taxonID	type	description	language	source
03F387B2FF895346CD2451F4FAF3A101.taxon	diagnosis	Diagnosis. See Klimaszewski & Pelletier (2002).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF895346CD2451F4FAF3A101.taxon	type_taxon	Type species. Tinotus cavicollis Sharp, 1883, by subsequent designation (Fenyes, 1918).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF895346CD2451F4FAF3A101.taxon	discussion	Discussion. The genus Tinotus Sharp, 1883 currently includes about 30 valid species (Klimaszewski & Pelletier 2002; Hanley 2002). Recently Hanley (2002) published a world catalogue of the genus. Klimaszewski and Pelletier (2002) revised the types of Nearctic species of Tinotus, synonymized some species names and provided a key for identification of Tinotus of America north of Mexico. Casey (1911 a) described the genus Acrimea, including three species described in the same paper. He placed Acrimea in the tribe Aleocharini and considered it related to Thiasophila Kraatz, 1856 and Ischnoglossa Kraatz, 1856. Subsequently Acrimea was considered to be a member of the tribe Oxypodini (Fenyes 1918, 1920; Bernhauer & Scheerpeltz 1926; Seevers 1978; Newton et al. 2000). My revision of the types of the three species of Acrimea revealed that in all three the tarsal formula is in fact 4 ­ 5 ­ 5 and all three belong to the genus Tinotus. The type species of Acrimea, Acri. resecta, is a synonym of Ti. acerbus (see below). Therefore, Acrimea is a junior synonym of Tinotus. Because Acrimea was erroneously considered to have the tarsal formula 5 ­ 5 ­ 5, the genus was incorrectly listed in key B to the Nearctic genera of aleocharines (Newton et al. 2000: 303). Couplet 19 should be deleted from key B, and couplet 18 should be modified as follows: 18 (16). Mesosternum carinate medially at base .............................................................. 20 ­ Mesosternum not at all carinate medially ............................................................ 21	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF8A5347CD2450CEFB88A409.taxon	description	(Figs. 2, 7, 23, 28 – 29 in Klimaszewski & Pelletier 2002)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF8A5347CD2450CEFB88A409.taxon	materials_examined	Type material. Holotype of Tinotus trisectus: UNITED STATES: Idaho: Kootenai Co.: ♀, Coeur d'Alene, vi (H. F. Wickham) (NMNH). Lectotype of Acrimea fimbriata (here designated): ♂, “ The Dalles, Oreg. [on (Wickham)] ”, “ fimbriata Csy. ”, “ TYPE USNM 39738 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype (♀; designated by Hanley (2002 )) and paralectotype (♂) of Tinotus pallidus: UNITED STATES: Iowa: Linn Co.: Cedar Rapids (NMNH). Lectotype of Tinotus brunneus (designated by Hanley (2002 )): UNITED STATES: Pennsylvania: ♂, near Philadelphia (NMNH). Holotype of Tinotus binarius: UNITED STATES: Arizona: Pima Co.: ♀, Tucson (NMNH). Additional material. UNITED STATES: California: Kern Co.: ♂, 12 mi. S Bakersfield, 20. v. 1958 (E. I. Schlinger) (CNCI); Tennessee: Lake Co.: ♂, Reelfoot Lk., 7 km NE Tiptonville, at black light, 8. v. 1986 (J. M. Campbell) (CNCI).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF8A5347CD2450CEFB88A409.taxon	diagnosis	Diagnosis. See Klimaszewski & Pelletier (2002).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF8A5347CD2450CEFB88A409.taxon	discussion	Discussion. The lectotype of Acrimea fimbriata is similar to the holotype of Ti. trisectus in external characters. The shape of the aedeagus, including the structure of the internal sac, is identical in the lectotype of Acri. fimbriata and in examined males of Ti. trisectus. Klimaszewski et al. (2002) placed Ti. pallidus in synonymy with Ti. caviceps. Unfortunately, the male paralectotype of Ti. pallidus lacks the head and pronotum which does not allow the comparison of these parts of the body with the males of Ti. caviceps which have strong medial impressions of the head disk and pronotum. However, both types of Ti. pallidus are more glossy than the types of Ti. caviceps and resemble Ti. trisectus in that respect. The aedeagus (including the structures of the internal sac) of the paralectotype of Ti. pallidus is similar to the aedeagus of Ti. trisectus, and not Ti. caviceps. This fact is clear when Figures 21, 23 and 17 – 20 in Klimaszewski & Pelletier (2002) are compared. Therefore, Ti. pallidus is removed from synonymy with Ti. caviceps and placed in synonymy with Ti. trisectus Casey, 1906. Hanley (2002) designated the lectotype of Ti. trisectus. However, when Casey (1906) described Ti. trisectus he indicated that he had “ the single type ” of this species. Therefore this type is the holotype, and the lectotype designation by Hanley (2002) is redundant. Hanley (2002) also designated the lectotype of Ti. binarius. When Casey (1911 a) described Ti. binarius he indicated that he had “ only one example ” of this species. Therefore this type is the holotype, and the lectotype designation by Hanley (2002) is redundant.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF8A5347CD2450CEFB88A409.taxon	distribution	Distribution. Ti. trisectus is widespread in the United States. It is known from Oregon, California, Arizona, Idaho, Iowa, Pennsylvania and Tennessee.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF8B5342CD2455C6FE5BA361.taxon	description	(Figs. 1 – 9)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF8B5342CD2455C6FE5BA361.taxon	materials_examined	Type material. Lectotype of Acrimea acerba (here designated): ♂, “ N. [orth] Yakima, Wash. [ington state (Wickham)] ”, “ acerba Csy. ”, “ TYPE USNM 39737 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Acrimea resecta (here designated): ♀, “ Coeur d'Alene, IDAHO [Wickham] ”, “ Acrimea resecta Csy. ”, “ TYPE USNM 39736 ” (red label), “ CASEY bequest 1925 ” (NMNH). Additional material. CANADA: British Columbia: ♂, ♀, 4 mi. W Midway, ex river debris, 6. vi. 1968 (Campbell & Smetana) (CNCI, SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF8B5342CD2455C6FE5BA361.taxon	diagnosis	Diagnosis. Tinotus acerbus differs from the other Nearctic species of Tinotus in its brown body, the weak and poorly visible microsculpture of its glossy pronotum, the strong asperate punctation of its elytra, its lack of a medial impression on the male head and pronotum, its unmodified male sternum 8, and the shape of its aedeagus (Figs. 5 – 8), in particular the straight apex of its median lobe (in lateral view (Fig. 7 )). Additionally, Ti. acerbus differs from Ti. morion in its brown body color; from Ti. caviceps in its lack of a medial impression on the male head and pronotum; from Ti. imbricatus in its glossier pronotum, its lack of a medial impression on the male pronotum, and its unmodified male sternum 8; from Ti. parvicornis in its glossier pronotum.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF8B5342CD2455C6FE5BA361.taxon	discussion	Discussion. The lectotypes of Acri. acerba and Acri. resecta are similar in all external characters. These lectotypes are identical in the shape of their genitalia to the male and female specimens listed above in Material and found in the same locality in British Columbia. Tinotus acerbus is a valid species which was not included in the review of Nearctic Tinotus by Klimaszewski & Pelletier (2002).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF8B5342CD2455C6FE5BA361.taxon	distribution	Distribution. Tinotus acerbus is known from Washington, Idaho and British Columbia.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF8E5342CD245154FE55A566.taxon	discussion	Discussion. Casey (1910 a) did not specifically state in his description of Ancillota that Anc. sollemnis was the type species of Ancillota; however, while designating the type species of Noverota Casey, 1910 a in the same paper, Casey stated (1910 a, p. 90): “ The first species may be regarded as the type, as in all other cases where the type is not specifically named ”. The type species of Ancillota, Anc. sollemnis, has a tarsal formula 5 ­ 5 ­ 5 and is synonymous with Oxypoda amica Casey, 1906 (see below). Therefore, Ancillota is placed in synonymy with Oxypoda. Casey (1911 a) described the monotypic genus Moluciba in the subtribe Athetina. My examination of the holotype of Mol. grandipennis revealed that the tarsal formula in the type is 5 ­ 5 ­ 5, and the species has all characters of the nominotypical subgenus of the genus Oxypoda. When describing Moluciba, Casey (1911 a) compared it with Acrotona and related athetine genera, and failed to realize that Mol. grandipennis in fact belonged to Oxypodini.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF8F535CCD24524EFC84A1E1.taxon	description	(Figs. 10 – 14)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF8F535CCD24524EFC84A1E1.taxon	materials_examined	Type material. Holotype of Moluciba grandipennis: ♀, “ Metlakatla B. [ritish] Col. [umbia] Keen ”, “ Moluciba grandipennis Csy. ”, “ TYPE USNM 39020 ” (red label), “ CASEY bequest 1925 ” (NMNH). Additional material. CANADA: Québec: ♀, Chic­Chocs, 17. viii. 1994; ♂, 50 km E Québec­City, Armagh, 46 º 44 ’ N 70 º 35 ’ W, 2. viii. 1994 (SPSU); 2 specimens, 170 km NE Québec­City, Pohénégamook, 47 º 28 ’ N 69 º 14 ’ W, 16. viii. 1994 (LFC, SPSU); 3 specimens, ditto but 2. viii. 1994 (LFC); 1 specimen, Lac Mitis, 48 º 18 ’ N 67 º 48 ’ W, 22. viii. 1994 (SPSU); 1 specimen, ditto but 15. viii. 1994 (LFC); UNITED STATES: Alaska: ♀, Kenai Peninsula, 10 km S Ninilchik, 59 º 57.26 –. 25 ’ N 151 º 43.81 ’ W, 50 m, forest litter, Picea, 22. vii. 1998 (V. I. Gusarov); ♂, Richardson Hwy., NNW Willow Creek, 61 º 50.17 ’ N 145 º 13.81 ’ W, 500 m, in human feces, 13. vii. 1998 (V. I. Gusarov) (SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF8F535CCD24524EFC84A1E1.taxon	diagnosis	Diagnosis. Oxypoda grandipennis can be distinguished from other Nearctic members of the nominotypical subgenus by a dark brown body with the brownish yellow pronotum, humeral angles and posterior margin of the elytra, abdominal terga 3 – 5 and posterior margin of tergum 7; and the distinct shape of the aedeagus (Figs. 10 – 13) and spermatheca (Fig. 14).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF8F535CCD24524EFC84A1E1.taxon	distribution	Distribution. Oxypoda grandipennis is a boreal species with a transnearctic distribution. It is known from Alaska and Québec.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF90535CCD24576EFD83A539.taxon	description	(Fig. 2 in Strand & Vik 1966 (as O. lividipennis ))	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF90535CCD24576EFD83A539.taxon	materials_examined	Examined material. FRANCE: Le Loiret: ♂, ♀, 15 km de Montargis, domaine des Barres, 13 – 20. x. 1998 (A. Gurov) (SPSU); UNITED STATES: New York: Orange Co.: ♂, ♀, 8 km S New Windsor, Black Rock Forest, 9. v. 1998 (V. I. Gusarov) (SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF90535CCD24576EFD83A539.taxon	diagnosis	Diagnosis. See Lohse (1974; as O. lividipennis).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF90535CCD24576EFD83A539.taxon	discussion	Discussion. This is a first North American record of this species (cf. Muona 1984).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF90535CCD24576EFD83A539.taxon	distribution	Distribution. Oxypoda acuminata is widespread in Europe. In North America it is known only from New York.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF91535FCD24524EFDF9A639.taxon	description	(Figs. 15 – 18)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF91535FCD24524EFDF9A639.taxon	materials_examined	Type material. Lectotype of Oxypoda amica (here designated): ♂ (mounted with a Formica specimen), “ Iowa City Wickham ”, “ [ant] Nest IV ”, “ amica Csy. ”, “ TYPE USNM 39786 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Ancillota sollemnis (here designated): ♂, “ Mo. [Missouri, St. Louis] ”, “ Ancillota sollemnis Csy. ”, “ TYPE USNM 39018 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Oxypoda vetula (here designated): ♀, “ N. [ew] J. [ersey] ”, “ vetula ­ 2 PARATYPE USNM 39835 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: ♀, “ N. [ew] J. [ersey] ”, “ vetula Csy. ”, “ TYPE USNM 39835 ” (red label), “ CASEY bequest 1925 ”; ♀, “ N. [ew] J. [ersey] ”, “ vetula ­ 3 PARATYPE USNM 39835 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Oxypoda neptis (here designated): ♂, “ Iowa City, Ia [Iowa] Wickham ”, “ neptis Csy. ”, “ TYPE USNM 39844 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Oxypoda schaefferi (here designated): ♂, “ Windsor, Broome Co., N. [ew] Y. [ork] H. Notman 29 May 1918 ”, “ TYPE ” (pink label), “ Oxypoda schaefferi ” (SIIS). Paralectotype: ♀, “ Windsor, Broome Co., N. [ew] Y. [ork] H. Notman 29 May 1918 ”, “ PARATYPE ” (pink label), “ schaefferi paratype ” (SIIS). Additional material. UNITED STATES: Kansas: Douglas Co.: 8 specimens, 1.5 km N Lawrence, right bank of the Kansas River, 38 º 58.96 ’ N 95 º 14.62 ’ W, 200 m, in pile of rotting wood chips, 1. v. 1999 (V. I. Gusarov) (KSEM, SPSU); 8 specimens, 2.5 km N Lawrence, flood refuse at the Kansas River, 38 º 59.59 ’ N 95 º 14.59 ’ W, 200 m, 18. iv. 1999 (V. I. Gusarov); Jefferson Co.: 3 ♂♂, ♀, 15 km SSW Oskaloosa, Perry State Park – Delaware Area, 39 º 09.36 ’ N 98 º 28.40 ’ W, 300 m, sifting forest litter, 16. i. 1999 (V. I. Gusarov) (SPSU); Kentucky: Bell Co.: 5 specimens, 12 km NWW Middlesboro, Hwy. 74 & 3485, 36 º 37.8 ’ N 83 º 51.9 ’ W, 800 m, forest litter, Quercus, Acer, 16. iii. 2001 (V. I. Gusarov) (SPSU); Louisiana: Natchitoches Par.: ♂, 1.5 mi. N Readhimer, 32 º 08 ’ N 92 º 59 ’ W, Geomys breviceps burrow, baited (gopher feces and malt extract) pitfall, 16. i – 4. ii. 2000 (P. Kovarik, A. Tishechkin & R. Turnbow) (SPSU); North Carolina: Yancey Co.: ♀, 21 km S Burnsville, Hwy. 128, 35 º 43.42 ’ N 82 º 16.97 ’ W, 1600 m, forest litter, Picea, Abies, Betula, Acer, Majanthemum, 25. vi. 2001 (V. I. Gusarov) (SPSU); North Carolina / Tennessee: Mitchell Co. / Carter Co.: ♀, Roan Road Pass, 29 km SSE Elizabethton, Hwy. 143 / 261, Cherokee / Pisgah Nat. Forest, 36 º 06.3 ’ N 82 º 06.8 ’ W, 1700 m, forest litter, Picea, Rhododendron, 17. iii. 2001 (V. I. Gusarov) (SPSU); Tennessee: Campbell Co.: 8 specimens, 19.5 km N La Follette, Pine Mountain, 36 º 33.5 ’ N 84 º 07.7 ’ W, 600 m, forest litter, Quercus, 16. iii. 2001 (V. I. Gusarov); Carter Co.: ♂, 29 km SSE Elizabethton, Hwy. 143, Cherokee Nat. Forest, 36 º 06.8 ’ N 82 º 06.1 ’ W, 1600 m, forest litter, Betula, Acer, 17. iii. 2001 (V. I. Gusarov); Fentress Co.: 1 specimen, 20 km NEE Jamestown, Hwy. 297, 36 º 28.39 ’ N 84 º 43.05 ’ W, 500 m, forest litter, Quercus, 16. iii. 2001 (V. I. Gusarov); Scott Co.: 6 specimens, 16 km W Oneida, Hwy. 297, Bandy Creek, 36 º 28.5 ’ N 84 º 41.3 ’ W, 400 m, forest litter, Quercus, Pinus, Rhododendron, 16. iii. 2001 (V. I. Gusarov); Sevier Co.: ♂, ♀, 7 km SW Gatlinburg, Little River Rd., Malonei Point, 35 º 40.40 ’ N 83 º 34.47 ’ W, 700 m, on flowers of Aruncus dioicus, 1. vi. 2001 (V. I. Gusarov) (SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF91535FCD24524EFDF9A639.taxon	diagnosis	Diagnosis. Oxypoda amica belongs to a group of Nearctic species with the copulatory piece of the internal sac extended in a long flagellum. Externally the species of this group are very similar, and O. amica can be recognized among them only by the distinct shape of the aedeagus (Figs. 15 – 17) and spermatheca (Fig. 18).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF91535FCD24524EFDF9A639.taxon	discussion	Discussion. The types of O. amica, Anc. sollemnis, O. vetula, O. neptis and O. schaefferi are similar in external characters and in the shape of the aedeagus (including the sclerites of the internal sac) and spermatheca.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF91535FCD24524EFDF9A639.taxon	distribution	Distribution. Oxypoda amica is widespread in the eastern United States. It is known from Iowa, Kansas, Missouri, Kentucky, New York, New Jersey, Tennessee, North Carolina and Louisiana.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF93535ACD24568EFCA0A6C9.taxon	description	(Figs. 22 – 23 in Lohse & Smetana 1985 (as De. prospera and De. congruens ))	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF93535ACD24568EFCA0A6C9.taxon	materials_examined	Type material. Lectotype of Oxypoda prospera (here designated): “ Germ. bor. Schüppel ”, “ 5514 ”, “ prospera Er. Germ. bor. Schüpp ” (ZMHB). Lectotype of Oxypoda congruens (here designated): ♀, “ Helena Mon. [tana] ”, “ Oxyp. [oda] congruens ”, “ TYPE USNM 39766 ” (red label), “ CASEY bequest 1925 ”, “ CASEY determ. prospera ­ 19 ” (NMNH). Paralectotypes: ♂, “ Mic. [higan] ”, “ congruens PARATYPE USNM 39766 ” (red label), “ CASEY bequest 1925 ”, “ CASEY determ. prospera ­ 14 ”; 2 ♀♀, “ Mich. [igan] ”, “ congruens PARATYPE USNM 39766 ” (red label), “ CASEY bequest 1925 ”, “ CASEY determ. prospera ”; ♀, “: Mon [tana, Helena (Wickham)] ”, “ congruens PARATYPE USNM 39766 ” (red label), “ CASEY bequest 1925 ”, “ CASEY determ. prospera ­ 16 ” (NMNH). Additional material. CANADA: Alberta: 18 specimens, Athabasca Riv. at Rocky Riv., 13. viii. 1965 (J. & W. Ivie); 1 specimen, Little Smokey River, 6 mi. S Guy, 4. viii. 1965 (J. & W. Ivie) (AMNH); British Columbia: 1 specimen, 9 mi. NW Wonowon, 56 º 50 ’ N 121 º 57 ’ W, 3. viii. 1968 (W. Ivie) (AMNH); Labrador: 2 specimens, Carter Basin, willow humus, 7. ix. 1958; 1 specimen, ditto but 4. viii. 1958; 1 specimen, ditto but moss under willow, 7. ix. 1958; 2 specimens, ditto but moss on willow, 17. viii. 1958 (BMNH); Manitoba: ♀, Aweme (N. Criddle) (NMNH (Casey collection )); 2 specimens, Riding Mt. Nat. Park, Whirlpool River at Rt. 19, 23. vi. 1975 (L. & N. Herman) (AMNH); Ontario: 2 specimens, Ottawa (NMNH (Casey collection )); ♂, Nepigon, 6. vi. 1918 (A. Fenyes) (NMNH (Casey collection )); CZECK REPUBLIC: 2 specimens, Moravia (Reitter) (NMNH (Casey collection )); RUSSIA: Magadan Reg.: 7 specimens, 25 km N Magadan, in grass, 3. vii. 1985 (Yu. M. Marusik); 10 specimens, 30 km N Magadan, Snezhnaya Dolina, viii­ix. 1996 (Yu. M. Marusik) (SPSU); Ul’yanovsk Reg.: ♂, ♀, Ul’yanovsk, near the Sviyaga River, 5. v. 1983 (Isayev) (SPSU); UNITED STATES: Alaska: 2 specimens, env. of Fairbanks, Chena Ridge, Chena Ridge Rd., 64 º 50.57 ' N 147 º 52.83 ' W, 100 m, forest litter (V. I. Gusarov), 20. vii. 1998 (SPSU); 1 specimen, Big Delta, 64 º 09 ’ N 145 º 50 ’ W, 10. viii. 1968 (W. Ivie); 13 specimens, Chatanika, 65 º 07 ’ N 147 º 30 ’ W, 13. viii. 1968 (W. Ivie); 6 specimens, Fairbanks, 13. viii. 1968 (W. Ivie) (AMNH); California: Sierra Co.: 4 specimens, 6.5 mi. E Sierra City, N. Yuba River, 5600 ’, 25. vi. 1976 (L. & N. Herman); Siskiyou Co.: 2 specimens, 1.5 mi. S McCloud, Soda Springs, 3200 ’, 1. vii. 1976 (L. & N. Herman) (AMNH); Colorado: Boulder Co.: 6 specimens, Boulder (NMNH (Casey collection )); Chaffee Co.: ♀, Buena Vista, 7900 – 8000 ’, 1 – 6. vii. 1896 (H. F. Wickham) (NMNH (Casey collection )); Dolores Co.: 5 specimens, near Fish Creek, 27 mi. NE Dolores, 8200 ’, 24. vii. 1976 (L. & N. Herman) (AMNH); Michigan: Washtenaw Co.: 2 specimen, Ann Arbor (AMNH); Minnesota: Carlton Co.: 6 specimens, 14 km SW Carlton, Hwy. 35, 46 º 33.56 ' N 92 º 35.98 ' W, 200 m, in forest litter, Betula, Populus, 10. vii. 1999 (V. I. Gusarov) (SPSU); New Mexico: Colfax Co.: 9 specimens, Cimarron Canyon W of Ute Park, 6. x. 1965 (J. & W. Ivie) (AMNH); Oregon: Klamath Co.: 3 specimens, Kimball State Park, Wood River Spring, 2 mi. N Fort Klamath, 20. vi. 1978 (J. Schuh, L. & N. Herman); 3 specimens, Wood River Spring, willow duff, 20. vi. 1978 (J. Schuh & L. Herman); 2 specimens, ditto but 2. v. 1974 (J. Schuh); 22 specimens, 7.5 mi. NW Klamath Falls, Geary Canal, Howard Bay of Upper Klamath Lake, 19. vi. 1978 (J. Schuh, L. & N. Herman) (AMNH); South Dakota: Custer Co.: 36 specimens, 7 mi. WSW Custer, North Pole Spring, 5500 ’, 15. vi. 1981 (L. Herman) (AMNH); Utah: San Juan Co.: 1 specimen, Monticello, 10. vi. 1987 (L. Herman); Summit Co.: 1 specimen, 10 mi. ESE Kamas, Shingle Cr., 7700 ’, 10. vi. 1981 (L. Herman); 15 specimens, 7 mi. ESE Kamas, Beaver Cr., 7300 ’, 10. vi. 1981 (L. Herman) (AMNH); Washington: Lewis Co.: 3 specimens, Packwood, Cowlitz Riv., leaf litter, 21 – 23. vii. 1969 (L. Herman) (AMNH); Wisconsin: Ashland Co.: 4 specimens, 9 mi. SW Marengo, near Beaver Lake, 23. vi. 1981 (L. Herman); 4 specimens, ditto but 7. viii. 1978 (L. & N. Herman); 8 specimens, 6 mi. S Marengo, near Brunsweiler Riv., 1400 ’, 7. viii. 1978 (L. & N. Herman) (AMNH); Bayfield Co.: 8 specimens, Bayfield (Wickham) (NMNH (Casey collection )); Wyoming: Fremont Co.: 12 specimens, 10 mi. SW Lander, Middle Fork Popo Agie Riv., 7100 ’, 12. vi. 1981 (L. Herman); 4 specimens, 16 mi. SW Lander Roaring Fork Creek, 8400 ’, 12. vi. 1981 (L. Herman) (AMNH).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF93535ACD24568EFCA0A6C9.taxon	diagnosis	Diagnosis. See Lohse 1974 c (as Dasyglossa prospera); Lohse et al. 1990 (as De. congruens).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF93535ACD24568EFCA0A6C9.taxon	discussion	Discussion. There is some controversy on how many species of Devia occur in North America. Bernhauer and Scheerpeltz (1926) listed Das. prospera as a single valid species of Dasyglossa and considered it as a Holarctic species. They listed Das. angularis (Mäklin in Mannerheim, 1853) and Das. congruens (both described from North America) as synonyms of Das. prospera. Moore and Legner (1975) followed Bernhauer and Scheerpeltz (1926). Seevers (1978) listed De. congruens and De. prospera as separate valid species, both represented in North America. Muona (1984) confirmed that De. prospera occurred in North America but did not comment on the status of De. congruens, perhaps because he did not have an opportunity to examine the type of the latter species. Lohse and Smetana (1985) designated the lectotype of Myrmedonia angularis Mäklin in Mannerheim, 1853 and transferred it to the genus Lypoglossa Fenyes, 1918 in the tribe Athetini. They considered De. prospera and De. congruens as separate species and illustrated the aedeagi for both. Lohse and Smetana (1985) were mostly interested in clarifying the status of My. angularis and provided no information on the localities of the illustrated specimens; nor did Lohse and Smetana discuss the geographical distribution of De. prospera and De. congruens. In a later paper (Lohse et al. 1990) dealing with the Arctic North American aleocharines Lohse mentioned only De. congruens. Unfortunately, he did not explicitly state whether, in his opinion, De. prospera occurred in North America. Ashe (Newton et al. 2000) states that only one species of Devia (De. congruens) occurs in North America. My examination of the types of Oxypoda prospera and O. congruens, as well as additional specimens did not reveal any differences between the specimens from the Nearctic and Palaearctic Regions in the shape of the aedeagus and the sclerites of its internal sac. The drawings in the paper by Lohse and Smetana (1985: Figs. 22 – 23) are very schematic and the difference between them results mostly from different extent of the evertion of the internal sac in the two preparations. I consider De. congruens to be a synonym of De. prospera, a species with Holarctic distribution and the single valid species in the genus Devia.