taxonID	type	description	language	source
03DECB44FFA5FFAF0CDCFC55FB3DF909.taxon	materials_examined	Type species: Viperinus orbiosus Park & Koo, sp. nov. Type locality: Kenya.	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFA5FFAF0CDCFC55FB3DF909.taxon	diagnosis	Diagnosis. The new genus Viperinus has similar external morphological characters to Protolychnis: the antenna much shorter than the forewing length; the forewing ground color dark fuscous, with a distinct orange-white or creamy-white spot near the apical end of discal cell; M 2 in the hindwing absent. However, the male genitalia can be clearly distinguished by having a uniquely specialized pair of arched processes arising from base of each valva. The presence of a polygonal signal plate in the female genitalia of the new genus is also a diagnostic character when it is compared to Protolychnis species having a crescent signal plate.	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFA5FFAF0CDCFC55FB3DF909.taxon	description	Description. Head: Antenna shorter than the forewing length. Labial palpus with 2 nd segment thickened; 3 rd segment slender, nearly same length as 2 nd segment. Thorax: Forewing is slightly broadened distally, ground color dark fuscous with a distinct orange-white spot at apical end of discal cell; blackish elliptical spots at 2 / 5 of the wing medially; apex obtuse; fringe concolorous with ground color. Hindwing broader than forewing, brownish gray; termen slightly concave medially. Abdomen: Spinous zones forming a row along posterior margin of each dorsal tergite. Male genitalia: Uncus broad basally, narrowed distally. Basal plate of gnathos produced distally; median process bent downwardly. Tegumen concave into wide V-shaped, with heavily sclerotized anterior margin. Valva with short basal part, slightly convex dorsally near base of costa; cucullus elongated, setose; a pair of specialized, arched processes arising one from each base of valva. Juxta poorly developed. Aedeagus with semi-ovate ventral lobe medially; dorsal margin heavily sclerotized.	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFA5FFAF0CDCFC55FB3DF909.taxon	etymology	Etymology. The generic name is derived from the Latin, vipera (= snake) with a Latin suffix, - inus, referring to the snake-like long processes in the male genitalia. The gender of the generic name is masculine.	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFA5FFAC0CDCF8AAFCD1F84E.taxon	description	(Figs 1 A – H)	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFA5FFAC0CDCF8AAFCD1F84E.taxon	materials_examined	Type specimens. • Holotype: male, Kenya, Rift Valley Province, Rumuruti, Ol Maisor S 1,840 m, 37 NBA 3616 3993, 25 – 26. xi. 2008, L. Aarvik, D. Agassiz & A. Kingston, gen. slide no. CIS- 7155, in NHMO. Paratypes 5 ♂, 2 ♀: • 1 ♂, Rift Valley, Rumuruti, 1,840 m, 0 ° 21 ′ 57 ″ N 36 ° 7 ′ 41 ″ E, 25 xi 2008, D. Agassiz, L. Aarvik & A. J. Kingston; • 1 ♂, Rift Valley, Rumuruti, 1,760 m, 0 ° 24 ′ 32 ″ N 36 ° 40 ′ 9 ″ E, 24 xi 2008, D. Agassiz, L. Aarvik & A. J. Kingston; • 2 ♂, Laikipia, Mpala R. C., 1,700 m, 0 ° 17 ′ 27 ″ N 36 ° 53 ′ 52 ″ E, 7. xi. 2012, Agassiz, Beavan, Heckford, & Ngugi, gen. slide no. CIS- 7438, wing slide no. CIS- 7443, COI barcode CBNU 259, CBNU 214; • 2 ♀, Laikipia, Mpala Res. Centre, 0 ° 17 ′ 27 ″ N 36 ° 53 ′ 52 ″ E, 1,700 m, 13. iv. 2010, Agassiz, Kioko, Mugambi & Ngugi, gen. slide no. CIS- 7439 (♀), wing slide no. CIS- 7477; • 1 ♂, Laikipia, Mpala R. C., 1,700 m, 0 ° 17 ′ 27 ″ N 36 ° 53 ′ 52 ″ E, 9. xi. 2012, Agassiz, Beavan, Heckford & Ngugi. All paratypes are deposited in NHMUK.	