taxonID	type	description	language	source
C27E87FFFF9E6107FF14DE0E25C3FB1C.taxon	type_taxon	Type species. Acalles nodulosus Boheman 1837: 330, by original designation. Gender masculine.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF9E6107FF14DE0E25C3FB1C.taxon	diagnosis	Diagnosis. Distinguishable among Tylodina by the metepisternum distinctly visible; the elytra clothed with elongate scales, moderately dense to sparse, more dense on the middle of disc and on the base, becoming more scattered toward the flanks; intervals weakly convex, less convex to flat toward the flanks; striae shallow, tubercles round or laminate; internal sac of male genitalia with diminutive round and acute sclerites.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF9E6107FF14DE0E25C3FB1C.taxon	description	Description. Length 2.0 – 11.5 mm. Width 0.9 – 7.0 mm. Body oblong-ovate, 1.8 – 2.7 x longer than wide, clothed with elongate scales, short or long. Head with red, dark red or black integument; scales on vertex dense to contiguous or overlapping, same color as head scales, yellow, light brown or whitish; frons concave to moderately concave; rostrum moderately robust or narrow, surface moderately carinate to rugose, punctures small or large, deep or shallow, longer or as long as head punctures, with an apical area glabrous (male with apical 1 / 2, female with apical 2 / 3 (Figure 12 )), puncticulate (very fine and widely spaced punctures), punctate (fine impressed punctures) or smooth; antennal scape in resting position not reaching anterior margin of eye, funicle with articles 1 and 2 subequal in length, articles 3 – 4 slightly longer than wide, articles 5 – 7 submoniliform; eyes separated by about the width of an eye, moderately convex or flush with outline of head. Pronotum with red or black integument; in lateral view with one of three patterns: 1) outline with anterior constriction and without posterior constriction (Figure 10), 2) outline with anterior and posterior constrictions (Figure 11), and 3) outline slanting downward from highest point to the apex and the base (Figure 9), feebly, moderately or strongly convex; in dorsal view widest at about midlength; with one of two patterns: 1) sides slightly divergent from base to midlength, then convergent to apex, constricted beyond the middle on the sides (Figure 14), or 2) similar to previous pattern but not constricted beyond the middle (Figure 13); apical margin in lateral view extended over hind portion of head; disc with one of four patterns: 1) deeply excavate (Figures 28, 30, 32), 2) distinctly hollowed (Figures 34, 38, 76), 3) somewhat hollowed (Figures 64, 66, 78), or 4) lacking impressions (Figures 44, 48); surface smooth, weakly carinate to rugose, without granules or granulate, with large (Figure 16), medium (Figure 14) or small punctures (Figure 17), deep or shallow, with one of two patterns: 1) contiguous at the base and progressively more widely spaced, smaller and shallower to the apex (Figures 14, 16 and 17), or 2) dense to contiguous on lateral sides, progressively more widely spaced and less conspicuous in the median, anterior and posterior sections (Figure 15); scales sparse, moderately dense to dense, distributed in one of three patterns: 1) light scales (generally yellow, light brown or white) very dense on lateral sides and flanks, anterior area and forming a thick longitudinal fringe along the middle and a narrow strip that diverges from the middle base to the flanks, and with dark scales on the middle and posterior areas of the disc (dark brown or black) (Figures 14, 17), occasionally the pattern is not well defined (Figure 48, 56, 78), 2) similar to last pattern but without dark brown scales (Figure 76), and 3) when excavation is present, scales dense on edge of pronotal excavation, more widely separated towards lateral and median areas of disc (Figures 28, 30, 32); scales dense (Figure 78), moderately dense (Figure 64) to scattered (Figure 76). Elytra with the basal margin straight to very sinuate (Figures 2 – 5); widest just posterior to basal margin then convergent to apex; apex narrowly truncate, clothed with sparse to moderately dense scales, more dense on middle of disc and on base, becoming more scattered toward the flanks; elytral declivity with scales denser on interval 1 – 2 (Figures 56 – 57, 58 – 59, 72 – 73, 76 – 77) or with dense patch on intervals 1 – 3 (Figures 46 – 47, 74 – 75) and on intervals 1 – 5 (Figures 48 – 49, 78 – 79) at base of declivity, base with scales similar those of disc or with light brown or white scales on tubercles or only on first tubercle; integument color with one of three patterns: 1) black (Figures 52 – 63), 2) black with red coloration at the base and along intervals 1 – 2 and on declivity of intervals 1 – 5 (Figures 74 – 77), and 3) black with red coloration at the base and along middle line, elytral declivity and flanks (Figures 44 – 51, 64 – 65). Intervals weakly convex, less convex to flat toward flanks, intervals 2, 3, 7, 8 and 9 convex on declivity near apex; striae shallow; stria 10 complete, incomplete or absent; tubercles on elytra disc variously developed, laminate (Figures 29, 31, 33) or round; declivity with tubercle on interval 4 laminate (Figures 29,31, 33 and 97), acute (Figure 18) or round; tubercle on interval 2 absent, longer, smaller or same size as tubercle in same position at interval 4; surface with or without granules. Scutellar shield absent. Wings absent. Metasternum concave, moderately concave or flat, glabrous on median section or squamous; metepimeron visible; metasternal tooth present (Figure 19) or absent. Abdomen with scales narrow and elongate (Figure 20 – 25), scales wider, longer and dense at sides; male with ventrites 3 – 5 in same plane as ventrites 1 – 2 or not (Figures 6 – 7); female with ventrites 3 – 5 not in same plane as 1 – 2 (Figure 8); ventrite 1 concave, moderately concave, feebly concave or weakly convex, glabrous on median section, squamous or with scattered scales or with an anterior fringe of scales; ventrite 2 feebly concave, concave or weakly convex, longer than or as long as ventrites 3 – 4 combined, glabrous (Figure 25), or with two transverse lines of scales interrupted medially (Figure 23) or not interrupted, female of some species with transverse line of scales interrupted medially (Figure 24); female with ventrites 1 – 2 strongly convex; ventrites 3 – 4 squamous; ventrite 5 convex along median line or convex at anterior 1 / 2 and flat on posterior 1 / 2 or flat, as long or longer than ventrite 1, with one of three patterns: 1) with an apical impression, 2) with apical and lateral impression or 3) without impression, in some species, female with acute projections (Figures 21 – 22). Legs narrowly elongate (3.6 – 5 x longer than wide); with feeble punctures or densely, deeply punctuate throughout, punctures elongate-oval and linearly arranged giving legs (especially femora) a striated appearance, with fine granules between punctures; hind femur with length variable, from not reaching elytral apex to reaching beyond the elytra apex; femora armed with a small, distally inclined tooth ventrally or unarmed; middle and hind tibiae with outer margin subparallel or an external tooth (Figure 26); front tibia straight to sinuate or very deeply excavate apically (Figure 27); premucro present or absent. Genitalia greatly varied, male with median lobe in lateral view with curvature and width varied, from weakly to strongly curved, very slender to stout, apex straight to abruptly curved downward; width of apical process very slender to robust; apical process form almost linear, slightly curved, sinuate or curved upward; occasionally with a longitudinal notch on lateral base. Median lobe length in ventral view from longer than to ½ as long as the length of the apodemes, sides straight or convex near the base (less than of 0.25 of length from the base) to near the middle (> 0.27 to 0.5 of length from the base), convergent (base of median lobe wider than subbasal portion of apex), divergent (subbasal portion of apex wider than base of median lobe) or subparallel (base of median lobe as wide as subbasal portion of apex), apex of median lobe acute (Figure 98 d – e), truncate (Figure 98 c) or blunt (Figure 98 a); apical process constricted beyond the median orifice (Figure 98 a, c, d – e) or not, sides sub-parallel, convergent and sometimes rhomboid-shaped subapically (Figure 98 e). Female with tergite eight trapezoidal or constricted, apex invaginate, obtuse or slightly curved; sternite eight oblong, doliform, or trapezoidal, with the length of median membranous area from 1 / 3 as long as basal plate to subequal in length, spermatheca very broad to very shallowly curved, corpus convex to slightly convex, ramus and nodulus distinct or indistinct.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF9E6107FF14DE0E25C3FB1C.taxon	discussion	Comments. Members of this genus can be confused with other genera of the subtribe Tylodina that also have tuberculate elytra, such as Tylodes, Xenosomus, or Phymatophosus. Species of Tylodinus are most commonly confused with Acalles, because so many species were erroneously originally placed in this genus (Champion 1904, O´Brien and Wibmer 1982); however, the metepisternum of these genera is not externally visible, and this character separates these genera from Tylodinus. Fiedler (1940) placed Tylodinus near Canistes and Pseudoacalles because these genera have the metepisternum distinctly visible, but other characters given in the diagnosis will separate Tylodinus from Canistes and Pseudoacalles. In a forthcoming paper on the phylogenetic relationships of Tylodinus, a number of new genera, closely related to Tylodinus, will be described. Derivation of generic name. C hampion (1905) did not indicate the derivation of the name Tylodinus but it appears to be derived from the combination of two words: Tylodes = derived from Greek “ tylodes ” = tubercle, probably referring to the tubercles on the body of the genus Tylodes and from the Latin suffix inus = like; Tylodinus = like a Tylodes. The name refers apparently to possible confusion between Tylodinus and Tylodes. Natural history. Very few details are available on the natural history of any species of Tylodinus. Adults have been collected almost exclusively in Berlese or Winkler extractions of sifted leaf litter and other ground substrate debris. Some specimens have been collected beating dead vegetation and numbers of larger-sized species have been hand collected on dead wood, apparently in association with Xylariaceae fungi, especially those with the crustose fruiting bodies, which the black, tuberculate body form of the weevil greatly resembles. Immature stages are unknown. Given what is known of the habits of related taxa, it is likely that the larvae mine in dead, decaying wood or other plant debris on the forest floor. Nearly all species are associated with a variety of montane wet forest habitats.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF9E6107FF14DE0E25C3FB1C.taxon	distribution	Distribution. Tylodinus occur throughout the Sierra Madre Oriental from Tamaulipas south to Panama with many species undescribed. At present, 20 species have been described with an additional 32 species described herein, and numerous other new species from other geographic areas are known. Thus, at present there 52 species known to date, 40 are known to occur in México and of these, 38 species have only been recorded from México and two extend from México south into Central America. As discussed, Tylodinus species in Chiapas are well sampled and were collected at elevations from 1080 to 2970 m in 20 municipalities; the species numbers in these municipalities are detailed in Table 1. Checklist of included species	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF976101FF14DBA521A3FAC1.taxon	discussion	Key to species and species groups of Tylodinus Champion from Chiapas	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF966102FF14DE6A21A3FCB0.taxon	description	Recognition. The species of this group are distinguished from other Tylodinus by having the pronotum in lateral view strongly convex, in outline slanting downward from the highest point to the apex (Figure 9), anterior constriction absent; in dorsal habitus, sides slightly divergent from base to midlength, then convergent to apex (Figure 13); disc deeply medially excavate (Figures 28, 30 and 32). Rostrum slender. Elytra with tubercles laminate or elongate (Figures 29, 31, 33).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF966102FF14DE6A21A3FCB0.taxon	diagnosis	Diagnosis. Body 1.8 – 2.1 x longer than wide, black, clothed with obovate, short scales, light brown to ferruginous. Head with dense contiguous scales; vertex with a patch of overlapping scales, yellow or same color as scales of head; rostrum slightly convex, slender, not carinate, surface with fine, shallow punctures, scattered, similar to punctures on head, apical area puncticulate; frons moderately concave, eyes flush with outline of head. Pronotum in lateral view strongly convex, outline slanting downward from highest point to apex and base, anterior constriction absent (Figure 9); in dorsal view with sides slightly divergent from base to midlength, then convergent to apex; disc deeply excavate, with protuberance on median section of disc (two species) or not (one species); surface with punctures more or less uniformly distributed, granulate and rugose; scales dense on edge of pronotal excavation, more widely separated towards lateral and median areas of disc (Figure 13). Elytra black, basal margin strongly sinuate (two species) to slightly sinuate (one species); scales denser on higher tubercles, along sutural interval and interval 2 on declivity, becoming more separated toward the flanks; surface granulate, tubercles laminate (Figures 29, 31, 33). Metasternum moderately concave, with the median section glabrous and smooth (one species) or with scales (two species), metasternal tooth absent. Abdomen with ventrites 3 – 5 not in same plane as ventrites 1 – 2, ventrite 1 moderately concave, squamous, ventrite 2 concave (one species) or weakly concave (two species), as long as ventrites 3 – 4 combined (one species) or longer (two species), squamous (one species) or with transverse line of scales (two species), ventrite 5 as long as ventrite 1, without impressions (two species) or with anterior impression (one species). Femora armed, with fine granules between punctures, punctures feeble on hind femur; hind femora length extending beyond elytral apex; tibiae elongate and narrow, subparallel, slightly curved at apical one-quarter, with large punctures, deep and dense; premucro present or only visible on front tibiae (one species). Male genitalia (Figures 99 – 100) with median lobe (not examined for T. nodulosus) in lateral view stout, weakly curved; in ventral view as long as apodemes, sides straight, convergent or subparallel, apex blunt; apical process in lateral view weakly curved, abruptly curved downward, moderate in width, in ventral view small or moderately large, constricted beyond the median orifice, lateral sides subparallel.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF966102FF14DE6A21A3FCB0.taxon	discussion	Key to species of Tylodinus nodulosus species group	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF956102FF14D8F420B0F88C.taxon	description	(Figures 28 – 29)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF956102FF14D8F420B0F88C.taxon	diagnosis	Diagnosis. Length male 8.6 mm. Width male 5.2 mm. Body 1.8 x longer than wide, clothed with decumbent brown scales and suberect yellow to light brown scales. Rostrum with short scales, as long as or slightly longer than punctures, punctures contiguous on head, becoming more widely spaced towards rostrum, vertex with scales longer and narrower than those of head, yellow to light brown, overlapping; smooth area of rostrum puncticulate. Pronotal disc with broad excavation at anterior section, encompassing most of median section, narrow at posterior section, with protuberance in middle part of pronotal disc, with longitudinal carina along median line that divides protuberance into two parts. Elytra with basal margin strongly sinuate, surface granulate; punctures small, moderately shallow. Elytra with tubercles as follows: I 2 with two small tubercles: first at middle of elytra, second at base of elytral declivity; I 3 with two large, elongate laminate tubercles, first subbasal and second in front of declivity, merged with tubercle on I 2; I 5 with three tubercles: first small and subbasal, second at middle of elytra and slightly longer than tubercle on I 2, third very small, in front of elytral declivity, I 6 with tiny tubercle at base of elytral declivity, I 7 with small subbasal tubercle, I 9 with very small tubercle in same position as tubercle on I 7. Abdomen with suture between ventrites 1 and 2 carinate, ventrite 2 as long as 3 – 4 combined, 3 – 4 convex. Male genitalia not examined.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF956102FF14D8F420B0F88C.taxon	discussion	Comments. We examined a single male of this species from the collections of the Natural History Museum, London. Despite examination of extensive collections from throughout México no other specimens were available. Geographic distribution. México, no other locality data on label. Habitat and elevation: No information. Derivation of the specific name. Boheman (1837) did not indicate the derivation of the specific name nodulosus, but it probably derived from Latin meaning having small nodes or knots.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF956102FF14D8F420B0F88C.taxon	materials_examined	Material examined. Total 1 male (BMNH). “ Mexico ”. No other locality data on label.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF956103FF14DCB92751FB1D.taxon	description	(Figures 9, 30 – 31, 99, 130)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF956103FF14DCB92751FB1D.taxon	diagnosis	Diagnosis. Length male 7.9 – 9.9 mm, female 9.2 mm. Width male 4.6 – 5.0 mm, female 5.0 mm. As for T. nodulosus with following exceptions: body 2.1 x longer than wide. Vertex with two patches of yellow scales. Pronotum without protuberance in middle part of pronotal disc, with longitudinal carinae, sometimes absent or feeble, punctures inconspicuous, granules weak, more widely spaced on posterior area, longer and denser on anterior area. Elytra strongly granulate; with tubercles as follows: I 1 with subbasal swelling; I 2 with two tubercles: first small and elongate, subbasal, second slightly larger and more round than first, at middle of elytra; I 3 with two large, elongate tubercles, first subbasal, second at base of elytral declivity, wide, encompassing interval 4; I 5 with three tiny tubercles, first subbasal, second at middle of elytra and third at base of elytral declivity; I 7 with small subbasal tubercle. Metasternum and ventrite 1 with setigerous scales, ventrite 1 with large punctures and two protuberances forming lobes, less high in females, suture between ventries 1 and 2 carinate, ventrite 2 with large punctures only at anterior margin and on flanks, surface with fine granules. Femora weakly armed, male with front femora unarmed. Male genitalia (Figure 99) in ventral view with lateral sides convergent; apical process 0.08 x as long as median lobe, sides parallel. Female genitalia with tergite eight trapezoidal, apex obtuse; sternite eight trapezoidal, apex not constricted, median membranous area one-half as long as basal plate, spermatheca shallowly curved, outer margin of corpus not curved or slightly curved, ramus indistinct, nodulus slender.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF956103FF14DCB92751FB1D.taxon	discussion	Comments. In 2011 the second author examined a paratype of this species housed in the Landcare Collections in Auckland, New Zealand. It matches the Honduras specimens listed below and we have no doubts as to species identity. Geographic distribution. El Salvador and adjacent Honduras Habitat and elevation: Specimens were collected in cloud forest between 1600 – 1900 m. Derivation of the specific name. Patronym, named by Guillermo Kuschel in honor of Dr. A. Zilch, a weevil collector.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF956103FF14DCB92751FB1D.taxon	materials_examined	Material examined. Total 6 males, 1 female. Honduras: Lempira, Gracias, Cerro de Puca, elev. 1600 m (14 ° 43 ' 22.8 '' N, 88 ° 34 ' 37.2 '' W), 2. vii. 1994, coll. Roberto Cordero — 1 male (CMNC); Ocotepeque, El Portillo, Guisayote, elev. 6000 feet, 7. x. 1993, coll. Robert H. Turnbow Jr. — 1 male (CMNC); same locality, 21 – 22. v. 1995, coll. Roy Morris — 1 male (CMNC); same locality, 26. ix. 1995, coll. Frank T. Hovore — 1 male (CMNC); Guisayote, 21. v. 1995, coll. James E. Wappes — 1 female (CMNC); Reserva Biologica Guisayote, elev. 2180 m (14 ° 25 ' 46.86 '' N, 89 ° 3 ' 44.06 '' W), 2. vii. 1994, coll. B. Ratcliffe, M. L. Jameson, R. Cave — 1 male (CMNC); Sinuapa, Guisayote, 6. viii. 1994, coll. Roberto Cordero — 1 male (CMNC).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF946103FF14DE1C208FF837.taxon	description	(Figures 13, 32 – 33, 100, 130) Diagnosis. Length male 8.3 mm. Width male 4.4 mm. As for T. nodulosus with the following exceptions: body 1.9 x longer than wide. Vertex with scales darker than scales of head. Pronotum with fine granules, scales and granules longer and denser in anterior section, posterior section of lateral area divided forming two small protuberances. Elytra with basal margin slightly sinuate, surface with fine granules, longer toward middle of elytra, striae with punctures small and moderately deep, intervals very weak; tubercle distribution as follows: I 2 with small tubercle at middle of elytra; I 3 with two longer tubercles: first subbasal, second very elongate, laminate, extending from middle of elytra to base of elytral declivity; I 4 with small tubercle at middle of elytra, merged with tubercle on I 3; I 5 and I 6 with tubercle merged, small, situated at middle of elytra; I 7 with small subbasal tubercle. Metasternum and ventrite 1 with setigerous scales, broader at posterior section in ventrite 1; ventrite 5 with a weak anterior impression. Legs with premucro only on front tibiae. Male genitalia (Figure 100) in ventral view with lateral sides subparallel; apical process very reduced, 0.02 x as long as median lobe. Female unknown. Geographic distribution. México (Chiapas) Habitat and elevation. The holotype was hand collected in high elevation cloud forest at 2050 m. Derivation of the specific name. Patronym, named after Robert S. Anderson (Canada) of the Canadian Museum of Nature, Canada.