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF93535ACD24568EFCA0A6C9.taxon	distribution	Distribution. Devia prospera has a circumpolar distribution. In North America De. prospera is known from Canada (British Columbia, Northwest Territories (Lohse et al. 1990), Alberta, Manitoba, Ontario and Labrador) and the United States (Alaska, Washington, Oregon, California, Montana, Wyoming, Utah, Colorado, New Mexico, South Dakota, Minnesota, Wisconsin and Michigan).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF96535BCD245606FD8EA3B9.taxon	type_taxon	type species: Atheta densissima Bernhauer, 1909; fixed by Fenyes (1918), by subsequent designation).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF96535BCD245606FD8EA3B9.taxon	diagnosis	Diagnosis. See Bernhauer (1909) and Fenyes (1920).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF96535BCD245606FD8EA3B9.taxon	discussion	Discussion. My examination of the type species of Paradilacra revealed that its aedeagus lacks the athetine bridge. Based on the shape of the aedeagus and the sclerites of its internal sac the genus Paradilacra is transferred to the subtribe Tachyusina Thomson, 1859 of the tribe Oxypodini.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF975350CD24528BFDA7A3B9.taxon	description	(Figs. 19 – 30)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF975350CD24528BFDA7A3B9.taxon	materials_examined	Type material. Lectotype of Atheta densissima (here designated): ♂, “ S. Anselmo Cal. [ifornia] ”, “ June 04.16 [4. vi. 1916] ”, “ 166. ”, “ ambigua Er.? Fenyes. det. Bernhauer ”, “ densissima Bernh. Typus. Fenyes ” (yellow label), “ not ambigua Erichson det. Lohse ”, (FMNH). Paralectotype: ♂, “ Willist’n [Williston], N. [orth] D. [akota] Jun. [e] 8 – 9 Wickham ”, “ 7614 ”, “ densissima Bernh. Cotypus. Fenyes ” (yellow label) (FMNH). Lectotype of Paradilacra persola (here designated): ♂, “ Cal. [ifornia, Monterey] ”, “ Paradilacra persola Csy. ”, “ TYPE USNM 39377 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotype: ♂, “ Cal. [ifornia, Monterey] ”, “ persola PARATYPE USNM 39377 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Paradilacra willametta (here designated): ♂, “ Portland Oreg. [on] ”, “ willametta Csy. ”, “ TYPE USNM 39380 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Paradilacra uintana (here designated): ♂, “ Milford Ut. [ah] Jul. [y] Wickha [m] ”, “ uintana Csy. ”, “ TYPE USNM 39379 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: ♂, “ Milford Ut. [ah] Jul. [y] Wickha [m] ”, “ uintana ­ 2 PARATYPE USNM 39379 ” (red label), “ CASEY bequest 1925 ”; ♂, “ Elko, Nevad [a] Wickham. ”, “ uintana ­ 3 PARATYPE USNM 39379 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Paradilacra glenorica (here designated): ♂, “ Glenora B. [ritish] C. [olumbia] ”, “ glenorica Csy. ”, “ TYPE USNM 39378 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotype: ♀, “ Kamloops, B. [ritish] C. [olumbia] Wickham. ”, “ glenorica ­ 3 PARATYPE USNM 39378 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Paradilacra symbolica (here designated): ♀, “ Cal. [ifornia, Napa Junction] ”, “ symbolica Csy. ”, “ TYPE USNM 39385 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Paradilacra erebea (here designated): ♀, “ Cal. [ifornia, Napa Junction] ”, “ erebea Csy. ”, “ TYPE USNM 39381 ” (red label), “ CASEY bequest 1925 ”. Paralectotype: ♂, “ Cal. [ifornia, near Mt. Diablo] ”, “ erebea ­ 2 PARATYPE USNM 39381 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Paradilacra subaequa (here designated): ♂, “ Cal. [ifornia, Santa Rosa] ”, “ subaequa Csy. ”, “ TYPE USNM 39382 ” (red label), “ CASEY bequest 1925 ”. Paralectotype: ♂, “ Cal. [ifornia, Santa Rosa] ”, “ subaequa ­ 2 PARATYPE USNM 39382 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Paradilacra sinistra (here designated): ♂, “ Cal. [ifornia, San José and Santa Clara Valley] ”, “ sinistra Csy. ”, “ TYPE USNM 39383 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Paradilacra memnonia (here designated): ♀, “ Cal. [ifornia, Cloverdale, north of Santa Rosa] ”, “ memnonia Csy. ”, “ TYPE USNM 39384 ” (red label), “ CASEY bequest 1925 ” (NMNH). Holotype of Paradilacra vulgatula: ♂, “ Cal. [ifornia, Jountville, Napa Co.] ”, “ vulgatula Csy. ”, “ TYPE USNM 39386 ” (red label), “ CASEY bequest 1925 ”. Paratype: ♀, “ Cal. [ifornia, Jountville, Napa Co.] ”, “ vulgatula ­ 2 PARATYPE USNM 39386 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Paradilacra deserticola (here designated): ♀, “ Elko Nev. [ada] ”, “ deserticola Csy. ”, “ TYPE USNM 39387 ” (red label), “ CASEY bequest 1925 ” (NMNH). Additional material. UNITED STATES: California: ♀, “ Cal ” with a black dot inside “ C ” (San Francisco and immediate vicinity as far south as Redwood City and Purissima (FitzGerald 1962 )) (NMNH (Casey collection )); San Bernardino Co.: 32 specimens, 4 km NEE Big Bear City, Hwy. 18, San Bernardino Nat. Forest, 34 º 16.46 ’ N 116 º 48.07 ’ W, 2050 m, Baldwin Lake bank, 11. viii. 1999 (V. I. Gusarov) (KSEM); San Diego Co.: 3 ♂♂, 5 ♀♀, Cuyamaca, 11. vi. 1961 (I. Moore) (CNCI, SPSU); San Luis Obispo Co.: ♂, Atascadero, 15. viii. 1956 (I. Moore) (CNCI); Ventura Co.: 14 specimens, 10 km N Ojai, Los Padres Nat. Forest, Rose Valley Rd., 34 º 32.51 ’ N 119 º 11.42 ’ W, 1150 m, reservoir bank, 10. viii. 1999 (V. I. Gusarov) (KSEM); Montana: ♂, Glendive (This specimen was labeled by curators as a paratype of Pa. glenorica but it had not been included in the type series of that species by Casey (1910 a) (NMNH (Casey collection )); CANADA: Alberta: ♂, ♀, Waterton Lakes N. P., bes. Hwy. 6, 4.4 km SE jct. Hwy. 5, tread beaver pond, 6. vi. 1980 (CNCI); British Columbia: ♂, Thompson River, Monte Creek, 27. vi. 1988 (L. LeSage); ♂, 16 mi. W Osoyoos, 5. vi. 1968 (Campbell & Smetana) (CNCI).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF975350CD24528BFDA7A3B9.taxon	diagnosis	Diagnosis. Paradilacra densissima is the only valid Nearctic species of the genus. When describing Paradilacra, Bernhauer (1909) included in it an additional Nearctic species (Atheta ambigua (Erichson, 1839 )) and one species from North Africa (At. hyperbolica Bernhauer, 1905). In this paper At. ambigua is placed in the genus Strigota Casey, 1910 a (see below). The aedeagus and spermatheca of Pa. densissima are shown in Figs. 23 – 30.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF975350CD24528BFDA7A3B9.taxon	discussion	Discussion. My comparison of the types of the species described by Casey (1910 a, 1911 a) in the genus Paradilacra with the types of Pa. densissima confirmed the synonymies established by Fenyes (1920). The body size and the length of the female spermatheca (Figs. 29 – 30) varies in examined specimens but this variation is continuous. In all examined male specimens the shape of the aedeagus and the sclerites of its internal sac were identical.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF975350CD24528BFDA7A3B9.taxon	distribution	Distribution. Paradilacra densissima is widely distributed in the western United States (California, Nevada, Oregon, Utah, Montana and North Dakota) and Canada (British Columbia and Alberta).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF9C5350CD24517CFC8CA4E6.taxon	type_taxon	(type species: Aleochara graminicola Gravenhorst, 1806, by monotypy, fixed by the International Commission on Zoological Nomenclature (ICZN 1961 )).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF9C5350CD24517CFC8CA4E6.taxon	discussion	Discussion. The magnitude of the genus Atheta varies in different contemporary works (Benick & Lohse 1974; Muona 1979 b; Lohse et al. 1990). In the narrowest sense (e. g., Lohse et al. 1990, where it is equivalent to Atheta s. str. of Benick and Lohse 1974) Atheta includes the athetines with pronotal setation of type I (Benick & Lohse 1974), with hypomera fully visible in lateral view and with distinctly shaped spermatheca with thick distal portion and thin coiled proximal portion (Figs. 150 – 155 in Strand & Vik 1964). In other interpretations (Benick & Lohse 1974; Muona 1979 b), which can be traced back to Ganglbauer (1895), Atheta is a much wider group. If this wider interpretation of the genus is accepted it is hard to say what the autapomorphies are which define it. In the key by Benick and Lohse (1974; pp. 72 – 79) one arrives at Atheta at the very end of the key, after having eliminated all other athetine genera, and essentially the genus is characterized by the features it lacks. Further analysis is needed to determine whether Atheta in the broad sense can be maintained as a monophyletic group.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF9D536CCD24524EFEA9A431.taxon	description	(Fig. 155 in Strand & Vik 1964)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF9D536CCD24524EFEA9A431.taxon	materials_examined	Type material. Lectotype of Aleochara graminicola (here designated): ♂, “ 5349 ”, “ graminicola Gr. ”, “ Zool. Mus. Berlin ” (ZMHB). Paralectotypes: 2 ♂♂, 2 ♀♀, without labels; ♀, “ Angl. Rudd ”; ♀, “ Nordeur. Schupp. ” (ZMHB). Lectotype of Homalota granulata (designated by Lohse and Smetana (1985 )): UNITED STATES: Alaska: ♀, Unalaska Island (Cygnaeus) (MZHF). Lectotype of Atheta lanei (here designated): ♀, “ Lane Co., Or. [egon] ”, “ lanei Csy. ”, “ TYPE USNM 39462 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta nomadica (here designated): ♀, “ Nome, Alaska ”, “ nomadica Csy. ”, “ TYPE USNM 39478 ” (red label), “ CASEY bequest 1925 ” (NMNH). Additional material. CANADA: Alberta: ♂, ♀, Banff (Fenyes); 3 ♀♀, 2 mi. S Ponaka, Battle Riv. marsh, 6. viii. 1976 (P. M. Hammond); ♂, 2 ♀♀, Waterton N. P., Cameron Lake, 540 ’, 2 – 4. viii. 1976 (P. M. Hammond) (BMNH); British Columbia: ♂, Alta Lake, 1. viii. 1975 (J. M. & B. A. Campbell) (CNCI); Labrador: ♀, L’Anse au Loup, 9. viii. 1972 (J. M. Campbell); 2 ♂♂, Hebron, 9. vii. 1954 (J. F. McAlpine) (CNCI); Manitoba: ♀, Churchill, 29. vi. 1937 (W. J. Brown); ♀, ditto but 20. vii. 1937; ♂, ditto but 18. viii. 1937; ♂, 2 ♀♀, ditto but 16. vi. 1937; 2 ♂♂, 6 ♀♀, ditto but 4. vi. 1952 (J. G. Chillcott); 4 ♂♂, 4 ♀♀, ditto but 13. vi. 1952; ♂, 3 ♀♀, ditto but 17. vi. 1952; ♂, ♀, ditto but 2. viii. 1952; 3 ♂♂, ♀, ditto but 5. viii. 1952; ♂, ditto but 15. viii. 1952; ♀, ditto but 27. viii. 1948 (W. R. Richards) (CNCI); Newfoundland: ♂, Long Range Mts., 50 º 34 ’ N 56 º 56 ’ W, 1400 ’, 24. vii. 1969 (M. J. D. Brendell) (BMNH); Northwest Territories: 2 ♀♀, Ft. Simpson, Harris River, 15. vi. 1972 (A. Smetana); ♂, ditto but 10. vi. 1972; 5 ♂♂, 19 ♀♀, Trail River, 32 mi. NW Ft. Simpson, 16. vi. 1972 (A. Smetana); 3 ♂♂, 4 ♀♀, Hwy. 3, 5 mi. SE Ft. Simpson, 17. vi. 1972 (A. Smetana); 3 ♂♂, 2 ♀♀, ditto but 21. vi. 1972; ♀, Hwy. 3, 8 mi. SE Ft. Simpson, 18. vi. 1972 (A. Smetana); ♂, Spence River, 38 mi. SE Ft. Simpson, 19. vi. 1972 (A. Smetana); 3 ♂♂, 13 ♀♀, Reindeer Sta., Caribou Hills, 30. vi. 1972 (A. Smetana); ♀, Yellowknife, 20. v. 1953 (J. G. Chillcott); ♂, ditto but 29. v. 1953; 2 ♀♀, Aklavik, 9. vi. 1956 (R. E. Leech); 2 ♀♀, ditto but 12. vi. 1956; ♂, ditto but 14. vi. 1956; ♂, ditto but 3. vii. 1956; ♀, ditto but 6. vii. 1956; ♂, ditto but 13. vi. 1956 (E. F. Cashman); ♀, ditto but 16. vi. 1953 (C. D. Bird); 10 ♀♀, Inuvik, Shell Lk., 27. vi. 1972 (A. Smetana); ♀, ditto but 1. vii. 1972 (A. Smetana); 6 ♀♀, ditto but 25. vi. 1972; 2 ♂♂, ♀, Reindeer Depot, Mackenzie Delta, 8. viii. 1948 (W. J. Brown); ♀, Kidluit Bay, N. Richards Is., 26. vii. 1948 (T. J. R. Vockeroth); ♀, Muscox L., 64 º 45 ’ N 108 º 10 ’ W, 2. viii. 1953 (J. G. Chillcott); 3 ♂♂, 1 specimen, ditto but 21. vii. 1953 (J. G. Chillcott); ♂, Salmita Mines, 64 º 05 ’ N 111 º 15 ’ W, 20. vi. 1953 (J. G. Chillcott); 2 ♀♀, Unnamed Lake, 18 mi. NW Inuvik via East Channel, 26. vi. 1972 (A. Smetana) (CNCI); Ontario: ♂, 2 ♀♀, Butterfly Lk., 13 mi. S Sioux Lookout, 17. vi. 1973 (Campbell & Parry) (CNCI); Québec: 7 ♂♂, 15 ♀♀, Blanc Sablon, 9. viii. 1972 (J. M. Campbell); ♂, 2 ♀♀, Koroc Riv., in leaf litter beside stream, 3. viii. 1986 (A. V. A. & S. Morgan); ♂, Whale Riv., 8. vii. 1949 (J. R. Vockeroth) (CNCI); Yukon Territory: ♀, Caribou Bar Creek, 20. vi. 1972; 2 ♂♂, 5 ♀♀, McQuesten Lk., 15 mi. SW Keno, 19. vii. 1968 (Campbell & Smetana); 2 ♂♂, 3 ♀♀, Mi. 1034, Alaska Hwy., near Kloo Lake, 5. vii. 1968 (Campbell & Smetana); 4 ♀♀, Mi. 1120, Alaska Hwy., 6. vii. 1968 (Campbell & Smetana); ♀, Mi. 1192, Alaska Hwy., near Snag Junction, 6. vii. 1968 (Campbell & Smetana); ♀, Mi. 1120, Alaska Hwy., 6. vii. 1968 (Campbell & Smetana); ♂, ♀, Mi. 1705, Alaska Hwy., Slim’s River, 7. vii. 1981 (Bright); ♂, 10 mi. E Dawson City, 11. vii. 1968 (Campbell & Smetana); 3 ♂♂, 2 ♀♀, Dempster Hwy., Mi. 236, Eagle Riv., 1400 ’, 12. vii. 1978 (Campbell & Smetana); 2 ♂♂, Dempster Hwy., Mi. 222, 1900 ’, 21. vii. 1978 (Smetana & Campbell); 3 ♂♂, 6 ♀♀, ditto but Mi. 147, 22. vii. 1978 (Campbell & Smetana); 2 ♀♀, ditto but Mi. 139.5, 20. vii. 1978 (Campbell & Smetana); ♀, ditto but Mi. 73, 3300 ’, 12. vii. 1968 (Campbell & Smetana); 2 ♂♂, 10 ♀♀, ditto but Mi. 71; ♂, 7 ♀♀, ditto but Mi. 65, 3300 ’, 13. vii. 1968; 4 ♂♂, ♀, ditto but Mi. 60, 3500 ’, 19. vii. 1978; 4 ♂♂, 4 ♀♀, ditto but 23. vii. 1978; ♂, 2 ♀♀, ditto but Mi. 55, 3300 ’, 13. vii. 1968; 5 ♂♂, 2 ♀♀, ditto but Mi. 53, North Fork Pass, 4200 ’, 24. vii. 1978; 3 ♂♂, ♀, ditto but Mi. 48.5, North Fork Pass, 4100 ’, 24. vii. 1978; ♂, 3 ♀♀, ditto but 19. vii. 1978; 5 ♂♂, 2 ♀♀, ditto but Mi. 45, 3500 ’, 14. vii. 1968; ♀, ditto but 3400 ’, 18 – 23. vii. 1978 (J. M. Campbell); 11 ♂♂, 13 ♀♀, ditto but Mi. 42, N Klondike Riv., 3300 ’, 18. vii. 1978 (Campbell & Smetana); ♂, ditto but Mi. 32, 2700 ’, 15. vii. 1968; ♂, ditto but Mi. 28, 15. vii. 1968; ♀, Ogilvie Mts., North Fork Pass, 4100 ’, 18. vi. 1962 (P. J. Skitsko); ♂, ♀, Rampart House, 28. v. 1951 (J. E. H. Martin); ♂, ditto but 1. vi. 1951; 4 ♀♀, British Mts., 340 m, 4 km NE Wolf Cr., 69 º 08 ’ N 140 º 14 ’ W, 23. vi. 1984 (Campbell); ♂, ♀, British Mts., 320 m, 3 km NW Firth Riv., 69 º 13 ’ N 140 º 05 ’ W, sifting Salix litter, 19. vi. 1984 (Campbell); ♂, ♀, British Mts., 180 m, Fish Creek, 69 º 27 ’ N 140 º 19 ’ W, sifting under tall Salix shrubs, 3. vii. 1984 (J. M. Campbell); 2 ♂♂, 3 ♀♀, Hansen Lk., 9 mi. SW Keno, 18. vii. 1968 (Campbell & Smetana); ♂, La Force Lake, 62 º 41 ’ N 132 º 20 ’ W, 3300 ’, 28. vi. 1960 (J. E. H. Martin); ♀, Swim Lakes, 62 º 13 ’ N 133 º 00 ’ W, 3200 ’, 15. vi. 1960 (J. E. H. Martin) (CNCI); ♀, Whitehorse, 21. vi. 1924 (MCZ); UNITED STATES: Alaska: 16 ♂♂, 17 ♀♀, Mi. 123, Taylor Hwy., 9. vii. 1968 (Campbell & Smetana); ♀, Mi. 23, Taylor Hwy., 8. vii. 1968 (Campbell & Smetana); ♀, Denali State Park, Mi. 165.5, Hwy. 3, 23. vi. 1978 (Smetana & Becker); 2 ♂♂, 3 ♀♀, Mi. 110, Denali Hwy., Seattle Cr., 15. vii. 1978 (Campbell & Smetana); 4 ♂♂, 5 ♀♀, Mi. 102, Denali Hwy., 12. vii. 1978 (Smetana & Campbell); 2 ♀♀, Mi. 94, Denali Hwy., Canyon Creek, 13. vii. 1978 (J. M. Campbell & A. Smetana); ♂, 12 mi. N Mi. 78, Denali Hwy., Windy Cr., 13. vii. 1978 (Smetana & Campbell); 6 ♂♂, 8 ♀♀, Mi. 1249, Alaska Hwy., Deadman Lk., 6 – 7. vii. 1968 (Campbell & Smetana); 2 ♂♂, 6 ♀♀, Mi. 1252, Alaska Hwy., 7. vii. 1968 (Campbell & Smetana); 2 ♂♂, ♀, Kenai Pen., Kenai, 24. vi. 1951 (W. J. Brown); ♂, ♀, Kenai Pen., 3 mi. SE Kenai, 7. vi. 1978 (Smetana & Becker); ♀, Kenai Pen., Hope, 24. v. 1951 (W. J. Brown); 5 ♂♂, 7 ♀♀, Kenai Pen., Anchor River at Hwy. 1, 450 ’, 2. vi. 1978 (Smetana & Becker); ♂, 20 ♀♀, ditto but 4. vi. 1978; ♀, ditto but 5. vi. 1978; ♂, 4 ♀♀, Kenai Pen., Anchor River Campground, 12 mi. N Homer, 450 ’, 5. vi. 1978 (Smetana & Becker); ♀, ditto but 3. vi. 1978; 4 ♂♂, 5 ♀♀, Kenai Pen., 1 mi. W Anchor Pt., 1. vi. 1978 (Smetana & Becker); 7 ♂♂, 3 ♀♀, Kenai Pen., Crooked Cr. at Hwy. 1, S Kasilof, 9. vi. 1978 (Smetana & Becker); 5 ♂♂, 11 ♀♀, Kenai Mts., 15 mi. N Seward, 400 ’, 29. v. 1978 (Smetana & Becker); 6 ♂♂, 9 ♀♀, Kenai Mts., 16 mi. N Seward, 500 ’ – 600 ’, 26. v. 1978 (Smetana & Becker); 4 ♂♂, 6 ♀♀, Kenai Mts., 22 mi. N Seward, 600 ’ – 800 ’, 27. v. 1978 (Smetana & Becker); 2 ♂♂, 5 ♀♀, Seward, 20. v. 1951 (W. J. Brown); 2 ♂♂, Kenai Pen., 15 mi. E Sterling, 14. vi. 1978 (Smetana & Campbell); 2 ♂♂, ♀, Kenai Pen., 2 mi. NE Soldotna, 10. vi. 1978 (Smetana & Becker); 4 ♂♂, Kenai Pen., Clam Gulch, 6. vi. 1978 (Smetana & Becker); 3 ♂♂, 5 ♀♀, Kenai Mts., Cooper Lk., 1000 ’, 15. vi. 1978 (Smetana & Becker); ♀, Kenai Mts., 8 mi. W Cooper Landing, 500 ’, 14. vi. 1978 (Smetana & Becker); ♀, Kenai Mts., Quarts Cr. At Hwy. 1, 500 ’, 15. vi. 1978 (Smetana & Becker); ♂, 3 ♀♀, Kenai Mts., Tern Lk. Campground, 700 ’, 18. vi. 1978 (Smetana & Becker); ♀, 8 mi. NW Haines, 3. vii. 1968 (Campbell & Smetana) (CNCI); 3 specimens, Skagway, 18 – 20. vi. 1924 (H. C. Fall) (CUIC); 4 ♂♂, ♀, ditto but 18. vi. 1924 (MCZ); 3 ♂♂, 8 ♀♀, Hess Cr., Mi. 24, Wales Hwy., 1. vii. 1978 (J. M. Campbell & A. Smetana); 3 ♂♂, ♀, ditto but 600 ’, 65 º 40 ’ N 149 º 10 ’ W, 10. vii. 1978 (J. M. Campbell & A. Smetana); 9 ♂♂, 5 ♀♀, Unalakleet, 4. vii. 1961 (R. Madge); 2 ♀♀, ditto but 23. vi. 1961; ♀, ditto but 17. vi. 1961; ♀, ditto but 11. viii. 1961; ♀, King Salmon, Naknek River, 6. vii. 1952 (W. R. Mason); 1 specimen, Nome, 10. vii. 1951 (D. P. Whillians); ♂, Mi. 220, George Parks Hwy., 14. vii. 1978 (A. Smetana) (CNCI); 2 ♀♀, Fairbanks, 7. vii. 1924 (MCZ); 2 ♀♀, Prudhoe Bay Rd., Bonanze Cr., 66 º 40 ’ N 150 º 40 ’ W, 900 ’, 2. vii. 1978 (A. Smetana & J. M. Campbell); ♀, Prudhoe Bay Rd., 4 mi. SW Chandalar Cmp., 68 º 04 ’ N 149 º 40 ’ W, 4000 ’, 7. vii. 1978 (Campbell & Smetana); 2 ♂♂, 4 ♀♀, Prudhoe Bay Rd., Coldfoot nr. Clara Cr., 67 º 16 ’ N 150 º 10 ’ W, 1100 ’, 3. vii. 1978 (A. Smetana & J. M. Campbell); 3 ♀♀, Prudhoe Bay Rd., 8 mi. N S. Fork Koyukuk Riv., 67 º 13 ’ N 150 º 07 ’ W, 1000 ’, 8. vii. 1978 (A. Smetana & J. M. Campbell); 4 ♂♂, 6 ♀♀, Prudhoe Bay Rd., 0.5 mi. N Yukon Riv., 65 º 52 ’ N 149 º 45 ’ W, 400 ’, 9. vii. 1978 (A. Smetana & J. M. Campbell) (CNCI); 2 specimens, Glenn Hwy. (Tok cut­off), NE of Gakona, 62 º 19.89 –. 88 ' N 145 º 12.60 –. 27 ' W, 550 m, a bog, lake shore, Carex, Comarum, 16. vii. 1998 (V. I. Gusarov); 4 specimens, Kenai Peninsula, 3 km SE Anchor Point, Sterling Hwy., Anchor River valley, 59 º 45.40 ' N 151 º 47.61 ' W, 0 m, forest litter, Alnus, 22. vii. 1998 (V. I. Gusarov) (SPSU); ♀, Dutch Harbor Unalaska I., 9. vii. 1907 (E. C. Van Dyke) (MCZ); RUSSIA: Chukotka Pen.: 7 specimens, NW part of the Krest Bay, along river, Salix grove with dense herbs at the margins, 28. vii. 1988 (Yu. M. Marusik) (SPSU); Moscow Reg.: ♂, Ramenskoye distr., Kraskovo, river bank, 30. vi. 1989 (V. I. Gusarov) (SPSU); Murmansk Reg.: 10 specimens, 150 km NE Murmansk, env. of Dalniye Zelentsy, Barentsevo Sea, Yarnyshnaya, seashore, under beach wrack, 4. viii. 1997 (A. Przhiboro); 5 specimens, Khibiny Mts., Bolshoy Vud'yavr Lake, 31. vii. 1984 (V. I. Gusarov) (SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF9D536CCD24524EFEA9A431.taxon	diagnosis	Diagnosis. See Benick & Lohse (1974). Aedeagus and spermatheca: Fig. 155 in Strand & Vik (1964).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF9D536CCD24524EFEA9A431.taxon	discussion	Discussion. The lectotype of At. nomadica represents a small specimen of At. graminicola, and falls within the range of intraspecific variation of At. graminicola in punctation, proportions of antennal segments and the shape of the spermatheca. The lectotype of At. lanei is similar to examined specimens of At. graminicola in external characters and in the shape of the spermatheca. Lohse and Smetana (1985) argued that At. graminicola and At. granulata are closely related but different species, citing the difference in the shape of the aedeagus and spermatheca and in punctation of the pronotum and elytra. The proximal portion of the spermatheca in At. graminicola is variable and the difference between Figs. 5 and 7 in Lohse and Smetana (1985) is within the range of variability of this species. I was unable to confirm the difference between Palaearctic specimens of At. graminicola and Nearctic specimens. I consider At. granulata to be synonymous with At. graminicola.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FF9D536CCD24524EFEA9A431.taxon	distribution	Distribution. Atheta graminicola is a Holarctic species. In North America it is known from Canada (British Columbia, Yukon Territory, Northwest Territories, Alberta, Manitoba, Ontario, Québec, Labrador, Newfoundland) and the United States (Alaska and Oregon).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFA05368CD245533FDC3A0F9.taxon	description	(Figs. 31 – 40)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFA05368CD245533FDC3A0F9.taxon	materials_examined	Type material. Lectotype of Homalota aemula (here designated): ♀, “ America sept. Zimmermann Nr. 5401 ”, “ Typus ” (red label), “ Zool. Mus. Berlin ” (ZMHB). Paralectotypes: ♀, “ 5401 ”, “ aemula Er. Am. Spt. Zimm. [ermann] ” (green label), “ Typus ” (red label), “ Zool. Mus. Berlin ”; ♀, “ America sept. Zimmermann Nr. 5401 ”, “ aemula Er. ”, “ Typus ” (red label), “ Zool. Mus. Berlin ”, “ Homalota aemula Er. [on red side] / Lectotypus Lohse fix. 1987 [on white side] ” [Lohse never published this lectotype designation] (ZMHB). Lectotype of Homalota polita (here designated): ♂, “ polita M. [elsheimer] Pa. [Pennsylvania] ”, “ polita ”, “ MCZ TYPE 34937 ” (red label) (MCZ) (In MCZ there is another poorly preserved specimen (♀) with a damaged head and abdomen, labeled “ U. [nited] S. [tates] ”, “ Melsh. [eimer] ”, “ polita Melsh. ”, a red piece of paper, “ MCZ TYPE 34937 ” (red label). This specimen is not conspecific with the lectotype of H. polita and belongs to Earota dentata (Bernhauer, 1906). The specimen was not labeled as paralectotype because its locality label is not Pennsylvania, the type locality of H. polita.) Lectotype of Atheta disjuncta (here designated): ♂, “ Alleghen [y] Pa [Pennsylvania] ”, “ Atheta disjuncta Csy. ”, “ TYPE USNM 39283 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 2 ♂♂, ♀, “ Alleghen [y] Pa [Pennsylvania] ”, “ disjuncta PARATYPE USNM 39283 ” (red label), “ CASEY bequest 1925 ”; 6 ♂♂, 2 ♀♀, “ · N. [orth] C. [arolina, Asheville] ”, “ disjuncta PARATYPE USNM 39283 ” (red label), “ CASEY bequest 1925 ”; (NMNH). Lectotype of Atheta replicans (here designated): ♂, “ Ia. [Iowa, Cedar Rapids (Brendel)] ”, “ Atheta replicans Csy. ”, “ TYPE USNM 39287 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 2 ♂♂, 3 ♀♀, “ Ia. [Iowa, Cedar Rapids (Brendel)] ”, “ replicans PARATYPE USNM 39287 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta spadix (here designated): ♂, “ N. [ew] Y. [ork, near New York City] ”, “ Atheta spadix Csy. ”, “ TYPE USNM 39288 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotype: ♀, “ N. [ew] Y. [ork, near New York City] ”, “ spadix ­ 2 PARATYPE USNM 39288 ” (red label), “ CASEY bequest 1925 ” (NMNH). Holotype of Atheta bucolica: ♀, “ · Miss [issippi, Vicksburg] ”, “ Atheta bucolica Csy. ”, “ TYPE USNM 39289 ” (red label), “ CASEY bequest 1925 ” (NMNH). Additional material. UNITED STATES: Iowa: Johnson Co.: ♂, Iowa City (Wickham) (NMNH (Casey Collection )); Kansas: Douglas Co.: 10 specimens, 2.5 km N Lawrence, 38 º 59.59 ’ N 95 º 14.59 ’ W, 200 m, flood refuse by the Kansas River (right bank), 18. iv. 1999 (V. I. Gusarov); 6 specimens, 1.5 km N Lawrence, 38 º 58.96 ’ N 95 º 14.62 ’ W, 200 m, in rotting wood chips, 1. v. 1999 (V. I. Gusarov); 2 ♂♂, 18 km SSE Lawrence, Breidenthal Preserve, 38 º 48.50 ’ N 95 º 11.52 ’ W, 250 m, on mushrooms, 23. v. 2000 (V. I. Gusarov) (SPSU); New Jersey: Burlington Co.: ♂, Pine Barrens, Whitesbog, sifting, 30. v. 1999 (A. Berkov) (SPSU); New York: Orange Co.: ♀, 8 km S New Windsor, Black Rock Forest, 9. v. 1998 (V. I. Gusarov) (SPSU); Texas: Blanco Co.: 8 specimens, 16 km NEE Johnson City, Pedernales Falls State Park, 30 º 18.87 ’ N 98 º 14.67 ’ W, 300 m, on mushrooms (Morchella), 1. iv. 2001 (V. I. Gusarov) (SPSU); CANADA: Québec: ♂, Chûte­aux­Galets, 18. vii. 1994 (Lindgren) (SPSU); ♂, Aylmer, 45 º 24 ’ N 75 º 51 W, 5. vii. 1994 (LFC).