taxonID	type	description	language	source
AE44F266FFDB340E6C92F8B3FD635293.taxon	description	Figs. 1, 2 A, 3, 4 A, 5, 6.	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFDB340E6C92F8B3FD635293.taxon	materials_examined	Non-sequenced specimens examined. Paralectotypes: MNHN: 254, 254 A, 254 B, “ Van Diemen’s Land ”. Additional topotypic specimens from Peron Peninsula, Shark Bay Western Australia): WAM: R 54636, 1 km S Monkey Mia, Shark Bay, WA (25 º 48 ’ S, 113 º 42 ’ E), R 54639 – 40, Peron Homestead, Peron Peninsula, WA (25 º 50 ’ S, 113 º 33 ’ E), R 54654, Monkey Mia, Shark Bay, WA (25 º 48 ’ S, 113 º 43 ’ E), R 54717, Peron Hills, 28 km NW Denham, WA (25 º 43 ’ S, 113 º 26 ’ E), R 54638, 54806 – 07, 10 km NE Denham, Shark Bay, WA (25 º 52 ’ S, 113 º 36 ’ E), R 54873, 13 km S Nanga Peron Peninsula, WA (26 º 20 ’ S, 113 º 54 ’ E), R 55003, 55067, 18 km SE Nanga, Peron Peninsula, WA (26 º 20 ’ S, 113 º 58 ’ E), R 55069 - 72, 8 km SE Nanga, Peron Peninsula, WA (26 º 17 ’ S, 113 º 51 ’ E), R 60428, 3 km N Peron Homestead, WA (25 º 48 ’ S, 113 º 33 ’ E), R 123646, 3 km NW Peron Homestead, WA (25 º 49 S’, 113 º 33 ’ E).	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFDB340E6C92F8B3FD635293.taxon	diagnosis	Diagnosis. Distinguished from other Australian Gehyra (except G. versicolor) by a combination of 7 or 8 divided scansors under the expanded portion of the fourth toe, moderate size, generally two pairs of enlarged chin shields, second infralabial notched and a dorsal pattern in which dark lines and white markings coordinate to produce a pattern of dark lines and bars with white trailing edges. Not readily distinguishable by external morphology from G. versicolor sp. nov. (see below), but distinguished karyotypically by the unique 2 n = 40 b arrangement (King 1979). Otherwise most similar to G. montium but distinguished by grey to brown rather than more rufous dorsal colouring, with white markings that form a posterior highlight or margin on the trailing edge of the dark dorsal lines, rather than small poorly contrasting dots that are not coordinated with the dark markings. This diagnosis applies to populations of Gehyra genetically assignable to the “ variegata clade ” of Sistrom et al. (2013).	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFDB340E6C92F8B3FD635293.taxon	description	Description (based on the lectotype and recent, genotyped material) Adult snout-vent length 34 – 49 mm (mean = 44.2 mm, n = 27). Length of original tail 43 - 48 mm (mean 107 % of SVL, n = 3). Sarre (1998) recorded that individuals in some Western Australian Wheatbelt populations can reach 60 mm SVL. Nostril bordered by rostral, first supralabial, supranasal and two subequal post nasals. Most frequently, a single moderate internasal separates the supranasals above the rostral, but supranasals often (f = 0.41) in medial contact. Supralabials 8 – 10 (mode 9). Infralabials 7 – 10 (mode 9). Usually two pairs chin shields (four specimens with 3 pairs), anterior pair in contact with only the first infralabial. Chin shields separated from the third and succeeding infralabials by the interpolation of a series of enlarged scales (parinfralabials, King 1982) that margin the ventral edge of the infralabials. Second infralabial notched where this parinfralabial scale row starts (Fig 4 A). Scansors under pad of fourth toe divided, 6 – 8 (mode 8). Precloacal pores in males 10 – 15 (mean = 12.5, n = 11) arranged in a chevron with median pore anterior most. The karyotype shown in Fig. 2 A is 2 n = 40 b (King 1979, Moritz 1986). In life, dorsally light to medium grey or brown, generally with a complex pattern of white-edged black lines (Figs 1 B, 5 A-B). These usually include several temporal streaks and often form continuous paravertebral and dorsolateral irregular lines, often with short cross bars that may form a vaguely ladder-like pattern. A trend is for specimens from rock-dwelling populations to have bolder and more contrasting markings, especially the white highlights which are expressed as spots (Fig. 5 A), while some animals from arboreal populations may have only weakly contrasting patterns (Fig. 5 B). However, colour pattern and intensity is variable both within and between populations, and individual animals can change colour intensity from pale to very dark in the course of a day. In preservative, the colour pattern is often greatly reduced in contrast and can be hard to discern. These patterns of extensive interpopulation, intrapopulation and temporal colour variation are characteristic of most species of Australian Gehyra.	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFDB340E6C92F8B3FD635293.taxon	distribution	Distribution. Widespread through the southern half of Western Australia, from the west coast as far north as the Pilbara, east to the Central Ranges and southeast to the western interior of South Australia (Fig. 3).	