identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
EF9858D7FB6956D49F44AD5CF026BDF4.text	EF9858D7FB6956D49F44AD5CF026BDF4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amblyiulus georgicus Lohmander 1932	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Amblyiulus georgicus Lohmander, 1932</p>
            <p>Figs 1A, 2, 3, 15A, 16</p>
            <p> Amblyiulus georgicus Lohmander, 1932a: 180-182, figs 10-12 (D). </p>
            <p> Amblyiulus georgicus - Lohmander 1936: 170 (M); Kobakhidze 1965: 395 (M);  Lokšina and Golovatch 1979: 385 (M). </p>
            <p> Syrioiulus georgicus -  Mauriès 1984: 43 (M); Kokhia and Golovatch 2020: 207 (M). </p>
            <p>Material examined.</p>
            <p>  Armenia: 3 ♂♂, 5 ♀♀, 2 juv. (ZMUM), SW of Shnokh halfway between Alaverdi and  Bagratashen , ca. 1500 m a.s.l.,  Carpinus forest, litter, 24.V.1987  ;   2 ♀♀ (ZMUM),  Odzun W of Alaverdi, 1500-1550 m a.s.l.,  Quercus ,  Fagus ,  Carpinus , etc. forest, litter and under stones with ants, 23-24.V.1987; all leg. S. Golovatch, K. Eskov. </p>
            <p>Diagnosis.</p>
            <p> Differs from  A. hirtus sp. nov., apparently the most similar and geographically the closest congener known to date, by the following combination of somatic and gonopodal characteristics. Head without frontal setae; collum and metazonae of body rings without setae. Male mandibular stipites expanded. Promere wide, with two apical denticles. Solenomere with a membranous lobe notched apically. Rod of opisthomere relatively short. See also Key below. </p>
            <p>Redescription.</p>
            <p> Length of adults 27-30 mm (♂♂) or 28-31 mm (♀♀), width 1.6-1.7 mm (♂♂) or 1.7-1.9 mm (♀♀). Number of body rings in adults, 65-67+2+T (♂♂) or 67-69+2+T (♀♀). Body subcylindrical (typical of  Julidae ), metazonae brownish grey, prozonae yellowish grey (Fig. 1A). Head, a few postcollum rings and telson lighter than other body rings. Collum more vividly red-brown. Antennae, mouthparts, and legs yellow (Fig. 2A-C). Eyes absent. Metazonae with weak and irregular striations, 14-16 striae between dorsal axial line and ozopore (Fig. 2D). Ozopores relatively large, with a stria in front and lying behind suture without touching it (Fig. 2H). </p>
            <p>Antennae relatively long, in situ reaching segment 4. Head without frontal setae, but with 8+8-9+9 labral and 2+2 supralabral setae (Fig. 2A-C). Gnathochilarium with four setae on each lamella lingualis, stipites with a group of several short setae in medial part and three long setae at anterolateral margin (Fig. 2I). Collum and metazonae without setae at posterior margin (Figs 1A, 2A-G). Epiproct undeveloped (Fig. 2E, F). Hypoproct subtriangular, with several long setae (Fig. 2G). Telson and anal valves sparsely setose, setae being long (Fig. 2E-G).</p>
            <p>Male. Mandibular stipites expanded, slightly swollen in distal part (Fig. 2A). Leg pair 1 small, unciform, with a group of setae on each coxa and at base of telopodite; telopodite relatively long (Fig. 2J). Leg pair 2 with pads on postfemur and tibia (Fig. 2K). Penes short, bifurcate on top. Ventral edge of male segment 7 with elongated and rounded lamellae bordering the gonopodal aperture (Fig. 2L).</p>
            <p>Gonopods (Fig. 3). Promere spoon-shaped, relatively wide, constricted in basal third; mesal ridge in apical part forming a small mesal denticle (Fig. 3B, E). Lateral denticle large, well-developed. Mesomeral process simple, flattened, ribbon-shaped, notched on top (Fig. 3A, C, F). Opisthomere tripartite (Fig. 3D). Solenomere long, slightly curved, with caudomesal lamella notched apically; apical part with a fovea and a pointed membranous process. Anterior process of opisthomere appressed to solenomere, with a rounded apex. Anterolateral part of opisthomere with a helicoid rod.</p>
            <p>Female. First two leg pairs unmodified. Vulva rounded, operculum and bursa equal in height (Fig. 15A). Operculum at apical margin oblique, undivided. Bursa asymmetric, lateral valve higher than mesal one. Each valve with two rows of long setae. Median field of bursa very short, narrow; emargination of median field suboval.</p>
            <p>Remarks.</p>
            <p>This species was described from Borjom (= Borjomi), Georgia (Lohmander 1932a). The above samples represent the first formal records of this species from Armenia. It seems to populate high-montane deciduous forests in the western part of the Caucasus Minor (= Lesser Caucasus) (Fig. 16).</p>
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	https://treatment.plazi.org/id/EF9858D7FB6956D49F44AD5CF026BDF4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evsyukov, Aleksandr P.;Golovatch, Sergei I.;Antic, Dragan Z.	Evsyukov, Aleksandr P., Golovatch, Sergei I., Antic, Dragan Z. (2021): The millipede genera Amblyiulus Silvestri, 1896 and Syrioiulus Verhoeff, 1914 in the Caucasus, with notes on their distributions (Diplopoda, Julida, Julidae). ZooKeys 1048: 109-143, DOI: http://dx.doi.org/10.3897/zookeys.1048.68454, URL: http://dx.doi.org/10.3897/zookeys.1048.68454
072D01806F185A3294F01F7FD38A08C8.text	072D01806F185A3294F01F7FD38A08C8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amblyiulus hirtus Evsyukov & Golovatch & Antić 2021	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Amblyiulus hirtus sp. nov.</p>
            <p>Figs 1B, 4, 5, 15B, 16</p>
            <p>Material examined.</p>
            <p>  Holotype ♂ (ZMUM), Azerbaijan, NW above  Bash-Layski ca. 20 km NNW of Sheki, 1250 m a.s.l.,  Fagus ,  Carpinus ,  Acer , etc. forest, litter, 3.V.1987, leg. S. Golovatch, K. Eskov.  Paratypes: 5 ♂♂, 3 ♀♀ (ZMUM), same collection data as holotype . </p>
            <p>Non-type material.</p>
