taxonID	type	description	language	source
6E515339FFED8A3BD5FB7416FD447BB8.taxon	type_taxon	Type species: Atemnus politus (Simon, 1878).	en	Hu, Jun-fang, Zhang, Feng (2012): Notes on two species of the genus Atemnus Canestrini (Pseudoscorpiones: Atemnidae) from China. Journal of Threatened Taxa 4 (11): 3059-3066, DOI: 10.11609/JoTT.o3049.3059-66, URL: http://threatenedtaxa.org/index.php/JoTT/article/view/804
6E515339FFED8A3BD5FB7416FD447BB8.taxon	diagnosis	Diagnosis: See (Beier 1932 a, b; Murthy & Ananthakrishnan 1977).	en	Hu, Jun-fang, Zhang, Feng (2012): Notes on two species of the genus Atemnus Canestrini (Pseudoscorpiones: Atemnidae) from China. Journal of Threatened Taxa 4 (11): 3059-3066, DOI: 10.11609/JoTT.o3049.3059-66, URL: http://threatenedtaxa.org/index.php/JoTT/article/view/804
6E515339FFED8A3BD5FB7416FD447BB8.taxon	discussion	Remarks: Atemnus, the type genus of Atemnidae, is similar to Anatemnus in having a similar pedipalpal patella, but can be distinguished by the trichobothrial pattern on the fixed finger: the distance between est and esb is the same as that between ist and isb in Atemnus, but longer in Anatemnus. The definition of atemnid genera has long been based on external characters only. Klausen (2005) studied the male genitalia of 44 Atemninae and Miratemninae species and concluded that the present generic delimitation was not reliable. As a result, the male genital structures are also explored, in addition to the external characters. The major differences between the two genera Atemnus and Anatemnus can also be found in the male genital organ (Klausen 2005, p. 658, figs 9, 23 – 25).	en	Hu, Jun-fang, Zhang, Feng (2012): Notes on two species of the genus Atemnus Canestrini (Pseudoscorpiones: Atemnidae) from China. Journal of Threatened Taxa 4 (11): 3059-3066, DOI: 10.11609/JoTT.o3049.3059-66, URL: http://threatenedtaxa.org/index.php/JoTT/article/view/804
6E515339FFED8A39D4747126FBE47CCF.taxon	description	(Image 1, Fig. 1) For synonymies see Harvey (2011).	en	Hu, Jun-fang, Zhang, Feng (2012): Notes on two species of the genus Atemnus Canestrini (Pseudoscorpiones: Atemnidae) from China. Journal of Threatened Taxa 4 (11): 3059-3066, DOI: 10.11609/JoTT.o3049.3059-66, URL: http://threatenedtaxa.org/index.php/JoTT/article/view/804
6E515339FFED8A39D4747126FBE47CCF.taxon	materials_examined	Material examined: 02. viii. 2010, five males (Ps. - MHBU-SX 10080201 – 10080205) and five females (Ps. - MHBU-SX 10080206 – 10080210) from China, Shanxi Province, Licheng City, Huangyadong Forest Park (36.800 N & 113.450 E), 757 m, coll. Sheng-tao Guo leg.	en	Hu, Jun-fang, Zhang, Feng (2012): Notes on two species of the genus Atemnus Canestrini (Pseudoscorpiones: Atemnidae) from China. Journal of Threatened Taxa 4 (11): 3059-3066, DOI: 10.11609/JoTT.o3049.3059-66, URL: http://threatenedtaxa.org/index.php/JoTT/article/view/804
6E515339FFED8A39D4747126FBE47CCF.taxon	diagnosis	Diagnosis: Atemnus politus is characterized by the granulate medial face of the pedipalpal femur and patella, the presence of a round dorsal tubercle on the trochanter, and the relatively small size of chela with pedicel (male 1.15 – 1.33 mm, female 1.30 – 1.35 mm).	en	Hu, Jun-fang, Zhang, Feng (2012): Notes on two species of the genus Atemnus Canestrini (Pseudoscorpiones: Atemnidae) from China. Journal of Threatened Taxa 4 (11): 3059-3066, DOI: 10.11609/JoTT.o3049.3059-66, URL: http://threatenedtaxa.org/index.php/JoTT/article/view/804
