taxonID	type	description	language	source
BB20213D0230FF8CFF06FDFD7527FF67.taxon	description	(Figs 1 – 3)	en	Aguiar, Alexandre P., Jennings, John T. (2007): New Caledonia as the centre of origin of Parastephanellus Enderlein, with a phylogeny and description of the female of P. khogis Aguiar (Hymenoptera, Stephanidae). Zootaxa 1576 (1): 15-24, DOI: 10.11646/zootaxa.1576.1.2, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.2
BB20213D0230FF8CFF06FDFD7527FF67.taxon	materials_examined	Material examined. 1 ♀ (QMBA), verbatim label data: “ New Caledonia 11513, 21 º 34 ’ S x 166 º 06 ’ E., Col de Petchecara, middle, 28 Jan 2004. G. Monteith. Pyrethrum, trees & logs. ” Triangle mount, complete. Other specimens (QMBA): 3 ♂♂, label data ibidem; ♂ “ New Caledonia: 21 º 34 ’ S x 165 º 46 ’ E, 500 m, Table Unio road, 50453, 14 Nov 2000, C. J. Burwell, rainforest, sweeping. ” Female description. Body length 15.2 mm, ovipositor 9.0 mm; morphometric ratios Ov / Ptl 3.50; Ov / Tt 0.60; Ov / Hdl 8.25; Ov / Smw 9.19; Cxl / h 3.76; Cxl / Fml 0.87; FmSI 2.68; tw / iEE 0.61; btl / w 5.82; Pnl / Smw 0.74; tw / io 1.29; Pnl / Ew 1.13; Gsl / Ptl 1.16; Ptl / T 3 l 2.27; Hdl / Ew 1.69; Ew / tw 1.98; iEE / eEE 0.42; iEE / io 2.12; Ppl / Ew 1.62; io / oo 6.50. Head: postgenal bridge distinctly longitudinally rugulose. Frons finely transversely rugulose, paraocular strigation distinct only narrowly along eye margin, frontal carina absent. Antenna with 26 flagellomeres, first flagellomere ventrally with few tiny, punctate tyloids; flagellomeres 2 – 7 with large oval tyloids, 2 – 5 of them restricted to apical 0.25 – 0.60, flagellomeres 8 – 21 generally with several tyloids occurring along entire length except just one or two on last few of these flagellomeres, absent from last 5. Coronal area from obliquely strigate near ocellum to transverselly strigate towards anterior tubercle. Vertex with about 25 transverse strigae counted centro-longitudinally, strigae varying from coarse anteriorly to fine posteriorly, reaching occipital carina and extending to temple, ending at genal stripe; dorso-longitudinal impression very weak, visible only with tangent illumination. Temple and gena smooth and polished, genal angle in dorsal view distinctly prominent. Mesosoma: prosternum apex alutaceous, then weakly obliquely strigate, posterior half somewhat smooth. Pronotum (Fig. 2) entirely coarsely transversely strigate, on ventral area fine, oblique to longitudinal; neck centrally concave; pronotal fold stout, its anterior margin weakly collapsed and weakly emarginated; semiannular latero-posteriorly with sparse, inconspicuous, shallow punctures, postero-ventral corner subapically with distinct patch of white pilosity; femoral impression narrow, distinct, smooth and polished, ventral area longitudinally strigate. Mesoscutum (Fig. 4) anterior margin (area over which semiannular slides) matte, extremely finely alutaceous, difficult to distinguish, main area strongly transversely rugose; median sulcus and notauli weakly distinct, formed by aligned foveolae mixed in between main sculpturing. Axilla (Fig. 4) concentrically strigate, from strong mesally to fine antero-laterally. Scutellum (Fig. 4) from anteriorly transversely strigate-rugulose to apically finely alutaceous; a few very shallow foveolae laterally. Mesepisternum (Fig. 2) dorsally and on anterior 0.3 covered with dense white short pilosity, hairs centrally and posteriorly sparse; transversely rugulose. Mesopseudosternum obliquely weakly strigate, discrimen deep and crenulate. Propodeum (Fig. 2) densely transversely areolate rugose, this pattern changing from anteriorly, including antero-lateral corners, fine and somewhat alutaceous to posteriorly coarse; parapetiolar fovea indistinct, its position indicated by a broadly sculptured and concave area mesad of spiracle. Spiracular groove and carina (Fig. 2) barely distinct to indistinct, suggested by coalescence of areolae, barely reaching antero-lateral corner; both spiracular plates transversely alutaceous. Metapleuron laterally (Fig. 2) with few delicate, long hairs, from dorsally transversely strigate to laterally broadly areolate, ventrally smooth. Interfoveolar area smooth; post-foveolar area with three strong transverse carinae. Hind coxa transversely strigate, basally coarse, apically fine; hind femur alutaceous, basally very fine, otherwise distinct, this sculpturing about same on all sides; ventrally with three weak tubercles between base and median tooth, three between femoral teeth, and three small ones apical to apical tooth. Hind