taxonID	type	description	language	source
219A9352B86056889DBEB0947CC84DEA.taxon	description	Figs 1, 2, 12 A-C, 13 A, 14 A, F	en	Lantsov, Vladimir I., Pilipenko, Valentin E. (2021): A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group. ZooKeys 1048: 145-175, DOI: http://dx.doi.org/10.3897/zookeys.1048.67564, URL: http://dx.doi.org/10.3897/zookeys.1048.67564
219A9352B86056889DBEB0947CC84DEA.taxon	materials_examined	Material examined. Georgia • 1 male, 3 females; " ushch. Bani-Khevi bl. Borzhomi, Adzharo-Imeretinsk. khr. [Gorge Bani-Khevi near Borjomi, Adjaro-Imeretinsk. Ridge]; 26 Jun. 1958; Kurnakov leg.; ZISP • 1 male; " Tskhra-Tskharo bliz Bakuriani " [Tskhra-Tskharo near Bakuriani]; 29 Jun. 1958; Kurnakov leg.; ZISP • 1 male; " Leberde, Mengreliya, Gruziya " [Leberde, Mengrelia, Georgia]; 17 Jul. 1959; Savenko leg.; ZISP • 1 male; " Adzharo-Imeretinsk. khr. Gory u Sairme, Gruziya " [Adjaro-Imeretinsk. Ridge, mountains near Sairme, Georgia]; 23 May 1958; Kurnakov; ZISP • 2 males, 1 female; " ushch. Tsvaniat-Khevi Adzharo-Imeretinsk. khr. Gruziya " [Tsvaniat-Khevi Gorge, Adjaro-Imeretinsk. Ridge, Georgia]; 23 Jun. 1958; Kurnakov leg; / " Vysokotrave u verkhn. granitsy lesa u ruchya " [High grass at the edge of the border of the forest by the stream]; ZISP • 2 males; " s. Otkhara, Gudautsk. r-n, Abkhaziya, ber. rechki, kusty. " [village Otkhara, Gudautsk. District, Abkhazia, river bank, bushes]; 30 Jun. 1958; Kurnakov; ZISP • 2 males; " Abkhaziya, s. Otkhara predgorya Bzybskogo khr. " [Abkhazia, village Otkhara foothills of the Bzybsk Ridge]; 20 May 1958; Kurnakov; / " Dubovo-grabovyi les, kusty Asalia " [Oak-hornbeam forest, bushes Asalia "]; ZISP; Azerbaijan • 1 male; " Belokany, khr. Akhkemal. Azerb. 200 m " [Belokany, Akhkemal Ridge. Azerb. [aidjan] alt. 200 m]; 15 Jun. 1964; Pastukhov leg.; ZISP; Russia - Krasnodarskiy kray • 1 male; Sotshi prov. Pontica; 6 May 1932; B. Rohdendorf leg.; / " T. Tipula caucasica Ried det. Lacksch. " [terminalia in Canada balsam pinned with specimen]; ZISP • 1 female, Khosta, Caucasian Reserve, Tiso-samshitovaya rosha [Yew-and-Boxwood Tree Grove]; 43 ° 31 ' 656 " N, 39 ° 52 ' 467 " E; alt. 54 m; collected at light, 8 May 2018; V. Lantsov leg.; IEMT • 2 males, 2 females; same collection data as for preceding, 43 ° 31 ' 688 " N, 39 ° 52 ' 561 " E; alt. 35 m; 8 May 2018; V. Lantsov leg. / Lime-beech (Tilia begoniifolia and Fagus orientalis) forest with yew (Taxus baccata) and hornbeam (Carpinus betulus) in tree layer and butcher's-broom (Ruscus colchicus - dominant and Ruscus aculeatus) in ground layer [one of the females was collected in the same community in a damp place near a yew log]; IEMT • 2 males, 2 females, same collection data as for preceding, 43 ° 31 ' 780 " N, 39 ° 52 ' 518 " E; alt. 98 m; 9 May 2018; V. Lantsov leg.; / Hornbeam - ash (Carpinus orientalis, Fraxinus excelsior) forest with addition of yew (Taxus baccata) and lime (Tilia begoniifolia) with blackberry (Rubus sanctus) and fig (Ficus carica) in shrub layer and with butcher's-broom (Ruscus colchicus) in ground layer; ZISP • 4 males, 4 females, same collection data as for preceding, 43 ° 31 ' 775 " N, 39 ° 52 ' 423 " E; alt. 101 m; 9 May 2018; V. Lantsov leg.; / Hornbeam (Carpinus orientalis) forest with butcher's-broom (Ruscus colchicus) and fern (Asplenium scolopendrium) in ground layer; ZISP • 1 male and 1 female (in copula), 1 female; same collection data as for preceding, 43 ° 31 ' 913 " N, 39 ° 52 ' 463 " E; alt. 153 m; 9 May 2018, V. Lantsov leg. / Ash-lime (Fraxinus excelsior + Tilia begoniifolia) forest with hornbeam (Carpinus betulus) and with black berry (Rubus anatolicus) and miscellaneous herbs in ground layer; IEMT • 2 males, 1 female, same collection data as for preceding, 43 ° 31 ' 825 " N, 39 ° 52 ' 653 " E; alt. 99 m; 10 May 2018, V. Lantsov leg.; IEMT • 1 male, same collection data as for preceding, 43 ° 31 ' 656 " N, 39 ° 52 ' 467 " E; alt. 54 m; V. Lantsov leg.; ZISP • 2 males, 1 female, same collection data as for preceding, 43 ° 32 ' 257 " N, 39 ° 52 ' 682 " E; alt. 94 m; 10 May 2018, V. Lantsov leg.; / beech (Fagus orientalis) forest with addition of yew (Taxus baccata) and hornbeam (Carpinus betulus) with bunch grass (Calamagrostis arundinacea) in ground layer; IEMT • 6 males, 3 females; Krasnodar Territory, environs Shakhe River higher than Salokh-Aul; 5 - 8 Jun. 2011; V. Pilipenko leg.; VPMC; Russia - Dagestan • 1 female, " Khochal-Dag, alp. obl. Dagst. [Dagestan] " [Hochal-Dag, alp. Region Dagst. [Dagestan ]]; 29 Jun. 1909 Mlokosevich leg.; ZISP • 1 female; Tlyaratinsky district, near village Salda, stream on the right bank of the river Dzhurmut; 41 ° 58 ' 388 " N, 46 ° 30 ' 446 " E; alt. 1792 m; 2 Jul. 2016, V. Lantsov leg.; / " Oak (Quercus iberica) sparse forest with Populus tremula and Acer platanoides in tree layer, Rosa canina, Spiraea crenata and Cotoneaster integerrimus in shrub, and clover (Trifolium canescens) and miscellaneous herbs in ground layer; ZISP • 2 males, 1 female; near village Betelda, stream on the left bank of the river Dzhurmut; 41 ° 56 ' 853 " N, 46 ° 32 ' 424 " E; alt. 1850 m; 3 Jul. 2016, V. Lantsov leg.; / Willow (Salix caprea) shrub with herbaceous layer (Epilobium montanum) (dominant), Cardamine seidlitziana, Mentha caucasica, Veronica anagallis-aquatica and Myosoton aquaticum community near spring above the flood plain of the left bank of the river; ZISP • 2 males; near village Genekolob, on the high left bank of the river Dzhurmut; 41 ° 57 ' 392 " N, 46 ° 31 ' 276 " E; alt. 2011 m; 4 Jul. 2016, V. Lantsov leg.; / Oak (Quercus iberica) forest with mountain-ash (Sorbus aucuparia), wayfaring tree (Viburnum lantana) in shrub layer [the lower border of a forest and a forb-cereal meadow]; ZISP • 2 males; same collection data as for preceding, 41 ° 57 ' 449 " N, 46 ° 31 ' 228 " E; alt. 1977 m; 4 Jul. 2016, V. Lantsov leg.; ZISP • 2 males, between the villages of Gerel and Genekolob, the right bank of the river Dzhurmut, 41 ° 55 ' 619 " N, 46 ° 34 ' 468 " E; alt. 1925 m; 5 Jul. 2016, V. Lantsov leg.; / Above the floodplain terrace with a rare herbaceous birch (Betula raddeana) forest with willow (Salix caprea) (coppice forest); ZISP.	en	Lantsov, Vladimir I., Pilipenko, Valentin E. (2021): A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group. ZooKeys 1048: 145-175, DOI: http://dx.doi.org/10.3897/zookeys.1048.67564, URL: http://dx.doi.org/10.3897/zookeys.1048.67564
219A9352B86056889DBEB0947CC84DEA.taxon	diagnosis	Diagnosis. Dark stripes medially on lighter background extending entire length of scutum. Inner gonostylus posteriorly with wedge-shaped projection directed anteriorly. Tergite 9 at apex with three rounded notches, middle one usually deepest and widest.	en	Lantsov, Vladimir I., Pilipenko, Valentin E. (2021): A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group. ZooKeys 1048: 145-175, DOI: http://dx.doi.org/10.3897/zookeys.1048.67564, URL: http://dx.doi.org/10.3897/zookeys.1048.67564
