identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
0B3187C8FFA5894BFDFD3433FBBEAFAE.text	0B3187C8FFA5894BFDFD3433FBBEAFAE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Typhlamphiascus Lang 1944	<div><p>Typhlamphiascus Lang, 1944</p> <p>Type species</p> <p>Amphiascus typhlops Sars, 1906, by original designation.</p> <p>Other species and subspecies</p> <p>T. blanchardi (T. Scott &amp; A. Scott, 1895) (originally described in Stenhelia)</p> <p>T. bouligandi Soyer, 1971</p> <p>T. brevicornis (I.C. Thompson &amp; A. Scott, 1903) (originally described in Stenhelia)</p> <p>T. capensis Kunz, 1975 comb. nov.</p> <p>T. confusus (T. Scott, 1902) (originally described in Stenhelia)</p> <p>T. confusus erythraeicus Por, 1963</p> <p>T. confusus gullmaricus Por, 1963</p> <p>T. dentipes (I.C. Thompson &amp; A. Scott, 1903) (originally described in Stenhelia)</p> <p>T. gracilicaudatus (I.C. Thompson &amp; A. Scott, 1903) (originally described in Stenhelia)</p> <p>T. gracilis Por, 1963</p> <p>T. higginsi Chullasorn, 2009</p> <p>T. lamellifer (Sars, 1911) (originally described in Amphiascus)</p> <p>T. latifurca Por, 1968</p> <p>T. medici sp. nov.</p> <p>T. ovale Wells &amp; Rao, 1987</p> <p>T. pectinifer Lang, 1965</p> <p>T. tuerkayi Ma &amp; Li, 2017</p> <p>T. unisetosus Lang, 1965</p> <p>Species incertae sedis</p> <p>Stenhelia hirsuta Thompson, 1893</p> <p>T. accraensis (T. Scott, 1894) (described in Stenhelia)</p> <p>T. drachi Soyer, 1963</p> <p>T. longifurcatus Rouch, 1962</p> <p>T. lutincola Soyer, 1963</p> <p>T. typhloides (Sars, 1911) (described in Amphiascus)</p> <p>Species inquirendae</p> <p>T. typhlops (Sars, 1906) sensu Por (1963)</p> <p>T. aff. confusus confusus (T. Scott, 1902) sensu Moore (1976)</p> <p>T. confusus confusus (T. Scott, 1902) sensu Marinov &amp; Apostolov (1985)</p> <p>Typhlamphiascus sp. in Vilela 1965</p> <p>Typhlamphiascus sp. ? in Bodin 1964</p> <p>Typhlamphiascus sp. ? in Bodin 1964</p> <p>Typhlamphiascus sp. I (? confusus) in Marinov 1977</p> <p>Typhlamphiascus sp. II in Marinov 1977</p> <p>Emended diagnosis</p> <p>Miraciidae: Diosaccinae. Body fusiform; length: width ratio of caudal rami from about 1.0 to about 3.0. Rostrum not fused to cephalothorax, triangular, elongate, as long as first and second antennulary segments combined, with pointed tip, with one subdistal sensilla on each side. Female genital and third urosomites distinct dorsally and/or laterally, completely fused ventrally forming genital double-somite; genital and third urosomites separated in males. Length:width ratio of caudal rami variable, from 1:1 to 3:1. Female antennule seven- to nine-segmented, with aesthetasc and seta fused basally on fourth segment, and with apical acrothek on last segment. Male antennule haplocer, eight–ten-segmented, with aesthetasc and seta fused basally on fifth segment, and with apical acrothek on last segment. Antennulary setae typically smooth, except for few slightly setulose setae on penultimate and last articles in females and males. Antenna with allobasis; exopod three-segmented, armature formula 1-0-1,3,0. Mandibular palp biramous; endopod one-segmented, larger than exopod; exopod two-segmented. Maxillule biramous; rami one-segmented. Maxilla with three endites; endopod two-segmented. Maxilliped subchelate; (syn)coxa with two to four, basis with two setae; endopod one-segmented, with claw and variable number of accompanying setae. P1–P4 with three-segmented rami. Male P1 basis sexually dimorphic, with normal or modified (elongate, blunt) inner spine, without or with variable number of inner accessory spines (two–eight). Male P2 sexually dimorphic, two-segmented; first segment with one inner seta, second segment with two inner setae, two subdistal elements (one of which thick and blunt) and two elements (one medial, one apical) issuing from elongate cylindrical extension fused to segment basally. P5 EXP typically longer than wide, with four to six setae in the female and male; P5 endopodal lobe with four to five setae in the female, with two setae in the male; two innermost elements of the female P5 endopodal lobe and armature of the male P5 endopodal lobe bifurcate distally.</p> <p>Armature formula of P1–P5 as follows:</p> <p>1 The report of eight setae on the female P5 EXP of T. blanchardi is most probably erroneous. 2 Typhlamphiascus typhlops is known to lack inner armature on the P4 ENP2. The inner seta of P4 ENP2 of T. capensis comb. nov. might have become detached from the supporting segment as indicated by the interrupted cuticle (Kunz 1975).</p> <p>Sexual dimorphism expressed in the male antennule (see above), P1 basis, P2 ENP (see above), P5 (see above), structure of P6, and in the separated genital and third urosomites.</p></div> 	http://treatment.plazi.org/id/0B3187C8FFA5894BFDFD3433FBBEAFAE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Gómez, Samuel;Corgosinho, Paulo Henrique Costa;Rivera-Sánchez, Karen I.	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FFA98957FDC73676FD21A90A.text	0B3187C8FFA98957FDC73676FD21A90A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Typhlamphiascus medici Gómez & Corgosinho & Rivera-Sánchez 2021	<div><p>Typhlamphiascus medici sp. nov.</p> <p>urn:lsid:zoobank.org:act: 67A13A9C-4D1E-4F86-82F2-CEE5407BCD26</p> <p>Figs 1–10</p> <p>Etymology</p> <p>The specific epithet from the Latin ‘medicī’, ‘doctor’, ‘physician’, is dedicated in honour and to the memory of all physicians and health personnel for their self-sacrifice during the fight against the COVID-19 pandemic. It is in the genitive singular, gender masculine.</p> <p>Material examined</p> <p>Holotype MEXICO • ♀ (preserved in alcohol); Sinaloa State, Mazatlán, Urías coastal system, stn 2 (see also Gómez 2020a: 43, fig. 1); 23.1587° N, 106.3326° W; 1.8 m depth; 18 Jan. 2019; S. Gómez leg.; organic carbon content 3.99%, organic matter content 6.86%, sand 80.42%, clay 8.29%, silt 11.28%; ICML- EMUCOP-180119-40.</p> <p>Allotype MEXICO • ♂ (preserved in alcohol); same collection data as for holotype; ICML-EMUCOP-180119-43. Other paratypes</p> <p>MEXICO • 3 ♀♀ (preserved in alcohol); same collection data as for holotype; ICML-EMUCOP-180119- 103 • 2 ♀♀, 2 ♂♂ (preserved in alcohol); same collection data as for holotype; ICML-EMUCOP-180119- 46 • 1 ♀ (dissected); same collection data as for holotype; ICML-EMUCOP-180119-41 • 1 ♀ (dissected); same collection data as for holotype; ICML-EMUCOP-180119-42 • 1 ♂ (dissected); same collection data as for holotype; ICML-EMUCOP-180119-44 • 1 ♂ (P1 and P2 dissected, rest intact and preserved in alcohol); same collection data as for holotype; ICML-EMUCOP-180119-45.</p> <p>Other material</p> <p>MEXICO • 8 CV, 4 CIV, 1 CII (preserved in alcohol); same collection data as for holotype; ICML- EMUCOP-180119-47 • 1 ♀, 2 ♂♂, 2 CV (preserved in alcohol); Sinaloa State, Mazatlán, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-106.3579&amp;materialsCitation.latitude=23.184" title="Search Plazi for locations around (long -106.3579/lat 23.184)">Urías</a> coastal system, stn 4 (see also Gómez 2020a: 43, fig. 1); 23.1840° N, 106.3579° W; 0.7 m depth; 18 Jan. 2019; S. Gómez leg.; organic carbon content 1.13%, organic matter content 1.94%, sand 82.44%, clay 8.27%, silt 9.29%; ICML-EMUCOP-180119-48 • 1 CV, 1 CIV, 1 CI (preserved in alcohol); Sinaloa State, Mazatlán, Urías coastal system, stn 5 (see also Gómez 2020a: 43, fig. 1); 23.2056° N, 106.3715° W; 0.6 m depth; 18 Jan. 2019; S. Gómez leg.; organic carbon content 0.99%, organic matter content 1.71%, sand 78.61%, clay 6.72%, silt 14.67%; ICML-EMUCOP-180119-49 • 1 ♂, 1 CIII (preserved in alcohol); same collection data as for preceding; ICML-EMUCOP-180119-50 • 1 ♀, 2 CIV, 1 CII (preserved in alcohol); same collection data as for preceding; ICML-EMUCOP-180119-51 • 1 CV, 1 CIII, 2 CII (preserved in alcohol); Sinaloa State, Mazatlán, Urías coastal system, stn 6 (see also Gómez 2020a: 43, fig. 1); 23.2123° N, 106.3780° W; 1.4 m depth; 18 Jan. 2019; S. Gómez leg.; organic carbon content 1.65%, organic matter content 2.84%, sand 61.44%, clay 14.57%, silt 23.99%; ICML- EMUCOP-180119-52 • 1 ♂, 1 CV, 1 CIV (preserved in alcohol); same collection data as for preceding; ICML-EMUCOP-180119-53 • 1 ♂ (preserved in alcohol); same collection data as for preceding; ICML- EMUCOP-180119-54) • 1 ♀, 1 CIV (preserved in alcohol); Sinaloa State, Mazatlán, Urías coastal system, stn 10 (see also Gómez 2020a: 43, fig. 1); 23.1815° N, 106.4214° W; 6.0 m depth; 18 Jan. 2019; S. Gómez leg.; organic carbon content 1.2%, organic matter content 2.07%, sand 69.12%, clay 7.91%, silt 22.97%; ICML-EMUCOP-180119-55 • 3 CIV (preserved in alcohol); same collection data as for preceding; ICML-EMUCOP-180119-56.</p> <p>Description</p> <p>Female</p> <p>GENERAL. Total body length measured from tip of rostrum to posterior margin of caudal rami ranging from 705 to 850 µm (mean 762 µm; n = 5; total body length of holotype 850 µm); habitus fusiform, widest at posterior end of cephalothorax in dorsal view, tapering posteriad (Fig. 1A).</p> <p>PROSOME (Figs 1A–B, 2A). Consisting of cephalothorax with fused first pedigerous somite and second to fourth free pedigerous somites; prosomites without expansions nor spinular ornamentation; posterior hyaline frill of cephalothorax, second and third pedigerous somites broad, with posterior margins finely serrated, striated, of fourth pedigerous somite visibly narrower, finely serrated, without striations.