            <p> Azerbaijan: 2 ♂♂, 6 ♀♀ (ZMUM), SW of Kuba, 750 m a.s.l.,  Fagus ,  Quercus ,  Carpinus , etc. forest, litter and under bark, 23.IV.1987, leg. S. Golovatch, K. Eskov; Russia, Dagestan: 1 ♂, 2 ♀♀ (ZMUM), Kurush, 2550 m a.s.l., S slope, subalpine and alpine meadows, 20.VIII.1990, leg. G. Magomedov. </p>
            <p>Diagnosis.</p>
            <p> Assigned to the genus  Amblyiulus primarily because of the presence of a rod on the posterior gonopod opisthomere. Differs from  A. georgicus , perhaps the most similar congener known to date, by the following combination of somatic and gonopodal characters. Head with frontal setae; collum and metazonae of body rings each with a posterior whorl of setae. Promere narrow, with two side ridges. Solenomere apically with small filament-like processes. Rod of opisthomere relatively long. </p>
            <p>Name.</p>
            <p>To emphasise the presence of metazonal setae; adjective.</p>
            <p>Description.</p>
            <p> Holotype: length 27 mm, width 1.3 mm, number of body rings 51+2+T. Paratypes: length 25-30 mm, width 1.2-1.4 mm, number of body rings in adults, 45-67+1-3+T (♂♂); or length 27-28 mm, width 1.1-1.3 mm, number of body rings, 46-55+2-3+T (♀♀). Body subcylindrical (typical of  Julidae ), metazonae and prozonae yellowish grey (Fig. 1B). Head, a few postcollum rings and telson slightly lighter than other body rings. Collum slightly more vividly reddish. Antennae, mouthparts, and legs yellow (Fig. 4A-C). Eyes absent. Metazonae with weak, dense, and regular striations, 21-23 striae per quarter of metazonal surface, i.e., that between dorsal axial line and ozopore (Fig. 4A-G). Ozopores relatively large, with a stria in front, lying behind suture without touching it (Fig. 4H). </p>
            <p>Antennae relatively long, in situ reaching ring 4. Head with 1+1 frontal, 8+8-9+9 labral and 2+2 supralabral setae (Fig. 4A-C). Gnathochilarium with three thick setae on each lamella lingualis; stipites without setae in medial part, but with three long setae at anterolateral margin (Fig. 4I). Collum and each following metazona with a whorl of setae at posterior margin (Fig. 4A). Epiproct poorly developed, triangular, with several setae (Fig. 4E, F). Hypoproct subtriangular, covered with long setae (Fig. 4G). Telson and anal valves densely setose, setae being long.</p>
            <p> Male. Mandibular stipites unmodified (Fig. 4A). Leg pair 1 small, unciform, with a group of setae on coxa and at base of telopodite; telopodites curved anteriad, not anteromesad as in other species of  Julidae (Fig. 4J, K). Leg pair 2 with a large pad on tibia and a small one on postfemur (Fig. 4L). Penes short and bifurcate. Ventral edge of male pleurotergum 7 with small subtriangular lamellae bordering the gonopodal aperture (Fig. 4M). </p>
            <p> Gonopods (Fig. 5) with anterior and posterior pair equal in height. Promere spoon-shaped, relatively narrow, constricted in basal third; with two ridges: mesal ridge prominent all along; lateral ridge short, located only in apical part of promere (Fig. 5B, H). Mesomeral process simple, flattened, ribbon-shaped, with a small membranous lobe on top (Fig. 5A, C, G, I). Opisthomere tripartite (Fig. 5A, C, E, F). Solenomere long, slightly curved, with a caudomesal lamella notched apically; apical part with a fovea and short filament-like processes (Fig. 5A, C, G, I). Solenomere sometimes with an additional filiform process apically (see Remarks under  Syrioiulus taliscius ). Anterior process notched apically (Fig. 5A, C, E). Rod of solenomere relatively long, consisting of filament-like structures, lateral in position (Fig. 5A, D-G, I). </p>
            <p>Female. First two leg pairs unmodified. Vulva rounded, operculum higher than bursa (Fig. 15B) and bilobed apically. Bursa asymmetric, lateral valve higher than mesal one. Each valve with two rows of long setae. Median field of bursa very short, narrow; emargination of median field suboval.</p>
            <p>Remarks.</p>
            <p>This species seems to be endemic to the eastern part of the Caucasus Major within both northeastern Azerbaijan and the Republic of Dagestan, Russia (Fig. 16).</p>
            <p> It is the presence of a laterally positioned rod that brings both  A. georgicus and  A. hirtus sp. nov. particularly close together. However, the rod in these two species is located laterally, whereas that in  A. barroisi anteromesally (Enghoff 1992: fig. 11; Golovatch 2018: fig. 10C). These differences seem to be quite important, but because those three species share not only the presence of a rod, but also a small, but discernible fovea on top of the solenomere, for the time being it seems best to regard the trio as members of  Amblyiulus . </p>
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	https://treatment.plazi.org/id/072D01806F185A3294F01F7FD38A08C8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evsyukov, Aleksandr P.;Golovatch, Sergei I.;Antic, Dragan Z.	Evsyukov, Aleksandr P., Golovatch, Sergei I., Antic, Dragan Z. (2021): The millipede genera Amblyiulus Silvestri, 1896 and Syrioiulus Verhoeff, 1914 in the Caucasus, with notes on their distributions (Diplopoda, Julida, Julidae). ZooKeys 1048: 109-143, DOI: http://dx.doi.org/10.3897/zookeys.1048.68454, URL: http://dx.doi.org/10.3897/zookeys.1048.68454
9B65CB30B6B85DFBBE9206280BF3A26A.text	9B65CB30B6B85DFBBE9206280BF3A26A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amblyiulus Silvestri 1896	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Genus  Amblyiulus Silvestri, 1896</p>
            <p>Type species.</p>
            <p> Julus barroisi Porat, 1893, by original designation. </p>
            <p>Diagnostic remarks.</p>
