taxonID	type	description	language	source
F535879E7E61FFC6FF0DFEADD4A0FA18.taxon	description	(Figs. 2 A – D, 3 A – B, 9 A – B, 10 A – B, 23 A)	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E61FFC6FF0DFEADD4A0FA18.taxon	materials_examined	Type series. Holotype male, dissected and slide mounted, Malaysia, Borneo, stream Kemantis, Sayap, Mt Kinabalu, 6 º 09.841 N, 116 º 33.936 E, alt. 928 m asl., 16. ix. 2012 Smit. Paratypes: two females, same data as holotype, one female dissected and slide mounted. Further records. Malaysia, Borneo, Mahua stream, Mahua, Crocker Range, 5 º 47.838 N, 116 º 24.510 E, alt. 1052 m asl., 21. ix. 2012 Smit 1 / 0 / 0; unnamed stream Bansadon Trail, Inobong, Crocker Range, 5 º 51.456 N, 116 º 68.403 E, 18. ix. 2012 Smit 1 / 0 / 0; Kipungit River, Poring Hot Springs, Mt. Kinabalu, 6 º 02.776 N, 116 º 41.432 E, 568 m asl., 15. ix. 2012 Smit 2 / 1 / 0; Kibamabangan River, Crocker Range, 5 º 51.28 N, 116 º 08.417 E, 433 m asl., 18. ix. 2012 Smit 1 / 0 / 0.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E61FFC6FF0DFEADD4A0FA18.taxon	diagnosis	Diagnosis. Idiosoma roundish (dorsal shield L / W ratio 1.3); shoulder platelets fused to the large dorsal plate; Cxgl – 4 far posterior at margin of Cx-I / II, near I – L insertion; P- 2 with a laterally compressed, anteriorly directed ventrodistal extension covering less than 30 % of ventral margin.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E61FFC6FF0DFEADD4A0FA18.taxon	description	Description General features — Idiosoma roundish; shoulder platelets fused to the large dorsal plate; dorsal shield with colour pattern as illustrated in Figs. 9 A – B; gnathosomal bay U-shaped, proximally rounded; Cxgl – 4 far posterior at margin of Cx-I / II, near I – L insertion; suture lines of Cx-IV extending posteriorly beyond posterior margin of genital field, laterally curved; excretory pore away from the line of primary sclerotization, Vgl – 2 on the same level or slightly posterior from excretory pore; gnathosomal rostrum long and slender, ventral margin in lateral view curved (Fig. 2 D); P- 2 slightly shorter than P- 4, P- 2 ventral margin convex, distally with a laterally compressed, anteriorly directed and apically serrated hyaline extension (covering less than 30 % of ventral margin) and a very short, denticle-like seta laterally at base of projection; P- 3 with a shorter, subrectangular, apically serrated ventrodistal projection, and a long seta laterally at base of projection; P- 4 slender, with ventral tubercles pointed and separated, bearing one long and three short setae (Figs. 2 C, 3 B). Male: medial suture line of Cx-II + III relatively long; genital field subrectangular, ejaculatory complex with small proximal chamber (Fig. 23 A). Female: genital field pentagonal in shape. Measurements Male (holotype) — Idiosoma (ventral view: Figs. 2 B, 10 A) L 666, W 469; dorsal shield (Figs. 2 A, 9 A) L 525, W 413, L / W ratio 1.27; dorsal plate L 490; frontal plate L 129 – 131, W 45 – 46, L / W ratio 2.8 – 2.9. Gnathosomal bay L 128, Cx-I total L 250, Cx-I mL 122, Cx-II + III mL 120; ratio Cx-I L / Cx-II + III mL 2.1; Cx-I mL / Cx-II + III mL 1.02. Genital field L / W 123 / 97, ratio 1.27; distance genital field-excretory pore 122, genital field-caudal idiosoma margin 171. Gnathosoma vL 302; chelicera total L 334; palp total L 287, dL / H, dL / H ratio 287: P- 1, 29 / 25, 1.18; P 2, 92 / 55, 1.69; P- 3, 57 / 45, 1.26; P- 4, 94 / 24, 3.86; P- 5, 15 / 11, 1.43; P- 2 / P- 4 ratio 0.98. Female (paratype) — Idiosoma (ventral view: Figs. 3 A, 10 B) L 720, W 506; dorsal shield (Fig. 9 B) L 594, W 450, L / W ratio 1.32; dorsal plate L 556; frontal plate L 148 – 151, W 49 – 50, L / W ratio 3.03. Gnathosomal bay L 125, Cx-I total L 250, Cx-I mL 108, Cx-II + III mL 108; ratio Cx-I L / Cx-II + III mL 2.3; Cx-I mL / Cx-II + III mL 1.0. Genital field L / W 134 / 128, ratio 1.05; distance genital field-excretory pore 166, genital field-caudal idiosoma margin 266. Gnathosoma vL 335; chelicera total L 368; palp total L 311, dL / H, dL / H ratio: P- 1, 35 / 27, 1.32; P- 2, 100 / 62, 1.61; P- 3, 55 / 49, 1.13; P- 4, 106 / 24, 4.37; P- 5, 15 / 11, 1.43; P- 2 / P- 4 ratio 0.94.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E61FFC6FF0DFEADD4A0FA18.taxon	etymology	Etymology. Named after the island where the new species was detected.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E61FFC6FF0DFEADD4A0FA18.taxon	discussion	Discussion. Torrenticola borneoensis n. sp belongs to the former subgenus Rusetria Thor, 1897, characterized by the fusion of the shoulder platelets with the large dorsal plate. Together with Torrenticola indica Cook, 1967 (India; Cook 1967), T. flangia Wiles, 1997 (Sulawesi; Wiles 1997, Pešić & Smit 2011), T. kinabaluensis n. sp. (see below) and T. neoindica n. sp. (see below), T. borneoensis n. sp. belongs to a group characterized by a flanged palp (P- 2 with a laterally compressed and apically serrated ventrodistal extension). The combination of a flanged palp with Cxgl- 4 shifted close to I-L insertion makes the new species similar to T. kinabaluensis n. sp. (see below). The latter species can easily be distinguished by a more elongated dorsal shield and the flange on P- 2 more longish (> 30 % of ventral margin of P- 2). Further, Cxgl- 4 is shifted more posteriorly in T. kinabaluensis n. sp., lying between I- and – II – L insertions, but more approaching I – L. Torrenticola flangia resembles T. kinabaluensis n. sp. in the shape of the palp (flange on P- 2 longish,> 30 % of ventral margin) but clearly differs from both aforementioned new species from Borneo in a characteristic colour pattern on the dorsal shield (typically with a broad posterior and a narrow anterior band of blue pigment) and Cxgl – 4 more posterior, lying in close proximity to Cxgl – 2 (see Wiles 1997). Habitat. Sandy / bouldery streams, shaded by rain forest (Figs. 43 A – D).	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E61FFC6FF0DFEADD4A0FA18.taxon	distribution	Distribution. Borneo.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E63FFCBFF0DF998D4A0FD35.taxon	description	(Figs. 4 A – E, 5 A – B, 9 C – D, 10 C – E, 23 B)	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E63FFCBFF0DF998D4A0FD35.taxon	materials_examined	Type series. Holotype male, dissected and slide mounted, Malaysia, Borneo, Mahua stream, downstream of national park entrance, Crocker Range, 5 º 46.491 N, 116 º 25.770 E, alt. 865 m asl., 22. ix. 2012 Smit. Paratypes: one male, same data as holotype; two males, one female, Little Lumotok stream, Sayap, Mt Kinabalu, 6 º 09.497 N, 116 º 34.027 E, alt. 1065 m asl., 17. ix. 2012 Smit, one male and one female dissected and slide mounted; one male, Mahua river, upstream, 5 º 47.939 N, 116 º 24.317 E, 1050 m asl., 21. ix. 2012 Smit. Further records. Malaysia, Borneo: Great Lumotok stream, Mt Kinabalu, 6 º 09.336 N, 116 º 08.417 E, alt. 433 m asl., 18. ix. 2012 Smit 0 / 1 / 0; unnamed stream Bansadon Trail, Inobong, Crocker Range, 5 º 51.456 N, 116 º 68.403 E, 18. ix. 2012 Smit 0 / 1 / 0; Kipungit River, Poring Hot Springs, Mt. Kinabalu, 6 º 02.776 N, 116 º 41.432 E, 568 m asl., 15. ix. 2012 Smit 1 / 0 / 0; Kibamabangan River, Crocker Range, 5 º 51.28 N, 116 º 08.417 E, 433 m asl., 18. ix. 2012 Smit 0 / 2 [one juvenile] / 0.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E63FFCBFF0DF998D4A0FD35.taxon	diagnosis	Diagnosis. Idiosoma elongated (dorsal shield L / W ratio 1.5); shoulder platelets fused to the large dorsal plate; Cxgl – 4 far posterior at margin of Cx-I / II, between I – L and II – L insertions, but closer to I – L insertion; P- 2 with a laterally compressed, longish (> 30 % of ventral margin), anteriorly directed ventrodistal extension.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E63FFCBFF0DF998D4A0FD35.taxon	description	Description General features — Idiosoma elongated; shoulder platelets fused to the large dorsal plate; dorsal shield with colour pattern as illustrated in Figs. 9 C – D; gnathosomal bay U-shaped, proximally rounded; Cxgl – 4 far posterior at margin of Cx-I / II, between I – L and II – L insertions, but closer to I – L insertion; suture lines of Cx-IV extending posteriorly beyond posterior margin of genital field, laterally curved; excretory and Vgl- 2 pore away from the line of primary sclerotization, excretory pore slightly anterior, or on the level with Vgl- 2; gnathosomal rostrum long, ventral margin in lateral view curved (Fig. 4 E); P- 2 shorter than P- 4 (L P- 2 / P- 4 ratio 0.9), ventral margin of P- 2 distally with a laterally compressed, longish (> 30 % of ventral margin), anteriorly directed and apically serrated hyaline extension and a very short, denticle-like seta laterally at base of projection; P- 3 with a shorter, subrectangular, apically serrated ventrodistal projection, and a moderately long seta laterally at base of projection; P- 4 slender, with ventral tubercles pointed and separated, bearing one long and three short setae (Figs. 4 C – D). Male: medial suture line of Cx-II + III relatively long; genital field subrectangular; ejaculatory complex conventional in shape (Fig. 23 B). Female: genital field pentagonal in shape. Measurements Male (holotype) — Idiosoma (ventral view: Figs. 4 B) L 600, W 387; dorsal shield (Figs. 4 A, 9 C) L 492, W 336, L / W ratio 1.46; dorsal plate L 457; frontal plate L 119, W 41 – 43, L / W ratio 2.8 – 2.9. Gnathosomal bay L 108, Cx-I total L 214, Cx-I mL 106, Cx-II + III mL 109; ratio Cx-I L / Cx-II + III mL 2.0; Cx-I mL / Cx-II + III mL 0.97. Genital field L / W 103 / 87, ratio 1.19; distance genital field-excretory pore 128, genital field-caudal idiosoma margin 172. Gnathosoma vL 281; chelicera total L 319; palp total L 252, dL / H, dL / H ratio: P- 1, 28 / 23, 1.2; P- 2, 76 / 42, 1.8; P- 3, 48 / 38, 1.27; P- 4, 85 / 20, 4.2; P- 5, 15 / 11, 1.35; P- 2 / P- 4 ratio 0.90. Female (paratype from Little Lumotok stream) — Idiosoma (ventral view: Figs. 5 A, 10 E) L 703, W 418; dorsal shield (Fig. 9 D) L 553, W 364, L / W ratio 1.52; dorsal plate L 510; frontal plate L 131 – 137, W 45 – 47, L / W ratio 2.9. Gnathosomal bay L 151, Cx-I total L 270, Cx-I mL 119, Cx-II + III mL 103; ratio Cx-I L / Cx-II + III mL 2.62; Cx-I mL / Cx-II + III mL 1.16. Genital field L / W 136 / 114, ratio 1.19; distance genital field-excretory pore 143, genital field-caudal idiosoma margin 192. Gnathosoma vL 334; chelicera total L 378; palp total L 282, dL / H, dL / H ratio: P- 1, 32 / 28, 1.17; P- 2, 90 / 52, 1.73; P- 3, 51 / 42, 1.21; P- 4, 94 / 23, 4.1; P- 5, 15 / 12, 1.25; P- 2 / P- 4 ratio 0.96.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E63FFCBFF0DF998D4A0FD35.taxon	etymology	Etymology. Named after the mountain (Kinabalu) where the new species was found.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E63FFCBFF0DF998D4A0FD35.taxon	discussion	Discussion. See discussion under T. borneoensis. Habitat. Sandy / bouldery streams, shaded by rain forest (Figs. 43 A, C).	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E63FFCBFF0DF998D4A0FD35.taxon	distribution	Distribution. Borneo.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E6EFFCDFF0DFC95D4A0F9ED.taxon	description	(Figs. 6 A – E, 7 A – B, 9 G – I, 10 G – I, 23 D)	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E6EFFCDFF0DFC95D4A0F9ED.taxon	materials_examined	Type series. Holotype male, dissected and slide mounted, Malaysia, Borneo, Great Lumotok stream, Mt Kinabalu, 6 º 09.336 N, 116 º 08.417 E, alt. 433 m asl., 18. ix. 2012 Smit. Paratypes: one juvenile male, one female, same data as holotype, female dissected and slide mounted. Further records. Malaysia, Borneo: Little Lumotok stream, Sayap, 6 º 09.497 N, 116 º 34.027 E, alt. 1065 m asl., 17. ix. 2012 Smit 6 / 1 / 0 (1 / 0 / 0 mounted); Mahua stream, Mahua, Crocker Range, 5 º 47.838 N, 116 º 24.510 E, alt. 1052 m asl., 21. ix. 2012 Smit 1 / 1 / 0; first stream Minduk Sirung Trail, Alab, Crocker Range, 5 º 45.225 N, 116 º 26.085 E, alt. 752 m asl., 22. ix. 2012 Smit 0 / 1 / 0.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E6EFFCDFF0DFC95D4A0F9ED.taxon	diagnosis	Diagnosis. Idiosoma elongated-oval (dorsal shield L / W ratio 1.3 – 1.4); shoulder platelets fused to the large dorsal plate; dorsal shield without colour pattern; Cxgl – 4 anteriorly adjacent to Cxgl – 2, posterior at margin of CxI / II; suture lines of Cx-IV extending posteriorly beyond posterior margin of genital field; excretory pore on the level with Vgl – 2. Male: medial suture line of Cx-II + III moderately long.