present at base of palp. Humerosejugal porose organ Ah expressed as concave porose area, with distinct margin; Am diffuse porose area; Al porose. Six pairs of genital setae. Aggenital seta and its alveolus absent. Anal setae well spaced or together on anterior half of plate. Postanal porose area present. Legs monodactylous. Trochanter III with one seta; femur III with seta l’ present; seta v' of genua I and II present. Setae l” of genua I and II thicker, more heavily barbed and longer than setae l’ on these segments. Trochanter and femur IV each with ventral carina, that of femur IV with dens ventrodistally. One or two anterodorsal spines present on tibia I close to, or between, solenidia φ 1 and φ 2.</p> <p>Immatures. Unknown.</p> <p>Reproductive Mode. Based on data from the type series of F. marauni sp. nov., species of Fenestrobates are bisexual.</p> <p>Remarks. 1. The name Fenestrobates refers to the “fenestrate tectum” with “the large rounded foramina” that Balogh and Mahunka (1969) considered covered the epimeral region. This was a misinterpretation of the ventral depression between pedotectum I and the lateral body wall which is unusally developed in Fenestrobates. This distinct concave depression or indentation (pdp) is found in all genera of Oribatellidae. Grandjean (1956 a) illustrated this indentation for Joelia fiorii (Coggi, 1898) and then (Grandjean 1956 b) discussed it extensively, noting its position ventrally between pedotectum I and the lateral body wall. In Ophidiotrichus Grandjean, 1953, Joelia Oudemans, 1906 and Fenestrobates this depression can be seen to various extent with transmitted light in ventral mounts as a thickened, circular region. Grandjean (1956 b, his Figs B, C) gave the term ‘ tache pectorale ’ (Pe) for this region. In Fenestrobates this concave depression is very large and flask-shaped, extending ventral of the level of pedotectum II, almost reaching ventrally to the coxisternum, and giving appearance of foramen when viewed in transmitted light ventrally (compare pdp in Figs 2 and 3 and Figs 16 and 19). As Grandjean (1956 b) noted this depression does not conform to the shape of the femur or to leg setae. The position of this depression may be related to the opening of trachea I in the acetabulum of leg I, and it may function as part of a plastron system in a manner similar to those of other oribatid taxa, e.g., some Ameroidea (Chen et al. 2004). This depression is unknown outside Oribatellidae. It is absent from species of Adoribatella Woolley, 1967, previously considered a member of Oribatellidae (Behan-Pelletier 2013). Its absence in species of Tectoribates Berlese, 1910 is one of the arguments given for removal of that genus from Oribatellidae (Behan-Pelletier &amp; Walter 2013).</p> <p>2. Adults of all Oribatellidae, including Fenestrobates, have lamellae with large cusps. The type species, Fenestrobates capucinus has a small interlamellar pocket as is found in many species of Oribatella, but absent from others; it is very small, easily overlooked, and was not illustrated by Balogh and Mahunka (1969).</p> <p>3. Adults of oribatellid genera have a flask-shaped bothridium with an indentation laterally, as has Fenestrobates. This has been described clearly for Oribatella berlesei (Michael, 1898) (Bernini 1977, Plate I, Fig. 3), Fberninia helenae (Bernini, 1975) (Bernini 1975, Plate XIII, 1), Siciliotrichus siculus Bernini, 1983 (Bernini 1983, Fig. 2b), Joelia fiorii (Grandjean 1956 a, Fig. 2A), and Ophidiotrichus exastus Higgins, 1965 (Behan- Pelletier 2013).</p> <p>4. The humerosejugal porose organ Ah is usually developed as a discrete porose area in oribatellid species, including species of Fenestrobates, though it is a saccule in Oribatella quadridentata Banks, 1895. Similarly Al is developed in all oribatellids examined for this porose area, other than O. quadridentata (Behan-Pelletier 2011), and is found in Fenestrobates.</p> <p>5. A striking feature of Fenestrobates species is the long spinous form of epimeral setae 3b and 3c. In F. capucinus, seta 4b also is spinous. Such development of these setae is rare in Oribatella but is found in O. rossicus, redescribed below. In contrast, a long spinous seta 4c is found in Fberninia, Ferolocella and many species of Oribatella. This character is variable in other families of poronotic Brachypylina, e.g., in the chamobatid genus Xiphobates Pavlichenko, 1993, epimeral setae 4b and 4c can be spinose, but not setae of epimere III.</p> <p>6. The postanal porose area is expressed in all Oribatellidae, including Fenestrobates. This character also is expressed in all known Tegoribatidae and most representatives of Ceratozetoidea and Galumnoidea.</p> <p>7. The sharp dens anteroventrally on the pteromorph in Fenestrobates (Fig. 3) is a character state expressed to some extent in all Oribatellidae. In species of Joelia there is no distinct dens, rather the pteromorph is pointed anterventrally, similar to that in the ceratozetoid Lamellobates reticulatus Behan-Pelletier, 1998. Such a dens is not found in other Ceratozetoidea (species in Ceratozetidae, Punctoribatidae, Zetomimidae examined), in Tegoribatidae (Hypozetes, Tegoribates, Tectoribates examined), or in Achipteriidae (Achipteria, Parachipteria and Dentachipteria examined).