taxonID	type	description	language	source
0B3187C8FFA5894BFDFD3433FBBEAFAE.taxon	type_taxon	Type species Amphiascus typhlops Sars, 1906, by original designation.	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FFA5894BFDFD3433FBBEAFAE.taxon	diagnosis	Emended diagnosis Miraciidae: Diosaccinae. Body fusiform; length: width ratio of caudal rami from about 1.0 to about 3.0. Rostrum not fused to cephalothorax, triangular, elongate, as long as first and second antennulary segments combined, with pointed tip, with one subdistal sensilla on each side. Female genital and third urosomites distinct dorsally and / or laterally, completely fused ventrally forming genital double-somite; genital and third urosomites separated in males. Length: width ratio of caudal rami variable, from 1: 1 to 3: 1. Female antennule seven- to nine-segmented, with aesthetasc and seta fused basally on fourth segment, and with apical acrothek on last segment. Male antennule haplocer, eight – ten-segmented, with aesthetasc and seta fused basally on fifth segment, and with apical acrothek on last segment. Antennulary setae typically smooth, except for few slightly setulose setae on penultimate and last articles in females and males. Antenna with allobasis; exopod three-segmented, armature formula 1 - 0 - 1,3,0. Mandibular palp biramous; endopod one-segmented, larger than exopod; exopod two-segmented. Maxillule biramous; rami one-segmented. Maxilla with three endites; endopod two-segmented. Maxilliped subchelate; (syn) coxa with two to four, basis with two setae; endopod one-segmented, with claw and variable number of accompanying setae. P 1 – P 4 with three-segmented rami. Male P 1 basis sexually dimorphic, with normal or modified (elongate, blunt) inner spine, without or with variable number of inner accessory spines (two – eight). Male P 2 sexually dimorphic, two-segmented; first segment with one inner seta, second segment with two inner setae, two subdistal elements (one of which thick and blunt) and two elements (one medial, one apical) issuing from elongate cylindrical extension fused to segment basally. P 5 EXP typically longer than wide, with four to six setae in the female and male; P 5 endopodal lobe with four to five setae in the female, with two setae in the male; two innermost elements of the female P 5 endopodal lobe and armature of the male P 5 endopodal lobe bifurcate distally. Armature formula of P 1 – P 5 as follows: 1 The report of eight setae on the female P 5 EXP of T. blanchardi is most probably erroneous. 2 Typhlamphiascus typhlops is known to lack inner armature on the P 4 ENP 2. The inner seta of P 4 ENP 2 of T. capensis comb. nov. might have become detached from the supporting segment as indicated by the interrupted cuticle (Kunz 1975). Sexual dimorphism expressed in the male antennule (see above), P 1 basis, P 2 ENP (see above), P 5 (see above), structure of P 6, and in the separated genital and third urosomites.	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FFA98957FDC73676FD21A90A.taxon	description	urn: lsid: zoobank. org: act: 67 A 13 A 9 C- 4 D 1 E- 4 F 86 - 82 F 2 - CEE 5407 BCD 26 Figs 1 – 10	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FFA98957FDC73676FD21A90A.taxon	etymology	Etymology The specific epithet from the Latin ‘ medicī’, ‘ doctor’, ‘ physician’, is dedicated in honour and to the memory of all physicians and health personnel for their self-sacrifice during the fight against the COVID- 19 pandemic. It is in the genitive singular, gender masculine.	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FFA98957FDC73676FD21A90A.taxon	materials_examined	Material examined Holotype MEXICO • ♀ (preserved in alcohol); Sinaloa State, Mazatlán, Urías coastal system, stn 2 (see also Gómez 2020 a: 43, fig. 1); 23.1587 ° N, 106.3326 ° W; 1.8 m depth; 18 Jan. 2019; S. Gómez leg.; organic carbon content 3.99 %, organic matter content 6.86 %, sand 80.42 %, clay 8.29 %, silt 11.28 %; ICML- EMUCOP- 180119 - 40. Allotype MEXICO • ♂ (preserved in alcohol); same collection data as for holotype; ICML-EMUCOP- 180119 - 43. Other paratypes MEXICO • 3 ♀♀ (preserved in alcohol); same collection data as for holotype; ICML-EMUCOP- 180119 - 103 • 2 ♀♀, 2 ♂♂ (preserved in alcohol); same collection data as for holotype; ICML-EMUCOP- 180119 - 46 • 1 ♀ (dissected); same collection data as for holotype; ICML-EMUCOP- 180119 - 41 • 1 ♀ (dissected); same collection data as for holotype; ICML-EMUCOP- 180119 - 42 • 1 ♂ (dissected); same collection data as for holotype; ICML-EMUCOP- 180119 - 44 • 1 ♂ (P 1 and P 2 dissected, rest intact and preserved in alcohol); same collection data as for holotype; ICML-EMUCOP- 180119 - 45. Other material MEXICO • 8 CV, 4 CIV, 1 CII (preserved in alcohol); same collection data as for holotype; ICML- EMUCOP- 180119 - 47 • 1 ♀, 2 ♂♂, 2 CV (preserved in alcohol); Sinaloa State, Mazatlán, Urías coastal system, stn 4 (see also Gómez 2020 a: 43, fig. 1); 23.1840 ° N, 106.3579 ° W; 0.7 m depth; 18 Jan. 2019; S. Gómez leg.; organic carbon content 1.13 %, organic matter content 1.94 %, sand 82.44 %, clay 8.27 %, silt 9.29 %; ICML-EMUCOP- 180119 - 48 • 1 CV, 1 CIV, 1 CI (preserved in alcohol); Sinaloa State, Mazatlán, Urías coastal system, stn 5 (see also Gómez 2020 a: 43, fig. 1); 23.2056 ° N, 106.3715 ° W; 0.6 m depth; 18 Jan. 2019; S. Gómez leg.; organic carbon content 0.99 %, organic matter content 1.71 %, sand 78.61 %, clay 6.72 %, silt 14.67 %; ICML-EMUCOP- 180119 - 49 • 1 ♂, 1 CIII (preserved in alcohol); same collection data as for preceding; ICML-EMUCOP- 180119 - 50 • 1 ♀, 2 CIV, 1 CII (preserved in alcohol); same collection data as for preceding; ICML-EMUCOP- 180119 - 51 • 1 CV, 1 CIII, 2 CII (preserved in alcohol); Sinaloa State, Mazatlán, Urías coastal system, stn 6 (see also Gómez 2020 a: 43, fig. 1); 23.2123 ° N, 106.3780 ° W; 1.4 m depth; 18 Jan. 2019; S. Gómez leg.; organic carbon content 1.65 %, organic matter content 2.84 %, sand 61.44 %, clay 14.57 %, silt 23.99 %; ICML- EMUCOP- 180119 - 52 • 1 ♂, 1 CV, 1 CIV (preserved in alcohol); same collection data as for preceding; ICML-EMUCOP- 180119 - 53 • 1 ♂ (preserved in alcohol); same collection data as for preceding; ICML- EMUCOP- 180119 - 54) • 1 ♀, 1 CIV (preserved in alcohol); Sinaloa State, Mazatlán, Urías coastal system, stn 10 (see also Gómez 2020 a: 43, fig. 1); 23.1815 ° N, 106.4214 ° W; 6.0 m depth; 18 Jan. 2019; S. Gómez leg.; organic carbon content 1.2 %, organic matter content 2.07 %, sand 69.12 %, clay 7.91 %, silt 22.97 %; ICML-EMUCOP- 180119 - 55 • 3 CIV (preserved in alcohol); same collection data as for preceding; ICML-EMUCOP- 180119 - 56.	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FFA98957FDC73676FD21A90A.taxon	description	Description Female GENERAL. Total body length measured from tip of rostrum to posterior margin of caudal rami ranging from 705 to 850 µm (mean 762 µm; n = 5; total body length of holotype 850 µm); habitus fusiform, widest at posterior end of cephalothorax in dorsal view, tapering posteriad (Fig. 1 A). PROSOME (Figs 1 A – B, 2 A). Consisting of cephalothorax with fused first pedigerous somite and second to fourth free pedigerous somites; prosomites without expansions nor spinular ornamentation; posterior hyaline frill of cephalothorax, second and third pedigerous somites broad, with posterior margins finely serrated, striated, of fourth pedigerous somite visibly narrower, finely serrated, without striations. UROSOME (Figs 1 A – D, 2 A). Consisting of fifth pedigerous somite (first urosomite), genital doublesomite (genital – second urosomite – and third urosomites fused), two free urosomites, anal somite and caudal rami; urosomites without expansions. First urosomite narrower than preceding somites in dorsal view (Fig. 1 A), without spinular ornamentation, with posterior sensilla as depicted, with one lateral pore on each side (Fig. 1 C), posterior frill as