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E6EFFCDFF0DFC95D4A0F9ED.taxon	description	Description General features — Idiosoma elongated-oval; shoulder platelets fused to the large dorsal plate; dorsal shield without colour pattern (Figs. 9 G – I); gnathosomal bay U-shaped; Cxgl – 4 anteriorly adjacent to Cxgl – 2, posterior at margin of Cx-I / II; suture lines of Cx-IV extending posteriorly beyond posterior margin of genital field, laterally curved; excretory and Vgl – 2 pore away from the line of primary sclerotization, excretory pore on the level with Vgl – 2; gnathosoma with curved ventral margin (Fig. 6 E); P- 2 nearly equal in length as P- 4, P- 2 ventral margin concave, P- 2 ventrodistal protrusion bluntly pointed, curved towards distal; P- 3 ventrodistal protrusion pointed and cone-shaped; P- 4 with well developed ventral protuberances, ending in two tips separated by a concavity (Figs. 6 D, 7 B). Male: medial suture line of Cx-II + III moderately long; genital field subrectangular in shape; ejaculatory complex conventional in shape (Figs. 6 C, 23 D). Female: genital field pentagonal in shape. Measurements Male (holotype, in parentheses specimen from Little Lumotok stream) — Idiosoma (ventral view: Figs. 6 B, 10 G – H) L 678 (672), W 481 (478); dorsal shield (Figs. 6 A, 9 G – H) L 556 (539), W 405 (411), L / W ratio 1.37 (1.31); dorsal plate L 519 (503); frontal plate L 143 (144), W 50 (50 – 53), L / W ratio 2.9 (2.7 – 2.9). Gnathosomal bay L 131 (148), Cx-I total L 250 (272), Cx-I mL 119 (122), Cx-II + III mL 106 (94); ratio Cx-I L / Cx-II + III mL 2.36 (2.9); Cx-I mL / Cx-II + III mL 1.12 (1.3). Genital field L / W 130 (131) / 107 (109), ratio 1.2 (1.2); ejaculatory complex L 162 (149); distance genital field-excretory pore 125 (109), genital field-caudal idiosoma margin 192 (173). Gnathosoma vL 303 (306); chelicera total L 378 (351); palp total L 300 – 301 (302), dL / H, dL / H ratio: P- 1, 37 / 31, 1.2 (44 / 34, 1.29); P- 2, 92 / 48, 1.94 (92 / 48, 1.92); P- 3, 59 – 60 / 42, 1.42 (57 / 45, 1.28); P- 4, 94 / 30, 3.1 (89 / 31, 2.9); P- 5, 18 / 16, 1.1 (20 / 15, 1.3); P- 2 / P- 4 ratio 0.98 (1.03). Female (paratype from Great Lumotok stream) — Idiosoma (ventral view: Figs. 7 A, 10 I) L 755, W 559; dorsal shield (Fig. 9 I) L 616, W 469, L / W ratio 1.31; dorsal plate L 575; frontal plate L 152 – 155, W 56, L / W ratio 2.7 – 2.8. Gnathosomal bay L 172, Cx-I total L 305, Cx-I mL 133, Cx-II + III mL 73; ratio Cx-I L / Cx-II + III mL 4.2; Cx-I mL / Cx-II + III mL 1.8. Genital field L / W 156 / 146, ratio 1.07; distance genital field-excretory pore 144, genital field-caudal idiosoma margin 220. Gnathosoma vL 363; chelicera total L 450; palp total L 351 – 352, dL / H, dL / H ratio: P- 1, 51 / 38, 1.3; P- 2, 111 / 57, 1.95; P- 3, 68 - 69 / 48, 1.44; P- 4, 103 / 32, 3.2; P- 5, 18 / 17, 1.06; P- 2 / P- 4 ratio 1.07.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E6EFFCDFF0DFC95D4A0F9ED.taxon	etymology	Etymology. Named after the province (Sabah) where the new species was found.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E6EFFCDFF0DFC95D4A0F9ED.taxon	discussion	Discussion. The new species is most similar to Torrenticola longipalpis Wiles, 1997 (see below). The latter species can easily be separated from T. sabahensis n. sp. due to the less slender idiosoma, the characteristic colour pattern (see Figs. 9 E – F), Cxgl – 4 shifted more anteriorly, at margin of Cx-I / II, the longer ventral seta on P- 2, and in males, medial suture line of Cx-II + III shorter and suture lines of Cx-IV extending less posteriorly beyond posterior margin of genital field. Habitat. Sandy / bouldery streams, shaded by rain forest (Fig. 43 A).	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E6EFFCDFF0DFC95D4A0F9ED.taxon	distribution	Distribution. Borneo.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E68FFD1FF0DF9EED1DDFDE6.taxon	description	(Figs. 8 A – D, 9 E – F, 10 F, 23 C)	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E68FFD1FF0DF9EED1DDFDE6.taxon	description	Morphology Male. General features — Idiosoma roundish (dorsal shield L / W ratio 1.2); shoulder platelets fused to the large dorsal plate; dorsal shield with colour pattern as illustrated in Figs. 9 E – F; gnathosomal bay U-shaped, proximally pointed; Cxgl – 4 far posterior at margin of Cx-I / II, near insertion of II-L; medial suture line of Cx-II + III short; suture line of Cx-IV distinct, medially starting from posterior margin of genital field in a right angle to the main idiosoma axis; genital field subrectangular in shape; ejaculatory complex conventional in shape (with well developed anterior keel and proximal arms — Figs. 8 D, 23 C); excretory and Vgl – 2 pore away from the line of primary sclerotization, excretory pore anterior to Vgl – 2; gnathosoma with curved ventral margin; P- 2 longer than P- 4, P- 2 ventral margin straight, P- 2 and P- 3 ventrodistal protrusions bluntly pointed and cone-shaped, P- 4 with well developed ventral protuberances, ending in two tips separated by a concavity (Fig. 8 C). Measurements — Idiosoma (ventral view: Figs. 8 B, 10 F) L 713, W 556; dorsal shield (Figs. 8 A, 9 E – F) L 584, W 485, L / W ratio 1.2; dorsal plate L 558; frontal plate L 130 – 141, W 47, L / W ratio 2.8 – 3.0. Gnathosomal bay L 167, Cx-I total L 320, Cx-I mL 153, Cx-II + III mL 56; ratio Cx-I L / Cx-II + III mL 5.7; Cx-I mL / Cx-II + III mL 2.7. Genital field L / W 159 / 128, ratio 1.25; ejaculatory complex L 223; distance genital field-excretory pore 119, genital field-caudal idiosoma margin 173. Gnathosoma vL 364; chelicera total L 402; palp total L 335, dL / H, dL / H ratio: P- 1, 38 - 39 / 36, 1.07; P- 2, 115 / 65, 1.79; P- 3, 59 / 55, 1.08; P- 4, 100 / 37, 2.7; P- 5, 23 / 19, 1.22; P- 2 / P- 4 ratio 1.15.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E68FFD1FF0DF9EED1DDFDE6.taxon	discussion	Remarks. Wiles (1999) reported T. longipalpis from several sites in the catchment area of the Rivers Temburong and Belalong in North Brunei. He mentioned that his specimens differ from those from Sulawesi in having Cxgl – 4 closer to, but still anterior to Cxgl – 2 and a slightly longer medial suture line of Cx-II + III. However, he considered that to be a case of intraspecific variability.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E68FFD1FF0DF9EED1DDFDE6.taxon	distribution	Distribution. Sulawesi (Wiles 1997, Pešić & Smit 2011), Borneo (Wiles 1999, our study).	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E74FFD1FF0DFDD9D6BCF84C.taxon	description	(Figs. 11 A – D, 16 A – B, 17 A – B, Fig. 23 E)	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E74FFD1FF0DFDD9D6BCF84C.taxon	description	Morphology General features — Idiosoma roundish (dorsal shield L / W ratio 1.2); shoulder platelets fused to the large dorsal plate; dorsal shield with colour pattern as illustrated in Figs. 16 A – B; gnathosomal bay U-shaped; Cxgl – 4 subapical, only slightly posterior of Cx-I tips; suture line of Cx-IV distinct, extending posteriorly beyond posterior margin of genital field; excretory and Vgl – 2 pore away from the line of primary sclerotization, excretory pore on the level with Vgl – 2; gnathosoma with curved ventral margin; P- 2 longer than P- 4, P- 2 ventral margin slightly convex, P- 2 ventrodistal protrusion, bluntly pointed, curved towards distal; P- 3 ventrodistal protrusion bluntly pointed; P- 4 with well developed ventral protuberances, ending in two tips separated by a concavity (Figs. 11 C – D). Male: medial suture line of Cx-II + III short; genital field subrectangular in shape; ejaculatory complex conventional in shape (Fig. 23 E). Female: genital field pentagonal in shape. Measurements Male (from stream Kemantis) — Idiosoma (ventral view: Figs. 11 A, 17 A) L 628, W 459; dorsal shield (Fig. 16 A) L 511, W 413, L / W ratio 1.24; dorsal plate L 481; frontal plate L 127 – 128, W 44, L / W ratio 2.9. Gnathosomal bay L 125, Cx-I total L 266, Cx-I mL 142, Cx-II + III mL 58; ratio Cx-I L / Cx-II + III mL 4.6; Cx-I mL / Cx-II + III mL 2.5. Genital field L / W 119 / 100, ratio 1.19; ejaculatory complex L; distance genital field-excretory pore 114, genital field-caudal idiosoma margin 183. Gnathosoma vL 293; chelicera total L 339; palp total L 289 – 291, dL / H, dL / H ratio: P- 1, 31 / 31, 1.0; P- 2, 101 - 102 / 55, 1.86; P- 3, 52 / 46, 1.13; P- 4, 88 - 89 / 28, 3.2; P- 5, 17 / 9, 1.85; P- 2 / P- 4 ratio 1.15. Female (from Great Lumotok stream) — Idiosoma (ventral view: Figs. 11 B, 17 B) L 625, W 459; dorsal shield (Fig. 16 B) L 497, W 412, L / W ratio 1.2; dorsal plate L 469; frontal plate L 128, W 41, L / W ratio 3.2. Gnathosomal bay L 130, Cx-I total L 266, Cx-I mL 136, Cx-II + III mL 37.5; ratio Cx-I L / Cx-II + III mL 7.1; Cx-I mL / Cx-II + III mL 3.6. Genital field L / W 129 / 114, ratio 1.13; distance genital field-excretory pore 113, genital field-caudal idiosoma margin 191. Gnathosoma vL 314; chelicera total L 358; palp total L 296 – 297, dL / H, dL / H ratio: P- 1, 34 / 29, 1.16; P- 2, 101 - 102 / 57, 1.78; P- 3, 54 / 48, 1.13; P- 4, 90 / 28, 3.25; P- 5, 17 / 10, 1.7; P- 2 / P- 4 ratio 1.13.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E74FFD1FF0DFDD9D6BCF84C.taxon	discussion	Remarks. Wiles (1999) reported T. minuta from several sites in the catchment area of the River Temburong in North Brunei. He mentioned that his specimens differ from those from Peninsular Malaysia in having Cxgl- 4 more posteriorly located, near I-L insertion, instead of being subapical (located just posterior of Cx-I tips).	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E74FFD1FF0DFDD9D6BCF84C.taxon	distribution	Distribution. Sulawesi (Wiles 1997), Borneo (Wiles 1999, our study).	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E77FFD4FF0DFF2DD4A0FEE5.taxon	description	(Figs. 12 A – D, 13 A – D, 16 C – D, 17 C – D, 23 F)	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E77FFD4FF0DFF2DD4A0FEE5.taxon	materials_examined	Type series. Holotype male, dissected and slide mounted, Malaysia, Borneo, Mahua stream, downstream of national park entrance, Crocker Range, 5 º 46.491 N, 116 º 25.770 E, alt. 865 m asl., 22. ix. 2012 Smit. Paratypes: 3 / 4 [one juvenile] / 0 (+ 5 / 0 in ethanol), same data as holotype, one female dissected and slide mounted. Further records. Borneo: Great Lumotok stream, Mt Kinabalu, 6 º 09.336 N, 116 º 08.417 E, alt. 433 m asl., 18. ix. 2012 Smit 2 / 1 [one juvenile] / 0 (1 / 0 / 0 mounted); Mahua stream, Mahua, Crocker Range, 5 º 47.838 N, 116 º 24.510 E, alt. 1052 m asl., 21. ix. 2012 Smit 0 / 3 / 1 (+ 4 / 0 in ethanol); Mahua stream at crossing with main road, Crocker Range, 5 º 45.225 N, 116 º 26.085 E, alt. 752 m asl., 22. ix. 2012 Smit 3 [one juvenile] / 0 / 1; Kibamabangan River, Crocker Range, 5 º 51.28 N, 116 º 08.417 E, alt. 433 m asl., 18. ix. 2012 Smit 1 [juvenile] / 0 / 0 (mounted); Little Lumotok stream, Sayap, 6 º 09.497 N, 116 º 34.027 E, alt. 1065 m asl., 17. ix. 2012 Smit 0 / 1 / 0; small stream waterfall trail Inobong, 5 º 51.306 N, 116 º 08.292 E, alt. 482 m asl., 19. ix. 2012 Smit 1 / 0 / 0; small stream Layang Layang, Mt Kinabalu, 6 º 02.711 N, 116 º 33.627 E, alt. 2697 m asl., 12. ix. 2012 Smit 0 / 1 / [juvenile] 0; first stream Minduk Sirung Trail, Alab, Crocker Range, 5 º 45.225 N, 116 º 26.085 E, alt. 752 m asl., 22. ix. 2012 Smit 4 [one juvenile] / 0 / 0; SilauSilau stream, upstream, Mt Kinabalu, 6 º 00.681 N, 116 º 32.417 E, alt. 1592 m asl., 2. ix. 2012 Smit 1 / 1 / 0 (damaged).	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E77FFD4FF0DFF2DD4A0FEE5.taxon	diagnosis	Diagnosis. Idiosoma elongated-oval (dorsal shield L / W ratio 1.2 – 1.4); shoulder platelets fused to the large dorsal plate; dorsal shield with colour pattern as illustrated in Figs. 16 C – D; Cxgl- 4 subapical, only slightly posterior of Cx-I tips; suture lines of Cx-IV extending posteriorly beyond posterior margin of genital field. Male: medial suture line of Cx-II + III short.