</p> <p>8. An anterodorsal spine between solenidia φ 1 and φ 2 on tibia I was incorrectly considered an apomorphy of oribatellid genera by Behan-Pelletier (2013). This character state is present in species of the tegoribatid genus Tectoribates, and is absent from the oribatellid genus Ophidiotrichus (its presence in Siciliotrichus is unknown).</p> <p>9. Two other species, both from Russia, have been ascribed to this genus: Fenestrobates rossicus Krivolutsky, 1974 and F. vicinus Ryabinin 1975. These are redescribed below and considered members of Oribatella sensu stricto. Therefore, Fenestrobates is only known from South America.</p> </div>	http://treatment.plazi.org/id/03FB6C167440FFF15A9BFE37E485FA57	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ermilov, Sergey G.;Behan-Pelletier, Valerie M.	Ermilov, Sergey G., Behan-Pelletier, Valerie M. (2014): Revision of Fenestrobates (Acari, Oribatellidae) with description of F. marauni sp. nov., from South America, and new diagnosis for Oribatellidae. Zootaxa 3827 (2): 258-272, DOI: http://dx.doi.org/10.11646/zootaxa.3827.2.7
03FB6C167443FFF05A9BF96EE469FA31.text	03FB6C167443FFF05A9BF96EE469FA31.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Fenestrobates capucinus Balogh and Mahunka. Differential 1969	<div><p>Fenestrobates capucinus Balogh and Mahunka, 1969</p> <p>(Figs. 19, 20)</p> <p>Oribatella (Fenestrobates) capucinus (Balogh and Mahunka, 1969) Subías, 2004, p. 172.</p> <p>Material examined. Two paratypes (male, female) kindly loaned from the Hungarian National History Museum, Budapest.</p> <p>Diagnosis. Body size 289–333 by 196–225 (measurements from Balogh &amp; Mahunka 1969). Interlamellar setae about 120, setiform, slightly thickened, barbed, inserted posterolaterally on lamella, just anterior of bothridium. Anterior margin of notogaster distinctly flattened between bothridia. Granular cerotegument evident in dorsosejugal scissure. Notogastral setae, barbed, long 45–50 (based on unbroken setae h 1, p 1, p 2). Epimeral setal formula 2–1–3–2; setae 3b and 3c thick, barbed, long, about 45–50; seta 4b thick, barbed, long, about 60. Other epimeral setae about 20–24, thin, barbed. Ventral depression between pedotectum I and the lateral body wall, about 21 long × 15 wide, when viewed in ventral aspect.</p> <p>Description. Integument: Micropunctate. Body dark brown. Lamellae, pedotectum I and anterior of coxisternal region with distinct longitudinal striae, striae on lamella longitudinal, striae on pedotectum I transverse, those of coxisternum transverse anteriorly, longitudinal laterally. Distinct transverse line anteriorly on epimere I extending between pedotecta I.</p> <p>Prodorsum. Rostrum with two blunt teeth, margin concave between teeth. Strong, medial, longitudinal ridge posterior to indentation, extending to level of rostral setae (Fig. 20). Lamella broad, straight, almost contiguous anteromedially, fused posteromedially, interlamellar pocket very small. Lamella about 129 long, lamellar cusp (measured from translamellar region) about 93, lateral and medial dentes subequal in width, tapered, smooth, about 45 long and about 53 long, respectively. Rostral seta about 85 long, barbed, curved anteromedially. Lamellar setae about 91, thick, heavily barbed, tapered, blunt, directed anteriorly. Interlamellar setae about 120, setiform, thick, heavily barbed, inserted posterolaterally on lamella, just anterior of bothridium. Sensillus thick, setiform (without developed head), barbed, about 143, thickest about 2/3 along length, tapered. Exobothridial seta not evident. Tutorium weakly sclerotized, narrow, curved ventromedially, about 115 long, of which cusp about 60 long, with three or four strong teeth distally, two to four small teeth dorsally, with tooth ventrally (Fig. 20).</p> <p>Notogaster. Anterior margin truncate, outline straight between bothridia. Granular cerotegument evident in dorsosejugal scissure. Dorsophragmata weakly visible (not illustrated in Balogh &amp; Mahunka 1969), pleurophragmata clearly evident. Four pairs of porose areas small, rounded. Notogastral setae, setiform, barbed, long; 45–50 (based on unbroken setae h 1, p 1, p 2).</p> <p>Lateral podosomal and epimeral regions. Genal teeth strong, triangular (Fig. 20). Pedotectum I (Pd I) large, convex, appearing weakly pointed anteriorly because of ridge extending laterally on epimere I between bases of pedotecta I. Pedotectum II (Pd II) of medium size, scale-like, posterolateral tooth not visible. Porose areas Am elongate, poorly visible; Ah oval, with distinct borders; Al not visible (only visible with dissection). Custodium short, blunt-ended, directed anteriorly. Epimeral setal formula 2–1–3–2; setae 3b and 3c thick, barbed, long, about 45–50; seta 4b thick, barbed, long, about 60; setae 3b, 3c, 4a, 4b directed anteriorly. Other epimeral setae about 20–24, thin, barbed. Depression in pedotectum I oblong, about 21 long × 15 wide.