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFA5FFAC0CDCF8AAFCD1F84E.taxon	diagnosis	Diagnosis. This species is superficially very similar to Protolychnis species, but the male genitalia of the new species distinctly differ by having a pair of uniquely specialized, upwardly curved processes arising from base of each valva (Figs 1 D, F). A polygonal signal plate in the female genitalia can also be a diagnostic character when compared to Protolychnis species having a crescent signal plate (Fig. 1 G). It can also be separated from the following new species, Viperinus pyknoistus by the shape of cucullus and processes at base of the valva: 1) the cucullus is broader than that of V. pyknoistus; 2) the process has a snake head-shaped distal part, whereas in V. pyknoistus, it has a broadened distal part.	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFA5FFAC0CDCF8AAFCD1F84E.taxon	description	Description. Male and female (Figs 1 A – C). Wingspan 17.5 – 18.5 mm. Head: Vertex dark brown with orange-white erect scales laterally. Antenna orange white, about 0.7 times shorter than the forewing length. Labial palpus strongly upturned; 2 nd segment thickened, covered with dark brown scales in basal 2 / 3, then gradually turning orange white toward apex on outer surface; 3 rd segment slender with sharply pointed apex, nearly same length as 2 nd segment, orange white in 2 / 3, then gradually turning dark brown toward apex. Thorax: Thorax and tegula dark brown. Forewing slightly broadened distally; ground color dark fuscous uniformly, with a distinct orange-white spot surrounded by blackish scales at apical end of discal cell; blackish elliptical spots well-developed at basal 2 / 5 across the wing medially, instead of antemedian fascia; costa slightly arched beyond 3 / 4; apex obtuse; fringe concolorous with ground color; venation (Fig. 1 B) with R 1 arising from before middle of discal cell; distance between origins of R 1 and R 2 about 1.6 times longer than that of R 2 and R 3; R 3 shortly stalked with R 4 + 5; R 4 and R 5 stalked beyond 1 / 2, R 5 reaching near apex; M 1 remote from R 3 + 4 at base; M 2 well-developed, nearly parallel to M 1; M 3 arising from near lower corner of discal cell; CuA 1 and CuA 2 shortly stalked for about basal 1 / 7; 1 A + 2 A narrowly forked at base; cell closed. Hindwing broader than forewing, brownish gray; termen slightly concave medially; venation with M 2 absent; M 3 and CuA 1 stalked for basal 1 / 5; CuP weakly developed, arising from near base of hindwing. Abdomen: Dark brown, spinous zones forming a row along posterior margin of each segment dorsally (Fig. 1 H). Male genitalia (Figs 1 D – F). Uncus broad at base, gradually narrowed distally, slightly bent beyond 3 / 4; apex more or less pointed. Basal plate of gnathos produced distally; median process small, narrow, gently curved downwardly beyond 2 / 3. Tegumen concave into inverted wide V-shaped, with heavily sclerotized anterior margin. Valva with short basal part; slightly convex dorsally near base of costa; cucullus long, with nearly straight costal margin and arched ventral margin, strongly upturned, apex narrowly produced; a pair of large, uniquely specialized, arched processes arising one from each base of valva, snake head-shaped distal part, with dense setae. Juxta poorly developed. Aedeagus rather small, shorter than valva, narrow, slightly arched in basal 2 / 5, slightly broadened beyond, with a large, semi-ovate lobe medially on ventral margin; cornutus absent. Female genitalia (Fig. 1 G). Eighth sternite slightly emarginated medially on posterior margin, emarginated medially in wide V-shaped on anterior margin. Apophyses posteriores about 2 times longer than apophyses anteriores. Antrum long, nearly same length as ductus bursae, narrower than ductus bursae, heavily sclerotized, gradually broadened slightly toward posterior end. Ductus bursae wider than antrum, membranous; ductus seminalis arising from near posterior end. Corpus bursae elliptical, about 3 times longer than ductus bursae; signal plate polygonal, positioned in basal 1 / 3 of corpus bursae.	