219A9352B86056889DBEB0947CC84DEA.taxon	distribution	Distribution. Endemic to the Caucasus. It was recently noted for Russia (Dagestan) for the first time (Lantsov 2020). Currently known from the West Caucasus (first record), East Caucasus and Transcaucasus.	en	Lantsov, Vladimir I., Pilipenko, Valentin E. (2021): A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group. ZooKeys 1048: 145-175, DOI: http://dx.doi.org/10.3897/zookeys.1048.67564, URL: http://dx.doi.org/10.3897/zookeys.1048.67564
35B73B80118E5247AD3A67AD3A54266C.taxon	description	Figs 3, 4, 5, 13 B, 14 B, G	en	Lantsov, Vladimir I., Pilipenko, Valentin E. (2021): A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group. ZooKeys 1048: 145-175, DOI: http://dx.doi.org/10.3897/zookeys.1048.67564, URL: http://dx.doi.org/10.3897/zookeys.1048.67564
35B73B80118E5247AD3A67AD3A54266C.taxon	materials_examined	Material examined. Holotype: Russia • 1 male; Krasnodarskiy Kray, Sochi env. Psekhako Mt., 43 ° 41 ' 28 " N, 40 ° 22 ' E; alt. ~ 2000 m; 14 - 18 Jun. 2008; K. Tomkovich leg.; ZISP. Holotype in good condition; however, left antenna, front right and left hind legs missing. Paratypes: Russia • 1 male; same data as for holotype • 2 males; Krasnodarskiy Kray, Kamyshanova Polyana env. [Biological station of Krasnodar State University], 44 ° 16 ' 91 " N, 40 ° 04 ' 46 " E; alt. 1200 m; 26 Jul. 2018; V .. Pilipenko leg.; ZISP.	en	Lantsov, Vladimir I., Pilipenko, Valentin E. (2021): A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group. ZooKeys 1048: 145-175, DOI: http://dx.doi.org/10.3897/zookeys.1048.67564, URL: http://dx.doi.org/10.3897/zookeys.1048.67564
35B73B80118E5247AD3A67AD3A54266C.taxon	diagnosis	Diagnosis. Male. General coloration grey with silver pruinescence. Tergite 9 with deep rounded notches either side of slightly grooved cone-shaped projection. Sternite 8 with a pair of small appendages, each bearing a long medially curved spine, broad base of appendages with fringe of long whitish hairs. Outer gonostylus small, triangular, slightly thickened distally, covered with long setae. Inner gonostylus with small rod-like outgrowth in middle of outer edge.	en	Lantsov, Vladimir I., Pilipenko, Valentin E. (2021): A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group. ZooKeys 1048: 145-175, DOI: http://dx.doi.org/10.3897/zookeys.1048.67564, URL: http://dx.doi.org/10.3897/zookeys.1048.67564
35B73B80118E5247AD3A67AD3A54266C.taxon	description	General description. Adult (Fig. 3 A). Male body length 15.7 mm, wing length 16.2 mm, haltere 2.5 mm, nasus 0.13 mm, rostrum 0.8 mm, scape 0.6 mm, pedicel 0.15 mm, 1 st flagellomere 0.5 mm. Length (mm) of leg segments, fore (1), mid (2), and hind (3); successively femur, tibia, 1 st and 2 nd tarsomeres: 1 (9.7; 10.4; 8.5; 3.8), 2 (9.8; 10.5; 9.0; 4.5) and 3 (10.0; 13.0; 9.0; 4.5). Length (mm) of 3 rd, 4 th, and 5 th tarsomeres of all three pairs of legs, approximately the same: 1.0; 0.6; 0.5 mm, respectively.	en	Lantsov, Vladimir I., Pilipenko, Valentin E. (2021): A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group. ZooKeys 1048: 145-175, DOI: http://dx.doi.org/10.3897/zookeys.1048.67564, URL: http://dx.doi.org/10.3897/zookeys.1048.67564
