taxonID	type	description	language	source
F12A87E8FFADFFD5FDBAFC5EFDECFDA6.taxon	description	Molecular analyses In the phylogenetic relationships among species of Taeniandrena (Fig. 1), all species represented by more than one individual formed well-supported monophyletic groups, with the exception of A. wilkella (Kirby, 1802), for which support was low (posterior probability, hereafter PP, of 0.74). These analyses confirm the distinctiveness of A. beaumonti Benoist, 1961 stat. rev. and A. wilkella, which form two separate clades with A. benoisti Wood & Praz sp. nov. sister to A. wilkella and A. beaumonti sister to A. benoisti + A. wilkella. The distinctiveness of A. gelriae s. tr. and A. gredana stat. nov. is also supported; these taxa do not form a monophyletic group. The sampled specimens of A. ovatula (Kirby, 1802) were similar to specimens of A. ovatula s. str. from northern Europe, a taxon that we consider distinct from most populations of ‘ A. ovatula auct. ’ from central and southern Europe (Praz & Wood, in prep.). Lastly, A. levante Wood & Praz sp. nov. was the sister species to A. gelriae s. str. (PP less than 0.5). Average within-species, uncorrected genetic distances were low for A. ovatula, A. gredana, A. levante sp. nov. and A. benoisti sp. nov. (0.022 %, 0.66 %, 0.0 % and 0.43 %, respectively) and considerably higher for A. wilkella (1.18 %) and A. similis Smith, 1849 (1.17 %). Uncorrected genetic distances between A. benoisti sp. nov. and A. wilkella were on average 3.30 % (range 2.86 – 3.86 %), those between A. benoisti sp. nov. and A. beaumonti stat. rev. 7.31 % (range 7.13 – 7.61); Andrena beaumonti was on average 6.89 % divergent from A. wilkella (range 6.62 – 7.26 %). Distances between A. gredana and A. gelriae were on average 3.02 % (range 2.98 – 3.07), those between A. levante sp. nov. and A. gelriae 1.67 % (range 1.51 – 1.75). The analyses of the Ptilandrena-Euandrena (Fig. 2) clade strongly support the recognition of A. impressa stat. nov. as a distinct species and not as a subspecies of A. angustior; these taxa were not closely related in our trees (Fig. 2). The placement of A. impressa (sister to a clade composed of A. fulvata Stoeckhert, 1930, A. angustior, A. allosa Warncke, 1975 and A. amieti Praz, Müller & Genoud, 2019) was surprising given that A. impressa shares numerous morphological features with A. fulvata and A. angustior, in particular the broadened gena and long male mandible, this character being absent in A. allosa and A. amieti, displaying the ‘ typical’ unbroadened gena found in Euandrena. Support for this arrangement was however very weak. Average genetic distances within A. angustior and A. fulvata were 0.20 % and 0.38 %; distances between these two taxa were on average 3.56 % (range 3.10 – 4.15). The distances between the single specimen of A. impressa and these two taxa were considerably higher: on average 8.22 % for distances with A. fulvata (range 8.08 – 8.40) and 8.61 % with A. angustior (range 8.39 – 9.22). With the exception of the difference between A. gelriae and A. levante sp. nov., these between species genetic distances are well above the 2 % divergence metric that typically indicates species-level differences (Schmidt et al. 2015).	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFAFFFD9FE5DF987FD39FB92.taxon	description	Figs 3 – 8	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFAFFFD9FE5DF987FD39FB92.taxon	materials_examined	Material examined Holotype SPAIN • ♂; Sierra de Guadarrama; Dusmet leg.; OÖLM. Other material PORTUGAL • 2 ♀♀; Minho, Ruivães, N 103; 12 May 2019; Wood leg.; TJWC • 1 ♀; Minho, Bastelo, Várzea Cova; 14 May 2019; Wood leg.; TJWC. SPAIN • 21 ♂♂, 2 ♀♀; Ávila, Sierra de Gredos, La Plataforma; 1700 m a. s. l.; 19 May 1995; H. and J. E. Wiering leg.; NMNL (illustrated Figs 4 – 8) • 1 ♂; Ávila, 2 km E of Hoyos Del Espino; 22 May 1995; H. and J. E. Wiering leg.; NMNL • 2 ♂♂; Ávila, Hoyos del Espino; 1400 m a. s. l.; 20 – 22 May 1995; H. and J. E. Wiering leg.; NMNL • 1 ♀; Madrid, Alto de los Leones; 22 May 1979; H. Teunissen leg.; NMNL • 1 ♂; Huesca, San Juan de la Peña; 14 May 1995; H. and J. E. Wiering leg.; NMNL. Literature records (Warncke 1976) PORTUGAL: Coimbra. SPAIN: Ávila, Puerto del Pico; Pontevedra, Tuy; León, Puerto de Leitariegos; Segovia, San Rafael; Segovia, Valsain; Cáceres, Banos; Madrid, Cercedilla; Madrid, El Escorial; Madrid, Ribas de Jarama.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFAFFFD9FE5DF987FD39FB92.taxon	discussion	Remarks Identification of female material from species close to A. gelriae is highly challenging without access to confidently determined comparative material. In Iberia, female A. gredana can be recognised by the dense punctures of the base of T 2 (smooth to weakly and sparsely punctate in A. similis Smith, 1853), faint punctures on the disc of T 1 (strongly punctate in A. wilkella (Kirby, 1802 )), fore part of clypeus shiny (Fig. 5), contrasting the central and basal areas which are dull (uniformly dull in other species), the sculpturing of the scutum (Fig. 6), which is shagreened but centrally has a circular area where the shagreenation is comparatively weaker when viewed dorsolaterally (other species uniformly shagreened or with a clearly shiny central circular area), and the comparatively wide hair bands on the terga (Fig. 7) that are complete on T 2 + 3 + 4 (either much thinner, or not complete on T 2 and / or T 3), Males are significantly easier to identify, and can initially be recognised by the length ratios of the antennal segments, being part of the group where A 3 is equal to A 4 in length. In this group, the genitalia of A. gredana are highly distinctive, showing gonocoxa that diverge only slightly at their apexes, and with a penis valve that is clearly narrowed basally and widened to its maximum width centrally before tapering to its apex (Fig. 8). This character allows separation from all other Iberian Taeniandrena. In fact, molecular data shows that this taxon is not closely allied to A. gelriae (Fig. 1), and the divergent structure of the penis valve would support this conclusion when compared to the species allied to A. gelriae (Figs 9 – 11). Given these differences and its phylogenetic placement, A. gredana is formally raised to species status.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFAFFFD9FE5DF987FD39FB92.taxon	distribution	Distribution Central and northern Spain and Portugal (Warncke 1976; Fig. 3). The distribution in Iberia is montane, and it may also occur in the French Pyrenees (see record from Huesca Province; Fig. 3), but no specimens have yet been reported from this region.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFA0FFDCFE37FBBBFE7DFC45.taxon	description	urn: lsid: zoobank. org: act: 6 B 3 F 3 AE 1 - AD 7 B- 4 A 22 - A 351 - 04 E 8 B 4 A 6 E 911 Figs 3, 10, 12 – 19	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFA0FFDCFE37FBBBFE7DFC45.taxon	diagnosis	Diagnosis Females of Andrena levante Wood & Praz sp. nov. are close to those of A. gelriae, with hair bands that are not as dense, as wide, or as complete as in A. gredana stat. nov. (Figs 7, 15). The clearest difference can be seen on the scutum which is strongly and densely shagreened and dull (Fig. 14), with dense punctures that are almost contiguous (except posteriorly), giving the overall surface a duller appearance than in either A. gelriae or A. gredana (Fig. 6) that have sparser punctation. The pubescence of the scutum and scutellum is also denser, shorter, and thicker than either comparison species (Figs 4, 12). As for other similar species of Taeniandrena, male identification is much easier. Males can be recognised as part of the group with A 3 equal to or slightly shorter than A 4 (A 3 1 – 1.03 times as long as A 4). The genitalia are distinctive, with the gonocoxa diverging from close to their base (without their inner margins parallel for at least 50 % of their length), the penis valve uniformly wide (not constricted medially), and with the blades of the gonostyli comparatively short, apically as wide as long (Fig. 10). Andrena levante sp. nov. differs from A. gelriae vocifera in numerous morphological features; this taxon will be characterised elsewhere (Praz & Wood, in prep.).	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFA0FFDCFE37FBBBFE7DFC45.taxon	etymology	Etymology The term ‘ El Levante ’ is the Spanish name for the eastern part of the Iberian Peninsula that constitutes the majority of the known range of this species (Almería, Granada, Málaga, Murcia, Valencia).	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFA0FFDCFE37FBBBFE7DFC45.taxon	materials_examined	Material examined Holotype SPAIN • ♂; 80 km SW of Valencia, Muela de Cortes reserve; [39.219 ° N, 0.957 ° W]; 14 May 2003; J. Halada leg.; BOLD accession number: HYMAA 245 - 21; OÖLM (illustrated Figs 10, 16 – 19). Paratypes SPAIN • 4 ♂♂, 23 ♀♀; Murcia, Sierra de Españula; 14 May 2003; J. Halada leg.; OÖLM • 2 ♂♂, 4 ♀♀; same collection data as for preceding; TJWC (illustrated Figs 12 – 15) • 2 ♂♂; Málaga, between Mijas and Benalmadena; 16 Apr. 1983; NMNL • 1 ♂; Almería, E-Sierra Nevada, near Alboloduy; 6 – 7 May 2003; J. Halada leg.; CPC • 1 ♂; same collection data as for preceding; OÖLM • 1 ♂; Murcia, 25 km SW of Cartagena; 12 May 2003; J. Halada leg.; OÖLM • 7 ♂♂, 2 ♀♀; Valencia, 80 km SW of Valencia, Muela de Cortes reserve; 14 May 2003; J. Halada leg.; OÖLM • 3 ♂♂, 1 ♀; same collection data as for preceding; TJWC • 1 ♀; Granada, Maitena, 9 km E of Granada; 1400 m a. s. l.; 1 Jun. 1970; M. J. and J. P. Duffels leg.; NMNL.