6E515339FFED8A39D4747126FBE47CCF.taxon	description	Description: Pedipalps and the anterior half of carapace deep brown, tergites yellow (Image 1). Setae on body and legs apically dentate. Carapace smooth (Fig. 1 A), without furrow and longer than broad (1.12 × in male and 1.04 – 1.07 × in female); with distinct eyespots; 38 – 42 setae in total, of which 6 on anterior margin and 6 – 8 on posterior margin; the anterior half darker than the posterior half. Abdomen tergites IV – X incompletely divided, half-tergites with 4 – 6 marginal setae. Tergal chaetotaxy: 6 – 8: 6 – 8: 8: 10 – 12: 10 – 11: 10 – 11: 11 – 12: 12: 12: 10 – 12 (4 T): 8 (2 T): 2; sternites IV – VIII divided, half-sternites with 5 – 8 setae, sternal chaetotaxy (IV – XI): 9 – 11: 11 – 14: 12 – 13: 12 – 14: 12 – 14: 12 – 15: 10 – 12 (4 T): 8 – 10 (4 T): 2. Anterior genital operculum of male (Fig. 1 J) with 9 – 10 setae on each side and 10 on posterior margin; that of female (Fig. 1 H) with 10 – 12 setae on anterior genital operculum and 2 – 3 lyrifissures, and with 9 – 10 setae on posterior margin. Male genital structure (Fig. 1 K): the distal part (l) of lateral apodemes is relatively smaller, and the hooked branch (b) is bowed distally and terminates in a plate-like tip, the proximal part has a distinct dark sclerotized bar (c); the longitudinal fold of medial diverticula (d) has a projection midway along its length; the ejaculatory canal atrium (e) not well-developed; the lateral rods (f) long and diverging proximally; the tip of dorsal apodeme (g) completely joined; the ventral diverticulum (h) bilobed. Female genital structure (Fig. 1 D) simple, spermathecae provided with separated median cribriform plates. Cheliceral hand with four setae, bs and es short and finely dentate, and is and ls long and acute, fixed finger with four retrorse teeth and three small apical teeth, movable finger with a long, tooth-like subapical lobe, male galea (Fig. 1 B) relatively short and with six branchelets and female galea (Fig. 1 C) long and with 5 – 6 distinct branchelets, of which the basal one rebranched, serrula exterior 21 – 25 lamellae, rallum of four setae, only the distal one dentate anteriorly. Pedipalp: trochanter 1.72 – 1.74, femur 2.50 – 2.67, patella 2.19 – 2.23, chela with pedicel 3.03 – 3.09, chela without pedicel 2.82 – 2.91, hand without pedicel 1.68 – 1.70 × longer than broad, and movable finger 0.59 – 0.71 × longer than hand (female trochanter 1.43 – 1.50, femur 2.67 – 2.86, patella 1.97 – 2.09, chela with pedicel 2.70 – 2.77, chela without pedicel 2.46 – 2.55, hand without pedicel 1.50 – 1.55 × longer than broad, and movable finger 0.75 – 0.77 × longer than hand). The prolateral face of pedipalpal femur and patella strongly granulate (Figs. 1 E – F); trochanter with distinct round dorsal protuberance, fixed finger (Fig. 1 G) with 29 – 30 cusped marginal teeth, movable finger with 35 marginal teeth, venom duct short, ramosus nodus between trichobothria et and it, but near et. Trichobothrial pattern: st on movable finger nearly between t than sb; on fixed finger distance of est removed from esb same as ist and isb, it from finger tip nearly equals to distance between ist and isb. Legs fairly stout, without granular, claws simple and longer than arolia, subterminal seta acute and curved. Tactile seta on tarsus IV (Fig. 2 I) situated near the