35B73B80118E5247AD3A67AD3A54266C.taxon	description	Description. Head (Figs 3, 13 B) grey with silvery pubescence. Rostrum grey dorsally, ventrally yellow-brown with sparse light rusty setae in black sockets. Nasus distinct, appearing triangular in dorsal aspect, covered with long, sparse procumbent setae. Frons and gula bare. Vertex (Fig. 13 B) framed by rows of sparse setae alongside eye, glabrous in center with indistinct narrow longitudinal dark grey line. Genae, including lateral part of rostrum, with very sparse fine pale bristles. Tempora and postgenae with sparse black setae. Antennae, bent backwards, reaches base of abdomen. Scape grey with silvery pruinescence, pedicel dirty yellow brown. Flagellomeres dark brown with verticils on slightly thickened bases, longest verticils subequal to length of respective flagellomere. Thorax (Fig. 3 A, B) grey with silvery pruinescence, and short sparse whitish hairs. Pronotum dark brown with a weak dark stripe medially, and short sparse whitish bristles. Scutum (Fig. 3 B) with four brownish grey longitudinal stripes, lateral stripe shorter, medial stripes narrowly separated by dirty yellow area. Katepisternum, anepisternum, katepimeron, anepimeron and meron grey, bare, with silvery pruinescence; scutellum and mediotergite light grey with silvery pruinescence; scutal lobes with wide dark grey stripe, bordered by sparse whitish setae and with long yellow setae anterolaterally at wing base. Scutellum framed anteriorly with brown, with sparse yellow setae laterally, and with vague narrow dark grey medial line. Mediotergite light grey with sparse yellow setae and with silvery pruinescence. Laterotergite grey with silvery pruinescence and sparse erect short brown setae. Wings (Fig. 3 A) with typical venation for the subgenus, transparent with brownish tin, (viewed in transmitted light). Stigma dirty yellow. Oblique lunule proximal to stigma. Setae on costal and subcostal veins; group of short black macrotrichia distally on R and base of R 1. Distal section of Rs, R 4 + 5, r-m, base of pentagonal discal cell lighter and weakened. Stem of cell m 1 approximately half length of cell. Apex of vein Cu 1 sharply curved at wing margin. Halteres. Stem grey with base covered with short white setae, knob dark grey. Legs. Coxae light grey with silvery pruinescence; laterally with long sparse whitish yellow setae; medially with very short yellowish setae. Trochanters yellow with long yellow setae. Femora brown, narrowly yellow at base, covered with adpressed short brown setae; tibiae brownish; tarsal segments dark brown with short, adpressed brown setae. Tibial spur formula 1 - 2 - 2. Tarsal claws with a small tubercle at the base (magnification 10 x 4.5); with short but clearly visible arolium between ones. Abdomen (Figs 3 A, 4 A). Tergites and sternites grey with silvery pruinescence, and sparse fine short golden setae. Tergites with lateral dark grey intermittent stripe, no distinct dorsal stripe. Tergites laterally and distally edged with whitish-yellow, proximal segments with thinner distal edge, becoming wider on caudal segments. Terminalia (Figs 4, 5, 14 B, G). Hypopygium grey to dark grey, moderately thickened, with silvery pruinescence (Fig. 4 A-C). Tergite 9 with two deep rounded notches flanking a median cone-shaped projection, the apex of which has a small groove (Figs 4 E, F; 5 G-L, 14 B). The dorsolateral edges of the tergite 9 convex, curved inward. Sternite 8 (Fig. 4 D) with a pair of small appendages bearing long medially curved spine, base of appendages with fringe of long whitish hairs not obscuring gap between. Appendages of sternite 9 (Fig. 4 K, L) abundantly pubescent. Outer gonostylus (Figs 4 G, 5 A-F, 14 G) small, triangular, slightly thickened distally, covered with long bristles. Inner gonostylus (Figs 4 G, H; 5 A-F, 14 G) with small rod-shaped outgrowth in middle of outer edge (Figs 4 G, 14 G), twice as long as wide at base, slightly narrower medially, tip obliquely truncate. Posterior part of inner gonostylus a wide triangular plate with short sparse brown bristles in inner surface, long whitish setae in outer one. Gonocoxite (Figs 4 G, 5 A-C) with two wide dentate outgrowths, ventral one longer, pointed, dorsal one rounded; long yellowish setae laterally on gonocoxite. Gonocoxal fragment (Fig. 4 I) with short forked extension at base. Semen pump and aedeagus as in Figure 4 J. Female unknown.	