Zwicknia turkestanica turkestanica (Kimmins, 1950) comb. n.

(Figs. 102–105, 113, 126, 147, 164, 196)

Capnia nigra (Pictet, 1833) — Morton 1896: 60. (Fig. 3 on Plate 2 refers to the holotype of Z. turkestanica).

Capnia turkestanica Kimmins, 1950 — Kimmins 1950a: 187. (original description of the adult); Zhiltzova 1969: 603. (complementary description and comparison with Z. tuberculata); Zhiltzova 2003: 349. (complementary description).

Diagnosis. Male epiproct: Ep-scl narrow and acute in dorsal view, short, tip slightly downcurved in lateral view, membranous portion long; ventral membranous section reaching base in lateral view, apical spines long, distributed also on the apex of Ep-scl. Process of male Tg 9: high, perpendicularly elevated, <2× wider than Ep-scl, rounded and with continuously narrow sides caudally.

Material examined. KAZAKHSTAN: East Kazakhstan Province, Tarbagatai Mts., River Ayagos 270km SW of Öskemen (Ust-Kamenogorsk), 05.04.200 3, leg., det. V. I. Devyatkov: 6m 6f (PZC; used for drawings Figs. 102–105, 126, 147, 164; one male terminalia prepared for SEM to Fig. 113) .

Description. Head, thorax, appendages and basal segments of the abdomen generotypic. Males micropterous or macropterous, females macropterous. Dimensions of the presently examined specimens: body length: males 5.5–6.5, females 6.0–7.5 mm; forewing length: males 1.0–6.0 mm, females 6.5–7.5 mm.

Male terminalia (Figs. 102–105, 126): Process of Tg 9 high, perpendicularly elevated, apex is <2× wider than the medial section of Ep-scl; its shape is rounded, the apex smooth; sides continuously narrow in caudal view, the membranous portion narrowest apically (Fig. 164). Tg 10, B-scl and Lb-scl generotypic. Ep-scl narrow and acute in dorsal view, short, medially not swollen, its medial width is ~½ of basal width; tip slightly downcurved in lateral view, divided section long. Ventral membranous portion between the division of Ep-scl reaching base in lateral view; apical spines moderatelly long, extending to the Ep-scl (Figs. 113, 147). I-scl generotypic, Ec long and contorted on the available specimens. St 9 not projecting medially, vesicle vestigial. Sg wide and rounded not as pronounced triangular shape, tip rounded. Pp, Fp, Rp and cerci generotypic.

Female subgenital plate: Rectangular, posterior margin rounded, equal to the segment`s posterior margin. Antero-lateral recess distinct, the plate brown, sometimes with paler lateral areas; lateral sclerites small (see Zhiltzova 2003: Fig. 585).

Affinities. The relationship of Z. turkestanica with closely related Z. tuberculata and Z. sevanica is discussed above. Males are easily distinguishable from all other Zwicknia species by high, but rounded process of Tg 9. Females are difficult to distinguish from other species of the genus. The larva is unknown.

Distribution and ecology. The species was described from a single male from`Turkestan` without exact locality. It is known from Tajikistan, Kyrgyzstan and Kazakhstan (Fig. 196). Adults apparently are active from March to April associated with mountain streams as high as 2,400 m.