</p> <p>Anogenital region. Genital setae barbed; anterior two pairs about 16; other four pairs shorter. Anal and adanal setae barbed, anal seta about 16, adanal setae about 20. Anal and adanal setae directed anteriorly. Anal setae spaced medially on anterior and posterior of anal plates. Lyrifissure iad longitudinally oriented, positioned anterolateral to anal aperture, anterior of seta ad 3 (one paratype specimen with iad unilaterally positioned anterolateral of ad 3). Postanal porose area about 7 × 3.</p> <p>Legs. As for Fenestrobates marauni sp. nov. (see below).</p> </div>	http://treatment.plazi.org/id/03FB6C167443FFF05A9BF96EE469FA31	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ermilov, Sergey G.;Behan-Pelletier, Valerie M.	Ermilov, Sergey G., Behan-Pelletier, Valerie M. (2014): Revision of Fenestrobates (Acari, Oribatellidae) with description of F. marauni sp. nov., from South America, and new diagnosis for Oribatellidae. Zootaxa 3827 (2): 258-272, DOI: http://dx.doi.org/10.11646/zootaxa.3827.2.7
03FB6C167442FFF55A9BFA3FE66DF800.text	03FB6C167442FFF55A9BFA3FE66DF800.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Fenestrobates marauni Ermilov & Behan-Pelletier 2014	<div><p>Fenestrobates marauni sp. nov.</p> <p>(Figs 1–15, 16–18)</p> <p>Material examined. The study material (eight specimens) was collected by Dorotee Sandmann from upper organic soil layer in mostly undisturbed rain forest in Southern Ecuador. Holotype and five paratypes: 4°60' S, 78°58'–79°10' W, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.96667&amp;materialsCitation.latitude=-5.0" title="Search Plazi for locations around (long -78.96667/lat -5.0)">Cajanuma</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.96667&amp;materialsCitation.latitude=-5.0" title="Search Plazi for locations around (long -78.96667/lat -5.0)">Podocarpus National Park</a>, 3000 m. a.s.l., 01.04.2009; two paratypes: 4°70' S, 78°58' W, Bombuscaro, Podocarpus National Park, 1050 m a.s.l., 01.10.2008 and 01.04.2009.</p> <p>Diagnosis. Distinguished from the type species by a combination of the following character states: body size 266–340 by 172–229; tutorium wide subrectangular with four to seven teeth distally, two to three teeth on dorsal side and with or without one to two teeth on ventral side; interlamellar seta positioned posteromedially on lamella; notogastral setae short, thin, smooth; epimeral setal formula 2(3)–1–3–2, 2 or 3 pairs of setae on epimere I, only setae 3b and 3c thick, barbed, others short, smooth; anogenital setae short; anal setae inserted in anterior part of anal plate.</p> <p>Description. Measurements. Body length: 307 (holotype, male), 266–340 (five paratypes: one male, four females); notogaster width: 192 (holotype male), 172–229 (five paratypes: one male, four females).</p> <p>Integument. Body dark brown to light brown. Body microfoveolate dorsally, laterally and in anogenital region (including anal plates); diameter microfoveae up to 1. Lamella, pteromorph laterally and pedotectum I with long longitudinal striae.</p> <p>Prodorsum. Rostrum with two teeth and indentation between them (clearly visible in dorso-frontal and frontal views). Strong, longitudinal ridge (r) located posteriorly to indentation. Lamella broad, straight, contiguous anteromedially, connected posteromedially. Lamella with two long (longer than half of lamella), tapered, smooth dentes. Median dens straight; lateral dens slightly shorter, slightly directed anteromedially. Rostral seta (ro, 73–82) setiform, curved anteromedially, with dense cilia in medioproximal part and with fewer barbs in distal part unilaterally. Lamellar seta (le, 73–82) thick, heavily barbed, tapered, parallel to midline. Interlamellar seta (in, 118–127) setiform, slightly thickened, barbed. Sensillus (ss, 110–118) rather thick, setiform (without developed head), barbed. Exobothridial seta (ex) absent, represented by alveolus. Tutorium (tu) long, with four to seven strong teeth distally, two to three small teeth on dorsal side and without or with one or two small teeth on ventral side.</p> <p>Notogaster. Slightly longer than width. Anterior margin undulating. Dorsophragmata (D) oval, located close to each other. Four pairs of porose areas small, rounded, often poorly visible: Aa (4–6, diameter) little larger than others (A1, A2, A3, 2–4). Notogastral setae short (8–12), setiform, thin, smooth. Three pairs of setae (p 1 – p 3) inserted in posteromarginal position. Distance p 2 – p 3 shorter than p 1 – p 2. Opisthonotal gland opening (gla) located lateroposteriorly to A1. Lyrifissures im and ip distinctly visible; ia, ih and ips indistinct.