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFA05368CD245533FDC3A0F9.taxon	diagnosis	Diagnosis. Atheta aemula differs from other Nearctic species of Atheta s. str. in the following characters: body brown, elytra brownish yellow with darker area around scutellum and in postero­lateral angles, legs and 1 – 2 basal antennal articles yellow; antennal article 10 slightly transverse, male abdominal tergum 8 as in Figs. 31, 33, aedeagus as in Figs. 36 – 39, spermatheca as in Fig. 40.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFA05368CD245533FDC3A0F9.taxon	discussion	Discussion. The types of H. aemula, H. polita, At. disjuncta, At. replicans, At. spadix and At. bucolica are all similar in external characters and in the shape of the aedeagus and spermatheca. All these names are considered to be synonyms.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFA05368CD245533FDC3A0F9.taxon	distribution	Distribution. Atheta aemula is widely distributed in the eastern United States (New York, New Jersey, Pennsylvania, Iowa, Kansas, Texas and Mississippi), and is also known from Canada (Québec).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFA4536ACD245076FB2FA7D9.taxon	description	(Fig. 97 in Klimaszewski & Winchester 2002; Figs. 41 – 46 in this paper)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFA4536ACD245076FB2FA7D9.taxon	materials_examined	Type material. Lectotype of Atheta keeni (here designated): ♂, “ Metlakatla, B. [ritish] Col. [umbia] Keen ”, “ Lamiota keeni Csy. ”, “ TYPE USNM 39480 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 5 ♂♂, 5 ♀♀, “ Metlakatla, B. [ritish] Col. [umbia] Keen ”, “ keeni PARATYPE USNM 39480 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta innocens (here designated): ♀, “ Lane Co., Or. [egon] ”, “ Atheta innocens Csy. ”, “ TYPE USNM 39270 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta achromata (here designated): ♂ (without head and prothorax), “ Metlakatla, B. [ritish] Col. [umbia] Keen ”, “ achromata Csy. ”, “ TYPE USNM 39481 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta profecta (here designated): ♂, “ Metlakatla, B. [ritish] Col. [umbia] Keen ”, “ profecta Csy. ”, “ TYPE USNM 39482 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotype (see discussion) of Homalota vasta Mäklin in Mannerheim, 1853: ♀, “ Sitka ”, “ Holmberg ”, “ Homalota vasta m. Sitkha ”, “ Typus ” (green label), “ Mus. Zool. H: fors. Spec. Typ. No. 2250. Homalota vasta Mäklin ”, “ Homalota vasta Mäkl. Holotypus / Liogluta vasta. Holotypus Lohse fix. 1985 ” (MZHF). Additional material. UNITED STATES: Alaska: ♀, Valdez, 61 º 09.05 ' N 146 º 21.80 ' W, 20 m, in forest litter, Alnus, ferns, Verathrum, 12. vii. 1998 (V. I. Gusarov) (SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFA4536ACD245076FB2FA7D9.taxon	diagnosis	Diagnosis. See Klimaszewski and Winchester (2002; as Atheta vasta).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFA4536ACD245076FB2FA7D9.taxon	discussion	Discussion. The lectotype of At. innocens agrees with the types of At. keeni in external characters and the shape of the spermatheca. The lectotype of At. profecta has no head and prothorax but agrees with the types of At. keeni in the shape of the aedeagus and abdominal tergum 8. The lectotype of At. profecta is poorly pigmented, but otherwise it agrees completely with At. keeni in external characters, including male secondary features, and in the shape of the aedeagus. The type series of Homalota vasta Mäklin in Mannerheim, 1853 (examined) includes two specimens (Lohse & Smetana 1985). One of them is a female conspecific with Atheta keeni and the other is a female belonging to the genus Liogluta and illustrated by Lohse and Smetana (1985). This second female bears the following labels: “ Sitka ”, “ Holmberg ”, “ Mus. Zool. H: fors. Spec. Typ. No. 2251. Homalota vasta Mäklin ”. From the description and illustrations in Lohse and Smetana (1985) it is clear that they meant to designate the second specimen (a female of Liogluta) as the lectotype of Homalota vasta. Unfortunately, when they cited the labels of the lectotype, they listed the labels of the first specimen (a female of Atheta keeni). Moreover, it is the first specimen that had been labeled as the lectotype. The latter circumstance is especially misleading and originally I was going to accept what the lectotype label said, and consider Atheta keeni and Homalota vasta to be synonymous. While identifying some athetines from British Columbia for Jan Klimaszewski (Klimaszewski & Winchester 2002) I labeled the specimens of Atheta keeni as At. vasta. However, Lohse and Smetana are very explicit in that they meant the specimen of Liogluta, illustrated and redescribed by them, to be the lectotype of Homalota vasta. Therefore I accept their intent to designate the Liogluta specimen as the lectotype, disregard their misplaced lectotype label and consider At. keeni to be a valid species, different from Liogluta vasta. In the type series of Homalota vasta I labeled the specimen of Liogluta as lectotype and the specimen of At. keeni as paralectotype. Atheta keeni is the type species of the subgenus Lamiota Casey, 1910 a (by original designation). Subgeneric assignment of At. keeni and the status of the name Lamiota requires further study. Atheta keeni has the pronotal pubescence of the type I and cannot be placed in the genus Liogluta Thomson, 1858.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFA4536ACD245076FB2FA7D9.taxon	distribution	Distribution. Atheta keeni is known from Alaska, British Columbia and Oregon.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFA6536BCD245516FD42A159.taxon	description	(Figs. 143 – 145, 148 – 151 in Lohse et al. 1990)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFA6536BCD245516FD42A159.taxon	materials_examined	Type material. Holotype of Dimetrota prudhoensis: ♂, UNITED STATES: Alaska: Prudhoe Bay Rd., Bonanze Cr., 900 ', 150 º 40 ' 66 º 40 ' (A. Smetana & J. M. Campbell), 2. vii. 1978 (CNCI). Holotype of Dimetrota dempsterensis: ♂, CANADA: Yukon Territory: Dempster Hwy. Mi. 75, 3400 ' (A. Smetana & J. M. Campbell), 23. vii. 1978 (CNCI). Additional material: UNITED STATES: New York: Orange Co.: 10 specimens, 8 km S New Windsor, Black Rock Forest, 23 – 25. v. 1998 (V. I. Gusarov) (SPSU); Vermont: Windsor Co.: ♀, 8 km SEE Springfield, Hwy. I­ 91, 43 º 14.00 ' N 72 º 26.81 ' W, 250 m, forest litter, Acer, Betula, Quercus, Tsuga, 21. ix. 1998 (V. I. Gusarov) (SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFA6536BCD245516FD42A159.taxon	diagnosis	Diagnosis. See Lohse et al. (1990).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFA6536BCD245516FD42A159.taxon	discussion	Discussion. The holotype of Dim. prudhoensis is similar to the holotype of Dim. dempsterensis in both external characters and the shape of the aedeagus, including the structures of the internal sac.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFA6536BCD245516FD42A159.taxon	distribution	Distribution. Atheta prudhoensis seems to be widespread in the northern Nearctic Region but its distribution is poorly documented. It is known from Alaska, Yukon Territory, Vermont and New York.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFA75365CD2450E4FEA1A721.taxon	description	(Figs. 3, 32 in Lohse & Smetana 1985; Figs. 63 – 66 in Klimaszewski & Winchester 2002)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFA75365CD2450E4FEA1A721.taxon	materials_examined	Type material. Lectotype of Homalota picipennis (designated by Lohse and Smetana (1985 )): UNITED STATES: Alaska: ♂, Sitka (MZHF). Lectotype of Atheta carlottae (here designated): ♂, “ QCI [Queen Charlotte Islands] ”, “ Atheta carlottae Csy. ”, “ TYPE USNM 39247 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 2 ♂♂, 2 ♀♀, “ QCI [Queen Charlotte Islands] ”, “ carlottae PARATYPE USNM 39247 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta aperta (here designated): ♂, “ Cal. [ifornia, Humboldt Co., Arcata] ”, “ Atheta aperta Csy. ”, “ TYPE USNM 39250 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotype: ♀, “ Cal. [ifornia, Humboldt Co., Arcata] ”, “ aperta ­ 2 PARATYPE USNM 39250 ” (red label), “ CASEY bequest 1925 ” (NMNH). Holotype of Atheta wrangelica: ♀, “ Ft. Wrangel [Wrangell], Alaska. Wickham ”, “ ♀ ”, “ wrangelica Csy. ”, “ TYPE USNM 39248 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta morbosa (here designated): ♀, “ Cal. [ifornia, Saucelito near San Francisco] ”, “ Atheta morbosa Csy. ”, “ TYPE USNM 39293 ” (red label), “ CASEY bequest 1925 ” (NMNH). Holotype of Atheta intacta: ♀, “ Cal. [ifornia, Marin Co., Saucelito] ”, “ intacta Csy. ”, “ TYPE USNM 39299 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta alaskana (here designated): ♀, “ Alask. [Alaska, “ without further indication, but probably collected on the mainland near Fort Wrangel ” (Casey 1910 a)] Wick. [ham] ”, “ alaskana Csy. ”, “ TYPE USNM 39281 ” (red label), “ CASEY bequest 1925 ” (NMNH). Additional material: CANADA: British Columbia: ♂, Upper Carmanah Valley, 31. vii – 11. viii. 1991 (N. Winchester); ♀, ditto but 28. viii – 9. ix. 1991 (SPSU); UNITED STATES: California: San Francisco Co.: ♀ (NMNH (Casey collection )).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFA75365CD2450E4FEA1A721.taxon	diagnosis	Diagnosis. See Lohse & Smetana (1985); Klimaszewski & Winchester (2002).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFA75365CD2450E4FEA1A721.taxon	discussion	Discussion. The name Atheta picipennis (Mannerheim, 1843) (ex Homalota) is a secondary junior homonym of the name Atheta picipennis (Stephens, 1832) (ex Aleochara) which is itself a primary junior homonym of the name Aleochara picipennis Gyllenhal, 1827. Atheta picipennis (Stephens, 1832) has been considered a synonym of Atheta amicula (Stephens, 1832) since at least 1868 (Gemminger & Harold 1868) and to the best of my knowledge has not been cited as valid after 1899 (Article 23.9.1.1). In the last 50 years at least 31 works by at least 42 authors have been published (Hatch 1957; Gensicke 1960; Hansen et. al. 1960; Strand & Vik 1964; Koch 1968; Scheerpeltz 1968; Easton 1970; Palm 1970; Blackwelder 1973; Bordoni 1973; Tikhomirova 1973; Benick & Lohse 1974; Moore & Legner 1975; Yosii & Sawada 1976; Hanski & Koskela 1977; Pope 1977; Hanski & Koskela 1978; Muona 1979 b; Burakowski et al. 1981; Uhlig & Vogel 1981; Benick 1982; Sawada 1982; Muona 1984; Lohse 1985; Segers 1986; Kahlen 1987; Muona 1987; Silfverberg 1988; Alexandrovitch et al. 1996; Derunkov & Melke 2001; Klimaszewski & Winchester 2002) listing Atheta picipennis as valid (Article 23.9.1.2). Atheta picipennis (Mannerheim, 1843) is a nomen protectum and Atheta picipennis (Stephens, 1832) is a nomen oblitum (Article 23.9.2). Atheta picipennis was believed to have a Holarctic distribution (e. g., Bernhauer & Scheerpeltz 1926) until Lohse and Smetana (1985) demonstrated that the Palaearctic specimens belonged to a separate species, At. aeneipennis (Thomson, 1856). Females of At. picipennis and At. altaica Bernhauer, 1901 have very similar spermathecae. Klimaszewski and Winchester (2002) were unable to find any difference between females of the two species. Although in average At. picipennis has a larger body than At. altaica, females of these two species can be reliably identified only by association with males. In comparison to At. altaica, males of At. picipennis have a narrower apex of the median lobe of the aedeagus.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFA75365CD2450E4FEA1A721.taxon	distribution	Distribution. Atheta picipennis and At. altaica appear to be allopatric: Atheta picipennis is known from southeast Alaska (Sitka, Wrangell), coastal British Columbia and California, while At. altaica is known (in North America) from the Kenai Peninsula (Alaska), central Alaska, Yukon Territory, Northwest Territories and Labrador (Lohse et al. 1990).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFA95366CD24563EFE8CA669.taxon	description	(Fig. 160 in Strand & Vik 1964; Figs. 164 – 167 in Lohse et al. 1990 (as Dimetrota ))	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFA95366CD24563EFE8CA669.taxon	materials_examined	Type material. Lectotype of Atheta altaica (here designated): ♂, “ Altaigebirge Mittelsibirien ”, orange square, “ altaica Brnh. leg. Leder. Reitter. Type ” (yellow label), white square, “ Chicago NHMus M. Bernhauer Collection ” (FMNH). Paralectotype: ♂, white square, “ Altaigebirge leg. Leder ”, “ altaica Brh. Type. det. Reitter ” (yellow label), “ Chicago NHMus M. Bernhauer Collection ” (FMNH). Additional material: UNITED STATES: Alaska: 2 ♂♂, Kenai Peninsula, Anchor River at Hwy. 1, 450 ’, 4. vi. 1978 (Smetana & Becker); ♂, Kenai Peninsula, Clam Gulch, 6. vi. 1978 (Smetana & Becker); 2 ♀♀, Kenai Mts., 2 mi. S Moose Pass, 500 ’, 30. v. 1978 (A. Smetana & E. C. Becker); 18 specimens, Denali Hwy. Mi. 110, Seattle Creek, 15. vii. 1978 (J. M. Campbell & A. Smetana); ♂, ♀, 12 mi. N Denali Hwy. Mi. 78, Windy Creek, 13. vii. 1978 (Smetana & Campbell); ♀, Richardson Hwy. Mi. 206, Isabel Pass, 2900 ’, 16. vii. 1962 (R. E. Leech); ♀, Prudhoe Bay Rd., S Fork Koyukuk River, 67 º 12 ’ N 150 º 07 ’ W, 1100 ’, 8. vii. 1978 (Campbell & Smetana) (CNCI); CANADA: Northwest Territories: ♀, East Channel, MacKenzie Delta, 6. vii. 1973 (CNCI); Yukon Territory: ♂, Dawson City, 16. vii. 1968 (Campbell & Smetana); ♀, Dempster Hwy. Mi. 48.5, North Fork Pass, 4100 ’, 19. vii. 1978 (J. M. Campbell & A. Smetana); 2 ♂♂, ♀, Dempster Hwy. Mi. 45, 3400 ’, 18 – 23. vii. 1978 (J. M. Campbell); 2 ♂♂, 3 ♀♀, Dempster Hwy. Mi. 42, North Klondike River, 3300 ’, 18. vii. 1978 (Smetana & Campbell); 2 ♂♂, ♀, Mi. 1034 Alaska Hwy., Nr. Kloo Lake, 5. vii. 1968 (Campbell & Smetana) (CNCI).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFA95366CD24563EFE8CA669.taxon	diagnosis	Diagnosis. See Lohse et al. (1990; as Dimetrota).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFA95366CD24563EFE8CA669.taxon	discussion	Discussion. To clarify the status of Atheta altaica and At. picipennis, the types of both species were examined. The types of At. altaica correspond to the accepted concept of this species (e. g., Strand & Vik 1964; Lohse et al. 1990).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFA95366CD24563EFE8CA669.taxon	distribution	Distribution. Atheta altaica has a circumpolar distribution. In North America it is known from Alaska, Yukon Territory, Northwest Territories and Labrador (Lohse et al. 1990).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFAA5360CD2457E6FAF5A571.taxon	description	(Figs. 13 – 14 in Lohse & Smetana 1985; Figs. 71 – 74 in Klimaszewski & Winchester 2002)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFAA5360CD2457E6FAF5A571.taxon	materials_examined	Type material. Lectotype of Homalota moesta (designated by Lohse and Smetana (1985 )): UNITED STATES: Alaska: ♂, Sitka (MZHF). Lectotype of Atheta hampshirensis (here designated): ♂, “ N. [ew] H. [ampshire], Fall ”, “ 7603 ”, “ hampshirensis Bernh. Typus. Fenyes! ” (yellow label), “ Chicago NHMus M. Bernhauer Collection ” (FMNH). Lectotype of Atheta leviceps (here designated): ♀, “ N. [ew] Y. [ork] [Catskill Mts.] ”, “ leviceps Csy. ”, “ TYPE USNM 39362 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 2 ♂♂, ♀, “ N. Y. [Catskill Mts.] ”, “ leviceps PARATYPE USNM 39362 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Dimetrota sectator (here designated): ♂, “ Cal. [ifornia] [Gilroy Hot Springs, Santa Clara Co.] ”, “ sectator Csy. ”, “ TYPE USNM 39118 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 2 ♂♂, 2 ♀♀, specimens, “ Cal. [ifornia] [Gilroy Hot Springs, Santa Clara Co.] ”, “ sectator PARATYPE USNM 39118 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Dimetrota retrusa (here designated): ♂, “ Metlakatla, B. [ritish] Col. [umbia] ”, “ retrusa Csy. ”, “ TYPE USNM 39120 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: ♂, 5 ♀♀, “ Metlakatla, B. [ritish] Col. [umbia] ”, “ retrusa PARATYPE USNM 39120 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Datomicra hebescens (here designated): ♂, “ N. [ew] Y. [ork] [Catskill Mts.] ”, “ hebescens Csy. ”, “ TYPE USNM 39176 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 8 specimens, “ N. [ew] Y. [ork] [Catskill Mts.] ”, “ hebescens PARATYPE USNM 39176 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Datomicra insolida (here designated): ♂, “ Cal. [ifornia] [Gilroy Hot Springs, Santa Clara Co.] ”, “ insolida Csy. ”, “ TYPE USNM 39119 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 2 specimens, “ Cal. [ifornia] [Gilroy Hot Springs, Santa Clara Co.] ”, 7 specimens (1 without head and prothorax), “ Cal. [ifornia] [San Francisco] ”, “ insolida PARATYPE USNM 39119 ” (red label), “ CASEY bequest 1925 ” (NMNH). Holotype of Datomicra pellax: ♀, “ R. [hode] I. [sland] [Boston Neck] ”, “ pellax Csy. ”, “ TYPE USNM 39171 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Pseudota vana (here designated): ♂, “ Q. [ueen] C. [harlotte] I. [slands] [Massett (Keen)] ”, “ vana Csy. ”, “ TYPE USNM 39157 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotype: ♀, “ Q. [ueen] C. [harlotte] I. [slands] [Massett (Keen)] ”, “ vana PARATYPE USNM 39157 ” (red label), “ CASEY bequest 1925 ” (NMNH). Additional material: UNITED STATES: Alaska: ♀, Kenai (Holmberg) (MZHF); New York: Orange Co.: 4 specimens, 8 km S New Windsor, Black Rock Forest, 23 – 25. v. 1998 (V. I. Gusarov) (SPSU); North Carolina: Haywood Co.: 13 specimens, Cold Springs Creek, Forest Road 148, 29 km N Waynesville, Pisgah National Forest, 35 º 44.6 ' N 83 º 00.3 ' W, 750 m, 4. vi. 2001 (V. I. Gusarov) (SPSU); Pennsylvania: ♀, 41 º 08.39 ' N 75 º 32.24 ' W, in mushrooms, 16. viii. 1998 (V. I. Gusarov) (SPSU); CANADA: Québec: ♂, St. ­ Jacques­de­Leeds, 46 º 16 ' N 71 º 23 ' W, maple forest, 8. vii. 1994 (SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFAA5360CD2457E6FAF5A571.taxon	diagnosis	Diagnosis. See Lohse & Smetana (1985).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFAA5360CD2457E6FAF5A571.taxon	discussion	Discussion. The types of Homalota moesta, At. hampshirensis, At. leviceps, Dim. sectator, Dim. retrusa, Dat. hebescens, Dat. insolida, Dat. pellax and Pseudota vana are similar in external characters and the shape of the aedeagus and spermatheca.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFAA5360CD2457E6FAF5A571.taxon	distribution	Distribution. Atheta hampshirensis is widespread in the United States (Alaska, Washington (Lohse & Smetana 1985), Oregon, California, New Hampshire, Rhode Island, New York, Pennsylvania and North Carolina) and Canada (British Columbia and Québec).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFAD5361CD24524EFB07A1D4.taxon	description	(Figs. 67 – 70 in Klimaszewski & Winchester 2002)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFAD5361CD24524EFB07A1D4.taxon	materials_examined	Type material. Lectotype of Atheta brumalis (here designated): ♂, “ Metlakatla, B. [ritish] Col. [umbia] Keen ”, “ Atheta brumalis Csy. ”, “ TYPE USNM 39246 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 14 specimens, “ Metlakatla, B. [ritish] Col. [umbia] Keen ”, “ brumalis PARATYPE USNM 39246 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta repexa (here designated): ♀, “ Mts. Near Claremont Cal. Baker ”, “ repexa Csy. ”, “ TYPE USNM 39297 ” (red label), “ CASEY bequest 1925 ” (NMNH).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFAD5361CD24524EFB07A1D4.taxon	diagnosis	Diagnosis. See Klimaszewski & Winchester (2002).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFAD5361CD24524EFB07A1D4.taxon	discussion	Discussion. The lectotype of At. repexa is similar to the types of At. brumalis in both external characters and the distinct shape of the spermatheca.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFAD5361CD24524EFB07A1D4.taxon	distribution	Distribution. Atheta brumalis is known from British Columbia and California.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFAD537CCD24576CFC1CA569.taxon	description	(Figs. 47 – 55)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFAD537CCD24576CFC1CA569.taxon	materials_examined	Type material. Lectotype of Atheta fenyesi (here designated): ♂, “ Pasadena Cal. Dr. A. Fenyes ”, “ Dec. [ember] ”, “ 781. ”, “ ♂ ”, “ 107. ”, “ fenyesi Brh. Typus ” (yellow label), “ Chicago NHMus M. Bernhauer Collection ” (FMNH). Paralectotypes: ♀, “ Pasadena Cal. [ifornia] Dr. A. Fenyes ”, “ 1. ”, “ fenyesi Brh. Cotyp. ” (yellow label), “ Chicago NHMus M. Bernhauer Collection ”; ♂, “ Pasadena Cal. [ifornia] Dr. A. Fenyes ”, “ 1. ”, “ fenyesi Brh. Cotypus. ” (yellow label), “ Chicago NHMus M. Bernhauer Collection ”; ♀, “ Pasadena Cal. [ifornia] Dr. A. Fenyes ”, “ Dec. [ember] ”, “ 801. ”, “ ♀ ”, “ fenyesi Brh. Cotypus. ” (yellow label), “ Chicago NHMus M. Bernhauer Collection ” (FMNH). Lectotype of Atheta querula (here designated): ♂, “ Calaveras Co. Cal. [ifornia] ”, “ Atheta querula Csy. ”, “ TYPE USNM 39263 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Dimetrota resima (here designated): ♂, “ San Mateo Cal. [ifornia] Ba [ker] ”, “ resima Csy. ”, “ TYPE USNM 39113 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: ♂ (with the head and prothorax missing), “ San Mateo Cal. [ifornia] Bake [r] ”, “ resima ­ 2 PARATYPE USNM 39113 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Dimetrota vigilans (here designated): ♂, “ Pom. [ona, Los Angeles Co.] Cal. [ifornia] 2 / 2 / 96 [2. ii. 1896 (Fall)] ”, “ vigilans Csy. ”, “ TYPE USNM 39121 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta socors (here designated): ♀, “ Cal. [ifornia, San Diego] ”, “ socors Csy. ”, “ TYPE USNM 39296 ” (red label), “ CASEY bequest 1925 ” (NMNH). Holotype of Dimetrota immerita: ♂, “ Riv. [erside] Cal. [ifornia] 3 ­ 20 [?] ­ 90 [20. iii. 1890] ”, “ immerita Csy. ”, “ TYPE USNM 39126 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Dimetrota incredula (here designated): ♂, “ Cal. [ifornia, Saucelito, Marin Co.] ”, “ incredula Csy. ”, “ TYPE USNM 39128 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Dimetrota opinata (here designated): ♂, “ Cal. [ifornia, Berkeley, Alameda Co.] ”, “ opinata Csy. ”, “ TYPE USNM 39129 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Dimetrota cerebrosa (here designated): ♂, “ Cal. [ifornia, Saucelito, Marin Co.] ”, “ cerebrosa Csy. ”, “ TYPE USNM 39127 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotype: ♀, “ Cal. [ifornia, Monterey] ”, “ CASEY determ.? cerebrosa ­ 2 ”, “ CASEY bequest 1925 ” (NMNH).