</p> <p>UROSOME (Figs 1A–D, 2A). Consisting of fifth pedigerous somite (first urosomite), genital doublesomite (genital – second urosomite – and third urosomites fused), two free urosomites, anal somite and caudal rami; urosomites without expansions. First urosomite narrower than preceding somites in dorsal view (Fig. 1A), without spinular ornamentation, with posterior sensilla as depicted, with one lateral pore on each side (Fig. 1C), posterior frill as in preceding somite dorsally and laterally (Fig. 1A, C). Genital double-somite (Figs 1A, C, 2A) rectangular, slightly longer than wide, with sensilla and minute surface spinules dorsolaterally as shown; anterior and posterior halves (second – genital – urosomite and third urosomite, respectively) distinct dorsally and laterally (Fig. 1A, C), completely fused ventrally (Fig. 2A); dorsolateral posterior margin of anterior half serrated and with two large spinules on each side (Fig. 1A, C); posterior half of genital double-somite with dorsal (Fig. 1A) and lateral (Figs 1C, 2A) large posterior spinules, posterior hyaline frill deeply incised and striated, the latter continuous dorsolaterally (Fig. 1A, C) and interrupted ventrally (Fig. 2A); P6 at anterior third of genital somite (anterior half of genital double-somite) and with genital field as shown (Fig. 2A). Fourth urosomite largely as in posterior half of genital double-somite, but with additional paired lateroventral pores and posterior medial transverse spinular row (Figs 1A, C, 2A). Fifth urosomite without sensilla, with one lateral row of small spinules (Fig. 1C) and with paired lateroventral pores (Figs 1C, 2A), with short medial row of large spinules ventrally (Fig. 2A), with continuous deeply incised posterior hyaline frill, the latter with medial part slightly crescentic dorsally (Fig. 1A). Anal somite slightly wider than long in dorsal view (Fig. 1A–B), without anal operculum, cleft medially dorsally (Fig. 1A–B) and ventrally (Fig. 2A), with two posterior sensilla dorsally (Fig. 1A–D), with lateral and ventral pores as shown (Figs 1C–D, 2A), and with small spinules close to joint with caudal rami. Caudal rami well-developed (Figs 1A–D, 2A), about 1.7 times as long as wide and with straight outer and slightly convex inner margin (Fig. 1A–B), with pores as shown; with seven setae (Fig. 1B) as follows: setae I and II situated subdistally on outer margin, the former very small and ventral to the latter; seta III displaced ventrally close to outer margin; setae IV and V normal, with fracture plane, the former shorter; seta VI arising at inner distal corner; dorsal seta VII issuing subdistally, close to inner margin, triarticulated.</p> <p>ROSTRUM (Figs 1A, C, 3A–B). Not fused to cephalothorax, triangular, pointed, with one sensilla arising at distal third on each side, reaching distal margin of second antennulary segment.</p> <p>ANTENNULE (Fig. 3A–B). Eight-segmented; all segments smooth; first and second segments with tube pore (arrowed in Fig. 3A); all setae smooth except for two and three biarticulated setulose setae on seventh and eight segments, respectively; outer subdistal seta of eighth segment biarticulated basally, neighbouring seta swollen basally. Armature formula: 1(1); 2(11); 3(7); 4(3+(1+ae)), 5(2); 6(4); 7(4); 8(5+acro). Acrothek consisting of two setae and one aesthetasc fused basally.</p> <p>ANTENNA (Fig. 3C–D). With small unornamented coxa. Allobasis with remains of former division between basis and first endopodal segment, slightly longer than free endopodal segment, with outer slender spinules proximally, without abexopodal seta. Exopod three-segmented; first segment longer than second and third segments combined, second segment smallest; first and second segments unornamented, the former with one, the latter without seta; third segment with subapical row of spinules, armed with three setae, of which one bare, the other two bipinnate. Free endopodal segment with proximal and subdistal inner strong spinules as shown, and with two outer frills; armed with two lateral inner flagellate spines and two slender setae (the latter set close to each other), one inner apical flagellate spine, three single geniculate setae, one slender bipinnate short seta and one strongly spinulose geniculate outer seta fused basally to slender pinnate element.</p> <p>MANDIBLE (Fig. 4A). With well-developed gnathobasis bearing strong multi- and bicuspidate teeth as shown, subdistally with two small slender setae and two spinules medially, and with one strongly spinulose element. Basis massive, with spinules as depicted, with three inner setae. Exopod twosegmented; first segment longer than second, the former with one lateral and one distal seta, the latter with one apical element. Endopod one-segmented, visibly larger that exopod; with two lateral and six distal setae.</p> <p>MAXILLULE (Fig. 4B). With arthrite ornamented with some slender spinules and armed with two surface setae, seven distal spines as shown, one spinulose element and one recurved stiff pinnate seta. Coxal endite with three setae. Basis with spinular row as depicted and armed with two lateral, one subdistal and three distal elements, one of which visibly stronger. Rami one-segmented, exopod with two, endopod with four elements – two distal, one inner subdistal and one inner medial seta.</p> <p>MAXILLA (Fig. 5A). With large syncoxa ornamented with small outer spinules; with three endites; proximal endite smallest, with three setae; middle and distal endites nearly subequal in length, the former with two, the latter with three setae. Basis drawn out into strong claw with minute pinnae, additionally</p> <p>with strong spine bearing one long spinule and two slender setae – one anterior, one posterior. Endopod two-segmented; first segment with two, second segment with one lateral and four distal setae.</p> <p>MAXILLIPED (Fig. 5B). Subchelate. Praecoxa small, trapezoidal, ornamented with slender outer spinules. Coxa trapezoidal, with proximal, medial and subdistal spinules as shown; with two inner small setae issuing at distal fourth and two long apical setae. Basis longer than coxa, oval, with outer slender longitudinal spinules, and one posterior and one anterior row of spinules as depicted, with one medial inner and one subdistal inner seta. Endopod one-segmented, elongate, with three setae – one lateral, one subdistal and one small distal – and apical claw.</p> <p>P1 (Fig. 6A–B). With broad unornamented intercoxal sclerite (Fig. 6B). Coxa massive, with several spinular rows as shown. Basis with inner and outer spinulose spines; with slender long inner spinules proximally and close to insertion of inner spine, and with shorter but stronger spinules at base of inner and outer seta, and distally on medial distal rounded protrusion. Exopod three-segmented, arising at a lower level than that of the endopod and reaching slightly beyond ENP1; ENP: EXP length ratio 1.5; exopodal segments subequal in length, with strong outer spinules as shown; first segment without, second segment with one inner seta with comb tip; third segment with five elements. Endopod three-segmented, arising from pedestal; ENP1 about four times as long as wide, as long as second and third segments combined, with outer and inner longitudinal rows of slender spinules, with one inner stiff, long, strongly pectinate seta; second and third segments ornamented with strong outer spinules as shown, the former shorter, nearly as long as wide and half as long as the latter, with one inner, slender, short seta; third segment elongate with one inner distal slender seta, one apical long element and one outer distal spine.</p> <p>P2 (Fig. 6B). Narrow intercoxal sclerite (not shown) with acute distal projection on each side, largely as in P3. Praecoxa (not shown) as in P3. Coxa with three anterior rows of spinules – one proximal and two medial transverse – and with posterior small spinules close to proximal outer margin. Basis with long slender spinules proximally close to inner margin, with small spinules between rami, and with comparatively larger spinules at base of exopod and at base of outer pinnate spine. Rami threesegmented, endopod and exopod subequal in absolute length. Exopod three-segmented, arising at a lower level than that of the endopod, not reaching tip of ENP; exopodal segments with strong outer spinules as shown; EXP1 and EXP2 with outer distal acute projection, on EXP1 shorter, both segments with coarse inner distal hyaline frill and inner seta with comb tip, on EXP1 visibly shorter; EXP3 with six elements – three outer spines, two apical elements and one inner seta with comb tip. Endopod three-segmented, arising from pedestal; endopodal segments with strong outer spinules as shown; ENP1 shortest, as long as wide, with small distal spinules close to small inner projection, without outer distal projection nor inner hyaline frill, with one short inner seta; ENP2 elongate, 1.7 times as long as wide and 0.6 times as long as ENP3, with strong outer spinules and with small spinules at base of outer distal acute projection, with coarse inner distal frill, with two inner setae of which proximal one short; ENP3 longest, elongate, 4.8 times as long as wide, with medial outer pore, with four elements (one inner and two apical setae, and one outer spine), with acute projection between outer spine and inner apical seta.</p> <p>P3 (Fig. 7A). Narrow intercoxal sclerite with acute distal projection on each side. Praecoxa small, with transverse distal row of slender spinules close to outer margin. Coxa and basis as in P2 except for outer setiform element of basis on P3. Rami three-segmented, endopod and exopod subequal in absolute length. Exopod largely as in P2 except for additional posterior pore on P3 EXP1. Endopod largely as in P2 except for one inner seta only on P3 ENP2 and two inner setae on P3 ENP3.</p> <p>P4 (Fig. 7B). With intercoxal sclerite, praecoxa, coxa and basis as in P3. Exopod longer than endopod; EXP: ENP length ratio 1.3; largely as in P3 except for two inner setae on P4 EXP3 of which distalmost one visibly thicker and strongly pectinate. Endopod reaching slightly beyond middle of EXP3; largely as in P3 except for stiffer inner seta on P4 ENP2, and one inner seta only on P4 ENP3.