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFA0FFDCFE37FBBBFE7DFC45.taxon	description	Description Female MEASUREMENTS. Body length 11 – 12 mm (Fig. 12). HEAD. 1.3 times as wide as long (Fig. 13). Clypeus dark, flattened over most of its area, densely and uniformly punctate with exception of raised central impunctate line, punctures separated by <0.5 puncture diameters, underlying surface shagreened, weakly shining, particularly apically. Face, gena, vertex, and scape with light brownish hair, longest not exceeding half of scape in length. Antennae dark, A 4 – 12 lightened to light brown below. Foveae broad, occupying almost all area between lateral ocellus and top of compound eye, filled with short brown hairs. MESOSOMA. Scutum densely punctate, punctures separated by <0.5 puncture diameters over majority of surface except becoming slightly sparser centrally and posteriorly, underlying surface shagreened, weakly shining (Fig. 14). Scutellum with sparser punctures separated by 1 puncture diameter, shagreenation weaker, generally shining. Episternum and propodeum with dense raised reticulation, underlying surface dull, propodeal triangle weakly indicated by weak carina, little differentiated from general reticulation. Scutum and scutellum with short, orange-brown, semi-squamiform hairs, episternum with longer light brownish to white hairs, becoming orange-brown on propodeum. Legs dark, hind tibiae and tarsi of mid and hind legs orange, general pubescence light brown basally, becoming orange apically, flocculus, femoral and tibial scopae light brown to golden. Wings hyaline, venation dark orange, stigma orange, nervulus interstitial. METASOMA. Terga dark, finely shagreened and weakly shining, apical part of marginal areas lightened semi-translucent brown (Fig. 15). T 1 very finely and subtly punctured, punctures on disc scarcely visible against shagreenation, those on margin more visible, separated by 1 puncture diameter. T 2 – 4 more densely and visibly punctate, punctures separated by 0.5 puncture diameters. Terga with whitish hairbands, on T 1 represented by two very widely separated spots (separated by almost entire width of tergal margin), T 2 widely interrupted, T 3 + 4 complete. Remaining tergal surface covered with short, fine brown to ferruginous hairs visible when viewed obliquely or in profile. Terminal fringe of T 5 and hairs flanking pygidial plate golden, pygidial plate rounded, flat, without raised margin. Male MEASUREMENTS. Body length 10 – 11 mm (Fig. 16). HEAD. 1.3 times as wide as long (Fig. 17). Clypeus flattened and densely punctate, punctures separated by <0.5 puncture diameter, sculpturing as in female. Gena and lower part of face with white hairs, becoming light brown on scape and vertex, longest equalling length of scape. Antennae dark, A 4 – 13 extensively lightened to dark brown below. A 3 as long as A 4. MESOSOMA. Scutum, scutellum, episternum, and propodeum structurally as in female (Fig. 18). Scutum and scutellum with fine light brown to golden hairs that equal length of scape, becoming light brown to whitish on propodeum and episternum. Legs dark, apical tarsal segments lightened dark red, pubescence whitish to light brownish. Wings hyaline, venation dark orange, nervulus slightly postfurcal. METASOMA. Terga dark, finely shagreened and weakly shining, apical part of marginal areas lightened semi-translucent brown (Fig. 19). Terga finely but clearly punctate, puncture separated by 0.5 – 1 puncture diameter. T 2 – 5 with hairbands, on T 2 medially interrupted, complete on T 3 – 5. S 8 strap-like, slightly broadened apically, uniformly hairy. Genitalia elongated oval-shaped in dorsal view, gonocoxa with inner margins clearly diverging, not parallel, forming 90 ° angles apically (Fig. 10). Penis valve moderately broad, basally parallel sided before tapering apically. Gonostyli comparatively short, apical blades as wide as long.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFA0FFDCFE37FBBBFE7DFC45.taxon	distribution	Distribution Areas broadly near the coast in southeastern Spain, from Málaga to Valencia (Fig. 3). All sites are mountainous (Sierra de Mijas, Sierra Nevada, Sierra de Españula, Muela de Cortes, Sierra de la Muela, Cabo Tiñoso y Roldán).	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFA5FFDFFE52FBEDFD6BF9A6.taxon	description	Figs 21, 23, 25, 27, 29, 31, 33, 35	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFA5FFDFFE52FBEDFD6BF9A6.taxon	materials_examined	Material examined MOROCCO • 1 ♀; Tizi-n-Test, S; 1900 m a. s. l.; 29 Jun. 1987; M. Schwarz leg.; OÖLM • 1 ♀; same collection data as for preceding; TJWC • 1 ♂, 1 ♀; Oukaimeden; 2700 m a. s. l.; 8 May 2015; K. Deneš leg.; OÖLM (illustrated Figs 21, 23, 25, 27, 29, 31, 33, 35).	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFA5FFDFFE52FBEDFD6BF9A6.taxon	description	Description Male MEASUREMENTS. Body length 11 mm (Fig. 29). HEAD. Dark, slightly as wide as long. Clypeus flattened, densely punctate, punctures separated by 0.5 puncture diameters, underlying surface shiny, without microsculpture. Gena, face, vertex, and scape covered with long whitish hairs that equal or exceed scape in length. Antennae dark, A 4 – 13 slightly lightened grey below, A 3 1.15 times as long as A 4 (Fig. 31). MESOSOMA. Scutum shagreened and dull over majority of disc, becoming weaker centrally and posteriorly, here weakly shining, surface densely but shallowly punctured, punctures apically separated by 0.5 puncture diameters, centrally becoming sparser, here separated by 1 – 2 puncture diameters. Scutellum more uniformly shiny and densely punctate, punctures separated by 1 puncture diameter. Episternum and propodeum weakly reticulate, underlying surface weakly shining, propodeal triangle basally comparatively more strongly reticulate, this dispersing apically. All parts of mesosoma with long brownish (scutum and scutellum) to whitish (episternum and propodeum) hairs that exceed length of scape. Legs dark, apex of hind tibiae and entirety of hind basitarsi orange, pubescence whitish. Wings hyaline, venation orange, nervulus interstitial. METASOMA. Terga dark, apical margins impressed, strongly impressed on T 3 – 5, covered in short, thick, and complete white hair bands (Fig. 33). Terga densely punctate, punctures separated by 0.5 – 1 puncture diameters, underlying surface shagreened and dull basally, becoming weaker apically, T 4 – 5 weakly shining. S 8 columnar, parallel sided, apically slightly notched. Genitalia simple, gonocoxa with internal margins diverging apically, laterally with weak shagreenation, apical corners rounded (Fig. 35). Penis valve relatively narrow, apically tapering, gonostyli with broad bases, apical blades clearly longer than wide.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFA5FFDFFE52FBEDFD6BF9A6.taxon	discussion	Remarks In his original description, Benoist (1961) drew a comparison between A. beaumonti stat. rev. and Andrena flavipes Panzer, 1799, presumably because of the dense hair bands that help characterise this species. Warncke (1967) noted that this association was clearly incorrect at a subgeneric level, associating the bee with members of the subgenus Taeniandrena instead. On the basis of its large and dense puncturing, its golden hairs flanking the pygidial plate, and the orange metatarsi of the second pair of legs, Warncke (1967) associated this taxon with Andrena wilkella, but as a subspecies because of the unbroken hair bands on T 3 + 4. He then went on to report A. wilkella beaumonti from Iberia (Warncke 1976; see also Ortiz-Sánchez 2011, 2020), giving it a distribution of Morocco, Spain, and Portugal (see also Lhomme et al. 2020). However, genetic barcoding shows that A. beaumonti is distinct, and Moroccan material is also morphologically different from Warncke’s concept of A. wilkella beaumonti in Iberia, most obviously by lacking a shiny spot on the scutum that is well differentiated from the surrounding shagreenation. It can also be separated by the finer hairs on the scutum, the generally paler pubescence, and by the thicker and more complete hair bands on terga 2 – 4 (Figs 20 – 27), meaning that A. wilkella beaumonti sensu Warncke in Iberia is actually undescribed (see below). The previously undescribed male of A. beaumonti can also be distinguished using the same characters (colour of pubescence, nature of tergal hair bands) and also by the length of A 4 which is 1.15 times as long as A 3, whereas in A. benoisti Wood & Praz sp. nov. (see below) it is 1.3 times longer (Figs 30 – 31). There are also slight differences in the genitalia, with a comparatively narrower penis valve and more clearly separated gonocoxa (Figs 34 – 35) in A. beaumonti.