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFAD537CCD24576CFC1CA569.taxon	diagnosis	Diagnosis. Atheta fenyesi can be recognized among the other species of the subgenus Dimetrota auct. by the presence of four projections at the posterior margin of the male tergum 8 (Figs. 47, 49), by the distinct shape of the median lobe (Figs. 52 – 54) (particularly the fused and expanded ostial lamellae forming a wide rhomboid sclerite on the abparameral side of the median lobe (Fig. 54 )) and the distinct shape of the spermatheca (Fig. 55).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFAD537CCD24576CFC1CA569.taxon	discussion	Discussion. The types of At. fenyesi, At. querula, Dim. resima, Dim. vigilans, At. socors, Dim. immerita, Dim. incredula, Dim. opinata and Dim. cerebrosa are similar in external characters and in the shape of the aedeagus and spermatheca. All these names are considered to be synonyms.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFAD537CCD24576CFC1CA569.taxon	distribution	Distribution. Atheta fenyesi is known from California.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFB15379CD24524EFB7BA4E7.taxon	description	(Figs. 56 – 69)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFB15379CD24524EFB7BA4E7.taxon	materials_examined	Type material. Lectotype of Homalota modesta (here designated): ♂, “ U. [nited] S. [tates] ”, “ modesta Melsh. ”, “ Melsh. [eimer] ”, “ modesta ”, a red piece of paper, “ MCZ TYPE 34936 ” (red label) (MCZ) (Two paralectotypes of H. modesta (with labels “ modesta ”, “ Melsh. ”, “ MCZ TYPE 34936 ”) are not conspecific with the lectotype and belong to Stethusa spuriella (Casey, 1910 a) (MCZ )). Lectotype of Atheta crassicornis var. virginica (here designated): ♂, “ White Sulphur W. [est] Va. [Virginia] ”, “ Aug. [ust] ”, yellow circle, “ pacifica Bh. Fenyes det. Bernhauer ”, “ v. virginica Brh. Typus ” (yellow label), “ Chicago NHMus M. Bernhauer Collection ” (FMNH) (The paralectotype of this species (♂, “ E. Machias, Me. [Maine] ”, “ June ”, white circle, “ crassicornis Gvh. Fenyes det. Bernhauer ”, “ v. virginica Brh. Cotypus ” (yellow label), “ Chicago NHMus M. Bernhauer Collection ” (FMNH )) is not conspecific with the lectotype. It belongs to At. crenuliventris Bernhauer, 1907). Lectotype of Atheta rhodeana (here designated): ♂, “ R. [hode] I. [sland, Boston Neck] ”, “ rhodeana ­ 2 PARATYPE USNM 39215 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: ♂, “ R. [hode] I. [sland, Boston Neck] ”, “ ♂ ”, “ Atheta rhodeana Csy. ”, “ TYPE USNM 39215 ” (red label), “ CASEY bequest 1925 ” (NMNH); 6 ♀♀, “ R. [hode] I. [sland, Boston Neck] ”, “ rhodeana PARATYPE USNM 39215 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta capella (here designated): ♂, “ R. [hode] I. [sland, Boston Neck] ”, “ capella ­ 2 PARATYPE USNM 39216 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: ♂ (with the aedeagus missing), “ R. [hode] I. [sland, Boston Neck] ”, “ Atheta capella Csy. ”, “ TYPE USNM 39216 ” (red label), “ CASEY bequest 1925 ” (NMNH); 2 ♂♂, ♀, “ R. [hode] I. [sland, Boston Neck] ”, “ capella PARATYPE USNM 39216 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta ducens (here designated): ♂, “ N. [ew] Y. [ork, Catskill Mts.] ”, “ ducens Csy. ”, “ TYPE USNM 39218 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 17 specimens, “ N. [ew] Y. [ork, Catskill Mts.] ”, “ ducens PARATYPE USNM 39218 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta temperans (here designated): ♀, “ D. [istrict of] C. [olumbia] ”, “ Atheta temperans Csy. ”, “ TYPE USNM 39225 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta logica (here designated): ♀, “ Penn. [sylvania, near Philadelphia] ”, “ logica Csy. ”, “ TYPE USNM 39489 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotype: ♀, “ Penn. [sylvania, near Philadelphia] ”, “ logica PARATYPE USNM 39489 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta tradita (here designated): ♀, “ N. [ew] Y. [ork, Catskill Mts.] ”, “ tradita Csy. ”, “ TYPE USNM 39253 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotype: ♂, “ N. [ew] Y. [ork, Catskill Mts.] ”, “ tradita PARATYPE USNM 39253 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta fenisex (here designated): ♀, “ R. [hode] I. [sland, Boston Neck] ”, “ fenisex Csy. ”, “ TYPE USNM 39256 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta vierecki (here designated): ♂, “ Double Beach, Ct. [Connecticut] 19 Aug. 1905 H. L. Viereck ”, “ Fungus ”, “ vierecki Csy. ”, “ TYPE USNM 39255 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 2 ♂♂, ♀, “ Double Beach, Ct. [Connecticut] 19 Aug. 1905 H. L. Viereck ”, “ Fungus ”, “ vierecki PARATYPE USNM 39255 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta auguralis (here designated): ♂, “ N. [ew] Y. [ork, Catskill Mts.] ”, “ auguralis Csy. ”, “ TYPE USNM 39257 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta bifaria (here designated): ♀, “ D. [istrict of] C. [olumbia] ”, “ bifaria Csy. ”, “ TYPE USNM 39258 ” (red label), “ CASEY bequest 1925 ” (NMNH). Additional material. UNITED STATES: Michigan: Newaygo Co.: ♂, 2 ♀♀, 16 km SEE White Cloud, Newaygo State Park, 43 º 29.84 ' N 85 º 34.85 ' W, 150 m, in forest litter, Pinus, Quercus, 16. vii. 1999 (V. I. Gusarov) (SPSU); New York: Orange Co.: ♀, 8 km S New Windsor, Black Rock Forest, 9. v. 1998 (V. I. Gusarov); ♀, ditto but 23 – 25. v. 1998; ♂, ditto but pitfall traps, vi. 1998 (V. Ovcharenko) (SPSU); Sullivan Co.: ♀, Jeffersonville, 11 – 12. ix. 1999 (A. Berkov) (SPSU); Pennsylvania: 3 ♂♂, 4 ♀♀, “ Penn. [sylvania, near Philadelphia] ”, (NMNH (Casey collection )); Vermont: Windsor Co.: 2 ♀♀, 8 km SEE Springfield, Hwy. I­ 91, 43 º 14.00 ' N 72 º 26.81 ' W, 250 m, in forest litter, Acer, Betula, Tsuga, Quercus, 21. ix. 1998 (V. I. Gusarov) (SPSU). CANADA: New Brunswick: Sunbury Co.: ♂, ♀, Acadia Research Forest, pitfall traps, 4. viii. 1999 (G. Gesner); ♀, ditto but 13. vii. 1999 (SPSU); Ontario: ♂, ♀, 22 km NNE Gananoque, env. of Charleston Lake Prov. Park, 44 º 30.03 ' N 76 º 03.96 ' W, 0 m, in forest litter, Acer, Ulmus, Quercus, Pinus, ferns, 15. vii. 1999 (V. I. Gusarov) (SPSU); Québec: ♂, Parc de la Gatineau, Lac Lapêche, 9 km SWW Sainte­François­de­Masham, 26. ix. 1998 (V. I. Gusarov); ♂, St. ­ Jaques­de­Leeds, 46 º 16 ' N 71 º 23 ' W, maple forest, 2. viii. 1993 (SPSU); 5 specimens, ditto but 15. vii. 1994 (LFC); ♀, ditto but 15. vii. 1994; ♀, ditto but 19. vii. 1994; ♂, 170 km NE Québec­City, Pohénégamook, 47 º 28 ' N 69 º 14 ' W, maple forest, 19. vii. 1994 (SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFB15379CD24524EFB7BA4E7.taxon	diagnosis	Diagnosis. Atheta modesta is closely related to the Palaearctic At. crassicornis (Fabricius, 1793) but differs in having the pronotal setation pattern of the type III, the distinct shape of the male tergum 8 (Fig. 56) and in the shape of the aedeagus (Figs. 60 – 68). The spermathecae of the two species are very similar (Fig. 69).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFB15379CD24524EFB7BA4E7.taxon	discussion	Discussion. Bernhauer (1907) described At. virginica as a variety of At. crassicornis. He noted the similarity between the examined North American specimens and the European specimens, but listed several differences between the two. He also noted that when additional Nearctic specimens become available they might prove to represent a distinct species and not a variety of At. crassicornis. The types of H. modesta, At. crassicornis var. virginica, At. rhodeana, At. capella, At. ducens, At. temperans, At. logica, At. tradita, At. fenisex, At. vierecki, At. auguralis and At. bifaria are similar in external characters and in the shape of the aedeagus and spermatheca. All these names are considered to be synonyms.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFB15379CD24524EFB7BA4E7.taxon	distribution	Distribution. Atheta modesta is widely distributed in the eastern United States (Michigan, Vermont, Rhode Island, Connecticut, New York, Pennsylvania, West Virginia and District of Columbia) and Canada (New Brunswick, Ontario and Québec).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFB65377CD245517FCBBA0A1.taxon	description	(Figs. 70 – 80)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFB65377CD245517FCBBA0A1.taxon	materials_examined	Type material. Lectotype of Atheta frosti (here designated): ♀, “ N. [ew] H. [ampshire] Fall ”, “ frosti Brh. Cotypus ”, “ 7597 ”, “ Chicago NHMus M. Bernhauer Collection ”. Paralectotypes: ♂, “ Frmghm. [Framingham] Mass. [achusetts] Frost ”, “ frosti Brh. Typus ” (yellow label), “ 7008 ”, “ Chicago NHMus M. Bernhauer Collection ” (FMNH); ♀, “ Penn. [sylvania] ”, “ Aug. [ust] [18] 96 ”, red circle, “ nigritula Amer. b. Gvh. ”, “ frosti Brh. Cotypus ” (yellow label), “ Chicago NHMus M. Bernhauer Collection ”; ♂ (with the head and the apex of the abdomen missing), “ Brooklyn, N. [ew] Y. [ork] ”, “ July ”, white circle, “ frosti Brh. Cotypus ” (yellow label), “ Chicago NHMus M. Bernhauer Collection ” (FMNH). Lectotype of Atheta comitata (here designated): ♂, “ R. [hode] I. [sland, Boston Neck] ”, “ Atheta comitata Csy. ”, “ TYPE USNM 39217 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 3 ♂♂, 5 ♀♀, “ R. [hode] I. [sland, Boston Neck] ”, “ comitata PARATYPE USNM 39217 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta gnoma (here designated): ♂, “ N. [ew] Y. [ork, Catskill Mts.] ”, “ Atheta gnoma Csy. ”, “ TYPE USNM 39234 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta elota (here designated): ♂, “ N. [ew] Y. [ork, Catskill Mts.] ”, “ Atheta elota Csy. ”, “ TYPE USNM 39219 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 4 ♂♂, 2 ♀♀, “ N. [ew] Y. [ork, Catskill Mts.] ”, “ elota PARATYPE USNM 39219 ” (red label), “ CASEY bequest 1925 ” (NMNH) (one additional paralectotype of Atheta elota (“ N. [ew] Y. [ork, Catskill Mts.] ”, “ elota – 5 PARATYPE USNM 39219 ” (red label), “ CASEY bequest 1925 ” (NMNH )) belongs to Atheta klagesi Bernhauer). Lectotype of Atheta insidiosa (here designated): ♂, “ N. [ew] Y. [ork, Catskill Mts.] ”, “ Atheta insidiosa Csy. ”, “ TYPE USNM 39220 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 7 ♂♂, 3 ♀♀, 1 specimen with the tip of the abdomen missing, “ N. [ew] Y. [ork, Catskill Mts.] ”, “ insidiosa PARATYPE USNM 39220 ” (red label), “ CASEY bequest 1925 ” (NMNH) (9 additional paralectotypes (“ N. [ew] Y. [ork, Catskill Mts.] ”, “ insidiosa PARATYPE USNM 39220 ” (red label), “ CASEY bequest 1925 ” (NMNH) belong to Atheta klagesi Bernhauer). Lectotype of Pseudota puricula (here designated): ♂, “ N. [ew] Y. [ork, Catskill Mts.] ”, “ puricula Csy. ”, “ TYPE USNM 39159 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotype: ♂, “ N. [ew] Y. [ork, Catskill Mts.] ”, “ puricula PARATYPE USNM 39159 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta candidula (here designated): ♀, “ N. [ew] Y. [ork, Catskill Mts.] ”, “ candidula Csy. ”, “ TYPE USNM 39232 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta diffisa (here designated): ♂, “ N. [ew] Y. [ork, Catskill Mts.] ”, “ diffisa Csy. ”, “ TYPE USNM 39233 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta nata (here designated): ♀, “ R. [hode] I. [sland, Boston Neck] ”, “ nata Csy. ”, “ TYPE USNM 39228 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta modiella (here designated): ♂, “ R. [hode] I. [sland, Boston Neck] ”, “ modicella [sic!] Csy. ”, “ TYPE USNM 39235 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta vacillans (here designated): ♀, “ R. [hode] I. [sland, Boston Neck] ”, “ vacillans Csy. ”, “ TYPE USNM 39222 ” (red label), “ CASEY bequest 1925 ” (NMNH). Additional material. UNITED STATES: North Carolina: Haywood Co.: ♂, 3 ♀♀, Cold Springs Creek, 29 km N Waynesville, 35 º 44.56 ' N 83 º 00.39 ' W, 650 m, 20. viii. 1998 (V. I. Gusarov); Macon Co.: 3 ♂♂, ♀, Nantahala River, 26 km NWW Franklin, 35 º 15.37 ' N 83 º 39.24 ' W, 650 m, 21. viii. 1998 (V. I. Gusarov) (SPSU); Pennsylvania: ♂, near Philadelphia, (NMNH (Casey collection )); Adams Co.: ♂, 3 ♀♀, 15 km E Chambersburg, Michaux State Forest, 39 º 54.87 ' N 77 º 28.33 ' W, 400 m, in mushrooms, 18. viii. 1998 (V. I. Gusarov) (SPSU); Vermont: Windsor Co.: ♀, 8 km SEE Springfield, Hwy. I­ 91, 43 º 14.00 ' N 72 º 26.81 ' W, 250 m, in forest litter, Acer, Betula, Tsuga, Quercus, 21. ix. 1998 (V. I. Gusarov) (SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFB65377CD245517FCBBA0A1.taxon	diagnosis	Diagnosis. Atheta frosti is closely related to Palaearctic At. nigritula (Gravenhorst, 1802) but differs in the shape of the male tergum 8 (Figs. 70, 72), the aedeagus (Figs. 75 – 79) and spermatheca (Fig. 80).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFB65377CD245517FCBBA0A1.taxon	discussion	Discussion. The types of At. frosti, At. comitata, At. gnoma, At. elota, At. insidiosa, Ps. puricula, At. candidula, At. diffisa, At. nata, At. modiella and At. vacillans are similar in external characters and in the shape of the aedeagus and spermatheca. All these names are considered to be synonyms.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFB65377CD245517FCBBA0A1.taxon	distribution	Distribution. Atheta frosti is widely distributed in the eastern United States (Vermont, New Hampshire, Massachusetts, Rhode Island, New York, Pennsylvania and North Carolina) but its distribution is poorly documented.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFBB5370CD2451AEFD71A641.taxon	description	(Fig. 117 in Strand & Vik, 1964; Fig. 7, p. 154 in Benick & Lohse 1974)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFBB5370CD2451AEFD71A641.taxon	materials_examined	Type material. Lectotype of Aleochara nigritula (here designated): ♀ (with the head and pronotum missing), “ var. c. A. nigritula Gr. ”, “ nigritula Gr. ” (ZMHB). Additional material. UKRAINE: Crimea: Simferopol’ distr.: 10 specimens, Krasnoles’ye, 400 m, in rotting wood and on mushrooms, 27. v. 1990 (V. I. Gusarov) (SPSU); Ivano­Frankovsk Reg.: 7 specimens, Nadvornaya, on polypore mushrooms, 6. viii. 1978 (S. Mosyakin) (SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFBB5370CD2451AEFD71A641.taxon	diagnosis	Diagnosis. See Benick & Lohse (1974).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFBB5370CD2451AEFD71A641.taxon	discussion	Discussion. The types of Ale. nigritula were examined to clarify the status of At. frosti and At. nigritula. The historic collection at ZMHB which contains the Gravenhorst types has several specimens arranged as Aleochara nigritula in the following order: the label at the bottom of the drawer “ var. c. A. nigritula Gr. ” is followed by 11 specimens: 1) ♀ (with the head and pronotum missing), labeled “ nigritula Gr. ”; 2) ♀, without labels; 3) ♀, without labels; 4) ♂, without labels; 5) ♀, without labels; 6) a specimen labeled “ fungicola Kraatz ”; 7) a specimen without labels; 8) a specimen labeled “ var. d ”; 9) a specimen without labels; 10) a specimen labeled “ Sard. Géné ”; 11) a specimen labeled “ Ural Er. ”. These 11 specimens complete the row. The next row starts with a label at the bottom of the drawer “ var.? ” which is followed by 7 specimens: 12 – 15) four unlabeled specimens, 16) a specimen labeled “ Aust. … [illegible] ”; 17 – 18) two specimens mounted on the same card labeled “ Pallau H. socialis var. nigra Gve. ”. Apparently, the specimens listed above had been arranged by Erichson, who described Homalota socialis Erichson, 1839 and listed Aleochara nigritula Gravenhorst as “ Var. c ” of H. socialis. Therefore only the specimens (Nos. 1 – 7) which follow the label “ var. c ” qualify as potential syntypes of Aleochara nigritula. My examination of these seven specimens demonstrated that only specimen No. 1 belongs to At. nigritula in the commonly used sense (e. g., Strand & Vik 1964; Benick & Lohse 1974); specimens No. 2 and No. 5 belong to At. pallidicornis (Thomson, 1856); specimen No. 3 belongs to Dalotia coriaria (Kraatz, 1856); specimen No. 4 belongs to At. gagatina (Baudi, 1848); specimen No. 6 is not conspecific with At. nigritula and has the pronotal pubescence of type I; specimen No. 7 is not conspecific with At. nigritula and has the pronotal pubescence of type II. Specimen No. 1 was selected as the lectotype of Aleochara nigritula. The spermatheca of that specimen matches the illustrations by Strand and Vik (1964: Fig. 117) and Benick and Lohse (1974, p. 154: Fig. 7). Atheta nigritula (Gravenhorst, 1802) is the types species of the subgenus Tetropla Mulsant & Rey, 1874 a (by subsequent designation, fixed by Blackwelder (1952 )).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFBB5370CD2451AEFD71A641.taxon	distribution	Distribution. Atheta nigritula is a common European species associated with fungi. It has been reported from North America (Bernhauer 1907; Bernhauer & Scheerpeltz 1926) but, to the best of my knowledge, does not occur there. The examined North American specimens formerly identified as At. nigritula turned out to be At. frosti Bernhauer, 1909, a closely related but separate species.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFBC5373CD24578EFD34A558.taxon	description	(Figs. 81 – 92)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFBC5373CD24578EFD34A558.taxon	materials_examined	Type material. Lectotype of Atheta ventricosa (here designated): ♀, “ Mt. Arlngtn [Mount Arlington] N. [ew] J. [ersey] ”, “ Sept. ”, white circle, “ pedicularis Melsh.? v. Fenyes det. Bernhauer ”, “ ventricosa Brh. Typus ”, “ Chicago NHMus M. Bernhauer Collection ”. Paralectotypes: ♀, “ Ithaca, N. [ew] Y. [ork] Wickham ”, “ 129. ”, “ ventricosa Brh. Cotypus ” (yellow label), “ Chicago NHMus M. Bernhauer Collection ”; ♀ (had been mounted on the same card as the lectotype), “ Mt. Arlngtn [Arlington] N. [ew] J. [ersey] ”, “ Sept. ”, white circle, “ pedicularis Melsh.? v. Fenyes det. Bernhauer ”, “ ventricosa Brh. Typus ”, “ Chicago NHMus M. Bernhauer Collection ”; ♂, “ Blue Ridge Mts. 9 ­ 04 [ix. 1904?] N. [orth] C. [arolina] ”, “ Beyer ”, “ ventricosa Brh. Cotypus ” (yellow label), “ Chicago NHMus M. Bernhauer Collection ”; 1 destroyed specimen with only metatrochanters remaining, “ Blue Ridge Mts. 9 ­ 04 [ix. 1904?] N. [orth] C. [arolina] ”, “ Beyer ”, “ 128 ”, “ ventricosa Brh. Cotypus ” (yellow label), “ Chicago NHMus M. Bernhauer Collection ” (FMNH). Lectotype of Atheta mollicula (here designated): ♂, “ Metlakatla B. [ritish] Col. [umbia] Keen ”, “ Atheta mollicula Csy. ”, “ TYPE USNM 39266 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 2 ♀♀, “ Metlakatla B. [ritish] Col. [umbia] Keen ”, “ mollicula PARATYPE USNM 39266 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Sableta phrenetica (here designated): ♂, “ Metlakatla B. [ritish] Col. [umbia] Keen ”, “ phrenetica Csy. ”, “ TYPE USNM 39140 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta callens (here designated): ♀, “ D. [istrict of] C. [olumbia] ”, “ callens Csy. ”, “ TYPE USNM 39491 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta franklini (here designated): ♀, “ Buena Vista Spg. [Springs] Franklin Co. Pa. [Pennsylvania] ”, “ Ath. franklini Csy. ”, “ TYPE USNM 39314 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 3 ♀♀, “ Buena Vista Spg. [Springs] Franklin Co. Pa. [Pennsylvania] ”, “ franklini PARATYPE USNM 39314 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta postulans (here designated): ♀, “ Metlakatla B. [ritish] Col. [umbia] Keen ”, “ postulans Csy. ”, “ TYPE USNM 39307 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 2 ♀♀, “ Metlakatla B. [ritish] Col. [umbia] Keen ”, “ postulans PARATYPE USNM 39307 ” (red label), “ CASEY bequest 1925 ” (NMNH). Additional material. UNITED STATES: Alaska: ♀, 16 km NW Lituya Bay, pitfalls in hemlock forest, 15 m, vi­ix [. 1977] (D. Mann); 3 specimens, 10 km SE Lituya Bay, pitfalls in hemlock forest, 35 m, vi­ 4. ix. 1977 (D. Mann); 7 specimens, 16 km NW of Lituya Bay, pitfalls in edge of spruce forest, 10 m, vi­ 4. ix. 1977 (D. Mann); ♂, Baranof Is., 5 mi. ESE of Sitka, 19. vii. 1980 (L. Herman); 3 ♂♂, Baranof Island, NNE of Sitka, Indian River, 20. vii. 1980 (L. Herman) (AMNH); New York: Orange Co.: ♂, ♀, 5 specimens, 8 km S New Windsor, Black Rock Forest, 23 – 25. v. 1998 (V. I. Gusarov) (SPSU); Vermont: Windsor Co.: ♂, ♀, 3 specimens, 8 km SEE Springfield, Hwy. I­ 91, 43 º 14.00 ' N 72 º 26.81 ' W, 250 m, in forest litter, Acer, Betula, Tsuga, Quercus, 21. ix. 1998 (V. I. Gusarov) (SPSU); CANADA: New Brunswick: Sunbury Co.: ♂, Acadia Research Forest, pitfall trap, 28. vii. 1999 (G. Gesner); ♀, ditto but 21. vii. 1999 (SPSU); Ontario: ♂, 74 km NWW Sault Ste. Marie, Hwy. 17, slope facing Lake Superior, 47 º 09.61 ' N 84 º 41.90 ' W, 80 m, in forest litter, Betula, Picea, Populus, Acer, Pinus, 11. vii. 1999 (V. I. Gusarov) (SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFBC5373CD24578EFD34A558.taxon	diagnosis	Diagnosis. Atheta ventricosa is very similar to At. sodalis (Erichson, 1837) but differs in the shape of the aedeagus (Figs. 85 – 88, 90 – 92) (particularly the apex of the median lobe (Figs. 85 – 86) and the sclerites of the internal sac (Figs. 90 – 92 )), and the posterior margin of the male tergum 8 (Fig. 81).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFBC5373CD24578EFD34A558.taxon	discussion	Discussion. The types of At. ventricosa, At. mollicula, Sa. phrenetica, At. callens, At. franklini and At. postulans are similar in external characters and the shape of the aedeagus and spermatheca. All these names are considered to be synonyms.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFBC5373CD24578EFD34A558.taxon	distribution	Distribution. Atheta ventricosa is a widespread species but its distribution is poorly documented. It is known from the United States (Alaska, Vermont, New York, New Jersey, Pennsylvania, District of Columbia and North Carolina) and Canada (British Columbia, Ontario and New Brunswick).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFC0530DCD2457E4FC49A189.taxon	description	(Fig. 68 in Strand & Vik 1964; Fig. 2, p. 175 in Benick & Lohse 1974)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFC0530DCD2457E4FC49A189.taxon	materials_examined	Type material. Lectotype of Homalota sodalis (here designated): ♂ (the specimen No. 3 among the three potential syntypes of H. sodalis; see Discussion) (ZMHB). Additional material. GERMANY: ♂, env. of Bayreuth, 30. ix. 1992; ♀, Bavaria, Mittelfranken, env. of Feuchtwangen, vi. 1991 (Th. Blick) (SPSU); RUSSIA: Samara Reg.: 2 specimens, Zhiguli Nature Reserve, 25. vii – 15. viii. 1988 (Krasnobayev) (SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFC0530DCD2457E4FC49A189.taxon	diagnosis	Diagnosis. See Benick & Lohse (1974).