base (TS = 0.12). Dimensions (length / breadth or depth, in mm): Male. Body length: 2.55 – 2.61. Carapace 0.75 – 0.84 / 0.67 – 0.75 (1.12); pedipalp: trochanter 0.40 – 0.43 / 0.23 – 0.25 (1.72 – 1.74), femur 0.70 – 0.80 / 0.28 – 0.30 (2.50 – 2.67), patella 0.70 – 0.78 / 0.32 – 0.35 (2.19 – 2.23), chela with pedicel 1.15 – 1.33 / 0.38 – 0.43 (3.03 – 3.09), chela without pedicel 1.07 – 1.25 (2.82 – 2.91), hand without pedicel 0.64 – 0.73 (1.68 – 1.70), movable finger length 0.43 – 0.52 (0.59 – 0.71 × hand); leg I: femur 0.23 – 0.25 / 0.15 – 0.18 (1.39 – 1.53), patella 0.38 – 0.40 / 0.14 – 0.18 (2.22 – 2.71), tibia 0.40 / 0.10 – 0.11 (3.64 – 4.00), tarsus 0.38 – 0.40 / 0.08 – 0.10 (4.00 – 4.75); leg IV: femur + patella 0.70 – 0.80 / 0.23 – 0.25 (3.04 – 3.20), tibia 0.53 – 0.60 / 0.15 – 0.18 (3.33 – 3.53), tarsus 0.40 – 0.43 / 0.10 – 0.12 (3.58 – 4.00). Female. Body length: 3.38 – 3.47. Carapace 0.80 / 0.75 – 0.77 (1.04 – 1.07); pedipalp: trochanter 0.42 – 0.43 / 0.28 – 0.30 (1.43 – 1.50), femur 0.80 / 0.28 – 0.30 (2.67 – 2.86), patella 0.73 – 0.75 / 0.35 – 0.38 (1.97 – 2.09), chela with pedicel 1.30 – 1.35 / 0.47 – 0.50 (2.70 – 2.77), chela without pedicel 1.20 – 1.23 (2.46 – 2.55), hand without pedicel 0.73 – 0.75 (1.50 – 1.55), movable finger length 0.55 – 0.58 (0.75 – 0.77 × hand); leg I: femur 0.28 – 0.35 / 0.15 – 0.18 (1.87 – 1.94), patella 0.38 – 0.40 / 0.15 – 0.18 (2.22 – 2.53), tibia 0.38 – 0.43 / 0.10 – 0.13 (3.31 – 3.80), tarsus 0.33 – 0.40 / 0.08 – 0.10 (4.00 – 4.13); leg IV: femur + patella 0.73 – 0.80 / 0.23 – 0.25 (3.17 – 3.20), tibia 0.55 – 0.60 / 0.15 – 0.18 (3.33 – 3.67), tarsus 0.43 – 0.45 / 0.10 – 0.12 (3.75 – 4.30).	en	Hu, Jun-fang, Zhang, Feng (2012): Notes on two species of the genus Atemnus Canestrini (Pseudoscorpiones: Atemnidae) from China. Journal of Threatened Taxa 4 (11): 3059-3066, DOI: 10.11609/JoTT.o3049.3059-66, URL: http://threatenedtaxa.org/index.php/JoTT/article/view/804
6E515339FFED8A39D4747126FBE47CCF.taxon	distribution	Distribution: China (Shanxi). Habitat: The specimens of A. politus were collected from leaf litter in a temperate deciduous forest.	en	Hu, Jun-fang, Zhang, Feng (2012): Notes on two species of the genus Atemnus Canestrini (Pseudoscorpiones: Atemnidae) from China. Journal of Threatened Taxa 4 (11): 3059-3066, DOI: 10.11609/JoTT.o3049.3059-66, URL: http://threatenedtaxa.org/index.php/JoTT/article/view/804
6E515339FFED8A39D4747126FBE47CCF.taxon	discussion	Remarks: Atemnus politus was first reported from China by Beier (1967). The male genital structures of the specimens from Shanxi Province fit A. politus illustrated by Klausen (2005). In addition, the dimensions and proportions are rather consistent with those of A. politus given by Beier (1932 a, b; 1963) and Dashdamirov & Schawaller (1992). Based on the external morphological characters and male genital characters, we believe the material from Shanxi, China belonging to A. politus.	en	Hu, Jun-fang, Zhang, Feng (2012): Notes on two species of the genus Atemnus Canestrini (Pseudoscorpiones: Atemnidae) from China. Journal of Threatened Taxa 4 (11): 3059-3066, DOI: 10.11609/JoTT.o3049.3059-66, URL: http://threatenedtaxa.org/index.php/JoTT/article/view/804
6E515339FFEF8A3CD6ED71B1FC127B83.taxon	description	(Image 2, Fig. 2)	en	Hu, Jun-fang, Zhang, Feng (2012): Notes on two species of the genus Atemnus Canestrini (Pseudoscorpiones: Atemnidae) from China. Journal of Threatened Taxa 4 (11): 3059-3066, DOI: 10.11609/JoTT.o3049.3059-66, URL: http://threatenedtaxa.org/index.php/JoTT/article/view/804