in preceding somite dorsally and laterally (Fig. 1 A, C). Genital double-somite (Figs 1 A, C, 2 A) rectangular, slightly longer than wide, with sensilla and minute surface spinules dorsolaterally as shown; anterior and posterior halves (second – genital – urosomite and third urosomite, respectively) distinct dorsally and laterally (Fig. 1 A, C), completely fused ventrally (Fig. 2 A); dorsolateral posterior margin of anterior half serrated and with two large spinules on each side (Fig. 1 A, C); posterior half of genital double-somite with dorsal (Fig. 1 A) and lateral (Figs 1 C, 2 A) large posterior spinules, posterior hyaline frill deeply incised and striated, the latter continuous dorsolaterally (Fig. 1 A, C) and interrupted ventrally (Fig. 2 A); P 6 at anterior third of genital somite (anterior half of genital double-somite) and with genital field as shown (Fig. 2 A). Fourth urosomite largely as in posterior half of genital double-somite, but with additional paired lateroventral pores and posterior medial transverse spinular row (Figs 1 A, C, 2 A). Fifth urosomite without sensilla, with one lateral row of small spinules (Fig. 1 C) and with paired lateroventral pores (Figs 1 C, 2 A), with short medial row of large spinules ventrally (Fig. 2 A), with continuous deeply incised posterior hyaline frill, the latter with medial part slightly crescentic dorsally (Fig. 1 A). Anal somite slightly wider than long in dorsal view (Fig. 1 A – B), without anal operculum, cleft medially dorsally (Fig. 1 A – B) and ventrally (Fig. 2 A), with two posterior sensilla dorsally (Fig. 1 A – D), with lateral and ventral pores as shown (Figs 1 C – D, 2 A), and with small spinules close to joint with caudal rami. Caudal rami well-developed (Figs 1 A – D, 2 A), about 1.7 times as long as wide and with straight outer and slightly convex inner margin (Fig. 1 A – B), with pores as shown; with seven setae (Fig. 1 B) as follows: setae I and II situated subdistally on outer margin, the former very small and ventral to the latter; seta III displaced ventrally close to outer margin; setae IV and V normal, with fracture plane, the former shorter; seta VI arising at inner distal corner; dorsal seta VII issuing subdistally, close to inner margin, triarticulated. ROSTRUM (Figs 1 A, C, 3 A – B). Not fused to cephalothorax, triangular, pointed, with one sensilla arising at distal third on each side, reaching distal margin of second antennulary segment. ANTENNULE (Fig. 3 A – B). Eight-segmented; all segments smooth; first and second segments with tube pore (arrowed in Fig. 3 A); all setae smooth except for two and three biarticulated setulose setae on seventh and eight segments, respectively; outer subdistal seta of eighth segment biarticulated basally, neighbouring seta swollen basally. Armature formula: 1 (1); 2 (11); 3 (7); 4 (3 + (1 + ae )), 5 (2); 6 (4); 7 (4); 8 (5 + acro). Acrothek consisting of two setae and one aesthetasc fused basally. ANTENNA (Fig. 3 C – D). With small unornamented coxa. Allobasis with remains of former division between basis and first endopodal segment, slightly longer than free endopodal segment, with outer slender spinules proximally, without abexopodal seta. Exopod three-segmented; first segment longer than second and third segments combined, second segment smallest; first and second segments unornamented, the former with one, the latter without seta; third segment with subapical row of spinules, armed with three setae, of which one bare, the other two bipinnate. Free endopodal segment with proximal and subdistal inner strong spinules as shown, and with two outer frills; armed with two lateral inner flagellate spines and two slender setae (the latter set close to each other), one inner apical flagellate spine, three single geniculate setae, one slender bipinnate short seta and one strongly spinulose geniculate outer seta fused basally to slender pinnate element. MANDIBLE (Fig. 4 A). With well-developed gnathobasis bearing strong multi- and bicuspidate teeth as shown, subdistally with two small slender setae and two spinules medially, and with one strongly spinulose element. Basis massive, with spinules as depicted, with three inner setae. Exopod twosegmented; first segment longer than second, the former with one lateral and one distal seta, the latter with one apical element. Endopod one-segmented, visibly larger that exopod; with two lateral and six distal setae. MAXILLULE (Fig. 4 B). With arthrite ornamented with some slender spinules and armed with two surface setae, seven distal spines as shown, one spinulose element and one recurved stiff pinnate seta. Coxal endite with three setae. Basis with spinular row as depicted and armed with two lateral, one subdistal and three distal elements, one of which visibly stronger. Rami one-segmented, exopod with two, endopod with four elements – two distal, one inner subdistal and one inner medial seta. MAXILLA (Fig. 5 A). With large syncoxa ornamented with small outer spinules; with three endites; proximal endite smallest, with three setae; middle and distal endites nearly subequal in length, the former with two, the latter with three setae. Basis drawn out into strong claw with minute pinnae, additionally with strong spine bearing one long spinule and two slender setae – one anterior, one posterior. Endopod two-segmented; first segment with two, second segment with one lateral and four distal setae. MAXILLIPED (Fig. 5 B). Subchelate. Praecoxa small, trapezoidal, ornamented with slender outer spinules. Coxa trapezoidal, with proximal, medial and subdistal spinules as shown; with two inner small setae issuing at distal fourth and two long apical setae. Basis longer than coxa, oval, with outer slender longitudinal spinules, and one posterior and one anterior row of spinules as depicted, with one medial inner and one subdistal inner seta. Endopod one-segmented, elongate, with three setae – one lateral, one subdistal and one small distal – and apical claw. P 1 (Fig. 6 A – B). With broad unornamented intercoxal sclerite (Fig. 6 B). Coxa massive, with several spinular rows as shown. Basis with inner and outer spinulose spines; with slender long inner spinules proximally and close to insertion of inner spine, and with shorter but stronger spinules at base of inner and outer seta, and distally on medial distal rounded protrusion. Exopod three-segmented, arising at a lower level than that of the endopod and reaching slightly beyond ENP 1; ENP: EXP length ratio 1.5; exopodal segments subequal in length, with strong outer spinules as shown; first segment without, second segment with one inner seta with comb tip; third segment with five elements. Endopod three-segmented, arising from pedestal; ENP 1 about four times as long as wide, as long as second and third segments combined, with outer and inner longitudinal rows of slender spinules, with one inner stiff, long, strongly pectinate seta; second and third segments ornamented with strong outer spinules as shown, the former shorter, nearly as long as wide and half as long as the latter, with one inner, slender, short seta; third segment elongate with one inner distal slender seta, one apical long element and one outer distal spine. P 2 (Fig. 6 B). Narrow intercoxal sclerite (not shown) with acute distal projection on each side, largely as in P 3. Praecoxa (not shown) as in P 3. Coxa with three anterior rows of spinules – one proximal and two medial transverse – and with posterior small spinules close to proximal outer margin. Basis with long slender spinules proximally close to inner margin, with small spinules between rami, and with comparatively larger spinules at base of exopod and at base of outer pinnate spine. Rami threesegmented, endopod and exopod subequal in absolute length. Exopod three-segmented, arising at a lower level than that of the endopod, not reaching tip of ENP; exopodal segments