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFC0530DCD2457E4FC49A189.taxon	discussion	Discussion. The types of Homalota sodalis were examined to clarify the status of At. ventricosa and At. sodalis. The historic collection at ZMHB which contains the Erichson types has 5 specimens arranged as Homalota sodalis in the following order: 1) ♀, labeled “ sodalis Er. Berol. [Berlin] Er. [Erichson] ”, “ 5411 ”; 2) ♂, without labels; 3) ♂, without labels; 4) ♂, labeled “ Aust. ”; 5) ♀, without labels. My examination of these five specimens demonstrated that only specimens Nos. 1 – 4 belonged to At. sodalis in the commonly used sense (e. g., Strand & Vik 1964; Benick & Lohse 1974). In specimen No. 5 the shape of the antennal articles is different and that specimen is not conspecific with At. sodalis. Specimen No. 4 is not a syntype because it was not collected in the type locality of At. sodalis (Mark Brandenburg). Specimen No. 3 was selected as the lectotype. It agrees with the commonly accepted concept of this species in external characters, including the shape of the tergum 8. The lectotype was not dissected. Specimens Nos. 1 and 2 were labeled as paralectotypes.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFC0530DCD2457E4FC49A189.taxon	distribution	Distribution. Atheta sodalis is a common European species. Although formerly reported from North America (Bernhauer & Scheerpeltz 1926), to the best of my knowledge, At. sodalis does not occur there. In North America At. sodalis is replaced by a closely related species, At. ventricosa Bernhauer, 1907.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFC15309CD245746FB0DA361.taxon	description	(Figs. 93 – 102)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFC15309CD245746FB0DA361.taxon	description	Lectotype of Atheta nympha (here designated): ♂, “ N. [ew] Y. [ork, Catskill Mts.] ”, “ Atheta nympha Csy. ”, “ TYPE USNM 39276 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 5 ♂♂, 3 ♀♀, “ N. [ew] Y. [ork, Catskill Mts.] ”, “ nympha PARATYPE USNM 39276 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta discreta (here designated): ♂, “ N. [ew] Y. [ork, Catskill Mts.] ”, “ Atheta discreta Csy. ”, “ TYPE USNM 39278 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Pseudota dissensa (here designated): ♂, “ Penn. [sylvania, Philadelphia] ”, “ Pseudota dissensa Csy. ”, “ TYPE USNM 39147 ” (red label), “ CASEY bequest 1925 ” (NMNH). Holotype of Atheta villica: ♀, “ Penn. [sylvania, near Philadelphia] ”, “ villica Csy. ”, “ TYPE USNM 39231 ” (red label), “ CASEY bequest 1925 ” (NMNH).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFC15309CD245746FB0DA361.taxon	materials_examined	Additional material. UNITED STATES: Minnesota: Lake Co.: ♂, 22 km NE Two Harbors, Split Rock Lighthouse State Park, Hwy. 61, 47 º 10.99 ' N 91 º 24.52 ' W, 70 m, in forest litter, Betula, Picea, Populus, Rubus, Majanthemum, ferns, 10. vii. 1999 (V. I. Gusarov) (SPSU); New Jersey: Warren Co.: 2 ♀♀, 24 km NW Hackettstown, Dunnfield Creek Nat. Area, 40 º 58.33 ' N 75 º 07.49 ' W, 20 m, Quercus, Acer, 5. iv. 1999 (V. I. Gusarov) (SPSU); New York: Orange Co.: 3 ♂♂, 2 ♀♀, 8 km S New Windsor, Black Rock Forest, 9. v. 1998 (V. I. Gusarov); 5 ♂♂, 2 ♀♀, ditto but 23 – 25. v. 1998 (SPSU); CANADA: Québec: ♂, Parc de la Gatineau, 11 km E Chelsea, Champlain, 26. ix. 1998 (V. I. Gusarov); ♂, Chûte­aux­Galets, 1. viii. 1994; ♂, 190 km NW Montréal, Mont­Laurier, 46 º 34 ' N 75 º 29 ' W, 5. vii. 1994 (SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFC15309CD245746FB0DA361.taxon	diagnosis	Diagnosis. Atheta klagesi can be recognized by the following combination of characters: head and abdominal terga 5 – 6 black; pronotum, abdominal terga 3 – 4 and 7 – 8, and antennal articles 4 – 11 or 3 – 11 brown; elytra yellow with vaguely defined darker areas around scutellum and in postero­lateral angles; antennal articles 1 – 2 or 1 – 3, legs and mouthparts yellow; antennal articles 5 – 10 strongly transverse, article 10 twice as wide as long; pronotal pubescence of type V (Benick & Lohse 1974); mesotibial macrosetae thin, as long as tibial width; posterior margin of male tergum 8 convex and crenulate (Figs. 93, 95); aedeagus as in Figs. 99 – 102; and spermatheca as in Fig. 98.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFC15309CD245746FB0DA361.taxon	discussion	Discussion. The types of At. klagesi, At. cephalina, At. nympha, At. discreta, Ps. dissensa and At. villica are similar in external characters and the shape of the aedeagus and spermatheca. All these names are considered to be synonyms. Atheta cephalina is the type species of the subgenus Delphota Casey, 1910 a by original designation. Pseudota dissensa is the type species of the genus Pseudota Casey, 1910 a by original designation (Casey first species rule). Casey did not specifically state in his description of Pseudota that Ps. dissensa was the type species of Pseudota; however, while designating the type species of Noverota Casey, 1910 a in the same paper, Casey stated (1910 a, p. 90): “ The first species may be regarded as the type, as in all other cases where the type is not specifically named ”. Since both At. cephalina and Ps. dissensa are synonyms of At. klagesi, the names Delphota and Pseudota are subjective synonyms of each other, irrespective of what subgenera in the genus Atheta are accepted as valid. The problem of subgeneric placement of At. klagesi within Atheta is outside the scope of the present paper. The matter will be addressed in a separate work when the generic revision of the tribe Athetini is completed.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFC15309CD245746FB0DA361.taxon	distribution	Distribution. Atheta klagesi is known from the eastern United States (Minnesota, Iowa, Pennsylvania, New Jersey, New York, Maine) and eastern Canada (Québec).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFC5530ACD2452FBFB48A571.taxon	description	(Figs. 50 – 55, 78 – 82 in Klimaszewski & Peck 1986)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFC5530ACD2452FBFB48A571.taxon	materials_examined	Type material. Lectotype of Atheta annexa (designated by Klimaszewski & Peck (1986 )): UNITED STATES: North Carolina: Buncombe Co.: ♂, Asheville (NMNH). Lectotype of Atheta citata (here designated): ♂, “ N. [ew] Y. [ork, Catskill Mts.] ”, “ Atheta citata Csy. ”, “ TYPE USNM 39275 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta evecta (here designated): ♂, “ Miss [issippi, Vicksburg] ”, “ Atheta evecta Csy. ”, “ TYPE USNM 39286 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotype: ♂, “ Vic. [ksburg] Miss [issippi] ”, “ evecta ­ 2 PARATYPE USNM 39286 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta propitia (here designated): ♀, “ · N. [orth] C. [arolina, Asheville] ”, “ propitia Csy. ”, “ TYPE USNM 39239 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta palpator (here designated): ♀, “ Cin. [cinnati] O. [hio (Dury)] ”, “ palpator Csy. ”, “ TYPE USNM 39240 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta burra (here designated): ♀, “ Bayfld [Bayfield], Wis [consin] Wickham. ”, “ burra Csy. ”, “ TYPE USNM 39241 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta nacta (here designated): ♀, “ Mo. [Missoury, St. Louis] ”, “ Fungus ”, “ nacta Csy. ”, “ TYPE USNM 39242 ” (red label), “ CASEY bequest 1925 ” (NMNH). Additional material. UNITED STATES: Illinois: Carroll Co.: ♀, Raccoon Den Cave, 27. v. 1992 (S. Taylor, J. Krejca & G. Gardner) (SPSU); Union Co.: ♂, Apis Annex [Cave], 8. v. 1992 (S. Taylor & J. Krejca) (SPSU); Kansas: Douglas Co.: 2 ♀♀, 18 km SSE Lawrence, Breidenthal Preserve, 38 º 48.50 ' N 95 º 11.52 ' W, 250 m, on mushrooms, 23. v. 2000 (V. I. Gusarov); 5 specimens, ditto but on rotting fruits of Asimina triloba, 1. x. 2000; Jefferson Co.: ♂, 12 km NNE Lawrence, 39 º 04.57 ' N 95 º 11.88 ' W, 250 m, in hay, 8. vi. 1999 (V. I. Gusarov) (SPSU); Louisiana: Natchitoches Par.: ♂, 1.5 mi. N Readhimer, 32 º 08 ' N 92 º 59 ' W, Geomys breviceps burrow, baited (gopher feces and malt extract) pitfall, 16. i – 4. ii. 2000 (P. Kovarik, A. Tishechkin, R. Turnbow) (SPSU); 3 specimens, ditto but 4. ii – 1. iii. 2000 (LSAM); Winn Par.: ♂, ♀, 2 mi. SE St. Maurice, 31 º 44 ' N 92 º 56 ' W, Geomys bursarius burrow, baited (malt and gopher feces) pitfall, 4. ii – 1. iii. 2000 (P. Kovarik, A. Tishechkin, R. Turnbow) (LSAM, SPSU); CANADA: Québec: ♀, Parc de la Gatineau, 11 km E Chelsea, Champlain, 26. ix. 1998 (V. I. Gusarov) (SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFC5530ACD2452FBFB48A571.taxon	diagnosis	Diagnosis. See Klimaszewski & Peck 1986.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFC5530ACD2452FBFB48A571.taxon	discussion	Discussion. Klimaszewski and Peck (1986) designated the lectotype of Atheta annexa but did not attach the lectotype label to the Casey type. In the Casey collection there is only one type of At. annexa. Apparently this is the type that was examined and designated as the lectotype by Klimaszewski and Peck. I attached the lectotype label to the specimen. Klimaszewski and Peck (1986) stated that At. annexa is practically indistinguishable from At. cephalina Casey, 1910 a and At. lymphatica Casey, 1911 a. My examination of the types did not confirm that opinion. Atheta cephalina is conspecific with At. klagesi Bernhauer, 1909, while At. lymphatica differs from At. annexa in details of its microsculpture and its body color.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFC5530ACD2452FBFB48A571.taxon	distribution	Distribution. Atheta annexa is widespread in the eastern United States (New York, Virginia, West Virginia, Kentucky, Tennessee, North Carolina, Georgia, Florida, Alabama, Mississippi, Louisiana, Missoury, Kansas, Iowa, Illinois, Wisconsin, Indiana and Ohio (see Klimaszewski & Peck 1986 )) and is also known from Canada (Québec).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFC7530BCD24524EFBB4A6D6.taxon	description	(Figs. 85 – 89 in Klimaszewski & Winchester 2002)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFC7530BCD24524EFBB4A6D6.taxon	materials_examined	Type material. Lectotype of Atheta concessa (here designated): ♂, “ Metlakatla, B. [ritish] Col. [umbia] Keen ”, “ concessa Csy. ”, “ TYPE USNM 39483 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 2 ♂♂, 2 ♀♀, “ Metlakatla, B. [ritish] Col. [umbia] Keen ”, “ concessa PARATYPE USNM 39483 ” (red label), “ CASEY bequest 1925 ” (NMNH). Holotype of Atheta sumpta: ♀, “ Metlakatla, B. [ritish] Col. [umbia] Keen ”, “ sumpta Csy. ”, “ TYPE USNM 39249 ” (red label), “ CASEY bequest 1925 ” (NMNH). (NMNH).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFC7530BCD24524EFBB4A6D6.taxon	diagnosis	Diagnosis. See Klimaszewski & Winchester 2002.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFC7530BCD24524EFBB4A6D6.taxon	discussion	Discussion. The holotype of At. sumpta is similar to the types of At. concessa in external characters. The spermatheca is identical in the female paralectotypes of At. concessa and the holotype of At. sumpta.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFC7530BCD24524EFBB4A6D6.taxon	distribution	Distribution. Atheta concessa is known only from British Columbia.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFC75306CD245614FC8FA0F9.taxon	description	(Figs. 103 – 106)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFC75306CD245614FC8FA0F9.taxon	materials_examined	Type material. Lectotype of Homalota festinans (here designated): ♀, “ 5430 ”, “ festinans Er. Pensylv. [Pennsylvania] Zimm. [ermann] ” (green label), “ Typus ” (red label), “ Zool. Mus. Berlin ”, “ Homalota festinans Er. [on red side of the label] / Lectotypus Lohse fix. 1983 [on white side of the label] ” [Lohse never published this lectotype designation] (ZMHB). Paralectotypes: 4 ♀♀, “ Pennsylvania Zimmermann Nr. 5430 ”, “ Typus ” (red label), “ Zool. Mus. Berlin ” (ZMHB). Lectotype of Atheta punctata (here designated): ♀, “ TYPE ” (red label), “ Posey Co. Ind. [iana] W. S. B. [Blatchley] 3. vi. 1909 ”, “ 3638 det. A. Fenyes ”, “ Purdue Blatchley collection ”, “ Atheta punctata sp. nov. ” (PURC). Lectotype of Synaptina merica (here designated): ♀, “ Ia. [Iowa, Keokuk] ”, “ Synaptina merica Csy. ”, “ TYPE USNM 39185 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: ♀, “ Iowa City, Ia. [Iowa] Wickham ”, “ merica ­ 2 PARATYPE USNM 39185 ” (red label), “ CASEY bequest 1925 ”; ♀, “ Ari. [zona, Williams] ” (with two black dots after “ i ”), “? merica ­ 3 (set apart slightly) PARATYPE USNM 39185 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Synaptina consonens (here designated): ♀, “ R. [hode] I. [sland, Boston Neck] ”, “ consonens Csy. ”, “ TYPE USNM 39186 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: ♀, “ R. [hode] I. [sland, Boston Neck] ”, “ consonens ­ 2 PARATYPE USNM 39186 ” (red label), “ CASEY bequest 1925 ”; ♀, “ N. [ew] Y. [ork, near New York City] ”, “ consonens ­ 3 PARATYPE USNM 39186 ” (red label), “ CASEY bequest 1925 ”; ♀, “ L. [ong Island] ”, “ consonens ­ 4 PARATYPE USNM 39186 ” (red label), “ CASEY bequest 1925 ”; 4 ♀♀, “ Ia. [Iowa, Cedar Rapids] ”, “ consonens PARATYPE USNM 39186 ” (red label), “ CASEY bequest 1925 ”; 4 ♀♀, “ Ia. [Iowa, Keokuk] ”, “ consonens PARATYPE USNM 39186 ” (red label), “ CASEY bequest 1925 ” (NMNH). Additional material. UNITED STATES: Connecticut: New Haven Co.: ♀, New Haven, 26. ii. 1910 (A. Champlain) (NMNH (Casey collection )); Kansas: Douglas Co.: 36 ♀♀, 1.5 km N Lawrence, right bank of the Kansas River, 38 º 58.96 ' N 95 º 14.62 ' W, 200 m, in flood refuse, 1. v. 1999 (V. I. Gusarov) (KSEM, SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFC75306CD245614FC8FA0F9.taxon	diagnosis	Diagnosis. Atheta festinans can be recognized by the black head and abdomen, the brown pronotum, elytra, antennae and mouthparts, yellow legs, strongly transverse antennal articles 8 – 10, the pronotal pubescence of type II (Benick & Lohse 1974), the mesotibial macroseta thin, short and poorly visible among microsetae; and the distinct shape of the spermatheca (Figs. 105 – 106).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFC75306CD245614FC8FA0F9.taxon	discussion	Discussion. The types of H. festinans, At. punctata, Sy. merica and Sy. consonens are similar in external characters and the shape of the spermatheca. All these names are considered to be synonyms. Synaptina merica is the type species of the genus Synaptina Casey, 1910 a by original designation (Casey first species rule). Casey did not specifically state in his description of Synaptina that Sy. merica was the type species of Synaptina; however, while designating the type species of Noverota Casey, 1910 a in the same paper, Casey stated (1910 a, p. 90): “ The first species may be regarded as the type, as in all other cases where the type is not specifically named ”. When describing the genus Synaptina, Casey (1910 a) did not explicitly state in what characters it differed from other athetine genera. In my opinion, all characters mentioned by Casey in the description of Synaptina fit the genus Atheta in the broad sense. The type species of Synaptina is similar to other species of Atheta with pronotal pubescence of type II. I consider Synaptina to be a synonym of Atheta at the rank of genus. Atheta festinans appears to be a parthenogenetic species. All 53 examined specimens are females.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFC75306CD245614FC8FA0F9.taxon	distribution	Distribution. Atheta festinans is known from Rhode Island, Connecticut, New York, Pennsylvania, Indiana, Iowa, Kansas and Arizona.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFCA5307CD245076FE49A361.taxon	description	(Figs. 183 – 185 in Lohse et al. 1990 (as Boreostiba hudsonica ))	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFCA5307CD245076FE49A361.taxon	materials_examined	Type material. Lectotype of Atheta parvipennis (here designated): ♂, “ Banff Alta [Alberta] 06.6.24 [6. vi. 1924] Dr A Fenyes ”, “ 294. ”, “ 79 ”, “ parvipennis Brh. Typus. ” (yellow label), “ Chicago NHMus M. Bernhauer Collection ” (FMNH). Paralectotypes: ♀, “ Banff Alta [Alberta] 06.6.24 [6. vi. 1924] Dr A Fenyes ”, “ 285. ”, white square, “ Atheta parvipennis Brnh. ”, “ parvipennis Brh. Cotypus. ” (yellow label), “ Chicago NHMus M. Bernhauer Collection ” (FMNH). Holotype of Boreostiba hudsonica: CANADA: Québec: ♂, Great Whale River, 17. vi. 1949 (J. R. Vockeroth) (CNCI). Paratype: CANADA: Québec: ♀, Great Whale River, 16. vii. 1949 (J. R. Vockeroth) (CNCI). Additional material. UNITED STATES: Alaska: ♂, Sitka (Holmberg) (FMNH); New Hampshire: Coos Co.: 2 ♂♂, ♀, Mt. Washington, 3800 ’, 16. viii. 1976 (J. M. & B. A. Campbell) (CNCI, SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFCA5307CD245076FE49A361.taxon	diagnosis	Diagnosis. See Lohse et al. 1990 (as Boreostiba hudsonica Lohse, 1990).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFCA5307CD245076FE49A361.taxon	discussion	Discussion. The types of At. parvipennis and Boreostiba hudsonica are similar in external characters and the shape of the aedeagus and spermatheca. The two names are considered to be synonyms. The generic placement of this species is considered below (See Discussion for Boreostiba).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFCA5307CD245076FE49A361.taxon	distribution	Distribution. Atheta parvipennis is known from Alaska, Alberta, Québec and New Hampshire.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFCB5300CD2452EEFC26A0F9.taxon	description	(Figs. 82 – 83 in Lohse et al. 1990 (as Boreophilia chillcotti ))	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFCB5300CD2452EEFC26A0F9.taxon	materials_examined	Type material. Lectotype of Atheta caviceps (here designated): ♂ (with the head and thorax missing), “ TYPE ” (red label), “ Pine Ind. [iana] v: 19: 07 [19. v. 1907] ”, “ 3612. Det. A. Fenyes ”, “ Purdue Blatchley collection ”, “ Atheta caviceps sp. nov. 5243 ” (PURC). Paralectotypes: ♀ (with the head and prothorax missing), “ Marshall Co. Ind. [iana] W. S. B. [Blatchley] 5 ­ 20 ­ 09 [20. v. 1909] ”, “ 3616 det. A. Fenyes ”, “ Purdue Blatchley collection ” (PURC) (The second paralectotype of At. caviceps (♀ (with the head and prothorax missing), “ Marion Co. Ind. [iana] W. S. B. [Blatchley] 5 ­ 15 ­ 10 [15. v. 1910] ”, “ 3616 det. A. Fenyes ”, “ Purdue Blatchley collection ” (PURC )) is not conspecific with the lectotype). Holotype of Boreophilia chillcotti: CANADA: Manitoba: ♂, Churchill, 4. vi. 1952 (J. G. Chillcott) (CNCI). Additional material. UNITED STATES: Alaska: 2 ♂♂, 2 ♀♀, Kenai Mts., 3 mi. NW Jct. Hwy. 1 & 9, 400 ’, 18. vi. 1978 (Smetana & Becker) (CNCI, KSEM); ♂, Mi. 23 Taylor Hwy., 8. vii. 1968 (Campbell & Smetana) (CNCI); 2 ♂♂, 6 ♀♀, Mi. 1249 Alaska Hwy., Deadman Lake, 6 – 7. vii. 1968 (Campbell & Smetana) (CNCI, SPSU); ♂, Mi. 1252 Alaska Hwy., 7. vii. 1968 (Campbell & Smetana) (CNCI); ♀, Prudhoe Bay Rd., Bonanze Cr., 66 º 40 ’ N 150 º 40 ’ W, 900 ’, 2. vii. 1978 (A. Smetana & J. M. Campbell) (CNCI); CANADA: New Brunswick: ♂, Kouchibouguac N. P., 21. ix. 1977 (J. M. Campbell) (CNCI); Northwest Territories: ♂, 12 ♀♀, 5 mi. SE Fort Simpson, 21. vi. 1972 (A. Smetana) (CNCI, KSEM, SPSU); Yukon Territory: ♀, Dawson City, 10. vii. 1968 (Campbell & Smetana) (CNCI).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFCB5300CD2452EEFC26A0F9.taxon	diagnosis	Diagnosis. See Lohse et al. 1990.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFCB5300CD2452EEFC26A0F9.taxon	discussion	Discussion. The types of At. caviceps and Boreophilia chillcotti are identical in external characters and in the shape of the aedeagus, including the sclerites of the internal sac. Atheta blatchleyi has the pronotal setation pattern of Type II. The holotype of Boreophilia chillcotti has damaged setation on the pronotum, and that explains why Lohse (Lohse et al. 1990) could not describe the setation of that species correctly. Based on the pronotal setation pattern and the shape of the aedeagus and spermatheca, this species cannot be placed in Boreophilia. Pending generic revision of Athetini, the species is placed in Atheta.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFCB5300CD2452EEFC26A0F9.taxon	distribution	Distribution. Atheta blatchleyi is a transnearctic species but its distribution is poorly documented. It is known from the United States (Alaska and Indiana) and Canada (Yukon Territory, Northwest Territories, Manitoba and New Brunswick).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFCC5301CD24504BFCA5A6F1.taxon	description	(Fig. 105 in Strand & Vik 1964 (as At. celata); Figs. 15: 5 in Benick & Lohse 1974, p. 189)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFCC5301CD24504BFCA5A6F1.taxon	materials_examined	Type material. Lectotype of Datomicra decolorata (here designated): ♂, “ N. [ew] Y. [ork, Catskill Mts.] ”, “ decolorata Csy. ”, “ TYPE USNM 39163 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 11 specimens, “ N. [ew] Y. [ork, Catskill Mts.] ”, “ decolorata PARATYPE USNM 39163 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Datomicra inopia (here designated): ♂, “ N. [ew] Y. [ork, Catskill Mts.] ”, “ inopia Csy. ”, “ TYPE USNM 39164 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 21 specimens, “ N. [ew] Y. [ork, Catskill Mts.] ”, “ inopia PARATYPE USNM 39164 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Datomicra schematica (here designated): ♂, “ Penn. [sylvania, Philadelphia] ”, “ schematica Csy. ”, “ TYPE USNM 39165 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: ♂, 2 ♀♀, “ Penn. [sylvania, Philadelphia] ”, “ schematica PARATYPE USNM 39165 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Datomicra stilla (here designated): ♂, “ R. [hode] I. [sland, Boston Neck] ”, “ stilla Csy. ”, “ TYPE USNM 39170 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 6 specimens, “ R. [hode] I. [sland, Boston Neck] ”, “ stilla PARATYPE USNM 39170 ” (red label), “ CASEY bequest 1925 ” (NMNH). Additional material. BELARUS’: ♀, Belovezhskaya Pushcha Nature Reserve, on Ganoderma, 11. vii. 1994 (V. Tsinkevich) (SPSU); CANADA: Newfoundland: ♂, ♀, Long Range Mts., 50 º 34 ’ 40 ” N 56 º 57 ’ 35 ” W, 1500 ’, 26. vii. 1969 (M. J. D. Brendell); ♂, Trappers Cove, from fungi, 8. viii. 1969 (M. J. D. Brendell) (BMNH); UNITED STATES: New York: Orange Co.: ♀, 8 km S New Windsor, Black Rock Forest, 23 – 25. v. 1998 (V. I. Gusarov) (SPSU); 2 ♂♂, ditto but pitfall traps, vi. 1998 (V. Ovcharenko) (SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFCC5301CD24504BFCA5A6F1.taxon	diagnosis	Diagnosis. See Benick & Lohse (1974).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFCC5301CD24504BFCA5A6F1.taxon	discussion	Discussion. The types of Dat. decolorata, Dat. inopia, Dat. schematica and Dat. stilla do not differ from examined European specimens of At. dadopora.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFCC5301CD24504BFCA5A6F1.taxon	distribution	Distribution. Atheta dadopora is a common European species. In North America At. dadopora is known from Rhode Island, New York, Pennsylvania and Newfoundland. This is the first record of this species from North America.