6E515339FFEF8A3CD6ED71B1FC127B83.taxon	materials_examined	Type material: 07. v. 2011, holotype male (Ps. - MHBU-HN 11050701) from China, Hainan Province, Mt. Limu (19.170 N & 109.740 E), together with five males (Ps. - MHBU-HN 11050702 – 11050706), nine females (Ps. - MHBU-HN 11050707 – 11050716) paratypes, coll. Junfang Hu leg.	en	Hu, Jun-fang, Zhang, Feng (2012): Notes on two species of the genus Atemnus Canestrini (Pseudoscorpiones: Atemnidae) from China. Journal of Threatened Taxa 4 (11): 3059-3066, DOI: 10.11609/JoTT.o3049.3059-66, URL: http://threatenedtaxa.org/index.php/JoTT/article/view/804
6E515339FFEF8A3CD6ED71B1FC127B83.taxon	etymology	Etymology: The specific name is derived from the type locality.	en	Hu, Jun-fang, Zhang, Feng (2012): Notes on two species of the genus Atemnus Canestrini (Pseudoscorpiones: Atemnidae) from China. Journal of Threatened Taxa 4 (11): 3059-3066, DOI: 10.11609/JoTT.o3049.3059-66, URL: http://threatenedtaxa.org/index.php/JoTT/article/view/804
6E515339FFEF8A3CD6ED71B1FC127B83.taxon	diagnosis	Diagnosis: This new species can be recognized by the following combination of characters: carapace with two distinct eyespots; body in large size (male 4.38 – 4.88, female 5.40 – 6.25); retrolateral face of pedipalpal trochanter, prolateral face of pedipalpal femur and patella, and basal fingers distinctly granulate; dorsal and retrolateral face of trochanter with coniform tubercles; carapace covered with numerous setae.	en	Hu, Jun-fang, Zhang, Feng (2012): Notes on two species of the genus Atemnus Canestrini (Pseudoscorpiones: Atemnidae) from China. Journal of Threatened Taxa 4 (11): 3059-3066, DOI: 10.11609/JoTT.o3049.3059-66, URL: http://threatenedtaxa.org/index.php/JoTT/article/view/804
6E515339FFEF8A3CD6ED71B1FC127B83.taxon	description	Description: Pedipalps and the anterior half of carapace reddish-brown (Image 2), tergites brown. Setae on body and legs apically dentate. Carapace smooth, without furrow, longer than broad (1.16 – 1.20 × in male and 1.20 – 1.32 × in female); with distinct eyespots (Fig. 2 A); 68 – 72 setae, with four on anterior margin and 8 – 10 on posterior margin; the anterior half darker than the posterior half. Abdomen all tergites incompletely divided, halftergites with 4 – 9 marginal setae. Tergal chaetotaxy: 8 – 11: 8 – 11: 9 – 11: 12 – 16: 14 – 18: 16 – 17: 16 – 17: 16 – 17: 16 – 17: 14 – 17 (4 T): 11 – 13 (4 T): 2; sternites divided, half-sternites with 6 – 10 setae, sternal chaetotaxy (IV – XI): 16 – 17: 18 – 19: 17 – 18: 17 – 18: 18 – 20: 16 – 19: 12 – 18 (4 T): 13 (4 T): 2. Anterior genital operculum of male (Fig. 2 G) with 8 – 9 setae on each side and 10 – 11 on posterior margin; that of female (Fig. 2 J) with 8 – 10 setae on anterior genital operculum and 8 – 10 setae on posterior margin. Male genital structure (Fig. 2 K): the hooked branch (b) bowed distally, terminated in a relatively small platelike tip, with a distinct dark sclerotized bar proximally (c); the longitudinal folder of medial diverticula (d) with a projection medially and a prominent extension along the sagittal plane, forming M-shape bulge (k); the ejaculatory canal atrium (e) crescent-shaped on both sides and the distal end of the atrium procurved; the lateral rods (f) long and diverging proximally; the