en	Lantsov, Vladimir I., Pilipenko, Valentin E. (2021): A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group. ZooKeys 1048: 145-175, DOI: http://dx.doi.org/10.3897/zookeys.1048.67564, URL: http://dx.doi.org/10.3897/zookeys.1048.67564
35B73B80118E5247AD3A67AD3A54266C.taxon	distribution	Distribution. Endemic to the Caucasus: currently known from the West Caucasus.	en	Lantsov, Vladimir I., Pilipenko, Valentin E. (2021): A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group. ZooKeys 1048: 145-175, DOI: http://dx.doi.org/10.3897/zookeys.1048.67564, URL: http://dx.doi.org/10.3897/zookeys.1048.67564
35B73B80118E5247AD3A67AD3A54266C.taxon	etymology	Etymology. Tipula (Lunatipula) eleniya sp. nov. is named after the mother of the first author, Elena Nikolaevna Lantsova.	en	Lantsov, Vladimir I., Pilipenko, Valentin E. (2021): A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group. ZooKeys 1048: 145-175, DOI: http://dx.doi.org/10.3897/zookeys.1048.67564, URL: http://dx.doi.org/10.3897/zookeys.1048.67564
0273951C175D51F098B012BAF0F9EB34.taxon	description	Figs 6, 7, 13 C, 14 C, I	en	Lantsov, Vladimir I., Pilipenko, Valentin E. (2021): A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group. ZooKeys 1048: 145-175, DOI: http://dx.doi.org/10.3897/zookeys.1048.67564, URL: http://dx.doi.org/10.3897/zookeys.1048.67564
0273951C175D51F098B012BAF0F9EB34.taxon	materials_examined	Material examined. Holotype: Georgia • 1 male; " Zagor [Zagar Svanetiya], Gruz. SSR " [Zagor Pass Svanetiya, Gruz. SSR. Georgian Soviet Socialist Republic]; alt. 2623 m; 19 Jul. 1957; R. Savenko leg.; ZISP. / " Tipula 4 - dentata Tipula paupera ssp. n. det. Savchenko " / " Holotypus " [Holotype not initially marked; red label] / " Tipula (Lunatipula) paupera Sav., stat. nov. Lantsov, Pilipenko, 2020 " [white label]. Thorax (prescutum, scutum, scutellum) smeared, obscuring coloration. Preservation of legs: forelegs both with only trochanters; midlegs: left with part of femur, right only with femur; hind legs: right missing. Wings slightly crumpled. Paratypes. Georgia • 1 male; " ur. Laedil (Koruldash [Koruldashi] - Zagar) " [Laedil [territory] (Koruldash [Koruldashi] - Zagar)]; 5 Aug. 1957; Savenko leg.; ZISP. / " Paratypus " [Paratype not initially marked; red label] / " Tipula (Lunatipula) paupera Sav., stat. nov. Lantsov, Pilipenko, 2020 " [white label] • 2 males; " Spusk s perevala Bassa v Nakru [v dolinu r. Nakra], Stavropolskii kr. " [Descent from the Bassa Pass to Nakra [to the valley of the river Nakra], Stavropol kr. [Stavropol Territory - in error, Georgia, Svanetiya]; 5 Aug. 1956; L. Arens leg.; ZISP. / " Paratypus " [Paratype not initially marked; red label] / " Tipula (Lunatipula) paupera Sav., stat. nov. Lantsov, Pilipenko, 2020 " [white label].	en	Lantsov, Vladimir I., Pilipenko, Valentin E. (2021): A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group. ZooKeys 1048: 145-175, DOI: http://dx.doi.org/10.3897/zookeys.1048.67564, URL: http://dx.doi.org/10.3897/zookeys.1048.67564