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFA5FFAC0CDCF8AAFCD1F84E.taxon	distribution	Distribution. Kenya.	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFA5FFAC0CDCF8AAFCD1F84E.taxon	etymology	Etymology. The species name is derived from the Latin, orbis (= circle) with a Latin suffix, - osus, referring to the arched process at the base of valva in the male genitalia.	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFA0FFAA0CDCFF6CFD3AFB88.taxon	description	(Figs 2 A – F)	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFA0FFAA0CDCFF6CFD3AFB88.taxon	materials_examined	Type specimens. • Holotype: male, Uganda, Murchison Fall N. P., Sambia Lodge, 2,500 ft, 22. vii. 2000, DJL Agassiz, gen. slide no. CIS- 7490, in NHMUK.	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFA0FFAA0CDCFF6CFD3AFB88.taxon	diagnosis	Diagnosis. This species is superficially similar to the preceding new species, V. orbiosus, but it can be easily distinguished by the male genitalia having different shape of the cucullus and the modified processes at each base of the valva: 1) the cucullus is narrowly elongated distally; 2) the process arising from the base of the valva has a broadened distal part, whereas in V. orbiosus, it is produced sharply with a pointed distal part. (Fig. 2 D).	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFA0FFAA0CDCFF6CFD3AFB88.taxon	description	Description. Male (Figs 2 A – C). Wingspan 14.0 mm. Head and Thorax: No superficial characters separable from the preceding new species, V. orbiosus, are found in the head, thorax and forewing color patterns. Only a little difference is observed in the 2 nd segment of labial palpus with dark brown in basal 1 / 3, then orange white in distal 2 / 3 on outer surface, whereas in V. orbiosus, it is covered with dark brown scales in basal 2 / 3, then gradually turning orange white on outer surface. Abdomen: Dark brown, spinose zones in a row along posterior margin of each dorsal segment, well accordant with that of V. orbiosus (Fig. 2 F). Male genitalia (Figs 2 D, E): Uncus broad basally, gradually narrowed distally, with round apex. Basal plate of gnathos produced distally, nearly pentagon; median process small, strongly bent downwardly beyond 5 / 6. Tegumen deeply concave in wide inverted V-shaped, heavily sclerotized on anterior margin. Valva with short basal part; slightly expanded dorsally near base of costa; cucullus long, narrowly elongated distally; ventral margin nearly straight, arched medially; apex narrowly produced; a pair of large, unique, arched processes arising one from each base of valva, broadened apically, giraffe ossicone-shaped, with dense setae. Juxta weakly developed. Aedeagus nearly same length as valva, narrow in basal 2 / 5, distal 3 / 5 broadened, with semi-ovate ventral lobe, sharply produced ventrally; dorsal margin heavily sclerotized; cornutus absent.	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFA0FFAA0CDCFF6CFD3AFB88.taxon	distribution	Distribution. Uganda.	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFA0FFAA0CDCFF6CFD3AFB88.taxon	etymology	Etymology. The species name is derived from the Greek, pyknos (= thick) with Greek superlative ending, - istus, referring to the thickened flagellum of the antenna.	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFA0FFA90CDCFA78FE51FD31.taxon	description	(Figs 3 A – E, I)	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFA0FFA90CDCFA78FE51FD31.taxon	materials_examined	Type specimen. • Holotype: male, Kenya, Rift Valley, Kajaido North Dist., Maasai Lodge 1,865 m, 1 ° 23 ′ 4 ″ S 36 ° 49 ′ 51 ″ E, 1 v 2015, D. J. L. Agassiz, gen. slide no. CIS- 7441, COI barcode CBNU 260, in NHMUK.	