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E77FFD4FF0DFF2DD4A0FEE5.taxon	description	Description General features — Idiosoma elongated-oval; shoulder platelets fused to the large dorsal plate; dorsal shield with colour pattern as illustrated in Figs. 16 C – D; gnathosomal bay U-shaped; Cxgl- 4 subapical, only slightly posterior of Cx-I tips; suture line of Cx-IV distinct, extending posteriorly beyond posterior margin of genital field; excretory and Vgl- 2 pore away from the line of primary sclerotization, excretory pore slightly anterior or on the level with Vgl- 2; gnathosoma with curved ventral margin; rostrum well developed (Fig. 12 D); P- 2 longer than P- 4, P- 2 ventral margin slightly convex, distally with a subrectangular, anteriorly directed and apically serrated hyaline extension (covering less than 30 % of ventral margin) and a short seta laterally at base of projection; P- 3 with a shorter, subrectangular, apically serrated ventrodistal projection, and a long seta laterally at base of projection; P- 4 stout, with ventral tubercles pointed and separated, bearing one long and three short setae (Figs. 12 C, 13 C – D). Male: medial suture line of Cx-II + III short; genital field subrectangular in shape; ejaculatory complex normal in shape (Fig. 23 F). Female: genital field pentagonal in shape. Measurements Male (holotype, in parentheses from Great Lumotok stream) — Idiosoma (ventral view: Figs. 12 A, 13 B, 17 C) L 563 (596), W 411 (469); dorsal shield (Fig. 13 A, 16 C) L 477 (500), W 360 (404), L / W ratio 1.32 (1.24); dorsal plate L 453 (469); frontal plate L 105 – 111 (116 – 117), W 37 – 38 (44), L / W ratio 2.8 – 3.0 (2.6 – 2.7). Gnathosomal bay L 109 (109), Cx-I total L 231 (232), Cx-I mL 121 (123), Cx-II + III mL 62 (69); ratio Cx-I L / Cx-II + III mL 3.7 (3.4); Cx-I mL / Cx-II + III mL 2.0 (1.8). Genital field L / W 119 (130) / 94 (98), ratio 1.27 (1.32); ejaculatory complex L 143 (177); distance genital field-excretory pore 116 (103), genital field-caudal idiosoma margin 150 (163). Gnathosoma vL 265 (305); chelicera total L 297 (328); palp total L 250 (271), dL / H, dL / H ratio: P- 1, 28 / 26, 1.08 (32 / 29, 1.1); P- 2, 86 / 50, 1.72 (95 / 48 - 49, 1.95); P- 3, 46 / 45, 1.03 (48 / 45, 1.06); P- 4, 72 / 26, 2.73 (80 / 25, 3.28); P- 5, 18 / 12, 1.46 (16 / 11, 1.48); P- 2 / P- 4 ratio 1.2 (1.18). Female (paratype from Mahua stream, downstream of national park entrance) — Idiosoma (ventral view: Figs. 12 B, 17 D) L 609, W 411; dorsal shield (Fig. 16 D) L 508, W 350, L / W ratio 1.45; dorsal plate L 481; frontal plate L 115, W 40, L / W ratio 2.9 – 3.1. Gnathosomal bay L 113, Cx-I total L 229, Cx-I mL 116, Cx-II + III mL 55; ratio CxI L / Cx-II + III mL 4.2; Cx-I mL / Cx-II + III mL 2.1. Genital field L / W 134 / 116, ratio 1.16; distance genital fieldexcretory pore 122, genital field-caudal idiosoma margin 191. Gnathosoma vL 281; palp total L 271, dL / H, dL / H ratio: P- 1, 31 / 29, 1.06; P- 2, 93 / 52, 1.78; P- 3, 53 / 46, 1.16; P- 4, 79 / 26, 3.0; P- 5, 15 / 11, 1.44; P- 2 / P- 4 ratio 1.18.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E77FFD4FF0DFF2DD4A0FEE5.taxon	etymology	Etymology. Named after its similarity with T. indica.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E77FFD4FF0DFF2DD4A0FEE5.taxon	discussion	Discussion. The specimens from Borneo agree well in general morphology of the palp (P- 2 and - 3 with a subrectangular, anteriorly directed and apically serrated hyaline extensions) with Torrenticola indica Cook, 1967, a species described from India (Cook 1967). The latter species can be easily distinguished in a different colour pattern on the dorsal shield (see Cook 1967: fig. 222) and in the male having a longer medial suture line of CxII + III. Wiles (1999) reported T. indica from several sites in the catchment area of the Rivers Temburong and Belalong in Brunei Darussalem, without discussion on differences with the original description. Habitat. Sandy / bouldery streams, shaded by rain forest (Figs. 43 A, C).	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E77FFD4FF0DFF2DD4A0FEE5.taxon	distribution	Distribution. Borneo.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E71FFD9FF0DFED8D029FDE6.taxon	description	(Figs. 14 A – D, 16 E, 17 E)	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E71FFD9FF0DFED8D029FDE6.taxon	description	Male. General features — Idiosoma elongated-oval (dorsal shield L / W ratio 1.3 - 1.4); gnathosomal bay Ushaped, proximally pointed; Cxgl- 4 subapical, only slightly posterior of Cx-I tips; genital field subrectangular; posterior suture lines of Cx-IV starting at right angle from genital field; excretory pore away from the line of primary sclerotization, excretory pore on the level with Vgl- 2; gnathosomal rostrum shortened, ventral margin in lateral view curved (Fig. 14 D); P- 2 ventral margin with a denticulation also in proximal half of the segment, distally with a small, laterally compressed, anteriorly directed and apically serrated hyaline extension (rudimentary flange, Wiles 1997) and a very short, denticle-like seta laterally at base of projection; P- 3 with a subrectangular, apically serrated ventrodistal projection, and a very long seta (longer than P- 3) laterally at base of projection; P- 4 stout, with ventral tubercles pointed and separated, bearing one long and three short setae (Figs. 14 B-C). Measurements — Idiosoma (ventral view: Figs. 14 A, 17 E) L 522, W 397; dorsal shield (Fig. 16 E) L 438, W 322, L / W ratio 1.36; dorsal plate L 403; shoulder plate L 131 - 138, W 50 - 54, L / W ratio 2.5 - 2.6; frontal plate L 91 - 95, W 45 - 47, L / W ratio 2.0 – 2.03; shoulder / frontal plate L 1.44 – 1.45. Gnathosomal bay L 86, Cx-I total L 189, CxI mL 103, Cx-II + III mL 94; ratio Cx-I L / Cx-II + III mL 2.0; Cx-I mL / Cx-II + III mL 1.1. Genital field L / W 122 / 97, ratio 1.26; ejaculatory complex L 168; distance genital field-excretory pore 78, genital field-caudal idiosoma margin 111. Gnathosoma vL 246; chelicera total L 266; palp total L 246, dL / H, dL / H ratio: P- 1, 26 / 28, 0.94; P- 2, 69 / 51, 1.36; P- 3, 55 / 46, 1.2; P- 4, 75 / 29, 2.58; P- 5, 21 / 11, 1.9; P- 2 / P- 4 ratio 0.93.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E71FFD9FF0DFED8D029FDE6.taxon	discussion	Remarks. The specimens from this study matches the general morphology of Torrenticola tjiwalensis (K. Viets, 1935), a species originally described from Java (K. Viets 1935), and later on reported from Peninsular Malaysia (Wiles 1997) and Borneo (Wiles 1999). The differences with the original description (in parentheses, from K. Viets 1935) are found in slightly smaller dimensions, stouter P- 4 and the excretory pore lying on the same level as Vgl- 2 (shifted slightly posterior to Vgl- 2) of the specimen from Borneo. Thus, our assignment of the specimen from Borneo is tentative. Only with more material in the future it will be possible to decide whether this is really just a variable species or a complex of several similar species.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E71FFD9FF0DFED8D029FDE6.taxon	distribution	Distribution. Java (K. Viets 1935), Peninsular Malaysia (Wiles 1997), Borneo (Wiles 1999, our study)	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7CFFD8FF0DFDEBD734F9E8.taxon	description	(Figs. 15 A – D, 16 F, 17 F)	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7CFFD8FF0DFDEBD734F9E8.taxon	materials_examined	Type series. Holotype female, dissected and slide mounted, Malaysia, Borneo, Kibamabangan River, Crocker Range, 5 º 51.28 N, 116 º 08.417 E, alt. 433 m asl., 18. ix. 2012 Smit.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7CFFD8FF0DFDEBD734F9E8.taxon	diagnosis	Diagnosis (Male unknown). Idiosoma roundish (dorsal shield L / W ratio 1.2); Cxgl – 4 subapical, only slightly posterior of Cx-I tips; P- 2 with a laterally compressed, longish (> 30 % of ventral margin), anteriorly directed ventrodistal extension; P- 3 with a very long seta laterally at base of projection.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7CFFD8FF0DFDEBD734F9E8.taxon	description	Description Female. General features — Idiosoma roundish; gnathosomal bay U-shaped, proximally pointed; Cxgl – 4 subapical, only slightly posterior of Cx-I tips; genital field pentagonal; suture lines of Cx-IV extending back beyond posterior margin of genital field, laterally curved; excretory pore away from the line of primary sclerotization, excretory pore slightly posterior from Vgl- 2; gnathosomal rostrum long, ventral margin in lateral view slightly curved (Fig. 15 D); P- 2 slightly longer than P- 4, P- 2 ventral margin with a denticulation also in proximal half of the segment, distally with a laterally compressed, longish (> 30 % of ventral margin), anteriorly directed and apically serrated hyaline extension with and a very short, denticle-like seta laterally at base of projection; P- 3 with a longish, subrectangular, apically serrated ventrodistal projection, and a very long seta laterally at base of projection; P- 4 slender, with ventral tubercles pointed and separated, bearing one long and three short setae (Fig. 15 C). Measurements — Idiosoma (ventral view: Figs. 15 B, 17 F) L 578, W 434; dorsal shield (Figs. 15 A, 16 F) L 448, W 362, L / W ratio 1.24; dorsal plate L 403; shoulder plate L 148, W 59 – 93, L / W ratio 2.4 – 2.5; frontal plate L 116, W 53 – 56, L / W ratio 2.05 – 2.2; shoulder / frontal plate L 1.28. Gnathosomal bay L 119, Cx-I total L 256, Cx-I mL 137, Cx-II + III mL 42; ratio Cx-I L / Cx-II + III mL 6.1; Cx-I mL / Cx-II + III mL 3.3. Genital field L / W 124 / 120, ratio 1.04; distance genital field-excretory pore 106, genital field-caudal idiosoma margin 153. Gnathosoma vL 291; chelicera total L 326; palp total L 238 – 239, dL / H, dL / H ratio: P- 1, 28 / 25, 1.12; P- 2, 71 / 47, 1.49; P- 3, 55 / 39, 1.4; P 4, 68 – 69 / 20, 3.4; P- 5, 16 / 11, 1.5; P- 2 / P- 4 ratio 1.03. Male: unknown.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7CFFD8FF0DFDEBD734F9E8.taxon	etymology	Etymology. Named after Menno Schilthuizen, leader of the Mt Kinabula-Crocker Range expedition.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7CFFD8FF0DFDEBD734F9E8.taxon	discussion	Discussion. Due to the Cxgl – 4 subapical and P- 2 ventral margin distally with a laterally compressed, anteriorly directed ventrodistal extension and a denticulation also in proximal half of the segment, the new species resembles Torrenticola tjiwalensis (K. Viets, 1935) and T. dentifera Wiles, 1991. The latter species, originally described from Selangor, Peninsular Malaysia (Wiles 1991), and later on found in South Korea (Pešić et al. 2013 b), is known only from the male sex. It can easily be distinguished from the new species in having a short ventral seta on P- 3 and P- 4 with long and broadly rounded distal seta. Torrenticola tjiwalensis (Java, Peninsular Malaysia, Borneo) resembles the new species in having a long ventral seta on P- 3 (longer than P- 3) but differs in a small, rudimentary flange ventrally on P- 2 (see Wiles 1997). Further differences are found in a more elongated dorsal shield, excretory pore and Vgl- 1 shifted more away from the line of primary sclerotization and a less shallow gnathosoma with a relatively more shortened rostrum in the female of T. tjiwalensis (see K. Viets 1935) Habitat. Sandy / bouldery streams, shaded by rain forest (Fig. 43 C).	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7CFFD8FF0DFDEBD734F9E8.taxon	distribution	Distribution. Borneo; known only from the locus typicus. Subgenus Megapalpis Halbert, 1944	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7DFFDAFF0DF9F6D70BFE61.taxon	description	(Figs. 18 A – D)	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7DFFDAFF0DF9F6D70BFE61.taxon	description	Morphology Deutonymph. General features — Idiosoma roundish; frontal platelets fused to each other forming an anterior plate (Fig. 18 A); gnathosomal bay U-shaped; Cxgl – 4 far posterior at margin of Cx-I / II, near insertion of II-L; provisional genital field with four pairs of acetabula (Fig. 18 B); gnathosomal rostrum long, long, bent dorsally (Fig. 18 D); cheliceral claw very long, ratio basal segment / claw 1.1; P- 2 and P- 3 without ventral setae (Fig. 18 C). Measurements — Idiosoma L 456, W 375; dorsal shield (Fig. 18 A) L / W 309 / 223, ratio 1.39, dorsal plate L 259; shoulder plate L 127, W 50 – 52, L / W ratio 2.5; anterior plate L 131, W 45, L / W ratio 2.9. Coxal field: L 331, W 326, gnathosomal bay L 97. Provisional genital field L / W 81 / 98, ratio 0.83; Gnathosoma vL 187; chelicera total L 195, basal segment L 114, claw L 100, L basal segment / claw ratio 1.14; palp total L 232, dL / H, dL / H ratio: P- 1, 22 / 29, 0.75; P- 2, 82 / 35, 2.32; P- 3, 37 / 28, 1.34; P- 4, 73 / 21.5, 3.4; P- 5, 18 / 17, 1.95; P- 2 / P- 4 ratio 1.13.