tibia laterally very strongly and densely alutaceous, more distinct and denser than on femur, matte; posteriorly with distinct, sharply carinate, oblique compression. Wings: front wing vein M + Cu with several setae along entire length, central and basal setae widely spaced, perpendicular to wing, apically with group of 6 setae distant from each other about their own length, last seta inserted past point of connection with vein 1 M; veins 2 r, 2 + 3 R very slightly irregular or sinuous; vein 1 Rs somewhat bent towards wing base; vein 2 Cua nearly entirely tubular, except very apex nebulous, from basally narrow to apically thick, apical 0.3 bent towards wing apex; vein 2 + 3 M tubular, except apex shortly nebulous to spectral, ending clearly before reaching wing margin, but not far from it; pterostigma not elongate, about 5.6 times as long as greatest width, apex curvilineous. Metasoma: petiole transversely strigate, basally coarse, apical 0.1 smooth and polished; ventrally more weakly sculptured than dorsally; T 3 basal end rugulose, then smooth and polished, centrally with very weak, inconspicuous alutaceous; T 4 – 7 with extremely fine and dense transverse strigulation, producing a noticeable dull texture; T 8 alutaceous. Pygidium narrow and emarginate; pygidial impression Y-shaped. Colour: dark brown to black, except as follows. Scape, pedicel and flagellomeres 1 – 3 light brown, pedicel lightest, remainder of antenna dark brown. Mandible, except apex, light brown; clypeus yellow. Face along eye margin with narrow yellow stripe from base of antenna to lateral tubercle; genal stripe pale yellow, reaching and crossing temple (Fig. 2), then bent backwards and widened, ending at occipital carina. Semiannular hind dorsal margin golden yellow. Wing membrane slightly but distinctly infuscate; veins brown. Coronal tubercles, except black apex, wide area behind coronal ocellus, and vertex laterally, along eye margin, red brown (Fig. 2). Mesonotum centrally and scutellum anteriorly and laterally with distinct red brown tone. Fore and mid legs mostly brown to red brown; fore tibia apical 0.1, mid tibia basal 0.2 and apical 0.1, and mid t 1 basal 0.3, pale yellow to whitish; hind leg dark brown, hind tibia basal 0.1 and hind t 1 entirely, white. Ovipositor sheath unicolorous, brown.	en	Aguiar, Alexandre P., Jennings, John T. (2007): New Caledonia as the centre of origin of Parastephanellus Enderlein, with a phylogeny and description of the female of P. khogis Aguiar (Hymenoptera, Stephanidae). Zootaxa 1576 (1): 15-24, DOI: 10.11646/zootaxa.1576.1.2, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.2
BB20213D0230FF8CFF06FDFD7527FF67.taxon	discussion	Discussion. Aguiar & Jennings (2005) stated that the single specimen of P. khogis then known could perhaps prove to be the male of P. delicatus Aguiar or P. mouensis Aguiar, although this was considered unlikely. In fact, the identity of P. khogis is now clearly strengthened by the current study. Further, the results also provide a first indication about the degree of variability between males and females of stephanids in New Caledonia. In the present case, overall variation between the sexes is surprisingly small, occurring far less than that usually observed for stephanids in other biogeographic regions (e. g., see Aguiar 2001). Whether or not this pattern of variability extends to other New Caledonian stephanids, however, remains unknown. Male. The original description (in Aguiar & Jennings 2005) can be complemented as follows: length 5.5 – 11.6 mm. Morphometric ratios (n = 3; underlined values correspond to the holotype) Cxl / h 3.08 – 3.68; Cxl / Fml 0.86 – 0.91; FmSI 1.82 – 2.37; tw / iEE 0.63 – 0.68; btl / w 4.30 – 5.27; Pnl / Smw 0.71 – 0.79 [value of 2.21 published for holotype in Aguiar & Jennings 2005 is incorrect]; tw / io 1.10 – 1.16; Pnl / Ew 1.16 – 1.35; Gsl / Ptl 0.98 – 1.24; Ptl / T 3 l 1.90 – 2.58; Hdl / Ew 1.82 – 1.85; Ew / tw 1.49 – 1.72; EE / eEE 0.44 – 0.47; iEE / io 1.67 – 1.78; Ppl / Ew 1.66 – 1.80; io / oo 6.00 – 7.71. The scutellum, destroyed in the holotype, varies from fully distinctly alutaceous on smaller than average specimens (Fig. 3) to strigate basally, as in the female, for larger than average specimens. The following variation seems to be the most significant. Antenna with 26 flagellomeres, largest specimen with first flagellomere ventrally with a few tiny, punctual tyloids; flagellomere 2 with large oval tyloids apically, otherwise as in holotype. Although showing weak differences in much of their external morphology, the male and