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFA0FFA90CDCFA78FE51FD31.taxon	diagnosis	Diagnosis. This new species is similar in superficial and male genital characters to P. petiliella Park, 2020, but it can be distinguished by the male genitalia: uncus slightly narrowed toward apex, whereas that of P. petiliella is more clavate distally; valva narrower at base, more narrowly elongated; and saccus much shorter, about 0.5 times shorter than that of P. petiliella (Figs 3 F – H).	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFA0FFA90CDCFA78FE51FD31.taxon	description	Description. Male (Figs 3 A – C). Wingspan 14.0 mm. Head: Vertex dark brown with orange-white erect scales laterally. Antenna thick, orange white, about 0.6 times shorter than the forewing length. Labial palpus slightly upturned; 2 nd segment thickened, covered with dark brown scales in basal 3 / 4, then gradually turning orange white toward apex on outer surface; 3 rd segment slightly curved upwardly, slender with sharply pointed apex, nearly same length as 2 nd segment, orange white entirely. Thorax: Thorax and tegula dark brown, Forewing slightly broadened distally; ground color fuscous dark brown uniformly, with a distinct orange-white spot at apical end of discal cell; fainted blackish spots at basal 2 / 5 across wing medially; costa slightly arched at basal 1 / 6, nearly straight medially, then slightly oblique from beyond 5 / 6; apex more or less rounded; termen oblique with fringe concolorous with ground color. Hindwing broader than forewing, brownish gray. Abdomen: Dark brown, spinous zones forming a band on dorsal surface along posterior margin of each segment (Fig. 3 I). Male genitalia (Figs 3 D, E): Uncus elongated, slightly narrowed toward apex, with round apex. Gnathos strongly bent from before middle, with sharply pointed apex. Valva elongated, narrow at base, broadest at about basal 1 / 3; cucullus gradually narrowed toward apex, strongly upturned from 3 / 5; costa slightly expanded near base, then gently concave; ventral margin expanded in basal 1 / 3, then gently arched; apex rounded. Juxta broad at base, with nearly symmetrical caudal processes, longer than tegumen, pointed apically. Saccus rather short. Aedeagus slender, longer than valva, narrowed in basal 1 / 6; cornuti probably consisting of multiple broom-like setae but removed.	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFA0FFA90CDCFA78FE51FD31.taxon	etymology	Etymology. The species epithet is derived from the Latin, oculi (= eye) with Latin diminutive suffix, - ella, referring to the orange-white eye-like dot on the forewing.	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFA0FFA90CDCFA78FE51FD31.taxon	distribution	Distribution. Kenya.	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFA3FFA70CDCFCE3FAC9FDCB.taxon	description	(Figs 4 A – D, H)	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFA3FFA70CDCFCE3FAC9FDCB.taxon	materials_examined	Type specimen: • Holotype, female, Tanzania, Morogoro Distr. Town, 550 – 600 m, 2 xii 1992, leg. L. Aarvik, gen. slide no. CIS- 7153, in NHMO.	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFA3FFA70CDCFCE3FAC9FDCB.taxon	diagnosis	Diagnosis. The new species can be separated from the preceding new species, P. oculiella, by the antenna less thickened and the forewing with well-developed short blackish fascia which is obliquely presented near basal 1 / 3. It is also superficially similar to P. maculata which is widely distributed in Africa, including South Africa, Malawi, Tanzania, Kenya and DR Congo. It can be distinguished from P. maculata by female genitalia characters that are as follows: 1) antrum is broadened in posterior 1 / 4 with sclerotized lateral margins, and the caudal margin is nearly straight, whereas in P. maculata, antrum is broadened with sclerotized lateral margins in posterior half, and the caudal margin is concave, with sharply produced lateral apices; 2) ductus bursae is more or less broad