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFA5FFDFFE52FBEDFD6BF9A6.taxon	distribution	Distribution Morocco, from the High Atlas Mountains in the area south of Marrakesh. The village of Asni is found at a moderate elevation of 1200 m. It is not clear exactly where Arroud is because of changes in spelling practices, but it probably refers to the village Aroumd some 25 km SSE of Asni. Contemporary sampling locations of A. beaumonti stat. rev. are relatively close to Asni itself. Tizi-n-Test is some 60 km SW of Asni, and Oukaimeden is even closer, just 10 km to the SE. The site of Oukaimeden is at a much higher elevation of 3200 m, and the cluster of records in this region suggests that A. beaumonti is restricted to high-elevation sites flying in May and June.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFA6FFC1FE2BF983FAEEF867.taxon	description	urn: lsid: zoobank. org: act: 7 B 5423 C 4 - E 358 - 4 C 5 D- 8 C 92 - 6 A 6074033347 Figs 3, 20, 22, 24, 26, 28, 30, 32, 34	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFA6FFC1FE2BF983FAEEF867.taxon	diagnosis	Diagnosis Andrena benoisti Wood & Praz sp. nov. can be recognised within the subgenus Taeniandrena by the combination of dense punctures on the base of T 2, strong punctures on the disc of T 1 (very similar to A. wilkella), thick hairbands that are only narrowly interrupted on T 2 and complete on T 3 + 4 (Fig. 26, thin and widely interrupted on T 2 + 3 in A. wilkella), and the sculpturing of the scutum, which is shagreened except for the centre where it has a circular shining area that strongly contrasts with the remaining surface which is dull (Fig. 24). The males can be recognised by the ratio of the antennal segments where A 3 is clearly shorter than A 4 in length (Fig. 30), the terga are strongly and densely punctate, the tergal margins are depressed with a thin, shiny, puncture-free apical zone, and T 2 – 4 have thick hairbands, medially interrupted on T 2, complete on T 3 + 4 (Fig. 32). They are therefore closest to A. wilkella, but the thicker and more complete hair bands in combination with its larger body size (10 – 11 mm against 8 – 9 mm) allows for differentiation. Genetically, A. benoisti sp. nov. is placed as the sister of A. wilkella, a position that is corroborated by their morphological similarity.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFA6FFC1FE2BF983FAEEF867.taxon	etymology	Etymology Named after Raymond Benoist, the French botanist and entomologist who described Andrena beaumonti stat. rev. from Morocco, this name being incorrectly applied at the subspecific level to the taxon we describe here.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFA6FFC1FE2BF983FAEEF867.taxon	materials_examined	Material examined Holotype PORTUGAL • ♀; Minho, Confurco, Várzea Cova; 41.4978 ° N, 8.0765 ° W; 14 May 2019; Wood leg.; BOLD accession number: HYMAA 239 - 21; OÖLM (illustrated Figs 20, 22, 24, 26). Paratypes PORTUGAL • 1 ♂, 1 ♀; Minho, Ruivães, N 103; 12 May 2019; Wood leg.; OÖLM • 1 ♂; Minho, 1.5 km E of Lindoso; 13 May 2019; Wood leg.; TJWC • 1 ♂; Minho, Serra do Gerês, 5 km W of Paradela; 12 May 2019; Wood leg.; OÖLM (illustrated Figs 28, 30, 32, 34) • 1 ♀; Trás-os-Montes, Curalha, A 24 and N 103 intersection; 16 May 2019; Wood leg.; TJWC • 1 ♀; Trás-os-Montes, Chaves, Estr. Braga, Rio Tâmega; 16 May 2019; Wood leg.; TJWC. SPAIN • 1 ♂; Cáceres, W of Garganta la Olla; 1000 m a. s. l.; 9 May 1999; H. and J. E. Wiering leg.; NMNL • 4 ♂♂; Cáceres, Piornal; 1050 m a. s. l.; 13 May 1999; H. and J. E. Wiering leg.; NMNL • 1 ♂; Cáceres, Madrigal de la Vera; 500 m a. s. l.; 9 May 1999; H. and J. E. Wiering leg.; NMNL • 3 ♂♂; Ávila, Sierra de Gredos, La Plataforma; 1700 m a. s. l.; 19 May 1995; H. and J. E. Wiering leg.; NMNL • 1 ♂; Ávila, Navarredonda de la Sierra en Pico de Almanzor; 1400 – 1600 m a. s. l.; 5 Jun. 1976; P. Oosterbroek leg.; NMNL • 1 ♀; Ávila, Sierra de Gredos Puerto del Pico; 2 Jul. 1988; M. Schwarz leg. • 2 ♂♂; Ávila, 2 km E of Hoyos Del Espino; 22 May 1995; H. and J. E. Wiering leg.; NMNL • 11 ♂♂; Ávila, Hoyos del Espino; 1400 m a. s. l.; 18 – 22 May 1995; H. and J. E. Wiering leg.; NMNL • 1 ♂; Gredos, Hoyos de Collado; 6 Jun. 1983; H. Teunissen leg.; NMNL • 1 ♂; Ávila, 6 km E of Parador del Gredos; 1 Jun. 1976; P. Oosterbroek leg.; NMNL. Other material (unspecified, listed as Andrena wilkella beaumonti by Warncke 1976) PORTUGAL • Coimbra, Pinhal de Marrocos; Ponte da Portella. SPAIN • Parador Nacional de Gredos; Zamora, Alcubilla de Nogales; Cáceres, Banos do Montemayor; Cáceres Tornavacas; Madrid, Cercedilla; Madrid, Ciempozuelos; Madrid, El Chaparral; Madrid, El Escorial; Madrid, El Paular; Madrid, Sierra de Guadarrama; Madrid, Vaciamadrid.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFA6FFC1FE2BF983FAEEF867.taxon	description	Description Female MEASUREMENTS. Body length 11 – 12 mm (Fig. 20). HEAD. 1.3 times as wide as long (Fig. 22). Clypeus flattened, densely but shallowly and weakly punctured, punctures separated by 0.5 puncture diameters, faint impunctate line visible centrally, underlying surface shagreened, dull, slightly shining apically. Lower face and gena with whitish hairs, becoming light brown to golden on scape, upper face, and vertex, hairs not exceeding length of scape. Antennae dark, A 5 – 12 slightly lightened grey below. Foveae broad, occupying almost all area between lateral ocellus and top of compound eye, filled with short brown hairs. MESOSOMA. Scutum densely punctured over most of surface, underlying surface dull, punctures separated by 1 puncture diameter except becoming sparser centrally and posteriorly forming circular shape, here punctures separated by 2 puncture diameters, underling surface weakly shagreened, shining, contrasting remaining surface (Fig. 24). Scutellum shiny, densely punctate centrally, separated by 0.5 puncture diameters, punctures sparser laterally. Episternum with slightly raised reticulation, underlying surface dull, reticulation reaches lateral faces of propodeum but weakly, disappearing before propodeal triangle, this marked by change in surface sculpturing, internal surface with fine, sparse, and slightly raised reticulation. Scutum and scutellum with short, orange-brown, semi-squamiform hairs, episternum with longer light brownish to white hairs, becoming orange-brown on propodeum. Legs dark, hind basitarsi fully and hind tibiae partially orange, general pubescence light brown. Flocculus and femoral scopa whitish, tibial scopa golden. Wings hyaline, venation and stigma golden, nervulus postfurcal. METASOMA. Terga dark, apical part of marginal areas lightened semi-translucent brown (Fig. 26). T 1 with clear punctures on disc, separated by 0.5 puncture diameters, extending to tergal margin, becoming sparser, separated by 1 – 1.5 puncture diameters. T 2 – 4 densely and uniformly punctate, punctures separated by 0.5 – 1 puncture diameters. T 1 with very widely separated hair patches on margin, T 2 – 4 with thick hair bands, on T 2 interrupted, on T 3 + 4 complete. Terminal fringe of T 5 and hairs flanking pygidial plate golden, pygidial plate rounded with slightly raised longitudinal area centrally. Male MEASUREMENTS. Body length 10 – 11 mm (Fig. 28). HEAD. 1.3 times as wide as long. Clypeus flattened, densely punctate, punctures separated by 0.5 puncture diameters with exception of clear longitudinal impunctate line, underlying surface shiny. Gena, face, vertex, and scape with long whitish to light brownish hairs, equalling or exceeding scape in length. Antennae dark, A 3 1.3 times as long as A 4 (Fig. 30). MESOSOMA. Scutum, scutellum, episternum, and propodeum structurally as in female, though propodeal reticulation a little more extensive. All parts of mesosoma with long light brown to golden hairs, equalling or exceeding scape in length. Legs dark, Hind tibiae entirely and hind tarsi apically coloured dark orange, pubescence whitish to light brown. Wings hyaline, venation dark orange, nervulus postfurcal. METASOMA. Terga dark, apical margins slightly depressed, apical part of marginal areas lightened semitranslucent brown (Fig. 32). T 1 – 5 with discs increasing densely punctate, on T 1 punctures separated by 1 puncture diameter, almost contiguous on T 5, underlying surface shagreened, weakly shining. Tergal discs with loose light to dark brown hairs, T 2 – 4 with thick hair bands, on T 2 widely interrupted, on T 3 + 4 complete. S 8 broadened apically, densely and uniformly hairy. Genitalia simple, gonocoxa with inner margins parallel, only slightly deviating apically, apical corners rounded (Fig. 34). Penis valve moderately broad, parallel sided basally before strongly tapering apically, gonostyli with broad bases, apical blades clearly longer than wide.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFA6FFC1FE2BF983FAEEF867.taxon	distribution	Distribution From the Sistema Central around Madrid westwards through the Sierra de Gredos into central Portugal, and north into northern Portugal, Galicia, and Zamora (Fig. 3). Both A. benoisti sp. nov. and A. gredana stat. nov. show a remarkably similar distribution and occurring at many of the same localities (see also Warncke 1976), particularly in the Sistema Central from which true A. wilkella seems to be absent.