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFDB340E6C92F8B3FD635293.taxon	discussion	Comments. General. The combination Gehyra variegata has long been applied Australia-wide to populations of morphologically similar Gehyra species with similarly generalist habits. Sistrom et al. (2013) clearly distinguished two major genetic clades within these populations, and confirmed that the karyotypic differences between western and eastern populations first noted by King (1979) reflect the existence of at least two distinct species. Indeed, the two are not even sister lineages. The two populations have generally similar colour patterns but are subtly different in proportions. Sistrom et al. (2013) could distinguish samples of the two species using multivariate analysis but extensive overlap of character variation is such that identification of an individual from morphology alone is problematic. Allocation of the name variegata to the western species centres on the correct identification of the type locality and evidence that specimens from the vicinity of this locality are conspecific with the more widespread western populations.	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFDB340E6C92F8B3FD635293.taxon	materials_examined	Type locality. The syntypes of Hemidactylus variegatus in the MNHN collection are from two localities, Shark Bay, Western Australia (one specimen, MNHN 2295) and Tasmania (three specimens, MNHN 254, 254 A and 254 B). Duméril & Bibron (1836) reported that the collectors of MNHN 2295 were [Jean-René] Quoy and [Joseph Paul] Gaimard, naturalists who travelled with Freycinet’s 1817 – 1820 expedition. The Shark Bay region (specifically the Peron Peninsula and Dirk Hartog Island) was the only location where the expedition spent any time (a week in September 1818) collecting in an area of Australia that would have yielded specimens of Gehyra (Arago, 1823). By contrast, the paralectotypes, MNHN 254, 254 A and 254 B, cannot be associated with a collecting locality. The given locality of Van Diemen’s Land (Tasmania) is in error as no Gehyra species (in fact, no gekkotans) occur in Tasmania, Hutchinson et al. 2001). The three specimens were collected on the Baudin expedition (1800 – 1803), which visited several localities along the west coast of Western Australia, including Shark Bay, where specimens of Gehyra might have been obtained (Bonnemains et al. 1988), but there is no more detailed information that would allow the locality to be identified. All four syntypes are in poor condition, very faded, and soft; MNHN 254 B is almost macerated. Without seeing any of these specimens, and under the misapprehension that all were from Shark Bay, Wells & Wellington (1983) named “ the largest ” specimen registered under MNHN 2295 as the lectotype. In fact there is only a single specimen registered under this number. The lectotype is distinctive in being very small (SVL 35 mm) for an apparently adult male (highly active precloacal pores). A series of 16 Western Australian Museum specimens from the Peron Peninsula was examined and revealed only one Gehyra morphotype, similar to the “ variegata clade ” in colour pattern. All specimens are noticeably small in size, the three largest individuals (WAM R 54640, 54654 and 55070) being 40 mm SVL. The smallest male with active precloacal pores (WAM R 54636) measured 34 mm SVL; the smallest females with an enlarging ovarian follicle or a fully yolked egg measured 38 mm SVL. By comparison, in our random sample of genetically typed specimens from across the species range the smallest male with active pores and smallest ovigerous female both measured 41 mm SVL and the largest animals were 49 mm SVL. Thus the Shark Bay specimen (MNHN 2295) is consistent in its small adult size with recently collected specimens from the same area, and we are confident that the type locality for Hemidactylus variegatus is the Peron Peninsula and that MNHN 2295 is correctly designated as lectotype. Conspecifity of topotypic and “ variegata clade ” populations. Molecular data (Sistrom et al. 2013) show that all of the samples that have been DNA sequenced from the vicinity of Shark Bay (specimens from Peron homestead, Cape Rose, Babbage Island, Bush Bay, Faure Island, Baudin Island, and the North-west Highway near ‘ Billabong’ roadhouse) are members of the “ variegata clade ”. Our two sequenced Peron Peninsula specimens (Cape Rose and Peron homestead) are closely related, and some other Shark Bay area specimens are relatively close genetically to them (e. g., the Carnarvon area specimens from Babbage Island and Bush Bay), while others are more distant (e. g., the Faure Island specimens). Equally, some geographically remote specimens are close genetic relatives of Shark Bay area specimens; the nearest relative of the Peron Peninsula specimens is a specimen from near Dampier (mouth of the Yanyare River) more than 600 km to the northwest, while the Carnarvon specimens are even more closely related to Pilbara specimens (Pannawonica, Muggon Station), than to the Peron Peninsula specimens, and one of the Faure Island specimens is closest to one from Neale Junction on the edge of the Great Victoria Desert, over 1200 km distant. These patterns of genetic relatedness are consistent with a single species with subpopulations that have undergone dispersal and introgression subsequent to a period of isolation and differentiation. Genetic studies in the Western Australian wheatbelt have suggested that this species of Gehyra can disperse widely over relatively short time scales (Hoehn et al. 2007). However the species varies in body size and