</p> <p>Gnathosoma. Subcapitulum longer than wide (86–94 × 61–69). Subcapitular setae (h, m, a) setiform, slightly barbed, similar in length (10–12). Two pairs of adoral setae (or 1, or 2, 8) setiform, straight, slightly barbed. Palps (69) with setation 0–2–1–3–9(+ω); solenidion thickened, straight. Chelicerae (102) with two setiform, barbed setae; cha (36) longer than chb (16). Trägårdh’s organ (Tg) distinct. Axillary saccule about 4 × 2.</p> <p>Lateral podosomal and epimeral regions. Genal tooth strong, triangular. Pedotectum I (Pd I) large, concave, weakly pointed anteriorly (visible in lateral view). Ventral depression between pedotectum I and the lateral body wall deep flask-shaped (pdp). Pedotectum II (Pd II) of medium size, scale-like, with postero-lateral blunt tooth (visible in ventro-lateral view). Porose area Am elongate, poorly visible; Ah oval, with distinct borders, Al poorly visible, overlain by base of lamella. Custodium (cus) long, thin, blunt-ended, directed anteriorly. Discidium (dis) triangular, pointed. Circumpedal carina (cp) well developed. Epimeral setal formula 2(3)–1–3–2. Pair of setae 1c present in two specimens. Setae 3b (49–57) and 3c (36–41) thick, barbed. Medial (1a, 2a, 3a) and other setae (1b, 1c, 4a, 4b, 8–10) short, thin, smooth.</p> <p>Anogenital region. Anogenital setae short, thin. Anterior two pairs of genital setae (g 1, g 2, 6–8) slightly barbed; other four pairs shorter (g 3 – g 6, 4–6), smooth. Aggenital setae and their alveoli absent. Two pairs of anal (an 1, 2–4; an 2, 4–6) and three pairs of adanal (ad 1 – ad 3, 4–6) setae smooth. Anal setae inserted close to each other, in anterior part of anal plate. Lyrifissures iad longitudinally oriented, located lateroanteriorly to anal aperture. Postanal porose area about 10 × 5.</p> <p>Legs. Femora IV and trochanters IV with ventral ridges. Porose areas on femora and trochanters III, IV present, but poorly visible. Formulae of leg setae and solenidia: I (1–5–3–4–20) [1–2–2], II (1–5–3–4–15) [1–1–2], III (1–3–1–3–15) [1–1–0], IV (1–2–2–3–12) [0–1–0]; homology of setae and solenidia indicated in Table 1. Trochanters III with only one seta. Tibiae and genua III, IV without thick setae. Genua I, II with thick seta, l”. Famulus short, straight, blunt-ended. Most setae (except smooth p) barbed, some ventral setae on tarsi and tibiae with strong teeth. Solenidia simple.</p> <p>Roman letters refer to normal setae (e to famulus), Greek letters to solenidia. Single prime (') marks setae on anterior and</p> <p>double prime (") setae on posterior side of the given leg segment. Parentheses refer to a pseudosymmetrical pair of setae.</p> <p>Immatures. Unknown.</p> <p>Type deposition. The holotype is deposited in the collection of the Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia; two paratypes are deposited in the collection of the Siberian Zoological Museum, Novosibirsk, Russia; three paratypes are deposited in the collection of the Tyumen State University Museum of Zoology, Tyumen, Russia; two paratypes are in the personal collection of the second author.</p> <p>Etymology. We name this species in honour of Dr. Mark Maraun, who has contributed extensively and significantly to the ecology of Oribatida.</p> <p>Remarks. Fenestrobates marauni sp. nov. is similar to F. capucinus in body size and shape of sensillus. It differs from the type species by the following character states (those of F. capucinus in parentheses): interlamellar seta positioned posteromedially on lamella (positioned posterolaterally on lamella); notogastral setae short, smooth, 8–12 (notogastral setae long, barbed, 45–50); epimeral seta 4b short, thin, smooth, 8–10 (4b thick, spinous, long, about 60).</p> </div>	http://treatment.plazi.org/id/03FB6C167442FFF55A9BFA3FE66DF800	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ermilov, Sergey G.;Behan-Pelletier, Valerie M.	Ermilov, Sergey G., Behan-Pelletier, Valerie M. (2014): Revision of Fenestrobates (Acari, Oribatellidae) with description of F. marauni sp. nov., from South America, and new diagnosis for Oribatellidae. Zootaxa 3827 (2): 258-272, DOI: http://dx.doi.org/10.11646/zootaxa.3827.2.7
03FB6C167448FFFA5A9BFF30E4F1F969.text	03FB6C167448FFFA5A9BFF30E4F1F969.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oribatella rossicus (Krivolutsky 1974)	<div><p>Oribatella rossicus (Krivolutsky, 1974)</p> <p>Fenestrobates rossicus Krivolutsky, 1974 p. 1883</p> <p>Material examined. Cotypes (male, female, on single slide) kindly loaned from the Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia.</p> <p>Diagnosis. Total length of adult 630 (measurement from Krivolutsky 1974). Integument of prodorsum, notogaster, venter and mentum micropunctate. Integument of coxisternum with distinct striae. Rostrum medially rounded with pair of small lateral teeth on margin, as in Oribatella brevicornuta Jacot, 1934 (see Fig. 2D, E, F in Behan-Pelletier 2011). Lamella (with cusp) about 191 long, cusp about 128 long, 63 wide at level of insertion of seta le. Lamellar cusps diverging anteromedially; separated posteromedially, leaving prodorsum visible through oval opening × about 12 at widest. Medial dens of lamellar cusps about 61, subequal or slightly shorter than lateral dens (68 long); medial dens smooth; lateral dens with about 3–4 small teeth on lateral edge (varying on either side of type female specimen). Long striae extending longitudinally along outer margin of lamella. Translamella without tooth, about 12 at greatest width, about 22 deep. Seta ro about 132 long, strongly barbed along length, acuminate, directed anteromedially. Seta le about 110 long, thick, heavily barbed, tapered. Setae in 213 long, thick (less so than le), heavily barbed, tapered; mutual distance of pair about 104 µm. Sensillus 127 long, subequal in width to seta le, except head slightly fusiform, stalk with small barbs distally; head directed slightly anteromedially. Exobothridial setae not visible. Genal tooth broad, subtriangular, rounded distally, without dens ventrodistally, with longitudinal ridge ventrally. Tutorium including cusp about 235 long, rectangular, distal margin with about eight dens. Ventral depression between pedotectum I and the lateral body wall similar to that in O. brevicornuta. Custodium about 56 long, Anterior margin of notogaster undulating, convex lateral of bothridium, with about 10 transverse ridges. Pteromorph with dens anteroventrally. Four pairs of porose areas present, Aa oval about 34 at widest. Eleven pairs of long, barbed notogastral setae present (data from Krivolutsky 1974); setae barbed, tapered, 14–30 long. Distance h 1 –h 1 and p 1 –p 1 could not be established. Lenticulus subtriangular. Epimeral setation 3(?)–1–3–3; about 12 to 48, barbed, tapered, except 4a, 4b, 4c, 3b, 3c long, spinous; 3b about 96, reaching anteriorly 3c about 80, 4a, 4b about 77, 4c about 48. Setae 3b, 4a, 4b, 4c strongly directed anteriorly to level of indentation of pedotectum I, seta 3b in longitudinal alignment with long spinous seta l’ of tibia IV when legs folded. Genital setae six pairs, two seta on anterior of genital plate; genital and aggenital setae about 11 long. Mentum with anterior tectum covering base of gena, reflexed anteriorly, similar in size and shape to that for Oribatella yukonensis Behan-Pelletier and Walter, 2012 (their Fig. 14G). Seta l” of tibia I about 54, seta l’ of tibia IV about 54. Tarsi heterotridactylous.</p> <p>Remarks. The description of this species by Krivolutsky (1974) is brief and lacks measurements other than for body length. The expanded description above is based on the slide mounted type specimens. These are mounted dorsal and ventral and thus some characters could not be re-evaluated. For example, Krivolutsky (1974) noted “Trochanters III, IV with long, thin, finger-like structure, directed anteriorly”; we were unable to evaluate this further.</p> <p>Krivolutsky (1974) assigned this species to Fenestrobates on the basis of “Pore-like structure on coxes of legs II rounded, clearly visible in ventral view”. This is referring to the ventral depression between pedotectum I and the lateral body wall as seen through the coxisternal integument. However, we do not consider this depression to be as extensive as in the species of Fenestrobates, or to be flask-shaped. This species shares the long spinous setae 3b with species of Fenestrobates, but seta 4c is present in O. rossicus, whereas this seta is absent in species of Fenestrobates. Furthermore O. rossicus has about 10 ridges on the anterior margin of the notogaster, ridges that are present in all described species of Oribatella, but absent in Fenestrobates.</p> </div>	http://treatment.plazi.org/id/03FB6C167448FFFA5A9BFF30E4F1F969	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ermilov, Sergey G.;Behan-Pelletier, Valerie M.	Ermilov, Sergey G., Behan-Pelletier, Valerie M. (2014): Revision of Fenestrobates (Acari, Oribatellidae) with description of F. marauni sp. nov., from South America, and new diagnosis for Oribatellidae. Zootaxa 3827 (2): 258-272, DOI: http://dx.doi.org/10.11646/zootaxa.3827.2.7
03FB6C167448FFF95A9BF8C1E3E6FA85.text	03FB6C167448FFF95A9BF8C1E3E6FA85.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oribatella vicinus (Ryabinin 1975)	<div><p>Oribatella vicinus (Ryabinin, 1975)</p> <p>Fenestrobates vicinus Ryabinin, 1975 p. 540</p> <p>Material examined. Holotype male, kindly loaned from the Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia.</p> <p>Diagnosis. Total length of adult male about 576. Integument of prodorsum, notogaster, venter and mentum micropunctate. Integument of lenticulus and coxisternum with distinct striae, as in Oribatella jacoti Behan- Pelletier 2011. Rostrum medially rounded with pair of small lateral teeth on margin, as in O. brevicornuta Jacot, 1934 (see Fig. 2D, E, F in Behan-Pelletier 2011). Lamella (with cusp) about 162 long, cusp about 108 long, 62 wide at level of insertion of seta le. Lamellar cusps diverging anteromedially; separated posteromedially, leaving prodorsum visible through oval opening about 14 at widest. Medial dens of lamellar cusps about 52 – 55 µm, subequal to length lateral dens; medial dens smooth; lateral dens with about three small teeth on lateral edge. Long striae extending longitudinally along outer margin of lamella. Translamella without tooth, about 14 at greatest width, about 26 deep. Setae ro strongly barbed along length, acuminate, strongly directed medially. Setae le about 79 long, thick, heavily barbed, tapered. Setae in 172 long, thick (less so than le), heavily barbed, tapered; mutual distance of pair about 81 µm. Sensillus 141 long, subequal in width to seta le, except head slightly fusiform, tapered, stalk with small barbs distally; head directed slightly anteromedially, as in O. brevicornuta (see Fig. 3, insert in Behan-Pelletier 2011). Exobothridial seta not visible. Genal tooth broad, subtriangular, rounded distally, without dens ventrodistally, with longitudinal ridge ventrally. Tutorium including cusp about 168 long, rectangular, distal margin with eight to 10 dentes. Ventral depression between pedotectum I and the lateral body wall less defined than that in O. brevicornuta. Custodium about 58 long, extending to anterior margin of pedotectum II. Anterior margin of notogaster undulating, convex lateral of bothridium, with about 10 transverse ridges. Pteromorph with dens anteroventrally. Four pairs of porose areas present, Aa circular about 12 at widest. Ten pairs of long, barbed notogastral setae present; setae barbed, acuminate, almost flagellate 52 – 79 long (Fig. 4 of Ryabinin 1975). Setae h 1 directed posteriorly and parallel as in O. jacoti (Behan-Pelletier, 2011). Distance h 1 –h 1 and p 1 –p 1 subequal, about 23 – 26. Lenticulus subtriangular. Epimeral setation 3(?) – 1 – 3 – 3; barbed, tapered, length not established except 3b, about 72, 4a, 4b about 29 – 36, long, spinous. Setae 3b, 4a, 4b, strongly directed anteriorly (Fig. 5 of Ryabinin 1975). Genital setae 6 pairs, two seta on anterior of genital plate. Mentum with anterior tectum covering base of gena, reflexed anteriorly, similar in size and shape to that for Oribatella yukonensis Behan- Pelletier and Walter, 2012 (their Fig. 14G). Seta l” of tibia I about 53. Tarsi heterotridactylous.</p> <p>Remarks. The description of this species by Ryabinin (1975) is brief and lacks measurements. We redescribe this species based on the slide mounted holotype male. This is a dorsal mount which precludes evaluation of ventral and lateral characters. Ryabinin (1975) assigned this species to Fenestrobates on the basis of its similarity to O. rossicus. He stated “The new species is similar to F. rossicus from Primorye territory (Krivolutsky 1974)”. The species are similar but can be easily separated by the long, acuminate notogastral seta in O. vicinus vs. the short notogastral setae of O. rossicus; the very long epimeral seta 3b (96) in O. rossicus vs. shorter seta (72) in O. vicinus; setae h 1 directed posteriorly and parallel in O. vicinus vs. curved h 1 in O. rossicus. We were unable to verify whether O. rossicus had 11 pairs of notogastral setae vs. the 10 pairs found in O. vicinus.</p> </div>	http://treatment.plazi.org/id/03FB6C167448FFF95A9BF8C1E3E6FA85	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ermilov, Sergey G.;Behan-Pelletier, Valerie M.	Ermilov, Sergey G., Behan-Pelletier, Valerie M. (2014): Revision of Fenestrobates (Acari, Oribatellidae) with description of F. marauni sp. nov., from South America, and new diagnosis for Oribatellidae. Zootaxa 3827 (2): 258-272, DOI: http://dx.doi.org/10.11646/zootaxa.3827.2.7
03FB6C16744BFFFF5A9BF8B1E537FCEB.text	03FB6C16744BFFFF5A9BF8B1E537FCEB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oribatellidae Jacot 1925	<div><p>Oribatellidae Jacot, 1925</p> <p>Diagnosis. Adult. Poronotic Brachypylina with the following character states. Prodorsum. Lamellae long, broad, with large cusps, medially converging, diverging or contiguous; cusps with or without medial and lateral dentes; translamella variable in width and depth, with or without medial tooth, or translamella absent. Bothridium flaskshaped with indentation laterally. Notogaster. Immovable pteromorphs with small dens anteroventrally. Notogaster anteriorly flattened to convex, with or without ridges lateral of bothridium. Lenticulus present or absent. Pair of cavities on notogaster present or absent. Octotaxic system as four pairs of porose areas or saccules; usually without sexual dimorphism, when present that of male modified, and number of porose areas reduced. Notogaster with 10 to 14 pairs of short to very long setae. Lateral podosomal and epimeral regions. Genal tooth subtriangular to subrectangular in shape, with or without dens ventrodistally, with or without ridge ventrally. Tutorium lamelliform, narrow to wide, cusp rectangular or subrectangular, with dentes distally, lying parallel to dorsal contour of prodorsum in lateral aspect, extending or not anterior to insertion of rostral seta, or cusp of tutorium absent. Humerosejugal porose organ Ah discrete, expressed as porose area or saccule; Am diffuse porose area; Al when present expressed as porose area