with strong outer spinules as shown; EXP 1 and EXP 2 with outer distal acute projection, on EXP 1 shorter, both segments with coarse inner distal hyaline frill and inner seta with comb tip, on EXP 1 visibly shorter; EXP 3 with six elements – three outer spines, two apical elements and one inner seta with comb tip. Endopod three-segmented, arising from pedestal; endopodal segments with strong outer spinules as shown; ENP 1 shortest, as long as wide, with small distal spinules close to small inner projection, without outer distal projection nor inner hyaline frill, with one short inner seta; ENP 2 elongate, 1.7 times as long as wide and 0.6 times as long as ENP 3, with strong outer spinules and with small spinules at base of outer distal acute projection, with coarse inner distal frill, with two inner setae of which proximal one short; ENP 3 longest, elongate, 4.8 times as long as wide, with medial outer pore, with four elements (one inner and two apical setae, and one outer spine), with acute projection between outer spine and inner apical seta. P 3 (Fig. 7 A). Narrow intercoxal sclerite with acute distal projection on each side. Praecoxa small, with transverse distal row of slender spinules close to outer margin. Coxa and basis as in P 2 except for outer setiform element of basis on P 3. Rami three-segmented, endopod and exopod subequal in absolute length. Exopod largely as in P 2 except for additional posterior pore on P 3 EXP 1. Endopod largely as in P 2 except for one inner seta only on P 3 ENP 2 and two inner setae on P 3 ENP 3. P 4 (Fig. 7 B). With intercoxal sclerite, praecoxa, coxa and basis as in P 3. Exopod longer than endopod; EXP: ENP length ratio 1.3; largely as in P 3 except for two inner setae on P 4 EXP 3 of which distalmost one visibly thicker and strongly pectinate. Endopod reaching slightly beyond middle of EXP 3; largely as in P 3 except for stiffer inner seta on P 4 ENP 2, and one inner seta only on P 4 ENP 3. Armature formula of P 1 – P 4 as follows: P 5 (Fig. 2 B). With outer seta of BENP arising from long setophore. Endopodal lobe triangular, reaching middle of exopod; with small spinules along outer margin and at base of inner setae; with five elements – one outer subdistal, one apical and one inner subdistal normal seta, and two inner bifurcate elements. Exopod elongate, 2.8 times as long as wide; with spinules along inner margin and with few proximal outer spinules; with six elements – three outer slender, short setae, two apical elements, of which outermost one shorter, and one inner seta. P 6 (Fig. 2 A). With three setae – one short bipinnate element, and one medial and one inner bare seta of which medial one shorter. Male GENERAL. Total body length measured from tip of rostrum to posterior margin of caudal rami ranging from 675 to 900 µm (mean 731 µm; n = 5; total body length of allotype 680 µm). PROSOME (Fig. 8 A). As in female. UROSOME (Fig. 8 A – B). As in female except for second – P 6 - bearing somite – and third urosomites separated, and for continuous spinular rows and hyaline frill of third and fourth urosomites ventrally. SEXUAL DIMORPHISM. Expressed in the ventral somatic ornamentation, antennule, basis of P 1, P 2 ENP, P 5 and P 6. ANTENNULE (Fig. 9 A – B). Ten-segmented, haplocer. All segments smooth; first and second segments with tube pore. All setae smooth except for two and three biarticulated setulose setae on ninth and tenth segments, respectively; outer subdistal seta of tenth segment biarticulated basally, neighbouring seta swollen basally. Armature formula: 1 (1); 2 (11); 3 (5); 4 (3); 5 (7 + (1 + ae )); 6 (1); 7 (1 + 2 spines); 8 (1 + 2 spines); 9 (4); 10 (5 + acro). Acrothek consisting of two setae and one aesthetasc fused basally. ANTENNA, MANDIBLE, MAXILLULE, MAXILLA AND MAXILLIPED (not shown). As in female. P 1. As in female except for basis without slender inner, long spinules, and with three inner accessory spines and a proximal inner, nose-like outgrowth in the male (Fig. 9 C). P 2 EXP (not shown). As in female. Endopod (Fig. 10 A) sexually dimorphic, two-segmented; ENP 1 0.7 times length of ENP 2, with sinuous inner margin, with proximal and medial outer slender spinules, with one short inner seta; ENP 2 oval, with longitudinal row of outer slender, long spinules, with two short inner setae arising from pedestal, subdistally with one posterior strong, blunt element, and with one anterior stiff, bare seta with subapical rounded expansion and pointed tip, with one lateral and one apical seta arising from elongate cylindrical extension fused to segment basally. P 3 AND P 4 (not shown). As in female. P 5 (Fig. 10 B). With outer seta of baseoendopod arising from long setophore. Endopodal lobe reaching middle of exopod, with inner and outer spinules as shown, with two distal bifurcate setae of which outermost one slightly shorter. Exopod rectangular, 2.5 times as long as wide, with longitudinal row of inner spinules and medial pore, with few proximal outer spinules, with six setae – three outer elements of which proximalmost visibly longer, one subdistal outer seta arising from setophore and two distal setae of which innermost one ⅓ length of outermost and with bifurcate tip. P 6 (Fig. 8 B). Asymmetrical, each leg with three setae of which innermost one shortest and spiniform, medial and outermost ones slender, medial one longest. Variability The left dorsal sensilla of the anal somite of the holotype is situated more proximally than the right sensilla. One female possesses five instead of six setae on one mandibular endopod and two instead of three maxillary endites on one maxilla.	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FFBB895AFD17324AFD1DAAC0.taxon	description	urn: lsid: zoobank. org: act: B 8 A 6 B 755 - 8375 - 48 B 0 - BC 31 - EBCCB 4 B 9 F 9 DE	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FFBB895AFD17324AFD1DAAC0.taxon	type_taxon	Type and only species Spinodiosaccus primus gen. et sp. nov., by monotypy.	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FFBB895AFD17324AFD1DAAC0.taxon	diagnosis	Diagnosis (based on Spinodiosaccus primus gen. et sp. nov.) Miraciidae: Diosaccinae. Body fusiform. Rostrum not fused to cephalothorax, elongate, almost as long as the first, second and third antennulary segments combined, with one subdistal sensilla on each side issuing at the third quarter of its length. Pro- and urosomites without extensions; with a flexible unsclerotized cuticle between prosome and urosome, the latter seemingly more evident in the male. Female genital double-somite (genital – second urosomite – and third urosomites fused) completely fused ventrally, with a dorsolateral cuticular rib marking former division between somites. Anal somite trapezoidal, without anal operculum. Caudal rami short, 1.6 times as wide as long in dorsal view, unornamented, with slender tube pore ventrally; with seven setae. Female antennule eight-segmented, with aesthetasc and seta fused basally on fourth segment and with apical acrothek on last segment. Male antennule haplocer, 11 - segmented, with aesthetasc and seta fused basally on fifth segment and with apical acrothek on last segment. Antennulary setae typically smooth. Antenna with allobasis; exopod three-segmented, armature formula 1 - 1 - 1,3,0. Mandibular palp biramous; endopod one-segmented, exopod two-segmented. Maxillule biramous; rami one-segmented. Maxilla with three endites; ENP one-segmented. Maxilliped subchelate; (syn) coxa with four, basis with two setae; ENP one-segmented, with claw and three accompanying setae. P 1 – P 4 with three-segmented rami. P 1 – P 2 EXP shorter than endopod, P 3 EXP and ENP of approximately the same length, P 4 ENP shorter than EXP. Male P 1 basis sexually dimorphic, with inner unipinnate spine and inner modified accessory spine. Male P 2 sexually dimorphic, two-segmented; first segment with one inner seta; second segment with two inner, one inner subdistal almost straight element, and two setae (one medial, one apical) issuing from elongate cylindrical extension fused to segment basally, and one strong outer subdistal spine, the latter strongly tapering distally. P 2 – P 4 EXP 1 without, EXP 2 with inner seta; P 2 – P 4 ENP 1 with inner seta; P 2 ENP 2 with two, P 3 – P 4 ENP 2 with one inner seta, inner distal seta of EXP 3 visibly shorter than other elements of same segment. Both P 5 EXP separated in the female, baseoendopods fused medially in the male; rami of P 5 distinct in both sexes; P 5 EXP with six setae in both sexes, whip-like in the female, two medial outer elements transformed into short spines in the male; P 5 endopodal lobe with five setae in the female, with two setae in the male.	