this morphological variation may be indicating very recent differentiation not reflected in the initial genetic survey. In contrast to the dwarfish populations on the Peron Peninsula, populations in the Western Australian Wheatbelt, can grow to large body sizes. How et al. (1988) and Sarre and coworkers (Sarre et al. 1995; Sarre 1996, 1998) who worked on these populations reported the smallest breeding female at 44 mm SVL and the maximum recorded SVL as much as 61 mm. As with the Peron Peninsula ‘ dwarfs’ these Wheatbelt ‘ giants’ are nested genetically well within the “ variegata clade ”. We note that body size alone need not be an indicator of speciation in Gehyra, as our data in this study on G. moritzi sp. nov. (q. v.), as well as striking size variation in G. lazelli (Sistrom et al. 2012) show that intraspecific selection for local differences in body size has happened more than once in the Australian Gehyra radiation. Consequently, we conclude that the observed body size variation alone does not overturn the idea of only a single species across these populations. Further work, integrating additional genetic and morphological data, will be necessary to better understand the variation seen among populations of this species as we have construed it. Contact between G. variegata and G. versicolor, sp. nov. The contact between our restricted concept of G. variegata and the eastern G. versicolor sp. nov. (2 n = 40 a / Clade 5 of Sistrom et al. 2013) appears to occur at the eastern margins of the Nullarbor Plain (Fig. 6). King (1979) mapped 2 n = 40 a individuals as far west as Bates on the transcontinental railway line. Sistrom et al. (2013 and unpublished) found only true G. variegata (DNA data from nine individuals from the far west of South Australia), and these findings have been supported by seven more individuals that we karyotyped from around Maralinga (40 km NW of Bates) and a site on the northeast Nullarbor Plain (23 km SE of Bates) which all proved to have the 2 n = 40 b karyotype. Thus our data, when combined those of King, appear to show an abrupt changeover in this area from G. variegata to the eastern species. It is noteworthy that we have found only G. variegata (i. e,. 2 n = 40 b) in this area while King found only G. versicolor sp. nov. (2 n = 40 a), one from Bates, another five from several localities approximately 160 – 180 km further east. Further detailed sampling in this area with both karyotypic and DNA sequence data would be very useful in exploring whether there is any gene flow between the two.	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFD2340A6C92FADCFD8D57F3.taxon	description	Figs 2 C – D, 3, 6, 7.	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFD2340A6C92FADCFD8D57F3.taxon	materials_examined	Holotype: SAMA R 51968 K, from 1.9 km SW of Reedy Hole Springs, Flinders Ranges, South Australia (30 ° 15 ’ 55 ˮ S, 138 ° 49 ’ 30 ˮ E), collected by J. Bice, on 19 November, 1998 (Fig. 7 A). Paratypes (n = 28; all genotyped as ‘ Clade 5 ’): SAMA: R 26185, Birdsville, Qld (25 ° 54 ’ S, 139 ° 21 ’ E), R 28201, 1 km S Mt Dutton, SA (27 ° 49 ’ S, 135 ° 43 ’ E), R 34249, 7 km E Mount Isa, Qld (20 ° 43 ’ S, 139 ° 43 ’ E), R 38930, Yudnamatana, SA (30 ° 10 ’ S, 139 ° 17 ’ E), R 38942, R 38945, Lancoona HS, NSW (32 ° 22 ’ S, 145 ° 53 ’ E), R 50277, 7 km SSE Mt Deception, Beltana Station, SA (30 ° 46 ’ S, 138 ° 17 ’ E), R 51609 K, 26.1 km ENE of Mimili, SA (26 ° 54 ’ 43 ” S, 132 ° 56 ’ 55 ” E), R 51637 K, 30.3 km WNW Indulkana, SA (26 ° 52 ’ 09 ” S, 133 ° 01 ’ 29 ” E), R 51760 K, 1.75 km W Yudnamutana Bore, SA (30 ° 10 ’ S, 139 ° 16 ’ E), R 51782 K, 10.4 km SW Yudnamutana Bore, SA (30 ° 14 ’ S, 139 ° 11 ’ E), R 51912, 0.5 km NW Nudlamutana Well, SA (30 ° 22 ’ S, 139 ° 21 ’ E), R 51962 K, 2.8 km W Moosha Bore, SA (30 ° 19 ’ S, 138 ° 47 ’ E), R 52366, 4.7 km W Parachilna Hill, SA (31 ° 08 ’ S, 138 ° 33 ’ E), R 54530, 4.5 km N Station Creek crossing, Prairie – Muttaburra road, Qld (22 ° 02 ’ S, 144 ° 37 ’ E), R 54546 – 47, 14 km NW Longreach on Landsborough Highway, Qld (23 ° 21 ’ S, 143 ° 12 ’ E), R 55133, 2.7 km E Gluepot HS, SA (33 ° 45 ’ S, 140 ° 09 ’ E), R 55268, Phosphate Hill mine, ‘ Snappy Site’, Qld (21 ° 53 ’ S, 139 ° 59 ’ E), R 55695 – 96, 13.4 km NNE Hughenden on Kennedy Developmental Road, Qld (20 ° 47 ’ S, 144 ° 19 ’ E), R 55905, 9 km N New South Wales / Queensland border on Mitchell Highway, Qld (28 ° 58 ’ S, 145 ° 44 ’ E), R 57970, 30.1 km SSW Memory Bore, SA (28 ° 53 ’ S, 132 ° 44 ’ E), R 63576, Mutawintja NP, NSW (31 ° 17 ’ S, 142 ° 18 ’ E), R 64097, Terrapinna Springs, SA (29 ° 55 ’ S, 139 ° 40 ’ E), R 64443, Mt Fitton HS, SA (28 ° 59 ’ S, 139 ° 33 ’ E), R 64447, 17 km E Mt Fitton HS, SA (29 ° 54 ’ S, 139 ° 25 ’ E), R 64549, 1.3 km WNW Nantawarrinna, SA (30 ° 49 ’ S, 138 ° 58 ’ E), R 64863, 6.9 km WNW Arkaroola HS (30 ° 18 ’ S, 139 ° 19 ’ E).	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFD2340A6C92FADCFD8D57F3.taxon	diagnosis	Diagnosis. Morphologically shares the external features of G. variegata (see above) but distinguished from that species by karyotype (2 n = 40 a or 2 n = 38 a) (Fig. 2 C – D). As with G. variegata, G. versicolor is most similar G. montium but distinguished by grey to brown rather than more rufous dorsal colouring, with white markings that form a posterior highlight or margin on the trailing edge of the dark dorsal lines, rather than forming discrete circular dots that are not coordinated with the dark markings. This diagnosis applies to populations of Gehyra genetically assignable to “ Clade 5 ” of Sistrom et al. (2013).	