or saccule. Custodium usually present, with short to long, free distal point. Epimeral setal formula 3–1–3–3 or 2(3)–1–3–2 or 3–1–3–2 or 2–1–3–2 or 3–1–2–2; seta 4c present or absent. Setae 3b, 3c and sometimes 4b (Fenestrobates capucinus), and 4c (most species of Oribatella) thick, spinous, very long. Pedotectum I large. Ventral depression between pedotectum I and the lateral body wall varying from small Ushaped depression to large flask-shaped pit depending on genus. Pedotectum II with or without blunt tooth posterolaterally or paraxially. Gnathosoma. Axillary saccule present at base of palp. Subcapitulum diarthric. Chelicera chelate-dentate, with 2 cheliceral setae; (chelicera elongated in one species of Oribatella and Joelia). Mentum with or without tectum, with or without recurved ridge distally. Palp setal formula 0–2–1–3–9(+ω) or 0–2–1–3–8(+ω); eupathidium acm subequal in length to solenidion, forming double horn with solenidion along length or distally (in Oribatella quadridentata Banks 1895 solenidion almost 2.5 × length of acm). Anogenital region. Six pairs of genital setae. Aggenital seta present or absent. Two pairs anal setae; three pairs adanal setae. Postanal porose area present (data for Siciliotrichus Bernini, 1983 unknown). Legs monodactylous, heterobidactylous or heterotridactylous. Trochanter III with seta l’ present or absent; femur III with seta l’ present or absent; seta v' of genua I and II present or absent. Genua I and II with or without small cusp ventrodistally. Setae l” of genua and tibiae I and II thicker, more heavily barbed and longer than setae l’ on these segments. One or two anterodorsal spines present or not on tibia I close to, or between, solenidia φ 1 and φ 2.</p> <p>Immature. Apopheredermous, with scalps of preceding instar maintained away from dorsal integument by modified setae da and dorsally directed h 1 (Oribatella), or modified setae da and dorsally directed setae dp and c 1 (Ophidiotrichus). Setae da usually showing modification for attachment of scalp of preceding instar, may be serpentine in shape with flattened tip (Ophidiotrichus). Axillary saccule present, where investigated. Body usually colorless, cuticle without plicae; hysterosomal sclerites present or absent. Gastronotal setation usually unideficient; larva with 11 or 12 pairs, protonymph with 14 or 15 pairs, deutonymph and tritonymph with 13 or 15 pairs. Setae dm and dp subequal in length to da in all immatures, or much shorter than da in deutonymph and tritonymph, or dm and dp (one species), or c 1 and dm (one species) absent from deutonymph and tritonymph. Pair of humeral organs present laterally in sejugal region. Without apodemato-acetabular tracheal system or porose homologues. Paraprocts atrichous in larva, protonymph and deutonymph. Genital setal formula (larva to adult): 0–1–3–5–6. Aggenital setal formula 0–0–1–1–1, or aggenital seta absent. Opisthonotal gland present in all instars. Cupule development normal. Bothridium and bothridial seta fully formed in all instars. Setation of protonymphal leg IV normal (0–0–0–0–7). Larva to deutonymph with circular line of dehiscence, such that dorsal scalp separates from ventral piece at ecdysis.</p> <p>Remarks. The adult diagnosis given is based on that of Grandjean (1953 b), the extensive research of Bernini (1975, 1978, 1983) publications by Shtanchaeva &amp; Subías (2009), Behan-Pelletier (2011, 2013), Behan-Pelletier &amp; Walter (2012), Ermilov &amp; Anichkin (2012) and Seniczak &amp; Seniczak (2013).</p> <p>The immature diagnosis is mainly based on immatures of Oribatella and Ophidiotrichus. Immatures of Ophidiotrichus tectus (Michael, 1884) were described by Grandjean (1953b). Immatures of Fenestrobates and other oribatellid genera, Fberninia, Ferolocella, Siciliotrichus are unknown, but the presumed deutonymph of Joelia fiorii (Coggi, 1898) was described by Grandjean (1956 b). The seminal paper by Grandjean (1953 a) on immatures of O. calcarata (C.L. Koch, 1835) is the framework for any evaluation of oribatellid immatures, which, as Behan-Pelletier and Walter (2012) noted, continue to surprise in their variation. For example, Seniczak and Seniczak (2013) describe immatures of Oribatella calcarata that do not bear scalps. Furthermore, some nymphs they examined lacked notogastral setae c 1 and dm.</p> <p>Ignorance of immatures of most oribatellid genera undoubtedly contributes to disagreement in the literature as to the number of genera that should be included in Oribatellidae, for example, Schatz et al. (2011; 10 genera) and Subías (2014; 8 genera). Subías (2014) included Cuspidozetes Hammer, 1962, Ferolocella Grabowski, 1971, Joelia Oudemans, 1906, Novoribatella Engelbrecht, 1986, Ophidiotrichus Grandjean, 1953, Oribatella, Prionoribatella Aoki, 1975 and Siciliotrichus Bernini, 1983. Fberninia Özdikmen, 2008 and Fenestrobates are considered subgenera of Oribatella by Subías (2014), though we treat them as genera, herein. Overall we follow Subías’ (2014) concepts with the following reservations.