in posterior 3 / 4, as wide as anterior 3 / 4 of antrum, but it is nearly parallel sided in P. maculata; 3) signal plate is crescent, broad medially and tapered to each termination, whereas in P. maculata, it is narrowed, heavily sclerotized as figured in 4 E′. The female genitalia of this new species can also be distinguished from P. bastini known from DR Congo and Kenya and P. petiliella known from Uganda by the different shape of antrum: 1) antrum is as long as ductus bursae in P. morogorensis, whereas in P. bastini, it is shorter than the ductus bursae; 2) antrum is sharply broadened from anterior 3 / 4, whereas in P. bastini and P. petiliella, it is gradually broadened from the anterior base and half, respectively; 3) the caudal margin of antrum is nearly straight in P. morogorensis, but concave in P. petiliella (Figs 4 F, G).	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFA3FFA70CDCFCE3FAC9FDCB.taxon	description	Description. Female (Figs 4 A – C). Wingspan 16.0 mm. Head: Vertex dark brown with orange-white erect scales laterally. Antenna thick, orange white, about 0.7 times shorter than the forewing length. Labial palpus strongly upturned; 2 nd segment thickened, covered with dark brown scales in basal 3 / 4, scattered with orange-white scales, then gradually turning orange white toward apex on outer surface; 3 rd segment slightly curved upwardly, slender with sharply pointed apex, almost same length as 2 nd segment, orange white entirely. Thorax: Thorax and tegula dark brown, Forewing slightly broadened distally; ground color fuscous dark brown uniformly, with a distinct orange-white spot at apical end of discal cell; blackish fascia at basal 1 / 3 across wing medially; costa slightly arched at basal 1 / 5, nearly straight medially, then slightly oblique from beyond 5 / 6; apex more or less rounded; termen oblique with fringe concolorous with ground color. Hindwing broader than forewing, brownish gray. Abdomen: Dark brown, band-like spinous zones forming on dorsal surface along posterior margin of each segment (Fig. 4 H). Female genitalia (Fig. 4 D): Eighth sternite deeply emarginated medially on posterior margin. Apophyses posteriores about 1.3 times longer than apophyses anteriores. Antrum long, nearly same length as ductus bursae, as wide as posterior 3 / 4 of ductus bursae, weakly sclerotized, widened and heavily sclerotized in posterior 1 / 4; caudal margin nearly straight. Ductus bursae broadened from anterior 1 / 4, as wide as anterior 3 / 4 of antrum; ductus seminalis arising from near posterior end. Corpus bursae ovate, about 3 times longer than ductus bursae; signal plate large, crescent, as wide as posterior 2 / 3 of corpus bursae.	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFA3FFA70CDCFCE3FAC9FDCB.taxon	distribution	Distribution. Tanzania.	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFA3FFA70CDCFCE3FAC9FDCB.taxon	etymology	Etymology. The species epithet is derived from the type locality.	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFA3FFA70CDCFCE3FAC9FDCB.taxon	discussion	Remarks. The female genitalia of this new species are compared with those of its allies, including P. maculata (Figs 4 E, E′), P. bastini (Figs 4 F, F′) and P. petiliella (Figs 4 G, G′). P. chlorotoma with the orange-white median band on the forewing and P. marginata with the orange-white streak along costa differ from the species above, and P. natalensis can be distinguished from its allies by large, distinct spot near the apical end of discal cell.	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFADFFA70CDCF939FCA2F89D.taxon	description	TD: Tervuren, Belgium; London, UK [Holotype ♂ and paratypes 1 ♂, in RMCA; 1 ♀, in NHMUK]. Distribution: DR Congo, Kenya (Park & Koo, 2020).	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFADFFA70CDCF9F6FC4BF965.taxon	materials_examined	Type species: Lecithocera maculata Walsingham, 1881: 276.	