</p> <p>Armature formula of P1–P4 as follows: P5 (Fig. 2B). With outer seta of BENP arising from long setophore. Endopodal lobe triangular, reaching middle of exopod; with small spinules along outer margin and at base of inner setae; with five elements – one outer subdistal, one apical and one inner subdistal normal seta, and two inner bifurcate elements. Exopod elongate, 2.8 times as long as wide; with spinules along inner margin and with few proximal outer spinules; with six elements – three outer slender, short setae, two apical elements, of which outermost one shorter, and one inner seta.</p> <p>P6 (Fig. 2A). With three setae – one short bipinnate element, and one medial and one inner bare seta of which medial one shorter.</p> <p>Male</p> <p>GENERAL. Total body length measured from tip of rostrum to posterior margin of caudal rami ranging from 675 to 900 µm (mean 731 µm; n = 5; total body length of allotype 680 µm).</p> <p>PROSOME (Fig. 8A). As in female.</p> <p>UROSOME (Fig. 8A–B). As in female except for second – P6-bearing somite – and third urosomites separated, and for continuous spinular rows and hyaline frill of third and fourth urosomites ventrally.</p> <p>SEXUAL DIMORPHISM. Expressed in the ventral somatic ornamentation, antennule, basis of P1, P2 ENP, P5 and P6.</p> <p>ANTENNULE (Fig. 9A–B). Ten-segmented, haplocer. All segments smooth; first and second segments with tube pore. All setae smooth except for two and three biarticulated setulose setae on ninth and tenth segments, respectively; outer subdistal seta of tenth segment biarticulated basally, neighbouring seta swollen basally. Armature formula: 1(1); 2(11); 3(5); 4(3); 5(7 +(1+ae)); 6(1); 7(1 +2 spines); 8(1 +2 spines); 9(4); 10(5+ acro). Acrothek consisting of two setae and one aesthetasc fused basally.</p> <p>ANTENNA, MANDIBLE, MAXILLULE, MAXILLA AND MAXILLIPED (not shown). As in female.</p> <p>P1. As in female except for basis without slender inner, long spinules, and with three inner accessory spines and a proximal inner, nose-like outgrowth in the male (Fig. 9C).</p> <p>P2 EXP (not shown). As in female. Endopod (Fig. 10A) sexually dimorphic, two-segmented; ENP1 0.7 times length of ENP2, with sinuous inner margin, with proximal and medial outer slender spinules, with one short inner seta; ENP2 oval, with longitudinal row of outer slender, long spinules, with two short inner setae arising from pedestal, subdistally with one posterior strong, blunt element, and with one anterior stiff, bare seta with subapical rounded expansion and pointed tip, with one lateral and one apical seta arising from elongate cylindrical extension fused to segment basally.</p> <p>P3 AND P4 (not shown). As in female.</p> <p>P5 (Fig. 10B). With outer seta of baseoendopod arising from long setophore. Endopodal lobe reaching middle of exopod, with inner and outer spinules as shown, with two distal bifurcate setae of which outermost one slightly shorter. Exopod rectangular, 2.5 times as long as wide, with longitudinal row of inner spinules and medial pore, with few proximal outer spinules, with six setae – three outer elements of which proximalmost visibly longer, one subdistal outer seta arising from setophore and two distal setae of which innermost one ⅓ length of outermost and with bifurcate tip.</p> <p>P6 (Fig. 8B). Asymmetrical, each leg with three setae of which innermost one shortest and spiniform, medial and outermost ones slender, medial one longest.</p> <p>Variability</p> <p>The left dorsal sensilla of the anal somite of the holotype is situated more proximally than the right sensilla. One female possesses five instead of six setae on one mandibular endopod and two instead of three maxillary endites on one maxilla.</p></div> 	http://treatment.plazi.org/id/0B3187C8FFA98957FDC73676FD21A90A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Gómez, Samuel;Corgosinho, Paulo Henrique Costa;Rivera-Sánchez, Karen I.	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FFBB895AFD17324AFD1DAAC0.text	0B3187C8FFBB895AFD17324AFD1DAAC0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spinodiosaccus Gómez & Corgosinho & Rivera-Sánchez 2021	<div><p>Spinodiosaccus gen. nov.</p> <p>urn:lsid:zoobank.org:act: B8A6B755-8375-48B0-BC31-EBCCB4B9F9DE</p> <p>Type and only species</p> <p>Spinodiosaccus primus gen. et sp. nov., by monotypy.</p> <p>Diagnosis (based on Spinodiosaccus primus gen. et sp. nov.)</p> <p>Miraciidae: Diosaccinae. Body fusiform. Rostrum not fused to cephalothorax, elongate, almost as long as the first, second and third antennulary segments combined, with one subdistal sensilla on each side issuing at the third quarter of its length. Pro- and urosomites without extensions; with a flexible unsclerotized cuticle between prosome and urosome, the latter seemingly more evident in the male. Female genital double-somite (genital – second urosomite – and third urosomites fused) completely fused ventrally, with a dorsolateral cuticular rib marking former division between somites. Anal somite trapezoidal, without anal operculum. Caudal rami short, 1.6 times as wide as long in dorsal view, unornamented, with slender tube pore ventrally; with seven setae. Female antennule eight-segmented, with aesthetasc and seta fused basally on fourth segment and with apical acrothek on last segment. Male antennule haplocer, 11-segmented, with aesthetasc and seta fused basally on fifth segment and with apical acrothek on last segment. Antennulary setae typically smooth. Antenna with allobasis; exopod three-segmented, armature formula 1-1-1,3,0. Mandibular palp biramous; endopod one-segmented, exopod two-segmented. Maxillule biramous; rami one-segmented. Maxilla with three endites; ENP one-segmented. Maxilliped subchelate; (syn)coxa with four, basis with two setae; ENP one-segmented, with claw and three accompanying setae. P1–P4 with three-segmented rami. P1–P2 EXP shorter than endopod, P3 EXP and ENP of approximately the same length, P4 ENP shorter than EXP. Male P1 basis sexually dimorphic, with inner unipinnate spine and inner modified accessory spine. Male P2 sexually dimorphic, two-segmented; first segment with one inner seta; second segment with two inner, one inner subdistal almost straight element, and two setae (one medial, one apical) issuing from elongate cylindrical extension fused to segment basally, and one strong outer subdistal spine, the latter strongly tapering distally. P2–P4 EXP1 without, EXP2 with inner seta; P2–P4 ENP1 with inner seta; P2 ENP2 with two, P3–P4 ENP2 with one inner seta, inner distal seta of EXP3 visibly shorter than other elements of same segment. Both P5 EXP separated in the female, baseoendopods fused medially in the male; rami of P5 distinct in both sexes; P5 EXP with six setae in both sexes, whip-like in the female, two medial outer elements transformed into short spines in the male; P5 endopodal lobe with five setae in the female, with two setae in the male.</p> <p>Armature formula of P1–P5 as follows:</p> <p>Sexual dimorphism expressed in the male antennule, P1 basis, P2 ENP, P5, P6, and in the genital and third urosomite separated in males.</p> <p>Etymology</p> <p>The prefix ‘spino’ from the Latin ‘spīna’, ‘spine’, refers to the two medial elements of the male P5 EXP modified into strong short spines. Gender masculine.</p> <p>Apomorphies of Spinodiosaccus gen. nov.</p> <p>This new genus can easily be recognised by i) the male A1 with 11 segments resulting from the reexpression of the plesiomorphic condition, ii) the absence of an inner seta on the P1 EXP2 and iii) the absence of inner armature on P2–P4 EXP1.</p></div> 	http://treatment.plazi.org/id/0B3187C8FFBB895AFD17324AFD1DAAC0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Gómez, Samuel;Corgosinho, Paulo Henrique Costa;Rivera-Sánchez, Karen I.	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FFB88965FDA63297FA9FA916.text	0B3187C8FFB88965FDA63297FA9FA916.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spinodiosaccus primus Gómez & Corgosinho & Rivera-Sánchez 2021	<div><p>Spinodiosaccus primus gen. et sp. nov.</p> <p>urn:lsid:zoobank.org:act: 4B5EB240-AC9A-4F68-A79F-BADEF42F6880</p> <p>Figs 11–18</p> <p>Etymology</p> <p>The specific epithet comes from the Latin ‘prīmus’, ‘first’, and refers to the first – and type – species described for Spinodiosaccus gen. nov. It is in the nominative singular, gender masculine.</p> <p>Material examined</p> <p>Holotype MEXICO • ♀ (preserved in alcohol); Sinaloa State, Mazatlán, Urías coastal system, stn 5 (see also Gómez 2020a: 43, fig. 1); 23.2056° N, 106.3715° W; 0.6 m depth; 18 Jan. 2019; S Gómez leg.; organic carbon content 0.99%, organic matter content 1.71%, sand 78.61%, clay 6.72%, silt 14.67%); ICML- EMUCOP-180119-57.</p> <p>Allotype MEXICO • ♂ (preserved in alcohol); same collection data as for holotype; ICML-EMUCOP-180119-58.</p> <p>Paratypes MEXICO • 7 ♀♀, 5 ♂♂ (preserved in alcohol); same collection data as for holotype; ICML- EMUCOP-180119-59 • 1 ♀ (preserved in alcohol); same collection data as for holotype; ICML- EMUCOP-180119-60 • 2 ♀♀, 3 ♂♂ (preserved in alcohol); same collection data as for holotype; ICML- EMUCOP-180119-61 • 1 ♀ (dissected); same collection data as for holotype; ICML-EMUCOP-180119- 64 • 1 ♀ (dissected); same collection data as for holotype; ICML-EMUCOP-180119-65 • 1 ♀ (dissected); same collection data as for holotype; ICML-EMUCOP-180119-66 • 1 ♂ (dissected); same collection data as for holotype; ICML-EMUCOP-180119-67 • 1 ♂ (dissected); same collection data as for holotype; ICML-EMUCOP-180119-68 • 1 ♂ (dissected); same collection data as for holotype; ICML- EMUCOP-180119-69 • 1 ♀ (preserved in alcohol); Sinaloa State, Mazatlán, Urías coastal system, stn 6 (see also Gómez 2020a: 43, fig. 1); 23.2123° N, 106.3780° W; 1.4 m depth; 18 Jan. 2019; S. Gómez leg.; organic carbon content 1.65%, organic matter content 2.84%, sand 61.44%, clay 14.57%, silt 23.99%; ICML-EMUCOP-180119-63.