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FFBB895AFD17324AFD1DAAC0.taxon	description	Armature formula of P 1 – P 5 as follows: Sexual dimorphism expressed in the male antennule, P 1 basis, P 2 ENP, P 5, P 6, and in the genital and third urosomite separated in males.	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FFBB895AFD17324AFD1DAAC0.taxon	etymology	Etymology The prefix ‘ spino’ from the Latin ‘ spīna’, ‘ spine’, refers to the two medial elements of the male P 5 EXP modified into strong short spines. Gender masculine.	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FFBB895AFD17324AFD1DAAC0.taxon	description	Apomorphies of Spinodiosaccus gen. nov. This new genus can easily be recognised by i) the male A 1 with 11 segments resulting from the reexpression of the plesiomorphic condition, ii) the absence of an inner seta on the P 1 EXP 2 and iii) the absence of inner armature on P 2 – P 4 EXP 1.	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FFB88965FDA63297FA9FA916.taxon	description	urn: lsid: zoobank. org: act: 4 B 5 EB 240 - AC 9 A- 4 F 68 - A 79 F-BADEF 42 F 6880 Figs 11 – 18	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FFB88965FDA63297FA9FA916.taxon	etymology	Etymology The specific epithet comes from the Latin ‘ prīmus’, ‘ first’, and refers to the first – and type – species described for Spinodiosaccus gen. nov. It is in the nominative singular, gender masculine.	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FFB88965FDA63297FA9FA916.taxon	materials_examined	Material examined Holotype MEXICO • ♀ (preserved in alcohol); Sinaloa State, Mazatlán, Urías coastal system, stn 5 (see also Gómez 2020 a: 43, fig. 1); 23.2056 ° N, 106.3715 ° W; 0.6 m depth; 18 Jan. 2019; S Gómez leg.; organic carbon content 0.99 %, organic matter content 1.71 %, sand 78.61 %, clay 6.72 %, silt 14.67 %); ICML- EMUCOP- 180119 - 57. Allotype MEXICO • ♂ (preserved in alcohol); same collection data as for holotype; ICML-EMUCOP- 180119 - 58. Paratypes MEXICO • 7 ♀♀, 5 ♂♂ (preserved in alcohol); same collection data as for holotype; ICML- EMUCOP- 180119 - 59 • 1 ♀ (preserved in alcohol); same collection data as for holotype; ICML- EMUCOP- 180119 - 60 • 2 ♀♀, 3 ♂♂ (preserved in alcohol); same collection data as for holotype; ICML- EMUCOP- 180119 - 61 • 1 ♀ (dissected); same collection data as for holotype; ICML-EMUCOP- 180119 - 64 • 1 ♀ (dissected); same collection data as for holotype; ICML-EMUCOP- 180119 - 65 • 1 ♀ (dissected); same collection data as for holotype; ICML-EMUCOP- 180119 - 66 • 1 ♂ (dissected); same collection data as for holotype; ICML-EMUCOP- 180119 - 67 • 1 ♂ (dissected); same collection data as for holotype; ICML-EMUCOP- 180119 - 68 • 1 ♂ (dissected); same collection data as for holotype; ICML- EMUCOP- 180119 - 69 • 1 ♀ (preserved in alcohol); Sinaloa State, Mazatlán, Urías coastal system, stn 6 (see also Gómez 2020 a: 43, fig. 1); 23.2123 ° N, 106.3780 ° W; 1.4 m depth; 18 Jan. 2019; S. Gómez leg.; organic carbon content 1.65 %, organic matter content 2.84 %, sand 61.44 %, clay 14.57 %, silt 23.99 %; ICML-EMUCOP- 180119 - 63. Other material MEXICO • 3 ♀♀, 6 ♂♂, 4 CV (preserved in alcohol); Sinaloa State, Mazatlán, Urías coastal system, stn 2 (see also Gómez 2020 a: 43, fig. 1); 23.1587 ° N, 106.3326 ° W; 1.8 m depth; 18 Jan. 2019; S. Gómez leg.; organic carbon content 3.99 %, organic matter content 6.86 %, sand 80.42 %, clay 8.29 %, silt 11.28 %; ICML-EMUCOP- 180119 - 70 • 1 ♀, 1 ♂ (preserved in alcohol); Sinaloa State, Mazatlán, Urías coastal system, stn 4 (see also Gómez 2020 a: 43, fig. 1); 23.1840 ° N, 106.3579 ° W; 0.7 m depth; 18 Jan. 2019; S. Gómez leg.; organic carbon content 1.13 %, organic matter content 1.94 %, sand 82.44 %, clay 8.27 %, silt 9.29 %; ICML-EMUCOP- 180119 - 71 • 1 ♀ (preserved in alcohol); same collection data as for preceding; ICML-EMUCOP- 180119 - 72 • 2 CV (preserved in alcohol); same collection data as for preceding; ICML-EMUCOP- 180119 - 73 • 1 CV (preserved in alcohol); same collection data as for holotype; ICML-EMUCOP- 180119 - 62 • 1 CV, 1 CIII (preserved in alcohol); Sinaloa State, Mazatlán, Urías coastal system, stn 6 (see also Gómez 2020 a: 43, fig. 1); 23.2123 ° N, 106.3780 ° W; 1.4 m depth; 18 Jan. 2019; S. Gómez leg.; organic carbon content 1.65 %, organic matter content 2.84 %, sand 61.44 %, clay 14.57 %, silt 23.99 %; ICML-EMUCOP- 180119 - 74 • 2 ♂♂ (preserved in alcohol); same collection data as for preceding; ICML-EMUCOP- 180119 - 75 • 2 ♀♀, 2 ♂♂ (preserved in alcohol); same collection data as for preceding; ICML-EMUCOP- 180119 - 76 • 1 ♀, 1 ♂ (preserved in alcohol); Sinaloa State, Mazatlán, Urías coastal system, stn 9 (see also Gómez 2020 a: 43, fig. 1); 23.1904 ° N, 106.4121 ° W; 5.4 m depth; 18 Jan. 2019; S. Gómez leg.; organic carbon content 1.41 %, organic matter content 2.43 %, sand 64.81 %, clay 8.09 %, silt 27.11 %; ICML-EMUCOP- 180119 - 77 • 4 ♂♂, 1 CV (preserved in alcohol); same collection data as for preceding; ICML-EMUCOP- 180119 - 78 • 3 ♂♂, 1 CV (preserved in alcohol); same collection data as for preceding; ICML-EMUCOP- 180119 - 79 • 1 ♂ (preserved in alcohol); Sinaloa State, Mazatlán, Urías coastal system, stn 10 (see also Gómez 2020 a: 43, fig. 1); 23.1815 ° N, 106.4214 ° W; 6.0 m depth; 18 Jan. 2019; S. Gómez leg.; organic carbon content 1.2 %, organic matter content 2.07 %, m sand 69.12 %, clay 7.91 %, silt 22.97 %; ICML-EMUCOP- 180119 - 80 • 1 ♂ (preserved in alcohol); same collection data as for preceding; ICML-EMUCOP- 180119 - 81.	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FFB88965FDA63297FA9FA916.taxon	description	Description Female GENERAL. Total body length measured from tip of rostrum to posterior margin of caudal rami ranging from 510 to 695 µm (mean 605.4 µm; n = 13; total body length of holotype 695 µm); habitus fusiform, widest at posterior end of cephalothorax in dorsal view, tapering posteriad (Fig. 11 A). PROSOME (Fig. 11 A, C). Consisting of cephalothorax, with fused first pedigerous somite, and second to fourth free pedigerous somites; prosomites without expansions nor spinular ornamentation; posterior hyaline frill of cephalothorax, second and third pedigerous somites broad, of P 4 - bearing somite visibly narrower, with posterior margins plain; posterior frill of second and third pedigerous somites with transverse row of minute spinules. UROSOME (Figs 11 A – D, 12 A). Consisting of fifth pedigerous somite (first urosomite), genital doublesomite (genital – second urosomite – and third urosomites fused), two free urosomites, anal somite and caudal rami; urosomites without expansions. First urosomite narrower than preceding somites in dorsal view (Fig. 11 A), without spinular ornamentation, with posterior sensilla as depicted, with one lateral pore on each side (Fig. 11 C), posterior frill as in preceding somite dorsally and laterally (Fig. 11 A, C). Genital double-somite (Figs 11 A, C, 12 A) rectangular, slightly longer than wide, with sensilla as shown, without spinular ornamentation; original division between anterior and posterior halves (second – genital – urosomite and