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7DFFDAFF0DF9F6D70BFE61.taxon	discussion	Remarks. The single deutonymph from this study matches the general morphology of Torrenticola pugionirostris (K. Viets, 1939), a species known from Java (K. Viets 1939, Lundblad 1971). This species is characterized by Cxgl- 4 shifted far posterior at margin of Cx-I / II, near insertion of II-L, and I – and – II-L insertions are anterior to the dorsal shield when viewed from above (Wiles 1997). The deutonymph has been described by Lundblad (1971) from Java. However, Lundblad mentioned a provisional genital field with six Ac instead of 8 in the specimen from our study. Thus, our assignment of the specimen from Borneo is tentative, and only with more material of adult specimens in the future will it be possible to clarify the status of this species.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7DFFDAFF0DF9F6D70BFE61.taxon	distribution	Distribution. Java (K. Viets 1939, Lundblad 1971). Genus Neoatractides Lundblad, 1941 Subgenus Allotorrenticola Cook, 1967	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7FFFDEFF0DFE79D7FCFF4C.taxon	description	(Figs. 19 A – F, 22 A – B)	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7FFFDEFF0DFE79D7FCFF4C.taxon	materials_examined	Type series. Holotype female, dissected and slide mounted, Malaysia, Borneo, stream Kemantis, Sayap, Mt Kinabalu, 6 º 09.841 N, 116 º 33.936 E, alt. 928 m asl., 16. ix. 2012 Smit.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7FFFDEFF0DFE79D7FCFF4C.taxon	diagnosis	Diagnosis (Male unknown). Shoulder platelets fused to the large dorsal plate; P- 1 separate from P- 2, P- 2 and - 3 without ventral projections, P- 2 with relatively long and slender seta in distal half of segment, dorsal margin proximally convexly protruding; P- 3 with relatively long, hair-like ventral seta inserted in the proximal half of the segment; gnathosoma with long and slender rostrum, without conspicuous oral papillae	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7FFFDEFF0DFE79D7FCFF4C.taxon	description	Description Female. General features — Shoulder platelets fused to the large dorsal plate (Figs. 19 B, 22 A); gnathosomal bay shallow; tips of Cx-I truncated; Cxgl- 4 far posterior at margin of Cx-I / II, halfway between I-L and II-L insertions; genital field pentagonal in shape; suture line of Cx-IV distinct, extending posteriorly beyond posterior margin of genital field; excretory pore away from the line of primary sclerotization, Vgl- 2 posterior from excretory pore; P- 1 separate from P- 2, P- 2 and - 3 without ventral projections, P- 2 relatively long, ventral margin concave, with relatively long and slender seta inserted in distal half of the segment, dorsal margin proximally convexly protruding, ventral margin of P- 3 slightly concave, with relatively long, hair-like seta inserted in proximal half of the segment (Fig. 19 E); gnathosoma with short posterodorsal projections, lacking conspicuous oral papillae, rostrum long and slender (Fig. 19 C); chelicera long with relatively short cheliceral claw (L basal segment / claw ratio 6.7). Legs: I-L (Fig. 19 F) with I-L- 6 L / H ratio 2.8. Measurements — Idiosoma (ventral view: Figs. 19 A, 22 B) L 684, W 503; dorsal shield (Figs. 19 B, 22 A) L 596, W 447, L / W ratio 1.33; dorsal plate L 563; frontal plate L 139, W 44, L / W ratio 3.18. Gnathosomal bay L 63, Cx-I total L 245, Cx-I mL 182, Cx-II + III mL 47; ratio Cx-I L / Cx-II + III mL 5.2; Cx-I mL / Cx-II + III mL 3.9. Genital field L / W 152 / 133, ratio 1.14; distance genital field-excretory pore 153, genital field-caudal idiosoma margin 242. Gnathosoma vL 378; chelicera total L 406, basal segment L 377, claw L 56, L basal segment / claw ratio 6.7; palp total L 258, dL / H, dL / H ratio: P- 1, 30 / 52, 0.57; P- 2, 107 / 49 - 54, 2.0 – 2.2; P- 3, 75 / 37, 2.04; P- 4, 31 / 26, 1.19; P- 5, 15 / 18.5, 0.79; P- 2 / P- 4 ratio 3.43. Legs: dL of I-L- 5 – 6: 88, 89; I-L- 6 H 32, dL / H I-L- 6 ratio 2.76. Male: unknown.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7FFFDEFF0DFE79D7FCFF4C.taxon	etymology	Etymology. Named after Sunda Shelf; major landmasses on the shelf include the Malay Peninsula, Sumatra, Borneo, Java, Madura, Bali and their surrounding smaller islands.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7FFFDEFF0DFE79D7FCFF4C.taxon	discussion	Discussion. The subgenus Allotorrenticola includes six species from Asia, i. e., Neoatractides abnormipalpis (Lundblad, 1941) (Burma, Malaysia, Brunei; Wiles 1997), N. bahtilli (Wiles, 1991) (Malaysia, Thailand; Wiles 1997, Pešić & Smit 2009 a), N. malayensis (Wiles, 1991) (Malaysia, Thailand; Wiles 1997, Pešić & Smit 2009 a), N. suvarna (Cook, 1967) (India, Cook 1967), N. farmerae Wiles, 1999 (Borneo, Wiles 1999) and N. calidus Pešić & Smit, 2012 (South Iran, Pešić et al. 2012 a), and N. erato Pešić & Smit, 2014 (West Africa, Pešić & Smit 2014). The shape of dorsal shield with the shoulder platelets fused to the large dorsal plate, makes the new species most similar to Neoatractides erato, a species recently described from Ghana (Pešić & Smit 2014). The latter species can easily be distinguished by the series of different character states: gnathosoma with conspicuous oral papillae (sensu Wiles 1997), P- 2 maximum height at distal margin, P- 3 stout with short ventral seta located in the distal half, P- 4 almost equal in length with P- 3, cheliceral claw relatively long, gnathosomal bay moderately deep (see Pešić & Smit 2014). Two Allotorrenticola species previously reported from Borneo (Wiles 1999), N. abnormipalpis and N. farmerae can easily be distinguished from N. sundaensis n. sp. in addition to the shoulder platelets not fused to the large dorsal plate, by P- 1 partially fused with P- 2 (see Wiles 1999). Habitat. Sandy / bouldery streams, shaded by rain forest (Fig. 43 D).	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7FFFDEFF0DFE79D7FCFF4C.taxon	distribution	Distribution. Borneo; known only from the locus typicus. Subgenus Heteratractides Lundblad, 1941	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7BFFDEFF0DFF6BD716F8FD.taxon	description	(Figs. 20 A – F, 22 C – D, 23 G)	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7BFFDEFF0DFF6BD716F8FD.taxon	materials_examined	Type series. Holotype male, dissected and slide mounted, Malaysia, Borneo, unnamed stream Bansadon Trail, Inobong, Crocker Range, 5 º 51.456 N, 116 º 68.403 E, 18. ix. 2012 Smit.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7BFFDEFF0DFF6BD716F8FD.taxon	diagnosis	Diagnosis (Female unknown). Frontal and shoulder plates fused with the large dorsal plate, but suture lines well visible; excretory pore and Vgl- 1 embedded in the area of primary sclerotization; gnathosomal rostrum broad and upturned, narrower than remainder of the gnathosoma; P- 4 stout (dL / H ratio 2.0).	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7BFFDEFF0DFF6BD716F8FD.taxon	description	Description Male. General features — Frontal and shoulder plates fused with the large dorsal plate, but suture lines well visible (Figs. 20 A, 22 C); Cxgl – 4 posterior to Cxgl – 2; medial suture line of Cx-II + III moderately long; suture line of Cx-IV in medial part directed posteriorly, laterally curved; genital field subrectangular in shape; ejaculatory complex as photographed in Figure 23 G; excretory pore and Vgl – 2 embedded in the area of primary sclerotization; P- 1 with dorsal seta, P- 2 and - 3 without ventral projections, P- 2 long with relatively short and slender ventral seta, lateral distal margin of P- 3 deeply indented, P- 4 stout, longer than P- 3, dorsal margin strongly bowed, ventral margin concave with one long and one shorter seta (Figs. 20 C – D). Gnathosoma without conspicuous oral papillae and short posterodorsal projections, rostrum broad and upturned, narrower than remainder of the gnathosoma (Fig. 20 E); chelicera long with relatively short cheliceral claw (L basal segment / claw ratio 4.4). Legs: I-L with I-L- 6 stout, L / H ratio 2.0, ventral margin distally strongly protruding (Fig. 20 F). Measurements — Idiosoma (ventral view: Figs. 20 B, 22 D) L 556, W 414; dorsal shield (Figs. 20 A, 22 C) L 455, W 328, L / W ratio 1.39. Gnathosomal bay L 131, Cx-I total L 259, Cx-I mL 128, Cx-II + III mL 106; ratio Cx-I L / Cx-II + III mL 2.4; Cx-I mL / Cx-II + III mL 1.2. Genital field L / W 93 / 70, ratio 1.32; distance genital field-excretory pore 78, genital field-caudal idiosoma margin 97. Gnathosoma vL 238 – 239; chelicera total L 263, basal segment L 220, claw L 50, L basal segment / claw ratio 4.4; palp total L 194 – 195, dL / H, dL / H ratio: P- 1, 32 / 31 – 32, 1.03; P- 2, 75 / 32, 2.36; P- 3, 34 / 38, 0.9; P- 4, 41 – 42 / 21, 2.0; P- 5, 12 / 15, 0.8; P- 2 / P- 4 ratio 1.82. Legs: dL of I-L- 5 – 6: 85, 96, I- L- 6 H 48, dL / H I-L- 6 ratio 2.0. Female: unknown.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7BFFDEFF0DFF6BD716F8FD.taxon	etymology	Etymology. The species name refers to the undivided dorsal shield of the new species.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7BFFDEFF0DFF6BD716F8FD.taxon	discussion	Discussion. Neoatractides uniscutatus n. sp. is the third representative of the subgenus Heteratractides Lundblad, 1941. Synapomorphies of the species included in this subgenus (i. e. subgeneric autapomorphies) include the following features: lateral distal margin of P- 3 deeply indented and gnathosomal rostrum broad and upturned (Wiles 1997). Two species are known so far, Neoatractides serratirostris (Lundblad, 1941) from Colombia (Lundblad 1941) and N. orientalis Wiles, 1991, described on the basis of two female specimens from Selangor, Peninsular Malaysia (Wiles 1991, 1997). From the latter species, Neoatractides uniscutatus n. sp. can be distinguished by the frontal platelets fused to the large dorsal plate. Neoatractides serratirostris resembles the new species due to the anterolateral platelets fused with the large dorsal plate forming an undivided dorsal shield, but differs from the new species in having a shorter and broader gnathosomal rostrum, a very long ventral seta on P- 2 exceeding the tip of P- 5, a conventionally shaped ejaculatory complex, relatively larger genital field and a shorter medial suture line of Cx-II + III (see Lundblad 1953). Habitat. Sandy / bouldery stream, shaded by rain forest (Fig. 43 B).	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7BFFDEFF0DFF6BD716F8FD.taxon	distribution	Distribution. Borneo; known only from the locus typicus. Genus Pseudotorrenticola Walter, 1906	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7BFFE3FF0DF885D71BFF06.taxon	description	(Figs. 21 A – D, 22 E – F, 23 H)	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7BFFE3FF0DF885D71BFF06.taxon	materials_examined	Type series. Holotype male, dissected and slide mounted, Malaysia, Borneo, Little Lumotok stream, Sayap, Mt Kinabalu, 6 º 09.497 N, 116 º 34.027 E, alt. 1065 m asl., 17. ix. 2012 Smit.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7BFFE3FF0DF885D71BFF06.taxon	diagnosis	Diagnosis (Female unknown). Large in size (idiosoma length 900 µm); anterolateral platelets of dorsal shield extending posteriorly to level of D 2; excretory pore on the margin of primary sclerotization; L / H P- 4 ratio 1.8.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7BFFE3FF0DF885D71BFF06.taxon	description	Description Male. General features — Dorsal shield with two pairs of anterolateral platelets, shoulder platelets much longer than frontal platelets, extending posteriorly to level of D 2 (Fig. 21 A); Cxgl – 4 absent; genital field subrectangular in shape; ejaculatory complex conventional in shape (Fig. 23 H); suture lines of Cx-IV extending posteriorly beyond posterior margin of genital field; excretory pore on the margin of primary sclerotization, embedded in the area of primary sclerotization, Vgl – 2 posterior to the line of primary sclerotization; gnathosoma long and narrow (Fig. 21 D), located at the end of an extrusible membranous tube. Palp segments short, ventral margin of P- 2 and P- 3 each with a short, slender ventral seta (Fig. 21 C). Measurements — Idiosoma (ventral view: Figs. 21 B, 22 F) L 906, W 531; dorsal shield (Figs. 21 A, 22 E) L 606, W 469, L / W ratio 1.29; dorsal plate L 525; shoulder plate L 288 – 297, W 75 – 78, L / W ratio 3.8; frontal plate L 184, W 84, L / W ratio 2.19; shoulder / frontal plate L 1.56 – 1.61. Gnathosomal bay L 48, Cx-I total L 356, Cx-I mL 308, Cx-II + III mL 125; ratio Cx-I L / Cx-II + III mL 2.8; Cx-I mL / Cx-II + III mL 2.5. Genital field L / W 166 / 128, ratio 1.29; ejaculatory complex L 172; distance genital field-excretory pore 184, genital field-caudal idiosoma margin 239. Gnathosoma vL 270; chelicera total L 295, basal segment L 249, claw L 49, L basal segment / claw ratio 5.1; palp total L 193 – 194, dL / H, dL / H ratio: P- 1, 21 – 22 / 26 - 29, 0.7 – 0.8; P- 2, 65 / 38.5, 1.68; P- 3, 35 / 32, 1.1; P- 4, 45 / 25, 1.77; P- 5, 27 / 14, 1.9; P- 2 / P- 4 ratio 1.45. Legs: dL of I-L- 2 – 6: 97, 88, 121, 150, 133; I-L- 6 H 40, dL / H I-L- 6 ratio 3.3; dL of IV-L: 100, 116, 103, 148, 175, 158. Female: Unknown.