</p> <p>Other material</p> <p>MEXICO • 3 ♀♀, 6 ♂♂, 4 CV (preserved in alcohol); Sinaloa State, Mazatlán, Urías coastal system, stn 2 (see also Gómez 2020a: 43, fig. 1); 23.1587° N, 106.3326° W; 1.8 m depth; 18 Jan. 2019; S. Gómez leg.; organic carbon content 3.99%, organic matter content 6.86%, sand 80.42%, clay 8.29%, silt 11.28%; ICML-EMUCOP-180119-70 • 1 ♀, 1 ♂ (preserved in alcohol); Sinaloa State, Mazatlán, Urías coastal system, stn 4 (see also Gómez 2020a: 43, fig. 1); 23.1840° N, 106.3579° W; 0.7 m depth; 18 Jan. 2019; S. Gómez leg.; organic carbon content 1.13%, organic matter content 1.94%, sand 82.44%, clay 8.27%, silt 9.29%; ICML-EMUCOP-180119-71 • 1 ♀ (preserved in alcohol); same collection data as for preceding; ICML-EMUCOP-180119-72 • 2 CV (preserved in alcohol); same collection data as for preceding; ICML-EMUCOP-180119-73 • 1 CV (preserved in alcohol); same collection data as for holotype; ICML-EMUCOP-180119-62 • 1 CV, 1 CIII (preserved in alcohol); Sinaloa State, Mazatlán, Urías coastal system, stn 6 (see also Gómez 2020a: 43, fig. 1); 23.2123° N, 106.3780° W; 1.4 m depth; 18 Jan. 2019; S. Gómez leg.; organic carbon content 1.65%, organic matter content 2.84%, sand 61.44%, clay 14.57%, silt 23.99%; ICML-EMUCOP-180119-74 • 2 ♂♂ (preserved in alcohol); same collection data as for preceding; ICML-EMUCOP-180119-75 • 2 ♀♀, 2 ♂♂ (preserved in alcohol); same collection data as for preceding; ICML-EMUCOP-180119-76 • 1 ♀, 1 ♂ (preserved in alcohol); Sinaloa State, Mazatlán, Urías coastal system, stn 9 (see also Gómez 2020a: 43, fig. 1); 23.1904° N, 106.4121° W; 5.4 m depth; 18 Jan. 2019; S. Gómez leg.; organic carbon content 1.41%, organic matter content 2.43%, sand 64.81%, clay 8.09%, silt 27.11%; ICML-EMUCOP-180119-77 • 4 ♂♂, 1 CV (preserved in alcohol); same collection data as for preceding; ICML-EMUCOP-180119-78 • 3 ♂♂, 1 CV (preserved in alcohol); same collection data as for preceding; ICML-EMUCOP-180119-79 • 1 ♂ (preserved in alcohol); Sinaloa State, Mazatlán, Urías coastal system, stn 10 (see also Gómez 2020a: 43, fig. 1); 23.1815° N, 106.4214° W; 6.0 m depth; 18 Jan. 2019; S. Gómez leg.; organic carbon content 1.2%, organic matter content 2.07%,m sand 69.12%, clay 7.91%, silt 22.97%; ICML-EMUCOP-180119-80 • 1 ♂ (preserved in alcohol); same collection data as for preceding; ICML-EMUCOP-180119-81.</p> <p>Description</p> <p>Female</p> <p>GENERAL. Total body length measured from tip of rostrum to posterior margin of caudal rami ranging from 510 to 695 µm (mean 605.4 µm; n = 13; total body length of holotype 695 µm); habitus fusiform, widest at posterior end of cephalothorax in dorsal view, tapering posteriad (Fig. 11A).</p> <p>PROSOME (Fig. 11A, C). Consisting of cephalothorax, with fused first pedigerous somite, and second to fourth free pedigerous somites; prosomites without expansions nor spinular ornamentation; posterior hyaline frill of cephalothorax, second and third pedigerous somites broad, of P4-bearing somite visibly narrower, with posterior margins plain; posterior frill of second and third pedigerous somites with transverse row of minute spinules.</p> <p>UROSOME (Figs 11A–D, 12A). Consisting of fifth pedigerous somite (first urosomite), genital doublesomite (genital – second urosomite – and third urosomites fused), two free urosomites, anal somite and caudal rami; urosomites without expansions. First urosomite narrower than preceding somites in dorsal view (Fig. 11A), without spinular ornamentation, with posterior sensilla as depicted, with one lateral pore on each side (Fig. 11C), posterior frill as in preceding somite dorsally and laterally (Fig. 11A, C). Genital double-somite (Figs 11A, C, 12A) rectangular, slightly longer than wide, with sensilla as shown, without spinular ornamentation; original division between anterior and posterior halves (second – genital – urosomite and third urosomite, respectively) marked by inner cuticular rib dorsally and laterally (Fig. 11A, C), completely fused ventrally (Fig. 12A); posterior half of genital double-somite with two dorsal pores and one lateroventral pore on each side, with posterior hyaline frill finely serrated and striated; P6 at anterior third of genital somite (anterior half of genital double-somite) and with genital field as shown (Fig. 12A). Fourth urosomite largely as in posterior half of genital double-somite, but with additional medial pore ventrally and posterior ventrolateral transverse spinular row on frill (Figs 11A, C, 12A). Fifth urosomite without sensilla or spinules, with two dorsal and two ventral pores, posterior hyaline frill as in preceding somites (Figs 1A, C, 12A). Anal somite wider than long in dorsal view (Fig. 11A–B), without anal operculum, cleft medially dorsally (Fig. 11A–B) and ventrally (Fig. 12A), with two posterior sensilla dorsally, two dorsolateral pores (Fig. 11A–C) and two ventral pores (Fig. 12A), dorsally without spinules, with one ventrolateral and one short ventral spinular row on each side close to joint with caudal rami as shown (Fig. 12A). Caudal rami short, twice as wide as long in dorsal view (Fig. 11A–B), unornamented, with slender tube pore ventrally; with seven setae (Figs 1B, D, 2A) as follows: setae I and II situated almost medially on outer margin, the former very small and ventral to the latter; seta III displaced ventrally close to outer margin; setae IV and V normal, with fracture plane, the former shorter; seta VI arising at inner distal corner; dorsal seta VII issuing subdistally, close to inner margin, triarticulated.</p> <p>ROSTRUM (Fig. 13A). Not fused to cephalothorax, elongate, triangular, with blunt tip, with one sensillum arising at distal third on each side, without pore, reaching distal margin of second antennulary segment.</p> <p>ANTENNULE (Fig. 13B–C). Eight-segmented; all segments smooth, except for one proximal and one subdistal short spinular row on first segment; all setae smooth; with two and four biarticulated setae on penultimate and last segments, respectively. Armature formula: 1(1); 2(10); 3(6); 4(3 +(1 +ae)); 5(2); 6(4); 7(4); 8(5+ acro). Acrothek consisting of two setae and one aesthetasc fused basally.</p> <p>ANTENNA (Fig. 13D). With small coxa ornamented with outer spinules. Allobasis with remains of former division between basis and first endopodal segment, as long as free endopodal segment, with outer spinules proximally, with one abexopodal seta. Exopod three-segmented; first and third ones subequal in length, second segment smallest; first and second segments unornamented, with one seta each; third segment unornamented, with four elements – one proximal bipinnate, one small subdistal and two distal spine-like elements. Free endopodal segment with proximal and subdistal inner strong spinules, and with row of outer small spinules as shown; armed with two lateral inner spines and two slender setae, the latter set close to each other, one inner apical spine, three single geniculate setae, one slender seta, and one strongly spinulose geniculate outer seta fused basally to slender pinnate element.</p> <p>MANDIBLE (Fig. 14A). With well-developed gnathobasis bearing strong multi- and bicuspidate teeth as shown, with two long spinules and one pinnate seta. Basis massive, with strong spinules, with three inner setae. Exopod two-segmented; first segment longer than second, the former with one lateral and one distal seta, the latter with three apical elements. Endopod one-segmented, visibly larger than exopod; with two lateral and six distal setae, of which three fused basally.</p> <p>MAXILLULE (Fig. 14B). With arthrite ornamented with some slender spinules and armed with two surface setae (the latter indicated with an asterisk on figure), seven distal spines as shown, one spinulose element and one recurved, stiff pinnate seta. Coxal endite with two setae. Basis with spinular rows as depicted and armed with three lateral, one subdistal and three distal elements, one of which visibly stronger. Rami one-segmented, exopod unornamented and with two setae, endopod with longitudinal spinular row and with four elements – one inner subdistal, two distal and two outer subdistal setae.</p> <p>MAXILLA (Fig. 14C). With large syncoxa ornamented with small outer spinules; with three endites; proximal endite smallest, with two setae; middle endite longer than proximal endite, with two setae; distal endite longest, with three setae. Basis drawn out into strong claw, additionally with pinnate spine, and two slender setae – one anterior, one posterior. Endopod one-segmented, with one proximal, one medial, one subdistal and four apical setae.</p> <p>MAXILLIPED (Fig. 14D). Subchelate. Praecoxa small. Coxa elongate, with proximal, medial and subdistal spinules as shown; with two inner small setae issuing at middle of segment and two long apical setae. Basis longer than coxa, elongate, with medial row of small spinules and with longitudinal row of minute outer spinules, with one medial inner and one subdistal inner seta. Endopod one-segmented, elongate, with apical claw and three setae of different lengths.</p> <p>P1 (Fig. 15A). With unornamented intercoxal sclerite. Coxa massive, with several spinular rows as shown. Basis with inner and outer spinulose spines; with strong spinules at top of crescentic medial extension, and with smaller spinules at base of inner spine. Exopod three-segmented, arising at a lower level than that of endopod and reaching tip of ENP2; ENP: EXP length ratio 1.2; exopodal segments with strong outer spinules as shown; second segment shortest, third segment longest; first and second segments without inner armature, outer spine of first segment longer than that of second segment; third segment with five elements of which outer proximal and medial spines shorter than those of preceding segments. Endopod three-segmented, arising from pedestal; ENP1 about 2.8 times as long as wide, 1.4 times as long as second and third segments combined, with outer longitudinal row of strong spinules, with one inner stiff, long, strongly pectinate seta; second and third segments with strong outer spinules as shown, the former shorter, slightly longer than wide, with one inner seta; third segment elongate, 1.5 times as long as preceding segment, with one inner distal slender seta, one apical long element and one outer distal spine.