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFADFFA70CDCFD75FD33FB12.taxon	materials_examined	Material examined. • 1 ♂, Tanzania, Tamga, Amani 5 ° 6 ′ 2 ″ S 38 ° 39 ′ 10 ″ E, 1,500 ft, 25. iv. 2001, DJL Agassiz, gen. slide no. CIS- 7476, in NHMUK.	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFADFFA70CDCFD75FD33FB12.taxon	diagnosis	Diagnosis. This species is superficially very similar to P. maculata, P. natalensis and P. oculiella, but it can be distinguished from the species mentioned above by the male genitalia: 1) uncus more clavate distally with rounded apex, but relatively less clavate distally in P. maculata, acute distally in P. natalensis and gradually narrowed toward rounded apex in P. oculiella; 2) two caudal processes of juxta symmetrical, much longer than tegumen, sickle-shaped with pointed apex, but relatively squat, nearly same length as teguman in P. maculata, asymmetrical, left one much longer than right one, relatively squat in P. natalensis; 3) saccus long, sharply narrowed toward basal 1 / 2, but much wider, gradually narrowed distally in P. maculata and P. natalensis and much shorter, about 0.5 times in P. oculiella; 4) aedeagus narrow and sharply arched in basal 2 / 5, whereas in the latter, gently arched.	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFADFFA70CDCFD75FD33FB12.taxon	discussion	Remarks. The species was described from Uganda and is herein reported for the first time from Tanzania. The habitus and genitalia of both sexes were recently illustrated by Park & Koo (2020).	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFADFFA70CDCFD75FD33FB12.taxon	distribution	Distribution. Uganda, Tanzania (new record).	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFADFFA40CDCF861FD40FF0F.taxon	description	TD: London, UK [Holotype ♀, in NHMUK]. Distribution: Malawi (Meyrick, 1914).	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFAEFFA40CDCFE04FDF8FD07.taxon	description	TD: London, UK [Holotype ♀ and paratype 1 ♂, in NHMUK]. Distribution: Rep. of Congo (Meyrick, 1920), DR Congo (Meyrick, 1938), Kenya (Meyrick, 1920), Malawi (Meyrick, 1920), Namibia (Janse, 1954), South Africa (Walsingham, 1881), Tanzania (Walsingham, 1891), Zimbabwe (Janse, 1954).	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFAEFFA40CDCFEDCFDB6FE27.taxon	description	TD: Beijing, China [Holotype ♂ and paratype 1 ♂, in IZCA]. Distribution: China (Wu, 1994).	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFAEFFA40CDCFCE4FA74FC3F.taxon	distribution	Distribution: Gambia (Walsingham, 1881), South Africa (Vári et al., 2002), Sudan (Rebel & Zerny, 1917).	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFAEFFA40CDCFC0CFD58FB88.taxon	description	TD: Oslo, Norway [Holotype ♀, in NHMO]. Distribution: Tanzania (present article).	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFAEFFA40CDCFA78FD7CF9FC.taxon	description	TD: London, UK [Holotype ♂, in NHMUK]. Distribution: Kenya (present article).	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFAEFFA40CDCF9CCFC14F914.taxon	description	TD: Incheon, Korea; London, UK [Holotype ♂ and paratypes 15 ♂, 6 ♀, in NIBR; 2 ♂, 2 ♀, in NHMUK]. Distribution: Uganda (Park & Koo, 2020), Tanzania (present article).	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
03DECB44FFAEFFA40CDCF8F4FD07F84C.taxon	description	TD: Cape Town, South Africa [Holotype ♀, in SAM]. Distribution: Australia (Nielsen et al., 1996).	en	Park, Kyu-Tek, Koo, Jun-Mo (2021): A new genus Viperinus Park with descriptions of two new species, and two new species of the genus Protolychnis Meyrick, 1925 (Lepidoptera, Gelechioidea, Lecithoceridae) in Kenya and Tanzania, with a checklist of the world species. Zootaxa 4985 (3): 359-370, DOI: https://doi.org/10.11646/zootaxa.4985.3.3