0273951C175D51F098B012BAF0F9EB34.taxon	diagnosis	Diagnosis. Male. Gonocoxite with two elongate, pointed teeth, one dorsally and one ventrally. Tergite 9 with two projections posteriorly, separated by deep wide notch. Paired appendages of sternite 8 widely spaced, base shorter than wide, gap between not masked by setae. Apical appendages of sternite 9 elongate, narrow distally, with dense bundle of relatively short golden yellow setae at tip.	en	Lantsov, Vladimir I., Pilipenko, Valentin E. (2021): A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group. ZooKeys 1048: 145-175, DOI: http://dx.doi.org/10.3897/zookeys.1048.67564, URL: http://dx.doi.org/10.3897/zookeys.1048.67564
0273951C175D51F098B012BAF0F9EB34.taxon	distribution	Distribution. Endemic to the Caucasus-currently known from the southern slopes of the Greater Caucasus (Georgia).	en	Lantsov, Vladimir I., Pilipenko, Valentin E. (2021): A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group. ZooKeys 1048: 145-175, DOI: http://dx.doi.org/10.3897/zookeys.1048.67564, URL: http://dx.doi.org/10.3897/zookeys.1048.67564
E104034BF2F251DB8C145337C9D914B4.taxon	description	Figs 8, 9, 12 D-F, 13 D, 14 D, J	en	Lantsov, Vladimir I., Pilipenko, Valentin E. (2021): A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group. ZooKeys 1048: 145-175, DOI: http://dx.doi.org/10.3897/zookeys.1048.67564, URL: http://dx.doi.org/10.3897/zookeys.1048.67564
E104034BF2F251DB8C145337C9D914B4.taxon	materials_examined	Material examined. Holotype. Russia • 1 male; " okolitsy Stavropolya, bairachn. [yi] les " [near Stavropol, bairak forest] [small forest in steppe ravines]; 25 May 1954; [S.] Medvedev leg.; ZISP. / " yuzhn. skl. pod pologom [lesa] " [southern slopes under the canopy forest] / " T. (Lunatipula) quadridenta sp. n., opr. E. Savchenko " [det. E. Savchenko]. The specimen is very badly damaged and glued together. The head is glued to the thorax; the abdomen is broken in half and glued together; two legs of uncertain position are glued to the specimen, one without coxa and trochanter, and the other missing the last four tarsal segments. Paratypes. Russia • 1 male; " Staryi les k yugu ot Stavropolya " [Old forest south of Stavropol]; 25 May 1954; [S.] Medvedev leg.; ZISP. / " nizh. chast sev. sklona " [lower part of northern slope] • 1 female, " Georgievskoe lesnichestvo Tuapsinsk. r-n " [Georgievskoe forestry Tuapse District]; 21 May 1954; K. Arnoldi leg.; ZISP • 1 male, same collection data as for preceding; 22 May 1954 • 3 males, 2 females; " g. Lysaya, 800 - 900 m, Tuapsinsk. r-n " [mount Lysaya, alt. 800 - 900 m, Tuapse District]; 26 May 1954; K. Arnoldi leg.; ZISP; / " vershinnaya lugovina i opushka lesa 800 - 900 m " [summit meadow and forest edge, alt. 800 - 900 m].	en	Lantsov, Vladimir I., Pilipenko, Valentin E. (2021): A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group. ZooKeys 1048: 145-175, DOI: http://dx.doi.org/10.3897/zookeys.1048.67564, URL: http://dx.doi.org/10.3897/zookeys.1048.67564
E104034BF2F251DB8C145337C9D914B4.taxon	materials_examined	Additional material. Russia • 9 males, 8 females; " kurort " Goryachii Klyuch " khr. Kotkh. [skii] Krasnodar. [skii] kr. dub. [ovyi] les " [Goryachiy Klyuch resort Koth [skiy] Ridge, Krasnodar. [sky] District oak. Forest]; 18 May 1956; Gilyarov leg.; ZISP / " T. (Lunatipula) Tipula quadridentata sp. n. opr. E. N. Savchenko " • 1 female, " m. [mys] Penai, k yugu ot Novorossiiska " [m. (cape) Penay, south of Novorossiysk] 24. V. [1] 956; Gilyarov leg.; ZISP • 1 male, 1 