</p> <p>P2 (Fig. 15B). With narrow intercoxal sclerite unornamented, with acute distal projection on each side. Coxa with three anterior rows of spinules. Basis with small inner acute outgrowth, with long slender inner spinules, with minute spinules at base of endopod, and with strong spinules between rami and at base of outer spine. Rami three-segmented, endopod longer than exopod. Exopod three-segmented, arising at a lower level than that of the endopod, reaching slightly above the middle of ENP3; exopodal segments with strong outer spinules as shown; EXP1 and EXP2 with outer distal acute projection, of EXP1 shorter, inner distal frill of first segment coarse, of second segment small, first segment without, second segment with inner seta; EXP3 with subdistal outer pore and seven elements – three outer spines, two apical elements and two inner setae of which distalmost one visibly shorter. Endopod three-segmented, arising from pedestal; endopodal segments with strong outer spinules as shown; ENP1 shortest, slightly longer than wide, with posterior small spinules proximally, with small inner distal spinules and with outer short acute projection, with one short inner seta; ENP2 elongate, twice as long as wide and 0.6 times as long as ENP3, with small inner distal spinules, with long outer distal acute projection, with two inner setae of which proximal shorter; ENP3 longest, elongate, six times as long as wide, with four elements (one inner and two apical setae, and one outer spine), with acute projection between outer spine and inner apical seta.</p> <p>P3 (Fig. 16A). With triangular praecoxa ornamented with transverse row of minute spinules. Intercoxal sclerite and coxa as in P2. Basis largely as in P2, but with smaller inner distal outgrowth and setiform outer element. Rami three-segmented, endopod slightly longer than exopod. Exopod largely as in P2. Endopod largely as in P2 except for lack of posterior spinules and with outer subdistal pore on ENP1, for inner seta of P3 ENP1 comparatively longer, for one long seta only on P3 ENP2, and for two inner setae on P3 ENP3.</p> <p>P4 (Fig. 16B). With intercoxal sclerite, praecoxa, and coxa as in P3. Basis as in P3 except for lack of inner distal outgrowth. Exopod longer than endopod; EXP: ENP length ratio 1.3; largely as in P3 except for three inner setae on P4 EXP3 of which medial one visibly thicker and strongly pectinate. Endopod reaching slightly beyond middle of EXP3; largely as in P3 except for one inner seta only on P4 ENP3.</p> <p>Armature formula of P1–P4 as follows: P5 (Fig. 12C). With outer seta of baseoendopod arising from setophore. Endopodal lobe triangular, reaching distal third of exopod; with small spinules along outer margin; with five elements – one outer and one inner apical and one inner subdistal long setae, and two inner spiniform elements. Exopod elongate, 2.2 times as long as wide; with spinules at base of innermost seta and on outer margin proximally; with six elements – three outer slender setae, two apical elements of which outermost one shorter and one inner seta.</p> <p>P6 (Fig. 12A). With three setae – one short bipinnate element, and one medial and one inner slender seta.</p> <p>Male</p> <p>GENERAL. Total body length measured from tip of rostrum to posterior margin of caudal rami ranging from 380 to 510 µm (mean 455 µm; n = 8; total body length of allotype 510 µm).</p> <p>PROSOME (Fig. 17A). As in female.</p> <p>UROSOME (Fig. 17A–B). As in female except for second – P6-bearing somite – and third urosomites separated, for medial row of eight ventral spinules on third urosomite, and for lack of spinules on hyaline frill of fourth urosomite.</p> <p>SEXUAL DIMORPHISM. Expressed in ventral somatic ornamentation (see above), antennule, and basis of P1, P2 ENP, P5 and P6.</p> <p>ANTENNULE (Fig. 18A–B). 11-segmented, haplocer. All segments smooth, except for proximal and subdistal spinular rows on first segment. All setae smooth except for two and four biarticulated setae on tenth and eleventh segments, respectively. Armature formula: 1(1); 2(11); 3(6); 4(1); 5(5+(1+ ae)); 6(2); 7(2 +2 spines); 8(0); 9(1); 10(4); 11(5+ acro). Acrothek consisting of two setae and one aesthetasc fused basally.</p> <p>ANTENNA, MANDIBLE, MAXILLULE, MAXILLA AND MAXILLIPED (not shown). As in female.</p> <p>P1. As in female except for basis without slender inner long spinules, without spinules at base of inner spine and with inner modified accessory spine (Fig. 18C).</p> <p>P2 EXP (not shown). As in female. Endopod (Fig. 18D) sexually dimorphic, two-segmented; ENP1 as long as ENP2 (excluding distal cylindrical extension of ENP2), with longitudinal row of outer spinules, with outer distal acute outgrowth, with one inner short seta; proximal part of ENP2 rectangular, distal extension cylindrical, 2.4 times as long as wide excluding distal cylindrical extension, with longitudinal row of outer spinules, with two inner setae arising from pedestal of which proximal (seta I on Fig. 18D) visibly shorter, subdistally with one strong inner and almost straight element (element III on Fig. 18D), with one lateral and one apical seta (setae IV and V on Fig. 18D) arising from distal elongate cylindrical extension fused to segment basally and with strong outer subdistal element strongly tapering distally (element VI on Fig. 18D).</p> <p>P3 AND P4 (not shown). As in female.</p> <p>P5 (Fig. 18E). With outer seta of baseoendopod arising from setophore. Endopodal lobe reaching middle of exopod, with outer spinules as shown, with two distal modified setae as shown of which outermost one shorter. Exopod triangular, 1.6 times as long as wide, with inner subdistal pore, with six setae – one proximal bipinnate seta, one medial and one subdistal short strong spines of which medial one shorter, one distal seta arising from setophore and two inner setae of which proximal one shorter.</p> <p>P6 (Fig. 17B). Asymmetrical, only one leg functional, each leg with three setae of which innermost one bipinnate, medial longest and bare, outermost shortest and bare.</p> <p>Variability</p> <p>Both caudal setae I spiniform in one female. One male possesses only one outer spine on P5 EXP.</p></div> 	http://treatment.plazi.org/id/0B3187C8FFB88965FDA63297FA9FA916	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Gómez, Samuel;Corgosinho, Paulo Henrique Costa;Rivera-Sánchez, Karen I.	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FF848968FDFA324AFC91AA8D.text	0B3187C8FF848968FDFA324AFC91AA8D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bulbamphiascus Lang 1944	<div><p>Bulbamphiascus Lang, 1944</p> <p>Type species</p> <p>Canthocamptus imus Brady, 1872, by original designation.</p> <p>Other species</p> <p>Bulbamphiascus denticulatus (Thompson, 1893), B. incus Gee, 2005, B. plumosus Mu &amp; Gee, 2000 and B. scilloniensis Gee, 2005.</p> <p>Species incertae sedis</p> <p>Bulbamphiascus chappuisi Rouch, 1962.</p> <p>Amended diagnosis</p> <p>Miraciidae: Diosaccinae. Body fusiform. Rostrum not fused to cephalothorax, triangular, elongate, with rounded apical margin, almost as long as first, second and third antennulary segments combined, with one subdistal sensilla on each side at third quarter of its length. Pro- and urosomites without extensions; with a flexible unsclerotized cuticle between prosome and urosome. First urosomite (P5-bearing somite) narrower than preceding somites in dorsal view. Female genital double-somite (genital – second urosomite – and third urosomites fused) completely fused ventrally, with a dorsolateral cuticular rib marking former division between somites. Anal somite trapezoidal, without anal operculum. Caudal rami 1.6 times as wide as long in dorsal view, with seven setae. Female antennule eight-segmented, with aesthetasc and seta fused basally on fourth segment and with apical acrothek on last segment. Male antennule haplocer, 10-segmented, with aesthetasc and seta fused basally on fifth segment, and with apical acrothek on last segment.Antennulary setae typically smooth. Antenna with allobasis and abexopodal seta; exopod threesegmented, armature formula 1-1-1,3,0. Mandibular palp biramous; endopod one-segmented, exopod two-segmented. Maxillule biramous; rami one-segmented. Maxilla with three endites, proximal and middle endites with two, distal endite with three elements; ENP one-segmented. Maxilliped subchelate; (syn)coxa with four, basis with two setae, ENP one-segmented with claw and three accompanying setae. P1–P4 with three-segmented rami. Male P1 basis sexually dimorphic, with inner unipinnate spine and inner modified accessory spine. P1 EXP shorter than ENP, reaching tip of ENP2; EXP2 with inner seta; ENP1 with inner seta; ENP3 longer than the ENP2, with one inner seta, a distal long and geniculate seta, and an outer spine. P2–P4 EXP and ENP of approximately the same length; P2–P4 EXP1 with inner seta; inner distal seta of EXP3 visibly shorter than other elements of same segment; P2 ENP2 with two setae, proximal shorter. Male P2 ENP sexually dimorphic, two-segmented; first segment with one inner seta; second segment with two inner setae, one inner subdistal sigmoid element, one subdistal and one distal seta arising from cylindrical extension fused basally to segment, and one strong distal outer element strongly tapering distally. P5 EXP with six setae in female and male; one or two medial outer elements transformed into short spines. P5 endopodal lobe with five setae in female, with two setae in male; armature of male P5 endopodal lobe strongly bipinnate.