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF946103FF14DE1C208FF837.taxon	materials_examined	Material examined. Total 1 male. Holotype male (CMNC). México: Chiapas, Ángel Albino Corzo, Reserva El Triunfo, Polígono 1, 2050 m (15 ° 39.428 ’ N, 92 ° 48.537 ’ W), 16 – 21. xi. 2001. R. Anderson.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF8A611DFF14DBAA21A3F93C.taxon	diagnosis	Diagnosis. Body 2.0 – 2.3 x longer than wide, black, head and rostrum reddish in some specimens. Head with dark brown scales; with patch of overlapping scales on middle of vertex, whitish, yellow or light brown; frons moderately concave or concave (one species); rostrum moderately robust, rugulate (one species), moderately convex, punctures small and deep, moderately dense, size similar to punctures on head, apical area puncticulate; eyes moderately convex. Pronotum in lateral view moderately or strongly convex (one species), outline with anterior and posterior constriction, these constrictions moderately to strongly depressed (Figure 11); in dorsal view with sides slightly divergent from base to midlength, then convergent to apex, constricted beyond the middle (Figures 14), with longitudinal and moderately deep impression; surface granulate, moderately carinate and / or rugose (one species); punctures contiguous or dense on lateral sides, progressively more spaced, smaller and shallower to the median section, and less conspicuous to the anterior and posterior section (Figure 15); with yellow or light brown scales very dense on lateral sides and flanks, anterior area forming a thick longitudinal fringe along the middle and a narrow strip that diverges from the middle base to the flanks, and with dark brown scales on the middle and posterior areas of the disc. Elytra black; with basal margin moderately to slightly sinuate (one species) (Figures 3 – 4); surface granulate, denser on tubercles; striae 10 complete; elytral scales sparse to moderately dense, dense at base and half of the disc, becoming more separated towards flanks, very dense at intervals of 1 – 2 of the elytral declivity; tubercles in I 2 on declivity smaller than tubercle in I 4 in same position or same size (one species). Metasternum with median section concave, smooth and glabrous; metasternal tooth present or absent (one species). Male abdomen with ventrites 1 – 2 not in the same plane as ventrites 2 – 5; ventrite 1 concave or moderately concave (one species), median section posteriorly smooth and glabrous; ventrite 2 longer than ventrites 3 – 4 combined, slightly concave, with two transverse lines of scales interrupted medially (Figure 23); ventrite 5 as large as ventrite 1, convex or flat in middle (one species), with an impression at anterior middle. Legs with dense scales, femora armed, hind femur almost reaching the elytral apex, front and middle tibiae (especially of male) bent inwards apically, sharply toothed on inner margin beyond middle, the space between tooth and apex deeply excavate (Figure 27); hind tibia with outer margin straight or with external tooth, premucro absent. Male genitalia (Figures 101 – 103) with median lobe in lateral view stout (one species) or moderately stout, weakly curved; in ventral view as long as half-length of apodemes (one species) or equal to apodeme length, lateral sides straight, subparallel, apex truncate or blunt (Figures 98 c, 103); apical process in lateral view weakly curved, not curved downward, in ventral view constricted beyond the median orifice, sides subparallel; tergite eight longer than width; sternite eight elongate. Female genitalia with the apex of the tergite eight straight, trapezoidal, the margin of apex smooth and slightly curved; spermatheca very shallowly curved, outer margin not convex to slightly convex, ramus slender and short, nodulus slender.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF8A611EFF14DC7927CFFC46.taxon	description	(Figures 34 – 35, 101, 130)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF8A611EFF14DC7927CFFC46.taxon	description	Habitat and elevation: Specimens have been collected between 2100 – 2750 m in wet high elevation cloud forest leaf litter and by beating dead branches. Derivation of specific name. Champion (1905) did not indicate the derivation of the specific name cavicrus, but it appears to be derived from two words combined from Latin cavi = hollow, excavated, open; and crus = leg; referring to the peculiar form of the leg.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF8A611EFF14DC7927CFFC46.taxon	materials_examined	Material examined. Total 30 males, 8 females. Holotype male (BMNH) Guatemala: Quiché Mountains 7000 – 9000 feet (Champion). Guatemala: El Progresso, 28 – 29 km N. San Augustin, 7000 – 8500 ’, 19 – 21 April 1990, coll. J. E. Wappes — 1 male, 1 female (CMNC); Cerro Pinalon (15.08392 ° N 89.93013 ° W), elev. 2750 m, 1. v. 2009, LLAMA (Wm-B- 01 - 1 - 04) — 1 male (CMNC); same locality (15.08411 ° N 89.93239 ° W), elev. 2715 m, 1. v. 2009, LLAMA (Wm-B- 01 - 1 - 05) — 1 male (CMNC); same locality (15.08432 ° N 89.93801 ° W), elev. 2640 m, 1. v. 2009, LLAMA (Wm-B- 01 - 1 - 06) — 1 male (CMNC); Cerro Pinalon, Finca Las Nubes (15.08385 ° N 89.94258 ° W), elev. 2574 m, 1 – 5. v. 2009, coll. R. S .. Anderson, cloud forest 2009 - 016 — 24 males, 7 females (BMNH, CMNC, CWOB, ECOS); Baja Verapaz, Biotopo El Quetzal (15.21274 ° N 90.21675 ° W), elev. 1700 – 1900 m, 7 – 10. v. 2009, coll. R. S. Anderson, cloud forest, 2009 - 025 — 1 male (CMNC).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF89611FFF14D8E6216AFDFB.taxon	description	(Figures 98 c, 36 – 37, 102, 130)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF89611FFF14D8E6216AFDFB.taxon	diagnosis	Diagnosis. Length male, 5.0 – 5.2 mm, female, 5.5 – 5.9 mm. Width male, 2.2 – 2.3 mm, female 2.5 – 2.6 mm. Body 2.2 x longer than wide. Head with patch of whitish to yellow or light brown scales on middle of vertex, surface with dense punctures, somewhat irregular; frons concave. Pronotum in lateral view moderately convex; in dorsal view with longitudinal carina near middle, flanks with upper half rugose, lower half with deep and long punctures. Elytra with basal margin sinuate (Figure 4); surface with punctures moderate, deep; tubercles distribution as follows: I 2 with elongate tubercle near base of declivity; I 3 with four tubercles, elongate subbasal tubercle, round tubercle at about midlength, two small elongate tubercles on declivity, one near middle, other near declivity apex; I 4 with two tubercles, elongate, small, between tubercles of I 3, other tubercle high, rounded at base of declivity, a little beyond I 2 (this slightly higher than others); I 5 with three elongate tubercles, first subbasal, second near midlength, both tubercles more or less the same size and shape as those on I 3 in same position and third tubercle little forward of base of declivity, much smaller than previous ones and that of neighboring tubercles, I 6 with two elongate tubercles, small one located between tubercles on I 5, and second near elytral declivity, I 7 with two tubercles, one subbasal and round, other near middle of elytra, elongate, very small; tubercle on I 2 on elytral declivity slightly smaller than that on I 4 in the same position. Metasternal tooth present. Male with ventrite 1 concave; ventrite 5 convex in middle, female with two apical lateral impressions, occasionally ventrite 2 depressed medially. Hind tibia with outer margin excavate, inner margin curved in apical ¼. Male genitalia (Fig. 102) moderately stout in lateral view; length in ventral view as long as apodemes, apex truncate; apical process moderately large. Geographic distribution. México (Chiapas). Habitat and elevation. Specimens were collected from leaf litter in high elevation cloud forest between 2400 – 2910 m. Derivation of specific name. Name derived from combining the Greek word pseudo = false, and the species name cavicrus.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF89611FFF14D8E6216AFDFB.taxon	materials_examined	Material examined. Total 9 males, 6 females. Holotype male (CMNC): México: Chiapas, San Cristóbal de las Casas, Cerro Huitepec (pico) 5 km W San Cristóbal, elev. 2750 m (16 ° 44 ' 33 '' N, 92 ° 41 ' 38.4 '' W), 18. ix. 1991, coll. R. S. Anderson. Paratypes: México: Chiapas, Huixtán, 15 km E San Cristóbal, elev. 2500 m (16 ° 44 ' 48.84 '' N, 92 ° 29 ' 25.3 '' W), 28. v. 2008, coll. LLAMA (Wa-A- 05 - 1 - 32) — 1 male (CMNC); San Cristóbal de las Casas, Cerro Huitepec (pico) 5 km W San Cristóbal, elev. 2750 m (16 ° 44 ' 33 '' N, 92 ° 41 ' 38.4 '' W), 18. ix. 1991, coll. R. S. Anderson — 2 females (CMNC); Cerro Huitepec ca. 5 km W San Cristóbal, elev. 2760 m (16 ° 44 ' 33 '' N, 92 ° 41 ' 38.4 '' W), 14. ix. 2008, coll. R. S. Anderson — 1 female (CMNC); Huitepec, S. Cristóbal, elev. 2480 m (16 ° 45 ' 1.4 '' N, 92 ° 40 ' 57.7 '' W), 29. v. 2001, coll. LLAMA (Wa-A- 05 - 2 - 32) — 1 male (CMNC); Los Alcanfores, elev. 2450 m (16 ° 44 ' 12.47 '' N, 92 ° 40 ' 46.93 '' W), 25. iv. 2006, coll. M. Girón — 1 male (ECOS); same locality, 6. vii. 2006, coll. J. L. Cozar — 1 male (ECOS); Reserva Huitepec, elev. 2500 m (16 ° 44 ' 42.7 '' N, 92 ° 40 ' 52 '' W), 4. viii. 1994, coll. B. Gomez — 1 male (ECOS); same locality, 9. vii. 2007, coll. M. G. Branstetter — 1 male (CMNC); same locality, 17. ix. 1991, coll. R. S. Anderson — 1 female (CMNC); San Juan Chamula, Cerro Tzontehuitz, elev. 2800 m (16 ° 49 ' 10.2 '' N, 92 ° 35 ' 3.8 '' W), 15. vii. 1994, coll. R. Jones — 1 female (ECOS); Tzontehuitz (Pico) ca. 10 km NE San Cristóbal, elev. 2910 m (16 ° 49 ' 10.2 '' N, 92 ° 35 ' 3.8 '' W), 16. ix. 1991, coll. R. S. Anderson — 1 female (CMNC); Cerro Tzontehuitz, elev. 2864 m (16 ° 48 ' 40.8 '' N, 92 ° 35 ' 2.4 '' W), 29. vii. 2005, coll. R. S. Anderson — 1 male (CMNC); Cerro Tzontehuitz, elev. 2677 m (16 ° 48 ' 47 '' N, 92 ° 34 ' 24.8 '' W), 29. vii. 2005, coll. R. S. Anderson — 1 male (CMNC).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF88611FFF14D9A2269AF91C.taxon	description	(Figures 11, 15, 27, 38 – 39, 103, 135)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF88611FFF14D9A2269AF91C.taxon	diagnosis	Diagnosis. Length male, 3.4 – 3.8 mm, female, 3.5 – 4.1 mm. Width male, 1.8 – 2.0 mm, female 1.7 – 2.3 mm. Body 2.0 x longer than wide. Head with white to pale yellow scales on middle of vertex; frons very concave; surface with rough appearance, punctures deep; rostrum strongly carinate. Pronotum in lateral view moderately convex; surface with longitudinal carina at median area. Elytra with basal margin slightly sinuate (Figure 3). Elytra with tubercles as follows: I 2 with tubercle on base of declivity; I 3 with three tubercles, first tubercle sub-basal, second tubercle near middle of elytra and third tubercle at middle of declivity, smaller than the previous tubercle; I 4 with two tubercles, one between tubercles of I 3 on disc, and the other at same position as tubercle on I 2 but smaller; I 5 with two tubercles similar and in the same position as I 3 tubercles on disc; I 6 with one tubercle similar and in the same position as I 4 tubercle; I 7 with two tubercles similar and in the same position as I 5 tubercles, but smaller. Metasternal tooth absent. Male with ventrite 1 moderately concave, ventrite 5 convex at middle area. Hind tibia with external tooth. Male genitalia in lateral view stout (Figure 78); in ventral view as long than half length of apodemes, apex blunt. Female genitalia with sternite eight doliform, with the apex not constricted, median membranous area as long as half the length of the basal plate. Geographic distribution. México (Chiapas). Habitat and elevation. Specimens were collected from leaf litter in cloud forest between 1300 – 2200 m. Derivation of specific name. Name derived from the Latin word rugosus = full of wrinkles, folds or creases.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF88611FFF14D9A2269AF91C.taxon	materials_examined	Material examined. Total 7 males, 9 females. Holotype male (CMNC): México: Chiapas, Villa Flores, Sierra Morena, La Peña, 23. vi. 2006, coll. J. L. Cozar (CMNC). Paratypes: México: Chiapas, Ejido Sierra Morena, R. B. La Sepultura, elev. 1497 m (16 ° 09.856 ' N, 93 ° 36.291 ' W), 13. v. 2008, coll. R. S. Anderson — 1 male, 2 females (CMNC); same locality, Cerro Bola, elev. 1948 m (16.13464 ° N, 93.60077 ° W), 14. v. 2008, coll. R. S. Anderson — 1 female (CMNC); same locality, elev. 1360 m (16.15938 ° N, 93.60495 ° W), 12. v. 2008, LLAMA (Wa-A- 01 - 1 - 11) — 1 female (CMNC); Coapilla, 5 km NE Coapilla, elev. 1990 m (17 ° 10 ' 32.16 '' N, 93 ° 9 '. 36 '' W), 26. v. 2008, coll. M. G. Branstetter — 1 female, 2 males (CMNC); La Concordia, 4 km SE Custepec, elev. 2163 m (15 ° 42 ' 11.5 '' N, 92 ° 55 ' 54.8 '' W), 18. v. 2008, coll. M. G. Branstetter — 1 female (CMNC); same locality (15.71119 ° N 92.92840 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 01) — 1 male (ECOS); 4 km SE Custepec, elev. 2140 m (15 ° 42 ' 38.48 '' N, 92 ° 55 ' 43.07 '' W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 03) — 1 male (ECOS); 4 km SE Custepec, elev. 2125 m (15.70635 ° N, 92.93125 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 34) — 1 male (CMNC); same locality (15.70644 ° N, 92.93124 ° W), LLAMA (Wa-A- 03 - 2 - 32) — 1 female (CMNC); same locality (15.70773 ° N, 92.93111 ° W), LLAMA (Wa-A- 03 - 2 - 02) — 1 female (ECOS); San Cristobal de las Casas, Reserva Huitepec, 10. xi. 1994, coll. R. Jones — 1 female (CMNC).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF886119FF14DC1921A3F835.taxon	diagnosis	Diagnosis. Body approximately 2.1 x longer than wide. Head with yellow (four species), light brown (one species) or yellow to light brown scales (one species), surface with punctures similar to punctures on rostrum; vertex with dense to contiguous scales (four species) or overlapping (two species), uniformly distributed along on vertex and yellow (one species), whitish to light brown (one species), whitish to yellow (two species) or similar in coloration and tonality to head scales (one species); frons concave (one species) or moderately concave; rostrum moderately robust, moderately carinate, apical area punctuate (four species) or puncticulate (two species); eyes moderately convex. Pronotum in lateral view strongly convex (four species) or moderately convex (two species), outline with anterior constriction, without posterior constriction (Figure 10); in dorsal with sides slightly divergent from base to midlength, then convergent to apex, constricted beyond the middle (Figure 14); without impression; surface not to weakly granulate, not carinate, punctures small or larger, dense to contiguous at the base and progressively more widely spaced, smaller and shallower to the apex (Figures 16 – 17); disc clothed with yellow or whitish to light brown scales (one species), very dense on lateral sides and flanks, anterior area forming a thick longitudinal fringe along the middle and a narrow strip that diverges from the middle base to the flanks, and with dark brown or black scales on the middle and posterior areas of the disc (Figure 14). Elytra black, with dark red at the base, along middle line, elytral declivity and flanks (Figures 40 – 51); with basal margin slightly sinuate (Figure 3) or straight (one species) (Figure 2), with dark scales on disc, scales dense and yellow on declivity at intervals 1 – 3 and on elytral base, fringe or band of white scales on tubercles of declivity and on intervals 1 – 7, denser on elytral base and sparsely on rest of elytra; striae 10 complete (four species) or incomplete (two species). Metasternum moderately concave or concave (one species), glabrous; metasternal tooth present (three species) or absent (three species). Male abdomen with ventrites 3 – 5 not in same plane as ventrite 1 – 2 (two species with male unknown), ventrite 1 concave (two species), feebly concave (one species) or convex (one species), median section glabrous (three species), with scatter scales (one species) or squamous (one specie); ventrite 2 longer (one species) or as long as ventrites 3 – 4 combined, median section glabrous (one species), squamous (two species), with a transverse line of scales (two species) or with the line of scales interrupted medially; ventrite 5 convex along median line, as long as ventrite 1, without impressions. Femora armed, hind femora length reaching (three species) or near to elytral apex (three species), tibiae subparallel (four species) or width at baseline (two species), premucro present (three species) or absent (three species). Male genitalia (Figures 104 – 107) with median lobe in lateral view weakly curved, very slender to stout; in ventral view as long as half length (one species) or equal (one species) to apodeme, lateral sides straight (one species) or convex at the sides at 0.27 from the base (two species) or concave near to middle (one species), convergent, apex acute; apical process in lateral view weakly (two species) to slightly curved (two species), abruptly curved downward, in ventral view not constricted beyond the median orifice, lateral sides convergent to apex. Female genitalia with tergite eight constricted (three species) or trapezoidal (two species), with the apex invaginate (one species) or slightly rounded (four species); sternite eight (missing in T. buchanani) oblong (one species), sub- trapezoidal (two species) or trapezoidal (one species), with the median membranous area with the median membranous area 1 / 3 (one species), ½ as long as basal plate (two species) or subapical (one species); spermatheca shallowly (three species) or very shallowly curved (two species), outer margin not convex, ramus indistinct (four species) or long and slender (one species); nodulus indistinct (three species) or slender (two species).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF8D611AFF14DBAA2546FC65.taxon	description	(Figures 40 – 41, 133)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF8D611AFF14DBAA2546FC65.taxon	diagnosis	Diagnosis. Length female, 3.7 mm. Width female, 1.9 mm. Head clothed with yellow scales; vertex with dense to contiguous scales, mostly white with yellow scales intermixed, yellow coloration dominant toward lateral side, uniformly distributed along on vertex; frons concave; rostrum with the apical area punctuate. Pronotum in lateral view strongly convex, disc clothed with yellow and dark brown scales, surface with small punctures. Elytra with basal margin slightly sinuate; tubercles distribution as follows: tubercle on I 2 in front of declivity, I 3 with two tubercles: the first tubercle subbasal and second tubercle on middle of elytra, I 4 with two tubercles: the first very small, positioned between tubercles on I 3, and the second tubercle at declivity as long as tubercle on I 2, I 5 with two tubercles similar in shape and size as tubercles on I 3, I 6 with a very small tubercle in front of declivity, I 7 with two small tubercles at same position as tubercles on I 5; surface granulate, stria 10 incomplete. Metasternum moderately concave; metasternal tooth present. Male unknown. Female abdomen with ventrite 2 squamous. Femora length almost reaching elytral ápex, tibiae subparallel, slightly sinuate. Genitalia with apex of tergite eight invaginate, sternite eight missing, spermatheca shallowly convex, ramus and nodulus indistinct. Geographic distribution. México (Chiapas). Habitat and elevation. The holotype was collected from leaf litter in high elevation cloud forest at 1950 m. Derivation of specific name. Patronym, named after L. L. Buchanan (U. S. A.), a well-known worker on weevils.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF8D611AFF14DBAA2546FC65.taxon	materials_examined	Material examined. Total 1 female. Holotype female (CMNC): México: Chiapas, Unión Juárez, Volcán Tacaná, lower slopes, ca 4 km N, Unión Juarez, elev. 1950 m (15 ° 5 ' 47.29 '' N, 92 ° 5 ' 17.04 '' W), 18. ix. 1992, coll. R. S. Anderson (92 – 109).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF8D611AFF14D8C42123F950.taxon	description	(Figures 16, 42 – 43, 98 b, 104, 132)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF8D611AFF14D8C42123F950.taxon	diagnosis	Diagnosis. Length male, 2.1 mm. Width male, 1.0 mm. As for T. buchanani, with the following exceptions: head with the frons moderately concave, rostrum with apical area puncticulate. Pronotum in lateral view moderately convex; disc with large punctures, scales yellow with intermixed whitish scales. Tubercles on I 2 in front of elytral declivity smaller than tubercle on I 4 at same position, I 6 with very small tubercle between tubercles on I 5, without tubercles on I 7; surface with large punctures; striae 10 complete. Metasternal tooth absent. Male abdomen with ventrites 3 – 5 not in same plane as ventrite 1 – 2, ventrite 1 feebly concave, glabrous, as long as 3 – 4 combined, ventrite 2 with a transverse line of scales, feebly concave, ventrite 5 convex along median line, as long as ventrite 1 without impressions. Femora armed, hind femora length reaching elytral apex, front tibiae with apical ¼ curved, middle and hid tibia subparallel and slightly curved. Male genitalia (Figure 104) in lateral view stout, as long as half apodeme length; in ventral view with lateral sides straight, convergent; apical process weakly curved. Female unknown. Geographic distribution. México (Chiapas). Habitat and elevation. The holotype was collected from leaf litter in high elevation cloud forest at 2000 m. Derivation of specific name. Name derived from Latin word exiguus = small, scanty, short.