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFBBFFC5FE0EFEEDFE1FF9E5.taxon	description	urn: lsid: zoobank. org: act: A 2 AD 083 A-AB 1 A- 4 B 85 - AE 9 D-F 58 DA 0 B 653 A 7 Figs 36 – 39, 42 – 45	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFBBFFC5FE0EFEEDFE1FF9E5.taxon	diagnosis	Diagnosis In the female sex, A. fortipunctata Wood sp. nov. can be placed in the subgenus Euandrena because of the narrow, comma shaped fovea, antennal A 3 longer than A 4 + 5, metasomal integument brown haired, propodeal triangle weakly wrinkled, hind femur without thorn-like projections, and dorsolateral angle of propodeum without elevated transverse carina (Praz et al. 2019). The female resembles A. bicolor Fabricius, 1775 in general colour pattern, but differs by the combination of a coarsely and densely punctate clypeus (punctures separated by 0.5 puncture diameters, interspaces barely visible, in A. bicolor punctures separated by 0.5 – 1 puncture diameters, interspaces visible), strongly punctured tergal discs, and strongly depressed tergal margins (compare Figs 38 – 41). Males are harder to place through specific subgeneric characters, in common with other Euandrena males as the subgenus is largely defined by female characters (Praz et al. 2019). However, A. fortipunctata sp. nov. males are instantly recognisable in the Iberian fauna by the wide and extremely strongly depressed tergal margins (Figs 43, 45) that are strikingly more pronounced that in either A. granulosa Pérez, 1902 or A. vulpecula Kriechbaumer, 1873 (Figs 47, 49), the other two European species of A. (Euandrena) known for their distinctive depressed tergal margins. These three species also share similar facial pubescence which is centrally yellow and laterally dark (Figs 46, 48), whereas other Iberian species of Euandrena have male facial pubescence that is either predominantly grey (A. symphyti Schmiedeknecht, 1883), brown (A. rufula Schmiedeknecht, 1883), dark (A. bicolor), or a mixture of grey and black (A. allosa Warncke, 1975).	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFBBFFC5FE0EFEEDFE1FF9E5.taxon	etymology	Etymology The name ‘ forti ’ (‘ strong’) + ‘ punctata ’ (‘ punctured’) was chosen because of the pronounced punctures visible in both sexes that help separate this taxon from other Iberian Euandrena.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFBBFFC5FE0EFEEDFE1FF9E5.taxon	materials_examined	Material examined Holotype SPAIN • ♂; Oveido-León, Puerto de Pajares; 1350 – 1700 m a. s. l.; [42.994 ° N, 5.763 ° W]; 11 Jul. 1972; V. S. May d. Groot and J. A. W. Lucas leg.; NMNL (illustrated Figs 36 – 39). Paratype SPAIN • 1 ♀; Ávila, Sierra de Gredos, 12 km SSW of Hoyos del Espino; 1950 – 2100 m a. s. l.; 4 Jul. 1972; V. S. May d. Groot and J. A. W. Lucas leg.; NMNL (illustrated Figs 42 – 45).	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFBBFFC5FE0EFEEDFE1FF9E5.taxon	description	Description Female MEASUREMENTS. Body length 10 mm (Fig. 36). HEAD. 1.2 times as wide as long (Fig. 38). Clypeus weakly arched, very densely punctate, punctures separated by <0.5 puncture diameters to almost contiguous, underlying surface shiny. Gena, face, and scape with black hairs, face between antennal insertions with few grey hairs, vertex with intermixed black and golden hairs, hairs at most equalling length of scape. Antennae dark, A 4 – 12 lightened grey below. Fovea narrow, occupying ⅓ of area between lateral ocellus and compound eye, comma shaped, filled with black hairs. MESOSOMA. Scutum and scutellum densely punctured, punctures separated by 0.5 – 1 puncture diameters, underlying surface shagreened, weakly shining (Fig. 37). Episternum and propodeum microreticulate, dull, propodeum with large irregular shallow punctures, separated by 0.5 – 1 puncture diameters, these not extending onto propodeal triangle, therefore defining it. Scutum and scutellum with golden-brown hairs, episternum centrally with black hairs, anteroventrally with white hairs, propodeum with mixture of whitish and brownish hairs, some black hairs intermixed. Legs dark, slightly lightened to dark red apically, pubescence golden brownish. Flocculus a mixture of blackish and whitish hairs, femoral and tibial scopa golden. Wings hyaline, venation and stigma dark orange, nervulus interstitial. METASOMA. Terga dark, margins strongly depressed and lightened semi-translucent brown (Fig. 39). Terga densely and uniformly punctate, punctures separated by 0.5 – 1 puncture diameters, punctures not extending onto depressed tergal margins, excluding margins underlying tergal surface shagreened, weakly shining. T 2 – 4 with hair fringes arising from junction between disc and margin, fringes of sparse white hairs overlying marginal areas, not obscuring underlying surface. Terminal fringe of T 5 and hairs flanking pygidial plate dark brown, pygidial plate with centrally raised longitudinal area and raised apical rim, weakly punctate, dull. Male MEASUREMENTS. Body length 10 mm (Fig. 42). HEAD. 1.2 times as wide as long (Fig. 44). Clypeus weakly arched, densely punctate, punctures separated by 0.5 puncture diameters. Gena, vertex, scape, clypeus, and face centrally with yellow hairs, lower paraocular areas to frons with black hairs. Antennae dark, A 4 – 13 lightened brown below, A 3 0.9 times as long as A 4 + 5. MESOSOMA. Structurally as in female, entire surface with long light brown to golden hairs, exceeding scape in length. Legs dark, tarsi lightened, dark red apically, pubescence golden. Wings hyaline, venation and stigma orange, nervulus prefurcal. METASOMA. Terga dark, apical margins wide and very strongly depressed, semi-translucent brown, impunctate and shiny (Fig. 43). Tergal discs arched, strongly contrasting depressed marginal areas, strongly and densely punctate, punctures separated by 1 puncture diameter, surface covered with loose golden hairs (Fig. 45).	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFBBFFC5FE0EFEEDFE1FF9E5.taxon	discussion	Remarks The subgenus Euandrena is taxonomically challenging, and it is likely that we are just scratching the surface of hidden species diversity in southern European mountain chains (Praz et al. 2019). The Andrena of the Iberian alpine areas have received little recent attention, and within this context, the presence of an undetected alpine Euandrena in the high mountains of Spain is less surprising, but emphasises how much more there is to learn about European diversity of Andrena even in a comparatively well-studied region.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFBBFFC5FE0EFEEDFE1FF9E5.taxon	distribution	Distribution The Sierra de Gredos in the Sistema Central to the Cantabrian mountains in northwestern Spain in the province of Asturias.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFBCFFC8FE62F94DFD70F867.taxon	description	urn: lsid: zoobank. org: act: 73 F 5116 B- 23 D 0 - 4071 - BA 9 F- 6 B 54 A 54405 CC Figs 50, 52, 54, 56, 58, 60, 62, 64	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFBCFFC8FE62F94DFD70F867.taxon	diagnosis	Diagnosis Andrena hattorfiana (Fabricius, 1775) can be very easily recognised as it is the only known member of the Charitandrena Hedicke, 1933 in the West Palaearctic. Females have a unique combination of a conspicuously carinate hind femur, a hind tibial spur that is convexly broadened towards the middle, and an elongate glossa. Males are harder to recognise through a combination of unique characters, but their large body size, yellow clypeus, lack of a pronotal carina, and long glossa in combination with specific characters such as genital structure are sufficient to facilitate identification. Andrena hattorfiana nigricauda Wood subsp. nov. differs from the nominate form (Figs 51, 53, 55) because it has an almost completely darkened integument and pilosity. True Andrena hattorfiana normally shows extensive variation in the colour of its abdominal integument, with individuals from the same location varying from extensively red marked to completely black. Its variability has resulted in a relative large number of synonyms for an Andrena (Gusenleitner & Schwarz 2002). However, what is consistent is that the hairs of T 5 and those flanking the pygidial plate are uniformly golden-orange (Fig. 55). In the Spanish material presented here, these hairs are completely black (Fig. 54). In the male, there are no clear structural differences, but as in the female the pubescence is much darker across the whole body (Figs 58 – 63).	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFBCFFC8FE62F94DFD70F867.taxon	etymology	Etymology The name ‘ nigri ’ (‘ black’) + ‘ cauda ’ (‘ tail’) was chosen to reflect the black hairs at the apex of the metasoma in the female sex, a clear point of difference from the nominate form.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFBCFFC8FE62F94DFD70F867.taxon	materials_examined	Material examined Holotype SPAIN • ♀; Alicante, Puerto de Confrides; 1000 m a. s. l.; [38.686 ° N, 0.271 ° W]; 15 Jun. 1978; H. Teunissen leg.; NMNL (illustrated Figs 50, 52, 54, 56). Paratypes SPAIN • 2 ♂♂, 5 ♀♀; Alicante, Puerto de Confrides; 1000 m a. s. l.; 15 Jun. 1978; H. Teunissen leg.; NMNL (illustrated Figs 58, 60, 62, 64) • 1 ♀; same collection data as for preceding; OÖLM • 1 ♀; same collection data as for preceding; TJWC. Other material (Andrena hattorfiana) SPAIN • 1 ♂, 1 ♀; Madrid, Collado Mediano; 17 May 1995; H. and J. E. Wiering leg.; NMNL (illustrated Figs 59, 61, 63, 65) • 1 ♀; Pirineos Orient, 20 km NE of Ripoll; 1650 m a. s. l.; 31 Jul. 2011; J. Halada leg.; OÖLM (illustrated Figs 51, 53, 55, 57) • 1 ♀; Huesca, Mte Perdido; 1300 m a. s. l.; 1 Aug. 1977; P. M. F. Verhoeff leg.; NMNL • 1 ♀; Viella; 1100 – 1800 m a. s. l.; 27 Jul. 1963; H. Hamann leg.; OÖLM • 1 ♂; Cantabria, Picos de Europa, Camaleno; 600 m a. s. l.; 5 May 2014; D. W. Baldock leg.; TJWC • 1 ♀; Santander, Arredondo, Bustablado; 11 Jul. 1984; R. Leys leg.; NMNL.