            <p> Here we follow  Tabacaru’s (1978) opinion that the genera of the subfamily/tribe  Pachyiulinae /-ini are best to be diagnosed using a complex of characters, both gonopodal and somatic. In the latest review of this tribe (  Mauriès 1982), all genera are divided into three groups depending on the structure of the apical part of the opisthomere, viz., the presence/absence of a fovea and the presence/absence of a pseudoflagellum. He mistakenly assigned the genus  Amblyiulus to group 3 (along with many other genera like  Dolichoiulus ), which have neither a fovea nor a pseudoflagellum. However, in accordance with our and earlier descriptions (e.g., Golovatch 2018),  Amblyiulus has a fovea, however small, on the top of the solenomere. By the presence of a fovea and the absence of a pseudoflagellum, the genera  Parapachyiulus Golovatch, 1979 and  Dangaraiulus Golovatch, 1979 also join this group (Golovatch 1979, 2018). According to a number of other characters, such as the presence of frontal setae, apicoventral lobes on the male mandibles, and the mesomeral process being as high as the opisthomere,  Amblyiulus belongs to  Mauriès’ subgroup 3aa, together with the genera  Syrioiulus and  Promeritoconus . However, it seems noteworthy that sometimes frontal setae can be absent, while male mandibular stipites can remain unmodified. </p>
            <p> The promere in  Amblyiulus is narrowed in the basal third, in contrast to that in  Promeritoconus , which is narrowed apically; in the apical part it may have one or two denticles, but sometimes none. The head can be with or without frontal setae. The eyes are mostly absent. The opisthomere of the posterior gonopod is tripartite: a solenomere, an anterior process, and an anteromesal or lateral rod, vs. bipartite in  Syrioiulus . </p>
            <p>Species included.</p>
            <p> Amblyiulus barroisi (Porat, 1893),  Amblyiulus cedrophilus (Attems, 1910),  Amblyiulus festae (Silvestri, 1895),  Amblyiulus georgicus Lohmander, 1932,  Amblyiulus hirtus sp. nov., and possibly several others, but their identity requires verification (Golovatch 2018). </p>
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	https://treatment.plazi.org/id/9B65CB30B6B85DFBBE9206280BF3A26A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evsyukov, Aleksandr P.;Golovatch, Sergei I.;Antic, Dragan Z.	Evsyukov, Aleksandr P., Golovatch, Sergei I., Antic, Dragan Z. (2021): The millipede genera Amblyiulus Silvestri, 1896 and Syrioiulus Verhoeff, 1914 in the Caucasus, with notes on their distributions (Diplopoda, Julida, Julidae). ZooKeys 1048: 109-143, DOI: http://dx.doi.org/10.3897/zookeys.1048.68454, URL: http://dx.doi.org/10.3897/zookeys.1048.68454
F6BBFDA7159E5AE8A68E94EFFED49F93.text	F6BBFDA7159E5AE8A68E94EFFED49F93.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pachyiulini gen subsp. sp var. 1 gen. sp. 1	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Pachyiulini gen. sp. 1</p>
            <p>Fig. 16</p>
            <p>Material examined.</p>
            <p>  1 ♀ (ZMUM) Russia, Stavropol Area, Kochubeevsky Distr.,  Nevinnomyssk , mouth of  Bolshoy Zelenchuk River , floodplain forest, 17.IV.2016, leg. R.B. Zuev, A.V. Aulova. </p>
            <p>Brief description.</p>
            <p>Body grey, head and collum dark yellow, antennae and legs yellow. The following somatic characteristics seem to be the most important: absence of eyes, presence of frontal setae, presence of caudal whorls of setae on metazonae, absence of an epiproct, and setose anal valves.</p>
            <p>Remark.</p>
            <p>Unfortunately, the only female does not allow a closer generic allocation.</p>
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	https://treatment.plazi.org/id/F6BBFDA7159E5AE8A68E94EFFED49F93	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evsyukov, Aleksandr P.;Golovatch, Sergei I.;Antic, Dragan Z.	Evsyukov, Aleksandr P., Golovatch, Sergei I., Antic, Dragan Z. (2021): The millipede genera Amblyiulus Silvestri, 1896 and Syrioiulus Verhoeff, 1914 in the Caucasus, with notes on their distributions (Diplopoda, Julida, Julidae). ZooKeys 1048: 109-143, DOI: http://dx.doi.org/10.3897/zookeys.1048.68454, URL: http://dx.doi.org/10.3897/zookeys.1048.68454
D4B62F70F7AA55F2B52227A43D18661D.text	D4B62F70F7AA55F2B52227A43D18661D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pachyiulini gen subsp. sp var. 2 gen. sp. 2	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Pachyiulini gen. sp. 2</p>
            <p>Fig. 16</p>
            <p>Material examined.</p>
            <p>  Georgia: 1 ♀ (ZMUM), Ajaria, Kintrish Nature Reserve,  Zeraboseli , 450-600 m a.s.l., 1-3.VI.1981, leg. S. Golovatch, J. Martens  ;   1 ♀ (ZMUM), same locality, 800 m a.s.l.,  Rhododendron thicket, litter, 13.X.1981, leg. S. Golovatch  ;   1 ♀ (ZMUM), same locality, valley of  Khekpara River , 12.X.1984, leg. E. Kvavadze. </p>
            <p>Brief description.</p>
            <p>Body greyish yellow. Head, collum, a few postcollum rings and telson slightly lighter than other body rings. Ommatidia and frontal setae absent. Collum and each metazona of following rings with a whorl of long setae at posterior margin. Epiproct undeveloped. Anal valves densely setose.</p>
            <p>Remarks.</p>
            <p> These specimens differ from  Syrioiulus adsharicus in the absence of frontal setae and ommatidia. The absence of males makes it impossible to definitively identify the above samples. At least the taxonomic significance of frontal setae must not be overestimated, as they may be present or absent even within the same species, e.g.,  S. aharonii (see Golovatch 2018). </p>
            <p> Since the two unidentified species may well prove to represent  Amblyiulus or  Syrioiulus , we map their records in Figure 16, but omit them from the key below. </p>
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	https://treatment.plazi.org/id/D4B62F70F7AA55F2B52227A43D18661D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evsyukov, Aleksandr P.;Golovatch, Sergei I.;Antic, Dragan Z.	Evsyukov, Aleksandr P., Golovatch, Sergei I., Antic, Dragan Z. (2021): The millipede genera Amblyiulus Silvestri, 1896 and Syrioiulus Verhoeff, 1914 in the Caucasus, with notes on their distributions (Diplopoda, Julida, Julidae). ZooKeys 1048: 109-143, DOI: http://dx.doi.org/10.3897/zookeys.1048.68454, URL: http://dx.doi.org/10.3897/zookeys.1048.68454