</p> <p>Armature formula of P1–P5 as follows:</p> <p>Sexual dimorphism expressed in male antennule, P2 ENP, P5, P6, and in genital and third urosomites being separated.</p> <p>Apomorphies for Bulbamphiascus</p> <p>We did not detect any apomorphy for Bulbamphiascus. However, Bulbamphiascus is part of a larger monophyletic group composed of Spinodiosaccus gen. nov., Spinopedia gen. nov. and Pallarica gen. nov. (Fig. 19). The subdistal outer element of the male P2 ENP2 with almost parallel margins, viz slightly tapering distally, is considered here as part of the ground pattern of this monophyletic group and is regarded as a potential synapomorphy for this clade. The modification of this spine into a strong tapering element is regarded here as secondary and is a synapomorphy for Bulbamphiascus and Spinodiosaccus gen. nov. The combination of an outer subdistal tapering spine and a sigmoid inner subdistal element on the male P2 ENP2 is unique to Bulbamphiascus (the outer subdistal spine on the male P2 ENP2 of Spinodiosaccus gen. nov. also tapers distally, but the inner subdistal element on the same segment of this species is rather straight).</p> <p>Bulbamphiascus chappuisi is here relegated to incertae sedis due to the shape of the rostrum and furca, and to the morphology of the male P2 ENP, the illustrations of which (see Rouch 1962: 247, figs 29–33) are too schematic, preventing any further comparison.</p> </div>	http://treatment.plazi.org/id/0B3187C8FF848968FDFA324AFC91AA8D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Gómez, Samuel;Corgosinho, Paulo Henrique Costa;Rivera-Sánchez, Karen I.	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FF8A8969FD093160FAC3AF96.text	0B3187C8FF8A8969FD093160FAC3AF96.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spinopedia Gómez & Corgosinho & Rivera-Sánchez 2021	<div><p>Spinopedia gen. nov.</p> <p>urn:lsid:zoobank.org:act: F6DD1CA7-26EE-476D-AC00-2C4401B63014</p> <p>Bulbamphiascus Lang, 1944: 19, pro parte.</p> <p>Type and only species</p> <p>Bulbamphiascus spinulosus Mu &amp; Gee, 2000 (= Spinopedia spinulosa (Mu &amp; Gee, 2000) comb. nov.), by monotypy.</p> <p>Diagnosis</p> <p>Miraciidae: Diosaccinae. Body fusiform; body surface seemingly covered with minute denticles. Rostrum not fused to cephalothorax, triangular, elongate, with round apical margin, almost as long as first and second antennulary segments combined, with one subdistal sensilla on each side at third quarter of its length. Pro- and urosomites without extensions; with a flexible unsclerotized cuticle between prosome and urosome. First urosomite (P5-bearing somite) narrower than preceding somites in dorsal view. Genital double-somite (genital – second urosomite – and third urosomites fused) completely fused ventrally, with a dorsolateral cuticular rib marking original division between genital and third urosomites. Telson trapezoidal, without anal operculum. Caudal rami short, about 1.5 times as wide as long in dorsal view, with seven setae. Female antennule eight-segmented, with aesthetasc and seta fused basally on fourth segment and with apical acrothek on last segment. Male antennule haplocer, 10-segmented, with aesthetasc and seta fused basally on fifth segment and with apical acrothek on last segment. Setae of first and second antennulary segments typically bipinnate, smooth in remaining segments. Antenna with allobasis and abexopodal seta; exopod three-segmented, armature formula 1-1-1,3,0. Mandibular palp biramous; endopod one-segmented, larger than exopod, the latter two-segmented. Maxillule biramous; rami one-segmented. Maxilla with three endites, proximal and middle endites with two, distal endite with three elements; ENP one-segmented. Maxilliped subchelate; (syn)coxa with four, basis with two setae; ENP one-segmented, with claw and three accompanying setae. P1–P4 with three-segmented rami. Male P1 basis sexually dimorphic, with inner unipinnate spine and inner modified accessory spine. P1 EXP shorter than ENP, reaching tip of ENP2; P1 ENP1 with strong bipinnate inner seta; P1 ENP3 2.5 times as long as ENP2, with one small inner seta, a distal long geniculate seta and one outer spine; P1 EXP2 with inner seta; P2–P4 EXP and ENP of approximately the same length. P2–P4 EXP1 with inner seta; inner distal seta of P2–P4 EXP3 visibly smaller than other inner setae on same segment. P2 ENP2 with two setae, proximalmost shorter. P2–P4 basis with inner chitinous projection. Male P2 ENP sexually dimorphic, two-segmented; first segment with one inner seta; second segment completely covered by spinules and setules of different lengths, with two inner setae subequal in length, two elements (one medial, one apical) issuing from elongate cylindrical extension fused to segment basally, and with one inner subdistal sigmoid blunt element and one outer subdistal blunt spine with almost parallel margins, viz slightly tapering distally, of which the former is shorter. P5 EXP longer than wide, comparatively shorter in male, with six setae in both sexes; two medial outer elements modified into stiff short spines in male; distal outer spine foliaceous in female. P5 endopodal lobe with five setae in female, with two setae in male; armature of male P5 endopodal lobe strongly bipinnate.</p> <p>Armature formula of P1–P5 as follows:</p> <p>Sexual dimorphism expressed in male antennule, P2 ENP, P5, P6, and in separated genital and third urosomites.</p> <p>Etymology</p> <p>The prefix ‘spino’ from the Latin ‘spīna’, ‘spine’, and the sufix ‘pedia’ from the Latin ‘pēs’, ‘pedis’, ‘foot’, refers to the presence of a spinular patch on the male P2 ENP1, and to the presence of several spinules and setules covering the male P2 ENP2. Gender feminine.</p> <p>Apomorphies for Spinopedia gen. nov.</p> <p>Two apomorphies for Spinopedia have been detected: i) presence of an inner subdistal spinular patch on the male P2 ENP1 and ii) male P2 ENP2 completely covered with spinules and setules of different lengths. In addition, the subdistal inner seta of the female P5 EXP arises directly from the ramus (i.e., the seta does not arise from a setophore) (character 42 in Table 3; see also Fig. 19). The lack of a setophore for the subdistal inner seta of the female P5 EXP is regarded here as autapomorphic for the genus and is a result of reversal. The presence of a setophore for the inner subdistal seta of the female P5 EXP is autapomorphic for a larger group of genera to which Spinopedia gen. nov. belongs (Fig. 19) and seems to have evolved convergently in more than one evolutionary lineage within the Diosaccinae (Fig. 19).</p> </div>	http://treatment.plazi.org/id/0B3187C8FF8A8969FD093160FAC3AF96	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Gómez, Samuel;Corgosinho, Paulo Henrique Costa;Rivera-Sánchez, Karen I.	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FF8B896AFD3D364EFAF9AD5E.text	0B3187C8FF8B896AFD3D364EFAF9AD5E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pallarica Gómez & Corgosinho & Rivera-Sánchez 2021	<div><p>Pallarica gen. nov.</p> <p>urn:lsid:zoobank.org:act: 78F45B32-1B47-41BB-99E0-E7F8327E3333</p> <p>Bulbamphiascus Lang, 1944: 19, pro parte.</p> <p>Type and only species</p> <p>Bulbamphiascus cibimae Pallares, 1982 (= Pallaricia cibimae (Pallares, 1982) comb. nov.), by monotypy.</p> <p>Diagnosis</p> <p>Miraciidae: Diosaccinae. Body fusiform. Rostrum not fused to cephalothorax, triangular, elongate, with round apical margin, almost as long as first, second and third antennulary segments combined, with one subdistal sensilla at third quarter of its length, on each side. Pro- and urosomites without expansions. Genital double-somite (genital – second urosomite – and third urosomites fused) completely fused ventrally, with a dorsolateral cuticular rib marking original division between somites. Caudal rami about 1.5 times as wide as long in dorsal view; with seven setae. Female antennule eight-segmented, with aesthetasc and seta fused basally on fourth segment, and with apical acrothek on last segment. Antennulary setae typically smooth. Antenna with allobasis and abexopodal seta; exopod threesegmented, armature formula 1-1-1,3,0. Mandibular palp biramous; endopod one-segmented, larger than exopod, the latter two-segmented. Maxillule biramous; rami one-segmented. Maxilla with three endites, proximal and middle endites with two, distal endite with three elements; ENP two-segmented. Maxilliped subchelate; (syn)coxa with three, basis with two setae; ENP one-segmented, with claw and three accompanying setae. P1–P4 with three-segmented rami. Male P1 basis sexually dimorphic, with inner unipinnate spine and inner modified accessory spine. P1 EXP shorter than ENP, reaching tip of ENP2; P1 ENP1 with strong bipinnate inner seta; ENP3 2.5 times as long as ENP2, with one small inner seta, a long distal geniculate seta, and an outer spine; EXP2 with inner seta. P2–P4 EXP and ENP of approximately same lengths; P2–P4 EXP1 with inner seta; inner distal seta of P2–P4 EXP3 visibly shorter than other inner setae on same segment. P2 ENP2 with two setae, proximalmost small. Male P2 ENP sexually dimorphic, two-segmented; first segment with one inner seta; second segment with two inner setae, two elements (one medial, one apical) issuing from elongate cylindrical extension fused to segment basally, and inner subdistal element almost straight and outer subdistal element slightly thicker and longer than the former and with almost parallel margins (i.e., slightly tapering distally). P5 EXP longer than wide in female, comparatively shorter in male, with six setae in female and seven setae in male; two medial outer elements stiff short spine-like in male, slender and smooth in female.</p> <p>P5 endopodal lobe with five setae in female, with two setae in male; armature of male P5 endopodal lobe strongly bipinnate.</p> <p>Armature formula of P1–P5 as follows:</p> <p>Sexual dimorphism expressed in male antennule, P2 ENP, P5, P6, and in separated genital and third urosomites.