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFBCFFC8FE62F94DFD70F867.taxon	description	Description Female MEASUREMENTS. Body length 16 – 17 mm (Fig. 50). HEAD. Black, 1.2 times as wide as long (Fig. 52). Clypeus domed, slightly flattened centrally and apically, underlying surface shagreened and dull basally and laterally, becoming weaker apically, weakly shining. Entire surface densely punctured with exception of faint impunctate central line, punctures separated by 0.5 – 1 puncture diameters. Process of labrum triangular, twice as broad as long, deeply emarginate apically. Gena, vertex, face, and scape with moderately long dark brown to black hairs, longest equalling length of scape. Antennae dark, A 3 long, exceeding length of A 4 + 5 + 6, A 5 – 12 lightened orange below. Fovea of a uniform width, occupying half of area between lateral ocelli and top of compound eye. Vertex wide, as wide as three ocellar diameters. MESOSOMA. Scutum and scutellum finely shagreened, weakly shining, densely and evenly punctured, punctures separated by 1 puncture diameter. Episternum and propodeum evenly and finely rugose, propodeal triangle clearly marked by a lateral carina, internal surface with more strongly produced rugosity, broadly similar. Scutum and scutellum with short, episternum and propodeum with longer dark brown hair, at its longest not exceeding length of scape. Legs dark, apical tarsal segments lightened red-brown, pubescence dark brown to dark red. All scopal hairs dark brown, those of femoral and tibial scopa ventrally finely plumose, otherwise simple. Wings weakly infuscate, venation dark brown, nervulus interstitial. METASOMA. Terga dark brown, only apical margins slightly lightened brown (Fig. 54). Tergal surface very weakly shagreened, generally shining, centrally punctured, punctures separated by 1 – 2 puncture diameters, punctures becoming sparse laterally, here separated by 3 – 4 puncture diameters. Hairs of T 5 and those flanking pygidial plate dark brown to black, pygidial plate rounded, smooth, very weakly convex, impunctate. Male MEASUREMENTS. Body length 15 mm (Fig. 58). HEAD. Black, 1.2 times as wide as long (Fig. 60). Characters as in female, but clypeus yellow with exception of two dark spots laterally, A 3 slightly subequal to A 4 + 5 + 6, and A 4 – 13 lightened orange below. MESOSOMA. As in female. METASOMA. As in female (Fig. 62). S 7 deeply emarginate apically, S 8 columnar, apical half hairy with two broad lateral tufts centrally and one tuft apically, apex truncate. Genitalia with pronounced but truncate gonocoxal teeth, penis valve broadly triangular, tapering apically, gonostyli long and narrow with slightly flattened apexes with short, white, sparse hairs (Fig. 64).	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFBCFFC8FE62F94DFD70F867.taxon	discussion	Remarks The presence of A. hattorfiana nigricauda Wood subsp. nov. in the mountains of Alicante represents, at the very least, an isolated population showing a unique colour pattern. Other records of A. hattorfiana in Spain come from the Sistema Central around and to the west of Madrid, the Picos de Europa, and the Pyrenees (Gusenleitner & Schwarz 2002). The location in Alicante is therefore around 400 km from the nearest known populations of A. hattorfiana that conform to the typical hair colour pattern. However, separating this population as specifically distinct is premature without molecular investigation, and overall differences are slight outside of colouration; the male genitalia of the two colour forms are essentially indistinguishable. Andrena hattorfiana nigricauda subsp. nov. could simply be a melanic form, and molecular analysis could show whether it is nested within A. hattorfiana when considered across its whole range. Indeed, material of A. hattorfiana from the most southerly parts of the Balkan Peninsula has hairs flanking the pygidial plate that are dark brown, but otherwise this material shows typical red terga (treated as A. h. dimidiata Brullé, 1832 by Warncke, see Gusenleitner & Schwarz 2002). Pollen removed from the scopae of four females of A. hattorfiana nigricauda subsp. nov. (Fig. 56) consisted of Knautia - type pollen (formerly Dipsacaceae Juss., now Caprifoliaceae Juss.). Andrena hattorfiana is a very well-studied specialist of this family (Fig. 57; Westrich 1989; Larsson & Franzén 2007), suggesting that the dietary niche itself is unchanged.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFBCFFC8FE62F94DFD70F867.taxon	distribution	Distribution Andrena hattorfiana is distributed from central and northern Spain across Europe to Greece, Turkey, and the Caucasus (Gusenleitner & Schwarz 2002).	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFB3FFCBFDEEFEEDFDBAFBD3.taxon	description	Figs 66 – 69	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFB3FFCBFDEEFEEDFDBAFBD3.taxon	materials_examined	Material examined PORTUGAL • 1 ♂; Algarve, Tareja; [37.173 ° N, 7.879 ° W]; 13 Apr. 1991; J. v. Corstanje leg.; NMNL (illustrated Figs 66 – 69).	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFB3FFCBFDEEFEEDFDBAFBD3.taxon	description	Description Male MEASUREMENTS. Body length 9 mm (Fig. 66). HEAD. Black, 1.1 times as wide as long (Fig. 67). Clypeus black, slightly domed and slightly flattened centrally, densely punctured laterally with punctures separated by 0.5 puncture diameters, becoming sparser centrally, separated by 1 puncture diameter, narrow central impunctate line present. Gena, clypeus, and inter-antennal area with white hairs, vertex with a mixture of white, brown, and black hairs, and frons and inner margin with black hairs, all hairs variable in length, longest exceeding length of scape. Antennae uniformly dark, A 3 slightly shorter than A 4 + 5. MESOSOMA. Scutum shagreened, very weakly shining, densely punctured, punctures separated by 0.5 – 1 puncture diameter, slightly sparser centrally. Scutellum comparatively less shagreened, weakly shining, very densely punctured, punctures separated by <0.5 puncture diameters. Episternum and propodeum reticulate, dull. Propodeal triangle large, clearly marked with raised carina, internal surface with weak longitudinal rugae. All parts of mesosoma with long white hairs which exceed length of scape. Legs dark, apical tarsal segments lightened orange, including hind basitarsi. Wings hyaline, venation orange, nervulus interstitial. METASOMA. Terga dark, apical margins lightened dark red, very densely and uniformly punctate, punctures separated by 0.5 puncture diameters, underlying surface weakly shagreened, shining (Fig. 68). T 1 – 4 with loose apical hair bands, those on T 1 – 3 interrupted, on T 4 complete. S 8 short, projecting area as wide as long, apical margin notched. Genitalia long, gonocoxa inflated, therefore forming a channel along their internal margins, apical teeth broadly rounded (Fig. 69). Penis valve triangular, narrowing apically to a rounded point. Gonostyli narrow and elongate, dorsal face flattened and bearing sparse, short hairs.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFB3FFCBFDEEFEEDFDBAFBD3.taxon	discussion	Remarks Andrena baetica is morphologically close to A. mocsaryi Schmiedeknecht, 1883, but differs in the shorter, squamous hairs on the scutum (Wood et al. 2020). The discovery of this male material further confirms the proximity of A. baetica to A. mocsaryi through genital structure (Fig. 69, see illustrations in Schmid-Egger & Scheuchl 1997), but also confirms its specific status as it possesses an entirely black clypeus, whereas A. mocsaryi has a yellow clypeus, and the apex of S 8 is truncate to slightly emarginate, whereas in A. mocsaryi it is slightly pointed (Schmid-Egger & Scheuchl 1997).	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFB3FFCBFDEEFEEDFDBAFBD3.taxon	distribution	Distribution Southern Spain (Cadiz, Los Alcornocales; Cazorla, Sierra Pozo) and southern Portugal (Alentejo; Algarve) (Wood et al. 2020).	