female; " okr. st. Smolenskoi, Siverskogo r-na, Krasnodar. kr. " [near Smolensk station, Siversky Region, Krasnodar District] 20 May 1963; Savchenko leg.; / " opush. [ka] smesh. [annogo] predgorn. [ogo] lesa " [edge of mixed foothill forest]; ZISP • 1 male; " okr. st. Krivenkovskoi, Tuaps. [inskogo] r-na Krasnodar. [skogo] kraya " [around village Krivenkovskaya, Tuapse Region of Krasnodar District]; 25 May 1963; Savchenko leg.; / " opushka listv. [ennogo] lesa u reki, vdol gorn. [ogo] potoka " [the edge of the foliage forest near the river, along the mountain stream]; ZISP • 1 male; Khosta, Krasnodarskiy Kray, Caucasian Reserve, Tiso-samshitovaya rosha [Yew-and-Boxwood Tree Grove]; 43 ° 32 ' 014 " N, 39 ° 52 ' 621 " E; alt. 135 m; 10 May 2018; V. Lantsov leg.; / Fern-butcher community on the rocky slopes of the right side of the gorge of the Khosta River; IEMT • 1 male; Khosta, Krasnodarskiy Kray, Caucasian Reserve, Tiso-samshitovaya rosha [Yew-and-Boxwood Tree Grove]; 43 ° 31 ' 656 " N, 39 ° 52 ' 467 '' E; alt. 54 m; 12 May 2018, V. Lantsov leg.; / collected on light; IEMT • 2 males (in alcohol); North Caucasus, Krasnodarskiy Kray, near village Mezmay; 44 ° 11 ' 291 " N, 39 ° 58 ' 090 " E; alt. 808 m; 15 May 2018; V. Lantsov leg.; / Beech (Fagus orientalis), fruit tree (Pyrus sp. and Malus sp.) forest with Fraxinus excelsior in under growth, Sambucus nigra, Cornus mas, Rosa canina, Crataegus sp., in shrub layer and sedge Carex pendula, cereals and herbs in ground layer [a leveled area of the light part of the forest, possibly a site of a wild pear-apple orchard 50 - 70 years old]; IEMT • 1 male; Krasnodar Territory, Apsheron District, environs of village Mezmay, Guam Gorge, left slope, 44 ° 12 ' 409 " N, 39 ° 55 ' 056 " E, alt. 547 m; 24 May 2019; V. Lantsov leg.; / Landslide foot, community with butterbur [Petasites albus] as dominant along the banks of the stream, open places at the edge of the forest; ZISP • 8 males (in alcohol); Krasnodar Territory, Seversky District, in vicinity of village Thamaha, 3 May 2016; S. Kustov leg.; ZISP • 7 males, 1 female (in alcohol); Krasnodar Territory, Seversky District, environs of village Plancheskaya, 3 May 2016. S. Kustov leg.; ZISP • 5 males (in alcohol); Krasnodar Territory, environs of village Bolshoy Utrish; 1 May 2008; E. Hachikov. leg.; ZISP • 3 males (in alcohol); Krasnodar Territory, municipality Anapa, environs of village Sukko, Kvashin's Gorge, Dolgaya Niva territory; 44 ° 47 ' 20 " N, 37 ° 28 ' 33 " E; alt. 67 m; 6 - 8 May 2016; S. Kustov, V. Gladun. leg.; ZISP • 5 males (in alcohol); Krasnodar Territory, river Shakhe Gorge; 43 ° 52 ' 46 " N, 39 ° 50 ' 00 " E; 2 May 2012; V. Pilipenko leg.; VPMC • 3 males; (in alcohol); Khosta, Krasnodarskiy Kray, Caucasian Reserve, Tiso-samshitovaya rosha [Yew-and-Boxwood Tree Grove]; 43 ° 32 ' 014 " N, 39 ° 52 ' 621 " E; 8 May 2012; V. Pilipenko leg.; VPMC • 4 males (in alcohol); Krasnodar Territory, 13 km to the N from Sochi, Sukhoy Canyon; 43 ° 32 ' N, 39 ° 56 ' E; 5 May 2014; V. Pilipenko leg.; VPMC; RUSSIA - Dagestan • 5 males; Makhachkala, Tarki Distr.; 42 ° 56 ' 57 " N, 47 ° 29 ' 41 " E; alt. 220 m; 1 May 2019., V. Pilipenko leg.; cemetery on the hillside; VPMC • 1 male, 1 female; same locality; 10 May 2019., V. Pilipenko leg.; VPMC • 1 male, Tarki-Tau Mt.; 42 ° 56 ' 28 " N, 47 ° 28 ' 08 " E; alt. 450 m; 2 May 2019; V. Pilipenko leg.; oak forest; VPMC.	en	Lantsov, Vladimir I., Pilipenko, Valentin E. (2021): A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group. ZooKeys 1048: 145-175, DOI: http://dx.doi.org/10.3897/zookeys.1048.67564, URL: http://dx.doi.org/10.3897/zookeys.1048.67564