</p> <p>Etymology</p> <p>This genus is dedicated to Rosa E. Pallares for her contribution to the taxonomy of harpacticoid copepods from Argentina. Gender feminine.</p> <p>Apomorphies for Pallarica gen. nov.</p> <p>This genus is unique by the possession of seven setae on the male P5 EXP, which is regarded here as apomorphic for the genus. The presence of an additional seta is regarded here as a character reversal.</p></div> 	http://treatment.plazi.org/id/0B3187C8FF8B896AFD3D364EFAF9AD5E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Gómez, Samuel;Corgosinho, Paulo Henrique Costa;Rivera-Sánchez, Karen I.	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FF88896CFD203506FBB9A8F8.text	0B3187C8FF88896CFD203506FBB9A8F8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dinetia Gómez & Corgosinho & Rivera-Sánchez 2021	<div><p>Dinetia gen. nov.</p> <p>urn:lsid:zoobank.org:act: 570211A6-5E4D-42CA-A26D-61E2F0E6BC6A</p> <p>Syn. Bulbamphiascus Lang, 1944: 19, pro parte.</p> <p>Type and only species</p> <p>Bulbamphiascus minutus Dinet, 1971 (= Dinetia minuta (Dinet, 1971) comb. nov), by monotypy.</p> <p>Diagnosis</p> <p>Miraciidae: Diosaccinae. Body fusiform. Width: length ratio of caudal rami 0.8–1.2. Rostrum not fused to cephalothorax, elongate, with rounded tip. Female antennule eight-segmented, with aesthetasc and seta fused basally on fourth segment. Antennulary setae typically smooth. Antenna with allobasis and abexopodal seta; EXP three-segmented, armature formula 1-1-1,3,0. Mandibular palp extremely reduced. Maxillule biramous; rami one-segmented. Maxilla with three endites. Maxilliped subchelate; (syn)coxa with two, basis with two setae; ENP one-segmented, with claw and two accompanying setae. P1–P4 with three-segmented rami. P1 EXP shorter than ENP1, reaching its distal third; P1 ENP1 with strong bipinnate inner seta; ENP3 about twice as long as ENP2, with one small inner seta, a long distal geniculate seta and an outer spine; P1 EXP2 with inner seta, EXP3 with three outer spines, proximal one shortest, and two geniculate distal setae. P2–P4 EXP1 with inner seta; P2–P4 EXP3 with reduced inner armature. P2 ENP1 with inner setae; ENP2 with two setae of approximately same length. Female P5 EXP longer than wide, with six setae – three outer, two distal and one inner element. P5 endopodal lobe with five setae.</p> <p>Armature formula of P1–P5 as follows:</p> <p>Male unknown.</p> <p>Etymology</p> <p>This genus is dedicated to Alain Dinet for his contribution to the taxonomy of harpacticoid copepods from France. Gender feminine.</p> <p>Apomorphies for Dinetia gen. nov.</p> <p>The exclusion of Bulbamphiascus minutus from that genus and from any other genus presented here, and its reallocation into Dinetia gen. nov., is supported by, for example, the lack of the short inner distal seta on P2–P4 EXP3. Bulbamphiascus minutus probably bears a sister-group relationship with Pseudamphiascopsis Lang, 1944 and Rhyncholagena Lang, 1944, with which it forms a monophyletic group (see Fig. 19). These three genera share the longer P1 ENP1 relative to the P1 EXP. Given that B. minutus could not be included in Bulbamphiascus or into any other diosaccin genus, and pending the discovery of the male of this species, we propose a new genus, Dinetia gen. nov., for B. minutus. The assessment and significance of character 43 for D. minuta comb. nov. (see Table 3) is still pending until the male is described.</p> <p>Phylogenetic statistics and resolution</p> <p>Our partial phylogenetic analysis resulted in 12 most parsimonious topologies with 126 steps, CI of 42, and RI of 77 (see Supplementary File 1). The strict consensus topology (nine branches collapsed) returned the following clades of unclear phylogenetic affinities between each other: a) a small clade composed by Dinetia gen. nov., Pseudamphiascopsis and Ryncholagena; b) a clade formed by Antiboreodiosaccus Lang, 1944, Diosaccopsis Brian, 1925, Goffinella Wilson, 1932, Ialysus Brian, 1927, Pseudodiosaccopsis Lang, 1944 and Pseudodiosaccus T. Scott, 1906; c) a clade formed by Typhlamphiascus Lang, 1944, Monardius Huys, 2009, Tydemanella A. Scott, 1909, Robertsonia Brady, 1880, Amphiascus Sars, 1905, Pararobertsonia Lang, 1944, Sinamphiascus Mu &amp; Gee, 2000, Pallarica gen. nov., Spinopedia gen. nov., Bulbamphiascus and Spinodiosaccus gen. nov.; d) a clade composed of Amonardia Lang, 1944, Diosaccus Boeck, 1872, Pholenota Vervoort, 1964, Parialysus Nicholls, 1941, Amphiascoides Nicholls, 1941b, Paramphiascella Lang, 1944, Robertgurneya Apostolov &amp; Marinov, 1988, Miscegenus Wells, Hicks &amp; Coull, 1982, Neomiscegenus Karanovic &amp; Ranga Reddy, 2004, Paramphiascoides Wells, 1967, Schizopera Sars, 1905, Actopsyllus Wells, 1967, Balucopsylla Rao, 1972, Eoschizopera Wells &amp; Rao, 1976, Haloschizopera Lang, 1944, Helmutkunzia Wells &amp; Rao, 1976, Schizoperoides Por, 1968, Protopsammotopa Geddes, 1968 and Psammotopa Pennak, 1942. The phylogenetic position of the genera Dactylopodamphiascopsis Lang, 1944, Metamphiascopsis Lang, 1944 and Sarsamphiascus Huys, 2009 remains uncertain. It is important to highlight the high level of character convergence within the Diosaccinae.</p> <p>Clades a–d above are present in the 12 most parsimonious topologies (see Supplementary File 1), but their relationships are not consistent. The positions of the clade Dinetia gen. nov. – Rhyncholagena and the genera Sarsamphiascus, Metamphiascopsis and Dactylopodamphiscopsis are unclear, sometimes forming a clade with Typhlamphiascus – Spinodiosaccus gen. nov. (6 times), sometimes forming a clade with Amonardia – Psammotopa (6 times). The same happens to the clade Antiboreodiosaccus – Pseudodiosaccus which appears either as the sister group of the remaining Diosaccinae or as the sister group of a clade including Dinetia gen. nov. – Rhyncholagena, Sarsamphiascus, Metamphiascopsis, Dactylopodamphiscopsis and Typhlamphiascus – Spinodiosaccus gen. nov.</p></div> 	http://treatment.plazi.org/id/0B3187C8FF88896CFD203506FBB9A8F8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Gómez, Samuel;Corgosinho, Paulo Henrique Costa;Rivera-Sánchez, Karen I.	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
0B3187C8FF8E896FFF2930A5FAD1A853.text	0B3187C8FF8E896FFF2930A5FAD1A853.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Typhlamphiascus Lang 1944	<div><p>Key to the species of Typhlamphiascus Lang, 1944</p> <p>1. P4 ENP3 with two inner setae (five setae in all). Female: P5 EXP with tip normal or hook-like. Male: P1 basis with inner seta normal or modified, elongate.........2 [Por’s group I (Pors 1963)]</p> <p>– P4 ENP3 with one inner seta (four setae in all). Female: P5 EXP with tip normal. Male: P1 basis with inner seta normal, not modified....................................11 [Por’s group II (Pors 1963)]</p> <p>2. Female P5 EXP with tip hook-like; endopodal lobe with apical outer seta very small............3</p> <p>– Female P5 EXP with tip normal......................................................................................................4</p> <p>3. Female: antennule seven-segmented; P5 EXP and endopodal lobe with five setae each, endopodal lobe with apical inner element spiniform; P4 EXP3 with three inner setae (male unknown).............................................................. T. dentipes (I.C. Thompson &amp; A. Scott, 1903)</p> <p>– Female: antennule eight-segmented; P5 EXP and endopodal lobe with five setae each, endopodal lobe with apical inner element setiform; P4 EXP3 with two inner setae (male unknown).................................................................. T. blanchardi (T. Scott &amp; A. Scott, 1895) 1</p> <p>4. Female: antennule eight-segmented; P5 EXP with five, endopodal lobe with four setae. Male: P1 basis with inner spine not modified, with three to five inner accessory spines; P5 EXP with four, endopodal lobe with two bifurcate setae..... T. gracilicaudatus (I.C. Thompson &amp; A. Scott, 1903) 6</p> <p>– These characters not combined........................................................................................................5</p> <p>5. Female: antennule eight-segmented; P5 EXP with four, endopodal lobe with five setae. Male: P1 basis with inner element modified, elongate; P5 EXP with four, endopodal lobe with two bifurcate setae.................................................................................... T. longifurcatus Rouch, 1962</p> <p>– These characters not combined.......................................................................................................6</p> <p>6. Female: P5 EXP and endopodal lobe with five setae each............................................................7</p> <p>– Female: P5 EXP with six setae, endopodal lobe with five setae.....................................................8</p> <p>7. Female: caudal rami 1.4–1.5 times as long as wide, shorter than anal somite; P5 second exopodal seta from outer to inner margin 1.5 times as long as third element; third seta 1.7 times as long as first seta and longer than exopod; innermost (fifth) seta 1.2 times as long as fourth seta. Male: P1 basis with inner spine normal, with seven to eight inner accessory spines; P5 EXP with five seate, endopodal lobe with two setae; P5 second exopodal seta from outer to inner margin shorter than first seta....................................................................................... T. gracilis Por, 1963 7</p> <p>– Female: caudal rami twice as long as wide, as long as anal somite; P5 second exopodal seta from outer to inner margin 4.5 times as long as third element; third seta as long as first seta and shorter than exopod; innermost (fifth) seta 2.3 times as long as fourth seta. Male: P1 basis with inner spine modified, elongate, with three inner accessory spines; P5 EXP with five setae, endopodal lobe with two setae; P5 second exopodal seta from outer to inner margin longer than first seta.............................................................................................................................. T. drachi Soyer, 1963</p> <p>8. P4 EXP3 without inner armature, formula 0,2,3. Female: antennule nine-segmented (male unknown)............................................................................................... T. accraensis (T. Scott, 1894)</p> <p>– P4 EXP3 with two inner setae, formula 2,2,3. Female: antennule eight-segmented............9</p> <p>9. Caudal rami 1.2 times as long as wide, shorter than anal somite... T. capensis Kunz, 1975 comb. nov.