</p> <p>We consider Cuspidozetes a member of Ceratozetidae. Hammer (1962) noted that femora I and II of the type species, C. armatus Hammer, 1962, are similar to those of Melanozetes and Fuscozetes. Furthermore, her illustration of tibia and tarsus I of this species shows a setiform seta l”, whereas this seta is spinous in all genera of Oribatellidae.</p> <p>Aoki (1975) described Prionoribatella as being similar to Oribatella but with: the lamellar cusp having a small inner tooth and a large and broad outer tooth; lamellar seta inserted closer to the inner tooth; legs monodactylous; and the rostrum pointed, without incisions. These characters states have been found in other species of Oribatella (Behan-Pelletier 2011, Shtanchaeva &amp; Subías 2009), and we consider Prionoribatella a subgenus of Oribatella.</p> <p>Engelbrecht (1986) proposed Novoribatella with Tectoribates transcriptus Mahunka, 1985 as type species, and described three additional species, all well illustrated. None shows the ventral depression between pedotectum I and the lateral body wall, the character state that is a distinct apomorphy for Oribatellidae. Species of Novoribatella have a morphology closely related to that of Tectoribates, recently revised for North America by Behan-Pelletier and Walter (2013). It is possible that Novoribatella is also a member of the Tegoribatidae, as has been proposed by the latter authors for Tectoribates.</p> </div>	http://treatment.plazi.org/id/03FB6C16744BFFFF5A9BF8B1E537FCEB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ermilov, Sergey G.;Behan-Pelletier, Valerie M.	Ermilov, Sergey G., Behan-Pelletier, Valerie M. (2014): Revision of Fenestrobates (Acari, Oribatellidae) with description of F. marauni sp. nov., from South America, and new diagnosis for Oribatellidae. Zootaxa 3827 (2): 258-272, DOI: http://dx.doi.org/10.11646/zootaxa.3827.2.7
03FB6C16744DFFFF5A9BFC5CE2BEF989.text	03FB6C16744DFFFF5A9BFC5CE2BEF989.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oribatellidae Jacot 1925	<div><p>Key to genera of Oribatellidae</p> <p>1. Rostral setae spinose. Tutorium without cusp (Bernini 1983, Fig. 2), not extending anterior of rostral seta...... Siciliotrichus</p> <p>- Rostral seta setose with large barbs. Tutorium with cusp, extending anterior of rostral seta........................... 2</p> <p>2(1) Interlamellar pocket large, box-like, reaching posterior of dorsosejugal scissure. Medial longitudinal ridge on coxisternum present.................................................................................... Ferolocella</p> <p>- Interlamellar pocket present or absent, if present, never large, box-like, never reaching posterior of dorsosejugal scissure. Medial longitudinal ridge on coxisternum absent............................................................. 3</p> <p>3(2) Notogastral setae lm positioned lateral to porose area Aa and lp positioned lateral to A1 (Behan-Pelletier 2013 Fig. 10). Dentes of lamellar cusps short and broadly tapered..................................................... Ophidiotrichus</p> <p>- Notogastral setae lm positioned medial or posterior to porose area Aa and lp positioned lateral or anterior to A1. Dentes of lamellar cusps long and sharply tapered................................................................... 4</p> <p>4(3) Lamellar cusp with ventral keel (Behan-Pelletier 2013 Fig. 2F). Translamella without tooth...................... Joelia</p> <p>- Lamellar cusp without ventral keel. Translamella with or without tooth........................................... 5</p> <p>5(4) Notogastral cavities present (Bernini 1975 Fig. 16b). Custodium absent................................... Fberninia</p> <p>- Notogastral cavities absent. Custodium present.............................................................. 6</p> <p>6(5) Anterior margin of notogaster distinctly undulating with 5 or more fine ridges lateral of bothridium (Behan-Pelletier 2011 Fig. 20A). Pouch-like depression positioned ventrally between pedotectum I and the lateral body wall absent. Epimeral setae 1c and 4c present................................................................................ Oribatella</p> <p>- Anterior margin of notogaster weakly undulating with no ridges lateral of bothridium. Pouch-like depression positioned ventrally between pedotectum I and the lateral body wall present (Fig. 3, 16). Epimeral seta 1c present or absent, 4c absent.............................................................................................. Fenestrobates</p></div> 	http://treatment.plazi.org/id/03FB6C16744DFFFF5A9BFC5CE2BEF989	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ermilov, Sergey G.;Behan-Pelletier, Valerie M.	Ermilov, Sergey G., Behan-Pelletier, Valerie M. (2014): Revision of Fenestrobates (Acari, Oribatellidae) with description of F. marauni sp. nov., from South America, and new diagnosis for Oribatellidae. Zootaxa 3827 (2): 258-272, DOI: http://dx.doi.org/10.11646/zootaxa.3827.2.7