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFB2FFCFFDEEFB7FFBDFF8A3.taxon	description	urn: lsid: zoobank. org: act: D 61 A 664 D- 0821 - 4470 - 8807 - 6 CC 3 C 7 B 93 E 9 F Figs 70 – 72, 74, 76, 78 – 83	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFB2FFCFFDEEFB7FFBDFF8A3.taxon	diagnosis	Diagnosis This taxon can be placed in the subgenus Notandrena in the female sex because of its broad head (clearly broader than wide), the weakly rugose (not shagreened) propodeal triangle, the clearly punctured metasoma, and by the dorsolateral angle of the pronotum with a weak transverse ridge. In the male sex, recognition is easy because of the greatly enlarged and carinate gena in combination with the broadened apex of the gonostyli. Eight species of the subgenus Notandrena are known from Iberia (Wood et al. 2020). Female Andrena juliana Wood sp. nov. can be recognised in the group of A. nitidiuscula Schenck, 1853 because of its small size (7 – 8 mm, excluding A. erythrocnemis Morawitz, 1870, A. langadensis Warncke, 1965, and A. urdula Warncke, 1965 which average 11 – 12 mm in length) and dark hind tibiae (orange in A. chrysosceles (Kirby, 1802 )). Within this group, it lacks the distinctive shortened dorsal scopal hairs of A. pallitarsis Pérez, 1903, the gena and vertex are normal (vertex clearly less than the diameter of a lateral ocelli, clearly greater than this distance in A. foeniculae Wood, 2020), and there is no deeply impressed line on the front half of the scutum (clearly impressed in A. nitidiuscula). In an Iberian context, it is therefore closest to A. fulvicornis Schenck, 1853 (alternative character state in parentheses), but differs in the sparser punctures on T 1, separated by 2 – 3 puncture diameters (separated by one puncture diameter, Fig. 77), by the shinier scutellum (scutellum shagreened and clearly dull, Fig. 73), and by the hind tibiae which are dark (orange). Outside of Iberia, A. juliana Wood sp. nov. is also close to A. curvana Warncke, 1965 which is distributed from southern Germany southwards and eastwards to Romania, Bulgaria, Greece, and the European part of Turkey (Gusenleitner & Schwarz 2002; Schwenninger 2013). It can be separated by the sculpturing of the clypeus which is shiny centrally (centrally shagreened) and by the sculpturing of the scutellum which is shiny (strongly shagreened and dull). Males can be quickly recognised by their yellow clypeus in combination with their small size (7 mm), a character that is also found in A. chrysosceles and A. pallitarsis (A. erythrocnemis, A. langadensis, and A. urdula with the clypeus yellow marked, but larger, averaging 10 mm). In both these species, the yellow markings cover the entire clypeus and extend onto the lower paraocular areas, whereas in A. juliana sp. nov. this marking is restricted to the centre of the clypeus. The gena is noticeably more carinate, and as in the female the scutum is also noticeably shinier. The genitalia conform to the typical shape found in members of the nitidiuscula group (Fig. 83; see illustrations in Schmid-Egger & Scheuchl 1997; Schwenninger 2013). As for other members of this group, the gonocoxa are weakly shagreened in a similar fashion to that found in the Zonandrena Hedicke, 1933. However, the outer margin of the gonostyli is almost straight, whereas there is a clearer obtuse angle here in A. chrysosceles, A. pallitarsis, and A. fulvicornis (see illustrations in Schmid-Egger & Scheuchl 1997; Schwenninger 2013). As in the female sex, the male is also similar to the eastern A. curvana. However, apart from the yellow clypeus (black in A. curvana), the genitalia are different, with less pronounced gonocoxal teeth, and lacking the pronounced impressed channel in the basal section of the gonostyli (see illustration of this character in Schwenninger 2013).	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFB2FFCFFDEEFB7FFBDFF8A3.taxon	etymology	Etymology The name is taken from the locus typicus, San Julián.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFB2FFCFFDEEFB7FFBDFF8A3.taxon	materials_examined	Material examined Holotype SPAIN • ♀; Málaga, San Julián 8 km SW of Málaga; [36.666 ° N, 4.476 ° W]; 25 May 1962; Jeekel & Wiering leg.; NMNL (illustrated Figs 70 – 72, 74, 76). Paratype PORTUGAL • 1 ♂; Algarve, Gambello [Gambelas]; 5 Mar. 1986; H. Teunissen leg.; NMNL (illustrated Figs 78 – 83).	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFB2FFCFFDEEFB7FFBDFF8A3.taxon	description	Description Female MEASUREMENTS. Body length 8.5 mm (Fig. 70). HEAD. Dark, broad, 1.4 times as wide as long (Fig. 71). Clypeus domed, laterally and dorsally shagreened and densely punctate, punctures separated by 0.5 puncture diameters, centrally shiny and less densely punctate, punctures separated by 0.5 – 1.5 puncture diameters, weakly-marked impunctate line present. Process of labrum small, rectangular, front margin very weakly emarginate. Gena as wide as compound eye, gena, vertex, and face with whitish hairs, these not exceeding length of scape. Antennae dark, A 5 – 12 lightened orange below, A 3 equalling A 4 + 5, both A 4 and A 5 clearly broader than long. Facial fovea neither wide nor narrow, occupying half space between lateral ocellus and compound eye. Ocelloccipital distance short, less than half diameter of lateral ocellus. MESOSOMA. Scutum weakly shagreened, gently shining, densely but shallowly punctured, punctures separated by 0.5 – 1 puncture diameter centrally (Fig. 72). Scutum in fore half with barely impressed mid line. Scutellum more clearly shiny, punctures dense only at margin and forming a central longitudinal line, otherwise sparse and separated by 3 – 4 puncture diameters. Episternum and propodeum finely reticulate, dull, propodeal triangle barely indicated by very fine carina, internal structure essentially unchanged. Scutum and scutellum with short whitish pubescence through which longer hairs protrude, these approaching length of scape. Episternum and propodeum with longer whitish hairs, longest equalling length of scape. Legs dark, apical tarsal segments and apex of basitarsi coloured orange, this extending slightly onto apex of hind tibiae. Wings hyaline, venation orange, nervulus interstitial. METASOMA. Terga dark, apical margins lightened yellow-brown, underlying surface weakly shagreened, shining (Fig. 74). T 1 sparsely and finely punctate on disc, punctures separated by 2 – 4 puncture diameters (Fig. 76), strongly contrasting with following terga, T 2 – 4 densely punctate on discs, punctures separated by 0.5 – 1 puncture diameter. Marginal areas of T 1 – 4 less densely punctate, punctures separated by 1 – 2 puncture diameters. T 2 – 4 with fringes of white hair, interrupted on T 2 – 3, complete on T 4. T 5 and hairs flanking pygidial plate light brown, pygidial plate smooth, apically rounded, weakly shining. Male MEASUREMENTS. Body length 7 mm (Fig. 78). HEAD. Dark, broad, 1.3 times as wide as long (Fig. 79). Clypeus yellow over majority of disc, all marginal areas black with two lateral triangular black marks. Underlying surface weakly shagreened, shining, centrally sparsely punctate, punctures separated by 2 puncture diameters, marginal areas densely punctate, punctures separated by 0.5 – 1 puncture diameter. Apical part of clypeus extended and upturned, process of labrum trapezoidal, apical margin upturned forming rounded ridge, apex emarginate. Gena exceeding width of compound eye, weakly carinate (Fig. 80). Gena, vertex, face, and scape with long brownish-white hairs, not exceeding length of scape. Antennae dark, A 4 – 13 lightened orange below, A 3 exceeding A 4 but shorter than A 4 + 5. MESOSOMA. Scutum and scutellum laterally shagreened and dull, centrally shining, sparsely punctate, punctures separated by 2 – 3 puncture diameters (Fig. 81). Episternum, propodeum, and vestiture as in female. Legs dark, apical tarsal segments lightened orange. Wings as in female. METASOMA. Terga dark, apical margins lightened brown, underlying surface very weakly shagreened, shining (Fig. 82). T 1 almost impunctate, T 2 – 4 with sparse and inconspicuous punctures, separated by 1 – 2 puncture diameters. S 8 short, apically slightly broadened and rounded, basally with short and thick golden bristles that project laterally. Genitalia simple, gonocoxa with faint shagreenation, apical corners rounded, diverging (Fig. 83). Penis valve triangular, strongly narrowed apically. Gonostyli with straight external margin, weakly pointed apically, with slightly raised internal margin.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFB2FFCFFDEEFB7FFBDFF8A3.taxon	discussion	Remarks Using the key to Iberian Notandrena of Wood et al. (2020), females of A. juliana Wood sp. nov. key to couplet seven which separates A. fulvicornis and A. nitidiuscula. It can be separated from A. nitidiuscula by the absence of an impressed longitudinal line on the front half of the scutum, and from A. fulvicornis by the less dense punctures of T 1 and the shinier scutellum. No male key was produced because the male of A. foeniculae is unknown, but males of A. juliana sp. nov. should be identifiable by the combination of their genital structure and yellow clypeus. Additionally, A. foeniculae flies in August and September, and so based on the limited number of specimens collected to date, both taxa are unlikely to fly together at the same time. The similar species A. curvana and A. fulvicornis are bivoltine and fly in both the spring and the summer. Since A. juliana sp. nov. is known only from spring material, further study is needed to clarify its full period of activity.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFB2FFCFFDEEFB7FFBDFF8A3.taxon	distribution	Distribution Southern Spain (Málaga) and southern Portugal (Algarve). Both localities are very close to the coast, adjacent to estuaries, specifically those of the Guadalhorce (San Julián) and Ribeira de São Lourenço (Gambelas). This habitat type should be searched during the spring, as it may contain habitat elements or flowering plants that are important for the ecology of A. juliana Wood sp. nov.