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFCD531CCD24567EFD07A101.taxon	description	(Figs. 107 – 115)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFCD531CCD24567EFD07A101.taxon	materials_examined	Type material. Lectotype of Atheta pennsylvanica (here designated): ♂, “ Marion, Mass. [achusetts] Bowditch. ”, “ pennsylvanica Brh. Cotypus ” (yellow label), “ Chicago NHMus M. Bernhauer Collection ” (FMNH). Paralectotypes: ♂, “ Penn. [sylvania] ”, “ July [18] 96 ”, “ red circle ”, “ flaveola Melsh. Fenyes det. Bernhauer ”, “ pennsylvanica Brh. Typus. Fenyes. ” (yellow label), “ Chicago NHMus M. Bernhauer Collection ” (FMNH); ♀, “ Penn. [sylvania] ”, “ July [18] 96 ”, “ white circle ”, “ pennsylvanica Brh. Cotypus ” (yellow label), “ Chicago NHMus M. Bernhauer Collection ” (FMNH). Holotype of Atheta fontis: ♀, “ Buena Vista Spg. [Springs] Franklin Co. Pa. [Pennsylvania] ”, “ fontis Csy. ”, “ TYPE USNM 39357 ” (red label), “ CASEY bequest 1925 ” (NMNH). Additional material. UNITED STATES: Minnesota: Lake Co.: ♀, 22 km NE Two Harbors, Split Rock Lighthouse State Park, Hwy. 61, 47 º 10.99 ’ N 91 º 24.52 ’ W, 70 m, 10. vii. 1999 (V. I. Gusarov) (SPSU); New York: Orange Co.: ♂, ♀♀, 8 km S New Windsor, Black Rock Forest, 23 – 25. v. 1998 (V. I. Gusarov) (SPSU); Pennsylvania: Clinton Co.: 12 specimens, 26 km SEE Lock Haven, 41 º 03.23 ’ N 77 º 09.65 ’ W, 350 ’, 5. iv. 1999 (V. I. Gusarov) (SPSU); Vermont: Windsor Co.: 23 specimens, 8 km SEE Springfield, Hwy. I­ 91, 43 º 14.00 ’ N 72 º 26.81 ’ W, 250 ’, 21. ix. 1998 (V. I. Gusarov) (SPSU); CANADA: Ontario: ♀, 74 km NWW Sault Ste. Marie, Hwy. 17, slope facing Lake Superior, 47 º 09.61 ’ N 84 º 41.90 ’ W, 80 ’, in forest litter, Betula, Picea, Populus, Acer, Pinus, 11. vii. 1999 (V. I. Gusarov); 2 ♀♀, 24 km SWW Wawa, Hwy. 17, Dead Woman Bay, 47 º 47.51 ’ N 84 º 53.77 ’ W, 60 ’, in forest litter, Picea, Betula, 11. vii. 1999 (V. I. Gusarov) (SPSU); Québec: ♀, Anti­Costi, 16. vi. 1993 (LFC); ♀, La Tuque, 13. vi. 1994 (LFC); ♀, Armagh, 46 º 44 ’ N 70 º 35 ’ W, 11. viii. 1994 (LFC); ♀, ditto but 7. vi. 1994 (LFC); ♂, ditto but 16. viii. 1994 (SPSU); ♀, Lac Princeton, 18. vii. 1994 (LFC); ♀, Lac Mitis, 20. vi. 1994 (LFC); ♀, Pohénégamook, 47 º 28 ’ N 69 º 14 ’ W, 19. vii. 1994 (SPSU); ♂, Pelegrin, 5. vii. 1994 (SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFCD531CCD24567EFD07A101.taxon	diagnosis	Diagnosis. Atheta pennsylvanica belongs to a group of distinctly colored species within Microdota with dark brown head and abdominal terga 5 – 8, light brown elytra, and reddish yellow pronotum and abdominal terga 3 – 4. Atheta pennsylvanica can be distinguished from other species of this group by having a larger body size and the distinct shape of the aedeagus (Figs. 114 – 115) and spermatheca (Figs. 112 – 113).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFCD531CCD24567EFD07A101.taxon	discussion	Discussion. The types of At. fontis are similar to the types of At. pennsylvanica in the body size, in external characters and in the shape of the spermatheca. Therefore, the synonymy of the two species established by Fenyes (1920) and Bernhauer & Scheerpeltz (1926) is confirmed. I agree with Bernhauer & Scheerpeltz (1926) that of all species of Microdota described by Casey (1910 a, 1911 a) only At. fontis is synonymous with At. pennsylvanica.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFCD531CCD24567EFD07A101.taxon	distribution	Distribution. Atheta pennsylvanica is widespread in the eastern United States (Vermont, Massachusetts, New York, Pennsylvania and Minnesota) and is also known from eastern Canada (Ontario, Québec).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD0531DCD2457E4FB34A17C.taxon	description	(Figs. 90 – 92 in Lohse et al. 1990 (as Boreophilia caseyiana ))	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD0531DCD2457E4FB34A17C.taxon	materials_examined	Type material. Lectotype of Dinaraea nomensis (designated by Lohse et al. (1990 )): UNITED STATES: Alaska: ♂, Nome (NMNH). Holotype of Boreophilia caseyiana: CANADA: Yukon Territory: ♂, North Fork Pass, Ogilvie Mts., 3500 ’, 17. vi. 1962 (R. E. Leech) (CNCI).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD0531DCD2457E4FB34A17C.taxon	diagnosis	Diagnosis. See Lohse et al. (1990).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD0531DCD2457E4FB34A17C.taxon	discussion	Discussion. Lohse (Lohse et al. 1990) considered the single type of Din. nomensis in the Casey collection as the holotype. However, Casey (1910 a) neither used the word “ holotype ” nor stated that he had the single specimen of that species. Therefore, Lohse designated the lectotype of Din. nomensis by inference of holotype (Article 74.6; ICZN 1999). Lohse (Lohse et al. 1990) stated that “ the aedeagi of B. caseyiana and B. nomensis are extremely similar ”. However, based on a larger body size (3.8 mm versus 3.0 mm) and a less transverse pronotum (width to length ratio 11 / 5 versus 11 / 3) of the holotype of Boreophilia caseyiana in comparison to the lectotype of Din. nomensis, Lohse considered the two as separate species. The aedeagi of the types of both species, including the sclerites of the internal sac, are in fact identical. According to my measuring, in the holotype of Boreophilia caseyiana the body length is 3.6 mm and pronotal width to length ratio is 1.22. In the lectotype of Din. nomensis the body length is 3.3 mm, and pronotal width to length ratio is 1.29. In Athetini, such difference in body length and pronotal width is not unusual within the same species. Considering the identity of the aedeagi I place Boreophilia caseyiana in synonymy with Boreophilia nomensis.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD0531DCD2457E4FB34A17C.taxon	distribution	Distribution. Boreophilia nomensis is known from Alaska and Yukon Territory.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD2531FCD24524EFB70A161.taxon	description	(Figs. 116 – 124)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD2531FCD24524EFB70A161.taxon	materials_examined	Type material. Lectotype of Atheta angusticornis (here designated): ♂ (with broken antennae), “ Mt. Washgtn [Washington] N. [ew] H. [ampshire] ”, “ July ”, white circle, “ angusticornis Brh. Typ. ”, “ angusticornis Brh. Typus ” (yellow label), “ Chicago NHMus M. Bernhauer Collection ” (FMNH). Paralectotype: ♀ (with missing head and prothorax), “ Mt. Washgtn [Washington] N. [ew] H. [ampshire] Fenyes ”, “ angusticornis Brh. Cotypus ” (yellow label), “ Chicago NHMus M. Bernhauer Collection ” (FMNH). Lectotype of Metaxya plutonica (here designated): ♀, “ Subalpine Mt. Wn. [Washington] N. [ew] H. [ampshire] vii. 7.1899 ”, “ plutonica Csy. ”, “ TYPE USNM 39064 ” (red label), “ CASEY bequest 1925 ” (NMNH).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD2531FCD24524EFB70A161.taxon	diagnosis	Diagnosis. Boreophilia angusticornis is closely related to Palaearctic Boreophilia subplana (J. Sahlberg, 1880) but differs in the shape of the aedeagus (Figs. 120 – 123), particularly a narrower apex of the median lobe (in parameral view) (cf. Fig. 73 in Brundin 1953 and Fig. 120 in this paper).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD2531FCD24524EFB70A161.taxon	discussion	Discussion. The lectotype of Me. plutonica and the paralectotype of At. angusticornis are similar in external characters and also in the shape of the spermatheca. The two species were described from the same locality. I consider Me. plutonica to be a synonym of Boreophilia angusticornis.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD2531FCD24524EFB70A161.taxon	distribution	Distribution. Boreophilia angusticornis is known only from the Mount Washington, New Hampshire. This area was affected by the last glaciation and is unlikely to have relict endemic species. Boreophilia angusticornis may occur in other high mountains of the north­eastern United States and also in Canada, at lower altitudes further north.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD3531FCD2450EEFE15A456.taxon	diagnosis	Diagnosis. See Lohse et al. (1990).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD3531FCD2450EEFE15A456.taxon	discussion	Discussion. Lohse et al. (1990) included in the genus Boreostiba five species. Two of them, Boreostiba sibirica (Mäklin, 1880) (the type species) and Boreostiba frigida (J. Sahlberg, 1880) (see Discussion for that species), have a long, completely fused and tubelike bridge of the median lobe and enlarged condilites of the parameres. Two other species included in Boreostiba by Lohse (Lohse et al. 1990), Boreostiba hudsonica Lohse, 1990 (a junior synonym of Atheta parvipennis Bernhauer, 1907; see above) and Boreostiba campbelliana Lohse, 1990 lack these features of the aedeagus and are very similar to a species (Dimetrota smetanai Lohse, 1990) placed by Lohse in a different genus. The males of Boreostiba lagunae Lohse, 1990 are unknown but this species is more similar to Boreostiba campbelliana than to Boreostiba sibirica. Until a generic revision of Athetini is completed, Boreostiba hudsonica, Boreostiba campbelliana and Boreostiba lagunae are placed in Atheta.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD35319CD2455EEFCB1A049.taxon	description	(Figs. 87 – 89 in Brundin 1940; Figs. 190 – 193 in Lohse et al. 1990 (as Boreostiba lamellifera ))	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD35319CD2455EEFCB1A049.taxon	materials_examined	Type material. Holotype of Boreostiba lamellifera: CANADA: Québec: ♂, Payne Bay, 19. viii. 1958 (W. R. M. Mason) (CNCI). Paratypes: CANADA: Québec: 4 ♀♀, Payne Bay, 19. viii. 1958 (W. R. M. Mason) (CNCI); Yukon Territory: 2 ♂♂, British Mts., “ Sunday Mt. ”, 950 m, 69 º 17 ’ N 140 º 03 ’ W, under rocks at edge of snowfield, 17. vi. 1984 (J. M. Campbell) (CNCI). Additional material. RUSSIA: Chukotskiy Aut. Distr.: ♂, 1 specimen, 174 th km of the road Egvekinot – Iul’tin, 67 º 42 ’ N 178 º 40 ’ W, 17. vii. 1989 (Yu. M. Marusik); ♂, 2 specimens, 161 th km of the road Egvekinot – Iul’tin, 24. vi. 1989 (Yu. M. Marusik); 2 ♂♂, ♀, 3 specimens, 121 th km of the road Egvekinot – Iul’tin, 67 º 18 ’ N 178 º 40 ’ W, vi. 1989 (Yu. M. Marusik) (SPSU); Magadan Region: ♂, 2 specimens, upper Kolyma River, Tundrovyy Creek, mountain tundra, under Salix, 1200 m, 11. vii. 1986 (Yu. M. Marusik); 5 specimens, upper Kolyma River, Bol’shoy Annachag Ridge, env. of Sibit­Tyellah, 1400 m, the Tundrovaya River valley, small Salix bush with moss, 6. ix. 1986 (Yu. M. Marusik); 3 ♂♂, ♀, 5 specimens, upper Kolyma River, Bol’shoy Annachag Ridge, env. of Sibit­ Tyellah, 800 m, Olen’ River, Alnus, Pinus pumila, in leaf litter, 15. vii. 1985 (Yu. M. Marusik); SWEDEN: ♂, Abisko (L. Brundin) (MZHF).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD35319CD2455EEFCB1A049.taxon	diagnosis	Diagnosis. See Brundin (1940; as At. frigida) and Lohse et al. (1990; as Boreostiba lamellifera).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD35319CD2455EEFCB1A049.taxon	discussion	Discussion. Lohse (Lohse et al. 1990) did not examine the types of At. frigida and based his judgement on the status of this species on the drawings by Brundin (1940) who had examined the types of both At. frigida and H. sibirica (the type species of Boreostiba). Lohse (Lohse et al. 1990: 197) believed that the difference in Brundin’s drawings for At. frigida and At. sibirica resulted from “ the variable extent of the evagination of the internal sac ”. I do not agree with this interpretation of Brundin’s drawings (Brundin 1940: Figs. 85 – 88) which are, in my opinion, more accurate than the drawings in Lohse et al. (1990: Figs. 190 – 191, 194 – 196). When Brundin’s drawings of the median lobe of the aedeagus of At. sibirica (Figs. 85 – 86) and At. frigida (Figs. 87 – 88) are compared, the difference between the two species in the length of the athetine bridge (cf. Fig. 85 and Fig. 87 in Brundin 1940) and the shape of the apex of the median lobe (cf. Fig. 86 and Fig. 88 in Brundin 1940) is very clear. Both the athetine bridge and the apex of the median lobe are rigid structures which do not change their shape when the internal sac is everted. The differences in Brundin’s drawings for At. sibirica and At. frigida allow to easily separate both species (the first species has a shorter athetine bridge than the second species) and cannot be attributed to a different extent of the evagination of the internal sac. My examination of the holotype of Boreostiba lamellifera demonstrated that it was conspecific with examined Palaearctic specimens of Boreostiba frigida. The drawings of the aedeagus of Boreostiba lamellifera (Figs. 190 – 191, 193 in Lohse et al. 1990) match the corresponding drawings of Boreostiba frigida (Figs. 87 – 88 in Brundin 1940). Therefore I remove Boreostiba frigida from synonymy with Boreostiba sibirica and place Boreostiba lamellifera in synonymy with Boreostiba frigida.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD35319CD2455EEFCB1A049.taxon	distribution	Distribution. Boreostiba frigida has circumpolar distribution. In North America it is known from Canada (Québec and Yukon Territory).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD5531ACD24574BFE0EA129.taxon	description	(Figs. 7 – 9 in Benick 1954)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD5531ACD24574BFE0EA129.taxon	materials_examined	Type material. Lectotype of Atheta laurentiana (here designated): ♀, (with the head and prothorax missing), “ TYPE ” (red label), “ Lawr. [ence] Co. Ind. [iana] W. S. B. [Blatchley] 5 ­ 11 ­ 04 [11. v. 1904] ”, “ 3631 det. A. Fenyes ”, “ 5 [?] 8 ”, “ Purdue Blatchley collection ”, “ Atheta laurentiana sp. nov. 5680 ” (PURC). Additional material. UKRAINE: Crimea: 8 specimens, Simferopol distr., 5 km SW Krasnoles’ye, near creek, 400 m, 19. v. 1990 (V. I. Gusarov); 6 specimens, ditto but 21. v. 1990; 9 specimens, Crimean Game Reserve, Alma River banks, 600 m, 3. vi. 1986 (V. I. Gusarov) (SPSU); UNITED STATES: Illinois: Monroe Co.: 2 ♂♂, ♀, Krueger Dry Run Cave, 3. iv. 1992 (S. Taylor & J. Krejca) (INHS, SPSU); New Hampshire: Grafton Co.: ♀, 2 km NNE Warren, Hwy. 118, Baker River, 43 º 56.42 ’ N 71 º 52.52 W, 300 m, river bank, under stones and in moss, 21. ix. 1998 (V. I. Gusarov) (SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD5531ACD24574BFE0EA129.taxon	diagnosis	Diagnosis. See Benick 1954; Benick & Lohse 1974.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD5531ACD24574BFE0EA129.taxon	discussion	Discussion. The lectotype of At. laurentiana is similar to examined European specimens of Alo. sulcifrons in external characters and in the shape of spermatheca. Both names are considered synonymous. The illustrations of the male tergum 8, the aedeagus and spermatheca provided by Klimaszewski and Peck (1986: Figs. 92 – 95) correspond to Alo. sulcifrons and not Alo. insecta (Thomson, 1856) (cf. Figs. 7 – 9 and Fig. 6 in Benick 1954).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD5531ACD24574BFE0EA129.taxon	distribution	Distribution. Aloconota sulcifrons is a common European species. According to Benick (1954), Alo. sulcifrons has a cosmopolitan distribution, but the old records not based on examination of genitalic characters need to be reconfirmed. In North America Alo. sulcifrons is known from New Hampshire, New York, Indiana, Illinois, Missouri, Kentucky, Virginia, West Virginia, Tennessee and Alabama (Klimaszewski & Peck 1986 (as Alo. insecta )).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD6531ACD245026FD2DA702.taxon	type_taxon	type species: Atheta massettensis Casey, 1910 a, by original designation).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD6531ACD245026FD2DA702.taxon	discussion	(For additional references see Gusarov 2003 c)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD6531BCD2456CBFCDAA0A1.taxon	description	(Figs. 1 – 16, 18 – 30 in Gusarov 2003 c)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD6531BCD2456CBFCDAA0A1.taxon	materials_examined	Type material. Lectotype of Homalota maritima (designated by Lohse and Smetana (1985 )): UNITED STATES: Alaska: Sitka (MZHF). Additional material. UNITED STATES: Alaska: 14 specimens, Kenai Peninsula, Homer, S shore of Homer spit, 59 º 36.33 ' N 151 º 25.71 ' W, sandy seashore, in seaweed (V. I. Gusarov), 22. vii. 1998 (AMNH, SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD6531BCD2456CBFCDAA0A1.taxon	diagnosis	Diagnosis. See Gusarov (2003 c).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD6531BCD2456CBFCDAA0A1.taxon	discussion	Discussion. The description and illustrations (Easton 1971) of At. immigrans agree completely with the lectotype and examined specimens of Ad. maritima. Therefore At. immigrans is a synonym of Ad. maritima.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD6531BCD2456CBFCDAA0A1.taxon	distribution	Distribution. Adota maritima is known from the Pacific coast of North America, from Alaska to California. The recent records from Britain (Easton 1971, Hammond 2000) apparently represent an introduction.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD75314CD2451AEFBA5A70C.taxon	description	(Fig. 55 in Yosii & Sawada 1976; Figs. 125 – 126 in this paper)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD75314CD2451AEFBA5A70C.taxon	type_taxon	type species: Dimetrota (Dalotia) pectorina Casey, 1910 a, by original designation).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD75314CD2451AEFBA5A70C.taxon	diagnosis	Diagnosis. Dalotia can be recognized by the combination of the following characters: body broad; sensilla a of epipharynx long; antennal article 2 as long as article 3, articles 4 – 10 slightly elongate or subquadrate; ligula with narrow base and split apically; labial palpus with setae, and present; ligula with two long setae; hypopharynx with long setae; infraorbicular carina complete, pronotum strongly transverse, with microsetae directed anteriad along the midline; in lateral portions of the disc microsetae directed laterad and obliquely posteriad (Type I, Benick & Lohse 1974); pronotal macrosetae long; pronotal hypomera fully visible in lateral view; medial macroseta of mesotibia strong, longer than tibial width; mesothoracic process narrow; tarsal formula 4 ­ 5 ­ 5; metatarsal segment 1 as long as segment 2; with one empodial seta, shorter than claws; abdominal terga 3 – 5 with transverse basal impression; medial lamellae of internal sac present; copulatory piece with pointed apex. Dalotia differs from Atheta in having its ligula with two long setae (Fig. 125) and long hypopharyngeal setae (Fig. 126).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD75314CD2451AEFBA5A70C.taxon	discussion	Discussion. The type species of Dalotia is Dimetrota pectorina Casey, 1910 a by original designation. Casey did not specifically state in his description of Dalotia that Dimetrota pectorina was the type species of Dalotia; however, while designating the type species of Noverota Casey, 1910 a in the same paper, Casey stated (1910 a, p. 90): “ The first species may be regarded as the type, as in all other cases where the type is not specifically named ”. Dimetrota pectorina is a synonym of the species known as Atheta coriaria (Kraatz, 1856) (see below). Muona (1979 a, 1979 b, 1987) pointed out that the species well known under the name Atheta coriaria does not in fact belong to Atheta because of the bisetose glossa and the distinct complex structure of the internal sac of the aedeagus. Because this species occurs in North America (Muona 1979 a, 1984) Muona did not propose a new name for the genus which would include this species. He suggested that one of the Casey names might be used when the types of Casey species are revised. My revision of Casey types of Aleocharinae suggests that Dalotia should be the valid name for this genus.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD85316CD2456C2FD31A019.taxon	description	(Fig. 129 in Strand & Vik 1964)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD85316CD2456C2FD31A019.taxon	materials_examined	Type material. Lectotype of Pseudota miscella (here designated): ♂, “ N. [ew] J. [ersey] ”, “ miscella Csy. ”, “ TYPE USNM 39147 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Dimetrota pectorina (here designated): ♂, “ Sta. Cruz Mts. Cal. [ifornia] ”, “ Dalotia pectorina Csy. ”, “ TYPE USNM 39131 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Dimetrota crucialis (here designated): ♂, “ Sta. Cruz Mts. Cal. [ifornia] ”, “ crucialis Csy. ”, “ TYPE USNM 39132 ” (red label), “ CASEY bequest 1925 ” (NMNH). Additional material. UNITED STATES: Louisiana: East Baton Rouge Parish: ♀, Baton Rouge, Louisiana State University campus, 1 – 15. vi. 1998 (A. Tishechkin); ♂, ditto but light trap, 7 – 17. v. 1998 (SPSU); New York: Manhattan: ♂, ♀, 6 th Street & Avenue B Garden, 40 º 43.45 ' N 73 º 58.91 ' W, compost, 3. xii. 1998 (A. Berkov & V. R. Coffman); 4 specimens, ditto but 5. ix. 1999 (A. Berkov) (SPSU); UKRAINE: Crimea: 10 specimens, env. of Yalta, Botanicheskoye, in rotting hay, 150 m, 2. ix. 1990 (V. I. Gusarov) (SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD85316CD2456C2FD31A019.taxon	diagnosis	Diagnosis. See Benick & Lohse (1974; as Atheta coriaria). Aedeagus and spermatheca: Fig. 129 in Strand & Vik (1964).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD85316CD2456C2FD31A019.taxon	discussion	Discussion. The lectotypes of Ps. miscella, Dim. pectorina and Dim. crucialis agree completely with the specimens of Dal. coriaria from Europe in external characters, including male secondary features and the shape of the aedeagus. Muona (1979 a, 1987) pointed out that Atheta coriaria is conspecific with Taxicera academica based on the detailed illustrations of the internal sac of the aedeagus of the latter species by Yosii and Sawada (1976). I accept Muona's interpretation.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFD85316CD2456C2FD31A019.taxon	distribution	Distribution. In North America Dal. coriaria is known from California, Florida (Frank 1980), Louisiana, New Jersey and New York. Because Dal. coriaria is restricted to areas on the West and East coasts, and not known from inland states of the United States, it is probably an introduction from Europe.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFDA5317CD2450ECFBA8A159.taxon	description	(Fig. 48 in Strand & Vik 1964)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFDA5317CD2450ECFBA8A159.taxon	materials_examined	Type material. Holotype of Dimetrota revoluta: ♂, “ Id. [aho] [Coeur d'Alene] ”, “ revoluta Csy. ”, “ TYPE USNM 39122 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Datomicra vaciva (here designated): ♂, “ Mass. [achusetts] [Blanchard leg.] ”, “ vaciva Csy. ”, “ TYPE USNM 39173 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 2 ♀♀, “ Mass. [achusetts] [Blanchard leg.] ”, “ vaciva PARATYPE USNM 39173 ” (red label), “ CASEY bequest 1925 ” (NMNH). Additional material. CANADA: Newfoundland: 2 ♂♂, 4 ♀♀, W Newfoundland, Deer Lake, 30. v. 1951 (Lindroth) (MZHF); Northwest Territories: ♂, Fort Simpson, 10. vi. 1972 (A. Smetana); ♂, ditto but Harris River, 15. vi. 1972; ♂, 2 ♀♀, 5 mi. SE Fort Simpson, Hwy. 3, 21. vi. 1972 (A. Smetana); ♀, 8 mi. SE Fort Simpson, 18. vi. 1972 (A. Smetana) (CNCI); Yukon Territory: ♀, Dempster Hwy. Mi. 152.5, 1900 ’, 20. vii. 1978 (A. Smetana & J. M. Campbell) (CNCI).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFDA5317CD2450ECFBA8A159.taxon	diagnosis	Diagnosis. See Benick & Lohse (1974; as Atheta rudiventris). Aedeagus and spermatheca (Fig. 48: Strand & Vik, 1964).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFDA5317CD2450ECFBA8A159.taxon	discussion	Discussion. The types of Dimetrota revoluta and Datomicra vaciva agree completely with European specimens of Dochmonota rudiventris in external characters, and in the shape of the aedeagus and spermatheca.