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF8D611AFF14D8C42123F950.taxon	materials_examined	Material examined. Total 1 male. Holotype male (CMNC): México: Chiapas, Motozintla, 7 km SSW Motozintla de Mendoza, elev. 2000 m (15 ° 19 ' 40.8 '' N, 92 ° 16 ' 34.7 '' W), 17. ix. 1992, coll. R. S. Anderson.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF8D611BFF14DDDB20A3FDDE.taxon	description	(Figures 44 – 45, 105, 131)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF8D611BFF14DDDB20A3FDDE.taxon	diagnosis	Diagnosis. Length male, 2.4 – 2.7 mm, female, ca. 2.8 mm (damaged). Width male, 1.1 – 1.2 mm, female ca. 1.3 mm (damaged). As for T. exiguus with following exceptions: head with dark brown scales; vertex with yellow scales. Pronotum in lateral view moderately convex; surface with small punctures. Metasternum concave. Ventrite 1 concave with scattered scales, ventrite 2 with line scales interrupted medially. Hind femur length extending near to elytral apex, front tibiae with apical 1 / 4 curved, middle and hind tibiae parallel. Male genitalia (Figure 105) with median lobe in lateral view very slender; in ventral view as long as apodemes, lateral sides convex beyond 0.27 of length from the base; apical process slightly curved. Female with sternite eight sub-trapezoidal, with the membranous median area ½ as long as basal plate, ramus indistinct, nodulus slender. Geographic distribution. México (Chiapas). Habitat and elevation. Specimens were collected from leaf litter in high elevation cloud forest at 2000 m. Derivation of specific name. Noun in apposition: after the word ixchel, Mayan name of the Moon goddess, also the goddess of childbirth, medicine and of rainbows.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF8D611BFF14DDDB20A3FDDE.taxon	materials_examined	Material examined. Total 5 males, 1 female. Holotype male (CMNC): México: Chiapas, Unión Juárez, Volcán Tacaná, lower slopes, ca 4 km N, Unión Juárez, elev. 2000 m (15 ° 5 ' 47.29 '' N, 92 ° 5 ' 17.04 '' W), 20. ix. 1992, coll. R. S. Anderson. Paratypes: Same data as holotype, 4 males, 1 female (CMNC, ECOS).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF8C611BFF14D9582063F870.taxon	description	(Figures 17, 46 – 47, 106, 134)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF8C611BFF14D9582063F870.taxon	diagnosis	Diagnosis. Length male, 3.2 – 4.2 mm, female, 3.4 – 3.8 mm. Width male, 1.6 – 2.3 mm, female 1.8 – 2.1 mm. As for T. buchanani with the following exceptions: head with scales on vertex similar to head scales. Tubercles on I 2 in front of elytral declivity, very small, I 4 with tubercle on declivity, longer than tubercle on I 2, occasionally small tubercles between tubercles on I 3 present, without tubercles on I 6, I 7 with subbasal tubercle; striae 10 incomplete; elytral declivity with lateral black polished area. Male abdomen with ventrite 1 concave, median section glabrous, ventrite 2 concave, as long as 3 – 4 combined, concave, median section glabrous. Hind femur length not reaching elytral apex, premucro absent. Male genitalia (Figure 106) with median lobe in lateral view slender; in ventral view as long as apodemes, laterally convex at sides beyond 0.27 of length from the base; apical process in lateral view slightly curved. Female genitalia with tergite eight constricted, apex slightly curved, sternite eight trapezoidal, median area membranous, 1 / 3 as long as basal plate. Geographic distribution. México (Chiapas). Habitat and elevation. Specimens were collected from leaf litter in high elevation cloud forest between 1800 – 2220 m. Derivation of specific name. Patronym, named after Robert W. Jones (México) of the Universidad Autónoma de Querétaro, Mexico.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF8C611BFF14D9582063F870.taxon	materials_examined	Material examined. Total 18 males, 5 females. Holotype male (CMNC): México: Chiapas, Angel Albino Corzo, Reserva de la Biosfera el Triunfo, Campamento el Quetzal, 1800 m (15 ° 43.239 ' N, 92 ° 56.405 ' W) 18. vii. 2007, coll. R. S. Anderson. Paratypes: México: Chiapas, La Concordia, 3.5 km ESE Custepec, Campamento El Quetzal, elev. 1800 m (15 ° 43 ' N, 92 ° 56 ' W), 17. vii. 2007, coll. M. G. Branstetter — 2 males (CMNC); 4 km SE Custepec, elev. 2140 m (15 ° 42 ' 38.48 '' N, 92 ° 55 ' 43.07 '' W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 03) — 1 male (CMNC); same locality, 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 02) — 1 male (CMNC); 4 km SE Custepec, elev. 2125 m (15 ° 42 ' 27.7 '' N, 92 ° 55 ' 52 '' W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 02) — 1 female (CMNC); same locality (15.71049 ° N 92.92872 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 18) — 1 female (CMNC); same locality (15.70671 ° N 92.93127 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 26) — 1 male (CWOB); same locality (15.70729 ° N 92.93123 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 12) — 1 male (CMNC); same locality (15.70738 ° N 92.93121 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 10) — 3 males (CMNC, ECOS); same locality (15.70751 ° N 92.93117 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 07) — 1 male (CMNC); same locality (15.71107 ° N 92.92846 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 04) — 1 male (CMNC); same locality (15.71111 ° N 92.92844 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 03) — 1 male (CMNC); same locality (15.71053 ° N 92.92870 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 17) — 1 male (ECOS); same locality (15.71061 ° N 92.92867 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 15) — 1 male (ECOS); same locality (15.71107 ° N 92.92846 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 04) — 1 male (ECOS); Reserva de la Biósfera El Triunfo, Campamento El Quetzal, elev. 2160 m (15 ° 42 ' 18.4 '' N, 92 ° 55 ' 53.8 '' W), 22. v. 2008, coll. R. S. Anderson — 2 males (BMNH, CMNC); same locality elev. 2220 m (15.70819 ° N 92.93070 ° W), 21. v. 2008 — 2 females (CMNC); same locality elev. 2150 m (15.70775 ° N 92.93121 ° W), 21. v. 2008 — 1 female (ECOS).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF826117FF14D87027D0FF3E.taxon	description	(Figures 2, 50 – 51, 107, 132)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF826117FF14D87027D0FF3E.taxon	diagnosis	Diagnosis. Length male, 3.4 – 4.0 mm, female, 3.2 – 4.0 mm. Width male, 1.6 – 1.7 mm, female 1.5 – 2.0 mm. As for T. buchanani with the follow exceptions: body clothed with short scales, light scales on vertex and pronotum frequently whitish, but in some specimens light brown. Head with overlapping scales on vertex; front moderately concave. Elytra with basal margin straight (Figure 2), stria 10 complete, surface granulate, tubercles similar to T. buchanani. Metasternal tooth absent. Abdomen squamous, scales setigerous, ventrite 1 convex; ventrite 2 longer than 3 – 4 combined, convex. Hind femora length almost reaching elytral apex, front tibiae sinuate, middle and hind tibiae subparallel, premucro very small, hind tibiae with apical ¼ curved. Male genitalia (Fig. 107) in lateral view slender; in ventral view with lateral sides concave, subparallel; apical process in lateral view weakly curved. Female genitalia with tergite eight trapezoidal, apex slightly curved, sternite eight oblong, median membranous area subapical; spermatheca shallowly curved, ramus long and slender, nodulus slender. Geographic distribution. México (Chiapas). Habitat and elevation. Specimens were collected from leaf litter in high elevation cloud forest between 2300 – 2750 m. Derivation of specific name. Name derived from the Latin word variabilis = variable.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF826117FF14D87027D0FF3E.taxon	materials_examined	Material examined. Total 95 males, 90 females. Holotype male (CMNC): México: Chiapas, San Cristóbal de las Casas, Cerro Huitepec (pico) 5 km W San Cristóbal, elev. 2750 m (16 ° 44 ' 33 '' N, 92 ° 41 ' 38.4 '' W), 15. ix. 1991, coll. R. S. Anderson. Paratypes: México: Chiapas, Huixtán, 15 km E San Cristóbal, elev. 2500 m (16 ° 44 ' 49.6 '' N, 92 ° 29 ' 24.22 '' W), 28. v. 2008, coll. LLAMA (Wa-A- 05 - 1 - 24) — 1 female (CMNC); same locality (16.74717 ° N 92.49017 ° W), 28. v. 2008, coll. LLAMA (Wa-A- 05 - 1 - 48) — 1 female (CMNC); same locality (16.74698 ° N 92.49013 ° W), 28. v. 2008, coll. LLAMA (Wa-A- 05 - 1 - 16) — 1 male (CMNC); Bazom, elev. 2450 m (16 ° 44 ' 18.96 '' N, 92 ° 29 ' 18.24 '' W), 9. vii. 2003, coll. R. S. Anderson — 1 male (CMNC); San Cristobal de las Casas Airport, elev. 2350 m (16 ° 41 ' 25.08 '' N, 92 ° 31 ' 48 '' W), 25. vii. 2005, coll. R. S. Anderson — 1 female (CMNC); San Cristóbal de las Casas, Cerro Huitepec (pico) 5 km W San Cristóbal, elev. 2750 m (16 ° 44 ' 33 '' N, 92 ° 41 ' 38.4 '' W), 15. ix. 1991, coll. R. S. Anderson — 6 males, 6 females (CMNC); same locality, 18. ix. 1991, coll. R. S. Anderson — 4 males, 2 females (CMNC); Cerro Huitepec 5 km W San Cristóbal, elev. 2650 m (16 ° 44 ' 37 '' N, 92 ° 41 ' 23.3 '' W), 17. ix. 1992, coll. R. S. Anderson — 2 males, 5 females (BMNH, CMNC); Corazón de María, elev. 2449 m (16 ° 41 ' 7.4 '' N, 92 ° 32 ' 11.4 '' W), 8. vi. 2006, coll. J. L. Cozar — 1 male (ECOS); Huitepec, elev. 2466 m (16 ° 44 ' 54.2 '' N, 92 ° 41 ' 6.4 '' W), 5. v. 2006, coll. J. L. Cozar — 2 males, 2 females (ECOS); same locality, 15. vi. 2006, coll. J. L. Cozar — 1 male (ECOS); same locality, 3. vii. 2006, coll. J. L. Cozar — 2 males (ECOS); same locality (16.75181 ° N 92.68270 ° W), elev. 2480 m, 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 01) — 2 males, 1 female (CWOB); same locality (16.75104 ° N 92.68270 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 18) — 1 male, 1 female (CMNC); same locality (16.75109 ° N 92.68270 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 17) — 1 female (CMNC); same locality (16.75131 ° N 92.68270 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 12) — 1 female (CMNC); same locality (16.75176 ° N 92.68270 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 02) — 3 males, 1 female (CMNC); same locality (16.75163 ° N 92.68270 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 05) — 3 males, 5 females (CMNC, ECOS); same locality (16.74689 ° N 92.48990 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 1 - 01) — 1 male (CMNC); same locality (16.75018 ° N 92.68280 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 39) — 2 males (CMNC); same locality (16.75018 ° N 92.68284 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 40) — 1 male (CMNC); Huitepec Ocotal, Cerro La Cruz, elev. 2970 m (16 ° 44 ' 31.2 '' N, 92 ° 41 ' 37.7 '' W), 12. vi. 2006, coll. J. L. Cozar — 3 males, 2 females (ECOS); Huitepec, 5 km W San Cristóbal, elev. 2750 m (16 ° 44 ' 33 '' N, 92 ° 41 ' 38.4 '' W), 14 - 25. ix. 1992, coll. R. S. Anderson — 2 females (CMNC); same locality, 14. ix. 1992, coll. R. S. Anderson — 5 males, 2 females (CMNC); Huitepec, S. Cristóbal, elev. 2480 m (16.75041 ° N 92.68270 ° W), 29. v. 2001, coll. LLAMA (Wa-A- 05 - 2 - 32) — 1 male, 1 female (CMNC); same locality (16.75032 ° N 92.68270 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 34) — 2 females (ECOS); same locality (16.75045 ° N 92.68270 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 31) — 1 male, 2 females (CMNC); same locality (16.75163 ° N 92.68270 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 05) — 1 female (CMNC); same locality (16.75050 ° N 92.68270 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 30) — 1 male, 1 female (ECOS); same locality (16.75032 ° N 92.68270 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 34) — 1 female (CMNC); same locality (16.75077 ° N 92.68270 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 24) — 1 female (CMNC); same locality (16.75018 ° N 92.68298 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 43) — 1 female (CMNC); same locality (16.75018 ° N 92.68303 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 44) — 1 female (CMNC); same locality (16.75018 ° N 92.68308 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 45) — 1 male (CWOB); same locality (16.75018 ° N 92.68317 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 47) — 1 male (CMNC); same locality (16.75018 ° N 92.68327 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 49) — 1 female (CMNC); same locality (16.75054 ° N 92.68270 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 29) — 1 male (CMNC); same locality (16.75082 ° N 92.68270 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 23) — 1 male (CMNC); Los Alcanfores, elev. 2450 m (16 ° 44 ' 12.47 '' N, 92 ° 40 ' 46.93 '' W), 25. iv. 2006, coll. M. Girón — 1 male (ECOS); Reserva Huitepec, elev. 2400 m (16 ° 44 ' 12.5 '' N, 92 ° 40 ' 46.9 '' W), 18. xi. 1993, coll. R. Jones — 4 males, 1 female (ECOS, UAQM); same locality, 18. xi. 1993, coll. R. Jones — 1 male (ECOS); same locality, 16. ii. 1994, coll. R. Jones — 1 male (ECOS); same locality, 16. ii. 1994, coll. R. Jones — 2 males, 2 females (ECOS); same locality, 7. iv. 1994, coll. R. Jones — 1 females (ECOS); same locality, 27. v. 1994, coll. B. Gómez-G y O. L. Gómez-N — 2 males (ECOS); same locality, 25. viii. 1994, coll. B. Gómez-G — 2 males (ECOS); same locality, 26. x. 1994, coll. B. Gómez-G — 1 male, 1 female (ECOS); same locality, 24. i. 1995, coll. R. Jones y B. Gómez-G — 1 female (ECOS); same locality, 24. i. 1995, coll. R. Jones — 1 male (ECOS); same locality, 24. i. 1995, coll. R. Jones-W y B. Gómez-G — 1 male (ECOS); same locality, 27. iv. 1995, coll. B. Gómez-G y M. Girón-I — 1 female (ECOS); same locality, 14. vi. 1995, coll. B. Gómez- G, M. Girón-I y A. Méndoza-R — 1 male (ECOS); same locality, 27. vi. 1995, coll. B. Gómez-G y M. Girón-I — 1 female (ECOS); same locality, 5. vii. 1995, coll. M. Girón — 1 male (UAQM); same locality, 5. vii. 1995, coll. R. Jones — 1 male (UAQM); same locality, 10. vii. 1995, coll. R. Jones — 1 female (UAQM); same locality, 25. vii. 1995, coll. R. W. Jones — 4 males, 3 females (ECOS, UAQM); same locality, 1. viii. 1995, coll. A. Mendoza y M. Girón — 1 male (UAQM); same locality, 15. viii. 1995, coll. M. Girón — 1 female (UAQM); same locality, 3. xi. 1995, coll. B. Gómez y M. Girón — 1 male, 2 females (UAQM); same locality, 14. xi. 1995, coll. B. Gómez, M. Girón — 1 male, 1 female (UAQM); same locality, 5. xii. 1995, coll. B. Gómez y M. Girón — 1 male, 2 females (UAQM); same locality, 6. ii. 1996, coll. L. Ruiz-M y M. Girón-I — 3 males, 4 females (ECOS, UAQM); same locality, 14. ii. 1996, coll. L. Ruiz-M y M. Girón-I — 1 male (ECOS); same locality, 14. ii. 1996, coll. L. Ruiz-M y M. Girón-I — 2 males, 2 females (UAQM); same locality, 7. vii. 2003, coll. R. S. Anderson — 1 female (CMNC); same locality, 12. ii. 2004, coll. J. L. Cozar, M. Girón — 1 female (ECOS); same locality, 9. vi. 2004, coll. J. L. Cozar, M. Girón — 1 male, 1 female (ECOS); same locality, 11. vi. 2004, coll. J. L. Cozar — 1 female (ECOS); same locality, 2. vii. 2004, coll. J. L. Cozar — 1 female (ECOS); same locality, 12. vii. 2004, coll. J. L. Cozar — 1 female (ECOS); same locality, 13. vii. 2004, coll. M. Girón — 1 male (ECOS); same locality, 15. vii. 2005, coll. R. S. Anderson — 1 male (CMNC); same locality, 9. vi. 2006, coll. U. Caballero — 1 female (ECOS); same locality, 12. vi. 2006, coll. J. L. Cozar — 1 female (ECOS); same locality, 17. vi. 2006, coll. U. Caballero — 3 females (ECOS); same locality, 11. vii. 2007, coll. J. Longino — 1 female (CMNC); same locality, 9. vii. 2007, coll. R. S. Anderson — 4 males, 4 females (CMNC); same locality, 9. vii. 2007, coll. M. G. Branstetter — 2 females, 1 male (CMNC); same locality, 11. vii. 2007, coll. R. S. Anderson — 3 males, 2 females (CMNC); Zoológico, elev. 2379 m (16 ° 43 ' 5.5 '' N, 92 ° 42 ' 9.7 '' W), 6. vi. 2006, coll. J. L. Cozar — 1 male (ECOS); San Juan Chamula, Cerro Tzontehuitz, elev. 2800 m (16 ° 49 ' 10.2 '' N, 92 ° 35 ' 3.8 '' W), 15. vii. 1994, coll. R. Jones — 1 male (UAQM).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF826115FF14DBAA2123FD29.taxon	description	(Figures 48 – 49, 132)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF826115FF14DBAA2123FD29.taxon	diagnosis	Diagnosis. Length female, 3.8 mm. Width female, 1.9 mm. As for T. buchanani with the following exceptions: head with yellow scales, vertex with scales similar in coloration and tonality as head scales; frons moderately concave. Elytra with basal margin sinuate, tubercles similar to T. jonesi, surface with fine granules, stria 10 complete. Metasternal tooth weak. Abdomen with ventrite 2 with a transverse line of scales. Hind femur length extended near to elytral apex, front tibiae sinuate, narrowly carinate on internal margin, middle and hind tibiae subparallel, wide at 1 / 3 basal on external margin, premucro present. Male unknown. Female genitalia with tergite eight constricted, apex slightly curved, sternite eight sub-trapezoidal, median membranous area ½ as long as length of basal plate, spermatheca very shallowly curved, ramus and nodulus indistinct. Geographic distribution. México (Chiapas). Habitat and elevation. The holotype was collected from leaf litter in high elevation cloud forest at 2000 m. Derivation of specific name. Patronym, named after Guillermo J. Wibmer (U. S. A.), a well-known worker on weevils.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF826115FF14DBAA2123FD29.taxon	materials_examined	Material examined. Total 1 female. Holotype female (CMNC): México: Chiapas, Motozintla, 7 km SSW Motozintla de Mendoza, elev. 2000 m (15 ° 19 ' 40.8 '' N, 92 ° 16 ' 34.7 '' W), 17. ix. 1992, coll. R. S. Anderson.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF806117FF14DA7E21A3FA0C.taxon	diagnosis	Diagnosis. Body black, approximately 2.2 x longer than wide. Head with brown or dark brown scales, vertex with white scales or scales similar in coloration and tonality to head scales, dense to contiguous or overlapping, distributed along vertex or in a patch on middle of vertex; frons concave or moderately concave; rostrum moderately robust, surface with large, deep punctures, longer than or similar to punctures on head, carinate or moderately carinate, apical area punctuate, eyes moderately convex or flush with outline of head. Pronotum in lateral view moderately convex, outline with anterior constriction, without posterior constriction (Figure 10); in dorsal view with sides slightly divergent from base to midlength, then convergent to apex, constricted beyond the middle (Figure 14), somewhat hollowed longitudinally; punctures contiguous or dense on lateral sides, progressively more widely spaced, smaller and shallower towards median section, less conspicuous towards anterior and posterior sections (Figure 15); whitish, moderately dense scales on lateral sides and flanks, anterior area forming thick longitudinal fringe along middle and narrow strip that diverges from middle base to flanks, and with dark brown scales on middle and posterior areas of disc; surface without granules. Elytra black, with basal margin slightly sinuate (Figure 3), with dense whitish scales subbasally and on tubercles at elytral declivity and widely scattered on middle of elytra, with patch of white scales on tubercles in front of elytra declivity or not; striae 10 absent. Metasternum flat or moderately concave, median section glabrous; metasternal tooth absent. Abdomen with ventrite 2 glabrous or with transverse line of scales, flat or convex along median line. Hind femur length not reaching elytra apex, femora armed, tibiae subparallel or wider towards base, premucro absent. Female with tergite eight trapezoidal, apex slightly curved, sternite eight doliform, apex not constricted, median membranous area 2 / 3 as long as basal plate length, spermatheca very shallowly curved, with the outer margin not convex, ramus and nodulus indistinct.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF806111FF14DD292601FE4E.taxon	description	(Figures 18, 52 – 53, 134)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF806111FF14DD292601FE4E.taxon	diagnosis	Diagnosis. Length female 2.3 – 3.0 mm. Width female 1.1 – 1.3 mm. Head and rostrum with brown scales; vertex with two patches of white scales; frons concave; rostrum moderately convex, surface moderately carinate, punctures similar to those of head. Elytra with small, shallow punctures, without granules; tubercle distribution as follows: I 2 with small spine-shaped tubercle in front of base of elytral declivity, I 3 with two tubercles: first tubercle subbasal, second tubercle smaller than first tubercle at middle of elytra, I 4 with tubercle at base of declivity, I 5 with tubercles similar in shape and position as tubercles on I 3, I 7 with two tiny tubercles: first tubercle between tubercles on I 3, second tubercle in front of base of declivity. Metasternum flat. Abdomen with aciculate scales, shorter on sides; ventrite 2 with a transverse line of scales; ventrite 5 with aciculate scales on anterior half, with long setae on posterior half, becoming short and narrow to apex. Tibiae subparallel, front tibiae with internal margin moderately sinuate. Male unknown. Geographic distribution. México (Chiapas). Habitat and elevation. Specimens were collected from leaf litter in cloud forest between 1900 – 2000 m. Derivation of specific name. Named from Latinized adjective meaning ‘‘ coapilla, ’’ after the type locality. Material examined. Total 3 females. Holotype female (CMNC): México: Chiapas, Coapilla, ca. 10.5 km NE Coapilla, elev. 1900 m (17 ° 9 ' 55.1 " N, 93 ° 8 ' 20.4 " W), 13. vii. 2007, coll. R. S. Anderson. Paratype: Same data as holotype, 1 female (CMNC); 5 km NE Coapilla, elev. 1990 m (17.17550 ° N, 93.13198 ° W), 25. v. 2008, LLAMA (Wa-A- 04 - 1 - 29) — 1 female (CMNC).