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFD2340A6C92FADCFD8D57F3.taxon	description	Description. Adult snout-vent length 37 – 54 mm (mean = 46.7 mm, n = 29). Length of original tail 40 – 58 mm (mean = 110 % SVL, n = 6). Nostril bordered by rostral, first supralabial, supranasal and two subequal postnasals. Usually a single internasal scale (occasionally 2 or none) separates the supranasals above the rostral. Supralabials 8 – 11 (mode 9). Infralabials 7 – 10 (mode 9). Usually (f = 0.6) two pairs of chin shields; if a third pair is visible it is markedly smaller than the second or asymmetric, anterior pair in contact with only the first infralabial. Chin shields separated from the third and succeeding infralabials by the interpolation of a series of enlarged scales (parinfralabials) that margin the ventral edge of the infralabials. Second infralabial notched where this parinfralabial scale row starts. Scansors under pad of fourth toe divided, 7 – 8 (mode 8). Precloacal pores in males 10 – 14 (mean = 11.9, n = 15). Most populations uniformly have the 2 n = 40 a karyotype first reported by King (1979), but within central Australia, geographically interspersed with 2 n = 40 a individuals, are individuals with 2 n = 38 karyotypes. These specimens all belong to the same clade on the basis ofsequence data. Moritz (1986) reported two variants of the 2 n = 38 karyotype that differ only in that the 2 n = 38 a karyotype had pair 8 acrocentric while the 2 n = 38 b karyotype had this pair symmetrically metacentric. Our karyotype (Fig. 2 B) from Native Gap (a population scored as 2 n = 38 a by Moritz 1986) has a pair 8 that is submetacentric, neither obviously acrocentric nor symmetrically metacentric, and in fact King (1979) described and illustrated his 2 n = 38 karyotype (later termed the “ a ” variant by Moritz) as having submetacentric pairs 7 and 8, as ours does. The discrepancy may be artifactual, given the variation in condensation and clarity that is typical across karyotypic preparations. In case further cryptic species are demonstrated among these populations, we have confined our type series for G. versicolor to animals from 2 n = 40 a populations only. In life, dorsally light to medium grey or grey-brown, generally with a complex pattern based on one or two pairs of more or less continuous dark longitudinal irregular dorsal stripes with numerous cross links and reticulations, the dark markings all coordinated with white highlights that can form irregular spots but are consistently in contact with the darker markings as in G. variegata. In general, specimens of G. versicolor, even when strongly patterned, tend to have less distinct and contrasting patterns (Fig. 7) than do well marked G. variegata. Common variant patterns include one where some of the dark dorsal markings form back-swept angular or crescentic bars, each with a whitish spot contained in the angle of the bar, and another where the body is densely stippled with low contrast darker grey bars and blotches with only weakly apparent paler edges. None of these variations is unvarying and the colour patterns show continuous variation both within and between populations.	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFD2340A6C92FADCFD8D57F3.taxon	distribution	Distribution. Widespread from the Murray Valley of northern Victoria north and east through New South Wales west of the Great Dividing Range and similar areas of Queensland north to about the latitude of Hughenden. Extends west into most of South Australia, with the exception of the southern and western Eyre Peninsula and the Great Victoria Desert, and north-west into southern and central Northern Territory. Not currently known to occur in Western Australia. Found in both rocky and arboreal situations, as well as on buildings.	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFD2340A6C92FADCFD8D57F3.taxon	discussion	Comments. This species is the only one where we find two karyotypic groups appearing to belong to a single taxon. Moritz (1986) reported both 2 n = 40 a and 2 n = 38 (a and b) from the MacDonnell Ranges and adjacent central Northern Territory, and at present our data suggests all belong to a single species, G. versicolor. As with the variable populations of G. moritzi, further detailed study combining the same multiple approaches used here are desirable to clarify the gene flow among these chromosomally different populations. Similar detailed studies are needed in central and western Queensland to better understand the distribution and variation of G. versicolor and Clade 4 where they co-occur (Sistrom et al. 2013), and the potential contact or overlap between G. variegata and G. versicolor in central western South Australia. However, it is clear that over the great majority of its distribution this is a single species, consistently different from other Gehyra. Most of the literature pertaining to the biology of “ Gehyra variegata ” (e. g., Bustard 1965, 1967, 1968, 1969; Henle 1990) actually applies to this species, as does recent population genetic work by Duckett and Stow (2010, 2013). Swan et al. (2004) report the maximum SVL for this species (as G. variegata) as 57 mm.	