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFDA5317CD2450ECFBA8A159.taxon	distribution	Distribution. In North America Dochmonota rudiventris is known from Yukon Territory, Northwest Territories, Newfoundland, Massachusetts and Idaho.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFDB5317CD245096FC8BA424.taxon	diagnosis	Diagnosis. See Pace (1982).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFDB5317CD245096FC8BA424.taxon	discussion	Discussion. The type species of Pseudousipalia and Emmelostiba are similar in their mouthparts, particularly the shape of the ligula which is split into two separate lobes (Fig. 12 in Pace 1982); in their pronotal setation pattern (Fig. 11 in Pace 1982); and in the distinct shape of the aedeagus, particularly the deeply split apex of the median lobe (Fig. 16 in Pace 1982; Fig. 54 in Lohse et al. 1990). Based on this similarity, I place Pseudousipalia in synonymy with Emmelostiba. The genus Emmelostiba was previously known from the Palaearctic and Oriental Regions.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFDB5310CD24552BFAD5A129.taxon	description	(Figs. 54 – 55 in Lohse et al. 1990; Fig. 1 in Lohse 1991)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFDB5310CD24552BFAD5A129.taxon	materials_examined	Type material. Holotype of Pseudousipalia microptera: CANADA: Yukon Territory: ♂, British Mts., “ June Creek ”, 320 m, 3 km NW Firth River, 69 º 13 ’ N 140 º 05 ’ W, sifting Salix and Alnus litter, 29. vi. 1984 (J. M. Campbell) (CNCI). Paratypes: CANADA: Yukon Territory: ♂, Keno, 5000 ’, 18. vii. 1968 (Campbell & Smetana) (CNCI); ♂, ♀, British Mts., “ Windy Ridge ”, 450 m, 69 º 27 ’ N 140 º 26 ’ W, sifting Salix litter, 2. vii. 1984 (J. M. Campbell); ♀, British Mts., Fish Creek, 180 m, 69 º 27 ’ N 140 º 19 ’ W, sifting litter under dwarf Populus, 1. vii. 1984 (J. M. Campbell) (CNCI). Additional material. CANADA: Yukon Territory: 3 ♂♂, ♀, 8 mi. NW Keno, 6000 ’, 19. vii. 1968 (Campbell & Smetana); 2 ♀♀, Keno, 5000 ’, 18. vii. 1968 (Campbell & Smetana) (CNCI); UNITED STATES: Alaska: ♂, Mi. 1252, Alaska Hwy., 7. vii. 1968 (Campbell & Smetana); ♂, Prudhoe Bay Road, creek near Minnie Creek, 1. vii. 1978 (J. M. Campbell); ♂, Mi. 43, Taylor Hwy., 8. vii. 1968 (Campbell & Smetana) (CNCI).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFDB5310CD24552BFAD5A129.taxon	diagnosis	Diagnosis. See Lohse et al. (1990).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFDB5310CD24552BFAD5A129.taxon	distribution	Distribution. Emmelostiba microptera is known from Alaska and Yukon Territory.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFDC5312CD2457D4FEBFA0F9.taxon	description	(Fig. 12 in Lohse & Smetana 1985; Figs. 98 – 100 in Klimaszewski & Winchester 2002)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFDC5312CD2457D4FEBFA0F9.taxon	materials_examined	Type material. Lectotype of Homalota nitens (designated by Lohse and Smetana (1985 )): UNITED STATES: Alaska: ♂, Sitka (Holmberg) (MZHF). Lectotype of Atheta insolens (here designated): ♂, “ QCI [Queen Charlotte Islands] ”, “ insolens ­ 2 PARATYPE USNM 39479 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotype: ♀, “ QCI [Queen Charlotte Islands] ”, “ insolens Csy. ”, “ TYPE USNM 39479 ” (red label), “ CASEY bequest 1925 ” (NMNH). Holotype of Dimetrota resplendens: ♀, “ QCI [Queen Charlotte Islands] ”, “ resplendens Csy. ”, “ TYPE USNM 39123 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Atheta apposita (here designated): ♀, “ Metlakatla, B. [ritish] Col. [umbia] Keen ”, “ apposita Csy. ”, “ TYPE USNM 39308 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotype: ♀, “ Metlakatla, B. [ritish] Col. [umbia] Keen ”, “ apposita PARATYPE USNM 39308 ” (red label), “ CASEY bequest 1925 ” (NMNH). Additional material: UNITED STATES: Alaska: 7 ♀♀, Denali State Park, Byers Lake Campground, 23. vi. 1978 (Smetana & Becker); 8 ♀♀, ditto but 25. vi. 1978; ♀, ditto but above Byers Lake, 2500 ’, 24. vi. 1978 (Smetana & Becker); 3 ♀♀, Denali State Park at Hwy. 1, 20. vi. 1978 (Smetana & Becker); 4 ♂♂, 7 ♀♀, Kenai Mts., Ptarmigan Creek campground, 500 – 600 ’, 26. v. 1978 (Smetana & Becker); 5 ♀♀, Kenai Mts., 16 mi. N Seward, 500 – 600 ’, 26. v. 1978 (Smetana & Becker); 2 ♂♂, 15 ♀♀, Kenai Mts., 22 mi. N Seward, 600 – 800 ’, 27. v. 1978 (Smetana & Becker); 7 ♀♀, Kenai Mts., 23 mi. N Seward, 600 – 900 ’, 28. v. 1978 (Smetana & Becker); 10 ♀♀, Kenai Mts., 15 mi. N Seward, 400 ’, 29. v. 1978 (Smetana & Becker); ♀, Kenai Mts., 2 mi. S Moose Pass, 500 ’, 30. v. 1978 (A. Smetana & E. C. Becker); ♀, Kenai Peninsula, 1 mi. W Anchor Point, 1. vi. 1978 (Smetana & Becker); ♂, 4 ♀♀, Kenai Peninsula, Anchor River Campground, 12 mi. N Homer, 450 ’, 3. vi. 1978 (Smetana & Becker); 3 ♀♀, ditto but 5. vi. 1978; 11 ♀♀, Kenai Peninsula, Anchor River at Highway 1, 450 ’, 4. vi. 1978 (Smetana & Becker); 3 ♀♀, Kenai Mts., Crown Pt. Mine 4000 ’, 3 air mi. NE Crown Pt., 17. vi. 1978 (Smetana & Becker); ♀, Kenai Mts., Tern Lake campground, 7000 ’, 12. vi. 1978 (Smetana & Becker); ♀, Kenai Peninsula, Kasilof River at Hwy. 1, 12. vi. 1978 (Smetana & Becker); ♂, ♀, Kenai Peninsula, 8 mi. SE Kasilof, 9. vi. 1978 (Smetana & Becker); ♂, Kenai Peninsula, Kalifonsky Beach near Kenai, 8. vi. 1978 (Smetana & Becker); 2 ♂♂, Kenai Peninsula, Clam Gulch, 6. vi. 1978 (Smetana & Campbell); ♀, Alaska Range, Antimony Creek, 3500 ’, 2.5 mi. E mi. 181 George Parks Hwy., 27. vi. 1978 (Smetana & Becker); ♀, 8 mi. NW Haines, 3. vii. 1968 (Campbell & Smetana) (CNCI); Oregon: Benton Co.: 5 ♂♂, 2 ♀♀, Mary’s Peak, 3800 ’, 27. vii. 1979 (J. M. & B. A. Campbell) (CNCI); Washington: ♀, Olympic N. P., Obstruction Peak, 6000 ’, 10. viii. 1979 (J. M. Campbell & B. A. Campbell); ♂, ♀, Olympic N. P., 4.0 – 6.0 mi. SE Soleduck Campground, 2500 – 3100 ’, 15. viii. 1979 (J. M. Campbell & B. A. Campbell); ♂, Olympic N. P., Lunch Lake, 4400 ’, 9 mi. SE Soleduck R. S., 17. viii. 1979 (J. M. Campbell & B. A. Campbell); ♂, Olympic N. P., 7 mi. S Port Angeles, 2400 ’, 11. viii. 1979 (J. M. Campbell & B. A. Campbell) (CNCI); CANADA: British Columbia: ♂, ♀, 5 mi. E Whistler’s Mt., 29. v. 1968 (Campbell & Smetana); ♂, Slide Mt, 10 mi. E Barkerville, 4500 ’, 24. vi. 1968 (Campbell & Smetana) (CNCI).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFDC5312CD2457D4FEBFA0F9.taxon	diagnosis	Diagnosis. See Klimaszewski & Winchester (2002). This species is unusual in lacking a sclerotized spermatheca.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFDC5312CD2457D4FEBFA0F9.taxon	discussion	Discussion. The types of H. nitens, At. insolens, At. apposita and Dim. resplendens are similar in external characters, in the shape of the aedeagus (in male specimens) and in lacking a sclerotized spermatheca (in female specimens).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFDC5312CD2457D4FEBFA0F9.taxon	distribution	Distribution. Atheta nitens is known from Alaska, British Columbia, Washington and Oregon.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFDE5312CD245076FAD0A54E.taxon	description	(Figs. 1 – 23 in Hoebeke 1988)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFDE5312CD245076FAD0A54E.taxon	diagnosis	Diagnosis. See Lohse (1974 a) and Hoebeke (1988).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFDE5312CD245076FAD0A54E.taxon	discussion	Discussion. The type species of Fusalia has all diagnostic characters of the genus Thamiaraea (particularly, very distinct labial palpi), and is conspecific with Thamiaraea lira Hoebeke, 1988. Therefore, Fusalia is a junior subjective synonym of Thamiaraea.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFDF532DCD24524EFE1EA412.taxon	description	(Figs. 11 – 14, 20 – 23 in Hoebeke 1988 (as Th. lira); Figs. 1 – 8 in Hoebeke 1994 (as Th. paralira); Figs. 127 – 134 in this paper)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFDF532DCD24524EFE1EA412.taxon	materials_examined	Type material. Lectotype of Sableta brittoni (here designated): ♀, “ Westville Ct. [Connecticut] 27 July 1905 W. E. Britton ”, “ Fusalia brittoni Csy. ”, “ TYPE USNM 39146 ” (red label), “ CASEY bequest 1925 ” (NMNH). Holotype of Thamiaraea lira: UNITED STATES: Pennsylvania: Adams Co.: ♂, Arendtsville, 24. v. 1927 (S. W. Frost) (CUIC). Paratypes: UNITED STATES: Pennsylvania: Adams Co.: ♀ (allotype), Arendtsville, 10. v. 1927 (S. W. Frost); ♀, ditto but 18. iii. 1927; ♂, ditto but 7. vi. 1927; ♂, ditto but 2. vi. 1927; ♂, ditto but 10. v. 1927; ♂, ditto but 24. v. 1927 (CUIC). Holotype of Thamiaraea paralira: UNITED STATES: New York: Westchester Co.: ♂, Yonkers, in flowers of Calycanthus fertilis, 29. v. 1938 (H. Dietrich) (CUIC). Paratypes: UNITED STATES: New York: Westchester Co.: 4 ♂♂, same data as the holotype; Mississippi: Pontotoc Co.: ♂, 1 mi. SE Ecru, pitfall trap in deciduous woods, 18. xii. 1980 (W. H. Cross) (CUIC). Additional material. UNITED STATES: Connecticut: New London Co.: ♂, molasses trap, 9. vi. 1974 (L. E. Watrous) (FMNH); Kansas: Douglas Co.: ♂, ♀, 7 km S Lawrence, ex fermenting sap flow, 30. viii. 1992 (J. S. Ashe) (KSEM); New Jersey: Middlesex Co.: 5 ♂♂, 4 ♀♀, Avenel (FMNH); Ohio: Clark [?] Co.: ♂, “ Shadywilde Selma ”, Malaise trap, center of big woods, 19. v. 1972 (R. S. Boone); ♂, ditto but corner of big woods, 1. vi. 1971; ♀, ditto but 23. ix. 1971; ♂, ditto but by creek, 3. ix. 1968; ♂, ditto but 9. vii. 1968; ♂, ditto but 6. vii. 1968; ♀, ditto but 7. vii. 1968; Franklin Co.: ♂, ♀, Columbus, Ohio State University woods, ex banana trap, 20 – 27. x. 1979 (L. E. Watrous); 2 ♂♂, ♀, ditto but 28 – 31. x. 1979 (FMNH); Pennsylvania: Northampton Co.: ♀, Easton, 30. ix. 1932 (J. W. Green) (FMNH); CANADA: Ontario: Wentworth Co.: 2 ♂♂, 2 ♀♀, Stoney Cr., on rotted bananas in pitfall, 2. vi. 1973 (H. Frania) (KSEM); Québec: ♀, Downey, 19. vi. 1995 (SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFDF532DCD24524EFE1EA412.taxon	diagnosis	Diagnosis. See Hoebeke (1988 (as Th. lira), and 1994 (as Th. paralira )).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFDF532DCD24524EFE1EA412.taxon	discussion	Discussion. According to Hoebeke (1994), Th. paralira differs from Th. lira only in having a smaller body size, shorter teeth at the posterior margin of the male abdominal tergum 8, and a weaker medial swelling in the posterior half of the tergum. My examination of the types of both species confirmed that they are identical in the shape of the aedeagus. In one paratype of Th. paralira and in several specimens fitting the description of Th. lira (in details of the male tergum 8) the internal sac was everted, and rather elaborate sclerites of the internal sac (Figs. 133 – 134) were examined. No difference between the specimens was found. Within species, the shape of the aedeagus and the sclerites of internal sac are usually less variable than the shape of the male tergum 8. It is a common phenomenon in many groups of aleocharines for smaller male specimens to have less prominent secondary sexual features (e. g., in Stethusa dichroa (Gravenhorst, 1802) (cf. Figs. 24 – 26 in Gusarov 2003 d); in Tropimenelytron americanum Gusarov, 2002 e and Tr. californicum Gusarov, 2002 e (Gusarov 2002 e )). Considering that 1) the types of Th. lira and Th. paralira are identical in the shape of the aedeagus; 2) there is no gap between the male specimens with the sexual features fitting Th. lira on one side, and Th. paralira on the other (Figs. 127 – 132); 3) it is the smaller male specimens of Thamiaraea that have less pronounced sexual features on the tergum 8; 4) both morphs are sympatric and even were collected in the same samples (Hoebeke 1994); I synonymize Th. lira and Th. paralira. My examination of the lectotype of Sableta brittoni revealed that it in fact belongs to Thamiaraea and is conspecific with both Th. lira and Th. paralira. Thamiaraea brittoni is the valid name for this species. Unfortunately, Casey (1911 a) misplaced this species in Sableta Casey, 1910 a, a genus unrelated to Thamiaraea, and when Hoebeke (1988) published his review of Thamiaraea, he had no means of knowing that one species of Thamiaraea had already been described by Casey.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFDF532DCD24524EFE1EA412.taxon	distribution	Distribution. Thamiaraea brittoni is widely distributed in the eastern United States and Canada. It is known from Ontario, Québec, Connecticut, New York, New Jersey, Pennsylvania, Ohio, Wisconsin (Hoebeke 1994), Kansas, North Carolina (Hoebeke 1994) and Mississippi.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFE2532FCD24517EFDB3A4B9.taxon	description	(Fig. 12: 5, p. 182 in Benick & Lohse 1974; Fig. 4 in Topp 1975)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFE2532FCD24517EFDB3A4B9.taxon	materials_examined	Type material. Syntypes of Aleochara fungi: ♀, “ 5499 ”, “ Europa Nr. 5499 ”, “ Typus ” (red label); “ Zool. Mus. Berlin ”, “ fungi Gr. ”; 4 ♀♀, 1 specimen, “ Europa Nr. 5499 ”, “ Typus ” (red label), “ Zool. Mus. Berlin ”; ♀, “ Sue … Schüpp ”, “ Nr. 5499 ”, “ Typus ” (red label), “ Zool. Mus. Berlin ”; 1 specimen, “ Austr … ”, “ Nr. 5499 ”, “ Typus ” (red label), “ Zool. Mus. Berlin ” (ZMHB). Holotype of Achromata fusiformis: ♀, “ N. [ew] Y. [ork] [near New York City (H. H. Smith)] ”, “ Achromata fusiformis ”, “ TYPE USNM 38990 ” (red label), “ CASEY bequest 1925 ” (NMNH). Holotype of Dimetrota nuptalis: ♀, “ R. [hode] I. [sland] ”, “ nuptalis Csy. ”, “ TYPE USNM 39114 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Acrotona lividula (here designated): ♀, “ Portland, Oreg. [on] ”, “ lividula Csy. ”, “ TYPE USNM 38981 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 2 ♀♀, “ Portland, Oreg. [on] ”, “ lividula PARATYPE USNM 38986 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Acrotona adjuvans (here designated): ♀, “ W. H. H. Ottawa, Can. [ada, Carrington leg.] ”, “ adjuvans Csy. ”, “ TYPE USNM 38986 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 10 ♀♀ (3 broken), “ W. H. H. Ottawa, Can. [ada, Carrington leg.] ”, “ adjuvans PARATYPE USNM 38986 ” (red label), “ CASEY bequest 1925 ” (NMNH). Additional material. CANADA: New Brunswick: Sunbury Co.: ♀, Acadia Research Forest, pitfall trap, 21. ix. 1999 (G. Gesner) (LFC); Ontario: 3 ♀♀, environs of Ottawa, v – vi. 1998 (V. Grebennikov) (SPSU); Québec: ♀, Lac Dumont, 4. viii. 1994 (SPSU); ♀, Lac Mitis, 19. vii. 1993 (SPSU); ♀, St. Jacques­de­Leeds, 25. vi. 1992 (SPSU); 1 specimen, Bellam (LFC); 1 specimen, Anticosti, 22. vi. 1993 (LFC); 1 specimen, Lac à l’Epaule, 15. viii. 1994 (LFC); 1 specimen, Lowry (LFC); UNITED STATES: Maine: Lincoln Co.: ♀, Sheepscot River, 7. viii. 1992 (N. Yu. Klyuge) (SPSU); Massachusetts: Berkshire Co.: 12 km N Pittsfield, 42 º 33.12 ' N 73 º 13.97 ' W, 350 m, in forest litter, Acer, Pinus, Tsuga, Quercus, 23. ix. 1998 (V. I. Gusarov) (SPSU); Minnesota: Lake Co.: 3 ♀♀, 22 km NE Two Harbors, Split Rock Lighthouse State Park, Hwy. 61, 47 º 10.99 ' N 91 º 24.52 ' W, 70 m, in forest litter, Betula, Picea, Populus, Rubus, Mayanthemum, ferns, 10. vii. 1999 (V. I. Gusarov) (SPSU); UNITED KINGDOM: Staffordshire: 2 ♀♀, near Gailey, ponds, SJ 934101, 24. i. 1993 (V. I. Gusarov) (SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFE2532FCD24517EFDB3A4B9.taxon	diagnosis	Diagnosis. See Benick & Lohse (1974).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFE2532FCD24517EFDB3A4B9.taxon	discussion	Discussion. The holotypes of Achromata fusiformis, Dim. nuptalis and the lectotypes of Acrotona lividula and Acro. adjuvans agree completely with European specimens of Moc. fungi (and the syntypes of Aleochara fungi) in external characters and the shape of spermatheca.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFE2532FCD24517EFDB3A4B9.taxon	distribution	Distribution. In North America Moc. fungi is known from Newfoundland (Muona 1984), New Brunswick, Québec, Ontario, Maine, Rhode Island, Massachusetts, New York, Minnesota, and Oregon.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFE35329CD2455B6FEB1A431.taxon	description	(Figs. 17 – 18 in Lohse & Smetana 1985)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFE35329CD2455B6FEB1A431.taxon	materials_examined	Type material. Lectotype of Homalota breviuscula (designated by Lohse and Smetana (1985 )): UNITED STATES: Alaska: Sitka (MZHF). Lectotype of Acrotona digesta (here designated): ♂, “ Cal. [ifornia, Sonoma Co.: Duncan's Mills] ”, “ digesta Csy. ”, “ TYPE USNM 38982 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 5 specimens (1 with the apex of the abdomen missing), “ Cal. [ifornia, Sonoma Co.: Duncan's Mills] ”, “ digesta PARATYPE USNM 38982 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Acrotona severa (here designated): ♀, “ Siskiyou Co., Cal. [ifornia] ”, “ severa Csy. ”, “ TYPE USNM 38984 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Acrotona shastanica (here designated): ♂, “ Siskiyou Co., Cal. [ifornia] ”, “ shastanica Csy. ”, “ TYPE USNM 38983 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 8 specimens, “ shastanica PARATYPE USNM 38983 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Acrotona prudens (here designated): ♀, “ Q. [ueen] C. [harlotte] I. [slands] [Keen leg.] ”, “ prudens Csy. ”, “ TYPE USNM 38985 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 9 specimens (1 with the head, prothorax and abdomen missing), “ Q. [ueen] C. [harlotte] I. [slands] [Keen leg.] ”, “ prudens PARATYPE USNM 38985 ” (red label), “ CASEY bequest 1925 ”; 7 specimens, “ Metlakatla, B. [ritish] Col. [umbia] Keen ”, “ prudens PARATYPE USNM 38985 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Acrotona ardelio (here designated): ♂, “ Cal. [ifornia] [Lake Tahoe] ”, “ ardelio Csy. ”, “ TYPE USNM 38988 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Acrotona renoica (here designated): ♂, “ Nev. [ada, Reno] ”, “ renoica Csy. ”, “ TYPE USNM 38987 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotype: ♂, “ Nev. [ada, Reno] ”, “ renoica PARATYPE USNM 38987 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Acrotona malaca (here designated): 1 specimen (with the abdomen missing), “ Siskiyou Co., Cal. [ifornia] ”, “ malaca Csy. ”, “ TYPE USNM 38989 ” (red label), “ CASEY bequest 1925 ” (NMNH).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFE35329CD2455B6FEB1A431.taxon	diagnosis	Diagnosis. See Lohse & Smetana (1985).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFE35329CD2455B6FEB1A431.taxon	discussion	Discussion. The lectotypes of Acro. digesta, Acro. severa, Acro. shastanica, Acro. prudens, Acro. ardelio, Acro. renoica, Acro. malaca and H. breviuscula are similar in body proportions, punctation, in the proportions of the antennal segments and in the shape of the aedeagus and spermatheca.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFE35329CD2455B6FEB1A431.taxon	distribution	Distribution. Mocyta breviuscula is known from British Columbia, California and Nevada.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFE5532ACD24553EFEA3A089.taxon	diagnosis	Diagnosis. See Benick & Lohse (1974; as subgenus of Atheta) or Lohse et al. (1990).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFE5532ACD24553EFEA3A089.taxon	discussion	Discussion. The type species of Eustrigota, Strigota seclusa, is conspecific with Acrotona sonomana (see below). Therefore, Eustrigota is placed in synonymy with Acrotona.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFE65325CD245046FB16A0D1.taxon	description	(Figs. 135 – 141)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFE65325CD245046FB16A0D1.taxon	materials_examined	Type material. Lectotype of Colpodota sonomana (here designated): ♀, “ Cal. [ifornia, Cloverdale] ”, “ sonomana Csy. ”, “ TYPE USNM 39009 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotype: ♀, “ Cal. [ifornia, Mt. Diablo] ”, “ sonomana ­ 2 PARATYPE USNM 39009 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Colpodota inceptor (here designated): ♀, “ Cal. [ifornia, Santa Clara (Harford)] ”, “ inceptor Csy. ”, “ TYPE USNM 39010 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Colpodota abdicans (here designated): ♂, “ Ojai Cal. [ifornia] 3.8.92 [8. iii. 1892 (Fall)] ”, “ abdicans Csy. ”, “ TYPE USNM 39011 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Colpodota repentina (here designated): ♂, “ Pom. [ona, Los Angeles Co.] Cal. [ifornia] Nov. 7.90 [7. xi. 1890 (Fall)] “, “ repentina Csy. ”, “ TYPE USNM 39012 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotype: ♀, “ Pom. [ona, Los Angeles Co.] Cal. [ifornia] Mts. 11.7.91 [7. xi. 1891 (Fall)] ”, “ repentina ­ 2 PARATYPE USNM 39012 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Colpodota laxella (here designated): ♂, “ Cal. [ifornia, Santa Cruz] ”, “ laxella Csy. ”, “ TYPE USNM 39013 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Colpodota pupilla (here designated): ♂, “ Cal. [ifornia, Paraiso Hot Springs, Monterey Co.] ”, “ pupilla Csy. ”, “ TYPE USNM 39015 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Strigota seclusa (here designated): ♀, “ Catalina [Island] CAL. [California] 7.16.94 [16. vii. 1894 (Fall)] ”, “ Eustrigota seclusa Csy. ”, “ TYPE USNM 39048 ” (red label), “ CASEY bequest 1925 ” (NMNH). Additional material. UNITED STATES: Washington: Spokane Co.: ♀, env. of Spokane, the Spokane River, in leaf litter near water, 20. vi. 1998 (V. Yanushev) (SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFE65325CD245046FB16A0D1.taxon	diagnosis	Diagnosis. Acrotona sonomana can be distinguished from other Nearctic species of Acrotona by the distinct shape of the aedeagus (Figs. 139 – 140) and spermatheca (Fig. 141).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFE65325CD245046FB16A0D1.taxon	discussion	Discussion. The types of C. sonomana, C. inceptor, C. abdicans, C. repentina, C. laxella, C. pupilla and St. seclusa are similar in external characters and in the shape of the aedeagus (including the structures of the internal sac) and spermatheca.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFE65325CD245046FB16A0D1.taxon	distribution	Distribution. Acrotona sonomana is known from California and Washington.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFE95326CD245033FEB5A3EA.taxon	diagnosis	Diagnosis. As in Acrotona and Mocyta, in Strigota the pronotal hypomera are invisible in lateral view. However, in Strigota the punctation of abdominal terga is denser, and the pronotum is not as broad as in Mocyta and most examined species of Acrotona. Strigota also has the following unusual characters: in both sexes the basal line of the abdominal tergum 8 laterally joins the base of the tergum (Figs. 142, 145) while in other examined athetines the basal line is always separated from the tergum base (cf. Figs. 107, 110; 135, 137); in both sexes the base of the abdominal tergum 8 has two gland openings (Figs. 142, 145); and the apical margin of the male sternum 8 with a row of microsetae on its dorsal [sic!] side (Fig. 144) (like in Philhygra Mulsant & Rey, 1873).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFE95326CD245033FEB5A3EA.taxon	discussion	Discussion. Casey did not specifically state in his description of Strigota that St. oppidana was the type species of Strigota; however, while designating the type species of Noverota Casey, 1910 a in the same paper, Casey stated (1910 a, p. 90): “ The first species may be regarded as the type, as in all other cases where the type is not specifically named ”. The type species of Strigota (St. oppidana) and Anaduosternum (Ana. brevipennis) are conspecific (see below). Therefore, Anaduosternum is a junior subjective synonym of Strigota.