third urosomite, respectively) marked by inner cuticular rib dorsally and laterally (Fig. 11 A, C), completely fused ventrally (Fig. 12 A); posterior half of genital double-somite with two dorsal pores and one lateroventral pore on each side, with posterior hyaline frill finely serrated and striated; P 6 at anterior third of genital somite (anterior half of genital double-somite) and with genital field as shown (Fig. 12 A). Fourth urosomite largely as in posterior half of genital double-somite, but with additional medial pore ventrally and posterior ventrolateral transverse spinular row on frill (Figs 11 A, C, 12 A). Fifth urosomite without sensilla or spinules, with two dorsal and two ventral pores, posterior hyaline frill as in preceding somites (Figs 1 A, C, 12 A). Anal somite wider than long in dorsal view (Fig. 11 A – B), without anal operculum, cleft medially dorsally (Fig. 11 A – B) and ventrally (Fig. 12 A), with two posterior sensilla dorsally, two dorsolateral pores (Fig. 11 A – C) and two ventral pores (Fig. 12 A), dorsally without spinules, with one ventrolateral and one short ventral spinular row on each side close to joint with caudal rami as shown (Fig. 12 A). Caudal rami short, twice as wide as long in dorsal view (Fig. 11 A – B), unornamented, with slender tube pore ventrally; with seven setae (Figs 1 B, D, 2 A) as follows: setae I and II situated almost medially on outer margin, the former very small and ventral to the latter; seta III displaced ventrally close to outer margin; setae IV and V normal, with fracture plane, the former shorter; seta VI arising at inner distal corner; dorsal seta VII issuing subdistally, close to inner margin, triarticulated. ROSTRUM (Fig. 13 A). Not fused to cephalothorax, elongate, triangular, with blunt tip, with one sensillum arising at distal third on each side, without pore, reaching distal margin of second antennulary segment. ANTENNULE (Fig. 13 B – C). Eight-segmented; all segments smooth, except for one proximal and one subdistal short spinular row on first segment; all setae smooth; with two and four biarticulated setae on penultimate and last segments, respectively. Armature formula: 1 (1); 2 (10); 3 (6); 4 (3 + (1 + ae )); 5 (2); 6 (4); 7 (4); 8 (5 + acro). Acrothek consisting of two setae and one aesthetasc fused basally. ANTENNA (Fig. 13 D). With small coxa ornamented with outer spinules. Allobasis with remains of former division between basis and first endopodal segment, as long as free endopodal segment, with outer spinules proximally, with one abexopodal seta. Exopod three-segmented; first and third ones subequal in length, second segment smallest; first and second segments unornamented, with one seta each; third segment unornamented, with four elements – one proximal bipinnate, one small subdistal and two distal spine-like elements. Free endopodal segment with proximal and subdistal inner strong spinules, and with row of outer small spinules as shown; armed with two lateral inner spines and two slender setae, the latter set close to each other, one inner apical spine, three single geniculate setae, one slender seta, and one strongly spinulose geniculate outer seta fused basally to slender pinnate element. MANDIBLE (Fig. 14 A). With well-developed gnathobasis bearing strong multi- and bicuspidate teeth as shown, with two long spinules and one pinnate seta. Basis massive, with strong spinules, with three inner setae. Exopod two-segmented; first segment longer than second, the former with one lateral and one distal seta, the latter with three apical elements. Endopod one-segmented, visibly larger than exopod; with two lateral and six distal setae, of which three fused basally. MAXILLULE (Fig. 14 B). With arthrite ornamented with some slender spinules and armed with two surface setae (the latter indicated with an asterisk on figure), seven distal spines as shown, one spinulose element and one recurved, stiff pinnate seta. Coxal endite with two setae. Basis with spinular rows as depicted and armed with three lateral, one subdistal and three distal elements, one of which visibly stronger. Rami one-segmented, exopod unornamented and with two setae, endopod with longitudinal spinular row and with four elements – one inner subdistal, two distal and two outer subdistal setae. MAXILLA (Fig. 14 C). With large syncoxa ornamented with small outer spinules; with three endites; proximal endite smallest, with two setae; middle endite longer than proximal endite, with two setae; distal endite longest, with three setae. Basis drawn out into strong claw, additionally with pinnate spine, and two slender setae – one anterior, one posterior. Endopod one-segmented, with one proximal, one medial, one subdistal and four apical setae. MAXILLIPED (Fig. 14 D). Subchelate. Praecoxa small. Coxa elongate, with proximal, medial and subdistal spinules as shown; with two inner small setae issuing at middle of segment and two long apical setae. Basis longer than coxa, elongate, with medial row of small spinules and with longitudinal row of minute outer spinules, with one medial inner and one subdistal inner seta. Endopod one-segmented, elongate, with apical claw and three setae of different lengths. P 1 (Fig. 15 A). With unornamented intercoxal sclerite. Coxa massive, with several spinular rows as shown. Basis with inner and outer spinulose spines; with strong spinules at top of crescentic medial extension, and with smaller spinules at base of inner spine. Exopod three-segmented, arising at a lower level than that of endopod and reaching tip of ENP 2; ENP: EXP length ratio 1.2; exopodal segments with strong outer spinules as shown; second segment shortest, third segment longest; first and second segments without inner armature, outer spine of first segment longer than that of second segment; third segment with five elements of which outer proximal and medial spines shorter than those of preceding segments. Endopod three-segmented, arising from pedestal; ENP 1 about 2.8 times as long as wide, 1.4 times as long as second and third segments combined, with outer longitudinal row of strong spinules, with one inner stiff, long, strongly pectinate seta; second and third segments with strong outer spinules as shown, the former shorter, slightly longer than wide, with one inner seta; third segment elongate, 1.5 times as long as preceding segment, with one inner distal slender seta, one apical long element and one outer distal spine. P 2 (Fig. 15 B). With narrow intercoxal sclerite unornamented, with acute distal projection on each side. Coxa with three anterior rows of spinules. Basis with small inner acute outgrowth, with long slender inner spinules, with minute spinules at base of endopod, and with strong spinules between rami and at base of outer spine. Rami three-segmented, endopod longer than exopod. Exopod three-segmented, arising at a lower level than that of the endopod, reaching slightly above the middle of ENP 3; exopodal segments with strong outer spinules as shown; EXP 1 and EXP 2 with outer distal acute projection, of EXP 1 shorter, inner distal frill of first segment coarse, of second segment small, first segment without, second segment with inner seta; EXP 3 with subdistal outer pore and seven elements – three outer spines, two apical elements and two inner setae of which distalmost one visibly shorter. Endopod three-segmented, arising from pedestal; endopodal segments with strong outer spinules as shown; ENP 1 shortest, slightly longer than wide, with posterior small spinules proximally, with small inner distal spinules and with outer short acute projection, with one short inner seta; ENP 2 elongate, twice as long as wide and 0.6 times as long as ENP 3, with small inner distal spinules, with long outer distal acute projection, with two inner setae of which proximal shorter; ENP 3 longest, elongate, six times as long as wide, with four