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFD2340A6C92FADCFD8D57F3.taxon	etymology	Etymology. The specific name chosen here is from the Latin meaning ‘ variable in colour’, appropriate for a species that shows considerable individual and geographic variation. Given the very wide distribution of the species it is somewhat surprising that no older name appears to be available for it in the synonymy of G. variegata. Cogger et al. (1983) listed several synonyms of Gehyra variegata. Subsequently, these have proven to be based on specimens attributable to other Australian Gehyra species groups, especially eastern species related to G. australis (Bauer & Henle 1994; Shea 1995). Other possible synonyms were discussed by Sistrom et al. (2009) in reference to G. lazelli; none apply to G. versicolor.	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFD634086C92FE37FCDA5414.taxon	description	Figs 3, 8.	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFD634086C92FE37FCDA5414.taxon	diagnosis	Diagnosis. Distinguished from other Australian Gehyra by a combination of modally seven divided scansors under the expanded portion of the fourth toe, small to moderate size, generally two pairs of enlarged chin shields, second infralabial notched, a dorsal colour pattern combining pinkish brown to rufous colouring (in life) usually patterned by small pale spots interspersed in a continuous network of irregular, dark lines, and a karyotype of 2 n = 42 a (illustrated in King 1979, fig. 9; Moritz 1986, fig. 3). Distinguished from some other rock dwelling species (G. minuta, G. moritzi sp. nov. and G. pulingka sp. nov.) by more reticulate dorsal pattern and often weak contrast of pale spots. Most similar to G. variegata and G. versicolor sp. nov. but somewhat smaller and usually more rufous in colouring with dorsal white markings consisting of small poorly contrasting dots (readily lost in preservative) that are not consistently coordinated with the dark markings. This diagnosis applies to populations of Gehyra genetically assignable to the “ montium clade ” of Sistrom et al. (2013).	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFD634086C92FE37FCDA5414.taxon	description	Description. Adult snout-vent length 36 – 51 mm (mean = 41.7 mm, n = 29). Length of original tail 41 – 58 mm (mean = 113 % SVL, n = 9). Nostril bordered by rostral, first supralabial, supranasal and two post nasals, the upper usually markedly larger than the lower. None to three (modally 1) moderate internasal scales separate the supranasals above the rostral. Supralabials 8 – 11 (mode 8). Infralabials 7 – 9 (mode 8). Consistently two pairs of chin shields, anterior pair in contact with only the first infralabial. Chin shields separated from the third and succeeding infralabials by the interpolation of a series of enlarged scales (parinfralabials) that margin the ventral edge of the infralabials. Second infralabial notched where this parinfralabial scale row starts. Scansors under pad of fourth toe divided, 7 – 8 (mode 7). Precloacal pores in males 10 – 13, (mean = 11.3, n = 16). The karyotype is 2 n = 42 a, a karyotype that is shared with a number of Gehyra species, including G. minuta and some populations that are morphologically distinct from G. montium (King 1979, fig. 13 a) and currently assigned to G. punctata (Fry, 1914) in Western Australia. In life, dorsally light grey-brown to reddish brown to pinkish, the entire dorsal surface patterned by a strong to weak reticulum of blackish grey (Fig. 8). Scattered over the dorsal surface are small circular pale spots, often only contrasting weakly with the dorsal background colour. In preservative any rusty colour tones and pale spots tend to disappear leaving the specimens greyish with a dark reticulum.	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFD634086C92FE37FCDA5414.taxon	distribution	Distribution. The western portion of the Australian arid zone, from the north-west of South Australia, and south-east of the Northern Territory, north-west to the Pilbara Region of Western Australia.	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFD634086C92FE37FCDA5414.taxon	discussion	Comments. Throughout arid areas of central and northern Australia, rocky outcrops may harbour relatively small Gehyra species, typically with rufous colouring and a pattern including pale spots. The name G. montium has often been applied to many such populations, but our study reveals that this species only just extends east of the Western Australian border, to the Tomkinson Ranges and Birksgate Ranges (east to Mt Kintore) in South Australia and is probably restricted in the Northern Territory to the far south-west corner (Uluru and the Petermann Ranges). The fact that the species was not recognised hitherto as extending westward as far as the Pilbara possibly reflects a tacit assumption that G. montium was a central Australian species, as well as the similarity of colour pattern in preserved G. montium and G. variegata. This similarity is such that