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7BFFE3FF0DF885D71BFF06.taxon	etymology	Etymology. Named after the island where the new species was found.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7BFFE3FF0DF885D71BFF06.taxon	discussion	Discussion. In the general characters, Peudotorrenticola borneoensis n. sp. closely resembles P. sharpae Wiles, 1997. The latter species was described by Wiles (1997) based on material from Thailand (holotype male), Peninsular Malaysia (one male and one female) and North Brunei (one male). Wiles (1997) mentioned that the specimen from Brunei differs in having more slender palps and anterolateral platelets extending posteriorly to the level with D 2. He stated that this may be a separate species. In the light of the new record from northern Borneo, these morphological differences cannot be attributed to intraspecific variability. The specimen examined from Borneo differs from the original description of P. sharpae in larger dimensions (idiosoma length 670 – 740 µm in male of P. sharpae), anterolateral platelets extending more posteriorly, approaching to the level with D 2, a more slender palp especially P- 4, and excretory pore lying on the margin of primary sclerotization (away from the margin of primary sclerotization in P. sharpae, see Wiles 1997: fig. 20 f). In our opinion the specimen from this study warrant the erection of a new species. Habitat. Sandy / bouldery stream, shaded by rain forest.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E7BFFE3FF0DF885D71BFF06.taxon	distribution	Distribution. Borneo (“ P. sharpae ” Wiles 1997, our study). Genus Monatractides K. Viets, 1926 Subgenus Monatractides K. Viets, 1926	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E46FFE4FF0DFE90D590F9B9.taxon	description	(Figs. 28 A – B, 29 A – B, 30 A, Tab. 1)	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E46FFE4FF0DFE90D590F9B9.taxon	materials_examined	Compared material	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E46FFE4FF0DFE90D590F9B9.taxon	description	General features — Idiosoma elongated (dorsal shield L / W ratio 1.6 - 1.7); shoulder plates elongated (shoulder / frontal plate L ratio 1.2 - 1.3); frontal margin with flat protrusions medial to the eyes; Cxgl- 4 located far anteriorly, near tips of Cx-I; gnathosomal bay U-shaped; suture line of Cx-IV lightly accentuated, originating from lateral edge of genital field and extending posteriorly beyond posterior margin of genital field (Figs. 29 A-B); Vgl- 2 posterior to excretory pore; P- 2 and P- 3 distal margins without pointed tips, P- 4 without denticles near insertion of ventral setae. Male: medial margin of Cx-II / III relatively long; genital field subrectangular; ejaculatory complex normal in shape (proximal and distal arms and proximal chamber well developed, Fig. 30 A); excretory pore and Vgl- 2 slightly away from the line of primary sclerotization, near posterior margin of idiosoma. Female: genital field pentagonal, anteriorly enlarged, laterally straight, tapering posteriorly (Fig. 29 B); excretory pore and Vgl- 2 located posterior to line of primary sclerification.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E46FFE4FF0DFE90D590F9B9.taxon	discussion	Remarks. The specimens from Borneo agree well in general morphology with Monatractides longiventris, a species originally described from Java (K. Viets 1939). The original description was restricted to the male; a first description of the female from Central Java was given by Lundblad (1971). In comparison with the holotype, specimens examined in our study are smaller, but do not differ in proportions and general idiosoma and palp morphology (Table 1).	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E46FFE4FF0DFE90D590F9B9.taxon	distribution	Distribution. Java (“ Atractides longiventris ” K. Viets 1939, “ T. longiventris ” Lundblad 1971). New for Borneo.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E4DFFEAFF0DFF2DD68AFC5D.taxon	description	(Figs. 24 A – D, 26 B, 28 F – G, 29 F – G, 30 C – D, Tabs. 2 – 3)	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E4DFFEAFF0DFF2DD68AFC5D.taxon	materials_examined	Material examined. Malaysia, Borneo, unnamed stream Bansadon Trail, Inobong, Crocker Range, 5 º 51.456 N, 116 º 08.403 E, alt. 436 m asl, 18. ix. 2012 Smit 1 / 0 / 0 (mounted); small stream waterfall trail Inobong, 5 º 51.306 N, 116 º 08.292 E, alt. 482 m asl., 19. ix. 2012 Smit 1 / 1 / 0 (mounted). Compared material Monatractides macroporus (K. Viets, 1935): Holotype ♂, SMF 4422 Torrenticola macropora ♂, type, K. Viets 47766, Sumatra, Ranau Urwaldbaches bei Aer Pisaup 22.1.1929; SMF 4431 Torrenticola macropora ♀, type, K. Viets 47763, Sumatra, Musi bei Aer Simpang, 650 m, 6.5.1929. Monatractides macrognathus (K. Viets, 1935): SMF 4173 Torrenticola macrognatha 1 ♂, 2 ♀, K. Viets 47733, Java, Zufluβbaches des Ranoe Bedali, 2.11.1928. Monatractides major (K. Viets, 1935): SMF 4173 Torrenticola macrognatha 1 ♂, 2 ♀, K. Viets 47733, Java, Zufluβbaches des Ranoe Bedali, 2.11.1928. Morphology Male. General features — Idiosoma elongated-oval; frontal plates equal or slightly longer than shoulder plates (shoulder / frontal plate L ratio 0.9); frontal margin medially pointed, between anterolaterally pointed apodemes; Cxgl – 4 located far anteriorly, near tips of Cx-I; gnathosomal bay deep, narrow U-shaped (Fig. 24 B); suture line of Cx-IV distinct, originating from lateral edge of genital field and extending posteriorly beyond posterior margin of genital field; excretory pore and Vgl- 2 away from the line of primary sclerotization, Vgl – 2 slightly posterior to excretory pore; gnathosoma elongated in lateral view, rostrum truncated; ventral and dorsal setae of P- 2 and P- 3 relatively strong and long, P- 2 and P- 3 distal margins without pointed tips, P- 4 ventral setae short (Fig. 24 C). Male: medial margin of Cx-II / III relatively short; genital field subrectangular in shape; ejaculatory complex (Figs. 30 C – D): proximal and distal arms well developed, carina anterior long, proximal chamber small (Fig. 26 B). Measurements Male (from unnamed stream Bansadon Trail, in parentheses some measurements of specimen from stream waterfall trail Inobong) — Idiosoma (ventral view: Fig. 29 F) L 631 (637), W 394 (407); dorsal shield (Fig. 28 F) L 503 (506), W 365 (370), L / W ratio 1.38 (1.37); dorsal plate L 453 (453); shoulder plate L 111 – 114 (125 – 128), W 56 (59 – 62), L / W ratio 2.0 (2.1); frontal plate L 128 – 131 (123 – 128), W 56 – 58 (59 – 61), L / W ratio 2.2 – 2.3 (2.0 – 2.2); shoulder / frontal plate L ratio 0.87 (0.98 – 1.03). Gnathosomal bay L / W 140 (134) / 34 (31), ratio 4.1 (4.3), Cx-I total L 234 (239), Cx-I mL 94 (105), Cx-II + III mL 73 (61); ratio Cx-I L / Cx-II + III mL 3.2 (3.9); Cx-I mL / CxII + III mL 1.3 (1.7). Genital field L / W 116 (122) / 91 (91), ratio 1.28 (1.35); ejaculatory complex L 154 (153); distance genital field-excretory pore 156 (150), genital field-caudal idiosoma margin 213 (216). Gnathosoma vL 132 (136); chelicera total L 143 (145); palp total L 143, dL / H, dL / H ratio: P- 1, 20 / 16, 1.23; P- 2, 40 / 31, 1.3; P- 3, 31 / 26, 1.18; P- 4, 37 / 19, 2.0; P- 5, 15 / 9, 1.7; P- 2 / P- 4 ratio 1.08. Legs: dL of I-I- 2 – 6: 42 (45), 45 (45), 57 (57), 65 (66), 71 (71), I-L- 6 H 29 (28), L / H I-L- 6 ratio 2.5 (2.5). In addition we give locality records and measurements of female specimens suspected to belong to M. cf. macroporus. New records. Great Lumotok stream, Mt Kinabalu, 6 º 09.336 N, 116 º 34.056 E, alt. 1087 m asl., 17. ix. 2012 Smit 0 / 1 / 0; Little Lumotok stream, Sayap, Mt Kinabalu, 6 º 09.467 N, 116 º 34.027 E, alt. 1069 m asl., 17. ix. 2012 Smit 0 / 1 / 0; Kibamabangan River, at waterfall, Crocker Range, 5 º 51.280 N, 116 º 08.417 E, 433 m asl., 18. ix. 2012 Smit 0 / 1 / 0. Measurements Female (from Little Lumotok stream, n = 1) — Idiosoma (ventral view: Fig. 29 G) L 818, W 587; dorsal shield (Fig. 28 G) L 681, W 488, L / W ratio 1.4; dorsal plate L 619; shoulder plate L 167 – 169, W 64 – 69, L / W ratio 2.5 – 2.6; frontal plate L 138 – 140, W 77, L / W ratio 1.8; shoulder / frontal plate L ratio 1.19 – 1.23. Gnathosomal bay L / W 173 / 34, ratio 5.1, Cx-I total L 322, Cx-I mL 148, Cx-II + III mL 53; ratio Cx-I L / Cx-II + III mL 6.1; Cx-I mL / Cx-II + III mL 2.8. Genital field L / W 156 / 138, ratio 1.14; distance genital field-excretory pore 184, genital fieldcaudal idiosoma margin 184. Gnathosoma vL 208; chelicera total L 222; palp total L 179, dL / H, dL / H ratio: P- 1, 25 / 23, 1.07; P- 2, 52 / 37, 1.41; P- 3, 38.5 / 28, 1.39; P- 4, 41 / 18.5, 2.2; P- 5, 22 / 11, 2.1; P- 2 / P- 4 ratio 1.27. Legs: dL of I-I- 4 – 6: 88, 92, 91 – 92, I-L- 6 H 26.5, L / H I-L- 6 ratio 3.45.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E4DFFEAFF0DFF2DD68AFC5D.taxon	discussion	Remarks. The specimens from Borneo match the general morphology of Monatractides macroporus, a species described by K. Viets (1935) from Sumatra based on a single male and one female specimen under question mark. In the same paper Viets (1935) described two species, closely related to the latter, M. macrognathus (K. Viets, 1935) and M. macrognathus major (K. Viets, 1935). Lundblad (1971) considered the latter two species synonymous with M. macroporus. However, Wiles (1991) rejected the synonymization of the latter species with M. macroporus and raised M. macrognathus major to a full species. Monatractides major is characterized at first line in comparison with the two abovementioned species by having four dorsoglandularia incorporated in the area of primary sclerotization. Monatractides macroporus (as well the specimens examined from Borneo) has a narrower gnathosomal bay than M. macrognathus (compare measurements in Tables 2 and 3) but this character is known to be variable in this species (Lundblad 1971). At the time being we keep M. macrognathus as synonym of M. macroporus. In comparison with the holotype of M. macroporus and specimens of M. macrognathus from Java (see Tables 2 – 3), males examined in our study are notably smaller, and differ in having much shorter shoulder plates, more or less equal in dimensions to frontal plates, stouter I-L- 6 and anterior dorsoglandularia shifted far anterior from medial muscle scar (lying near anterior end of medial muscle scar in specimens from Java). Thus, our assignment of the specimens from Borneo to M. macroporus is tentative, and very probably specimens from Borneo belong to a new species. However, only with application of molecular techniques it will be possible to decide whether this is really just a variable species or a complex of several similar species. Furthermore, as already stated by Pešić & Smit (2009 b) understanding the taxonomic position of other species (M. macroporus (K. Viets, 1935), M. major (K. Viets, 1935), M. angulatus (Walter, 1928), M. minor Wiles, 1991, and M. nondescripta (Cook, 1966 )) tentatively included in the so-called M. macroporus- complex is not possible without additional material from a wide area and / or without the application of molecular techniques. Wiles (1999) reported a single specimen, which best fits the description of M. macrognathus in the River Temburong (North Brunei). He mentioned that “ there are other specimens that are not such a good fit and their status must await further investigation ”.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E4DFFEAFF0DFF2DD68AFC5D.taxon	distribution	Distribution. Sumatra (“ Atractides macroporus, A. macrognathus ” K. Viets 1935), Java (“ A. macrognathus ” K. Viets 1935, “ Torrenticola macropora ” Lundblad 1971). New for Borneo.