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF866112FF15DAEE2777FD43.taxon	description	(Figures 54 – 55, 108, 133)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF866112FF15DAEE2777FD43.taxon	diagnosis	Diagnosis. Length male 2.5 – 3.0 mm, female, 2.5 – 2.8 mm. Width male 1.3 – 1.5 mm, female, 1.3 – 1.5 mm. As for T. coapillensis with the following exceptions: rostrum with punctures longer than punctures on head, carinate. Elytra with tubercles as follows: I 2 with low rounded tubercle at base of elytral declivity, I 3 with two tubercles: first tubercle subbasal, second tubercle smaller than first tubercle on middle of elytra, I 4 with tubercle at base of declivity, smaller than tubercle at I 2, I 5 with small subbasal tubercle, I 6 with tubercle on declivity, smaller than tubercle on I 2 and I 4. Metasternum moderately concave. Abdomen with ventrites 3 – 5 not in same plane as ventrites 1 – 2, moderately concave, glabrous; ventrite 2 feebly concave, as long as 3 – 4 combined; ventrite 5 convex along median line, with antero-medial impression. Tibiae subparallel, middle and hind tibiae slightly wider at base, front tibiae with inner margin sinuate. Male genitalia (Figure 108) with aedeagus in lateral view strongly curved, moderately stout; in ventral view as long as apodemes, lateral sides straight, convergent, apex blunt; apical process in lateral view weakly curved, in ventral view constricted beyond median orifice, lateral sides convergent. Geographic distribution. México (Chiapas). Habitat and elevation. Specimens were collected from leaf litter in various forest types between 1080 – 2500 m. Derivation of specific name. Patronym, named after Jorge León Cortés (México) of El Colegio de la Frontera Sur, Chiapas (ECOSUR), México	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF866112FF15DAEE2777FD43.taxon	materials_examined	Material examined. Total 45 males, 33 females. Holotype male (CMNC): México: Chiapas, Pueblo Nuevo Solistahuacan, La Yerbabuena, 2.1 km NW Pueblo Nuevo, Solistahuacan, elev. 2100 m (17 ° 11 ' 0.2 " N, 92 ° 53 ' 52.4 " W), 23. ix. 1992, coll. R. Anderson. Paratypes: México: Chiapas, Coapilla, ca. 10.5 Km NE Coapilla, elev. 1900 m (17 ° 9 ' 55.1 " N, 93 ° 8 ' 20.4 " W), 13. vii. 2007, coll. R. Anderson — 1 male (CMNC); Huixtán, San Cristobal de las Casas Airport, elev. 2350 m (16 ° 41 ' 25.1 " N, 92 ° 31 ' 48 " W), 25. vii. 2005, coll. R. Anderson — 1 male (CMNC); Ocozocoautla, Laguna Bélgica, sendero montaña, elev. 1080 m (16 ° 52 ' 40.1 " N, 93 ° 27 ' 14.4 " W), 8. vii. 2003, coll. R. Anderson — 1 male (CMNC); Pueblo Nuevo Solistahuacan, La Yerbabuena, 2.1 km NW Pueblo Nuevo, Solistahuacan, elev. 2100 m (17 ° 11 ' 0.2 " N, 92 ° 53 ' 52.4 " W), 23. ix. 1992, coll. R. Anderson — 2 males, 4 females (CMNC, CWOB); Yerbabuena Reserve, elev. 1750 m, 19. ix. 1991, R. Anderson - 2 males (CMNC); Reserva La Yerbabuena, elev. 1950 m, 22. vii. 2003, coll. R. Anderson — 2 males, 2 females (BMNH, CMNC); Sán Cristóbal de las Casas, Corazón de María, 2449 m (16 ° 41 ' 7.4 " N, 92 ° 32 ' 11.4 " W), 8. vii. 2006, coll. J. L. Cozar - 1 male (ECOS); same locality, 3. v. 2006, coll. J. L. Cozar - 1 female (ECOS); Huitepec, elev. 2466 m (16 ° 44 ' 54.2 " N, 92 ° 41 ' 6.4 " W), 5. v. 2006, coll. J. L. Cozar - 1 male (ECOS); Los Alcanfores, elev. 2450 m (16 ° 44 ' 12.5 " N, 92 ° 40 ' 46.9 " W), 5. vi. 2006, coll. J. L. Cozar, 1 male (ECOS); same locality, 25. iv. 2006, coll. M. Giron - 1 male, 1 female (ECOS); Moxviquil, elev. 2147 m (16 ° 45 ' 20.2 " N, 92 ° 37 ' 58.8 " W), 26. iv. 2006, coll. J. L. Cozar - 1 male (ECOS); same locality, 26. iv. 2006, coll. M. Girón - 1 male (ECOS); same locality, 4. vii. 2006, coll. J. L. Cozar - 1 male, 1 female (ECOS); Zoológico, elev. 2379 m (16 ° 43 ' 5.5 " N, 92 ° 42 ' 9.7 " W), 4. v. 2006, coll. J. L. Cozar — 1 female (ECOS); Huitepec, S. Cristóbal, elev. 2480 m (16 ° 45 ' 0.7 " N, 92 ° 40 ' 58.1 " W), 28. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 26) — 1 male (CMNC); Huitepec, elev. 2480 (16 ° 45 ' 6.1 " N, 92 ° 40 ' 57.7 " W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 03, Wa- A- 05 - 2 - 07) — 2 males (CMNC); 3 km E de San Cristóbal de las Casas, elev. 2400 m (16 ° 44 ' 12.5 " N, 92 ° 40 ' 46.9 " W), 15. ii. 94, coll. R. Jones — 3 males, 1 female (ECOS); Reserva Huitepec, elev. 2400 m (16 ° 44 ' 12.5 " N, 92 ° 40 ' 46.9 " W), 04. viii. 94, coll. B. Gómez-G — 3 females (ECOS); same locality, 05. vii. 1995, coll. R. W. Jones — 1 male, 1 female (ECOS, UAQM); same locality, elev. 2645 m, 1. vii. 2004, coll. J. L. Cozar — 2 males (ECOS); same locality, 2520 m, 1. vii. 2004, coll. J. L. Cozar — 1 male (ECOS); same locality, 25. vii. 1995, coll. R. Jones — 1 male, 1 female (UAQM); same locality, 03. xi. 1995, coll. B. Gómez, M. Girón — 1 female (UAQM); same locality, elev. 2498 m, 11. vi. 2004, coll. J. L. Cozar — 1 male (ECOS); same locality, elev. 2400, 11. vii. 2007, coll. R. Anderson — 2 males (CMNC); same locality, elev. 2614 m, 12. vi. 2004, coll. J. L. Cozar — 1 female (ECOS); same locality, elev. 2580 m, 12. vi. 2004, coll. J. L. Cozar — 1 female (ECOS); same locality, elev. 2508 m, 12. vi. 2004, coll. J. L. Cozar — 1 female (ECOS); same locality, elev. 2560 m, 12. vi. 2004, coll. J. L. Cozar — 1 male (ECOS); same locality, elev. 2430 m, 14. vi. 2004, coll. J. L. Cozar — 2 males (ECOS); same locality, elev. 2400 m, 17. ix. 1994, coll. B. Gómez — 1 female (ECOS); same locality, 24. i. 1995, coll. B. Gómez — 2 female (ECOS); same locality, 25. viii. 94, coll. B. Gómez — 6 males, 3 females (ECOS); same locality, 27. v. 94, coll. B. Gómez — 1 female (ECOS); same locality, 29. v. 94, coll. B. Gómez — 1 female (ECOS); same locality, elev. 2400 m, 18. ix. 1993, coll. R. Jones — 1 male (ECOS); same locality, elev. 2328 m, 17. vi. 2004, coll. J. L. Cozar — 1 male, 2 females (ECOS); same locality, elev. 2520 m, 2. vii. 2004, coll. J. L. Cozar — 2 females (ECOS); same locality, elev. 2600 m, 9. vii. 2007, coll. M. G. Branstetter — 1 male (CMNC); Tenejapa, Ojo de Agua, elev. 1800 m, 21. vii. 2003, coll. R. Anderson — 1 male, 1 female (CMNC); Km. 30.5, Carretera cuota Tuxtla — San Cristóbal de las Casas, elev. 1763 m, 7. vii. 2008, coll. R. Anderson — 2 males (CNMC, CWOB).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF856113FF14D9EA21A3FEAE.taxon	description	Recognition. The species of this group are distinguished from other Tylodinus by having in combination the body robust and black and the hind and / or middle tibiae with an external tooth (Figure 26).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF856113FF14D9EA21A3FEAE.taxon	diagnosis	Diagnosis. Body 1.9 – 2.1 x longer than wide, black, clothed with dense, narrow to very narrow scales. Head and vertex with scales overlapping, similar in color, frons moderately concave; rostrum moderately convex and moderately robust, surface with large and deep (one species) or small and shallow (two species) punctures, similar in size to punctures on head, moderately carinate, apical area puncticulate; eyes moderately convex. Pronotum in lateral view moderately convex, outline with an anterior constriction, without a posterior constriction (Figure 10), in dorsal view with sides slightly divergent from base to midlength, then convergent to apex, constricted beyond the middle (Figure 14); weakly carinate (one species) or not carinate (two species); disc hollowed, moderately deep; punctures small and shallow, dense on the sides, becoming more spaced toward median section of disc, middle line sometimes smooth and glabrous (Figure 15); clothed with light brown to yellow scales, very dense on lateral sides, flanks, and anterior area, forming a thick fringe longitudinally along middle and narrow strip that diverges from middle base to flanks, dark brown scales on middle and posterior areas of disc. Elytra black, sometimes dark red on intervals 1 – 2, basal margin sinuate, finely granulate, sometimes indistinct, stria 10 complete, interrupted near middle of elytra; clothed with dark brown to light brown scales; tubercle distribution as follows: I 2 with a tubercle in front of the declivity, I 3 with two tubercles: first tubercle subbasal and second tubercle on middle of elytral, I 4 with two tubercles: first between tubercle on I 3, second tubercle on base of declivity, I 5 with two tubercles in same position as tubercles on I 3 but smaller in size, I 6 with two tubercles similar to tubercles on I 4 but smaller in size, I 7 with two tubercles similar to tubercles on I 5, smaller in size, I 9 with a subbasal swelling, tubercle on I 2 longer (one species) or same size (two species) as tubercle on I 4. Metasternum concave, glabrous on median section, metasternal tooth absent. Male abdomen with ventrites 3 – 5 not in same plane as 1 – 2, ventrite 1 concave, median section glabrous, ventrite 2 concave, as long (one species) or longer (two species) as 3 – 4 combined, with two transverse lines of scales along median section interrupted medially (Figure 23), ventrite 5 longer than ventrite 1, with a median-anterior impression. Femora armed, hind femur almost reaching elytral apex, hind tibiae with an external tooth (Figure 26), premucro absent. Male genitalia (not examined for T. canaliculatus) with the median lobe in lateral view slender, weakly curved, as long as apodemes (Figures 109 – 110); in ventral view with lateral sides straight, convergent, apex blunt; apical process in lateral view sinuate, moderately curved downward, in ventral view constricted beyond the median orifice, lateral sides subparallel. Female genitalia with tergite eight constricted, apex smooth, slightly curved and constricted, sternite eight doliform, median membranous area length as long (one species) to 2 / 3 (one species) as long as basal plate, spermatheca shallowly (one species) to very shallowly (one species) curved, ramus and nodulus indistinct.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF856113FF14D9EA21A3FEAE.taxon	discussion	Key to species of Tylodinus canaliculatus species group	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF846113FF14DA9E2139FB80.taxon	description	(Figure 130)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF846113FF14DA9E2139FB80.taxon	diagnosis	Diagnosis. Length male, 3.9 – 4.5 mm, female, 4.4 – 5.1 mm. Width male, 1.9 – 2.5 mm, female 2.1 – 2.4 mm. In addition to Champion’s description: As for T. triumforium with the following exceptions: rostrum with punctures large and deep. Pronotum weakly carinate. Female with ventrite 5 with large transversely ridged swelling. Male with ventrite 2 longer than 3 – 4 combined, ventrite 5 flat. Hind tibia with an external tooth.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF846113FF14DA9E2139FB80.taxon	discussion	Comment. The specimen from Tacaná, Chiapas has the scales more scattered and basal margin less sinuate than the specimens described by Champion. Females have a very distinctive transversely ridged swelling on abdominal ventrite 5. Geographic distribution. Guatemala and México. Habitat and elevation. Specimens were collected from leaf litter in cloud forest at 1800 m. Derivation of the specific name. Champion (1905) did not indicate the derivation of the specific name canaliculatus, but it appears to be derived from Latin meaning channeled or grooved.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF846113FF14DA9E2139FB80.taxon	materials_examined	Material examined. Total 3 males, 3 females. Guatemala: Cerro Zunil, elev. 4000 ft. — 2 males, 3 females (BMNH, syntypes); México: Chiapas, Unión Juárez, Volcán Tacaná, lower slopes, ca. 4 km N. Unión Juárez, elev. 1800 m (15 ° 5 ' 47.29 '' N, 92 ° 5 ' 17.04 '' W), 18. ix. 1992, coll. R. S. Anderson (92 – 108) — 1 male (CMNC).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF84612CFF14DFAB27DFFDDE.taxon	description	(Figures 23, 26, 56 – 57, 109, 130)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF84612CFF14DFAB27DFFDDE.taxon	diagnosis	Diagnosis. Length male, 3.9 – 4.5 mm, female, 3.5 – 4.2 mm. Width male, 2.3 – 2.1 mm, female 1.9 – 2.1 mm. Body approximately 1.9 x longer than wide. Rostrum with small and shallow punctures. Elytra clothed with dark brown to light brown scales on disc, with yellow scales intermixed, denser on sides and in patches on intervals 1 – 3 at declivity, covering apical part of tubercle on I 3 on middle of elytra and I 4 on base of declivity, and covering tubercle on I 2 almost completely. Male abdomen with ventrite 2 longer than 3 – 4 combined. Middle and hind tibiae with external tooth. Male genitalia with apical process in ventral view small (Figure 109). Female genitalia with median membranous area of sternite eight as long as basal plate, spermatheca shallowly curved. Geographic distribution. México (Chiapas). Habitat and elevation. Specimens were collected from leaf litter in cloud forest between 1500 – 2200 m. Derivation of specific name. Name derived from the Latin word triumphus = triumph, and named after the El Triunfo Biosphere Reserve, the protected natural area where the species was collected.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFF84612CFF14DFAB27DFFDDE.taxon	materials_examined	Material examined. Total 11 males, 13 females. Holotype male (CMNC): México: Chiapas, Mpio Angel Albino Corzo, Reserva Biosfera El Triunfo, Campamento El Quetzal, elev. 2220 m (15.70819 ° N 92.93070 ° W), 21. v. 2008, coll. R. S. Anderson, mossy oak forest litter. Paratypes: México: Chiapas, Angel Albino Corzo, Reserva Biosfera El Triunfo, Campamento El Quetzal, elev. 2220 m (15.70819 ° N 92.93070 ° W), 21. v. 2008, coll. R. S. Anderson — 1 female (CMNC); La Concordia, 4 km SE Custepec, elev. 2125 m (15.70658 ° N, 92.93126 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 29) — 1 male (CMNC); La Concordia, 2 km SE Custepec, elev. 1520 m (15 ° 43 ' 13.58 '' N, 92 ° 57 ' 4.18 '' W), 17. v. 2008, coll. LLAMA (Wa-A- 02 - 1 - 39) — 2 males, 1 female (CMNC, ECOS); same locality (15.72051 ° N, 92.95164 ° W), LLAMA (Wa-A- 02 - 1 - 42) — 1 male, 1 female (CMNC, BMNH); same locality, elev. 1700 m (15.71595 ° N, 92.93842 ° W), LLAMA (Wa-A- 02 - 2 - 03) — 1 female (CMNC); same locality (15.71603 ° N 92.93808 ° W) LLAMA (Wa-A- 02 - 2 - 19) — 1 female (ECOS); same locality (15.71580 ° N 92.93811 ° W), LLAMA (Wa-A- 02 - 2 - 24) — 1 male (ECOS); same locality (15.71576 ° N 92.93812 ° W), LLAMA (Wa-A- 02 - 2 - 25) — 1 female (CMNC); same locality, elev. 2125 m (15.70708 ° N, 92.93131 ° W), LLAMA (Wa-A- 03 - 2 - 17) — 1 male (CMNC); same locality, elev. 2140 m (15.70925 ° N, 92.92929 ° W), LLAMA (Wa-A- 03 - 1 - 48) — 1 female (CWOB); 3.5 km ESE Custepec, Campamento El Quetzal, elev. 1850 m (15 ° 43 '. 1 '' N, 92 ° 55 ' 59.9 '' W), 17. vii. 2007, coll. M. G. Branstetter — 1 male, 1 female (CMNC); Reserva de la Biósfera El Triunfo, Campamento El Quetzal, elev. 1830 m (15 ° 43 ' 17 '' N, 92 ° 56 ' 20.8 '' W), 20. v. 2008, coll. R. S. Anderson — 1 female (CMNC); same locality, 22. v. 2008, coll. R. S. Anderson — 1 female (CMNC); same locality, elev. 1530 m, 18. vii. 2007, coll. R. S. Anderson — 1 male (CMNC); same locality, elev. 1800 m, 18. vii. 2007, coll. R. S. Anderson — 1 female (CMNC); same locality (15.72188 ° N, 92.93677 ° W), elev. 1900 m, 19. v. 2008, coll. R. S. Anderson — 1 male (CMNC); same locality (15.72178 ° N, 92.94544 ° W), elev. 1700 m, 17. v. 2008, coll. R. S. Anderson — 2 females (CMNC, ECOS); same locality, elev. 2177 m, 23. v. 2008, coll. J. L. Cozar — 1 male (CMNC).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFBB612CFF14D95E20D4FA79.taxon	description	(Figures 10, 58 – 59, 110, 130)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFBB612CFF14D95E20D4FA79.taxon	diagnosis	Diagnosis. Length male, 4.2 – 4.5 mm, female, 4.0 – 4.5 mm. Width male, 2.1 – 2.3 mm, female 2.0 – 2.3 mm. As for T. triumforium, with the follow exceptions: body approximately 2.1 x longer than wide. Elytra without light scales on tubercles. Male genitalia (Figure 110) with apical process in ventral view longer. Female genitalia with median membranous area of sternite eight 2 / 3 as long as basal plate, spermatheca very shallowly curved. Geographic distribution. México (Chiapas). Habitat and elevation. Specimens were collected from leaf litter in oak and cloud forest between 1300 – 1400 m. Derivation of specific name. Name derived from the Latin word sepulturaensis = grave, sepulture, and La Sepultura, the name of the protected natural area where the species was collected.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFBB612CFF14D95E20D4FA79.taxon	materials_examined	Material examined. Total 7 males, 4 female. Holotype male (CMNC): México: Chiapas, Villa Corzo, Ejido Sierra Morena, Reserva de la Biósfera La Sepultura, (16 ° 09 ' N, 93 ° 36 ' 28.5 " W), elev. 1372 m, 14. vii. 2007. J. L. Cozar, hojarasca. Paratypes: México: Chiapas, Villa Corso, Ejido Sierra Morena, R. Biosfera La Sepultura, elev. 1300 m (16 ° 9 ' 21.2 '' N, 93 ° 36 ' 2.9 '' W), 17. vii. 2007, coll. R. S. Anderson — 1 female (CMNC); same locality, elev. 1367 m (16.16121 ° N, 93.60024 ° W), 12. v. 2008, coll. R. S. Anderson — 1 male, 1 female (CMNC); same locality, 1348 m, (16 ° 09.620 ’ N 93 ° 36.343 ’ W), 12. v. 2008, coll. R. S. Anderson — 1 male (CMNC); Villa Flores, Sierra Morena, elev. 1390 m (16 ° 9 ' 22.7 '' N, 93 ° 35 ' 57.1 '' W), 24. vi. 2006, coll. J. L. Cozar — 1 male (ECOS); same locality, elev. 1360 m (16.16036 ° N, 93.60511 ° W), 12. v. 2008, LLAMA (Wa-A- 01 - 1 - 42) — 1 male (ECOS); same locality, elev. 1330 m (16.15282 ° N, 93.60067 ° W), 12. v. 2008, LLAMA (Wa-A- 01 - 2 - 39) — 1 male (CMNC); same locality, elev. 1330 m (16.15296 ° N, 93.60042 ° W), 12. v. 2008, LLAMA (Wa-A- 01 - 2 - 45) — 1 female (CMNC); same locality, elev. 1310 m (16.15453 ° N, 93.60080 ° W), 12. v. 2008, LLAMA (Wa-A- 01 - 2 - 06) — 1 female (ECOS); same locality, elev. 1360 m (16.15948 ° N, 93.60516 ° W), 12. v. 2008, LLAMA (Wa-A- 01 - 1 - 04) — 1 male (CMNC);	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFBB612DFF14DD21267FFCD3.taxon	description	Recognition. The species of this group are distinguished from other Tylodinus by having in combination the body moderately robust, black, with the tubercles in front of the base of the elytral declivity similar in size, and median section of abdominal ventrites 1 – 2 glabrous.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFBB612DFF14DD21267FFCD3.taxon	diagnosis	Diagnosis. Body approximately 2.1 x longer than wide, black. Head with punctures similar to punctures on rostrum, scales yellow to light brown; vertex with yellow scales, overlapping, uniformly distributed along vertex; frons moderately concave; rostrum moderately robust, surface moderately carinate, apical area puncticulate, eyes moderately convex. Pronotum in lateral view strongly convex, outline with anterior constriction and without posterior constriction (Figure 10); in dorsal view with sides slightly divergent from base to midlength, then convergent to apex, constricted beyond the middle (Figure 14); disc with longitudinal impression; surface with dense and deep punctures on flanks, becoming smaller and scattered towards middle, with yellow to light brown scales, very dense on lateral area and flanks, anterior area, forming a thick longitudinal fringe along middle and narrow strip that diverges from middle of base to flanks, and with dark scales on middle and posterior areas of disc; surface without granules. Elytra black, basal margin slightly sinuate (Figure 3), surface without granules, striae 10 incomplete. Elytra with tubercles as follows: I 2 with weak tubercle at base of elytral declivity, I 3 with two tubercles, first subbasal and other on middle of elytral, I 4 with two tubercles very weak, first tubercle between tubercles on I 3 and other little in front on elytral declivity, not longer than tubercle on I 2, I 5 with two tubercles similar in size and position to tubercles on I 3, I 7 with only one sub basal and weak tubercle. Metasternum concave, median section glabrous, metasternal tooth absent. Male abdomen with ventrites 1 – 2 almost on same plane as ventrites 3 – 5; ventrite 1 concave, glabrous at median section; ventrite 2 concave to feebly concave, longer than ventrites 3 – 4 combined, glabrous at median section; ventrite 5 flat, as long or longer than ventrite 1, with an anterior medial impression, female with a spine-shaped projection on ventrite 5 (one species) or not. Legs narrow, femora armed, hind femora length not reaching the elytral apex, tibiae subparallel, premucro absent. Male genitalia (Figures 111 – 112) with median lobe in lateral view moderately or strongly curved, stout or moderately stout; in ventral view as long as half-length or equal to apodeme length, lateral sides straight, subparallel, apex truncate or blunt; apical process in lateral view not curved, straight, very small (<0.17 as long as median lobe body), in ventral view not constricted beyond the median orifice, lateral sides convergent to apex. Female genitalia with tergite eight trapezoidal, apex slightly curved, apical margin smooth; sternite eight doliform, with the apex not constricted, median membranous area 1 / 3 as long as basal plate length, spermatheca shallowly curved, outer margin of corpus convex, ramus and nodulus indistinct.