FD1AF8FACF675EFE91C4A30BE3A2CF21.text	FD1AF8FACF675EFE91C4A30BE3A2CF21.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Syrioiulus adsharicus (Lohmander 1936)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Syrioiulus adsharicus (Lohmander, 1936)</p>
            <p>Figs 1C, 6, 7, 15C, 16</p>
            <p> Amblyiulus (Heteropachyiulus) adsharicus Lohmander, 1936: 156-159, figs 131, 132 (D). </p>
            <p> Amblyiulus adsharicus - Kobakhidze 1964: 191 (M);  Lokšina and Golovatch 1979: 385 (M); Talikadze 1984: 143 (M); Kokhia and Golovatch 2018: 40 (M). </p>
            <p> Syrioiulus adsharicus - Vagalinski 2020: 89 (M); Kokhia and Golovatch 2020: 207 (M). </p>
            <p>Material examined.</p>
            <p>  Georgia: 10 ♂♂, 16 ♀♀, 5 juv. (ZMUM), 15 km W of  Adigeni ,  Abies ,  Picea ,  Fagus ,  Acer , etc. forest, 1500-1700 m a.s.l., litter, logs, under stones, 14-15.V.1983, leg. S. Golovatch  ;   4 ♂♂, 5 ♀♀ (ZMUM), near  Adigeni , 9.VI.1977, leg. V. Dolin. </p>
            <p>Diagnosis.</p>
            <p>Differs from all congeners by the following combination of somatic and gonopodal characters. Head with frontal setae. Collum and each metazona of following body rings with a whorl of long setae at caudal margin. Ommatidia present, but only a few ommatidia, all unpigmented and very small. Solenomere with small denticles apically. Anterior process of opisthomere with small filament-like spines apically.</p>
            <p>Redescription.</p>
            <p> Length of adults 17-30 mm (♂♂) or 18-31 mm (♀♀), width 1.2-1.4 mm (♂♂) or 1.3-1.7 mm (♀♀). Number of body rings in adults, 50-63+1-2+T (♂♂) or 52-60+1-2+T (♀♀). Body subcylindrical (typical of  Julidae ), metazonae brownish grey, prozonae violet grey (Figs 1C, 6A, C). Head, collum and body rings from yellow to greyish yellow. Antennae, mouthparts and first leg pairs yellow, other pairs brown (Fig. 6A, C-H). Eyes present, unpigmented, very small, composed of 3-7 ommatidia, unequal numbers on opposite sides of head (Fig. 6B). Metazonae with weak striations, 17-19 striae per quarter of metazonital surface, i.e., that between dorsal axial line and ozopore (Fig. 6E). Ozopores small, lying behind suture and touching it (Fig. 6I). </p>
            <p>Antennae relatively long, in situ reaching segment 4. Head with 1+1 frontal, 11+11-13+13 labral and 2+2 supralabral setae (Fig. 6C, D). Gnathochilarium with four long setae on each lamella lingualis, stipites with a medial group of 7-10 short setae, three long and two short setae at anterolateral margin (Fig. 6J). Collum and each metazona of following rings with a whorl of long setae at posterior margin (Fig. 6A, E, F-H). Epiproct undeveloped (Fig. 6F, G). Hypoproct subtriangular, with long setae (Fig. 6H). Anal valves densely setose, setae being long.</p>
            <p> Male. Mandibular stipites expanded, slightly swollen (Fig. 6A). Leg pair 1 small, unciform, telopodites curved anteromesad (as in most other  Julidae ), with a group of setae on each coxa and at base of telopodite (Fig. 6K). Leg pair 2 with pads on postfemur and tibia (Fig. 6L). Penes short, bifurcate. Ventral edge of male pleurotergum 7 with relatively wide and apically rounded lamellae bordering the gonopodal aperture (Fig. 6M). </p>
            <p>Gonopods (Fig. 7) with anterior and posterior pair equal in height. Promere spoon-shaped, constricted in basal third; a mesal ridge along basal 2/3 extent; two apical denticles well-developed, mesal one vertical and with a weakly bifurcate apex, lateral one short, wide and rounded apically (Fig. 7B, E). Mesomeral process simple, flattened, ribbon-shaped, bifurcate (Fig. 7C, F). Opisthomere bipartite (Fig. 7D, C). Solenomere long, erect, with small denticles apically, bearing a fovea at apex; caudomesal lamella wide with a notched apical margin (Fig. 7A, C, F). Anterior process apically with small filament-shaped spines (Fig. 7D).</p>
            <p>Female. First two leg pairs unmodified. Vulva elongated, covered with long setae (Fig. 15C). Operculum relatively low, deeply divided. Bursa asymmetric, lateral valve higher than mesal one. Median field of bursa narrow; emargination of median field narrow and elongated.</p>
            <p>Remark.</p>
            <p> This species was originally described from Batumi,  “Bortschacha” (Lohmander 1936). Our new record from near Adigeni is evidence of the species likely to be endemic to the southern part of the Colchidan biogeographic province, all within Georgia (Fig. 16). </p>
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	https://treatment.plazi.org/id/FD1AF8FACF675EFE91C4A30BE3A2CF21	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evsyukov, Aleksandr P.;Golovatch, Sergei I.;Antic, Dragan Z.	Evsyukov, Aleksandr P., Golovatch, Sergei I., Antic, Dragan Z. (2021): The millipede genera Amblyiulus Silvestri, 1896 and Syrioiulus Verhoeff, 1914 in the Caucasus, with notes on their distributions (Diplopoda, Julida, Julidae). ZooKeys 1048: 109-143, DOI: http://dx.doi.org/10.3897/zookeys.1048.68454, URL: http://dx.doi.org/10.3897/zookeys.1048.68454
74F6B9A623CB5EFBB4DE8FC15CFD2604.text	74F6B9A623CB5EFBB4DE8FC15CFD2604.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Syrioiulus armeniacus Evsyukov & Golovatch & Antić 2021	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Syrioiulus armeniacus sp. nov.</p>
            <p>Figs 1F, 13, 14, 15F, 16</p>
            <p>Material examined.</p>
            <p>  Holotype ♂ (ZMUM), Armenia,  Kafan Distr. , Shikahoh Nature Reserve, Shikahoh, 900-950 m a.s.l.,  Quercus ,  Fagus ,  Carpinus forest by spring, litter, logs and under stones, 28.IV.1983, leg. S. Golovatch.  Paratypes: 14 ♂♂, 24 ♀♀ (ZMUM), same collection data as holotype . </p>