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E4BFFF1FF0DFB47D671FC39.taxon	description	(Figs. 28 E, 29 E, 30 B, Tab. 4)	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E4BFFF1FF0DFB47D671FC39.taxon	materials_examined	Material examined. Malaysia, Borneo, Kipungit River, Poring Hot Springs, Mt Kinabalu, 6 º 02.776 N, 116 º 41.432 E, 568 m asl., 15. ix. 2012 Smit 1 / 0 / 0 (mounted); Kibamabangan River, at waterfall, Crocker Range, 5 º 51.280 N, 116 º 08.417 E, 433 m asl., 18. ix. 2012 Smit 0 / 1 / 0. Compared material Monatractides oxystomus (K. Viets, 1935): Lectotype ♀, here designated, SMF 4361 Torrenticola oxystoma ♀, type, K. Viets 47777, Java, Tjibodas, 1400 m, Kali Tjiwalen, 13.7.1929. Paralectotype ♀, on the same slide, here designated. Monatractides tobaensis (K. Viets, 1935): Lectotype ♂, here designated, SMF 4509 Torrenticola tobaensis ♂, type, K. Viets 47804, Sumatra, Toba-Meer, 5 m, 8.4.1929. Paralectotype ♀, on the same slide, here designated. Morphology Male. General features — Idiosoma elongated-oval; shoulder platelet elongated (shoulder / frontal plate L ratio 2.0); frontal margin medially with large anterolaterally pointed apodemes; Cxgl – 4 located far anteriorly, near tips of Cx-I; gnathosomal bay V-shaped, three pairs of knob-like protrusions at the lateral margins of gnathosomal bay; medial margin of Cx-II / III moderately long; genital field subrectangular; ejaculatory complex: proximal and distal arms well developed, carina anterior long, proximal chamber small (Fig. 30 B); suture line of Cx-IV distinct, originating from lateral edge of genital field, laterally curved; line of primary sclerotization close to posterior margin of genital field; excretory pore and Vgl – 2 well separated from the line of primary sclerotization, Vgl – 2 posterior to excretory pore; gnathosoma with long dorsal apodemes, rostrum truncated; distal margin of P- 3 and P 4 medially and laterally with several pointed extensions; P- 4 with two ventral setae, one very long and away from distal edge.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E4BFFF1FF0DFB47D671FC39.taxon	discussion	Remarks. The male specimen from Kipungit River agrees well in general morphology with Monatractides oxystomus (K. Viets, 1935), a species originally described from Sumatra (K. Viets 1935) based on two females. The male was described by Lundblad (1971) from Central Java. Monatractides oxystomus seems to be most similar to M. tobaensis (K. Viets, 1935), a species decribed from Toba Lake, in N Sumatra from at depth of 5 m (K. Viets 1935). The main difference between M. macroporus and M. tobaensis concerns the gnathosomal bay, characteristically V-shaped in the both species: in M. oxystomus three pairs of knob-like protrusions laying at the lateral margins of gnathosomal bay, while they are lacking in M. tobaensis. However there is one another important difference between these two species — the presence of swimming setae on the legs II – IV in M. tobaensis. The presence of swimming setae on legs was mentioned by K. Viets in the original description of M. tobaensis and its confirmed by our re-examination of the holotype of M. tobaensis. The numbers of swimming setae are as follows: II-L- 4, 1; II-L- 5, 1; III-L- 4, 1; III-L- 5, 1; IV-L- 4, 1; IVL- 5, 1. This is the second species of the genus characterized by the presence of swimming setae on legs. Recently, Pešić & Smit (2014) erected a new subgenus, Vietsclio, to accommodate Monatractides uniscutatus (K. Viets, 1925), a species known from Cameroon (K. Viets 1925) and Ghana (Pešić & Smit 2014). The latter species can easily be distinguished from M. tobaensis due to the anterolateral platelets fused with the large dorsal plate forming an undivided dorsal shield. Therefore we propose to shift M. tobaensis to the subgenus Vietsclio Pešić & Smit, 2014.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E4BFFF1FF0DFB47D671FC39.taxon	distribution	Distribution. Sumatra (“ Atractides oxystomus ” K. Viets 1935), Java (“ Torrenticola oxystoma ” Lundblad 1971), Thailand (Pešić & Smit 2009 b). New for Borneo.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E55FFF3FF0DF9E7D72DFBA6.taxon	description	(Figs. 28 C – D, 29 C – D, Tab. 5)	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E55FFF3FF0DF9E7D72DFBA6.taxon	materials_examined	Material examined. Malaysia, Borneo: unnamed stream Bansadon Trail, Inobong, Crocker Range, 5 º 51.456 N, 116 º 68.403 E, alt. 436 m asl., 18. ix. 2012 Smit 4 / 1 / 0 (1 / 1 / 0 mounted); Kibamabangan River, Crocker Range, 5 º 51.280 N, 116 º 08.417 E, 433 m asl., 18. ix. 2012 Smit 1 / 1 / 0; Mahua stream, Mahua, Crocker Range, 5 º 47.838 N, 116 º 24.510 E, alt. 1052 m asl., 21. ix. 2012 Smit 1 / 2 / 2; Mahua stream, upstream, Crocker Range, 5 º 47.939 N, 116 º 24.317 E, alt. 1050 m asl., 21. ix. 2012 Smit 2 / 1 / 0 (dorsal shield of female mounted); small stream waterfall trail Inobong, 5 º 51.306 N, 116 º 08.292 E, alt. 482 m asl., 19. ix. 2012 Smit 2 / 0 / 0; Kipungit River, Poring Hot Springs, Mt Kinabalu, 6 º 02.776 N, 116 º 41.432 E, 568 m asl., 15. ix. 2012 Smit 2 / 11 / 0; stream Kemantis, Sayap, Mt Kinabalu, 6 º 09.841 N, 116 º 33.936 E, alt. 928 m asl., 16. ix. 2012 Smit 1 / 2 / 0 (1 / 0 / 0 mounted); Great Lumotok stream, Mt Kinabalu, 6 º 09.336 N, 116 º 34.056 E, alt. 1087 m asl., 17. ix. 2012 Smit 0 / 2 / 0; Little Lumotok stream, Sayap, Mt Kinabalu, 6 º 09.467 N, 116 º 34.027 E, alt. 1069 m asl., 17. ix. 2012 Smit 8 / 15 / 2; Silau-Silau stream, upstream, Mt Kinabalu, 6 º 00.681 N, 116 º 32.417 E, alt. 1592 m asl., 11. ix. 2012 Smit 0 / 1 / 0 (mounted); Liwagu River, upstream, Mt Kinabalu, 6 º 01.868 N, 116 º 32.912 E, alt. 1919 m asl., 11. ix. 2012 Smit 0 / 1 / 0. Morphology General features — Idiosoma roundish; shoulder plates very short, shorter than frontal plates (shoulder / frontal plate L ratio 0.7 – 0.8); frontal margin with flat protrusions medial to the eyes; Cxgl – 4 located far anteriorly, near tips of Cx-I; gnathosomal bay U-shaped; suture line of Cx-IV distinct, originating from lateral edge of genital field and extending posteriorly beyond posterior margin of genital field (Figs. 29 C – D); excretory pore and Vgl – 2 away from the line of primary sclerotization, Vgl – 2 posterior to excretory pore; gnathosoma with long dorsal apodemes; P- 4 with well visible denticle near insertion of ventral setae. Male: medial margin of Cx-II / III short; genital field subrectangular; ejaculatory complex: proximal and distal arms well developed, carina anterior long, proximal chamber small. Female: genital field pentagonal, anteriorly enlarged, laterally straight, tapering posteriorly (Fig. 29 C).	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E55FFF3FF0DF9E7D72DFBA6.taxon	discussion	Remarks: The specimens examined from Borneo agree well in general morphology with Monatractides parviventris (K. Viets, 1935), a species originally described from South Sumatra (K. Viets 1935) based on a single female and one deutonymph. The male was described by Lundblad (1971) from Central Java. Wiles (1999) reported M. parviventris from several sites in the catchment area of the River Belalong in North Brunei. He mentioned variability of his specimens in the shape of the idiosoma (rounded to oval) and in the fusion between the median margins of frontal plates. We also found a mixed populations with separate (Fig. 28 C) and fused (Fig. 28 D) frontal plates. As stated by Wiles (1999) the taxonomy of this species needs further investigation, probably with application of molecular techniques. However, unfortunately our effort to obtain the mtCOI fragment from specimens of this species from Borneo, was not successful (see under Molecular results).	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E55FFF3FF0DF9E7D72DFBA6.taxon	distribution	Distribution. Sumatra (“ Atractides parviventris ” K. Viets 1935), Java (“ Torrenticola parviventris ” Lundblad 1971), Thailand (Pešić & Smit 2009 b).	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E57FFF5FF0DF893D634F86B.taxon	description	(Figs. 25 A – G, 26 A, 27 A – B, 28 H – I, 29 H – I, 30 E – F)	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E57FFF5FF0DF893D634F86B.taxon	materials_examined	Type series. Holotype male, dissected and slide mounted, Malaysia, Borneo, Kipungit River, Poring Hot Springs, Mt Kinabalu, 6 º 02.776 N, 116 º 41.432 E, 568 m asl., 15. ix. 2012 Smit. Paratypes: one male, same data as holotype, dissected and slide mounted. Other material: one female dissected and slide mounted same data as holotype.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E57FFF5FF0DF893D634F86B.taxon	diagnosis	Diagnosis. Male. Idiosoma elongated-oval (dorsal shield L / W ratio 1.3 – 1.4); frontal plates similar in dimensions to shoulder plates (shoulder / frontal plate L ratio 0.99 – 1.04); gnathosomal bay deep, narrow U-shaped (L / W ratio 4.6); medial margin of Cx-II / III relatively short; genital field elongated (L / W ratio 1.3 – 1.4); ejaculatory complex with large proximal chamber, proximal and distal arms short; excretory pore anterior to Vgl – 2 and away from the line of primary sclerotization; ventral setae on P- 4 short.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E57FFF5FF0DF893D634F86B.taxon	description	Description Male. General features — Idiosoma elongated-oval; frontal plates similar in dimensions to shoulder plates; frontal margin medially pointed, between anterolaterally pointed apodemes (Fig. 25 A); Cxgl – 4 located far anteriorly, near tips of Cx-I; gnathosomal bay deep, narrow U-shaped (Fig. 25 C); medial margin of Cx-II / III relatively short; genital field subrectangular in shape; ejaculatory complex characteristic (Figs. 26 A, 30 E – F), proximal chamber large, proximal and distal arms short, carina anterior short; suture line of Cx-IV distinct, originating from lateral edge of genital field and extending posteriorly beyond posterior margin of genital field; excretory pore and Vgl – 2 away from the line of primary sclerotization, Vgl – 2 slightly posterior to excretory pore; gnathosoma elongated in lateral view (Fig. 25 F), rostrum truncated; ventral and dorsal setae of P- 2 and P- 3 relatively strong and long, P- 2 and P- 3 distal margins without pointed tips, P- 4 ventral setae short (Figs. 25 D – E). Measurements Male (holotype; in parentheses paratype, n = 1) — Idiosoma (ventral view: Figs. 25 C, 29 H) L 618 (591), W 394 (369); dorsal shield (Figs. 25 B, 28 H) L 477 (453), W 363 (331), L / W ratio 1.31 (1.37); dorsal plate L 429 (408); shoulder plate L 124 – 127 (113), W 52 – 56 (50 – 53), L / W ratio 2.2 – 2.4 (2.1 – 2.3); frontal plate L 122 – 125 (111), W 53 – 55 (47 – 52), L / W ratio 2.3 (2.2 – 2.4); shoulder / frontal plate L ratio 0.99 – 1.04 (1.0). Gnathosomal bay L / W 148 (128) / 31 (28), ratio 4.8 (4.6); Cx-I total L 250 (227), Cx-I mL 101 (98), Cx-II + III mL 66 (58); ratio Cx-I L / CxII + III mL 3.8 (3.9); Cx-I mL / Cx-II + III mL 1.5 (1.7). Genital field L / W 119 (111) / 87 (83), ratio 1.37 (1.34); ejaculatory complex L (166); distance genital field-excretory pore 138 (141), genital field-caudal idiosoma margin 183 (195). Gnathosoma vL 147 (138); chelicera total L 160 (148); palp total L 149 (147), dL / H, dL / H ratio: P- 1, 21.5 / 15, 1.4 (21.5 / 17, 1.3); P- 2, 41.5 / 31, 1.35 (39 / 26, 1.5); P- 3, 33 / 26, 1.27 (32 / 23, 1.4); P- 4, 36 / 17, 2.14 (37 / 17, 2.19); P- 5, 17 / 9, 1.84 (17 / 9, 1.85); P- 2 / P- 4 ratio 1.15 (1.06). Legs: dL of I-L- 2 – 6: 45, 52 (52), 65 (65), 69 (69), 71 (68), I-L- 6 H 28.5 (26), L / H I-L- 6 ratio 2.5 (2.6). At present, it is impossible to give an unequivocal diagnosis for the female of M. epiales n. sp. In the following list, measurements are given for a female taken at the locus typicus of M. epiales n. sp. — Idiosoma (ventral view: Figs. 27 A, 29 I) L 700, W 430; dorsal shield (Fig. 28 I) L 553, W 409, L / W ratio 1.35; dorsal plate L 500; shoulder plate L 122 – 127, W 64 – 67, L / W ratio 1.9; frontal plate L 131, W 61 – 66, L / W ratio 2.0 – 2.2; shoulder / frontal plate L ratio 0.93 – 0.96. Gnathosomal bay L 148, Cx-I total L 253, Cx-I mL 106, Cx-II + III mL 46; ratio Cx-I L / Cx-II + III mL 5.5; Cx-I mL / Cx-II + III mL 2.3. Genital field pentagonal in shape, anteriorly enlarged, laterally straight, tapering posteriorly, L / W 150 / 130, ratio 1.16; distance genital field-excretory pore 175, genital field-caudal idiosoma margin 250. Gnathosoma vL 138 – 139; chelicera total L 159; palp (Fig. 27 B) total L 155, dL / H, dL / H ratio: P- 1, 22 / 17, 1.3; P- 2, 43 / 32, 1.35; P- 3, 33 / 26, 1.27; P- 4, 40 / 18.5, 2.16; P- 5, 17 / 9, 1.87; P- 2 / P- 4 ratio 1.08. Legs: dL of I-L- 4 – 6: 62, 69, 71, I-L- 6 H 30, L / H I-L- 6 ratio 2.4.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E57FFF5FF0DF893D634F86B.taxon	etymology	Etymology. The species is named after Epiales (Ancient Greek: Ἐπιάλης), who was a spirit of nightmares. The species name is a noun in apposition (in the nominative case), despite the Recommendation 31 A of the ICZN (1999) about avoidance of personal names as nouns in appositions, because there is no case for it being confusing or misleading.