</p> <p>– Caudal rami twice as long as wide, as long as anal somite.........................................................10</p> <p>10. Caudal rami as long as anal somite, rather cylindrical, with inner margin slightly concave. Female: second antennulary segment about 1.5 times as wide as long; P5 EXP with three outer setae subequal in length; outermost seta of P5 endopodal lobe long,reaching well beyond tip of exopod. Male: P1basis with three inner accessory spines, inner seta normal, not modified; P5 EXP with six setae, endopodal lobe with two bifurcate setae; P5 innermost exopodal seta bifurcate......... T. lutincola Soyer, 1963</p> <p>– Caudal rami 1.5 times as long as anal somite, elongate, elliptic. Female: second antennulary segment three times as long as wide; P5 EXP with outer proximal seta very long, medial and subdistal outer setae small and subequal in length. Male: second antennulary segment three times as long as wide; P1 basis without inner accessory spines, inner seta modified, elongate; P5 EXP with four setae, endopodal lobe with two bifurcate setae; P5 innermost exopodal seta normal... T. bouligandi Soyer, 1971</p> <p>11. P3 ENP3 with one inner seta, formula 1,2,1..............................................................................12</p> <p>– P3 ENP3 with two inner setae, formula 2,2,1..............................................................................13</p> <p>12. P2–P3 EXP3 without inner armature, formula 0,2,3; caudal rami nearly twice as long as wide (length: width ratio 1.8). Male: P1 basis with three inner accessory spines; P5 EXP with six normal (not bifurcated) setae, endopodal lobe with two bifurcate setae...... T. higginsi Chullasorn, 2009</p> <p>– P2–P3 EXP3 with one inner seta, formula 1,2,3; caudal rami slightly more than two times as long as broad (length: width ratio 2.3) (male unknown)..................... T. unisetosus Lang, 1965</p> <p>13. Caudal rami slightly more than three times as long as wide (length: width ratio 3.2). Female: P5 EXP with six setae, endopodal lobe with four setae. Male: P1 basis with eight inner accessory spines; P5 EXP with four normal setae, endopodal lobe with two bifurcated elements................................................................................................................................. T. tuerkayi Ma &amp; Li, 2017</p> <p>– Female: P5 EXP with six, endopodal lobe with five setae...........................................................14</p> <p>14. P4 ENP2 without inner seta; caudal rami from 1.7 to 2.2 times as long as wide. Male: P1 basis with four 3 or from seven to eight 4 inner accessory spines; P5 EXP with six normal setae, endopodal lobe with two bifurcated elements........................ T. typhlops (Sars, 1906) 2,8</p> <p>– P4 ENP2 with inner seta...............................................................................................................15</p> <p>15. P1 EXP3 with four elements; caudal rami three times as long as wide (male unknown)....................................................................................... T. brevicornis (I.C. Thompson &amp; A. Scott, 1903)</p> <p>– P1 EXP3 with five elements..........................................................................................................16</p> <p>16. Caudal rami less than 1.5 times as long as wide.....................................................................17</p> <p>– Caudal rami more than 1.5 times as long as wide...................................................................18</p> <p>17. Caudal rami about as long as wide. Female: P5 EXP elongate. Male: P1 basis with three inner accessory spines; P5 EXP with five setae..................................... T. latifurca Por, 1968</p> <p>– Caudal rami length: width ratio from 1.3 to 1.5. Female: P5 EXP oval. Male: P1 basis with eight inner accessory spines; P5 EXP with six setae.......................... T. ovale Wells &amp; Rao, 1987</p> <p>18. Caudal seta V characteristically deformed proximally, with inner blunt outgrowth.......................................................................................................................................... T. typhloides (Sars, 1911)</p> <p>– Caudal seta V normal.....................................................................................................................19</p> <p>19. Length: width ratio of caudal rami variable, from about 1.7 to 3.1 times as long as wide, with concave inner margins, with lenticular, biconvex thickening of external chitinous walls, with one spinule on inner distal margin. Female: ventral surface of posterior half of genital double-somite without medial spinules, with two posterior transverse sets of three to five spinules, each close to outer margin; ventral surface of fourth urosomite with one medial spinular row of 21 to 30 spinules and one posterior transverse row of 15 to 24 spinules; ventral surface of fifth urosomite with two medial sets of three to ten spinules on each side or with a medial continuous row of about 24 spinules, and with short posterior row of four to six medial spinules or with three sets of spinules of which medial one with ten spinules and marginal sets with three to four spinules; ventral surface of anal somite without spinular ornamentation. Male: ventral surface of third urosomite with two medial transverse sets of four spinules each or with medial continuous spinular row of eight to ten transverse spinules, posterior margin with transverse row of 14 to 26 spinules; ventral surface of fourth urosomite with two medial sets of four to eight transverse spinules or with short medial row of four transverse spinules, and with posterior transverse row of 16 to 24 spinules; ventral surface of fifth urosomite without medial spinules or with two medial sets of four transverse spinules, posterior margin with transverse row of four to 12 spinules; P1 basis with three to five inner accessory spines; P5 EXP with six setae, endopodal lobe with two bifurcate setae............................................................................................................................ T. confusus confusus (T. Scott, 1902) 5,7</p> <p>– Caudal rami from about 1.7 to 3.2 times as long as wide; ventral spinular pattern of female and male and number of inner accessory spines on male P1 basis different..........................20</p> <p>20. Caudal rami shorter than anal somite...........................................................................................21</p> <p>– Caudal rami longer than anal somite...........................................................................................22</p> <p>21. Rostrum with concave lateral margin proximally; length: width ratio of caudal rami 1.7. Female: P5 EXP with three outer setae subequal in length, fourth seta from outer to inner margin longer than preceding elements and of about same length as innermost seta. Male: P1 basis with four inner accessory spines......... T. confusus erythraeicus Por, 1963</p> <p>– Rostrum with convex lateral margin proximally; length: width ratio of caudal rami 1.7. Female: P5 EXP with outer proximal seta shorter than two neighbouring elements, fourth seta from outer to inner margin shorter than two preceding elements and shorter than innermost seta. Male: P1 basis with two inner accessory spines....................... T. lamellifer (Sars, 1911) 8</p> <p>22. Length: width ratio of caudal rami from 2.0 to 3.2. Female: three outer setae and innermost seta of P5 EXP long, subequal in length. Male: P1 basis with five to seven inner accessory spines................................................... T. confusus gullmaricus Por, 1963</p> <p>– Length: width of caudal rami from 1.7 to 2.2. Female: proximal seta of P5 EXP longer than two following elements, all three outer setae noticeably shorter than innermost element. Male: P1 basis with three inner accessory spines; innermost seta of P5 EXP bifurcate.........23</p> <p>23. Caudal rami with subdistal inner set of long spinules forming a comb. Female: both halves of genital double-somite without dorsolateral spinular ornamentation; setae of P6 subequal in length; with five midventral spinules on fourth urosomite. Male: genital, third and fourth urosomites without dorsolateral spinular ornamentation............................ T. pectinifer Lang, 1965</p> <p>– Caudal rami with few subdistal inner short spinules; inner setae of P6 noticeably longer than outer ones. Female: both halves of genital double-somite with a few posterior spinules dorsolaterally; with 13 midventral spinules on fourth urosomite. Male: genital, third and fourth urosomites with few a dorsolateral spinules......................................... T. medici sp. nov.</p></div> 	http://treatment.plazi.org/id/0B3187C8FF8E896FFF2930A5FAD1A853	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Gómez, Samuel;Corgosinho, Paulo Henrique Costa;Rivera-Sánchez, Karen I.	Gómez, Samuel, Corgosinho, Paulo Henrique Costa, Rivera-Sánchez, Karen I. (2021): Proposal of new genera and species of the subfamily Diosaccinae (Copepoda: Harpacticoida: Miraciidae). European Journal of Taxonomy 759 (1): 1-62, DOI: https://doi.org/10.5852/ejt.2021.759.1433, URL: http://dx.doi.org/10.5852/ejt.2021.759.1433