E104034BF2F251DB8C145337C9D914B4.taxon	diagnosis	Diagnosis. Tergite 9 with four widely spaced teeth and with three rounded notches distally; caudal margin of gonocoxite with two dentate projections. Paired appendages of sternite 8 with wide base bearing thick yellow setae distally covering gap between. Cercus long and straight; hypogynial valve only slightly longer than width at base.	en	Lantsov, Vladimir I., Pilipenko, Valentin E. (2021): A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group. ZooKeys 1048: 145-175, DOI: http://dx.doi.org/10.3897/zookeys.1048.67564, URL: http://dx.doi.org/10.3897/zookeys.1048.67564
E104034BF2F251DB8C145337C9D914B4.taxon	distribution	Distribution. Endemic to the Caucasus; currently known from the West Caucasus (northern and southern slopes; Krasnodar and Stavropol Territory) and from the East Caucasus (Dagestan; first record).	en	Lantsov, Vladimir I., Pilipenko, Valentin E. (2021): A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group. ZooKeys 1048: 145-175, DOI: http://dx.doi.org/10.3897/zookeys.1048.67564, URL: http://dx.doi.org/10.3897/zookeys.1048.67564
1D18358B839155CFB3A133D56252432A.taxon	description	Figs 10, 11, 12 G-I; 13 E, 14 E, K	en	Lantsov, Vladimir I., Pilipenko, Valentin E. (2021): A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group. ZooKeys 1048: 145-175, DOI: http://dx.doi.org/10.3897/zookeys.1048.67564, URL: http://dx.doi.org/10.3897/zookeys.1048.67564
1D18358B839155CFB3A133D56252432A.taxon	materials_examined	Material examined. Holotype: Azerbaijan • 1 male, " r-n Lerik, Azerbaidzh. SSR 26. VI. [1] 954 Dzhafarov " [Lerik region, Azerbaijan. SSR. [Azerbaijanskaya Soviet Socialist Republic] [alt. ~ 1115 m, 38 ° 46 ' 31 " N, 48 ° 24 ' 55 " E]; 26 Jun. 1954; Jafarov; / " Tipula talyshensis det. Savchenko sp. n. " [white label] / [Original red label without text] / " Holotypus " [red label]; ZISP. Holotype in good condition (Fig. 10 A, B), however, some legs missing: fore legs - left missing, right up to femur; mid legs - left present, right missing; hind legs - left and right up to femur. Paratypes. Azerbaijan • 2 males, 2 females; same data as for holotype [printed on white paper] / " Paratypus " [Paratype not initially marked; red label]; ZISP.	en	Lantsov, Vladimir I., Pilipenko, Valentin E. (2021): A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group. ZooKeys 1048: 145-175, DOI: http://dx.doi.org/10.3897/zookeys.1048.67564, URL: http://dx.doi.org/10.3897/zookeys.1048.67564
1D18358B839155CFB3A133D56252432A.taxon	diagnosis	Diagnosis. Male. Tergite 9 at apex with a denticle on either side of wide, shallow, flat median notch. Paired apical appendages of sternite 9 narrower distally, bearing tassel of long golden setae. Female. Hypogynial valve several times longer than width at base.	en	Lantsov, Vladimir I., Pilipenko, Valentin E. (2021): A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group. ZooKeys 1048: 145-175, DOI: http://dx.doi.org/10.3897/zookeys.1048.67564, URL: http://dx.doi.org/10.3897/zookeys.1048.67564
1D18358B839155CFB3A133D56252432A.taxon	distribution	Distribution. Endemic to the Caucasus - currently only known from Talysh (Azerbaijan).	en	Lantsov, Vladimir I., Pilipenko, Valentin E. (2021): A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group. ZooKeys 1048: 145-175, DOI: http://dx.doi.org/10.3897/zookeys.1048.67564, URL: http://dx.doi.org/10.3897/zookeys.1048.67564