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFBA612EFF14DFBB271AFEA3.taxon	description	(Figures 60 – 61, 111, 135)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFBA612EFF14DFBB271AFEA3.taxon	diagnosis	Diagnosis. Length male, 3.8 – 4.0 mm, female, 3.5 – 3.7 mm. Width male, 1.7 – 1.9 mm, female 1.5 – 1.8 mm. Head with light brown scales, vertex with patch of yellow scales, rostrum with yellow scales, moderately dense, surface with deep punctures. Metasternum, concave, clothed with setigerous scales. Male abdomen with ventrite 1 concave, ventrite 2 longer than 3 – 4 combined, weakly concave, ventrite 5 as long as ventrite 1; female lacking projection on ventrite 5. Male genitalia (Figure 111) with the median lobe in lateral view strongly curved, stout; in ventral view as long as half apodeme length. Geographic distribution. México (Chiapas). Habitat and elevation. Specimens were collected from leaf litter in high elevation cloud forest between 2000 – 2750 m. Derivation of specific name. Noun in apposition: after the Tzotzil word lum, meaning earth	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFBA612EFF14DFBB271AFEA3.taxon	materials_examined	Material examined. Total 8 males, 13 females. Holotype male (CMNC): México: Chiapas, San Cristóbal de las Casas, Cerro Huitepec (pico) 5 km W San Cristóbal, elev. 2750 m (16 ° 44 ' 33 '' N, 92 ° 41 ' 38.4 '' W), 15. ix. 1991, coll. R. S. Anderson. Paratypes: México: Chiapas, Coapilla, 7.3 km NE Coapilla, elev. 2200 m (17 ° 10 ' 59.88 '' N, 93 ° 7 '. 12 '' W), 12. vii. 2007, coll. M. G. Branstetter — 1 male (CMNC); Huixtán, San Cristobal de las Casas Airport, elev. 2350 m (16 ° 41 ' 25.08 '' N, 92 ° 31 ' 48 '' W), 25. vii. 2005, coll. R. S. Anderson — 1 female (CMNC); Bazoom, elev. 2450 m (16 ° 44 ' 19.0 '' N, 92 ° 29 ' 18.3 '' W), 9. vii. 2003, coll. R. S. Anderson — 1 male (CMNC); San Cristóbal de las Casas, 21 km W San Cristóbal, elev. 2000 m (16 ° 44 ' 32.6 '' N, 92 ° 48 ' 43.6 '' W), 20. ix. 1991, coll. R. S. Anderson — 1 female (CMNC); Cerro Huitepec (pico) 5 km W San Cristóbal, elev. 2750 m (16 ° 44 ' 33 '' N, 92 ° 41 ' 38.4 '' W), 15. ix. 1991, coll. R. S. Anderson — 1 male (CMNC); Reserva Huitepec, elev. 2500 m (16 ° 44 ' 42.7 '' N, 92 ° 40 ' 52 '' W), 24. i. 1995, coll. R. Jones, B. Gomez — 1 male (ECOS); same locality, 20. vi. 1995, coll. R. Jones — 2 females (UAQM); same locality, 14. xi. 1995, coll. B. Gómez, M. Girón — 2 females (ECOS); same locality, 17. vi. 2004, coll. J. L. Cozar, M. Girón — 1 female (ECOS); same locality, 3. vii. 2006, coll. J. L. Cozar — 1 female (ECOS); same locality, 2400 m (16 ° 44.795 ’ N 92 ° 41.050 ’ W), coll. R. S. Anderson — 1 female (CMNC); same locality, 2450 m (16 ° 45 ’ 84 ” N 92 ° 10 ’ 70 ” W), 11. vii. 2003, coll. R. S. Anderson — 2 females (CMNC); same locality, 2300 m, 15. vii. 2005, coll. R. S. Anderson — 1 female (CMNC); same locality, 2480 m (16.75018 ° N, 92.68284 ° W), 29. v. 2008, LLAMA (Wa-A- 05 - 2 - 40) — 1 male (CMNC); same locality, 2480 m (16.75100 ° N, 92.68270 ° W), 29. v. 2008, LLAMA (Wa-A- 05 - 2 - 19) — 1 female (CMNC).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFB9612FFF14DA8A20F4FEDB.taxon	description	(Figures 20 – 21, 62 – 63, 112, 135)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFB9612FFF14DA8A20F4FEDB.taxon	diagnosis	Diagnosis. Length male, 3.8 – 4.0 mm, female, 3.8 – 4.0 mm. Width male, 1.6 – 1.8 mm, female 1.7 – 1.9 mm. As for T. lum with the following exceptions: pronotum in lateral view moderately depressed on basal and anterior areas. Elytra with basal margin strongly sinuate. Mesosternum with cup-shaped impression, impression surface with fine granules. Male abdomen with ventrite 2 concave, ventrite 5 as long as ventrites 1 – 2 combined, female with thornshaped projection on ventrite 5. Male genitalia (Figure 112) with median lobe in lateral view moderately curved and moderately stout, in ventral view as long as apodemes. Geographic distribution. México (Chiapas). Habitat and elevation. Specimens were collected from leaf litter in cloud forest between 1500 – 2750 m. Derivation of specific name. Name derived from two words combined from Latin spinus, spini = thorn, spine; and ventris = venter, stomach, womb; belly.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFB9612FFF14DA8A20F4FEDB.taxon	materials_examined	Material examined. Total 39 males, 10 females. Holotype male (CMNC): México: Chiapas, San Cristóbal de las Casas, Reserva Huitepec, 2400 m, 11 / VII / 2007. R. Anderson. Paratypes: México: Chiapas, Huixtán, 15 km E San Cristóbal, elev. 2500 m (16 ° 44 ' 49.92 '' N, 92 ° 29 ' 24.47 '' W), 28. v. 2008, coll. LLAMA (Wa-A- 05 - 1 - 47) — 1 male (CMNC); San Cristobal de las Casas Airport, elev. 2350 m (16 ° 41 ' 25.08 '' N, 92 ° 31 ' 48 '' W), 25. vii. 2005, coll. R. S. Anderson — 1 male (CMNC); Rayón, 8.9 km E Rayón, elev. 1500 m (17 ° 10 ' 58.1 " N, 92 ° 55 ' 47.3 " W), 19. ix. 1991, coll. R. Anderson — 1 female (CMNC); Pueblo Nuevo Solistahuacán, Reserva La Yerbabuena, elev. 1950 m (17 ° 11 '. 2 '' N, 92 ° 53 ' 52.4 '' W), 22. vii. 2003, coll. R. S. Anderson — 1 male (CMNC); Reserva la Yerbabuena, 2.1 km NW de Pueblo Nuevo, Solistahuacán, elev. 2100 m (17 ° 11 ' 12.12 '' N, 92 ° 53 ' 43.44 '' W), 23. ix. 1992, coll. R. S. Anderson — 2 males (CMNC); Cerro El Calvario, near Tapalapa, elev. 2200 m (17 ° 11 ' 11.8 " N, 93 ° 7 ' 21 " W), 23. vii. 2003, coll. R. Anderson — 1 female (CMNC); Merced Bazom, elev. 2470 m (16 ° 44 ' 39.5 " N, 92 ° 29 ' 34.1 " W), 30. v. 2008, coll. R. Anderson — 1 male (CMNC); Bazom, elev. 2450 m (16 ° 44 ' 19.0 '' N, 92 ° 29 ' 18.3 '' W), 9. vii. 2003, coll. R. S. Anderson — 1 female (CMNC); San Cristóbal de las Casas, Cerro Huitepec (pico) 5 km W San Cristóbal, elev. 2750 m (16 ° 44 ' 33 '' N, 92 ° 41 ' 38.4 '' W), 18. ix. 1991, coll. R. S. Anderson — 1 male (CMNC); Reserva Huitepec, elev. 2200 m (16 ° 45 ' 50.4 " N, 92 ° 40 ' 41.9 " W), 7. vii. 2003, coll. R. Anderson — 1 male (CMNC); Huitepec, elev. 2466 m (16 ° 44 ' 54.2 '' N, 92 ° 41 ' 6.4 '' W), 5. v. 2006, coll. J. L. Cozar — 1 male (ECOS); same locality, 15. vi. 2006, coll. J. L. Cozar — 1 male (ECOS); same locality (16.75176 ° N 92.68270 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 02) — 1 male (CMNC); same locality (16.75127 ° N 92.68270 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 13) — 2 males, 2 females (CMNC, ECOS); same locality (16.75018 ° N 92.68275 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 38) — 2 males (CMNC, ECOS); same locality (16.75018 ° N 92.68284 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 40) — 1 male (CMNC); same locality (16.75122 ° N 92.68270 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 14) — 1 male (CMNC); same locality (16.75149 ° N 92.68270 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 08) — 1 male (CMNC); same locality (16.75167 ° N 92.68270 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 04) — 2 males (BMNH, CMNC); Reserva Huitepec, elev. 2600 m (16 ° 44.578 ’ N 92 ° 41.287 ’ W), 11. vii. 2008, coll. J. Longino — 1 male (CMNC); Los Alcanfores, elev. 2450 m (16 ° 44 ' 12.47 '' N, 92 ° 40 ' 46.93 '' W), 25. iv. 2006, coll. M. Girón — 1 female (ECOS); same locality, 25. iv. 2006, coll. J. L. Cozar — 1 male (ECOS); same locality, 5. vi. 2006, coll. M. Girón — 1 male (ECOS); Parque Educativo San José, elev. 2550 m (16 ° 43 ' 13.8 '' N, 92 ° 42 ' 4.32 '' W), 4. v. 2006, coll. M. Girón — 1 male (ECOS); same locality, 9. vii. 2007, coll. R. S. Anderson — 1 male (CMNC); Reserva Huitepec, elev. 2500 m (16 ° 44 ' 42.7 '' N, 92 ° 40 ' 52 '' W), 4. viii. 1994, coll. B. Gomez — 1 male (ECOS); same locality, 20. vi. 1995, coll. R. Jones — 1 female (ECOS); same locality, 5. vii. 1995, coll. M. Girón — 1 female (UAQM); same locality, 5. vii. 1995, coll. R. Jones — 1 female (UAQM); same locality, 5. vii. 1995, coll. R. Jones — 1 male (UAQM); same locality, 15. viii. 1995, coll. M. Girón — 1 male (ECOS); same locality, 14. xi. 1995, coll. B. Gómez, M. Girón — 1 male (UAQM); same locality, 14. vi. 2004, coll. J. L. Cozar — 2 males (ECOS); same locality, 17. vi. 2004, coll. J. L. Cozar — 1 male (ECOS); same locality, 2. vii. 2004, coll. J. L. Cozar — 1 male (ECOS); same locality, 9. vi. 2006, coll. U. Caballero — 1 female (ECOS); same locality, 9. vii. 2007, coll. M. G. Branstetter — 2 males (CMNC, CWOB); same locality, 11. vii. 2007, coll. R. S. Anderson — 3 males, 1 female (CMNC, BMNH).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFB8612FFF14DA4221A3F890.taxon	description	Recognition. The species of this group are distinguished from other Tylodinus by having in combination the pronotum in dorsal view weakly to feebly convex; elytra black, with dark red coloration at the base, along interval 2 and elytral declivity from interval 1 to interval 5, tubercles similar in size, scales yellow, dense on tubercles on intervals 1 – 5 at declivity base, and intervals 1 – 3 at middle of declivity (Figures 64 – 65); metasternal tooth present (Figure 19); hind tibiae with an external tooth (Figure 26).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFB8612FFF14DA4221A3F890.taxon	diagnosis	Diagnosis. Body 1.9 – 2.3 x longer than wide. Head with punctures similar to punctures on rostrum, clothed with dark brown or light brown to yellow scales; vertex with whitish or yellow scales, or similar in coloration and tonality to head scales, uniformly distributed along vertex or in patch on middle of vertex, overlapping or dense to contiguous; frons moderately concave; rostrum moderately robust, moderately carinate, apical area puncticulate; eyes moderately convex, scales between eyes similar to scales on head or with a patch of yellow scales. Pronotum in dorsal view weakly to feebly convex (Figures 65 – 67), outline with anterior constriction and without posterior constriction (Figure 10) or with anterior and posterior constriction (Figure 11); in dorsal view lateral sides slightly divergent from base to midlength, then convergent to apex, constricted beyond middle (Figure 14), disc hollowed or somewhat hollowed, punctures dense to contiguous at base and progressively more widely spaced, smaller and shallower to apex, clothed with yellow scales, very dense on lateral sides and flanks, anterior area forming thick longitudinal fringe along middle and narrow strip that diverges from middle base to flanks, and with dark brown scales on middle and posterior areas of disc, occasionally with whitish scales intermixed. Elytra black, with dark red coloration at base, along interval 2 and elytral declivity from interval 1 to interval 5 (Figures 64 – 65); tubercle distribution as follows: I 2 with tubercle on base of declivity, I 3 with two tubercles: first tubercle subbasal, second tubercle on middle of elytra, I 4 with two tubercles: first tubercle small, positioned between tubercles on I 3, second tubercle on base of declivity, similar in size or longer than tubercle on I 2, I 5 with two tubercles in same position as tubercles on I 3, I 7 with very small subbasal tubercle, tubercles weak, more or less homogenous in size; elytra with patch of yellow scales, dense on tubercles on intervals 1 – 5 at declivity base, and intervals 1 – 3 along declivity. Metasternum concave or flat, glabrous in median section, metasternal tooth present. Male abdomen concave or feebly concave, glabrous in median section; ventrite 2 feebly concave, as long as 3 – 4 combined, glabrous in median section or with a transverse line of scales along median line; ventrite 5 flat, longer than ventrite 1, with median-anterior impression. Femora armed, hind femur length reaching elytral apex, hind tibiae with external tooth (Figure 26), premucro present or absent. Male genitalia (Figures 113 – 114) with median lobe in lateral view weakly curved, stout or moderately stout, in ventral view as long as half length of apodemes, swollen at sides beyond 0.27 from base, subparallel, apex acute; apical process in lateral view large (0.35 – 0.42 as long as median lobe) or moderately large (0.26 – 0.34 as long as median lobe), slightly curved or sinuate, abruptly or moderately curved downward, in ventral view constricted beyond median orifice or not, lateral sides convergent to apex, in one species sharply widened subapically and immediately converging to the apex (triangle shaped).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFB8612FFF14DA4221A3F890.taxon	discussion	Key to species of Tylodinus mutabilis species group	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFBE6129FF14DBAA20DEF8A9.taxon	description	(Figures 64 – 65, 113, 132)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFBE6129FF14DBAA20DEF8A9.taxon	diagnosis	Diagnosis. Length male, 2.3 – 2.8 mm, female, 2.4 – 2.7 mm. Width male, 1.1 – 1.4 mm, female 1.2 – 1.3 mm. Body approximately 2.3 x longer than wide. Head with light brown to yellow scales; vertex with patch of overlapping whitish scales on middle of vertex. Pronotum in dorsal view feebly convex, outline with anterior and posterior constriction; disc somewhat hollowed. Elytra with basal margin sinuate; base of declivity with tubercle on I 2 similar in size to tubercle on I 4. Metasternum concave. Male abdomen with ventrite 1 concave, ventrite 2 with transverse line of scales along median line. Premucro absent. Male genitalia (Figures 113) with median lobe in lateral view weakly curved, moderately stout; apical process in lateral view moderately large (0.26 – 0.34 as long as median lobe), slightly curved, abruptly curved downward; in ventral view not constricted beyond the median orifice. Female genitalia with tergite eight trapezoidal, slightly curved at the apex, sternite eight trapezoidal; length of membranous median area 1 / 3 or ½ as long as basal plate; spermatheca very shallowly curved, ramus and nodulus indistinct.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFBE6129FF14DBAA20DEF8A9.taxon	discussion	Comments. This species is variable in the curvature of the front tibiae, sometimes being slightly sinuate; also, the external tooth on the middle tibia is weak or absent. Geographic distribution. México (Chiapas). Habitat and elevation. Specimens were collected from leaf litter in various types of forest between 1100 – 1900 m. Derivation of specific name. Name derived from Latin word mutabilis = changeable; inconstant.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFBE6129FF14DBAA20DEF8A9.taxon	materials_examined	Material examined. Total 23 males, 19 females. Holotype male (CMNC): México: Chiapas, Villa Corzo, Ejido Sierra Morena, Reserva de la Biósfera La Sepultura, (16 ° 09 ' 10.6 " N, 93 ° 35 ' 25.1 " W), elev. 1600 m, 19 – 20. vii. 2005. R. Anderson, oak pine ridge forest litter 2005 - 004 A. Paratypes: México: Chiapas, same locality, elev. 1800 m, 18 / 19. vii. 2007, R. Anderson — 1 male, 1 female (( CMNC); La Concordia, Reserva El Triunfo, Campamento el Quetzal, elev. 1850 m (15 ° 43 ' 17 " N, 92 ° 56 ' 21.8 " W), 17. v. 2008, coll. R. Anderson - 1 female (CMNC); same locality, 19. vii. 2007, coll. R. Anderson — 1 female (CMNC); same locality, elev. 1860 m, 17. v. 2008, coll. R. S. Anderson — 1 male (CMNC); same locality (15.72173 N 92.94059 W), elev. 1850 m, 20. v. 2008, coll. R. S. Anderson — 1 female (CMNC); 2 km SE Custepec, elev. 1520 m (15 ° 43 ' 13.8 " N, 92 ° 57 ' 1.1 " W), 17. v. 2008, coll. LLAMA (Wa-A- 02 - 1 - 13) - 1 female (ECOS); same locality (15.72078 ° N 92.95047 ° W), 17. v. 2008, coll. LLAMA (Wa-A- 02 - 1 - 18) — 1 male (ECOS); same locality (15.72098 ° N 92.95054 ° W), 17. v. 2008, coll. LLAMA (Wa-A- 02 - 1 - 22) — 1 male (CMNC); same locality (15.72053 ° N 92.95100 ° W), 17. v. 2008, coll. LLAMA (Wa-A- 02 - 1 - 41) — 1 female (ECOS); same locality (15.72051 ° N 92.95104 ° W), 17. v. 2008, coll. LLAMA (Wa-A- 02 - 1 - 42) — 1 male (CMNC); same locality, elev. 1700 m (15.71522 ° N 92.93794 ° W), 17. v. 2008, coll. LLAMA (Wa-A- 02 - 2 - 39) — 1 female (CMNC); same locality, elev. 1700 m (15.71530 ° N 92.93820 ° W), 17. v. 2008, coll. LLAMA (Wa-A- 02 - 2 - 40) — 1 male (CMNC); Villa Flores, Reserva La Sepultura, Ejido Sierra Morena, Parcela Escolar, elev. 1350 (16 ° 9 ' 10.4 " N, 93 ° 35 ' 25.1 " W), 23. vi. 2006, coll. J. L. Cozar — 1 male (CMNC); Villa Flores, Ejido Sierra Morena, R. Biosfera La Sepultura, elev. 1400 m (16 ° 9 ' 10.4 '' N, 93 ° 35 ' 25.1 '' W), 17 - 18. vii. 2003, coll. R. S. Anderson — 1 female, 1 male (CMNC); same locality (16 ° 09.351 ’ N 93 ° 36.045 ’ W), elev. 1450 m — 1 female (CMNC); same locality (16 ° 09.856 ’ N 93 ° 36.291 ’ W), elev. 1497 m — 3 males, 1 female (CMNC); same locality (16.16121 ° N 93.60024 ° W), elev. 1137 m — 2 males, 1 female (CMNC, ECOS); same locality, elev. 1235 m, 19 - 20. vii. 2005, coll. R. S. Anderson — 1 male, 1 female (CMNC); same locality, elev. 1350 m — 1 male, 2 females (CMNC); same locality, elev. 1450 m — 2 males, 1 female (CMNC); same locality, elev. 1600 m — 1 male, 1 female (BMNH, CMNC); same locality (16.15976 ° N 93.60505 ° W), 12. v. 2008, coll. LLAMA (Wa-A- 01 - 1 - 26) — 1 male (ECOS); same locality (16.16044 ° N 93.60516 ° W), 12. v. 2008, coll. LLAMA (Wa-A- 01 - 1 - 44) — 1 male (CMNC); same locality (16.15458 ° N 93.60080 ° W), 12. v. 2008, coll. LLAMA (Wa-A- 01 - 2 - 05) — 1 male (ECOS); same locality (16.15276 ° N 93.60079 ° W), 12. v. 2008, coll. LLAMA (Wa-A- 01 - 2 - 36) — 1 male (CMNC); same locality (16.15287 ° N 93.60059 ° W), 12. v. 2008, coll. LLAMA (Wa-A- 01 - 2 - 41) — 1 female (ECOS); same locality (16.15292 ° N 93.60051 ° W), 12. v. 2008, coll. LLAMA (Wa-A- 01 - 2 - 43) — 1 female (CMNC); Sierra Morena, elev. 1550 m (16.1529 ° 0 ' N, 93.5903 ° 0 ' 0 '' W), 17 - 18. vii. 2003, coll. R. S. Anderson — 1 female (CMNC).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFBD612AFF14DBAA26E2FD61.taxon	description	(Figures 66 – 67, 114, 133)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFBD612AFF14DBAA26E2FD61.taxon	diagnosis	Diagnosis. Length male, 2.0 – 2.5 mm. Width male, 1.1 – 1.2 mm. As for T. mutabilis with follow exceptions: body approximately 1.9 x longer than wide. Head with dark brown scales; vertex with scales similar in coloration and tonality to scales on head, dense to contiguous. Pronotum in lateral outline with anterior constriction and without posterior constriction; disc hollowed. Base of elytra declivity with tubercle on I 2 smaller than tubercle on I 4. Metasternum flat. Male abdomen with ventrite 1 feebly concave, ventrite 2 glabrous at median section. Male genitalia (Figures 114) with median lobe in lateral view stout. Female unknown. Geographic distribution. México (Chiapas). Habitat and elevation. The two known specimens were collected from leaf litter in cloud forest at 1500 m. Derivation of specific name. Name derived from Latin word parvus = small, little, smallest.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFBD612AFF14DBAA26E2FD61.taxon	materials_examined	Material examined. Total 2 males. Holotype male (CMNC): México: Chiapas, Trinitaria, Lagunas de Montebello, Cinco Lagos, elev. 1500 m (16 ° 6 ’ 13.32 ’’ N, 91 ° 40 ’ 46.2 ’’ W), 21. ix. 1991, coll. R. S. Anderson. Paratype: Same data as holotype — 1 male (CMNC).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFBD612AFF14D9C821A3F86C.taxon	description	Recognition. The species of this group are distinguished from other Tylodinus by having in combination the body size 2.3 mm or less; pronotum with punctures dense to contiguous at the base and progressively more widely spaced, smaller and shallower to the apex (Figures 14, 16 – 17); female with the median section of ventrite 2 with a transverse line of scales along midline (Figure 24).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFBD612AFF14D9C821A3F86C.taxon	diagnosis	Diagnosis. Body 1.9 – 2.1 x longer than wide. Head clothed with yellow scales, punctures similar to punctures on rostrum; vertex with dense to contiguous scales, similar in coloration and tonality to scales on head, uniformly distributed along vertex; rostrum moderately robust, moderately carinate, apical area puncticulate; eyes moderately convex. Pronotum in dorsal view moderately convex, outline with anterior constriction and without posterior constriction (Figure 10); in dorsal view with sides slightly divergent from base to midlength, then convergent to apex, constricted beyond middle (Figure 14); disc hollowed or somewhat hollowed, punctures dense to contiguous at base and progressively more widely spaced, smaller and shallower to apex (Figures 14, 16 – 17), clothed with yellow scales, very dense on lateral sides and flanks, anterior area and forming thick longitudinal fringe along the middle and narrow strip that diverges from middle base to flanks, and with dark brown scales on middle and posterior areas of disc (Figure 14). Elytra black, basal margin slightly sinuate; tubercle distribution as follows: I 2 with a tubercle on the base of declivity, I 3 with two tubercles: first tubercle subbasal, second tubercle on middle of elytra, I 4 with two tubercles: first tubercle small, positioned between tubercles on I 3, second tubercle on base of elytral declivity, similar in size or longer than tubercle on I 2, I 5 with two tubercles in same position as tubercles on I 3. Metasternum moderately concave, glabrous in median section; metasternal tooth absent. Male abdomen with ventrite 1 concave, median section glabrous; ventrite 2 feebly concave, male with median section glabrous, female with transverse line of scales along middle line (Figures 24 – 25), as long as or longer than ventrites 3 – 4 combined; ventrite 5 convex along median line, as long as ventrite 1, with an anterior-median impression. Femora armed or unarmed, hind femora length not reaching elytra apex, tibiae subparallel, premucro absent. Male genitalia (Figures 115 – 116) with the median lobe in lateral view weakly curved, stout or moderately stout; in ventral view as long as or ½ length that of apodeme, lateral sides straight, subparallel or convergent, apex acute; apical process large (0.35 – 0.42 as long as median lobe), slightly curved; in ventral view not constricted beyond the median orifice, convergent.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFBD612AFF14D9C821A3F86C.taxon	discussion	Key to species of Tylodinus pusillus species group	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFB36125FF14DBAA2584FB2E.taxon	description	(Figures 68 – 69, 115, 133)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFB36125FF14DBAA2584FB2E.taxon	diagnosis	Diagnosis. Length male 1.8 - 2.3 mm, female, 1.8 - 2.2 mm. Width male 0.9 - 1.0 mm, female, 1.0 - 1.1 mm. Body 2.1 x longer than wide, cuticle shiny, scales pale yellow. Pronotum somewhat hollowed. Base of elytra declivity with tubercle on I 2 longer than tubercle on I 4. Abdomen with ventrite 2 longer than ventrites 3 - 4 combined. Legs black, femora unarmed, occasionally with moderately to distinctly obtusely angulate carina. Male genitalia (Figure 115) with median lobe in lateral view stout; in ventral view as long as apodemes, lateral sides subparallel. Female genitalia with sternite eight trapezoidal, with median membranous area along the basal plate, ramus and nodulus indistinct. Geographic distribution. México (Chiapas). Habitat and elevation. Specimens were collected from leaf litter in high elevation cloud forest between 1800 - 2200 m. Derivation of specific name. Name derived from Latin word pusillus = tiny, wee, very small.