elements (one inner and two apical setae, and one outer spine), with acute projection between outer spine and inner apical seta. P 3 (Fig. 16 A). With triangular praecoxa ornamented with transverse row of minute spinules. Intercoxal sclerite and coxa as in P 2. Basis largely as in P 2, but with smaller inner distal outgrowth and setiform outer element. Rami three-segmented, endopod slightly longer than exopod. Exopod largely as in P 2. Endopod largely as in P 2 except for lack of posterior spinules and with outer subdistal pore on ENP 1, for inner seta of P 3 ENP 1 comparatively longer, for one long seta only on P 3 ENP 2, and for two inner setae on P 3 ENP 3. P 4 (Fig. 16 B). With intercoxal sclerite, praecoxa, and coxa as in P 3. Basis as in P 3 except for lack of inner distal outgrowth. Exopod longer than endopod; EXP: ENP length ratio 1.3; largely as in P 3 except for three inner setae on P 4 EXP 3 of which medial one visibly thicker and strongly pectinate. Endopod reaching slightly beyond middle of EXP 3; largely as in P 3 except for one inner seta only on P 4 ENP 3. Armature formula of P 1 – P 4 as follows: P 5 (Fig. 12 C). With outer seta of baseoendopod arising from setophore. Endopodal lobe triangular, reaching distal third of exopod; with small spinules along outer margin; with five elements – one outer and one inner apical and one inner subdistal long setae, and two inner spiniform elements. Exopod elongate, 2.2 times as long as wide; with spinules at base of innermost seta and on outer margin proximally; with six elements – three outer slender setae, two apical elements of which outermost one shorter and one inner seta. P 6 (Fig. 12 A). With three setae – one short bipinnate element, and one medial and one inner slender seta. Male GENERAL. Total body length measured from tip of rostrum to posterior margin of caudal rami ranging from 380 to 510 µm (mean 455 µm; n = 8; total body length of allotype 510 µm). PROSOME (Fig. 17 A). As in female. UROSOME (Fig. 17 A – B). As in female except for second – P 6 - bearing somite – and third urosomites separated, for medial row of eight ventral spinules on third urosomite, and for lack of spinules on hyaline frill of fourth urosomite. SEXUAL DIMORPHISM. Expressed in ventral somatic ornamentation (see above), antennule, and basis of P 1, P 2 ENP, P 5 and P 6. ANTENNULE (Fig. 18 A – B). 11 - segmented, haplocer. All segments smooth, except for proximal and subdistal spinular rows on first segment. All setae smooth except for two and four biarticulated setae on tenth and eleventh segments, respectively. Armature formula: 1 (1); 2 (11); 3 (6); 4 (1); 5 (5 + (1 + ae )); 6 (2); 7 (2 + 2 spines); 8 (0); 9 (1); 10 (4); 11 (5 + acro). Acrothek consisting of two setae and one aesthetasc fused basally. ANTENNA, MANDIBLE, MAXILLULE, MAXILLA AND MAXILLIPED (not shown). As in female. P 1. As in female except for basis without slender inner long spinules, without spinules at base of inner spine and with inner modified accessory spine (Fig. 18 C). P 2 EXP (not shown). As in female. Endopod (Fig. 18 D) sexually dimorphic, two-segmented; ENP 1 as long as ENP 2 (excluding distal cylindrical extension of ENP 2), with longitudinal row of outer spinules, with outer distal acute outgrowth, with one inner short seta; proximal part of ENP 2 rectangular, distal extension cylindrical, 2.4 times as long as wide excluding distal cylindrical extension, with longitudinal row of outer spinules, with two inner setae arising from pedestal of which proximal (seta I on Fig. 18 D) visibly shorter, subdistally with one strong inner and almost straight element (element III on Fig. 18 D), with one lateral and one apical seta (setae IV and V on Fig. 18 D) arising from distal elongate cylindrical extension fused to segment basally and with strong outer subdistal element strongly tapering distally (element VI on Fig. 18 D). P 3 AND P 4 (not shown). As in female. P 5 (Fig. 18 E). With outer seta of baseoendopod arising from setophore. Endopodal lobe reaching middle of exopod, with outer spinules as shown, with two distal modified setae as shown of which outermost one shorter. Exopod triangular, 1.6 times as long as wide, with inner subdistal pore, with six setae – one proximal bipinnate seta, one medial and one subdistal short strong spines of which medial one shorter, one distal seta arising from setophore and two inner setae of which proximal one shorter. P 6 (Fig. 17 B). Asymmetrical, only one leg functional, each leg with three setae of which innermost one bipinnate, medial longest and bare, outermost shortest and bare. Variability Both caudal setae I spiniform in one female. One male possesses only one outer spine on P 5 EXP.	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FF848968FDFA324AFC91AA8D.taxon	type_taxon	Type species Canthocamptus imus Brady, 1872, by original designation.	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FF848968FDFA324AFC91AA8D.taxon	diagnosis	Amended diagnosis Miraciidae: Diosaccinae. Body fusiform. Rostrum not fused to cephalothorax, triangular, elongate, with rounded apical margin, almost as long as first, second and third antennulary segments combined, with one subdistal sensilla on each side at third quarter of its length. Pro- and urosomites without extensions; with a flexible unsclerotized cuticle between prosome and urosome. First urosomite (P 5 - bearing somite) narrower than preceding somites in dorsal view. Female genital double-somite (genital – second urosomite – and third urosomites fused) completely fused ventrally, with a dorsolateral cuticular rib marking former division between somites. Anal somite trapezoidal, without anal operculum. Caudal rami 1.6 times as wide as long in dorsal view, with seven setae. Female antennule eight-segmented, with aesthetasc and seta fused basally on fourth segment and with apical acrothek on last segment. Male antennule haplocer, 10 - segmented, with aesthetasc and seta fused basally on fifth segment, and with apical acrothek on last segment. Antennulary setae typically smooth. Antenna with allobasis and abexopodal seta; exopod threesegmented, armature formula 1 - 1 - 1,3,0. Mandibular palp biramous; endopod one-segmented, exopod two-segmented. Maxillule biramous; rami one-segmented. Maxilla with three endites, proximal and middle endites with two, distal endite with three elements; ENP one-segmented. Maxilliped subchelate; (syn) coxa with four, basis with two setae, ENP one-segmented with claw and three accompanying setae. P 1 – P 4 with three-segmented rami. Male P 1 basis sexually dimorphic, with inner unipinnate spine and inner modified accessory spine. P 1 EXP shorter than ENP, reaching tip of ENP 2; EXP 2 with inner seta; ENP 1 with inner seta; ENP 3 longer than the ENP 2, with one inner seta, a distal long and geniculate seta, and an outer spine. P 2 – P 4 EXP and ENP of approximately the same length; P 2 – P 4 EXP 1 with inner seta; inner distal seta of EXP 3 visibly shorter than other elements of same segment; P 2 ENP 2 with two setae, proximal shorter. Male P 2 ENP sexually dimorphic, two-segmented; first segment with one inner seta; second segment with two inner setae, one inner subdistal sigmoid element, one subdistal and one distal seta arising from cylindrical extension fused basally to segment, and one strong distal outer element strongly tapering distally. P 5 EXP with six setae in female and male; one or two medial outer elements transformed into short spines. P 5 endopodal lobe with five setae in female, with two setae in male; armature of male P 5 endopodal lobe strongly bipinnate.	