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFB6FFF6FE22F88CFB1DFC2B.taxon	description	Figs 84 – 87, 89, 91, 93, 95, 97, 99	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFB6FFF6FE22F88CFB1DFC2B.taxon	materials_examined	Material examined (Andrena impressa) Holotype MOROCCO • ♂; Tangier; OÖLM (illustrated Fig. 86). Paratypes PORTUGAL • 1 ♂, 1 ♀; Cardigos; OÖLM • 1 ♀; Matto do Fundão; OÖLM (illustrated Fig. 85). SPAIN • 1 ♂; Barcelona; O. Schmiedeknecht leg.; OÖLM • 1 ♂; Canet de Mar; 17 May 1963; Vergés leg.; OÖLM. Other material ALGERIA • 1 ♂; El Kseur, F. t. d’Akfadou; 22 May 1981; OÖLM. FRANCE • 1 ♀; Landes, St-Justin, Arouille; 25 May 2013; I. Cross leg.; ICC • 2 ♀♀; Pyr. Or., Banyulsdels-Aspres; 5 – 7 Jun. 1997; H. Wiering and F. Kunst leg.; NMNL • 1 ♀; Pyr. Or., Villelongue; 28 May 1992; H. and J. E. Wiering leg.; TJWC • 1 ♀; Saint-Guilhem-le-Désert; 10 Apr. 1965; OÖLM. MOROCCO • 1 ♀; Tangier; OÖLM • 1 ♀; Azil. Taddert; 1750 m a. s. l.; 10 Mar. 1988; V. Lefeber leg.; NMNL • 1 ♀; Issaguen, 150 km SE of Tanger; 1550 m a. s. l.; 12 May 2015; Mucska leg.; OÖLM. PORTUGAL • 1 ♀; Bensafrim; 5 Mar. 2015; I. Cross leg.; ICC • 1 ♀; Rossas, Touça, N 205 x M 614; 14 May 2019; Wood leg.; TJWC (illustrated Figs 87, 89, 91) • 1 ♀; Vilarinha; 16 Apr. 2017; I. Cross leg.; ICC. SPAIN • 1 ♂; Almeria, Enix; 300 m a. s. l.; 10 May 1978; Diller leg.; OÖLM • 1 ♀; Almeria: Bayarcal; 24 Jun. 1988; M. Schwarz leg.; MSC • 1 ♀; Ávila, Guisando; 750 m a. s. l.; 27 May 1995; H. and J. E. Wiering leg.; TJWC • 1 ♂; Cáceres; 4 Apr. 1921; J. M. Dusmet y Alonso leg.; OÖLM • 1 ♂; Cáceres, Rivera de Gata, W of Villasbuenas; 16 Jun. 1984; W. Schacht leg.; OÖLM • 1 ♂; Canet de Mar; 12 Apr. 1965; Vergés leg.; OÖLM • 1 ♀; Granada, Ventas del Molinello; 20 Jun. 1987; M. Schwarz leg.; TJWC • 1 ♂; Granada, Sierra de Almijara, Pico Lopera; 25 Mar. 2009; I. Cross leg.; ICC • 1 ♀; Málaga, Alcuzcuz, nr San Pedro de Alcantara; 18 Apr. 1983; NMNL • 4 ♀♀; Málaga, Marbella; 14 May 1959; J. v. d. Vecht leg.; NMNL • 1 ♀; Málaga, Sierra Bermeja; 15 May 1959; J. v. d. Vecht leg.; NMNL • 1 ♀; Mallorca, Soller; 1957; N. Briedé leg.; NMNL • 1 ♂, 1 ♀; Mallorca, Tramuntana, Galilea; 22 May 2012; D. W. Baldock leg.; TJWC • 1 ♀; Murcia, Sierra de Españula; 11 May 2003; J. Halada leg.; TJWC • 1 ♀; N of Figueres; 2 May 2003; M. Snižek leg.; OÖLM • 1 ♂; Ronda env.; 26 Feb. 2015; P. Kylies leg.; TJWC (illustrated Figs 93, 95, 97, 99) • 1 ♀; Ronda; 800 m a. s. l.; 23 Mar. 1986; C. v. Achterberg leg.; NMNL • 1 ♀; Sierra Filabres Albanchez; 23 Apr. 2003; J. Halada leg.; OÖLM. Material examined (Andrena angustior) BELGIUM • 1 ♂, 1 ♀; Mons; 29 Apr. – 12 May 2019; W. Fiordaliso leg. • 1 ♀; Tromp, Stropersbos; 6 May 2020; Wood leg.; TJWC. FRANCE • 1 ♀; Bertry; 13 May 2019; A. Cozzani leg. • 1 ♀; Haute Pyr., 5 km N of Col de Pourtalet; 1700 m a. s. l.; 11 Jun. 1983; J. P. Duffels leg.; NMNL • 1 ♂; Landrecies; 13 May 2019; A. Cozzani leg. • 1 ♀; Le Plessis sur Autheuil; 29 Apr. 2019; Wood leg.; TJWC • 1 ♀; Ligny-en-cambrésis; 23 May 2019; C. Pellet leg. • 1 ♀; Pyr. Or., Eyne; 1600 m a. s. l.; 10 Jun. 1997; H. and J. E. Wiering leg.; NMNL • 1 ♀; Quineville, Manche; 1 Jun. 2006; D. W. Baldock leg.; TJWC • 1 ♀; Versigny, Les communaux; 10 May 2018; D. Top leg. GERMANY • 1 ♀; Osterwald; 25 May 1926; J. D. Alfken leg.; ZMHB • 1 ♀; Sababurg; 30 May 1936; J. D. Alfken leg.; ZMHB. PORTUGAL • 2 ♂♂, 2 ♀♀; Branca; 11 Mar. – 2 Apr. 2019; H. Gaspar leg. • 1 ♀; Confurco, Várzea Cova; 14 May 2019; Wood leg.; TJWC • 1 ♀; 1 km south of Paradela, M 308 – 4; 12 May 2019; Wood leg.; TJWC • 1 ♀; Pedraído, Fafe; 14 May 2019; Wood leg.; TJWC (illustrated Figs 88, 90, 92) • 1 ♀; Serra do Gerês, 5 km W of Paradela; 12 May 2019; Wood leg.; TJWC. SPAIN • 1 ♀; Ávila, Hoyos del Espino; 1400 m a. s. l.; 20 May 1995; H. and J. E. Wiering leg.; NMNL • 3 ♂♂, 4 ♀♀; Ávila, Sierra de Gredos, La Plataforma; 1700 m a. s. l.; 19 May 1995; H. and J. E. Wiering leg.; NMNL • 1 ♀; Ávila, Sierra de Gredos Puerto del Pico; 2 Jul. 1988; M. Schwarz leg.; MSC • 1 ♀; Benasque; 15 Jun. 1983; J. P. Duffels leg.; NMNL • 1 ♂, 1 ♀; Burgos, Hornilloyuso; 24 Jun. 1984; R. Leys leg.; NMNL • 1 ♂; Burguete; 4 Jun. 1987; E. A. M. Speijer leg.; NMNL • 2 ♀♀; Cáceres, Cuacos de Yuste; 500 m a. s. l.; 3 May 1996; H. and J. E. Wiering leg.; NMNL • 3 ♀♀; Cáceres, Piornal; 1050 m a. s. l.; 13 May 1999; H. and J. E. Wiering leg.; NMNL • 1 ♀; Cantabria, Argüébanes; 6 Jun. 2019; I. Cross leg.; ICC • 1 ♀; Cantabria, Picos de Europa, Camaleno; 600 m a. s. l.; 14 Jun. 2013; D. W. Baldock leg.; TJWC • 1 ♀; Cantabria, Picos de Europa, Camaleno; 600 m a. s. l.; 5 May 2014; D. W. Baldock leg.; TJWC • 1 ♀; Huesca, El Portalet; 1800 m a. s. l.; 2 Jun. 1995; H. and J. E. Wiering leg.; NMNL • 1 ♀; Oviedo, Cudillero; 26 May 2019; W. Klein leg. • 6 ♂♂; Oviedo, picos de Europa, Lago de la Ercina; 19 Apr. 1984; R. Leys leg.; NMNL • 1 ♂; Santander, Barcenaciones; 17 Apr. 1984; R. Leys leg.; NMNL • 2 ♀♀; Santander, Golbardo; 20 Apr. 1984; R. Leys leg.; NMNL • 1 ♂; Soria, Puerto de Santa Ines; 1 May 1999; H. and J. E. Wiering leg.; NMNL (illustrated Figs 94, 96, 98, 100). UNITED KINGDOM • 1 ♀; Angmering, Hammerpot; 26 May 2015; Wood leg.; TJWC • 1 ♀; Farnham Heath RSPB; 24 May 2016; Wood leg.; TJWC • 2 ♀♀; Goudhurst; 21 Apr. 2019; L. Hutchinson leg. • 1 ♂, 1 ♀; Petworth; 2 Jun. 2014; Wood leg.; TJWC • 1 ♂; Devon, Dartmoor; 6 May 1935; T. F. Perkins leg.; ZMHB. Material examined (Andrena fulvata) AUSTRIA • 1 ♀; Fürberg, Wolfgangsee; 16 Apr. 1946; ZMHB. BELGIUM • 1 ♂; Estinnes-Au-Mont, Les Trieux; 24 Mar. 2019; J. Dewaele leg. BOSNIA AND HERZEGOVINA • 1 ♂, 4 ♀♀; Gradiska, Turjak; 7 – 9 Apr. 2019; Mitrovic and Golubovic leg.; ULB. BULGARIA • 1 ♂; Šípka mont [Shipka]; 12 May 1994; K. Deneš leg.; OÖLM. CROATIA • 1 ♂; Zagreb; 22 Apr. 1897; ZMHB. FRANCE • 3 ♂♂, 1 ♀; Belval-Bois-des-Dames, Domaine de Belval; 22 Mar. – 21 Apr. 2019; C. Amy leg. • 1 ♀; Desvres; 15 May 2018; R. Vandeweghe leg. • 1 ♀; Eclusier-Vaux, Marais de Vaux; 5 Mar. – 1 Apr. 2019; D. Adam leg. • 11 ♀♀; Elincourt-Sainte-Marguerite; 18 May – 7 Jun. 2018; B. Piallat and C. Bocaux leg. • 1 ♀; Fresnoy-la-Rivière, Les petits Monts; 6 Apr. 2018; D. Top leg. • 1 ♀; Raismes, forêt de Saint-Amand; 13 Jun. 2005; J. - L. Vago leg. • 1 ♀; Xouaxange; 20 May 2019; C. Brelot and C. Filet leg. • 1 ♂; Ruy; 17 Mar. 2020; C. Triquet leg. GERMANY • 3 ♂♂, 2 ♀♀; Kaiserstuhl, Achkarren; 29 Mar. – 20 Apr. 1937; S. G. Bischoff leg.; ZMHB • 1 ♀; Kaiserstuhl, Büchsenberg; 1 Apr. 1937; S. G. Bischoff leg.; ZMHB • 2 ♂♂, 1 ♀; Kaiserstuhl, Liliental; 19 – 24 Apr. 1937; S. G. Bischoff leg.; ZMHB • 1 ♀; Kaiserstuhl, Vogtsburg; 20 – 21 May 1933; S. G. Bischoff leg.; ZMHB • 5 ♂♂; Kehl am Rh. [Rhine]; 1 – 5 Apr. 1937; L. Battes leg.; ZMHB. ITALY • 1 ♂, 1 ♀; Bologna, Parco di Villa Ghigi; 9 – 16 Apr. 2017; G. Ghisbain leg.; TJWC • 1 ♂, 1 ♀; Lazio, Acquapendente; 26 May 1991; J. Gusenleitner leg.; TJWC. SLOVENIA • 1 ♀; Styr, Podčetrtek; 28 Apr. 1932; Jaeger leg.; ZMHB • 1 ♀; Orechek [Orehek]; 28 May 2005; Egger leg.; OÖLM. SWITZERLAND • 1 ♀; Weiach, Kiesgrube Rüteren; 10 Jun. 2014; A. Müller leg.; ETHZ • 1 ♂; Savièse; 14 May 2013; S. Gerber leg.; ETHZ • 1 ♀; Haldenstein; 13 May 2005; M. Hermann leg.; ETHZ • 1 ♀; Bern, Kirchenfeld; 2 May 1920; T. Steck-Hofmann leg.; NMB • 1 ♂; Seedorf, Reussdelta, Auenwald; 20 Apr. 1999; L. Reser-Rezbanyai leg.; NMLU • 1 ♂; Meride, S. Antonio; 18 Apr. 1998; L. Reser- Rezbanyai leg.; NMLU. Material not directly examined (Andrena fulvata) AUSTRIA • 1 ♀; Oberösterreich, Kremsmünster; 6 May 1972; J. Gusenleitner leg.; Zobodat • 1 ♀; Oberösterreich, Losenstein a. d. Enns; 25 Mar. 1979; K. Kremslehner leg.; Zobodat • 1 ♂; Oberösterreich, Magdalenaberg SE of Pettenbach; 16 May 1979; J. Gusenleitner leg.; Zobodat. NETHERLANDS • 1 ♂; Limburg, Eys - Bronnen en Bronbos; 14 Apr. 2018; I. Raemakers leg.; Waarneming. nl • 1 ♀; Limburg, Savelsbos; 2 Apr. 2017; I. Raemakers leg.; Waarneming. nl. ROMANIA • Băile Herculane • Bocsa Montana • Mehadia • Timisoara, all B. Tomozii pers. comm.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFB6FFF6FE22F88CFB1DFC2B.taxon	discussion	Remarks Though placed in the subgenus Ptilandrena by Warncke (1967), the use of this subgenus should be restricted to the Nearctic (Pisanty et al. 2021). Therefore, A. impressa stat. nov. and related species are found within the Euandrena (see also Praz et al. 2019). Andrena angustior was originally described from England, and can rapidly be identified in the female sex by the combination of a central, longitudinal impression on the clypeus with the wide, depressed and shiny margins of the terga. In the male, in addition to the tergal sculpturing, the gena is enlarged and the mandibles are elongated in a manner similar to Andrena subgenus Andrena but without the presence of basal mandibular teeth. Andrena fulvata Stoeckhert, 1930 was later described from southern Germany. The two species are similar, but A. fulvata lacks the shiny depressed tergal margins, these are instead only weakly depressed, shagreened, and matt. Warncke considered A. fulvata to be a subspecies of A. angustior, and described A. a. impressa from Morocco, Portugal, and Spain, arguing that the variation across this geographic gradient merited a broad species concept (Warncke 1967). This form can easily be separated from A. angustior s. str. in the female sex by the structure of the terga which are weakly depressed, shagreened, only weakly shining, and also by the colouration of the hairs flanking the pygidial plate which are black (compare Figs 91 – 92), and the bee has an overall darker appearance (compare Figs 87 – 90). Males can be separated by the same differences in tergal structure (Figs 97 – 98) and overall darker pubescence, particularly on the head (Figs 93 – 96). The most obvious difference can be seen in the antennal ratios. In A. impressa, A 4 is only slightly