tip of dorsal apodeme (g) completely joined; the ventral diverticulum (h) bilobed. Female genital structure (Fig. 2 I) simple, spermathecae with separated median cribriform plate. Cheliceral hand with four setae (Fig. 2 B), bs and es short and finely dentated, is and ls long and acute, fixed finger with four retrorse teeth and 2 – 3 small apical teeth, movable finger with a long, tooth-like subapical lobe, male galea (Fig. 2 C) relatively short, with 6 – 7 branchelets, female galea (Fig. 2 D) long, with 5 – 6 distinct branchelets, of which the basal one rebranched, serrula exterior 27 – 28 lamellae, rallum (Fig. 2 E) of 4 setae, only the distal one dentated anteriorly. Pedipalp: trochanter 1.10 – 1.30, femur 2.22 – 2.55, patella 2.12 – 2.18, chela with pedicel 2.97 – 3.10, chela without pedicel 2.73 – 2.84, hand without pedicel 1.71 – 1.72 longer than broad, and movable finger 0.59 – 0.63 × longer than hand (female trochanter 1.43 – 1.50, femur 2.27 – 2.38, patella 1.92 – 2.02, chela with pedicel 2.77 – 2.92, chela without pedicel 2.49 – 2.58, hand without pedicel 1.58 – 1.67 longer than broad, and movable finger 0.63 × longer than hand). The retrolateral face of pedipalpal trochanter (Fig. 2 L), prolateral face of pedipalpal femur and patella, and the basal fingers strongly granulated; trochanter with distinct coniform and granulated protuberances, fixed finger (Fig. 2 H) with 29 – 30 cusped marginal teeth, movable finger with 35 – 36 marginal teeth, venom duct short, ramosus nodus medially between trichobothria et and it. Trichobothrial pattern: st on movable finger between t than sb; on fixed finger est at the level of ist, distance of est and esb same as ist and isb, it from finger tip nearly equals to distance between ist and isb. Legs fairly stout and smooth, claws simple, longer than arolia, subterminal seta acute and curved. Tactile seta on tarsus IV (Fig. 2 F) situated proximally (TS = 0.20). Dimensions (length / breadth or depth, in mm): Male: Body length: 4.38 – 4.88. Carapace 1.10 – 1.20 / 0.95 – 1.00 (1.16 – 1.20); pedipalp: trochanter 0.50 – 0.65 / 0.45 – 0.50 (1.10 – 1.30), femur 1.00 – 1.20 / 0.45 – 0.47 (2.22 – 2.55), patella 1.10 – 1.20 / 0.52 – 0.55 (2.12 – 2.18), chela with pedicel 1.80 – 1.90 / 0.58 – 0.64 (2.97 – 3.10), chela without pedicel 1.65 – 1.75 (2.73 – 2.84), hand without pedicel 1.00 – 1.10 (1.71 – 1.72), movable finger length 0.63 – 0.65 (0.59 – 0.63 × hand); leg I: femur 0.33 – 0.35 / 0.25 – 0.27 (1.30 – 1.32), patella 0.58 – 0.60 / 0.23 – 0.25 (2.40 – 2.52), tibia 0.50 – 0.55 / 0.15 – 0.17 (3.24 – 3.33), tarsus 0.38 – 0.45 / 0.12 – 0.14 (3.17 – 3.21); leg IV: femur + patella 1.05 – 1.15 / 0.38 – 0.40 (2.76 – 2.88), tibia 0.75 – 0.80 / 0.22 – 0.25 (3.20 – 3.41), tarsus 0.50 – 0.55 / 0.14 – 0.15 (3.57 – 3.67). Female: Body length: 5.40 – 6.20. Carapace 1.20 – 1.25 / 0.95 – 1.00 (1.25 – 1.26); pedipalp: trochanter 0.50 – 0.60 / 0.35 – 0.40 (1.43 – 1.50), femur 0.95 – 1.00 / 0.40 – 0.44 (2.27 – 2.38), patella 0.95 – 1.00 / 0.47 – 0.52 (1.92 – 2.02), chela with pedicel 1.75 – 1.80 / 0.60 – 0.65 (2.77 – 2.92), chela without pedicel 1.55 – 1.62 (2.49 – 2.58), hand without pedicel 1.00 – 1.03 (1.58 – 1.67), movable finger length 0.63 – 0.65 (0.63 × hand); leg I: femur 0.30 – 0.32 / 0.25 – 0.26 (1.20 – 1.23), patella 0.55 – 0.57 / 0.23 – 0.25 (2.28 – 2.39), tibia 0.50 – 0.55 / 0.15 – 0.18 (3.06 – 3.33), tarsus 0.42 – 0.45 / 0.10 – 0.12 (3.75 – 4.20); leg IV: femur + patella 1.05 – 1.10 / 0.40 – 0.45 (2.44 – 2.63), tibia 0.65 – 0.70 / 0.21 – 0.25 (2.80 – 3.10), tarsus 0.50 – 0.55 / 0.13 – 0.15 (3.67 – 3.85).	en	Hu, Jun-fang, Zhang, Feng (2012): Notes on two species of the genus Atemnus Canestrini (Pseudoscorpiones: Atemnidae) from China. Journal of Threatened Taxa 4 (11): 3059-3066, DOI: 10.11609/JoTT.o3049.3059-66, URL: http://threatenedtaxa.org/index.php/JoTT/article/view/804