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFB36125FF14DBAA2584FB2E.taxon	materials_examined	Material examined. Total 83 males, 62 females. Holotype male (CMNC): México: 4.3 Km SE Custepec, Campamento El Quetzal, elev. 2160 m (15 ° 42 ' N, 92 ° 56 ’ W), 18. vii. 2007, coll. M. G. Branstetter (# 679). Paratypes: México: Chiapas, 4 km SE Custepec, elev. 2140 m (15.70954 ° N 92.92916 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 41) — 1 male, 1 female (CMNC); Ángel Albino Corzo, Reserva El Triunfo, Campamento, elev. 2020 m (15 ° 39 ' 25.6 '' N, 92 ° 48 ' 40.3 '' W), 16 - 20. xi. 2002, coll. R. Jones — 1 male (CMNC); La Concordia, 4 km SE Custepec, elev. 2125 m (15.70777 ° N 92.93110 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 01) — 1 male (CMNC); 4.3 Km SE Custepec, Campamento El Quetzal, elev. 2160 m (15 ° 42 ' N, 92 ° 55 ' 59.9 '' W), 17. vii. 2007, coll. M. G. Branstetter — 3 females (CMNC); same locality, 18. vii. 2007, coll. M. G. Branstetter — 3 males, 4 females (BMNH, CMNC); 4 km SE Custepec, elev. 2125 m (15.70760 ° N 92.93115 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 05) — 1 male, 1 female (CMNC); same locality (15.70764 ° N 92.93114 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 04) — 3 males (CMNC); same locality (15.70755 ° N 92.93116 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 06) — 1 male (CMNC); same locality (15.70746 ° N 92.93118 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 08) — 1 male (CMNC); 4 km SE Custepec, elev. 2125 m (15.70721 ° N 92.93127 ° W), 20. v. 208, coll. LLAMA (Wa-A- 03 - 2 - 14) — 1 male, 1 female (ECOS); same locality (15.71119 ° N 92.92840 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 01) — 3 males, 1 female (ECOS); same locality (15.71115 ° N 92.92842 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 02) — 1 male (ECOS); same locality (15.71111 ° N 92.92844 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 03) — 1 male (CMNC); same locality (15.71078 ° N 92.92859 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 11) — 2 females (CMNC); same locality (15.71032 ° N 92.92880 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 22) — 1 male (ECOS); same locality (15.71028 ° N 92.92882 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 23) — 1 male, 3 females (CMNC); same locality (15.70941 ° N 92.92922 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 44) — 1 males, 2 females (ECOS); same locality (15.71024 ° N 92.92884 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 24) — 1 female (CMNC); same locality (15.70950 ° N 92.92918 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 42) — 1 female (CMNC); same locality (15.71016 ° N 92.92887 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 26) — 1 female (CMNC); same locality (15.70937 ° N 92.92924 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 45) — 2 males, 2 females (ECOS); same locality (15.70933 ° N 92.92925 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 46) — 1 female (CMNC); same locality (15.70929 ° N 92.92927 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 47) — 2 males, 1 female (CMNC); same locality (15.70925 ° N 92.92929 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 48) — 3 males, 5 females (BMNH, CMNC, CWOB); same locality (15.70917 ° N 92.92933 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 50) — 1 male, 1 female (ECOS); same locality (15.70777 ° N 92.93110 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 01) — 1 male, 1 female (CMNC); same locality (15.70773 ° N 92.93111 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 02) — 1 female (CMNC); same locality (15.70742 ° N 92.93120 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 09) — 1 male, 1 female (CMNC); same locality (15.70729 ° N 92.93123 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 12) — 1 male, 2 females (ECOS); same locality (15.70725 ° N 92.93125 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 13) — 1 female (CMNC); same locality (15.70708 ° N 92.93131 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 17) — 1 female (CMNC); same locality (15.70662 ° N 92.93126 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 28) — 1 male, 1 female (ECOS); same locality (15.70658 ° N 92.93126 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 29) — 1 male, 2 females (ECOS); same locality (15.70644 ° N 92.93124 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 32) — 1 female (CMNC); same locality (15.70635 ° N 92.93125 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 34) — 1 male, 1 female (CMNC); same locality (15.70631 ° N 92.93126 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 35) — 1 male, 2 females (CMNC); same locality (15.70595 ° N 92.93129 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 43) — 5 males, 4 females (CMNC, ECOS); same locality (15.70591 ° N 92.93129 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 44) — 2 females (CMNC); same locality (15.70568 ° N 92.93128 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 49) — 1 female (CMNC); same locality, elev. 2140 m (15.71065 ° N 92.92865 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 14) — 1 male (CMNC); same locality (15.70987 ° N 92.92901 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 33) — 1 male (CMNC); same locality (15.70983 ° N 92.92903 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 34) — 1 male (CMNC); same locality (15.70970 ° N 92.92908 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 37) — 3 males (ECOS); same locality (15.70962 ° N 92.92912 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 39) — 2 males (ECOS); same locality (15.70946 ° N 92.92920 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 43) — 1 male (CMNC); same locality, elev. 2125 m (15.70738 ° N 92.93121 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 10) — 1 male (CMNC); same locality (15.70733 ° N 92.93122 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 11) — 1 male (CMNC); same locality (15.70682 ° N 92.93136 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 23) — 1 male (CMNC); same locality (15.70679 ° N 92.93133 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 24) — 1 male (CMNC); same locality (15.70667 ° N 92.93127 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 27) — 2 males (CMNC); same locality (15.70653 ° N 92.93125 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 30) — 1 male (CMNC); same locality (15.70618 ° N 92.93128 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 38) — 1 male (ECOS); same locality (15.70609 ° N 92.93129 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 40) — 1 male (ECOS); same locality (15.70600 ° N 92.93130 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 42) — 1 male (CMNC); same locality (15.70582 ° N 92.93128 ° W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 2 - 46) — 2 males (CMNC); Reserva de la Biósfera El Triunfo, Campamento El Quetzal, elev. 2160 m (15 ° 42 ' 18.4 '' N, 92 ° 55 ' 53.8 '' W), 22. v. 2008, coll. R. S. Anderson — 1 male (CMNC); Reserva de la Biósfera El Triunfo, Campamento El Quetzal, elev. 1800 m (15 ° 43 ' 14.5 '' N, 92 ° 56 ' 24.4 '' W), 18. vii. 2007, coll. R. S. Anderson — 1 male (CMNC); same locality, elev. 2200 m (15.70819 ° N 92.93070 ° W) — 2 males (CMNC); same locality, elev. 2160 m (15.70510 N 92.93157 W) — 1 male (CMNC); same locality, 21. v. 2008, coll. R. S. Anderson — 2 males (CMNC); same locality, elev. 2180 m (15.71115 ° N 92.92832 ° W), 21. v. 2008, coll. R. S. Anderson — 3 females (CMNC); same locality, 21. v. 2008, coll. R. S. Anderson — 2 males (CMNC); Chiapas, Motozintla, 7 km SSW Motozintla de Mendoza, elev. 2000 m (15 ° 19 ' 40.8 '' N, 92 ° 16 ' 34.7 '' W), 17. ix. 1992, coll. R. S. Anderson — 2 females (CMNC); Unión Juárez, Volcán Tacaná, lower slopes, ca 4 km N, Unión Juárez, elev. 2000 m (15 ° 5 ' 47.4 '' N, 92 ° 5 ' 17.2 '' W), 18. ix. 1992, coll. R. S. Anderson — 1 male, 1 female (CMNC); same locality, 19. ix. 1992, coll. R. S. Anderson — 10 males, 4 females (CMNC, CWOB, ECOS).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFB26125FF14DE0E25EBF8A9.taxon	description	(Figures 14, 70 – 71, 116, 131)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFB26125FF14DE0E25EBF8A9.taxon	diagnosis	Diagnosis. Length male 2.3 mm. Width male 1.1 – 1.2 mm. As for T. pusillus with following exceptions: body 1.9 x longer than wide. Pronotum in dorsal view hollowed. Base of elytra declivity with tubercle on I 2 as long as tubercle on I 4. Abdomen with ventrite 2 as long as ventrites 3 – 4 combined. Legs reddish, femora armed. Male genitalia (Figure 116) with median lobe in lateral view moderately stout; in ventral view as long as half apodeme length, lateral sides convergent. Female unknown. Geographic distribution. México (Chiapas). Habitat and elevation. The two known specimens were collected from leaf litter in oak-pine-fir forest at 2800 m. Derivation of specific name. Named from Latinized adjective meaning ‘‘ porvenir, ’’ referring to the region in which this species is found and the suffix ‘‘ - ensis ’’ denoting place or locality.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFB26125FF14DE0E25EBF8A9.taxon	materials_examined	Material examined. Total 2 males. Holotype male (CMNC): México: Chiapas, El Porvenir, El Porvenir (2 km NE), elev. 2800 m (15 ° 28 ' 13.8 '' N, 92 ° 16 ' 47.64 '' W), 27. vii. 2005, coll. R. S. Anderson. Paratype: Same data as holotype — 1 male (CMNC).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFB16126FF14DBEF2783F8E3.taxon	description	(Figures 5, 19, 22, 72 - 73, 98 a, 117, 132)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFB16126FF14DBEF2783F8E3.taxon	diagnosis	Diagnosis. Length male, 10.0 - 10.1 mm, female, 10.0 mm. Width male, 3.4 - 3.5 mm, female 3.5 mm. Body approximately 2.7 x longer than wide, black. Head with dark brown scales; vertex with patch of yellow scales on middle, scales overlapping; frons moderately concave; rostrum moderately robust, moderately carinate, apical area puncticulate; eyes convex. Pronotum in lateral view strongly convex, outline with anterior constriction and without posterior constriction (Figure 10); in dorsal view with sides slightly divergent from base to midlength, then convergent to apex, constricted beyond middle; disc a longitudinal impression; surface with dense and deep punctures on flanks, becoming smaller and scattered towards middle, with yellow to light brown scales, very dense on lateral area and flanks, anterior area forming thick longitudinal fringe along middle and narrow strip that diverges from middle base to flanks, and with dark scales on middle and posterior areas of disc (Figure 14); surface without granules. Elytra with basal margin strongly sinuate (Figure 5), surface with shallow punctures, small and sparse; intervals more convex as follows: I 3 at declivity near the apex, I 7 at middle of elytra near the tubercle in this interval, I 8 at middle of elytra and I 9 at apical 2 / 3; tubercle distribution as follows: tubercle on I 2 in front of the declivity, I 3 with two tubercles, first tubercle sub basal and second tubercle on middle of elytra, I 4 with tubercle at base of elytra, smaller than tubercle on I 2, I 5 with two tubercles in same position as tubercles on I 3 but smaller in size, I 6 with swelling on declivity, I 7 with callous on humerus and subbasal tubercle, I 9 with subbasal swelling. Metasternum moderately concave, median section glabrous, metasternal tooth present (Figure 19). Abdomen very large, longer than length from head to metasternum on anterior area, male with ventrite 1 moderately concave, glabrous in median section; ventrite 2 as long as 3 - 4 combined, convex, glabrous in median section; ventrite 5 longer than 1, convex in middle area, almost glabrous, scales dense on ventrite 5, female with two acute projections at apex (Figure 22). Legs very elongate, clothed with dark brown scales, light brown on apical 1 / 3 on dorsal area and along inside of femora, with patch of light brown to yellow scales on femoral tooth, hind femur almost reaching elytral apex, front tibia subparallel, excavated on apical ½ on internal margin, middle and hind tibiae subparallel, internal margin slightly sinuate, premucro absent. Male genitalia (Figure 98 a, 117) with median lobe in lateral view weakly curved, very slender; in ventral view longer than apodemes, lateral sides straight, convergent, apex blunt; apical process in lateral view moderately large (0.26 - 0.34 as long as median lobe), moderately curved downward and then abruptly curved upward (Figure 117); in ventral view constricted beyond median orifice, lateral sides subparallel Female genitalia with tergite eight trapezoidal, apex slightly curved, apical margin smooth; sternite eight doliform, with apex not constricted, length of median membranous area 1 / 3 as long as basal plate, spermatheca very shallowly curved, outer margin of the corpus convex, ramus and nodulus indistinct.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFB16126FF14DBEF2783F8E3.taxon	discussion	Comments. This species resembles members of the T. spiniventris species group but is distinguished by having a very elongate body, abdomen longer than the length from head to metasternum. Geographic distribution. México (Chiapas). Habitat and elevation. The three specimens of this odd species were collected on dead tree limbs at night; they were observed to run rapidly along the branches. The habitat was a high elevation cloud forest between 2000 - 2100 m. Derivation of specific name. Name derived from the Latin word elongates = prolong, protract.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFB16126FF14DBEF2783F8E3.taxon	materials_examined	Material examined. Total 2 males, 1 female. Holotype male (CMNC): México: Chiapas, Ángel Albino Corzo, Reserva El Triunfo, Polígono 1, elev. 2050 m (15 ° 39 ' 25.56 '' N, 92 ° 48 ' 32.4 '' W), 16 - 21. xi. 2002, coll. R. S. Anderson. Paratype: México: Chiapas, Ángel Albino Corzo, El Triunfo Polygono # 1, elev. 2100 m (15 ° 39.33 ' N, 92 ° 48.44 ' W), 18. xi. 2001, coll. R. Jones — 1 female (UAQM); Reserva El Triunfo, Polígono 1, elev. 2050 m (15 ° 39 ' 25.56 '' N, 92 ° 48 ' 32.4 '' W), 16 - 21. xi. 2002, coll. R. S. Anderson — 1 male (CMNC).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFB06120FF14DBAA2023FA23.taxon	description	(Figures 24 – 25, 74 - 75, 118, 131)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFB06120FF14DBAA2023FA23.taxon	diagnosis	Diagnosis. Length male, 2.2 - 2.5 mm, female, 2.2 - 2.5 mm. Width male, 1.0 - 1.1 mm, female 1.1 - 1.3 mm. Body approximately 2.1 x longer than wide, cuticle sometimes shiny. Head with yellow scales; vertex with dense to contiguous scales, uniformly distributed along vertex, similar in coloration and tonality to scales on head; frons moderately concave; rostrum moderately robust, moderately carinate, apical area puncticulate, eyes moderately convex. Pronotum in dorsal view weakly convex, outline with anterior constriction and without posterior constriction (Figure 10); in dorsal view with sides slightly divergent from base to midlength, then convergent to apex, constricted beyond middle; disc hollowed, punctures dense to contiguous at base and progressively more widely spaced, smaller, shallower to apex, clothed with yellow scales, very dense on lateral sides and flanks, anterior area forming thick longitudinal fringe along middle and narrow strip that diverges from middle base to flanks, with dark brown scales on middle and posterior areas of disc (Figure 14, 74). Elytra black with red coloration on base and along intervals 1 - 2 and intervals 1 - 5 on declivity (Figure 74 - 75), basal margin slightly sinuate (Figure 3), clothed with scattered yellow scales, dense on tubercles on intervals 1 - 5 at declivity base, and on intervals 1 - 3 along declivity, stria 10 complete; tubercle distribution as follows: I 2 with a small and round tubercle on declivity base, I 3 with two tubercles, the first tubercle subbasal, the second tubercle on the middle of elytral declivity, I 4 with two small tubercles, first tubercle between tubercles on I 3, the other tubercle on declivity base, smaller than tubercle on I 2 (but in some specimens of same size), I 5 with two small tubercles in same position as tubercles on I 3, tubercles on I 3 longer than other tubercles, surface without granules. Metasternum flat, glabrous at median section, metasternal teeth absent. Male abdomen with ventrites 3 - 5 in same plane as ventrites 1 - 2, ventrite 1 weakly concave, almost flat, glabrous at median section, ventrite 2 as long as 3 - 4, median section glabrous (Figure 23), female with transverse line of scales interrupted medially (Figure 24), ventrite 5 longer than ventrite 1, flat, with anterior impression. Femora armed, hind femur length reaching elytral apex, middle and hind tibiae subparallel, premucro absent. Male genitalia (Figure 118) with median lobe in lateral view moderately curved, stout; in ventral view as long as half length of apodemes, lateral sides subparallel, convex at sides beyond 0.27 of length from base, apex acute; apical process sinuate, moderately curved downward, small (0.17 - 0.25 as long as body of median lobe); in ventral view constricted beyond median orifice, rhomboid shape apically (Figure 98 e). Female with tergite eight trapezoidal, sternite eight doliform, apex not constricted, median membranous area 2 / 3 as long as basal plate, spermatheca shallowly curved, ramus indistinct, nodulus slender.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFB06120FF14DBAA2023FA23.taxon	discussion	Comments. Male genitalia in T. complicatus, T. rufus, T. noctis and T. dominicus are very similar; however, T. complicatus is distinguished from its allies by the small size, elytra coloration and hind tibiae without external teeth. Geographic distribution. México (Chiapas). Habitat and elevation. Specimens were collected from leaf litter in cloud forest between 1800 - 2260 m. Derivation of specific name. Name derived from Latin complicatus = complicated, confused, intricate.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFB06120FF14DBAA2023FA23.taxon	materials_examined	Material examined. Total 64 males, 39 females. Holotype male (CMNC): México: Chiapas, Pueblo Nuevo Solistahuacán, La Yerbabuena, 2.1 km NW Pueblo Nuevo, Solistahuacan, 23. ix. 1992, Coll. R. Anderson. Paratypes: México: Chiapas, Bochil, 15.1 km NW de Bochil, elev. 1930 m (17 ° 1 ' 51.6 " N, 92 ° 52 ' 5.9 " W), 24. ix. 1992, coll. R. S. Anderson — 1 male, 1 female (CMNC); 17.3 km NW de Bochil, elev. 1800 m, (17 ° 1 ' 53.8 " N, 92 ° 50 ' 49.2 " W), 24. ix. 1992, coll. R. S. Anderson — 1 male (CMNC); Pueblo Nuevo Solistahuacán, Reserva la Yerbabuena, 2.1 km NW de Pueblo Nuevo, Solistahuacán, elev. 2100 m (17 ° 11 ' 0.2 " N, 92 °, 53 ' 52.4 " W), 23. ix. 1992, coll. R. S. Anderson — 4 males, 2 females (CMNC, ECOS); Coapilla, 5 km NNW Coapilla, elev. 1915 m, (17.18355 ° N 93.15222 ° W), 24. v. 2008, coll. LLAMA (Wm-A- 04 - 2 - 01) — 1 female (CMNC); 5 km NE Coapilla, elev. 1990 m (17 ° 10 ' 14.5 " N, 93 ° 8 ' 12.5 " W), 26. v. 2008, coll. R. S. Anderson (2008 - 035) — 3 males, 2 females (BMNH, CMNC); 1.8 km NE Jct Rds. Tapalapa / Ocotepeque, elev. 1810 m (17.17314 ° N 93.14607 ° W), 26. v. 2008, coll. R. S. Anderson — 1 male (CMNC); 2.5 km NW Jct Rds. Tapalapa / Ocotepeque, elev. 1960 m (17.17602 ° N 93.13293 ° W), 26. v. 2008, coll. R. S. Anderson — 2 females (CMNC); Cerro de Tapalapa, microwave towers, elev. 2260 m (17.18786 ° N 93.12308 ° W), 26. v. 2008, coll. R. S. Anderson — 6 males, 1 female (CMNC, ECOS); 6 km NE Coapilla, elev. 1990 m (17 ° 10 ' 33.6 " N, 93 ° 7 ' 58.4 " W), 26. v. 2008, coll. M. G. Branstetter — 1 male (CMNC); 5 km NE Coapilla, elev. 1990 m, 26. v. 2008, coll. M. G. Branstetter — 1 male (CMNC); 7.3 km NE Coapilla, elev. 2200 m (17 ° 10 ' 59.9 " N, 93 ° 7 ' 0.1 " W), 12. vii. 2007, coll. M. G. Branstetter — 1 male, 1 female (CMNC); 9 km NE Coapilla, elev. 1990, 24. v. 2008, coll. LLAMA (Wm-A- 04 - 1 - 01), 1 female (CMNC); ca. 10.5 km NE Coapilla, elev. 1990 (17 ° 9 ' 55.1 " N, 93 ° 8 ' 20.4 " W), 13. vii. 2007, coll. R. Anderson — 3 males, 1 female (ECOS); Cerro El Calvario, near Tapalapa, elev. 2000 m (17 ° 10 ' 25.7 " N, 93 ° 7 ' 52.3 " W), 23. vii. 2003, coll. R. Anderson — 2 males (CMNC); Coapilla (6 - 8 km NE), elev. 2030 m (17 ° 10 ' 25 " N, 93 ° 7 ' 52 " W), 26. vii. 2005, coll. R. Anderson — 1 male, 1 female (CMNC); Carretera Coapilla- Ocotepec, 26. v. 2006, coll. J. L. Cozar — 2 males (ECOS); El Bosque, 10 km W El Bosque, elev. 1475 m (17 ° 2 ' 36.6 " N, 92 ° 45 ' 43.6 " W), 15. ix. 1992, coll. R. Anderson — 1 male (CMNC); Rayón, 8.9 km E Rayón, elev. 1500 m (17 ° 10 ' 58.1 " N, 92 ° 55 ' 47.3 " W), 19. ix. 1991, coll. R. Anderson — 9 males, 8 females (BMNH, CMNC, CWOB); Tapalapa, Cerro El Calvario, ca. 12 km NE Coapilla, elev. 2000 m (17 ° 10 ' 28.2 " N, 93 ° 7 ' 53.4 " W), 13. vii. 2007, coll. R. Anderson — 1 female (CMNC); Cerro El Calvario, ca. 14 km NE Coapilla, elev. 2150 m (17 ° 10 ' 57.4 " N, 93 ° 7 ' 5.5 " W), 13 / vii / 2007, coll. R. Anderson — 2 males, 1 female (CMNC); Cerro El Calvario, near Tapalapa, elev. 2200 m (17 ° 11 ' 11.8 " N, 93 ° 7 ' 21 " W), 23. vii. 2003, coll. R. Anderson — 1 male, 1 female (ECOS); Carretera Tapalapa-Coapilla a 5.8 km NE de Coapilla, elev. 2000 m (17 ° 10 ' 33.22 " N, 93 ° 9 ' 51.9 " W), 13. vii. 2007, coll. J. L. Cozar — 1 male, 1 female (ECOS); Carretera Tapalapa-Coapilla a 7.3 Km NE de Coapilla, elev. 2212 m (17 ° 10 ' 57.94 " N, 93 ° 7 ' 5.29 " W), 13. vii. 2007, coll. J. L. Cozar — 1 male (ECOS); same locality, elev. 2190 (17 ° 10 ' 57.48 " N, 93 ° 7 ' 5.4 " W), 26. v. 2006, coll. J. L. Cozar — 1 male (ECOS); 5 km NE Coapilla, elev. 1990 m (17.17549 ° N 93.13251 ° W), 25. v. 2008, LLAMA (Wa-A- 04 - 1 - 17) – 1 female (ECOS); same locality, (17.17550 ° N 93.13227 ° W), 25. v. 2008, LLAMA (Wa-A- 04 - 1 - 23) – 1 male, 1 female (ECOS); same locality, (17.17550 ° N 93.13217 ° W), 25. v. 2008, LLAMA (Wa-A- 04 - 1 - 25) – 1 female (ECOS); same locality, (17.17550 ° N 93.13212 ° W), 25. v. 2008, LLAMA (Wa-A- 04 - 1 - 26) – 1 female (CMNC); same locality, (17.17550 ° N 93.13203 ° W), 25. v. 2008, LLAMA (Wa-A- 04 - 1 - 28) – 1 male (CMNC); same locality, (17.17586 ° N 93.13197 ° W), 25. v. 2008, LLAMA (Wa-A- 04 - 1 - 44) – 1 male (CMNC); same locality, (17.17585 ° N 93.13202 ° W), 25. v. 2008, LLAMA (Wa-A- 04 - 1 - 46) – 2 males (CMNC); same locality, (17.17585 ° N 93.13211 ° W), 25. v. 2008, LLAMA (Wa- A- 04 - 1 - 47) – 1 female (CMNC); 5 km NNW Coapilla, elev. 1915 m (17.18300 ° N 93.15196 ° W), 25. v. 2008, LLAMA (Wa-A- 04 - 2 - 14) — 2 males, 1 female (CMNC, ECOS); same locality, (17.18342 ° N 93.15215 ° W), LLAMA (Wa-A- 04 - 2 - 04) — 1 male (ECOS); same locality, (17.18334 ° N 93.15211 ° W), LLAMA (Wa-A- 04 - 2 - 06) — 1 female (CMNC); same locality, (17.18313 ° N 93.15202 ° W), LLAMA (Wa-A- 04 - 2 - 11) — 1 male, 1 female (ECOS); same locality, (17.18305 ° N 93.15199 ° W), LLAMA (Wa-A- 04 - 2 - 13) — 1 female (CMNC); same locality, (17.18296 ° N 93.15197 ° W), LLAMA (Wa-A- 04 - 2 - 15) — 1 female (CMNC); same locality, (17.18292 ° N 93.15195 ° W), LLAMA (Wa-A- 04 - 2 - 16) — 1 male (CMNC); same locality, (17.18287 ° N 93.15194 ° W), LLAMA (Wa-A- 04 - 2 - 17) — 2 males (CMNC); same locality, (17.18292 ° N 93.15195 ° W), LLAMA (Wa-A- 04 - 2 - 16) — 1 male (CMNC); same locality, (17.18255 ° N 93.15163 ° W), LLAMA (Wa-A- 04 - 2 - 27) — 1 male (ECOS); same locality, (17.18227 ° N 93.15139 ° W), LLAMA (Wa-A- 04 - 2 - 35) — 1 male (CMNC); same locality, (17.18248 ° N 93.15129 ° W), LLAMA (Wa-A- 04 - 2 - 40) — 1 female (CMNC); same locality, (17.18277 ° N 93.15116 ° W), LLAMA (Wa-A- 04 - 2 - 47) — 1 male (ECOS); same locality, (17.18281 ° N 93.15114 ° W), LLAMA (Wa-A- 04 - 2 - 48) — 1 female (CMNC); same locality, (17.18289 ° N 93.15110 ° W), LLAMA (Wa-A- 04 - 2 - 50) — 1 male (CMNC); Estación de Microondas, elev. 2260 m (17 ° 11 ' 16.4 " N, 93 ° 7 ' 23.2 " W), 26. v. 2006, coll. J. L. Cozar – 1 male, 1 female (ECOS); Tenejapa, Ojo de Agua, elev. 1800 m, 21. vii. 2003, coll. R. Anderson — 1 female (CMNC); Yashanal, elev. 1650 m (16 ° 49 ' 49 " N, 92 ° 27 ' 9 " W), 21. vii. 2003, coll. R. S. Anderson — 2 males (CMNC).