            <p>Non-type material.</p>
            <p> Armenia: 4 ♂♂, 3 ♀♀ (ZMUM), Shikahoh Nature Reserve, Nerkin And, old  Platanus stand along river, litter, in rotten wood, under stones, 30.IV.1983; 1 ♂, 1 ♀ (ZMUM), near Kajaran, Megri Mt. Ridge, N of Tashtun Pass, 2000 m a.s.l.,  Quercus forest on steep slope, litter, logs, 27.IV.1983; 5 ♂♂, 3 ♀♀, 2 juv. (ZMUM), Megri Distr., SSE of Lichk, Megri River valley,  Quercus forest, litter, under stones and in rotten wood, 25.IV.1983; 2 ♂♂, 16 ♀♀, 2 juv. (ZMUM), above Kuris, 1500 m a.s.l.,  Quercus and  Carpinus forest, litter, under bark and stones along spring, 26.IV.1983; 12 ♂♂, 11 ♀♀, 1 juv. (ZMUM), ca. 4 km NNW of Megri, Legvaz,  Juglans and  Quercus shrub with  Paliurus and Rosa, litter and under stones, 1000 m a.s.l., 24-25.IV.1983; 5 ♂♂, 2 ♀♀ (ZMUM), 6 km N of Shvanidzor, sparse  Quercus forest, 1200-1300 m a.s.l., litter, under stones and bark, 24.IV.1983; 5 ♂♂, 6 ♀♀, 2 juv. (ZMUM), environs of Megri, xeriphytous bare canyon, under stones, sparse  Juniperus and  Paliurus , ca. 1000 m a.s.l., 24.IV.1983, all leg. S. Golovatch; 2 ♀♀ (ZMUM), Odzun W of Alaverdi, 1500-1550 m a.s.l.,  Quercus ,  Fagus ,  Carpinus etc. forest, litter and under stones with ants, 23-24.V.1987, leg. S. Golovatch and K. Eskov; 1 ♀ (ZMUM), Nurkus, 7.VII.1985, leg. V.A. Zakharyan. </p>
            <p>Diagnosis.</p>
            <p> This new species belongs to the genus  Syrioiulus because of the presence of only two apices on the opisthomere. Differs from all regional congeners by the following combination of somatic and gonopodal characters. Head with frontal setae. Collum and metazonae of following body rings without setae. Eyes absent. Solenomere with a pointed process apically. Anterior process rounded on top. </p>
            <p>Name.</p>
            <p>The new species is named after its terra typica; adjective.</p>
            <p>Description.</p>
            <p>Holotype: length 25 mm, width 1.2 mm, number of body rings 50+2+T. Paratypes and non-type material: length 17-33 mm, width 1.2-1.6 mm, number of body rings, 50-68+1-2+T (♂♂); or length 20-29 mm, width 1.2-1.6 mm, number of body rings, 46-55+2-3+T (♀♀). Body subcylindrical, metazonae brownish yellow, prozonae brownish grey (Figs 1F, 13A-G). Head, collum and telson slightly lighter than body rings (Fig. 1F). Antennae, mouthparts, and legs yellow (Fig. 13A, C-E, G). Eyes absent. Metazonae with weakly developed, dense, and regular striations, 20-23 striae per quarter of metazonital surface, i.e., that between dorsal axial line and ozopore (Fig. 13D). Ozopores relatively large, situated between striae in touch with one of them, lying behind suture without touching it (Fig. 13H).</p>
            <p>Antennae relatively long, in situ reaching segment 3. Head with 1+1 frontal, 9+9-10+10 labral and 2+2 supralabral setae (Fig. 13A-C). Gnathochilarium with three thick setae on each lamella lingualis; stipites with a group of 6-9 setae in medial part and three long setae in anterolateral part (Fig. 13I). Collum and metazonae without setae (Fig. 13A, B). Epiproct poorly developed, triangular, with several long setae (Fig. 13E, F). Hypoproct subtriangular, covered with long setae (Fig. 13G). Telson and anal valves densely setose.</p>
            <p> Male. Mandibular stipites modified, slightly swollen in distal part (Fig. 13A). Leg pair 1 small, unciform, telopodites directed anteromesad (typical of  Julidae ), with a group of setae on coxa and telopodite (Fig. 13J). Leg pair 2 with small pads on postfemur and tibia (Fig. 13K). Penes short, bifurcate. Ventral edge of male pleurotergum 7 with wide curved lamellae bordering the gonopodal aperture (Fig. 13L). </p>
            <p>Gonopods (Fig. 14) with anterior and posterior parts both equal in height. Promere spoon-shaped, bowl being relatively wide, constricted in basal third; mesal ridge well-developed all along, with a small mesal denticle in apical part; lateral denticle large, rounded on top (Fig. 14B, F). Mesomeral process simple, flattened, ribbon-shaped, widened apically, with a group of small teeth (Fig. 14A, C, E, G). Opisthomere bipartite. Solenomere long, slightly curved; its apical part with a fovea and a pointed process (Fig. 14A, C, D, G). Anterior process as high as solenomere, rounded at tip (Fig. 14D).</p>
            <p>Female. First two leg pairs unmodified. Vulva rounded, operculum higher than bursa (Fig. 15F). Operculum slightly divided at apical margin. Bursa asymmetric, lateral valve higher than mesal one. Each valve with two rows of long setae. Median field of bursa narrow; emargination of median field elongated and suboval.</p>
            <p>Remark.</p>
            <p>This species seems to be endemic to the Caucasus Minor within Armenia, but most likely it also occurs in the adjacent parts of eastern Azerbaijan and northwestern Iran (Fig. 16).</p>
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	https://treatment.plazi.org/id/74F6B9A623CB5EFBB4DE8FC15CFD2604	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evsyukov, Aleksandr P.;Golovatch, Sergei I.;Antic, Dragan Z.	Evsyukov, Aleksandr P., Golovatch, Sergei I., Antic, Dragan Z. (2021): The millipede genera Amblyiulus Silvestri, 1896 and Syrioiulus Verhoeff, 1914 in the Caucasus, with notes on their distributions (Diplopoda, Julida, Julidae). ZooKeys 1048: 109-143, DOI: http://dx.doi.org/10.3897/zookeys.1048.68454, URL: http://dx.doi.org/10.3897/zookeys.1048.68454
80560408A12F52E390522CA9F91DFB0D.text	80560408A12F52E390522CA9F91DFB0D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Syrioiulus continentalis (Attems 1903)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Syrioiulus continentalis (Attems, 1903)</p>
            <p>Figs 1D, 8A, B, 9, 10, 15D, 16</p>
            <p> Pachyiulus (Dolichoiulus) continentalis Attems, 1903: 147, 148, figs 82-84 (D). </p>