6E515339FFEF8A3CD6ED71B1FC127B83.taxon	distribution	Distribution: China (Hainan Island). Habitat: Atemnus limuensis sp. nov. was collected from under an Ormosia tree bark in a humid tropical forest.	en	Hu, Jun-fang, Zhang, Feng (2012): Notes on two species of the genus Atemnus Canestrini (Pseudoscorpiones: Atemnidae) from China. Journal of Threatened Taxa 4 (11): 3059-3066, DOI: 10.11609/JoTT.o3049.3059-66, URL: http://threatenedtaxa.org/index.php/JoTT/article/view/804
6E515339FFEF8A3CD6ED71B1FC127B83.taxon	discussion	Remarks: Atemnus limuensis sp. nov. differs from A. letourneuxi in its large size and the presence of strong granules on the prolateral face of the chelal patella. Atemnus neotropicus from Puerto Rico is known only from a female, which differs from females of A. limuensis sp. nov. in having a small size (body length 4.10 mm), light in color, the presence of weak eyespots and rounded dorsal protuberances on the chelal trochanter. Atemnus strinatii can be distinguished from A. limuensis sp. nov. in having round dorsal tubercles on the chelal trochanter, without eyespots and with weakly granulate chelal femur and patella. Atemnus limuensis sp. nov. differs from A. syriacus by the slender pedipalp (male femur 2.22 – 2.55 ×, patella 2.12 – 2.18 ×, chela with pedicel 2.97 – 3.10 ×, chela without pedicel 2.73 – 2.84 ×, hand without pedicel 1.71 – 1.72 × versus femur 2.1 ×, patella 1.7 ×, chela with pedicel 2.6 ×, chela without pedicel 2.3 × as long as broad in A. syriacus) and the absence of a brown band on carapace. Atemnus politus can be easily be separated from A. limuensis sp. nov. by the slimmer male genital organ (Figs. 1 K, 2 K): the ejaculatory canal atrium (e) is not well-developed and may be aberrant in A. politus; but in A. limuensis sp. nov., the atrium is normal, crescent-shaped on either side, and the distal end of the atrium is typically procurved. Moreover, the distal part (l) of lateral apodemes (a) in A. limuensis sp. nov. is larger than that in A. politus. However, they have many congeneric characters in the male genitalia (the letters refer to those of Figs. 1 K and 2 K): (1) the sclerotized distal part of lateral apodemes; (2) the presence of a hooked branch on proximal part of the lateral apodemes; (3) the presence of a rugose darker area on proximal part of the lateral apodemes; (4) the presence of a round projection on medial diverticulum and a prominent extension along the sagittal plane in M-shaped bulge (k) on two medial diverticula; (5) the long and proximally diverging lateral rods; (6) the distally joined long dorsal apodeme; and (7) the presence of bilobed anterior wall on the ventral diverticulum.	en	Hu, Jun-fang, Zhang, Feng (2012): Notes on two species of the genus Atemnus Canestrini (Pseudoscorpiones: Atemnidae) from China. Journal of Threatened Taxa 4 (11): 3059-3066, DOI: 10.11609/JoTT.o3049.3059-66, URL: http://threatenedtaxa.org/index.php/JoTT/article/view/804