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFEA5320CD24515EFCE5A519.taxon	description	(Figs. 142 – 154)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFEA5320CD24515EFCE5A519.taxon	materials_examined	Type material. Lectotype of Homalota ambigua (here designated): ♀, “ 5507 ”, “ ambigua Er. Am. [erica] spt. Zimm. [ermann] ” (green label), “ Zool. Mus. Berlin ”, “ Typus ” (red label), “ Homalota ambigua Er. [on red side] / Lectotypus Lohse fix. 1983 [on white side] ” [Lohse never published this lectotype designation] (ZMHB). Paralectotypes: 2 ♂♂, 2 ♀♀, “ America Sept. Zimmermann Nr. 5507 ” (green label), “ Zool. Mus. Berlin ”, “ Typus ” (red label) (ZMHB). Lectotype of Strigota oppidana (here designated): ♀, “ N. [ew] Y. [ork, near New York City] ”, “ Strigota oppidana Csy. ”, “ TYPE USNM 39034 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 2 ♂♂, 2 ♀♀, “ N. [ew] Y. [ork, near New York City] ”, “ oppidana PARATYPE USNM 39034 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Strigota gnava (here designated): ♂, “ · N. [orth] C. [arolina, Asheville] ”, “ gnava Csy. ”, “ TYPE USNM 39035 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: ♂, ♀, “ · N. [orth] C. [arolina, Asheville] ”, “ gnava PARATYPE USNM 39035 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Strigota verecunda (here designated): ♂, “ Ia. [Iowa, Cedar Rapids (Brendel)] ”, “ verecunda Csy. ”, “ TYPE USNM 39037 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Strigota assueta (here designated): ♀, “ Mo. · [Missouri, St. Louis] ”, “ assueta Csy. ”, “ TYPE USNM 39038 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Strigota mediocris (here designated): ♀, “ · Tex. [as, El Paso] ”, “ mediocris Csy. ”, “ TYPE USNM 39039 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Strigota vapida (here designated): ♂, “ Salida Colo [rado] Wickham ”, “ vapida ­ 3 PARATYPE USNM 39040 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotype: ♂, “ Cöl. [Colorado, Greeley] ”, “ vapida ­ 2 PARATYPE USNM 39040 ” (red label), “ CASEY bequest 1925 ” (the second paralectotype of St. vapida (♂, “ Coolidge N. [ew] Mex. [ico] ”, “ vapida Csy. ”, “ TYPE USNM 39040 ” (red label), “ CASEY bequest 1925 ” (NMNH )) is not conspecific with St. vapida and belongs to St. seducens Casey, 1910 a or St. perplexa Casey, 1910 a). Lectotype of Strigota inculta (here designated): ♀, “ · Nev. [ada, Reno] ”, “ inculta Csy. ”, “ TYPE USNM 39041 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotypes: 2 ♂♂, ♀, “ · Nev. [ada, Reno] ”, “ inculta PARATYPE USNM 39041 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Strigota placata (here designated): ♂, “ Cal. [ifornia, Santa Cruz] ”, “ placata Csy. ”, “ TYPE USNM 39045 ” (red label), “ CASEY bequest 1925 ” (NMNH). Paralectotype: ♂, “ Cal. [ifornia, Santa Cruz] ”, “ placata PARATYPE USNM 39045 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Strigota recta (here designated): ♀, “ West Haven Ct [Connecticut] 27 June 1905 H. L. Viereck ”, “ recta Csy. ”, “ TYPE USNM 39036 ” (red label), “ CASEY bequest 1925 ” (NMNH). Holotype of Anaduosternum brevipennis: UNITED STATES: New Jersey: ♂, Newark, 24. vii (E. L. Dickerson) (SIIS). Paratype: UNITED STATES: New Jersey: ♂, Newark, 24. vii (E. L. Dickerson) (SIIS). Additional material. UNITED STATES: California: Kern Co.: ♂, ♀, 4 specimens, Lake Isabella, Rd. 155, 35 º 38.37 ' N 118 º 29 ' W, 800 m, Isabella Lake Main Dam, river bank below dam, 12. viii. 1999 (V. I. Gusarov) (KSEM); Kansas: Douglas Co.: 10 specimens, 2.5 km N Lawrence, flood refuse by the Kansas River (right bank), 38 º 59.59 ' N 95 º 14.59 ' W, 200 m, 18. iv. 1999 (V. I. Gusarov) (SPSU); New York: 2 ♀♀, “ L. [ong] I. [sland] ” (NMNH (Casey Collection )); Texas: Brewster Co.: 2 ♀♀, 19 km S Study Butte, 29 º 09.41 ' N 103 º 33.39 ' W, 650 m, Rio Grande, river banks, 31. iii. 2001 (V. I. Gusarov) (SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFEA5320CD24515EFCE5A519.taxon	diagnosis	Diagnosis. Strigota ambigua is very similar to other Nearctic species of Strigota and can be reliably identified only by the distinct shape of the aedeagus (Figs. 149 – 154) and spermatheca (Figs. 147 – 148).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFEA5320CD24515EFCE5A519.taxon	discussion	Discussion. The types of H. ambigua, St. oppidana, St. gnava, St. verecunda, St. assueta, St. mediocris, St. vapida, St. inculta, St. placata, St. recta and Ana. brevipennis are similar in external characters and in the shape of the aedeagus and spermatheca. All these names are considered to be synonyms. Moore and Legner (1975) proposed the name Atheta notmani to replace the name Atheta brevipennis (Notman, 1922) (ex Anaduosternum) which was preoccupied by the name Atheta brevipennis (J. Sahlberg, 1876) (ex Homalota), listed by Bernhauer & Scheerpeltz (1926) as a junior synonym of Atheta (Anopleta) arcana (Erichson, 1839). Atheta notmani is a junior synonym of Strigota ambigua.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFEA5320CD24515EFCE5A519.taxon	distribution	Distribution. Strigota ambigua is widespread across the United States. It is known from New York, Connecticut, New Jersey, North Carolina, Iowa, Missouri, Kansas, Texas, Colorado, New Mexico, Nevada and California.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFEF533CCD24524EFC08A699.taxon	description	(Figs. 155 – 156)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFEF533CCD24524EFC08A699.taxon	materials_examined	Type material. Holotype of Drusilla cavicollis: ♂, “ Sitk ”, “ Drusilla cavicollis Csy. ”, “ TYPE USNM 39494 ” (red label), “ CASEY bequest 1925 ” (NMNH). Additional material. CANADA: Ontario: 2 specimens, env. of Ottawa, sifting, iv – v. 1998 (V. Grebennikov) (SPSU); Québec: ♂, Dorval, 7. x. 1975 (C. Chantal); 3 ♀♀, Mirabel, 12. vii. 1984 (S. Tousignant); ♂, ditto but 20. vii. 1984; ♂, Montréal, 22. ix. 1968 (E. J. Kiteley); ♂, ditto but 31. v. 1969; ♂, ♀, ditto but 1. x. 1969; ♂, 2 ♀♀, ditto but 13. xi. 1969; 5 ♂♂, 3 ♀♀, ditto but 19. iv. 1971; ♂, ditto but 8. v. 1971; ♂, ditto but 21. v. 1971; ♂, ditto but 5. vi. 1971; ♀, ditto but 5. x. 1971; ♀, ditto but 20. v. 1973; ♂, 2 ♀♀, ditto but 12. x. 1975; ♀, ditto but 21. viii. 1977; ♂, ditto but 24. x. 1978; ♀, ditto but 23. v. 1979; ♂, ditto but 24. ix. 1979; ♀, ditto but 27. vii. 1980; 2 ♀♀, ditto but 22. viii. 1980; ♂, ditto but 28. iv. 1980 (A. Smetana); 2 ♂♂, 2 ♀♀, ditto but 5. v. 1980; 2 ♂♂, 4 ♀♀, Lucerne, pitfall in field, 14 – 21. v. 1983 (L. LeSage); ♀, Gatineau Park, Lac des Fées, pitfall, 14 – 20. v. 1982 (L. LeSage); ♀, ditto but 21 – 28. v. 1982 (F. Rickey, L. LeSage); ♂, Abercorn, Cté de Brome (Y. Bousquet) (CNCI); RUSSIA: Chelyabinsk Reg.: 3 specimens, Isakovo, 8. v. 2000 (R. Filimonov) (SPSU); UKRAINE: Kiev Reg.: 3 specimens, env. of Kiev, Pionerskaya metro station, 25. ix. 1988 (V. I. Gusarov) (SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFEF533CCD24524EFC08A699.taxon	diagnosis	Diagnosis. See Lohse (1974 b).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFEF533CCD24524EFC08A699.taxon	discussion	Discussion. The type material of Staphylinus canaliculatus was not examined but this is a well known European species. The holotype of Dr. cavicollis agrees completely with European specimens of Dr. canaliculata in external characters and the shape of the aedeagus, including the sclerites and diverticula of the internal sac (Figs. 155 – 156).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFEF533CCD24524EFC08A699.taxon	distribution	Distribution. In North America Dr. canaliculata is known from New York (Muona 1984), Ontario, Québec, Kentucky and Alaska and appears to be to be an introduction from Eurasia. In Europe and Siberia Dr. canaliculata is a common species in natural habitats. The fact that the only known Alaskan specimen is known from Sitka and Dr. canaliculata was never collected again anywhere in Alaska, despite numerous collecting trips by Smetana, Campbell and myself, suggests that this species was introduced to Sitka by early Russian colonists, perhaps in ship ballast and never was established in continental Alaska. In eastern North America Dr. canaliculata appears to be restricted to disturbed habitats. This fact also supports the introduction hypothesis.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF0533CCD24561CFDF0A51C.taxon	diagnosis	Diagnosis. See Pace (1989).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF0533CCD24561CFDF0A51C.taxon	discussion	Discussion. Pace (1989) redescribed and illustrated most of the Nearctic species of Leptusa described by the previous authors. Unfortunately, he could not obtain the types of some Nearctic species and did not validly designate the lectotypes for some other species. Below I establish some new synonyms and designate the lectotypes for some Nearctic species omitted by Pace.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF1533DCD24524EFB63A43E.taxon	materials_examined	Type material. Lectotype of Leptusa canonica (designated by Pace (1989 )): ♂, “ · Miss. [issipi, Vicksburg] ”, “ canonica Csy. ”, “ TYPE USNM 39583 ” (red label), “ CASEY bequest 1925 ”, “ vidit R. Pace 1981 ” (NMNH). Lectotype of Leptusa tricolor (here designated): ♀, “ Iowa City, Wickham ”, “ tricolor Csy. ”, “ TYPE USNM 39580 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Leptusa nebulosa (here designated): ♀, “ DURY, Cin. [cinnati] Ohio ”, “ nebulosa Csy. ”, “ TYPE USNM 39581 ” (red label), “ CASEY bequest 1925 ” (NMNH). Holotype of Leptusa iowensis: ♀, “ Iowa City, Wickham ”, “ iowensis Csy. ”, “ TYPE USNM 39582 ” (red label), “ CASEY bequest 1925 ”, “ Leptusa canonica Cas. Det. R. Pace 1981 ” (NMNH).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF1533DCD24524EFB63A43E.taxon	diagnosis	Diagnosis. See Pace (1989).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF1533DCD24524EFB63A43E.taxon	discussion	Discussion. The synonymy of Le. canonica, Le. tricolor, Le. nebulosa and Le. iowensis is confirmed. Leptusa obscura Blatchley, 1910, which was synonymized with Le. canonica by Casey (1989), is a valid species (see below).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF1533DCD24524EFB63A43E.taxon	distribution	Distribution. Leptusa canonica is known from Mississippi, Ohio and Iowa.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF1533ECD24550CFB40A129.taxon	materials_examined	Type material. Lectotype of Leptusa obscura (here designated): ♀, “ TYPE ” (red label), “ Marion Co. Ind. [iana] W. S. B. [Blatchley] 3 ­ 17 ­ 98 [17. iii. 1898] ”, “ 2383 ”, “ Purdue Blatchley collection ”, “ Leptusa obscura sp. nov. 4891 ” (PURC). Additional material. UNITED STATES: Pennsylvania: Clinton Co.: ♀, 26 km SEE Lock Haven, 41 º 03.23 ' N 77 º 09.65 ' W, 350 m, Quercus, Tsuga, Pinus, 5. vi. 1999 (V. I. Gusarov) (SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF1533ECD24550CFB40A129.taxon	discussion	Discussion. Pace (1989) synonymized Le. obscura with Le. canonica without having examined the type of the first species. My examination of the lectotype of Le. obscura demonstrated that it is not conspecific with Le. canonica. Leptusa obscura differs from Le. canonica in its uniform dark brown body color, its stronger pronotal punctation and its flatter pronotum. The lectotype of Le. obscura does not match the types of any other Casey species of Leptusa. This species will be redescribed when male specimens become available.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF1533ECD24550CFB40A129.taxon	distribution	Distribution. Leptusa obscura is known from Indiana and Pennsylvania.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF2533FCD24503BFC5EA3E9.taxon	materials_examined	Type material. Lectotype of Leptusa pusio (here designated): ♂, “ Cin. [cinnati] O. [hio] 5.19.6 [19. v. 1906] ”, “ pusio Csy. ”, “ TYPE USNM 39595 ” (red label), “ CASEY bequest 1925 ”, “ vidit R. Pace 1981 ” (NMNH). Paralectotype: “ Cin. [cinnati] O. [hio] 4.18.6 [18. iv. 1906] ”, “ pusio Csy. ”, “ pusio ­ 2 PARATYPE USNM 39595 ” (red label), “ CASEY bequest 1925 ”, “ vidit R. Pace 1981 ” (NMNH).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF2533FCD24503BFC5EA3E9.taxon	diagnosis	Diagnosis. See Pace (1989).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF2533FCD24503BFC5EA3E9.taxon	discussion	Discussion. In his list of examined type material Pace (1989) listed a single male specimen as the holotype of U. pusio. However, in the Casey collection U. pusio is represented by two specimens from the type locality. One specimen (the lectotype) was dissected by Pace and illustrated in Figs. 579 – 582 (Pace 1989). The second specimen (the paralectotype) was dissected by me. My examination of the genitalia preparations of both specimens demonstrated that the illustrations by Pace (1989; Figs. 579, 582) are incorrect. The distal of the two projections on the parameral side of the median lobe illustrated by Pace in Figs. 579 and 582 does not exist. Pace mistook a small piece of dirt encased in the Canada balsam for a projection of the aedeagus. The correct drawing of the median lobe is given in Figs. 157 – 158. The basal of the two projections illustrated by Pace (cf. Fig. 579 in Pace 1989 and Fig. 157 in this paper) does exist. This projection is shared by two other Nearctic species of the subgenus Dysleptusa Pace, 1982 (Le. carolinensis Pace, 1989 and Le. smetanaiella Pace, 1989) (see Figs. 1686, 1691) but not by the Palaearctic species included by Pace in Dysleptusa. The type species of Dysleptusa is Leptusa fuliginosa (Aubé, 1850), a Palaearctic species without a projection. Therefore, the three Nearctic species placed by Pace (1989) in this subgenus may belong to a different lineage of Leptusa.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF2533FCD24503BFC5EA3E9.taxon	distribution	Distribution. Leptusa pusio is known from Ohio.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF35338CD2456CCFE48A641.taxon	materials_examined	Type material. Holotype of Leptusa brevicollis: ♂, “ Pen. [nsylvania] ”, “ Leptusa brevicollis Csy. ”, “ TYPE USNM 39577 ” (red label), “ CASEY bequest 1925 ”, “ vidit R. Pace 1981 ” (NMNH). Holotype of Leptusa laticollis: UNITED STATES: New York: Chautauqua Co.: ♂ [only right middle and posterior legs remain of this specimen], Westfield (H. Notman), 19. v. 1919 (SIIS); paratype: UNITED STATES: New York: Chautauqua Co.: ♀ (allotype), Westfield (H. Notman), 19. v. 1919 (SIIS). Additional material. UNITED STATES: New Hampshire: Grafton Co.: ♂, 12 km E Orford, 43 º 54.04 ' N 71 º 58.96 ' W, 300 m, in forest litter, Acer, Pinus, Betula, moss, ferns, 21. ix. 1998 (V. I. Gusarov) (SPSU); New York: Essex Co.: 2 ♂♂, Keene Valley, 4. vii. 1917 (H. Notman) [although labeled by Notman (?) as paratypes these two males had not been included in the type series (Notman 1921)] (SIIS); Orange Co.: 9 specimens, 8 km S New Windsor, Black Rock Forest, 23 – 25. v. 1998 (V. I. Gusarov) (SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF35338CD2456CCFE48A641.taxon	diagnosis	Diagnosis. See Pace (1989).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF35338CD2456CCFE48A641.taxon	discussion	Discussion. Pace (1989) designated the lectotype of Le. brevicollis, however from the original description (Casey 1893) it is clear that Casey had a single male type of this species. This type is the holotype of Le. brevicollis and Pace's lectotype designation is redundant. Pace (1989) did not have an opportunity to examine the types of Le. laticollis. Unfortunately, the holotype of Le. laticollis, like some other Notman’s staphylinid types deposited at SIIS, is almost completely destroyed. The female paratype and two additional male specimens from New York, identified by Notman as Le. laticollis, are similar to the holotype of Le. brevicollis in external characters and in the shape of the aedeagus.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF35338CD2456CCFE48A641.taxon	distribution	Distribution. Leptusa brevicollis is known from Pennsylvania, New York and New Hampshire.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF45339CD2457E6FD73A781.taxon	materials_examined	Type material. Lectotype of Leptusa elegans (here designated), ♂, “ TYPE ” (red label), “ Pulasky Co. Ind. [iana] W. S. B. [Blatchley] 6 ­ 19 ­ 08 [19. vi. 1908] ”, “ Purdue Blatchley Collection ”, “ Leptusa elegans sp. nov. 4890 ” (PURC); paralectotype, ♀, was mounted on the same card as the lectotype (PURC). Lectotype of Sipalia fontana (here designated): ♂, “ Buena Vista Spg. [Spring] Franklin Co. Pa. [Pennsylvania] ”, “ fontana Csy. ”, “ TYPE USNM 39024 ” (red label), “ CASEY bequest 1925 ” (NMNH); paralectotypes: 2 ♀♀, “ Buena Vista Spg. [Spring] Franklin Co. Pa. [Pennsylvania] ”, “ fontana PARATYPE USNM 39024 ” (red label), “ CASEY bequest 1925 ” (NMNH). Lectotype of Pasilia virginica (designated by Pace (1989 )): ♂, “ Va. [Virginia, Norfolk] ”, “ Pasilia virginica Csy. ”, “ TYPE USNM 39586 ” (red label), “ CASEY bequest 1925 ”, “ Leptusa virginica (Cas.) det. R. Pace 1981 ” (NMNH); paralectotypes: 3 specimens, “ Va. [Virginia, Norfolk] ”, “ virginica PARATYPE USNM 39586 ” (red label), “ CASEY bequest 1925 ”, “ vidit R. Pace 1981 ” (NMNH). Additional material. UNITED STATES: New Jersey: Burlington Co.: ♀, Pine Barrens, Whitesbog, sifting, 30. v. 1999 (A. Berkov) (SPSU); New York: Orange Co.: 5 specimens, 8 km S New Windsor, Black Rock Forest, 23 – 25. v. 1998 (V. I. Gusarov); 7 specimens, ditto but 9. v. 1998 (SPSU); Pennsylvania: Clinton Co.: 12 specimens, 26 km SEE Lock Haven, 41 º 03.23 ' N 77 º 09.65 ' W, 350 m, Quercus, Tsuga, Pinus, 5. vi. 1999 (V. I. Gusarov) (SPSU); Vermont: Windsor Co.: 15 specimens, 8 km SEE Springfield, Hwy. I­ 91, 43 º 14.00 ' N 72 º 26.81 ' W, 250 m, forest litter, Acer, Betula, Quercus, Tsuga, 21. ix. 1998 (V. I. Gusarov) (SPSU).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF45339CD2457E6FD73A781.taxon	diagnosis	Diagnosis. See Pace (1989).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF45339CD2457E6FD73A781.taxon	discussion	Discussion. Pace did not have an opportunity to examine the type of Le. elegans. Without any argumentation he (Pace 1989) transferred this species to the genus Meronera Sharp, 1887. My examination of the lectotype of Le. elegans demonstrated that it belongs to Leptusa and is conspecific with Le. fontana and Le. virginica. Pace (1989) designated the lectotype of Pasilia virginica but failed to appropriately label any of the four types of this species in the Casey collection. Because only one specimen was dissected by Pace it seems reasonable to conclude that this specimen represents the lectotype illustrated by Pace in Figs. 756 – 758. I attached the lectotype label to this specimen. The female specimen from Arkansas illustrated by Pace (1989, Fig. 759) belongs to a closely related but different species. This species will be described elsewhere.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF45339CD2457E6FD73A781.taxon	distribution	Distribution. Leptusa elegans is known from Vermont, New York, New Jersey, Pennsylvania, Virginia and Indiana.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF5533ACD245543FCCFA721.taxon	description	(Figs. 750 – 754 in Pace 1989)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF5533ACD245543FCCFA721.taxon	materials_examined	Type material. Lectotype of Leptusa opaca (designated by Pace (1989 )): ♂, “ Penn. [sylvania] ”, “ Lept. opaca ”, “ TYPE USNM 39579 ” (red label), “ CASEY bequest 1925 ”, “ Leptusa opaca C. det. R. Pace 1981 ” (NMNH); paralectotypes: 2 ♀♀, “ Penn. [sylvania] ”, “ opaca PARATYPE USNM 39579 ” (red label), “ CASEY bequest 1925 ”, “ vidit R. Pace 1981 ” (NMNH). Lectotype of Leptusa seminitens (here designated): ♀, “ N. [ew] Y. [ork] ”, “ Lept. seminitens ”, “ TYPE USNM 39578 ” (red label), “ CASEY bequest 1925 ” (NMNH); Additional material. UNITED STATES: Arkansas: Crawford Co.: 3 specimens, 32 km NE Van Buren, Ozark National Forest, 35 º 39.30 ' N 94 º 06.19 ' W, 500 m, under bark, 16. x. 1999 (V. I. Gusarov); Franklin Co.: ♀, 30 km N Ozark, Ozark National Forest, 35 º 45.32 ' N 93 º 48.77 ' W, 500 m, in forest litter, Quercus, 17. x. 1999 (V. I. Gusarov) (SPSU); Pennsylvania: Franklin Co.: ♂, ♀, Buena Vista Spring (NMNH (Casey collection )); Rhode Island: ♀, Casey farm, Boston Neck, N Kingston (NMNH (Casey collection )); Wisconsin: Bayfield Co.: ♀, Bayfield (Wickham) (NMNH (Casey collection )).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF5533ACD245543FCCFA721.taxon	diagnosis	Diagnosis. See Pace (1989).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF5533ACD245543FCCFA721.taxon	discussion	Discussion. My examination of the lectotypes of Le. opaca and Le. seminitens confirmed the synonymy of the two species established by Fenyes (1920). Pace (1989) designated the lectotype of Le. opaca but failed to appropriately label any of the three syntypes present now in the Casey collection. These three syntypes include one male and two females. Since, in his revision, Pace (1989) indicated that the lectotype is a male, it seems reasonable to conclude that the male syntype represents the lectotype. I attached the lectotype label to this specimen.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF5533ACD245543FCCFA721.taxon	distribution	Distribution. Leptusa opaca is known from Pennsylvania, New York, Rhode Island, Wisconsin and Arkansas. The records from North Carolina (Pace 1989) are based on a closely related undescribed species which will be described elsewhere. The records from Georgia (Pace 1989) need to be reconfirmed.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF6533BCD24550BFB19A0F9.taxon	description	(Figs. 8, 16, 24, 31, 47 – 49, 76 – 78 in Klimaszewski et al. 2001)	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF6533BCD24550BFB19A0F9.taxon	materials_examined	Type material. Holotype of Placusa vaga: ♀, “ Sta. Cruz Mts. Cal. [ifornia] ”, “ vaga Csy. ”, “ TYPE USNM 39576 ” (red label), “ CASEY bequest 1925 ” (NMNH). Holotype of Pseudota cornicula: ♀, “ Metlakatla, B. [ritish] Col. [umbia] Keen ”, “ cornicula Csy. ”, “ TYPE USNM 39161 ” (red label), “ CASEY bequest 1925 ” (NMNH).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF6533BCD24550BFB19A0F9.taxon	diagnosis	Diagnosis. See Klimaszewski et al. (2001).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF6533BCD24550BFB19A0F9.taxon	discussion	Discussion. The holotype of Ps. cornicula agrees with the lectotype of Pl. vaga in external characters and in the shape of the spermatheca. Casey (1911 a) admitted that he was unable to examine the tarsal formula of the holotype of Ps. cornicula (it is in fact 4 ­ 4 ­ 5). This may explain why he failed to recognize that the species belonged to Placusa (tribe Placusini).	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
03F387B2FFF6533BCD24550BFB19A0F9.taxon	distribution	Distribution. Placusa vaga is known from British Columbia and California.	en	Gusarov, Vladimir I. (2003): Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes. Zootaxa 353 (1): 1-134, DOI: 10.11646/zootaxa.353.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.353.1.1