            <p> Amblyiulus continentalis - Lohmander 1932b: 40, 41, figs 33-35 (D); 1936: 156 (R); Rakhmanov 1971: 1412 (R); 1972: 116 (R); Samedov et al. 1972: 1245;  Lokšina and Golovatch 1979: 385 (M); Bababekova 1996: 90 (M). </p>
            <p> Syrioiulus continentalis -  Mauriès 1982: 441 (M); 1984: 43 (M); Vagalinski 2020: 89 (M). </p>
            <p>Material examined.</p>
            <p>
                  Azerbaijan: 3 ♂♂, 1 ♀ (ZMUM), Talysh Mts, Zuvand,  Joni , 1500 m a.s.l., 28-29.V.1976, leg. V.G. Dolin  ;   1 ♂ (ZMUM), Lenkoran, Hyrcan forest, Khan Bulan River near  Alexeevka , 22.IV.1985, leg. E.B. Kupriyanova  ;   7 ♂♂, 5 ♀♀, 2 juv. (ZMUM), Lenkoran,  Hyrcan Nature Reserve , litter, 26.I.-4.II.1985, leg. A. Druk  ;  2 ♂♂, 4 ♀♀ (ZMUM), same locality, 21.IX.1987, leg. S. Zonstein ; 2 ♂♂, 2 juv. (IZB);   1 ♂, 1 ♀, 1 juv. (SMNG),  Lǝnkǝran rayon, Hyrcan Nature Reserve,  Daştatük 1.3 km Xanbulan Reservoir,  Parrotia forest, diverse bushes, under leaves, 110 m a.s.l., 38.6747°N, 48.7622°E  ; 1 ♂ (IZB);   1 ♂ (SMNG), same locality,  
                <a title="Search Plazi for locations around (long 48.7362/lat 38.6726)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=48.7362&amp;materialsCitation.latitude=38.6726">
SW of 
Asagi
Apu
</a>
                 ,  Quercus forest, within leaves and rotten wood, 180 m a.s.l., 38.6726°N, 48.7362°E, all leg. F. Walther, H. Reip, D.  Antić ; 3 ♂♂, 2 ♀♀, 1 juv. (IZB);   5 ♂♂, 3 ♀♀ (SMNG), Lerik rayon,  
                <a title="Search Plazi for locations around (long 48.5819/lat 38.7638)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=48.5819&amp;materialsCitation.latitude=38.7638">Hyrcan Nature Reserve</a>
                 , road  Lǝnkǝran-Lerik at km 32, small side valley, forest of  Parrotia with some  Quercus , thick leaf layer, 400 m a.s.l., 38.7638°N, 48.5819°E  ;   1 juv. (IZB), Astara rayon,  
                <a title="Search Plazi for locations around (long 48.7623/lat 38.4493)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=48.7623&amp;materialsCitation.latitude=38.4493">Hyrcan Nature Reserve</a>
                 , SW of  Zünqülǝş , beginning of a small valley,  Parrotia and  Alnus bushes, in leaves, 60 m a.s.l., 38.4493°N, 48.7623°E  ; 2 ♂♂, 1 juv. (IZB);   2 ♂♂, 1 ♀ (SMNG), same locality, end of small valley, steep slope,  Parrotia ,  Quercus ,  Acer trees, under leaves and rotten tree trunks, 130 m a.s.l., 38.4480°N, 48.7597°E, all leg. F. Walther, H. Reip, D.  Antić ;   2 ♂♂ (ZMUM),  Azfilial , 100 m a.s.l., 31.V.-1.VI.1996  ;   1 ♂ (ZMUM), Apo below  Bilasar , 350 m a.s.l., 8-9.VI.1996  ;   2 ♂♂, 1 ♀, 1 juv. (ZMUM), Astara Distr.,  Istisu ca. 8 km WSW of Astara,  Quercus ,  Acer ,  Carpinus , etc. forest, 10-30 m a.s.l., litter, under stones and bark, 10.X.1983  ;   2 ♀♀ (ZMUM),  Istisu ca. 8 km SW of Masally,  Quercus ,  Acer ,  Carpinus etc. forest, 80-140 m a.s.l., under bark and stones, 19-20.X.1983  ;   2 ♀♀ (ZMUM),  Istisu W of Astara, 100 m a.s.l., 2-6.VI.1996, all leg. S. Golovatch. 
            </p>
            <p>Diagnosis.</p>
            <p> Differs from all congeners by the following combination of somatic and gonopodal characters. Head with frontal setae. Collum and each metazona of following body rings with a whorl of long setae at caudal margin. Eyes present. Solenomere with a group of small spines on top. Anterior process of opisthomere subtriangular apically. This species is clearly distinguished in the field from all other millipedes by its characteristic greyish yellow colouration with a yellow stripe dorsally, and its particularly strong odour clearly resembling that of  Pachyiulus krivolutskyi from the western Caucasus (= Colchis). </p>
            <p>Redescription.</p>
            <p>Length of adults 28-45 mm (♂♂) or 26-46 mm (♀♀), width 2.0-2.3 mm (♂♂) or 2.2-2.7 mm (♀♀). Number of body rings in adults, 46-66+1-2+T (♂♂) or 49-66+1-2+T (♀♀). Body subcylindrical, metazonae from greyish yellow to yellow, prozonae light yellow (Figs 1D, 8A, B); live specimens dorsally with a darker, vivid yellow stripe (Fig. 8B). Head, collum and telson slightly lighter than other body rings (Fig. 8A, B). Antennae grey, mouthparts and legs light yellow (Fig. 9A-G). Eyes present, black, oval, each composed of 19-23 ommatidia (Fig. 9A, C). Striations on metazonae deep, not reaching the caudal margin, 28-32 striae per quarter of metazonal surface, i.e., between dorsal axial line and ozopore (Fig. 9D). Ozopores large, lying behind suture without touching it (Fig. 9H).</p>
            <p>Antennae relatively long, in situ reaching segment 3. Head with 1+1 frontal, 8+8-9+9 labral and 2+2-4+4 supralabral setae (Fig. 9A-C). Gnathochilarium with three long setae on each lamella lingualis, groups of several small setae in median part of stipites and six or seven long setae at anterolateral margin (Fig. 9I). Collum and each following metazona with a whorl of long and thick setae at posterior margin (Fig. 9A-F). Epiproct undeveloped (Fig. 9E, F). Hypoproct subtriangular, with long setae (Fig. 9G). Telson and anal valves densely setose, setae being long.</p>
            <p>Male. Mandibular stipites expanded, with swollen lobes (Fig. 9A). Leg pair 1 small, unciform, telopodites directed anteromesad, with a group of long setae on each coxa; telopodite setose in basal part (Fig. 9J). Leg pair 2 with pads on postfemur and tibia (Fig. 9K). Penes short, bifurcate. Ventral edge of male pleurotergum 7 with narrow elongated lamellae bordering the gonopodal aperture (Fig. 9L).</p>
            <p>Gonopods (Fig. 10) with anterior (promere) part higher than posterior (opisthomere) one. Promere spoon-shaped, constricted in basal third; mesal ridge wide along 2/3 extent; with denticles in apical part: mesal denticle small and broadly rounded, lateral one well-expressed and long (Fig. 10B, E). Mesomeral process simple, slightly curved, flattened apically (Fig. 10A, C, F). Opisthomere bipartite (Fig. 10D). Solenomere elongated, with an apical membranous lobe, subtriangular at apex, with a fovea and a group of small spines on top; caudomesal lamella wide (Fig. 10A, C, D, F). Anterior process of opisthomere subtriangular apically (Fig. 10D).</p>