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FF848968FDFA324AFC91AA8D.taxon	description	Armature formula of P 1 – P 5 as follows: Sexual dimorphism expressed in male antennule, P 2 ENP, P 5, P 6, and in genital and third urosomites being separated. Apomorphies for Bulbamphiascus We did not detect any apomorphy for Bulbamphiascus. However, Bulbamphiascus is part of a larger monophyletic group composed of Spinodiosaccus gen. nov., Spinopedia gen. nov. and Pallarica gen. nov. (Fig. 19). The subdistal outer element of the male P 2 ENP 2 with almost parallel margins, viz slightly tapering distally, is considered here as part of the ground pattern of this monophyletic group and is regarded as a potential synapomorphy for this clade. The modification of this spine into a strong tapering element is regarded here as secondary and is a synapomorphy for Bulbamphiascus and Spinodiosaccus gen. nov. The combination of an outer subdistal tapering spine and a sigmoid inner subdistal element on the male P 2 ENP 2 is unique to Bulbamphiascus (the outer subdistal spine on the male P 2 ENP 2 of Spinodiosaccus gen. nov. also tapers distally, but the inner subdistal element on the same segment of this species is rather straight). Bulbamphiascus chappuisi is here relegated to incertae sedis due to the shape of the rostrum and furca, and to the morphology of the male P 2 ENP, the illustrations of which (see Rouch 1962: 247, figs 29 – 33) are too schematic, preventing any further comparison.	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FF8A8969FD093160FAC3AF96.taxon	description	urn: lsid: zoobank. org: act: F 6 DD 1 CA 7 - 26 EE- 476 D-AC 00 - 2 C 4401 B 63014	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FF8A8969FD093160FAC3AF96.taxon	type_taxon	Type and only species Bulbamphiascus spinulosus Mu & Gee, 2000 (= Spinopedia spinulosa (Mu & Gee, 2000) comb. nov.), by monotypy.	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FF8A8969FD093160FAC3AF96.taxon	diagnosis	Diagnosis Miraciidae: Diosaccinae. Body fusiform; body surface seemingly covered with minute denticles. Rostrum not fused to cephalothorax, triangular, elongate, with round apical margin, almost as long as first and second antennulary segments combined, with one subdistal sensilla on each side at third quarter of its length. Pro- and urosomites without extensions; with a flexible unsclerotized cuticle between prosome and urosome. First urosomite (P 5 - bearing somite) narrower than preceding somites in dorsal view. Genital double-somite (genital – second urosomite – and third urosomites fused) completely fused ventrally, with a dorsolateral cuticular rib marking original division between genital and third urosomites. Telson trapezoidal, without anal operculum. Caudal rami short, about 1.5 times as wide as long in dorsal view, with seven setae. Female antennule eight-segmented, with aesthetasc and seta fused basally on fourth segment and with apical acrothek on last segment. Male antennule haplocer, 10 - segmented, with aesthetasc and seta fused basally on fifth segment and with apical acrothek on last segment. Setae of first and second antennulary segments typically bipinnate, smooth in remaining segments. Antenna with allobasis and abexopodal seta; exopod three-segmented, armature formula 1 - 1 - 1,3,0. Mandibular palp biramous; endopod one-segmented, larger than exopod, the latter two-segmented. Maxillule biramous; rami one-segmented. Maxilla with three endites, proximal and middle endites with two, distal endite with three elements; ENP one-segmented. Maxilliped subchelate; (syn) coxa with four, basis with two setae; ENP one-segmented, with claw and three accompanying setae. P 1 – P 4 with three-segmented rami. Male P 1 basis sexually dimorphic, with inner unipinnate spine and inner modified accessory spine. P 1 EXP shorter than ENP, reaching tip of ENP 2; P 1 ENP 1 with strong bipinnate inner seta; P 1 ENP 3 2.5 times as long as ENP 2, with one small inner seta, a distal long geniculate seta and one outer spine; P 1 EXP 2 with inner seta; P 2 – P 4 EXP and ENP of approximately the same length. P 2 – P 4 EXP 1 with inner seta; inner distal seta of P 2 – P 4 EXP 3 visibly smaller than other inner setae on same segment. P 2 ENP 2 with two setae, proximalmost shorter. P 2 – P 4 basis with inner chitinous projection. Male P 2 ENP sexually dimorphic, two-segmented; first segment with one inner seta; second segment completely covered by spinules and setules of different lengths, with two inner setae subequal in length, two elements (one medial, one apical) issuing from elongate cylindrical extension fused to segment basally, and with one inner subdistal sigmoid blunt element and one outer subdistal blunt spine with almost parallel margins, viz slightly tapering distally, of which the former is shorter. P 5 EXP longer than wide, comparatively shorter in male, with six setae in both sexes; two medial outer elements modified into stiff short spines in male; distal outer spine foliaceous in female. P 5 endopodal lobe with five setae in female, with two setae in male; armature of male P 5 endopodal lobe strongly bipinnate.	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FF8A8969FD093160FAC3AF96.taxon	description	Armature formula of P 1 – P 5 as follows: Sexual dimorphism expressed in male antennule, P 2 ENP, P 5, P 6, and in separated genital and third urosomites.	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FF8A8969FD093160FAC3AF96.taxon	etymology	Etymology The prefix ‘ spino’ from the Latin ‘ spīna’, ‘ spine’, and the sufix ‘ pedia’ from the Latin ‘ pēs’, ‘ pedis’, ‘ foot’, refers to the presence of a spinular patch on the male P 2 ENP 1, and to the presence of several spinules and setules covering the male P 2 ENP 2. Gender feminine.	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FF8A8969FD093160FAC3AF96.taxon	description	Apomorphies for Spinopedia gen. nov. Two apomorphies for Spinopedia have been detected: i) presence of an inner subdistal spinular patch on the male P 2 ENP 1 and ii) male P 2 ENP 2 completely covered with spinules and setules of different lengths. In addition, the subdistal inner seta of the female P 5 EXP arises directly from the ramus (i. e., the seta does not arise from a setophore) (character 42 in Table 3; see also Fig. 19). The lack of a setophore for the subdistal inner seta of the female P 5 EXP is regarded here as autapomorphic for the genus and is a result of reversal. The presence of a setophore for the inner subdistal seta of the female P 5 EXP is autapomorphic for a larger group of genera to which Spinopedia gen. nov. belongs (Fig. 19) and seems to have evolved convergently in more than one evolutionary lineage within the Diosaccinae (Fig. 19).	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FF8B896AFD3D364EFAF9AD5E.taxon	description	urn: lsid: zoobank. org: act: 78 F 45 B 32 - 1 B 47 - 41 BB- 99 E 0 - E 7 F 8327 E 3333	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FF8B896AFD3D364EFAF9AD5E.taxon	type_taxon	Type and only species Bulbamphiascus cibimae Pallares, 1982 (= Pallaricia cibimae (Pallares, 1982) comb. nov.), by monotypy.	