individuals of the two species will continue to be difficult to separate by morphology alone In the adjacent rocky ranges of north-west South Australia and the southern and central Northern Territory, G. montium is replaced by geckos of Clades 2 and 1 (Sistrom et al. 2013), respectively, and these are described below as new species. Populations we have assigned to this species require further study in several areas. Colour and pattern as described here is based on central Australian populations for which we had freshly collected material of known genotype; preserved specimens examined by us suggest that this colour pattern extends to Pilbara populations, but in this region the species is more variable in colour pattern. Animals of known genotype from this area include not only animals with a dark dorsal network, but also almost plain individuals, grey in preservative but possibly rusty in life, with no more than scattered small black speckles. Three specimens genotyped as G. montium by Sistrom et al. (2013) are from areas to the south and west of the species’ main distribution, apparently in areas of sandy desert rather than rocky ranges. Further collection and typing of specimens from these populations is needed to confirm that these seemingly atypical records are correctly documented, and that there are populations of Gehyra from these areas that pertain to this species. When describing this species Storr (1982) suggested it might represent the 2 n = 38 karyotypic group of King (1979). However, the data of Moritz and our own checks of karyotype show that populations conspecific with the type population of montium have the 2 n = 42 a karyotype.	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFD434086C92FD4BFF3653A9.taxon	description	Fig. 3. Distinguished from other Australian Gehyra by a combination of modally 7 divided scansors under the expanded portion of the fourth toe, generally two pairs of enlarged chin shields, with the median (postmental) pair being relatively small, second infralabial notched and often in contact with postmental, low numbers of labial scales (nine or fewer supralabials, eight or fewer infralabials), a dorsal colour pattern combining pinkish brown to rufous colouring (in life) patterned by circular whitish spots and irregular short black flecks, and a karyotype of 2 n = 42 a (King 1982). This diagnosis applies to populations of Gehyra genetically assignable to the “ minuta clade ” of Sistrom et al. (2013).	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFD434086C92FD4BFF3653A9.taxon	discussion	Comments. King described his new species from a small number of localities and more recent knowledge has not suggested any broader distribution for this species. We did not have significant sampling of this species and so suggest that until further data prove the contrary, it should be regarded as an endemic inhabitant of the scattered rocky ranges centred around Tennant Creek, Northern Territory. King’s description reveals this species as very short-faced compared to the other species described here, with a reduced labial count (as few as six and not exceeding nine) and relatively small chin shields crowded by the short snout so that the postmentals are often in contact with the second infralabials (a very rare exception among the other species treated here). In colour pattern it is most similar to G. pulingka sp. nov. of the Central Ranges around Northern Territory-South Australia-Western Australia border region. That species is further distinguished from G. minuta in generally having three pairs of enlarged chin shields and the third infralabial notched. Nearby populations of G. moritzi sp. nov. can be distinguished by their finely spotted colour pattern (both black and white markings in the form of small circular spots),	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFD434096C92F9CEFCF35111.taxon	description	Figs 3, 9.	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFD434096C92F9CEFCF35111.taxon	materials_examined	Holotype: SAMA R 65941 K, adult male from Corroboree Rock, East MacDonnell Ranges, Northern Territory (23 ° 44 ’ 23.0 ˮ S, 133 ° 57 ' 02.5 " E), collected by M. Hutchinson, P. Oliver, G. Armstrong and S. South on 9 January 2011 (Fig 9 A – B). Paratypes (n = 17; all genotyped as ‘ Clade 1 ’): MAGNT: R 14356, R 15356, 6 km SSW Claraville HS, NT (23 ° 25 ’ S, 134 ° 44 ’ E), R 18310, Palm Valley Gas Well, Finke Gorge NP, NT (24 ° 01 ’ S, 132 ° 37 ’ E), R 20664, Finke Gorge, NT (24 ° 08 ’ S, 132 ° 49 ’ E). SAMA: R 65935 K – 36, Rainbow Valley, NT, R 65896 – 900, R 65937 K – 38, Emily Gap, NT (23 ° 44 ’ S, 133 ° 57 ’ E), R 65895, Corroboree Rock, NT (23 ° 44 ’ 23 ˮ S, 133 ° 57 ’ 03 ˮ E, R 65881, R 65945 K – 46, 20 km S of Barrow Creek, NT (21 ° 38 ’ S, 133 ° 44 ’ E).	