            <p>Female. First two leg pairs unmodified. Operculum of vulva without setae on caudal surface, apical margin relatively flat (Fig. 15D). Bursa subsymmetrical, lateral valve slightly larger than mesal one. Each valve with two rows of long setae. Median field of bursa narrow; emargination of median field suboval.</p>
            <p>Remarks.</p>
            <p> Probably one of the most common and apparently the largest species of the genus. The unusually strong odour and the chemical composition of the repugnatorial secretion are similar to those of  Pachyiulus krivolutskyi (Makarov et al., pers. obs.). This species inhabits various deciduous forests in Azerbaijan, also occurring in northern Iran (Lohmander 1932b), endemic to the Hyrcanian biogeographic province (Fig. 16). </p>
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	https://treatment.plazi.org/id/80560408A12F52E390522CA9F91DFB0D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evsyukov, Aleksandr P.;Golovatch, Sergei I.;Antic, Dragan Z.	Evsyukov, Aleksandr P., Golovatch, Sergei I., Antic, Dragan Z. (2021): The millipede genera Amblyiulus Silvestri, 1896 and Syrioiulus Verhoeff, 1914 in the Caucasus, with notes on their distributions (Diplopoda, Julida, Julidae). ZooKeys 1048: 109-143, DOI: http://dx.doi.org/10.3897/zookeys.1048.68454, URL: http://dx.doi.org/10.3897/zookeys.1048.68454
366D2F9E79FA5C68BFAE74A6DE9E6FBC.text	366D2F9E79FA5C68BFAE74A6DE9E6FBC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Syrioiulus taliscius (Attems 1927)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Syrioiulus taliscius (Attems, 1927)</p>
            <p>Figs 1E, 8C, D, 11, 12, 15E, 16</p>
            <p> Amblyiulus taliscius Attems, 1927: 243, 244, figs 336-338 (D). </p>
            <p> Amblyiulus taliscius - Lohmander 1932b: 182 (M); 1936: 170 (M); Rakhmanov 1972: 116 (R);  Lokšina and Golovatch 1979: 385 (M); Bababekova 1996: 90 (M). </p>
            <p> Syrioiulus taliscius -  Mauriès 1982: 441 (M); 1984: 43 (M); Vagalinski 2020: 92 (M). </p>
            <p>Material examined.</p>
            <p>
                  Azerbaijan: 4 ♂♂, 14 ♀♀, 1 juv. (ZMUM), Talysh,  Joni , 1500 m a.s.l., 28-29.V.1976, leg. V.G. Dolin  ;   2 ♂♂ (ZMUM), Lenkoran, Hyrcan Nature Reserve,  Telman , 28.IV.1984  ;   1 ♂, 1 ♀ (ZMUM), same locality,  Gaftoni , 9.V.1984, all leg. H. Aliev  ;   1 ♀ (ZMUM), Hyrcan forest, Khan Bulan River near  Alexeevka , 22.IV.1985, leg. E.B. Kupriyanova  ;   3 ♂♂, 12 ♀♀ (ZMUM), same locality, Avrora,  Moscow-Forest , 50 m a.s.l., 1.VI.1996  ;   3 ♂♂, 3 ♀♀, 1 juv. (ZMUM), same locality, Apo below  Bilasar , 350 m a.s.l., 8-9.VI.1996, all leg. S. Golovatch  ;   1 ♂ (SMNG),  Lǝnkǝran rayon,  Siyablı ,  Parrotia ,  Zelkova ,  Quercus coppice, steep slope, 110 m a.s.l., 38.7170°N, 48.7253°E  ;   1 ♂ (IZB), Lerik rayon,  
                <a title="Search Plazi for locations around (long 48.5819/lat 38.7638)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=48.5819&amp;materialsCitation.latitude=38.7638">Hyrcan Nature Reserve</a>
                 , road  Lǝnkǝran-Lerik at km 32, small side valley, forest of  Parrotia with some  Quercus , thick leaf litter, 400 m a.s.l., 38.7638°N, 48.5819°E  ; 7 ♀♀ (IZB),   7 ♀♀ (SMNG), Astara rayon,  
                <a title="Search Plazi for locations around (long 48.7597/lat 38.448)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=48.7597&amp;materialsCitation.latitude=38.448">Hyrcan Nature Reserve</a>
                 , SW of  Zünqüləş , end of small valley, steep slope,  Parrotia ,  Quercus ,  Acer trees, under leaves and rotten tree trunks, 130 m a.s.l., 38.4480°N, 48.7597°E  ; 2 ♂♂, 1 ♀, 1 juv. (IZB);   2 ♂♂, 2 ♀♀, 1 juv. (SMNG),  Lǝnkǝran rayon, Hyrcan Nature Reserve,  Daştatük 1.3 km Xanbulan Reservoir,  Parrotia forest, divers bushes, under leaves, 110 m a.s.l., 38.6747°N, 48.7622°E  ; 3 ♂♂, 2 ♀♀, 1 juv. (IZB); 5 ♂♂ (1 SEM), 3 ♀♀ (1 SEM),   1 juv. (SMNG)  Lǝnkǝran rayon,  
                <a title="Search Plazi for locations around (long 48.7362/lat 38.6726)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=48.7362&amp;materialsCitation.latitude=38.6726">Hyrcan Nature Reserve</a>
                 , SW of  Aşağı Apu,  Quercus forest, within leaves and rotten wood, 180 m a.s.l., 38.6726°N, 48.7362°E, all leg. F. Walther, H. Reip, D.  Antić ;   1 ♂ without gonopods (ZMUM), Shemakha Distr., farm  Guseinzade , foothills,  Vitis , summer 1982, leg. A. Ismailov  ;   1 ♀ (ZMUM), Zakataly Distr.,  Geyam , cornfield, 17.IV.1986, leg  .?;   7 ♂♂, 5 ♀♀ (ZMUM), Baku,  City parks, 18-21.V.1981  ;   1 ♂ (ZMUM), Belokani near  Zakatali , 600 m a.s.l., village garden, 24.V.1981  ;   1 ♀ (ZMUM), above  Akhsu 120 km W Baku, 900 m a.s.l.,  Quercus shrub, 22.V.1981, all leg. S. Golovatch and J. Martens  ;   1 ♀ (ZMUM), ca. 14 km W of Ismailly,  Galyhjakh , under bark, 1.V.1987  ;   1 ♀ (ZMUM),  Altyagach , 1050-1100 m a.s.l.,  Quercus ,  Fagus ,  Carpinus , etc. forest, litter, 20 and 26.IV.1987, all leg. S.Golovatch and K. Eskov  ; 3 ♂♂, 5 ♀♀, 1 juv. (IZB);   3 ♂♂, 5 ♀♀ (SMNG),  İsmayıllı rayon,  Topçu 7.8 km towards  Vəndam , flat area with old  Fagus forest, under leaves and dead wood, 630 m a.s.l., 40.9193°N, 48.0027°E  ;   1 ♂ (SMNG),  İsmayıllı rayon,  Xanəgah 2 rkm towards  İsmayıllı , flat area with old  Fagus forest, with channels, under leaves, 650 m a.s.l., 40.8233°N, 48.1518°E  ;   1 ♀ (SMNG),  İsmayıllı rayon, S of  Zərgəran , slope with  Corylus ,  Clematis and some  Prunus trees, stone heaps overgrown by moss, mainly in thick leaf litter and under stones, 880 m a.s.l., 40.7310°N, 48.3680°E, all leg. F. Walther, H. Reip, D.  Antić . 