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFB76123FF14DD0A266AFC2B.taxon	description	(Figures 98 d-e, 76 - 77, 119, 134)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFB76123FF14DD0A266AFC2B.taxon	diagnosis	Diagnosis. Length male, 2.8 - 3.3 mm, female, 2.7 - 3.9 mm. Width male, 1.3 - 1.5 mm, female 1.4 - 2.0 mm. As for T. complicatus with following exceptions: body approximately 2 x longer than wide, clothed only with yellow and narrowly elongate scales. Punctures on rostrum longer than punctures on head, apical area smooth. Pronotum red, in lateral view strongly convex. Elytra with basal margin slightly sinuate (Figure 3), tubercles on I 2 and I 3 red, declivity base with tubercle on I 4 distinctly longer than tubercle on I 2; striae 10 incomplete, surface with or without granules. Male genitalia (Figure 98 d-e, 119) with median lobe moderately stout, apical process in lateral view abruptly curved downward. Female with sternite eight oblong, median membranous area as long as basal plate.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFB76123FF14DD0A266AFC2B.taxon	discussion	Comments. The punctures on the pronotum are evenly distributed, but sometimes smooth and glabrous areas are present in the middle of the disc, which is occasionally weakly carinate. Male genitalia in T. complicatus, T. rufus, T. noctis and T. dominicus are very similar; however, T. rufus is recognizable from its allies by the pronotum coloration. Geographic distribution. México (Chiapas). Habitat and elevation. Specimens were collected from leaf litter in high elevation cloud forest between 1900 - 2950 m. Derivation of specific name. Name derived from Latin rufus = red	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFB76123FF14DD0A266AFC2B.taxon	materials_examined	Material examined. Total 77 males, 53 females. Holotype male (CMNC): México: Chiapas, San Cristóbal de las Casas, Cerro Huitepec (pico) ca. 5 km W San Cristobal, elev. 2750 m (16 ° 44 ' 33 " N, 92 ° 41 ' 38.4 " W), 15. ix. 1991, coll. R. Anderson cloud forest litter, 91 - 101. Paratypes: México: Chiapas, Coapilla, 5 km NE Coapilla, elev. 1990 m (17.17602 ° N 93.13293 ° W), 26. v. 2008, coll. LLAMA (Wm-A- 04 - 1 - 01) — 1 male (CMNC); same locality (17.17550 ° N 93.13227 ° W), LLAMA (Wa-A- 04 - 1 - 23) — 1 male, 1 female (CMNC); 5 Km NNW Coapilla, elev. 1915 m (17.18285 ° N 93.15112 ° W), 24. v. 2008, coll. LLAMA (Wa-A- 04 - 2 - 49) — 1 female (CMNC); Cerro El Calvario, near Tapalapa, elev. 2200 m (17 ° 11 ' 11.8 " N, 93 ° 7 ' 21 " W), 23. vii. 2003, coll. R. Anderson — 2 males (CMNC); same locality, elev. 2000 m (17 ° 10 ' 25.7 " N, 93 ° 7 ' 52.3 " W), 23. vii. 2003, coll. R. Anderson — 2 males (CMNC); Cerro de Tapalapa, microwave towers, elev. 2260 m (17.18786 ° N 93.12308 ° W), 27. v. 2008, coll. R. S. Anderson — 1 female (CMNC); Coapilla (6 - 8 Km NE), elev. 2030 m (17 ° 10 ' 25 " N, 93 ° 7 ' 52 " W), 26. vii. 2005, coll. R. Anderson — 2 females (CMNC); Chalchihuitán, Cerros de Chalchihuitán, elev. 2050 m (16 ° 59 ' 20.8 " N, 92 ° 37 ' 13.1 " W), 24. vii. 2003, coll. R. Anderson — 1 male (CMNC); Huixtán, Bazom, Barrio los Pinos, elev. 2350 m (16.74689 ° N 92.48985 ° W), 30. v. 2008, coll. J. L. Cozar — 2 females (ECOS); Merced Bazom, elev. 2470 m (16 ° 44 ' 39.5 " N, 92 ° 29 ' 34.1 " W), 30. v. 2008, coll. R. Anderson — 1 female (CMNC); Tapalapa, Cerro de la Danta, (17 ° 11 ' 14.3 " N, 93 ° 6 ' 31.3 " W), 20. vii. 94, coll. R. Jones — 1 female (ECOS); San Juan Chamula, Tzontehuitz (Pico) ca. 10 km NE San Cristóbal, elev. 2950 m (16 ° 49 ' 10.2 " N, 92 ° 35 ' 3.8 " W), 16. ix. 1991, coll. R. Anderson — 2 males (CMNC); San Cristóbal de las Casas, 15 km E San Cristóbal, elev. 2500 m (16.74708 ° N 92.48997 ° W), 28. v. 2008, coll. LLAMA (Wa-A- 05 - 1 - 20) — 1 male, 1 female (CMNC); same locality (16.74689 ° N 92.48990 ° W), 28. v. 2008, coll. LLAMA (Wa-A- 05 - 1 - 01) — 4 males, 3 females (CMNC); same locality (16.74687 ° N 92.48999 ° W), 28. v. 2008, coll. LLAMA (Wa-A- 05 - 1 - 03) — 1 male (ECOS); same locality (16.74686 ° N 92.49004 ° W), 28. v. 2008, coll. LLAMA (Wa-A- 05 - 1 - 04) — 1 male (CMNC); same locality (16.74703 ° N 92.49006 ° W), 28. v. 2008, coll. LLAMA (Wa-A- 05 - 1 - 18) — 2 males, 1 female (ECOS); same locality (16.74701 ° N 92.49021 ° W), 28. v. 2008, coll. LLAMA (Wa-A- 05 - 1 - 28) — 1 male (CMNC); same locality (16.74698 ° N 92.49024 ° W), 28. v. 2008, coll. LLAMA (Wa-A- 05 - 1 - 29) — 1 male (CMNC); same locality (16.74690 ° N 92.49036 ° W), 28. v. 2008, coll. LLAMA (Wa-A- 05 - 1 - 32) — 1 male (ECOS); same locality (16.74695 ° N 92.49039 ° W), 28. v. 2008, coll. LLAMA (Wa-A- 05 - 1 - 33) — 1 male (ECOS); same locality (16.74707 ° N 92.490 ° W), 28. v. 2008, coll. LLAMA (Wa-A- 05 - 1 - 38) — 2 males (ECOS); Cerro Huitepec (pico) ca. 5 km W San Cristobal, elev. 2750 m (16 ° 44 ' 33 " N, 92 ° 41 ' 38.4 " W), 15. ix. 1991, coll. R. Anderson — 1 female (CMNC); Cerro Huitepec ca. 5 km W San Cristóbal, elev. 2700 m (16 ° 44 ' 33 " N, 92 ° 41 ' 38.4 " W), 14. ix. 1992, coll. R. Anderson — 1 female (CMNC); Huitepec, elev. 2328 m (16 ° 44 ' 52.4 " N, 92 ° 40 ' 23.2 " W), 17. vi. 2004, coll. J. L. Cozar — 1 female (ECOS); same locality, elev. 2400 m (16 ° 44 ' 12.5 " N, 92 ° 40 ' 46.9 " W); 18. xi. 1993, coll. R. Jones — 1 male, 1 female (ECOS); same locality, 16. ii. 1994, coll. R. Jones — 3 males (ECOS); same locality, elev. 2480 m (16.75176 ° N 92.68270 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 02) — 1 female (CMNC); same locality (16.75167 ° N 92.68270 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 04) — 1 male (ECOS); same locality (16.75145 ° N 92.68270 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 09) — 2 males, 1 female (CMNC); same locality (16.75140 ° N 92.68270 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 10) — 2 males (ECOS); same locality (16.75127 ° N 92.68270 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 13) — 2 males, 2 females (CMNC, CWOB); same locality (16.75122 ° N 92.68270 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 14) — 1 female (ECOS); same locality (16.75113 ° N 92.68270 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 16) — 1 male (CMNC); same locality (16.75109 ° N 92.68270 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 17) — 1 male, 1 female (CWOB); same locality (16.75104 ° N 92.68270 ° W), 29. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 18) — 1 male (CMNC); same locality, elev. 2500 m (16 ° 44 ' 42.7 " N, 92 ° 40 ' 52 " W), 30. ix. 1994, coll. R. Jones — 2 males — 1 female (ECOS); same locality, 29. iv. 1994, coll. R. Jones — 1 male (ECOS); same locality, 29. iv. 1994, coll. R. Jones — 1 male (ECOS); same locality, 27. v. 1994, coll. B. Gómez-G y O. L. Gómez-N — 1 male (ECOS); same locality, 27. v. 1994, coll. O. L. Gómez-N y B. Gómez-G — 1 male (ECOS); same locality, 25. viii. 1994, coll. B. Gómez-G — 1 female (ECOS); same locality, 24. i. 1995, coll. R. Jones — 1 male (ECOS); same locality, 14. vi. 1995, coll. B. Gómez-G, M. Girón-I y A. Méndoza-R — 1 male (ECOS);); same locality, 6. iii. 1994, coll. R. Jones — 1 female (ECOS); same locality, 4. viii. 1994, coll. B. Gómez-G — 1 female (ECOS); same locality, elev. 2668 m (16 ° 44 ' 33 " N, 92 ° 41 ' 19.3 " W), 9. vi. 2004, coll. J. L. Cozar — 1 male (ECOS); same locality, elev. 2800 m (16 ° 44 ' 35.2 " N, 92 ° 41 ' 28.3 " W), 07. iv. 1994, coll. R. Jones — 1 female (ECOS); same locality, elev. 2600 m (16.75163 ° N 92.68270 ° W), 21. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 05) — 3 males, 2 females (CMNC, CWOB); Huitepec, S. Cristóbal, elev. 2480 m (16.75086 ° N 92.68270 ° W), 28. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 22) — 1 female (CMNC); same locality (16.75077 ° N 92.68270 ° W), 28. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 24) — 1 female (ECOS); same locality (16.75045 ° N 92.68270 ° W), 28. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 31) — 1 male (ECOS); same locality (16.75041 ° N 92.68270 ° W), 28. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 32) — 1 female (CMNC); same locality (16.75032 ° N 92.68270 ° W), 28. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 34) — 2 males (CMNC); same locality (16.75018 ° N 92.68317 ° W), 28. v. 2008, coll. LLAMA (Wa-A- 05 - 2 - 47) — 1 female (CMNC); Reserva Huitepec, 15. viii. 1995, coll. R. Jones — 1 male (UAQM); same locality, elev. 2200 m (16 ° 45 ' 50.4 " N, 92 ° 40 ' 41.9 " W), 7. vii. 2003, coll. R. Anderson — 1 male, 1 female (CMNC); same locality, elev. 2300 m (16 ° 45 ' 50.4 " N, 92 ° 40 ' 41.9 " W), 15. vii. 2005, coll. R. Anderson — 1 male, 1 female (BMNH); same locality, 13. ix. 2001, coll. R. Anderson — 2 males (CMNC); same locality, elev. 2400 m (16 ° 44 ' 19 " N, 92 ° 29 ' 18.2 " W), 11. vii. 2007, coll. R. Anderson — 5 males, 5 females (BMNH, CMNC, ECOS); same locality, elev. 2450 m (16 ° 44 ' 19 " N, 92 ° 29 ' 18.2 " W), 9. vii. 2003, coll. R. Anderson — 3 females (CMNC); same locality, 11. vii. 2007, coll. R. Anderson — 1 female (CMNC); same locality, elev. 2500 m (16 ° 44 ' 32 " N, 92 ° 41 ' 07 " W) 9. vi. 2006, coll. U. Caballero — 1 male, 1 female (ECOS); same locality, 17. vi. 2006, coll. U. Caballero — 1 male, 4 females (ECOS); same locality, elev. 2600 m (16 ° 45 ' 0 " N, 92 ° 40 ' 59.9 " W), 9. vii. 2007, coll. M. G. Branstetter — 6 males, 2 females (CMNC, CWOB); same locality, 9. vii. 2007, coll. R. Anderson — 2 males (CMNC); same locality, elev. 2450 m (16 ° 45 ' 84 " N, 92 ° 40 ’ 70 " W), 11. vii. 2003, coll. R. S. Anderson — 2 males (CMNC); same locality, elev. 2750 m (pico), 18. ix. 1991, coll. R. S. Anderson — 1 male (CMNC).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFB4613CFF14DF7227A9FF3E.taxon	description	(Figures 78 – 79, 120, 131)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFB4613CFF14DF7227A9FF3E.taxon	diagnosis	Diagnosis. Length male, 2.9 – 3.5 mm, female, 3.1 – 3.4 mm. Width male, 1.4 – 1.6 mm, female 1.4 – 1.6 mm. As for T. complicatus with following exceptions: body approximately 1.9 x longer than wide. Head with dark brown scales; vertex with overlapping yellow scales, uniformly distributed along vertex or in patch on middle of vertex, occasionally with intermixed whitish scales, with patch or yellow scales between eyes. Pronotum in lateral view moderately convex; disc somewhat hollowed. Elytral tubercles weak, similar in size. Metasternum concave; metasternal tooth absent. Abdomen with ventrite 1 concave; ventrite 2 glabrous at median section. Hind tibia with external tooth; premucro present. Male genitalia (Figure 120) with median lobe in lateral view moderately to strongly curved, slender; apical process in lateral view large (0.35 – 0.42 as long as body of median lobe), sinuate, moderately curved downward. Female with sternite eight with membranous median area ½ as long as basal plate.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFB4613CFF14DF7227A9FF3E.taxon	discussion	Comments. Male genitalia in T. complicatus, T. rufus, T. noctis and T. dominicus are very similar; however, T. dominicus is distinguished from its allies by elytra coloration and an external tooth on the tibiae. The external tooth on the middle and hind tibia is occasionally weak. Geographic distribution. México (Chiapas). Habitat and elevation. Specimens were collected from leaf litter in mixed forest between 1250 – 1350 m. Derivation of specific name. Species name dedicated to religious order of Dominicans or dominicus, meaning = pertaining to a lord, belonging to a master.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFB4613CFF14DF7227A9FF3E.taxon	materials_examined	Material examined. Total 6 males, 5 females. Holotype male (CMNC): México: Chiapas, Villa Corzo, Reserva de la Biósfera La Sepultura, elev. 1300 m, 17. vii. 2007. R. Anderson. Paratypes: México: Chiapas, Villa Corso, Ejido Sierra Morena, R. Biosfera La Sepultura, elev. 1322 m (16 ° 9 ' 36.7 " N, 93 ° 36 ' 20.5 " W), 12. v. 2008, coll. R. S. Anderson — 3 males (CMNC); Sierra Morena, elev. 1330 m (16.15271 ° N 93.60087 ° W), 12. v. 2008, LLAMA (Wa-A- 01 - 2 - 34) — 1 male (ECOS); same locality (16.15265 ° N 93.60089 ° W), 12. v. 2008, LLAMA (Wa- A- 01 - 2 - 32) — 1 male (CMNC); Sierra Morena elev. 1250 m (16 ° 9 ' 22.7 " N, 93 ° 35 ' 57.1 " W), 18. vii. 2005, coll. J. L. Cozar — 1 female (ECOS); Sierra Morena, La Peña, elev. 1250 m, 23. vi. 2006, coll. J. L. Cozar — 1 female (ECOS); Sierra Morena, Parcela de Don Dimas, elev. 1250 m, 23. vi. 2006, coll. J. L. Cozar — 1 female (ECOS); Sierra Morena Parcela Escolar, elev. 1250 m, 23. vi .. 2006, coll. J. L. Cozar — 1 female (ECOS); Sierra Morena, Piedra Bola, elev. 1350 m, 23. vi. 2006, coll. J. L. Cozar — 1 female (ECOS).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFAB613CFF14DA7E2566F989.taxon	description	(Figures 80 – 81, 121, 135)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFAB613CFF14DA7E2566F989.taxon	diagnosis	Diagnosis. Length male, 2.5 – 2.8 mm, female, 3.0 mm. Width male, 1.2 – 1.4 mm, female 1.6 mm. Body approximately 2.0 x longer than wide, black. Head with dark brown scales; vertex with patch of yellow scales on middle of vertex, dense to contiguous; frons moderately concave; rostrum moderately robust, moderately carinate, apical area puncticulate; eyes moderately convex. Pronotum in lateral view strongly convex, outline with anterior constriction, without posterior constriction (Figure 10); in dorsal view with sides slightly divergent from base to midlength, then convergent to apex, constricted beyond middle; disc with longitudinal impression; surface with dense and deep punctures on flanks, becoming smaller, scattered towards middle, with yellow to light brown scales, very dense on lateral area and flanks, anterior area forming a thick longitudinal fringe along middle and narrow strip that diverges from middle base to flanks, and with dark scales on the middle and posterior areas of disc (Figure 14); surface without granules. Elytra with basal margin slightly sinuate (Figure 3); tubercle distribution as follows: I 2 with small, round tubercle on declivity base, I 3 with two tubercles: first subbasal, second on middle of elytra, I 4 with two small tubercles, first between tubercles on I 3, second on declivity base, longer than tubercle on I 2 (but in some specimens of equal size), I 5 with two small tubercles in same position as tubercles on I 3, tubercles on I 3 longer than other tubercles, I 6 with subbasal tubercle; surface without granules. Metasternum flat, median section glabrous; metasternal tooth absent. Male abdomen with ventrites 3 – 5 in same plane as ventrites 1 – 2, ventrite 1 weakly concave, glabrous at median section; ventrite 2 feebly concave, as long as ventrites 3 – 4 combined, with transverse line of scales along middle; ventrite 5 longer than ventrite 1, flat, with anterior impression. Femora armed, hind femur length reaching elytral apex, middle and hind tibiae subparallel, premucro absent. Male genitalia (Figure 121) similar to that of T. rufus. Female with sternite eight trapezoidal, median membranous area ½ as long as basal plate length, spermatheca very shallowly curved.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFAB613CFF14DA7E2566F989.taxon	discussion	Comments. Male genitalia in T. complicatus, T. rufus, T. noctis and T. dominicus are very similar; however, T. noctis is distinguished from its allies by body coloration Geographic distribution. México (Chiapas). Habitat and elevation. Specimens were collected from leaf litter in mixed cloud forest between 1500 – 1900 m. Derivation of specific name. Name derived from Latin word noctis = night	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFAB613CFF14DA7E2566F989.taxon	materials_examined	Material examined. Total 7 males, 2 females. Holotype male (CMNC): México: Chiapas, Coapilla, ca. 10.5 km NE Coapilla, elev. 1900 m (17 ° 09.916 ' N, 93 ° 08.337 ' W), 13. vii. 2007, coll. R. S. Anderson, mixed Liquidambar, magnolia, pine forest litter 2007 - 010. Paratypes: México: Chiapas, ca. 10.5 km NE Coapilla, elev. 1900 m (17 ° 9 ' 55.08 '' N, 93 ° 8 ' 20.4 '' W), 13. vii. 2007, coll. R. S. Anderson — 2 males, 1 female (CMNC); 5 km NE Coapilla, elev. 1990 m (17.17575 ° N 93.13284 ° W), 25. v. 2008, LLAMA (Wa-a- 04 - 1 - 07) — 1 male (CMNC); Rayon, 8.9 km E Rayón, elev. 1500 m (17 ° 10 ' 58.08 '' N, 92 ° 55 ' 47.28 '' W), 19. ix. 1991, coll. R. S. Anderson — 3 males, 1 female (CMNC, ECOS).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFAB613DFF14DD912730FA93.taxon	description	(Figures 82 – 83, 122, 134)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFAB613DFF14DD912730FA93.taxon	diagnosis	Diagnosis. Length male, 3.9 – 4.2 mm, female 4.1 – 4.2 mm. Width male, 2.1 – 2.2 mm, female 2.3 mm. Body 2.1 x longer than wide, black. Head with punctures similar to punctures on rostrum, clothed with yellow scales; vertex with overlapping scales, uniformly distributed along vertex, similar in color and tonality to head scales; frons concave; rostrum moderately robust, moderately carinate, apical area puncticulate; eyes moderately convex. Pronotum in lateral view strongly convex, outline with anterior constriction, without posterior constriction (Figure 10, 83); in dorsal view with sides slightly divergent from base to midlength, then convergent to apex, constricted beyond middle; disc with longitudinal impression; surface with dense, deep punctures on flanks, becoming smaller, scattered towards middle, with yellow to light brown scales very dense on lateral area and flanks, anterior area forming a thick longitudinal fringe along middle and narrow strip that diverges from middle base to flanks, and with dark scales on middle and posterior areas of disc (Figure 14, 82); surface without granules. Elytra black, occasionally black with dark red color along intervals 1 – 2, basal margin slightly sinuate (Figure 3), with tubercles as follows: I 2 with tubercle slightly beyond declivity base; I 3 with three tubercles, first subbasal, second on middle of elytra, similar to subbasal tubercle, third on middle of elytral declivity, similar to tubercle on I 2; I 4 with two tubercles, first very weakly developed between first two tubercles on I 3, second on base of elytral declivity, longer than tubercle on I 2; I 5 with two tubercles at same position as first two tubercles on I 3; I 6 with two weak tubercles in same position as those on I 4; I 7 with two small tubercles in same position as tubercles on I 5; elytral declivity with tubercle on I 4 much longer than tubercle on I 2. Metasternum glabrous, moderately concave, metasternal tooth absent. Male abdomen with ventrites 3 – 5 not in same plane as ventrites 1 – 2, ventrite 1 moderately concave, squamous, ventrite 2 as long as 3 – 4 combined, with a transverse line of scales along middle, ventrite 5 longer than ventrite 1, flat, with anterior impression. Femora armed, hind femur reaching elytral apex, premucro absent. Male genitalia (Figure 122) with median lobe in lateral view weakly curved, moderately stout; in ventral view longer than half the length of apodemes, swollen at 0.25 length from base of median lobe, convergent, apex acute; apical process in lateral view large (0.35 – 0.42 as long as median lobe), slightly curved; in ventral view not constricted beyond median orifice, lateral sides convergent. Female not dissected.