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E57FFF5FF0DF893D634F86B.taxon	discussion	Discussion. Monatractides epiales n. sp. is rather similar and probably represent a sister species of M. macroporus (K. Viets, 1935) (see above) with respect to the general shape of idiosoma and palps. Main differences between M. macroporus and M. epiales n. sp. are found in the morphology of ejaculatory complex (normallydeveloped with small proximal chamber in M. macroporus vs. proximal chamber large, proximal and distal arms short, carina anterior short in M. epiales n. sp. — compare Figs. 26 A and - B). The female assigned to M. epiales n. sp. can be identified on the basis of a relatively larger extension of the genital field (L ratio genital / idiosoma 0.21). Habitat. Sandy / bouldery stream, shaded by rain forest	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E57FFF5FF0DF893D634F86B.taxon	distribution	Distribution. Borneo; known only from the locus typicus.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E51FFF6FF0DF94ED4A0FC5D.taxon	description	(Figs. 30 I – L, 31 A – G, 32 A – B, 33 A – H, 39 A – E, Tab. 5)	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E51FFF6FF0DF94ED4A0FC5D.taxon	materials_examined	Type series. Holotype male, dissected and slide mounted, Malaysia, Borneo, small stream Layang Layang, Mt Kinabalu, 6 º 02.711 N, 116 º 33.627 E, alt. 2697 m asl., 12. ix. 2012 Smit. Paratypes: 5 (2 juv.) / 11 / 0, same data as holotype, two males and three females of them dissected and slide mounted. Further records. Malaysia, Borneo: first stream Minduk Sirung Trail, Alab, Crocker Range, 5 º 49.249 N, 116 º 19.862 E, alt. 1781 m asl., 23. ix. 2012 Smit 2 / 2 / 0 (2 / 0 / 0 mounted); Kibamabangan River, Crocker Range, 5 º 51.280 N, 116 º 08.417 E, 433 m asl., 18. ix. 2012 Smit 2 / 3 / 0; Mahua stream, Mahua, Crocker Range, 5 º 47.838 N, 116 º 24.510 E, alt. 1052 m asl., 21. ix. 2012 Smit 3 / 2 (1 ov, 1 juv) / 0; stream Kemantis, Sayap, Mt Kinabalu, 6 º 09.841 N, 116 º 33.936 E, alt. 928 m asl., 16. ix. 2012 Smit 1 / 1 / 0 (1 / 0 / 0 mounted); Great Lumotok stream, Mt Kinabalu, 6 º 09.336 N, 116 º 34.056 E, alt. 1087 m asl., 17. ix. 2012 Smit 4 / 3 / 0; Little Lumotok stream, Sayap, Mt Kinabalu, 6 º 09.467 N, 116 º 34.027 E, alt. 1069 m asl., 17. ix. 2012 Smit 3 / 3 / 0 (2 / 2 / 0 mounted); Rheocrene along Little Lumotok stream, Sayap, Mt Kinabalu, 6 º 09.440 N, 116 º 34.026 E, 17. ix. 2012 Smit 1 / 0 / 0; Silau-Silau stream, upstream, Mt Kinabalu, 6 º 00.681 N, 116 º 32.417 E, alt. 1592 m asl., 11. ix. 2012 Smit 1 / 0 / 0 (mounted); Liwagu River, upstream, Mt Kinabalu, 6 º 01.868 N, 116 º 32.912 E, alt. 1919 m asl., 11. ix. 2012 Smit 2 / 0 / 0; spring near helipad, Summit Trail, Mt Kinabalu, 6 º 03.199 N, 116 º 33.957 E, alt. 3048 m asl., 14. ix. 2012 Smit 0 / 1 / 0; small stream Panar Laban Hut Mt Kinabalu, 6 º 03.641 N, 116 º 33.968 E, 3330 m asl., 13. ix. 2012 Smit 1 / 1 / 0; torrential stream crossing main road Alab-Kota Kinabalu, Crocker Range, 5 º 50.039 N, 116 º 20.156 E, alt. 1571 m asl., 24. ix. 2012 Smit 0 / 1 / 0.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E51FFF6FF0DF94ED4A0FC5D.taxon	diagnosis	Diagnosis. Idiosoma elongated (dorsal shield L / W ratio 1.4); shoulder plates only slightly longer than frontal plates (shoulder / frontal plate L ratio 1.03 – 1.04); gnathosomal bay U-shaped; excretory pore well anterior to Vgl- 2 and away from the line of primary sclerotization; ventral setae on P- 4 short. Male: medial margin of Cx-II / III relatively short; genital field, large and elongated (L / W ratio 1.4); ejaculatory complex with large proximal chamber, proximal and distal arms short.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E51FFF6FF0DF94ED4A0FC5D.taxon	description	Description General features — Idiosoma elongated; shoulder plates only slightly longer than frontal plates; frontal margin medially slightly convex between large anterolaterally pointed apodemes (Fig. 31 A); Cxgl- 4 located far anteriorly, near tips of Cx-I; gnathosomal bay deep, U-shaped; suture line of Cx-IV distinct, originating from lateral edge of genital field and extending posteriorly beyond posterior margin of genital field; excretory pore and Vgl- 2 well separated from the line of primary sclerotization, Vgl- 2 posterior to excretory pore; nearly triangular in lateral view, rostrum truncated (Fig. 31 F); P- 2 and P- 3 distal margins without pointed tips, P- 4 ventral setae short, not reaching the tip of P- 5 (Figs. 31 D – E). Male: medial margin of Cx-II / III relatively short; genital field subrectangular, large and elongated (L / W ratio 1.4); ejaculatory complex (Figs. 30 I – L, 33 F – G) characteristic, proximal chamber large, proximal and distal arms short. Female: genital field large and pentagonal in shape, anteriorly enlarged, laterally straight, tapering posteriorly. Measurements Male (holotype; in parentheses paratype from small stream Layang Layang, n = 1) — Idiosoma (ventral view: Figs. 31 C, 40 A) L 913 (797), W 681 (591); dorsal shield (Figs. 31 B, 39 A) L 806 (719), W 578 (503), L / W ratio 1.4 (1.4); dorsal plate L 756 (679); shoulder plate L 173 (166 – 169), W 72 – 74 (67 – 69), L / W ratio 2.3 – 2.4 (2.4 – 2.5), frontal plate L 168 – 169 (147 – 154), W 67 (63), L / W ratio 2.5 (2.4 – 2.5); L shoulder / frontal plate ratio 1.03 – 1.04 (1.08 – 1.15). Gnathosomal bay L 187 (170), Cx-I total L 350 (313), Cx-I mL 163 (142), Cx-II + III mL 45 (43); ratio Cx-I L / Cx-II + III mL 7.8 (7.3); Cx-I mL / Cx-II + III mL 3.6 (3.3). Genital field L / W 200 (195) / 143 (139), ratio 1.4; ejaculatory complex L 250 (244); distance genital field-excretory pore 169 (147), genital field-caudal idiosoma margin 301 (240). Gnathosoma vL 203 (188); chelicera total L 248 (211); palp total L 249 (226), dL / H, dL / H ratio: P- 1, 37 / 29, 1.25 (34 / 29, 1.16); P- 2, 72 / 46, 1.56 (63 / 47, 1.33); P- 3, 49 / 39, 1.26 (45 / 38, 1.17); P- 4, 63 / 29, 2.18 (59 / 27, 2.19); P- 5, 28 / 15, 1.8 (25 / 14, 1.8); P- 2 / P- 4 ratio 1.15 (1.07). Legs: dL of I-L- 2 – 6: 85, 98, 133, 129, 132, I-L- 6 H 48, dL / H I-L- 6 ratio 2.8. Female (paratype from small stream Layang Layang, n = 1) — Idiosoma (ventral view: Figs. 32 A, 40 B) L 881, W 663; dorsal shield (Fig. 39 B) L 773, W 556, L / W ratio 1.4; dorsal plate L 734; shoulder plate L 170 – 172, W 70, L / W ratio 2.4 – 2.5; frontal plate L 162 – 163, W 59, L / W ratio 2.7; L shoulder / frontal plate ratio 1.05 – 1.06. Gnathosomal bay L 169, Cx-I total L 306, Cx-I mL 138, Cx-II + III mL 31; ratio Cx-I L / Cx-II + III mL 9.9; Cx-I mL / Cx-II + III mL 4.5. Genital field L / W 226 / 188, ratio 1.21; distance genital field-excretory pore 172, genital fieldcaudal idiosoma margin 302. Gnathosoma vL 216; chelicera total L 241; palp total L 233, dL / H, dL / H ratio: P- 1, 32 / 30, 1.08; P- 2, 65.5 / 48, 1.36; P- 3, 48 / 41.5, 1.15; P- 4, 63 / 30, 2.09; P- 5, 24 / -; P- 2 / P- 4 ratio 1.04; Legs: dL of I-L 2 – 6: 80, 94, 122, 122, 119, I-L- 6 H 42, dL / H I-L- 6 ratio 2.8.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E51FFF6FF0DF94ED4A0FC5D.taxon	etymology	Etymology. The species is named after Morpheus (Ancient Greek: Μορφεύς), who was a spirit of dreams, and who takes the shape of humans. The species name is a noun in apposition (in the nominative case), despite the Recommendation 31 A of the ICZN (1999) about avoidance of personal names as nouns in appositions, because there is no case for it being confusing or misleading.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E51FFF6FF0DF94ED4A0FC5D.taxon	discussion	Discussion. Monatractides morpheus n. sp. resembles the M. luteus (K. Viets, 1935) — complex characterized in the first line in comparison with the species of M. macroporus – complex (see above) by the presence of anteriorly broad and short Cx-I and relatively wide gnathosomal bay (Pešić & Smit 2010). This group includes five similar species known from Java, i. e., M. luteus (K. Viets, 1935), M. roseus (Lundblad, 1941), M. landbergi (Lundblad, 1941), M. parvus (Lundblad, 1941) and M. longiusculus (Lundblad, 1941). From all abovementioned species the new species from Borneo can easily be distinguished by the characteristic morphology of the ejaculatory complex (proximal chamber large, proximal and distal arms short, carina anterior short versus normally-developed with small proximal chamber in the species from Java (see Lundblad 1956, 1971). There appear to be two forms of this species which can be identified in the female sex. At one extreme end, females with an enlarged genital field and less extended postgenital area (Fig. 40 B). At the other extreme end specimens with a less enlarged genital field and more extended postgenital area (Fig. 40 C). The figures 32 A – B illustrate two specimens collected from the locus typicus. The specimens from Little Lumotok stream and Kemantis stream (Table 5) differ from the type specimens by broader frontal plates (Fig. 33 B, 39 D – E), slightly longer medial suture line of Cx-II + III, smaller dimensions of genital field and broader I-L- 6 (Fig. 33 H) These differences could reflect some degree of genetical differentiation between these populations. Unfortunately, our effort to obtain sequences of the mtCOI fragment from different populations of this species from Borneo were not successful. On the other hand introducing a new species for both forms, in our opinion, will create more confusion given the present state of knowledge of this species. Habitat. Sandy / bouldery streams, shaded by rain forest (Figs. 43 A, C – D).	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E51FFF6FF0DF94ED4A0FC5D.taxon	distribution	Distribution. Borneo.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E5CFFF9FF0DFF2DD4A0F81A.taxon	description	(Figs. 30 G, 34 A – G, 35 A – D, 39 F – G, 40 F – G)	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E5CFFF9FF0DFF2DD4A0F81A.taxon	materials_examined	Type series. Holotype female, dissected and slide mounted, Malaysia, Borneo, stream Kemantis, Sayap, Mt Kinabalu, 6 º 09.841 N, 116 º 33.936 E, alt. 928 m asl., 16. ix. 2012 Smit. Paratypes: one male, Great Lumotok stream, Mt Kinabalu, 6 º 09.336 N, 116 º 34.056 E, alt. 1087 m asl., 17. ix. 2012 Smit, dissected and slide mounted.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E5CFFF9FF0DFF2DD4A0F81A.taxon	diagnosis	Diagnosis. Idiosoma elongated-oval (dorsal shield L / W ratio 1.3); frontal plates broad (L / W ratio 1.4), shorter than shoulder plates (shoulder / frontal plate L ratio 1.3); gnathosomal bay V-shaped; ventral setae on P- 4 short. Male: medial margin of Cx-II / III relatively short; ejaculatory complex conventional in shape with small proximal chamber.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E5CFFF9FF0DFF2DD4A0F81A.taxon	description	Description General features — Idiosoma elongated-oval; frontal plates broad (Fig. 34 B), shorter than shoulder plates; frontal margin medially slightly convex, between flat, anterolaterally pointed protrusions (Figs. 34 A, 35 A); Cxgl- 4 located far anteriorly, near tips of Cx-I; gnathosomal bay V-shaped, proximally pointed (Figs. 34 C, 12 C); suture line of Cx-IV distinct, originating from the lateral edge of the genital field and extending posteriorly beyond the posterior margin of the genital field; excretory pore and Vgl- 2 away from the line of primary sclerotization, Vgl- 2 slightly posterior to excretory pore; gnathosoma with long dorsal apodemes, rostrum truncated (Fig. 34 G); P- 2 and P- 3 distal margins without pointed tips, P- 4 ventral setae short, not reaching the tip of P- 5, with small denticles near insertion (Figs. 34 D – E). Male: medial margin of Cx-II / III relatively short; genital field subrectangular; ejaculatory complex: proximal and distal arms well developed, carina anterior long, proximal chamber small (Fig. 30 G). Female: genital field large and pentagonal in shape, anteriorly enlarged, laterally slightly convex, tapering posteriorly. Measurements Female (holotype) — Idiosoma (ventral view: Fig. 40 F) L 813, W 647; dorsal shield (Figs. 34 B, 39 F) L 741, W 572, L / W ratio 1.3; dorsal plate L 641; shoulder plate L 211 – 213, W 103, L / W ratio 2.05 – 2.06; frontal plate L 166, W 116, L / W ratio 1.43, L shoulder / frontal plate ratio 1.27 – 1.28. Gnathosomal bay L 111, Cx-I total L 289, Cx-I mL 178, Cx-II + III mL 43; ratio Cx-I L / Cx-II + III mL 6.7; Cx-I mL / Cx-II + III mL 4.1. Genital field L / W 198 / 175, ratio 1.13; distance genital field-excretory pore 163, genital field-caudal idiosoma margin 275. Gnathosoma vL 154; palp: total L 184, dL / H, dL / H ratio: P- 1, 29 / 21 - 22, 1.36; P- 2, 47 / 33.5, 1.4; P- 3, 35 / 28, 1.28; P- 4, 48 / 20, 2.4; P- 5, 25 / 11 – 12, 2.1; P- 2 / P- 4 ratio 0.98. Legs: dL of I-L- 2 – 6: 68, 72, 91, 100, 103, I-L- 6 H 37, dL / H I-L- 6 ratio 2.8. Male (paratype) — Idiosoma (ventral view: Fig. 40 G) L 781, W 634; dorsal shield (Figs. 35 B, 39 G) L 725, W 556, L / W ratio 1.3; dorsal plate L 640; shoulder plate L 213 – 222, W 100 – 109, L / W ratio 2.0 – 2.1; frontal plates fused to form an elongated platelet, L / W 319 / 123, ratio 2.6. Gnathosomal bay L 116, Cx-I total L 288, Cx-I mL 172, Cx-II + III mL 72; ratio Cx-I L / Cx-II + III mL 4.0; Cx-I mL / Cx-II + III mL 2.34. Genital field L / W 163 / 138, ratio 1.18; ejaculatory complex L 228; distance genital field-excretory pore 158, genital field-caudal idiosoma margin 256. Palp: total L 187, dL / H, dL / H ratio: P- 1, 30 – 31 / 21, 1.45; P- 2, 49 / 34, 1.45; P- 3, 35 / 26, 1.35; P- 4, 48 / 19, 2.6; P 5, 25 / 11, 2.3; P- 2 / P- 4 ratio 1.02. Legs: dL of I-L- 2 - 6: 65, 72, 88, 97, 99, I-L- 6 H 36, dL / H I-L- 6 ratio 2.8.