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFD434096C92F9CEFCF35111.taxon	diagnosis	Diagnosis. Distinguished from other Australian Gehyra by a combination of either 7 or 8 divided scansors under the expanded portion of the fourth toe, small to moderate size, generally two pairs of enlarged chin shields, second or third infralabial notched, dorsal colour pattern combining pinkish grey to rufous colouring (in life) patterned entirely by black and whitish spots, and a karyotype of 2 n = 44 (Moritz 1986). Distinguished from most central Australian species (G. versicolor, G. purpurascens and G. montium) by its strongly white-spotted pattern with dark markings also forming discrete spots rather than wavy lines or continuous networks. Distinguished from G. minuta and G. pulingka sp. nov. by having black spots rather than short black wavy lines, normally two rather than three pairs of chin shields (G. pulingka sp. nov.), and higher numbers of labial scales (G. minuta). This diagnosis applies to populations of Gehyra genetically assignable to “ Clade 1 ” of Sistrom et al. (2013).	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFD434096C92F9CEFCF35111.taxon	description	Description. Adult snout-vent length 36 – 49 mm (mean = 42.0 mm, n = 19). Length of original tail 38 – 51 mm (mean = 106 % SVL, n = 5). Nostril bordered by rostral, first supralabial, supranasal and two subequal postnasals. Either a single internasal scale separates the supranasals above the rostral (n = 9) or supranasals in median contact (n = 9). Supralabials 8 – 10 (mode 9). Infralabials 7 – 9 (mode 8). Two, less frequently (6 / 19) three, pairs of chin shields, anterior pair in contact with only the first infralabial. Chin shields separated from the fourth, or third, and succeeding infralabials by the interpolation of a series of enlarged scales (parinfralabials) that margin the ventral edge of the infralabials. Third, less frequently (7 / 19) second, infralabial notched where this parinfralabial scale row starts. Scansors under pad of fourth toe divided, 7 – 8 (mode 8). Precloacal pores in males 11 – 16 (mean = 14.4, n = 11). The karyotype is 2 n = 44, regarded as the plesiomorphic karyotype for the Gehyra variegata species group by King (1979) and Moritz (1986), but see Sistrom et al. (2013) for cautionary remarks on this assumption. In life, dorsally light pinkish grey to reddish brown, the entire dorsal surface patterned by spots. Dark spots are larger and more irregular, pale spots tend to be more precisely circular in shape.	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFD434096C92F9CEFCF35111.taxon	distribution	Distribution. Rocky mountain ranges of the south-central Northern Territory centred on the MacDonnell Ranges and south to the James Range, west to the Kings Canyon area and north to the Devils Marbles.	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFD434096C92F9CEFCF35111.taxon	discussion	Comments. The above description refers to specimens from the central and southern parts of the species' range. The northernmost sample (SAMA R 65875 - 80, 65943 - 44), from rocky hills south of the Devils Marbles, is distinctly different in morphology but is not distinguishable by either DNA sequence data or karyotype (SAMA R 65943: 2 n = 44). This series of eight specimens is consistently smaller (largest specimens only 40 mm SVL), males have fewer precloacal pores (range 8 – 11) and the spotted colour pattern consists of relatively very small spots, each covering only a few dorsal scales. All have seven enlarged scansors rather than the eight usual for the other populations. The small size, spotted pattern and rock dwelling habits of these animals are reminiscent of G. minuta which occurs close by, but that species has a dorsal pattern with larger pale spots and irregular black flecks rather than spots, lower infralabial counts and frequent contact between the postmental and second infralabial. For the present we refer this sample to G. moritzi but exclude it from the paratype series. Further genetic studies of gecko populations along the Northern Territory portion of the Stuart Highway would be useful to clarify the genetic relationships among G. moritzi populations and the pattern of occurrence of the rock dwelling Gehyra of this region. The prominently spotted colour pattern of G. moritzi is very similar to that of some Western Australia rockdwelling populations of Gehyra. They are quite genetically distinct (‘ punctata clade’ of Sistrom et al. 2013), but the species delineation modeling indicates that they are likely sister taxa. Current chromosomal data (King 1979, Moritz 1986) suggests a further distinction (2 n = 44 for G. moritzi, 2 n = 42 a for G. ‘ punctata’), but the precise relationships will require further work on Western Australian populations where multiple species are likely present in the Pilbara and adjacent areas. The strong similarity in colour pattern suggests selection for camouflage against rocks, but given their relationships this could be due to common ancestry rather than convergence.	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFD434096C92F9CEFCF35111.taxon	etymology	Etymology. The specific name recognises the contribution of Dr Craig Moritz (Australian National University, Canberra, and University of California, Berkeley) in revealing the high level of karyotypic and morphological diversity among central Australian populations of Gehyra.	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFCB34176C92FF70FD5751C3.taxon	description	Figs 2 B, 3, 4 B, 10.	