female of P. khogis are generally very similar to each other, and more so than usually observed for stephanids (e. g., Aguiar 2001). The most significant differences are as follows: female considerably larger than male, its vertex with fewer strigations than on the holotype (25 vs. 35) but same as in other male specimens; pronotum laterally more coarsely sculptured; scutellum basally with strigation (vs. usually fully alutaceous); mandibles brown (vs. yellow); yellow stripe along eye margin on face narrower and darker than observed for the male holotype, but same as in other male specimens; petiole ventrally strigate (vs. apical 0.7 smooth and polished). The key provided by Aguiar & Jennings (2005) will work adequately for the recognition of both male and female specimens of P. khogis.	en	Aguiar, Alexandre P., Jennings, John T. (2007): New Caledonia as the centre of origin of Parastephanellus Enderlein, with a phylogeny and description of the female of P. khogis Aguiar (Hymenoptera, Stephanidae). Zootaxa 1576 (1): 15-24, DOI: 10.11646/zootaxa.1576.1.2, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.2
BB20213D0230FF8CFF06FDFD7527FF67.taxon	distribution	Distribution. Southern half of New Caledonia (Fig. 1).	en	Aguiar, Alexandre P., Jennings, John T. (2007): New Caledonia as the centre of origin of Parastephanellus Enderlein, with a phylogeny and description of the female of P. khogis Aguiar (Hymenoptera, Stephanidae). Zootaxa 1576 (1): 15-24, DOI: 10.11646/zootaxa.1576.1.2, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.2
BB20213D0235FF8CFF06FE9872B8FC49.taxon	description	Female holotype. The following corrects some imprecision and makes relevant additions to the original description: antenna without any kind of sensilla on flagellomeres 1 – 7 and 28 – 36; flagellomeres 8 – 27 with punctate sensilla (not “ large oval tyloids, ” as in original description), which become progressively less numerous toward apical articles. Frontal carina distinct, particularly with tangent light (not “ indistinct but weak ”). Genal bridge smooth, polished. Hind tibia ventral side of dilated part strongly obliquely carinate on each side, generating a V-shaped pattern with vertices towards apex. Fore tibia uniformly brown.	en	Aguiar, Alexandre P., Jennings, John T. (2007): New Caledonia as the centre of origin of Parastephanellus Enderlein, with a phylogeny and description of the female of P. khogis Aguiar (Hymenoptera, Stephanidae). Zootaxa 1576 (1): 15-24, DOI: 10.11646/zootaxa.1576.1.2, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.2
BB20213D0235FF8CFF06FE9872B8FC49.taxon	discussion	Discussion. The punctate sensilla and their disposition seem to be unique among stephanids, and are therefore also diagnostic for the species. The elongated T 3 also somewhat unusual in this species, and is not observed in other New Caledonian stephanids. Parastephanellus mouensis Aguiar	en	Aguiar, Alexandre P., Jennings, John T. (2007): New Caledonia as the centre of origin of Parastephanellus Enderlein, with a phylogeny and description of the female of P. khogis Aguiar (Hymenoptera, Stephanidae). Zootaxa 1576 (1): 15-24, DOI: 10.11646/zootaxa.1576.1.2, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.2
BB20213D0235FF8CFF06FE9872B8FC49.taxon	description	Female holotype. Genal bridge mostly smooth and polished (not in the original description).	en	Aguiar, Alexandre P., Jennings, John T. (2007): New Caledonia as the centre of origin of Parastephanellus Enderlein, with a phylogeny and description of the female of P. khogis Aguiar (Hymenoptera, Stephanidae). Zootaxa 1576 (1): 15-24, DOI: 10.11646/zootaxa.1576.1.2, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.2
BB20213D0235FF8CFF06FC6D7503FAE2.taxon	description	Male holotype. The following can be added to the original description: postgenal bridge [sculpturing not visible due to orientation of specimen]; femoral impression indistinct, transversely strigate; mesoscutum anterior area strigate; propodeal spiracular groove almost reaching crenulate sulcus; interfoveolar area largely smooth; genal stripe reaching and crossing temple, then bent backwards and widened, ending at occipital carina; fore, mid, brown, hind tibia dark brown; mid basitarsomere brown.	en	Aguiar, Alexandre P., Jennings, John T. (2007): New Caledonia as the centre of origin of Parastephanellus Enderlein, with a phylogeny and description of the female of P. khogis Aguiar (Hymenoptera, Stephanidae). Zootaxa 1576 (1): 15-24, DOI: 10.11646/zootaxa.1576.1.2, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.2