shorter than A 3 (Fig. 99), whereas in A. angustior A 4 is less than half the length of A 3 (Fig. 100). Subsequent authors have not followed Warncke’s viewpoint, treating A. angustior and A. fulvata as good species (see Gusenleitner & Schwarz 2002; Nieto et al. 2014). These two taxa show different climatic affinities. For example, in Germany, A. angustior is present in the north, reaching as far south as Rhineland-Palatinate but not extending into Baden-Württemberg (Westrich 1989). To the east, the more northwestern A. angustior is completely replaced by the more continental A. fulvata. However, to date, the status of A. a. impressa has not been assessed, with the default position of Warncke being that A. angustior s. str. was absent from Iberia (Gusenleitner & Schwarz 2002; Ortiz-Sánchez 2011). Specimens collected from northern Portugal showed that both A. angustior and A. a. impressa were present in sympatry, with the two specimens barcoded here collected just 6 km apart. Molecular data strongly supports the species status of Andrena impressa stat. nov., as the Portuguese A. angustior sequence closely matched sequences from northern European populations, and the phylogeny suggests that A. impressa is more strongly differentiated than A. angustior and A. fulvata are from each other (Fig. 2). Though the molecular phylogeny suggests that A. impressa is more distantly related to A. angustior + A. fulvata than A. allosa + A. amieti, support for this placement is weak, and based on the morphology of the male head we consider A. angustior + A. fulvata + A. impressa to form a trio of related but distinct species. The diet of A. angustior in Britain was documented by Wood & Roberts (2017). Five additional analysed pollen loads from northwestern Iberia contained Caryophyllaceae Juss. (48.8 %, Arenaria L.), Ericaceae Juss. (15.8 %, Erica L.), Asparagaceae Juss. (10.5 %, Scilla L.), Crassulaceae J. St. Hil. (8.4 %, Sedum L.), Cistaceae Juss. (6.3 %, Cistus L.), Brassicaceae Burnett (5.7 %, Raphanus - type), Rosaceae Juss. (3.2 %, Potentilla L.), and Ranunculaceae Juss. (1.3 %, Ranunculus L.). The western distribution, the latespring flight period at the end of April into May, and these dietary preferences support the position that A. angustior is a species of Atlantic woodland edges, utilising spring flowering herbs and shrubs but interestingly not the trees themselves. Fewer data are available for Andrena impressa; the two analysed pollen loads contained Cistaceae (50.3 %, Cistus), Scrophulariaceae Juss. (42.9 %, Scrophularia L.), and Asteraceae Bercht. & J. Presl (6.9 %, Anthemis - type). Given its distribution, a diet more focused on Mediterranean herbs and shrubs is expected, but more study is required.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FFB6FFF6FE22F88CFB1DFC2B.taxon	distribution	Distribution The distribution map of Warncke (see Gusenleitner & Schwarz 2002) is broadly correct in the northern distribution of A. angustior, the continental distribution of A. fulvata, and the West Mediterranean distribution of A. impressa (France, Spain, Portugal, Algeria, Morocco), but incorrect in the omission of true A. angustior from cooler parts of Iberia (Fig. 84). This distribution indicates that A. angustior is not simply a northern species but rather one with an Atlantic affinity. Though we did not examine material from this region as part of this study, A. angustior is likely to be present in cooler parts of western France, joining the two distributions together. Warncke’s map also does not capture the sympatric presence of A. angustior and A. fulvata populations in northern Europe as this is a more recent and ongoing phenomenon, with A. fulvata a recent colonist of Belgium for example (compare Patiny & Terzo 2010; Drossart et al. 2019). The lack of recognition of true A. angustior from the mountains of Iberia is strange, as Warncke (1967) made reference to males from the Pyrenees with shiny tergal margins, but he seemed to consider these intermediate between A. angustior and A. impressa stat. nov ..	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FF8FFFF8FE17FBFCFEA2FA5E.taxon	description	Figs 101 – 108	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FF8FFFF8FE17FBFCFEA2FA5E.taxon	materials_examined	Material examined Holotype MOROCCO • 1 ♂; Marraquesh; Mar. 1907; Escalera leg.; OÖLM (illustrated Fig. 86). Paratype MOROCCO • 1 ♀; Marraquesh; Mar. 1907; Escalera leg.; OÖLM. Other material MOROCCO • 1 ♀; Massa-Tiznit; 26 Mar. 1988; V. Lefeber leg.; NMNL • 2 ♂♂, 2 ♀♀; Tamri, 70 km N of Agadir; 8 May 1995; Ma. Halada leg.; MSC • 1 ♂, 2 ♀♀; same collection data as for preceding; OÖLM (illustrated Figs 106, 108) • 1 ♀; 20 km W of Boudnib; 9 Apr. 1995; Ma. Halada leg.; OÖLM (illustrated Figs 102, 104). SPAIN • 1 ♂, 1 ♀; Sevilla, Los Pinares de Aznalcazar; 13 Mar. 2012; I. Cross leg.; ICC (illustrated Figs 105, 107) • 2 ♀♀; Huelva, Hinojos; 15 Mar. 2012; I. Cross leg.; Reseda phyteuma; ICC • 1 ♀; same collection data as for preceding; TJWC (illustrated Figs 101, 103).	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FF8FFFF8FE17FBFCFEA2FA5E.taxon	discussion	Remarks Material from southwestern Spain closely resembles Moroccan specimens, but are generally less hirsute, particularly in the single male Spanish specimen, although this may be simply due to the condition of the individual specimen (Figs 101 – 108). Structurally, there are no clear differences, and so they are considered to be conspecific. The pronouncedly Atlantic distribution of Morocco and southwestern Iberia is not unprecedented in bees, as the extremely rare Lasioglossum musculoides Ebmer, 1974 has a global distribution of the Souss Valley in southwestern Morocco and southwestern Portugal (Pauly 2015). Coastal areas of Huelva and Sevilla continue to produce records of range-restricted species thought to be endemic elsewhere, such as Flavipanurgus fuzetus Patiny, 1999 which was discovered there in 2018 (Cross & Wood 2018). Pollen loads taken from three specimens at a single Moroccan locality contained pure Reseda - type (Resedaceae Martinov) pollen (Table 2), so the floral observation data from Spain is consistent with this picture. Comparisons with other species are complicated because the former subgenus Poliandrena Warncke, 1968 in which it was placed (Warncke 1974) has been found to be strongly polyphyletic, and Poliandrena in a strict sense is now a synonym of the subgenus Ulandrena Warncke, 1968 (Pisanty et al. 2021). Excluding A. polita Smith, 1847 (now subgenus Ulandrena) and A. florea Fabricius, 1793 (the placement of this species necessitates the erection of a new subgenus) which clearly belong elsewhere (Pisanty et al. 2021), other members of the old concept of Poliandrena that can be found in Spain for which pollen data are available are predominantly associated with Reseda L., with the exception of A. farinosa Pérez, 1895 which is likely a specialist of Fabaceae (Table 2). These taxa fall into at least two clades (Pisanty et al. 2021), and so work is needed to establish their true subgeneric affinities in addition to their dietary choices.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FF8FFFF8FE17FBFCFEA2FA5E.taxon	distribution	Distribution Previously thought to be endemic to Morocco, now also including Spain (Warncke 1974; Lhomme et al. 2020).	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FF81FFF9FE15FA12FB90FDAF.taxon	materials_examined	Material examined SPAIN • 1 ♀, Andrena confinis; Cantabria, Picos de Europa, Camaleno; 600 m a. s. l.; 5 May 2014; E. Scheuchl det.; D. W. Baldock leg.; TJWC • 1 ♀, Andrena congruens; Pirineos Orient, 20 km NE of Ripoll; 1650 m a. s. l.; 31 Jul. 2011; J. Halada leg.; TJWC • 1 ♀, Andrena congruens; Pyrenees, Cavalers (exact location unclear); 11 Jul. 2002; H. Kohout leg.; OÖLM.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FF81FFF9FE15FA12FB90FDAF.taxon	discussion	Remarks This species pair is in need of molecular revision across Europe because of historical synonymy with A. congruens Schmiedeknecht, 1884 (e. g., Warncke 1967). Both species are present in central Europe (Schmid-Egger & Scheuchl 1997; Le Divelec 2020), and material examined from northern Europe (Belgium, northern France, United Kingdom) conforms to A. confinis, suggesting that it is the more northerly of the two. Currently, only A. congruens is listed for Spain (Ortiz-Sánchez 2011), but both taxa are present in mountainous parts of northern Spain, though given the limited material available it is difficult to come to firm conclusions concerning their true distributions.	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
F12A87E8FF81FFF9FE15FA12FB90FDAF.taxon	distribution	Distribution The distribution of A. confinis is unclear because of continued confusion and synonymy with Andrena congruens Schmiedeknecht, 1884. At this moment, it is not possible to give an accurate picture of its true distribution, but both species are present in central Europe, with A. congruens having more southern and A. confinis more northern tendencies (Schmid-Egger & Scheuchl 1997).	en	Wood, Thomas J., Ghisbain, Guillaume, Michez, Denis, Praz, Christophe J. (2021): Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy 758: 147-193, DOI: https://doi.org/10.5852/ejt.2021.758.1431, URL: http://dx.doi.org/10.5852/ejt.2021.758.1431