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFCB34176C92FF70FD5751C3.taxon	materials_examined	Holotype: SAMA R 65248, from Umuwa, Musgrave Ranges, South Australia (26 ° 28 ’ 45 ˮ S, 133 ° 57 ’ 02 ˮ E), collected by M. Hutchinson, on 26 May 2010, (Fig. 10 A – B). Paratypes (n = 16; all genotyped as ‘ Clade 2 ’): SAMA: R 28265, Kulgera, NT (25 ° 50 ’ S, 133 ° 18 ’ E), R 28322 – 23, between Victory Well and Betty Well, Everard Ranges, SA (27 ° 03 ’ S, 132 ° 28 ’ E), R 41876 – 77, 15 km W Mimili, SA (27 ° 01 ’ E, 132 ° 34 ’ E), R 42069, 29 km SW Illintjitja, SA (26 ° 20 ’ S, 130 ° 10 ’ E), R 44892, 8 km SE Mitchell Knob, SA (26 ° 11 ’ S, 131 ° 53 ’ E), R 50119, 0.9 km SE Sentinel Hill, SA (26 ° 05 ’ S, 132 ° 27 ’ E), R 51536 – 37 K, R 51540 K, R 51565 K, R 51574 K, 35 km ESE Amata, SA (26 ° 15 ’ S, 131 ° 29 ’ E), R 54751, Mt Howe, SA (26 ° 16 ’ S, 133 ° 26 ’ E), R 61926 K, Kurtjurntari Rockhole, WA (24 ° 53 ’ S, 128 ° 46 ’ E). WAM: R 166314, Kurtjurntari Rockhole, WA (24 ° 53 ’ S, 128 ° 46 ’ E).	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFCB34176C92FF70FD5751C3.taxon	diagnosis	Diagnosis. Distinguished from other Australian Gehyra by a combination of modally 7 or 8 divided scansors, small to moderate size, generally three pairs of enlarged chin shields, third infralabial notched, dorsal colour pattern a light to medium brown (in life) patterned by irregular thin black lines and circular pale spots, and a karyotype of 2 n = 42 b (Fig. 2 B) (Moritz 1986). Distinguished from most central Australian species (G. variegata, G. versicolor sp. nov., G. purpurascens and G. montium) by its strongly white-spotted pattern with dark markings forming short, wavy lines rather than discrete spots or continuous networks. Distinguished from G. moritzi by having short, black wavy lines rather than black spots, and from G. minuta, which has a similar colour pattern, by having the third, rather than second, infralabial notched. This diagnosis applies to populations of Gehyra genetically assignable to “ Clade 2 ” of Sistrom et al. (2013).	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFCB34176C92FF70FD5751C3.taxon	description	Description. Adult snout-vent length 38 – 49 mm (mean = 43.3 mm, n = 14). Length of original tail 43 – 56 mm (mean = 117 % SVL, n = 6). Nostril bordered by rostral, first supralabial, supranasal and two subequal postnasals. Usually a single internasal scale (occasionally two or none) separates the supranasals above the rostral. Supralabials 7 – 10 (mode 8). Infralabials 7 – 9 (mode 8). Three pairs of chin shields, outer (third) pair small (absent in three specimens), anterior pair in contact with only the first infralabial. Chin shields separated from the fourth and succeeding infralabials by the interpolation of a series of enlarged scales that margin the ventral edge of the infralabials. Third infralabial notched (all specimens) where this parinfralabial scale row starts (Fig. 4 B). Scansors under pad of fourth toe divided, 7 – 8 (mode 8). Precloacal pores in males 12 – 16 (mean = 13.9, n = 7). The karyotype (2 n = 42 b) is unique for this species, differing from the 2 n = 42 a karyotype via a secondary constriction on pair 11 (Moritz 1986). In life, dorsally pinkish grey to reddish brown, the entire dorsal surface patterned by light spots that are generally strongly contrasting and precisely circular in shape, sometimes forming rather regular transverse series. The pale spots are interspersed with short dark irregular lines, which are more elongate and irregular than the pale spots.	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFCB34176C92FF70FD5751C3.taxon	distribution	Distribution. Rocky mountain ranges of north-western South Australia, extending into adjacent areas of Western Australia (northwest to the eastern Rawlinson Range) and far southern Northern Territory (Kulgera area).	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFCB34176C92FF70FD5751C3.taxon	discussion	Comments. Long included in G. montium, G. pulingka is consistently distinguishable in morphology, karyotype and DNA sequence data. The two species show overlap in distribution, although no cases of strict syntopy are known as yet. In the field, the colour pattern of blackish squiggles and prominent spots can be used to distinguish this species from true G. montium, which has a more continuous black dorsal network and small, weakly contrasting spots. Additional distinctions in chin shields (three versus two pairs), having the third, rather than second, infralabial notched and higher male precloacal pore counts will provide extra support if genetic data are lacking.	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
AE44F266FFCB34176C92FF70FD5751C3.taxon	etymology	Etymology. The specific name is from the Pitjantjatjara language (Goddard 1996) from the roots puli, rock, or rocky hill, and the suffix - ngka meaning pertaining to, alluding to the habits of the species and its distribution, which is confined to the desert areas occupied by the speakers of Pitjantjatjara and related dialects. Specific name would not change with gender of the genus.	en	Hutchinson, Mark N., Sistrom, Mark J., Donnellan, Stephen C., Hutchinson, Rhonda G. (2014): Taxonomic revision of the Australian arid zone lizards Gehyra variegata and G. montium (Squamata, Gekkonidae) with description of three new species. Zootaxa 3814 (2): 221-241, DOI: http://dx.doi.org/10.11646/zootaxa.3814.2.4