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFAB613DFF14DD912730FA93.taxon	discussion	Comments. This species is closely related to T. pappi and T. branstetteri based on the body being clothed mainly with yellow to fulvous scales, moderately dense, dark brown scales that are less sparse, and the median lobe in ventral view distinctly wider at the apical 1 / 3 (at median orifice). Geographic distribution. México (Chiapas). Habitat and elevation. Specimens were collected from leaf litter in various wet forest types between 1300 – 1600 m. Derivation of specific name. Patronym, named after Guillermo (Willy) Kuschel (New Zealand), a wellknown worker on weevils.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFAB613DFF14DD912730FA93.taxon	materials_examined	Material examined. Total 4 males, 2 females. Holotype male (CMNC): México: Chiapas, Villa Corso, Ejido Sierra Morena, R. Biosfera La Sepultura, elev. 1487 m (16 ° 9.773 ’ N, 93 ° 3536.211 ’ W), 13. v. 2008, oak forest litter, coll. R. S. Anderson. Paratypes: México: Chiapas, Villa Flores, Ejido Sierra Morena, R. Biosfera La Sepultura, elev. 1400 m (16 ° 9 ' 10.4 '' N, 93 ° 35 ' 25.1 '' W), 19 – 20. vii. 2003, coll. R. S. Anderson — 1 male (ECOS); same locality, elev. 1600 m (16 ° 9 ' 10.6 '' N, 93 ° 35 ' 25.1 '' W) — 1 female (CMNC); same locality, elev. 1360 m (16.16028 ° N, 93.60507 ° N), 12. v. 2008, LLAMA (Wa-A- 01 - 1 - 40) — 1 female (CMNC); same locality (16.15969 ° N, 93.60516 ° N), 12. v. 2008, LLAMA (Wa-A- 01 - 1 - 23) — 1 male (ECOS); same locality, elev. 1310 m (16.15404 ° N, 93.60080 ° N), 12. v. 2008, LLAMA (Wa-A- 01 - 2 - 17) — 1 male (CMNC).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFAA613FFF14DE92278DFE86.taxon	description	(Figures 84 – 85, 123, 134)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFAA613FFF14DE92278DFE86.taxon	diagnosis	Diagnosis. Length male, 3.3 – 3.4 mm. Width male, 1.6 – 1.7 mm. As for T. kuscheli with following exceptions: body approximately 2 x longer than wide, black. Head clothed with light brown scales; vertex with yellow scales, dense to contiguous, forming patches on middle and lateral sides of vertex; frons concave. Pronotum in lateral view moderately convex; punctures dense to contiguous at base and progressively more widely spaced, smaller, shallower to apex. Elytra with tubercles as follows: I 2 with weak tubercle on declivity base, I 3 with two tubercles, first subbasal, other at elytral middle, I 4 with two very weak tubercles, first between tubercles on I 3, second slightly anterior to declivity, I 5 with two tubercles similar in size and position to those on I 3, I 7 with only one subbasal weak tubercle. Metasternum squamous. Abdomen with ventrite 5 convex along middle line, as long as ventrite 1. Hind femur length extended beyond elytral apex, front tibiae with inner margin moderately sinuate, hind tibia slightly curved. Male genitalia (Figure 123) with median lobe in ventral view as long as apodemes; apical process in lateral view very large (0.45 – 0.55 as long as median lobe), sinuate. Female unknown Geographic distribution. México (Chiapas). Habitat and elevation. Specimens were collected from leaf litter in cloud forest at 2000 m. Derivation of specific name. Patronym, named after Charles S. Papp (U. S. A.), a well-known worker on weevils.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFAA613FFF14DE92278DFE86.taxon	materials_examined	Material examined. Total 2 males. Holotype male (CMNC): México: Chiapas, Unión Juárez, Volcán Tacaná, lower slopes, ca 4 km N, Unión Juárez, elev. 2000 m (15 ° 5 ’ 47.4 ’’ N, 92 ° 5 ’ 17.2 ’’ W), 19. ix. 1992, coll. R. S. Anderson (92 - 111). Paratype: Same data as holotype, 1 male (CMNC).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFA8613FFF14DAA620C0FC2D.taxon	description	(Figures 86 – 87, 124, 131)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFA8613FFF14DAA620C0FC2D.taxon	diagnosis	Diagnosis. Length male, 3.2 mm. Width male, 1.8 mm. As for T. kuscheli with following exceptions: body 2.0 x longer than wide. Vertex scales dense to contiguous, frons moderately concave. Pronotum in lateral view moderately convex, outline with anterior and posterior constriction (Figures 11, 87). Base of elytral declivity with tubercle on I 2 longer than tubercle on I 4. Metasternum flat, glabrous. Male abdomen with ventrites 3 – 5 in same plane as ventrites 1 – 2, ventrite 1 feebly concave, glabrous at median section, ventrite 2 glabrous at median section. Male genitalia (Figure 124) with median lobe in ventral view very broadly swollen near 0.20 length from base; apical process in ventral view moderately large (0.26 – 0.34 as long as body of median lobe), sinuate. Female unknown. Geographic distribution. México (Chiapas). Habitat and elevation. The holotype was collected from leaf litter in high elevation cloud forest at 2140 m. Derivation of specific name. Patronym, named after Michael G. Branstetter (U. S. A.), a worker on Formicidae and LLAMA field companion.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFA8613FFF14DAA620C0FC2D.taxon	materials_examined	Material examined. Total 1 male. Holotype male (CMNC): México: Chiapas, La Concordia, 4 km SE Custepec, elev. 2140 m (15 ° 42 ' 38.48 '' N, 92 ° 55 ' 43.07 '' W), 20. v. 2008, coll. LLAMA (Wa-A- 03 - 1 - 01).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFA86138FF14DF0A2010FDA6.taxon	description	(Figures 88 – 89, 125, 131)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFA86138FF14DF0A2010FDA6.taxon	diagnosis	Diagnosis. Length male 2.5 – 3.3 mm, female, 2.9 – 3.2 mm. Width male 1.2 – 1.7 mm, female, 1.4 – 1.6 mm. Body 2.1 x longer than wide, black. Head clothed with yellow scales, punctures similar to those of rostrum; vertex with dense to contiguous scales, uniformly distributed along vertex, similar in color and tonality as scales on head; frons moderately concave; rostrum moderately robust, surface moderately carinate, apical area puncticulate. Pronotum in lateral view strongly convex, outline with anterior and posterior constriction (Figure 11, 89), in dorsal view with sides slightly divergent from base to midlength, then convergent to apex, constricted beyond middle; disc longitudinally hollowed, punctures dense to contiguous at base and progressively more widely spaced, smaller and shallower towards apex; disc clothed with yellow scales, very dense on lateral sides and flanks, anterior area forming thick longitudinal fringe along middle and narrow strip that diverges from middle base to flanks, and with dark brown or black scales on middle and posterior areas of disc (Figures 14, 88), surface with small punctures and without granules. Elytra black, sometimes with dark red coloration along interval 1 – 2, basal margin sinuate (Figure 4), surface granulate; tubercle distribution as follows: I 2 with small round tubercle on elytral declivity base, I 3 with two tubercles, first tubercle subbasal, second on middle of elytral declivity, I 4 with two small tubercles, first between tubercles on I 3, second on base of elytral declivity and same size as tubercle on I 2, I 5 with two small tubercles in same position as tubercles on I 3, I 6 with very small tubercle between tubercles on I 5, tubercles on I 3 longer than other tubercles. Metasternum flat, glabrous, metasternal tooth absent. Male abdomen with ventrites 3 – 5 not in same plane as 1 – 2, ventrite 1 weakly concave, with scattered scales, ventrite 2 as long as 3 – 4, feebly concave, completely squamous, ventrite 5 as long as ventrite 5, convex at middle, with an anterior impression. Femora armed, hind femur reaching elytral apex, middle and hind tibiae subparallel, premucro absent. Male genitalia (Figure 125) with median lobe in lateral view weakly curved, very slender; in ventral view as long as apodemes, lateral sides convex beyond 0.27 length from base, convergent, apex acute; apical process in lateral view moderately large (0.26 – 0.34 as long as median lobe body), sinuate, moderately curved downward; in ventral view constricted beyond median orifice, lateral sides convergent. Female with sternite eight doliform, apex not constricted, median membranous area as long as the basal plate, ramus short, slender, nodulus slender. Geographic distribution. México (Chiapas). Habitat and elevation. Specimens were collected from leaf litter in high elevation cloud forest between 2400 – 2760 m. Derivation of specific name. Name derived from the Latin word immudus = dirty, filthy, slovenly.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFA86138FF14DF0A2010FDA6.taxon	materials_examined	Material examined. Total 7 males, 12 females. Holotype male: (CMNC): México: Chiapas, San Cristóbal de las Casas Cerro Huitepec, ca. 5 km W San Cristóbal, elev. 2650 m (92 ° 41 ' 38.4 " W, 16 ° 44 ' 33 " N), 17. ix. 1991. R. Anderson, 91 – 106. Paratypes: México: Chiapas, San Cristóbal de las Casas, Cerro Huitepec (pico) 5 km W San Cristóbal, elev. 2750 m (16 ° 44 ' 33 '' N, 92 ° 41 ' 38.4 '' W), 15. ix. 1991, coll. R. S. Anderson — 1 males 1 female (CMNC); same locality, 18. ix. 1991, coll. R. S. Anderson — 1 male (CMNC); Cerro Huitepec ca. 5 km W San Cristóbal, elev. 2760 m (16 ° 44 ' 33 '' N, 92 ° 41 ' 38.4 '' W), 14. ix. 1992, coll. R. S. Anderson — 1 male, 2 females (CMNC); same locality, elev. 2650 m, 17. ix. 1991, coll. R. S. Anderson — 2 males, 8 females (BMNH, CMNC, CWOB, ECOS); same locality, 14 – 25. ix. 1992 — 1 female (CMNC); Reserva Huitepec, elev. 2400 m (16 ° 44 ' 47.76 '' N, 16 ° 45 ' 0 '' E), 11. viii. 2007, coll. R. S. Anderson — 1 male (CMNC).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFAF6138FF14D98624C7F88D.taxon	description	(Figures 90 – 91, 126, 132)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFAF6138FF14D98624C7F88D.taxon	diagnosis	Diagnosis. Length male 2.4 – 2.5 mm, female, 2.3 – 2.9 mm. Width male 1.3 – 1.4 mm, female, 1.4 – 1.6 mm. As for T. immundus with the following exceptions: body approximately 2.0 x longer than wide. Head with scale patches on middle and lateral sides on vertex; frons concave, rostrum carinate. Pronotum in lateral view feebly convex, outline with anterior constriction, without posterior constriction (Figure 10, 90); disc somewhat hollowed. Elytra sometimes with dark red coloration on intervals 1 – 4 on elytral declivity, surface granulate, basal margin slightly sinuate; tubercles absent on I 2; with dense patch of scales on intervals 1 – 3 and intervals 1 – 5 at base of elytral declivity. Metasternum moderately concave. Abdomen with ventrite 1 slightly convex, glabrous at median section, ventrite 2 glabrous at median section, ventrite 5 as long as ventrite 1, flat. Middle and hind tibiae wider basally. Male genitalia (Figure 126) with median lobe in lateral view stout, in ventral view longer than 1 / 2 length of apodemes (0.6 to 0.8), lateral sides straight, divergent from the base to the apex, apex acute; apical process in lateral view very large (0.45 – 0.55 as long as median lobe body), slightly curved; in ventral view not constricted beyond median orifice. Female with sternite eight trapezoidal, apex not constricted, membranous area as long as the basal plate, spermatheca shallowly curved, ramus and nodulus indistinct. Geographic distribution. México (Chiapas). Habitat and elevation. Specimens were collected from leaf litter in cloud forest between 1300 – 2100 m. Derivation of specific name. Patronym, named after Manuel Girón Intzin, of Ecosur (El Colegio de Frontera Sur), San Cristobal de las Casas, México.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFAF6138FF14D98624C7F88D.taxon	materials_examined	Material examined. Total 10 males, 4 females. Holotype male: (CMNC): México: Chiapas, Tenejapa, Yashanal, elev. 1800 m (16 ° 49 ' 6.6 '' N, 92 ° 26 ' 33.72 '' W), 21. vii. 2003, coll. R. S. Anderson. Paratypes: México: Chiapas, Bochil, 5.9 Km E. Bochil, elev. 1300 m (17 ° 0 ' 44.64 '' N, 90 ° 46 ' 31.8 '' W), 15. ix. 1992, coll. R. S. Anderson — 2 males, 2 females (CMNC, ECOS); Chalchihuitán, Cerros de Chalchihuitán, elev. 2050 m (16 ° 59 ' 20.76 '' N, 92 ° 37 ' 13.08 '' W), 24. vii. 2003, coll. R. S. Anderson — 1 male (CMNC); El Bosque, 10 km W El Bosque, elev. 1475 m (17 ° 2 ' 36.6 '' N, 92 ° 45 ' 43.56 '' W), 15. ix. 1992, coll. R. S. Anderson — 1 male (CMNC); Oxchuc, Cruz Wix, 13. vi. 2006, coll. M. Girón — 1 male (ECOS); Pueblo Nuevo Solistahuacán, Reserva la Yerbabuena, 2.1 km NW de Pueblo Nuevo, Solistahuacán, elev. 2100 m (17 ° 11 ' 12.12 '' N, 92 ° 53 ' 43.44 '' W), 23. ix. 1992, coll. R. S. Anderson — 2 males (CMNC); Rayon, 8.9 km E Rayón, elev. 1500 m (17 ° 10 ' 58.08 '' N, 92 ° 55 ' 47.28 '' W), 19. ix. 1991, coll. R. S. Anderson — 1 male (CMNC); Tenejapa, Ojo de Agua, elev. 1800 m (16 ° 49 ' 6.6 '' N, 92 ° 26 ' 33.72 '' W), 21. vii. 2003, coll. R. S. Anderson — 1 male (CMNC); Trinitaria, Parque Nacional Lagos de Montebello, a 50 mts Ote. de camino a 5 lagos, elev. 1594 m (16 ° 6 ' 21.88 '' N, 91 ° 40 ' 49.85 '' W), 22. vi. 2007, coll. J. L. Cozar — 2 females (ECOS).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFAE6139FF14DBAA20CAFCD5.taxon	description	(Figures 92 – 93, 127, 133)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFAE6139FF14DBAA20CAFCD5.taxon	diagnosis	Diagnosis. Length male 2.6 mm. Width male 1.3 mm. As for T. immundus with following exceptions: body 1.9. x longer than wide; densely clothed with whitish to gray scales. Frons moderately concave, rostrum moderately carinate, punctures shallow. Pronotum in lateral view moderately convex, outline with anterior constriction, without posterior constriction (Figure 10); disc somewhat hollowed, surface with small, dense punctures. Elytra strongly convex, basal margin slightly sinuate, punctures small, shallow, without granules, stria 10 incomplete, with dense patch on intervals 1 – 3 and intervals 1 – 5 at base of elytral declivity. Metasternum moderately concave. Ventrite 2 glabrous in median section. Hind femur extending beyond the apex elytral. Male genitalia (Figure 127) with median lobe in lateral view moderately stout; in ventral view with lateral sides straight, subparallel; apical process in lateral view almost as long as length of median lobe, slightly curved. Female unknown. Geographic distribution. México (Chiapas). Habitat and elevation. The holotype was collected from leaf litter in cloud forest at 1750 m. Derivation of specific name. Name from the Latin gibbosus derived from Latin word gibbus = hump, hunchback	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFAE6139FF14DBAA20CAFCD5.taxon	materials_examined	Material examined. Total 1 male. Holotype male (CMNC): México: Chiapas, Pueblo Nuevo Solistahuacán, Yerbabuena Reserve, elev. 1750 m (17 ° 11 '. 24 '' N, 92 ° 53 ' 52.44 '' W), 19. ix. 1991, coll. R. S. Anderson.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFAE6131FF14D86E2162FE86.taxon	description	(Figures 94 – 95, 128, 133)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFAE6131FF14D86E2162FE86.taxon	diagnosis	Diagnosis. Length male 4.2 – 4.6 mm, female, 4.2 mm. Width male 2.2 – 2.4 mm, female, 2.3 – 2.4 mm. Body approximately 2.2 x longer than wide, black. Head clothed with light brown scales, punctures similar to punctures on rostrum; vertex with whitish scales, dense to contiguous, uniformly distributed along vertex; frons concave, rostrum moderately robust, moderately carinate, apical area puncticulate; eyes moderately concave. Pronotum in lateral view moderately convex, outline with anterior constriction, without posterior constriction (Figures 10, 95); in dorsal view with sides slightly divergent from base to midlength, then convergent to apex, constricted beyond middle; disc with longitudinal impression, surface with dense and deep punctures on flanks, becoming smaller, scattered towards the middle, weakly carinate, disc clothed with whitish scales, dense on lateral sides and flanks, anterior area forming a thick longitudinal fringe along middle and narrow strip that diverges from middle base to flanks, and with dark brown or black scales on middle and posterior areas of disc (Figures 14, 94). Elytra strongly convex, with basal margin slightly sinuate (Figure 3); tubercles as follows: I 2 with small and round tubercle on declivity base (absent in one specimen), I 3 with two tubercles, first tubercle subbasal, second tubercle at middle of elytral declivity, I 4 with two small tubercles, first tubercle between tubercles on I 3, other tubercle at base of elytral declivity, longer than tubercle on I 2, I 5 with two small tubercles in same position as tubercles on I 3, I 6 with very small tubercle between tubercles on I 5. Metasternum moderately concave, glabrous, metasternal tooth present. Abdomen with ventrites 3 – 5 not in same plane as ventrites 1 – 2, ventrite 1, moderately concave, glabrous in median section, ventrite 2 feebly concave, as long as 3 – 4 combined, with a transverse line of scales along middle, ventrite 5 convex along median line, as long as ventrite 1, with anterior impression. Femora armed, hind femora length not reaching elytral apex, tibiae subparallel, front tibiae with inner margin sinuate, premucro absent. Male genitalia (Figure 128) with median lobe in lateral view moderately curved, moderately stout; in ventral view as long as half of apodeme length, lateral sides straight, apex acute; apical process in lateral view small (0.17 – 0.25 as long as body of median lobe), sinuate, moderately curved downward; in ventral view constricted beyond median orifice, sides convergent. Female genitalia with tergite eight trapezoidal, apex invaginate, smooth, sternite eight doliform, median membranous area 2 / 3 as long as basal plate, spermatheca shallowly curved, outer margin not convex, ramus long and slender, nodulus slender. Geographic distribution. México (Chiapas). Habitat and elevation. Specimens were collected from leaf litter in high elevation cloud forest between 1900 – 2200 m. Derivation of specific name. Patronym, named after David G. Kissinger (U. S. A), a well-known worker on	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFAE6131FF14D86E2162FE86.taxon	materials_examined	Material examined. Total 2 males, 2 females. Holotype male (CMNC): México: Chiapas, Tapalapa, ca. 14 km NE Coapilla, elev. 2150 m (17 ° 10.958 ' N, 93 ° 07.091 ' W), 13. vii. 2007, coll. R. S. Anderson, mixed cloud forest litter (steep), 2007 - 008. Paratypes: México: Chiapas, Coapilla: 5 km NE Coapilla, elev. 1990 m (17.17557 ° N 93.13268 ° W), 25. v. 2008, LLAMA (Wa-A- 04 - 1 - 13) — 1 female (CMNC); Chalchihuitán, Cerros de Chalchihuitán, elev. 2050 m (16 ° 59 ' 20.8 " N, 92 ° 37 ' 13.1 " W), 24. vii. 2003, coll. R. S. Anderson — 1 male, 1 female (CMNC).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFA66131FF14DAA627A1FA5D.taxon	description	(Figures 96 – 97, 129, 135)	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFA66131FF14DAA627A1FA5D.taxon	diagnosis	Diagnosis. Length male 4.8 – 4.9 mm. Width male 2.2 – 2.3 mm. Body 2.2 x longer than wide, black. Head clothed with dark brown scales, punctures similar to punctures on rostrum; vertex with two patches of overlapping scales, whitish or pale yellow color, light brown on lateral margin; frons moderately concave; rostrum moderately robust, weakly carinate, apical area punctuate, eyes moderately convex. Pronotum in lateral view strongly convex, outline with anterior constriction, without posterior constriction (Figures 10, 97), in dorsal view with sides slightly divergent from base to midlength, then convergent to apex, constricted beyond middle; disc somewhat hollowed medially, punctures dense to contiguous at base and progressively more widely spaced, smaller and shallower to apex (Figures 14, 96); surface granulate. Elytra with tubercles as follows: I 2 with round and large tubercle at declivity base; I 3 with two tubercles, first subbasal, second smaller, round at middle of elytra; I 4 with laminate tubercle beyond declivity; I 5 with two small tubercles in same position as tubercles on I 3; I 7 with two tubercles in same position as tubercles on I 5. Metasternum moderately concave, median section glabrous. Male abdomen with ventrite 1 moderately concave, squamous; ventrite 2 feebly concave, longer than ventrites 3 – 4 combined, squamous; ventrite 5 flat, without impressions, as long as ventrites 1 – 2. Femora armed, hind femur reaching elytral apex, tibiae with inner margin slightly sinuate, premucro present. Male genitalia (Figure 129) with median lobe in lateral view weakly curved, slender; in ventral view as long as apodemes, lateral sides moderately swollen at 0.25 length from the base, convergent, apex acute; apical process in lateral view moderately large (0.26 – 0.34 as long as body of median lobe), straight, moderately curved downward; in ventral view not constricted beyond median orifice, lateral sides convergent. Geographic distribution. México (Chiapas). Habitat and elevation. Specimens were collected from leaf litter in high elevation cloud forest between 2000 – 2200 m. Derivation of specific name. Name derived from the Latin word pinguis = fat.	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
C27E87FFFFA66131FF14DAA627A1FA5D.taxon	materials_examined	Material examined. Total 2 males. Holotype male (CMNC): México: Chiapas, Angel Albino Corzo, Reserva El Triunfo, Polígono 1, elev. 2050 m (15 ° 39 ' 25.56 '' N, 92 ° 48 ' 32.4 '' W), 16 – 21. xi. 2002, coll. R. S. Anderson. Paratype: México: Chiapas, La Concordia, 4.3 km SE Custepec, Campamento El Quetzal, elev. 2160 m (15 ° 42 ' N, 92 ° 55 ' 59.9 '' W), 18. vii. 2007, coll. M. G. Branstetter, 1 male (CMNC).	en	Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W., León-Cortés, Jorge L. (2014): A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico. Zootaxa 3788 (1): 1-63, DOI: http://dx.doi.org/10.11646/zootaxa.3788.1