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FF8B896AFD3D364EFAF9AD5E.taxon	diagnosis	Diagnosis Miraciidae: Diosaccinae. Body fusiform. Rostrum not fused to cephalothorax, triangular, elongate, with round apical margin, almost as long as first, second and third antennulary segments combined, with one subdistal sensilla at third quarter of its length, on each side. Pro- and urosomites without expansions. Genital double-somite (genital – second urosomite – and third urosomites fused) completely fused ventrally, with a dorsolateral cuticular rib marking original division between somites. Caudal rami about 1.5 times as wide as long in dorsal view; with seven setae. Female antennule eight-segmented, with aesthetasc and seta fused basally on fourth segment, and with apical acrothek on last segment. Antennulary setae typically smooth. Antenna with allobasis and abexopodal seta; exopod threesegmented, armature formula 1 - 1 - 1,3,0. Mandibular palp biramous; endopod one-segmented, larger than exopod, the latter two-segmented. Maxillule biramous; rami one-segmented. Maxilla with three endites, proximal and middle endites with two, distal endite with three elements; ENP two-segmented. Maxilliped subchelate; (syn) coxa with three, basis with two setae; ENP one-segmented, with claw and three accompanying setae. P 1 – P 4 with three-segmented rami. Male P 1 basis sexually dimorphic, with inner unipinnate spine and inner modified accessory spine. P 1 EXP shorter than ENP, reaching tip of ENP 2; P 1 ENP 1 with strong bipinnate inner seta; ENP 3 2.5 times as long as ENP 2, with one small inner seta, a long distal geniculate seta, and an outer spine; EXP 2 with inner seta. P 2 – P 4 EXP and ENP of approximately same lengths; P 2 – P 4 EXP 1 with inner seta; inner distal seta of P 2 – P 4 EXP 3 visibly shorter than other inner setae on same segment. P 2 ENP 2 with two setae, proximalmost small. Male P 2 ENP sexually dimorphic, two-segmented; first segment with one inner seta; second segment with two inner setae, two elements (one medial, one apical) issuing from elongate cylindrical extension fused to segment basally, and inner subdistal element almost straight and outer subdistal element slightly thicker and longer than the former and with almost parallel margins (i. e., slightly tapering distally). P 5 EXP longer than wide in female, comparatively shorter in male, with six setae in female and seven setae in male; two medial outer elements stiff short spine-like in male, slender and smooth in female. P 5 endopodal lobe with five setae in female, with two setae in male; armature of male P 5 endopodal lobe strongly bipinnate. Armature formula of P 1 – P 5 as follows: Sexual dimorphism expressed in male antennule, P 2 ENP, P 5, P 6, and in separated genital and third urosomites.	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FF8B896AFD3D364EFAF9AD5E.taxon	etymology	Etymology This genus is dedicated to Rosa E. Pallares for her contribution to the taxonomy of harpacticoid copepods from Argentina. Gender feminine.	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FF8B896AFD3D364EFAF9AD5E.taxon	description	Apomorphies for Pallarica gen. nov. This genus is unique by the possession of seven setae on the male P 5 EXP, which is regarded here as apomorphic for the genus. The presence of an additional seta is regarded here as a character reversal.	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FF88896CFD203506FBB9A8F8.taxon	description	urn: lsid: zoobank. org: act: 570211 A 6 - 5 E 4 D- 42 CA-A 26 D- 61 E 2 F 0 E 6 BC 6 A	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FF88896CFD203506FBB9A8F8.taxon	type_taxon	Type and only species	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FF88896CFD203506FBB9A8F8.taxon	diagnosis	Diagnosis Miraciidae: Diosaccinae. Body fusiform. Width: length ratio of caudal rami 0.8 – 1.2. Rostrum not fused to cephalothorax, elongate, with rounded tip. Female antennule eight-segmented, with aesthetasc and seta fused basally on fourth segment. Antennulary setae typically smooth. Antenna with allobasis and abexopodal seta; EXP three-segmented, armature formula 1 - 1 - 1,3,0. Mandibular palp extremely reduced. Maxillule biramous; rami one-segmented. Maxilla with three endites. Maxilliped subchelate; (syn) coxa with two, basis with two setae; ENP one-segmented, with claw and two accompanying setae. P 1 – P 4 with three-segmented rami. P 1 EXP shorter than ENP 1, reaching its distal third; P 1 ENP 1 with strong bipinnate inner seta; ENP 3 about twice as long as ENP 2, with one small inner seta, a long distal geniculate seta and an outer spine; P 1 EXP 2 with inner seta, EXP 3 with three outer spines, proximal one shortest, and two geniculate distal setae. P 2 – P 4 EXP 1 with inner seta; P 2 – P 4 EXP 3 with reduced inner armature. P 2 ENP 1 with inner setae; ENP 2 with two setae of approximately same length. Female P 5 EXP longer than wide, with six setae – three outer, two distal and one inner element. P 5 endopodal lobe with five setae.	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FF88896CFD203506FBB9A8F8.taxon	description	Armature formula of P 1 – P 5 as follows: Male unknown.	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FF88896CFD203506FBB9A8F8.taxon	etymology	Etymology This genus is dedicated to Alain Dinet for his contribution to the taxonomy of harpacticoid copepods from France. Gender feminine.	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FF88896CFD203506FBB9A8F8.taxon	description	Apomorphies for Dinetia gen. nov. The exclusion of Bulbamphiascus minutus from that genus and from any other genus presented here, and its reallocation into Dinetia gen. nov., is supported by, for example, the lack of the short inner distal seta on P 2 – P 4 EXP 3. Bulbamphiascus minutus probably bears a sister-group relationship with Pseudamphiascopsis Lang, 1944 and Rhyncholagena Lang, 1944, with which it forms a monophyletic group (see Fig. 19). These three genera share the longer P 1 ENP 1 relative to the P 1 EXP. Given that B. minutus could not be included in Bulbamphiascus or into any other diosaccin genus, and pending the discovery of the male of this species, we propose a new genus, Dinetia gen. nov., for B. minutus. The assessment and significance of character 43 for D. minuta comb. nov. (see Table 3) is still pending until the male is described. Phylogenetic statistics and resolution Our partial phylogenetic analysis resulted in 12 most parsimonious topologies with 126 steps, CI of 42, and RI of 77 (see Supplementary File 1). The strict consensus topology (nine branches collapsed) returned the following clades of unclear phylogenetic affinities between each other: a) a small clade composed by Dinetia gen. nov., Pseudamphiascopsis and Ryncholagena; b) a clade formed by Antiboreodiosaccus Lang, 1944, Diosaccopsis Brian, 1925, Goffinella Wilson, 1932, Ialysus Brian, 1927, Pseudodiosaccopsis Lang, 1944 and Pseudodiosaccus T. Scott, 1906; c) a clade formed by Typhlamphiascus Lang, 1944, Monardius Huys, 2009, Tydemanella A. Scott, 1909, Robertsonia Brady, 1880, Amphiascus Sars, 1905, Pararobertsonia Lang, 1944, Sinamphiascus Mu & Gee, 2000, Pallarica gen. nov., Spinopedia gen. nov., Bulbamphiascus and Spinodiosaccus gen. nov.; d) a clade composed of Amonardia Lang, 1944, Diosaccus Boeck, 1872, Pholenota Vervoort, 1964, Parialysus Nicholls, 1941, Amphiascoides Nicholls, 1941 b, Paramphiascella Lang, 1944, Robertgurneya Apostolov & Marinov, 1988, Miscegenus Wells, Hicks & Coull, 1982, Neomiscegenus Karanovic & Ranga Reddy, 2004, Paramphiascoides Wells, 1967, Schizopera Sars, 1905, Actopsyllus Wells, 1967, Balucopsylla Rao, 1972, Eoschizopera Wells & Rao, 1976, Haloschizopera Lang, 1944, Helmutkunzia Wells & Rao, 1976, Schizoperoides Por, 1968, Protopsammotopa Geddes, 1968 and Psammotopa Pennak, 1942. The phylogenetic position of the genera Dactylopodamphiascopsis Lang, 1944, Metamphiascopsis Lang, 1944 and Sarsamphiascus Huys, 2009 remains uncertain. It is important to highlight the high level of character convergence within the Diosaccinae. Clades a – d above are present in the 12 most parsimonious topologies (see Supplementary File 1), but their relationships are not consistent. The positions of the clade Dinetia gen. nov. – Rhyncholagena and the genera Sarsamphiascus, Metamphiascopsis and Dactylopodamphiscopsis are unclear, sometimes forming a clade with Typhlamphiascus – Spinodiosaccus gen. nov. (6 times), sometimes forming a clade with Amonardia – Psammotopa (6 times). The same happens to the clade Antiboreodiosaccus – Pseudodiosaccus which appears either as the sister group of the remaining Diosaccinae or as the sister group of a clade including Dinetia gen. nov. – Rhyncholagena, Sarsamphiascus, Metamphiascopsis, Dactylopodamphiscopsis and Typhlamphiascus – Spinodiosaccus gen. nov.	en	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