            </p>
            <p>Diagnosis.</p>
            <p>Differs from all congeners by the following combination of somatic and gonopodal characters. Head without frontal setae. Collum and metazonae of following body rings without setae. Eyes absent. Solenomere in apical part with a group of small spines. Anterior process of opisthomere subtriangular apically.</p>
            <p>Redescription.</p>
            <p> Length of adults 26-33 mm (♂♂) or 26-34 mm (♀♀), width 1.2-1.3 mm (♂♂) or 1.2-1.4 mm (♀♀). Number of body rings in adults, 50-65+1-2+T (♂♂) or 49-70+1-2+T (♀♀). Body subcylindrical (typical of  Julidae ), live specimens with brownish grey pro- and metazonae (Fig. 8C, D), after storage in alcohol prozonae grey, metazonae yellow (Fig. 1E). Head, collum, a few postcollum rings, last body rings, telson and anal valves yellow (Figs 1E, 8C, D). Antennae, mouthparts, and legs light yellow (Fig. 11A-G). Eyes absent. Striations of metazonae deep, not reaching the caudal margin, 23-25 striae per quarter of metazonital surface, i.e., that between dorsal axial line and ozopore (Fig. 11D). Ozopores small, lying behind suture without touching it (Fig. 11H). </p>
            <p>Antennae relatively long, in situ reaching segment 3. Head without frontal setae, 9+9-12+12 labral and 2+2 supralabral setae (Fig. 11A-C). Collum and metazonae without setae (Fig. 11A-G). Gnathochilarium with 3-4 long setae on each lamella lingualis, stipites with a medial curved row of 4-5 thick setae and three long setae at anterolateral margin (Fig. 11I). Epiproct undeveloped (Fig. 11E, F). Hypoproct rounded, with several setae (Fig. 11G). Telson covered with long setae, anal valves densely setose.</p>
            <p> Male. Mandibular stipites unmodified (Fig. 11A). Leg pair 1 small, unciform, telopodites directed anteromesad (as in most  Julidae ), with long setae on each coxa and in basal part of telopodite (Fig. 11J). Leg pair 2 with pads on postfemur and tibia (Fig. 11K). Penes short, bifurcate. Ventral edge of male pleurotergum 7 with wide curved lamellae bordering the gonopodal aperture (Fig. 11L). </p>
            <p>Gonopods (Fig. 12) with anterior and posterior parts both equal in height. Promere spoon-shaped, constricted in basal third; mesal ridge relatively narrow along 2/3 extent; with denticles in apical part: mesal denticle small and broadly rounded, lateral one well-expressed and long (Fig. 12B, H). Mesomeral process simple, slightly curved, with a wide subquadrate lamella apically (Fig. 12A, C, G, I). Opisthomere bipartite. Solenomere elongated, with a caudomesal, notched, membranous lobe, in apical part with a fovea and a group of small spines (Fig. 12A, C-F). Fovea may be equipped with a filiform process (Fig. 12E). Anterior process subtriangular apically.</p>
            <p>Female. First two leg pairs unmodified. Operculum of vulva without setae on caudal surface, apical margin poorly divided (Fig. 15E). Bursa mostly symmetric, lateral valve slightly larger than mesal one. Each valve with two rows of long setae. Median field of bursa narrow; emargination of median field suboval.</p>
            <p>Remarks.</p>
            <p> This species was described from the Talysh Mts, Lenkoran, Azerbaijan (Attems 1927). In the Caucasus, this is probably one of the most common and widespread congeners. Like  S. continentalis , it inhabits various deciduous forests, but it can only be considered as subendemic to the Hyrcanian biogeographic province (Fig. 16). </p>
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	https://treatment.plazi.org/id/366D2F9E79FA5C68BFAE74A6DE9E6FBC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evsyukov, Aleksandr P.;Golovatch, Sergei I.;Antic, Dragan Z.	Evsyukov, Aleksandr P., Golovatch, Sergei I., Antic, Dragan Z. (2021): The millipede genera Amblyiulus Silvestri, 1896 and Syrioiulus Verhoeff, 1914 in the Caucasus, with notes on their distributions (Diplopoda, Julida, Julidae). ZooKeys 1048: 109-143, DOI: http://dx.doi.org/10.3897/zookeys.1048.68454, URL: http://dx.doi.org/10.3897/zookeys.1048.68454
04C3854959AC5E77992CAD4439BDBD6E.text	04C3854959AC5E77992CAD4439BDBD6E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Syrioiulus Verhoeff 1914	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Genus  Syrioiulus Verhoeff, 1914</p>
            <p>Type species.</p>
            <p> Dolichoiulus polyzonus Attems, 1910, by subsequent designation of Jeekel (1971). </p>
            <p>Diagnosis.</p>
            <p> All characters as in  Amblyiulus , except as follows. Promere usually with two denticles in apical part. Head with or without frontal setae. Eyes present or absent. Opisthomere of posterior gonopod bipartite: a solenomere (with a distinct fovea on top) and an anterior process, vs. tripartite in  Amblyiulus . </p>
            <p>Species included.</p>
            <p> Syrioiulus adsharicus (Lohmander, 1936),  Syrioiulus andreevi Mauriès , 1984,  Syrioiulus aharonii (Verhoeff, 1914),  Syrioiulus armeniacus sp. nov.,  Syrioiulus continentalis (Attems, 1903),  Syrioiulus discolor (Lohmander, 1932),  Syrioiulus incarnatus (Lohmander, 1932),  Syrioiulus lohmanderi Vagalinski, 2020,  Syrioiulus persicus (Golovatch, 1983),  Syrioiulus polyzonus (Attems, 1910),  Syrioiulus taliscius (Attems, 1927), and several others provisionally listed by Vagalinski (2020). </p>
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	https://treatment.plazi.org/id/04C3854959AC5E77992CAD4439BDBD6E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evsyukov, Aleksandr P.;Golovatch, Sergei I.;Antic, Dragan Z.	Evsyukov, Aleksandr P., Golovatch, Sergei I., Antic, Dragan Z. (2021): The millipede genera Amblyiulus Silvestri, 1896 and Syrioiulus Verhoeff, 1914 in the Caucasus, with notes on their distributions (Diplopoda, Julida, Julidae). ZooKeys 1048: 109-143, DOI: http://dx.doi.org/10.3897/zookeys.1048.68454, URL: http://dx.doi.org/10.3897/zookeys.1048.68454