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E5CFFF9FF0DFF2DD4A0F81A.taxon	etymology	Etymology. The species is named after Phantasos (Ancient Greek: Φάντασος) who was a spirit of dreams of fantasy, and who takes the shape of inanimate objects. The species name is a noun in apposition (in the nominative case), despite the Recommendation 31 A of the ICZN (1999) about avoidance of personal names as nouns in appositions, because there is no case for it being confusing or misleading.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E5CFFF9FF0DFF2DD4A0F81A.taxon	discussion	Discussion. The combination of a broad frontal plates (L / W ratio 1.4) and moderately deep, V-shaped, gnathosomal bay, easily separates the new species from all other members of the genus. Due to the V-shaped, gnathosomal bay, Monatractides phantasos n. sp. somehow resembles M. oxystomus (K. Viets, 1935) and M. tobaensis (K. Viets, 1935) (see above). However these two species seem not to be closely related to the new species as they differ in a series of features: less broader frontal plates, frontal margin medially strongly concave between large anterolaterally pointed protrusions, gnathosomal bay more deep, moderately long medial margin of Cx-II / III, P- 4 with long ventral seta exceeding distal edge and excretory pore and Vgl- 2 shifted more away from the line of primary sclerotization. Variability. Frontal plates in male fused to form an elongated platelet, but traces of fusion visible (Figs. 35 B, 39 G). Habitat. Sandy / bouldery streams, shaded by rain forest (Fig. 43 D).	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E5CFFF9FF0DFF2DD4A0F81A.taxon	distribution	Distribution. Borneo.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E5EFFFAFF0DF9D7D634FBA6.taxon	description	(Figs. 30 H, 36 A – G, 39 H, 40 H)	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E5EFFFAFF0DF9D7D634FBA6.taxon	materials_examined	Type series. Holotype male, dissected and slide mounted, Malaysia, Borneo, Little Lumotok stream, Sayap, Mt Kinabalu, 6 º 09.467 N, 116 º 34.027 E, alt. 1069 m asl., 17. ix. 2012 Smit.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E5EFFFAFF0DF9D7D634FBA6.taxon	diagnosis	Diagnosis. Male. Idiosoma elongated (dorsal shield L / W ratio 1.4); frontal and shoulder plates fused with the large dorsal plate forming an undivided dorsal shield; gnathosomal bay V-shaped, proximally pointed, three pairs of knob-like protrusions at the lateral margins of gnathosomal bay; medial margin of Cx-II / III relatively long.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E5EFFFAFF0DF9D7D634FBA6.taxon	description	Description Male. General features — Idiosoma elongated; frontal and shoulder plates fused with the large dorsal plate forming an undivided dorsal shield (Fig. 36 A); frontal margin medially slightly convex, between flat protrusions (Fig. 36 C); gnathosomal bay V-shaped, proximally pointed, three pairs of knob-like protrusions at the lateral margins of gnathosomal bay; medial margin of Cx-II / III relatively long; genital field subrectangular; ejaculatory complex (Fig. 30 H) normal in shape (proximal and distal arms and proximal chamber well developed); suture line of Cx-IV distinct, originating from lateral edge of genital field; excretory pore and Vgl- 2 well separated from the line of primary sclerotization, Vgl- 2 slightly posterior to excretory pore; gnathosoma elongated in lateral view, rostrum truncated (Fig. 36 F); P- 2 shorter than P- 4, distal margin of P- 3 and P- 4 medially and laterally with several pointed extensions; P- 4 with two ventral setae, one very long and away from distal edge (Figs. 36 D – E). Measurements — Idiosoma (ventral view: Figs. 36 B, 40 H) L 727, W 553; dorsal shield (Figs. 36 A, 39 H) L 688, W 505, L / W ratio 1.36. Gnathosomal bay L 106, Cx-I total L 266, Cx-I mL 160, Cx-II + III mL 113; ratio Cx-I L / Cx-II + III mL 2.35; Cx-I mL / Cx-II + III mL 1.4. Genital field L / W 146 / 120, ratio 1.21; ejaculatory complex L 168; distance genital field-excretory pore 144, genital field-caudal idiosoma margin 200. Gnathosoma vL 137; chelicera total L 160; palp total L 191, dL / H, dL / H ratio: P- 1, 27 / 21, 1.3; P- 2, 46 / 35, 1.32; P- 3, 35 / 28, 1.46; P- 4, 51 / 19, 2.67; P- 5, 32 / 13.5, 2.4; P- 2 / P- 4 ratio 0.91. Legs: dL of I-L- 2 - 6: 55, 65, 79, 77, 85, I-L- 6 H 29, L / H I-L- 6 ratio 2.9. Female: unknown	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E5EFFFAFF0DF9D7D634FBA6.taxon	etymology	Etymology. The species is named after Phobetor (Ancient Greek: Φοβήτωρ) who was a spirit of nightmares and who takes the shape of animals. The species name is a noun in apposition (in the nominative case), despite the Recommendation 31 A of the ICZN (1999) about avoidance of personal names as nouns in appositions, because there is no case for it being confusing or misleading.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E5EFFFAFF0DF9D7D634FBA6.taxon	discussion	Discussion. Monatractides phobetor n. sp is closely related to M. synaptus (K. Viets, 1935), M. amabilis (Lundblad, 1941) and M. magnipharynx (Lundblad, 1941), all known from Java (K. Viets 1935, Lundblad 1971) characterized by the fusion of both frontal and shoulder platelets with the large dorsal plate. Due to the slender ventral setae on P- 2 and P- 3 the new species resembles M. amabilis (and differs from the two other species in having a denticle-like ventral seta on P- 2 and - 3). The new species can easily be distinguished from all abovementioned species by having V-shaped, proximally pointed gnathosomal bay with three pairs of knob-like protrusions at its lateral margins. Habitat. Sandy / bouldery stream, shaded by rain forest.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E5EFFFAFF0DF9D7D634FBA6.taxon	distribution	Distribution. Borneo; known only from the locus typicus.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E5FFFFCFF0DFB19D634F881.taxon	description	(Figs. 30 M, 37 A – C, 38 A – D)	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E5FFFFCFF0DFB19D634F881.taxon	materials_examined	Type series. Holotype male, dissected and slide mounted, Malaysia, Borneo, Little Lumotok stream, Sayap, Mt Kinabalu, 6 º 09.467 N, 116 º 34.027 E, alt. 1069 m asl., 17. ix. 2012 Smit.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E5FFFFCFF0DFB19D634F881.taxon	diagnosis	Diagnosis. Idiosoma large in size (> length 1300 µm) and roundish (dorsal shield L / W ratio 1.2); gnathosomal bay deep, narrow Y-shaped; medial margin of Cx-II / III relatively long; genital field anteriorly enlarged, laterally straight, tapering posteriorly; ejaculatory complex normal in shape (Fig. 30 M); palp with thick, pennate setae on P 2 and P- 3, distal margin of P- 2, P- 3 and P- 4 medially with several pointed extensions; ventral setae on P- 4 thickened and short.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E5FFFFCFF0DFB19D634F881.taxon	description	Description Male. General features — Idiosoma large in size and roundish; frontal plates broad (L / W ratio 1.8), shorter than shoulder plates (shoulder / frontal plate L ratio 1.16 – 1.2); frontal margin medially slightly convex between flat, anterolaterally directed protrusions (Fig. 37 A); Cxgl – 4 located far anteriorly, near tips of Cx-I; the tips of Cx-I orientated to the lateral side, gnathosomal bay deep, narrow Y-shaped (Fig. 37 C); medial margin of Cx-II / III relatively long; genital field anteriorly enlarged, laterally straight, tapering posteriorly; ejaculatory complex (Fig. 30 M) normal in shape (proximal and distal arms and proximal chamber well developed); posterior suture line of Cx-IV distinct, extending posteriorly beyond posterior margin of genital field, medially approaching and parallel to each other; line of primary sclerotization close to posterior margin of genital field; excretory pore and Vgl – 2 well separated from the line of primary sclerotization, Vgl – 2 posterior to excretory pore; gnathosomal rostrum truncated (Fig. 38 D); palp with thick, pennate setae on P- 2 and P- 3, distal margin of P- 2, P- 3 and P- 4 medially with several pointed extensions (Figs. 38 A – B); ventral setae on P- 4 short, not reaching the tip of P- 5. Measurements — Idiosoma (ventral view: Fig. 37 C) L 1363, W 1067; dorsal shield (Fig. 37 B) L 1131, W 944, L / W ratio 1.2; dorsal plate L 1006; shoulder plate L 322 – 325, W 147, L / W ratio 2.2; frontal plate L 272 – 277, W 153 – 158, L / W ratio 1.76 – 1.78; L shoulder / frontal plate ratio 1.16 – 1.2. Gnathosomal bay L 253, Cx-I total L 438, Cx-I mL 184, Cx-II + III mL 160; ratio Cx-I L / Cx-II + III mL 2.74; Cx-I mL / Cx-II + III mL 1.15. Genital field L / W 248 / 194, ratio 1.28; ejaculatory complex L 256; distance genital field-excretory pore 338, genital field-caudal idiosoma margin 497. Gnathosoma vL 204; chelicera total L 237; palp total L 294, dL / H, dL / H ratio: P- 1, 39 / 25, 1.6; P- 2, 75 / 46, 1.63; P- 3, 78.5 / 40, 1.96; P- 4, 67 / 23, 2.9; P- 5, 34 / 14, 2.44; P- 2 / P- 4 ratio 1.1. Legs: dL of I-L- 5 – 6: 126, 100, I-L- 6 H 25.4, dL / H I-L- 6 ratio 3.9. Female: unknown	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E5FFFFCFF0DFB19D634F881.taxon	etymology	Etymology. Named after Heracles (Ancient Greek: ρακλῆς), who was the greatest of the Greek heroes and a paragon of masculinity. The species name is a noun in apposition (in the nominative case), despite the Recommendation 31 A of the ICZN (1999) about avoidance of personal names as nouns in appositions, because there is no case for it being confusing or misleading.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E5FFFFCFF0DFB19D634F881.taxon	discussion	Discussion. The male of Monatractides heracles n. sp. is easily distinguished from other members of the genus due to the combination of a large idiosoma dimensions (> length 1300 µm) and a narrow Y-shaped gnathosomal bay. Further diagnostic features of the new species include the relatively long medial suture line of Cx-II + III, posterior suture line of Cx-IV extending beyond posterior margin of genital field and medially approaching to each other, and by the shape of the palp with thick, pennate setae on P- 2 and P- 3, distal margin of P 2 and P- 3 medially with several pointed extensions and a thickened and short ventral setae on P- 4. Habitat. Sandy / bouldery stream, shaded by rain forest.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E5FFFFCFF0DFB19D634F881.taxon	distribution	Distribution. Borneo; known only from the locus typicus.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E5AFFFFFF0DF914D72AF826.taxon	description	(Figs. 39 I, 40 I, 41 A – F)	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E5AFFFFFF0DF914D72AF826.taxon	materials_examined	Type series. Holotype female, dissected and slide mounted, Malaysia, Borneo, Great Lumotok stream, Mt Kinabalu, 6 º 09.336 N, 116 º 34.056 E, alt. 1087 m asl., 17. ix. 2012 Smit. Paratypes: three females, Mahua stream, Mahua, Crocker Range, 5 º 47.838 N, 116 º 24.510 E, alt. 1052 m asl., 21. ix. 2012 Smit.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
F535879E7E5AFFFFFF0DF914D72AF826.taxon	diagnosis	Diagnosis. Female. Idiosoma small in size (length <600 µm) and elongated (dorsal shield L / W ratio 1.4); suture line of Cx-IV nearly obliterated; P- 2 and P- 3 distal margins without pointed tips, P- 4 with ventral setae, one of them longer and reaching tip of P- 5.	en	Pešić, Vladimir, Smit, Harry (2014): Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo. Zootaxa 3840 (1): 1-72, DOI: http://dx.doi.org/10.11646/zootaxa.3840.1.1
