taxonID	type	description	language	source
03CDBD54EC5DFFFAFC22FDF3FA93FC03.taxon	diagnosis	Diagnosis. Distinguished from any other tribes in the subfamily Phylinae by the following combination of characters: Body elongate to elongate oval, sometimes more slender (e. g., Podullahas Schuh, 1984, Sohenus Distant, 1910) variable in size (2.2 – 7.0 mm), more or less antlike (Figs 9 – 24), with simple vestiture, lacking scalelike setae; brachypterous form often present mostly in female (coleoptery or staphylinoidy sensu SCHUH & SLATER 1995); basic coloration brownish or reddish, without green tinge; eyes small; antenna uniformly long, almost linear and often thickened, as long as or longer than body; pronotum usually constricted anteriorly, with distinct, flat collar; margin of exocorium (FWS) minutely notched or serrate and dorsal surface of metafemur furnished with minute bumps (plectrum, MFP) forming a stridulatory device in six genera; legs generally long; left paramere with developed sensory lobe tightly holding phallotheca; right paramere usually reduced, tiny; endosoma elongate in most members, often sigmoid or coiled, with more or less elaborated apical part and thick-rimmed secondary gonopore; sclerotized ring ovoid to elongate oval, clearly rimmed.	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC5DFFFAFC22FDF3FA93FC03.taxon	distribution	Distribution. Nearly cosmopolitan but majority of species known from the Old World tropics, subtropics and warm temperate zones; no reliable record existing from Pacific islands or atolls; only 13 species in three genera, Cyrtopeltocoris Reuter, 1876, Lapazphylus Carvalho & Costa, 1992 and Phoradendrepulus Polhemus & Polhemus, 1985 known in New World and Acrorrhinium spicatum (Distant, 1904) in Australian Region (SCHUH 2013). However, some New World genera treated by SCHUH & MENARD (2013) as incertae sedis in the Phylini could be true hallodapine members (K. L. Menard, pers. comm.).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC5DFFFAFC22FDF3FA93FC03.taxon	discussion	Discussion. SCHUH (1984) proposed Auricillocorini composed of five Oriental genera, Auricillocoris Schuh, 1984, Cleotomiris, Cleotomiroides, Wygomiris and Zaratus Distant, 1909. Although this classification was followed by YASUNAGA (2012), the Auricillocorini is now regarded as a junior synonym of Hallodapini, involving 14 valid genera in Asia (cf. SCHUH 2013, SCHUH & MENARD 2013). Possession of the stridulatory device is currently confirmed in members of six Old World genera. In addition to Asian Alloeomimella new genus, Cleotomiris, Hallodapus, and Wygomiris (Table 1), two Ethiopian genera, Laemocoris Reuter, 1879 and Trichophthalmocapsus Poppius, 1914, were reported to have the lateral hemelytral margins (FWS) and dorsobasal surface of metafemora (MFP) modified to form a stridulatory mechanism (SCHUH 1974). We have successfully confirmed that Hallodapus centrimaculatus (approx. 2,100 Hz) and H. ravenar (800 Hz) produced sounds, using the stridulatory device (FWS + MFP) during courtship behavior (Fig. 32). Nonetheless, our recent observations suggest that the presence of the stridulatory device may be homoplasious in some species of a certain genus. This character status is evidenced by closely related species of Wygomiris; in W. indochinensis (Fig. 145) vs. W. paveli (Figs 141 – 142) and W. kaliyahae (Fig. 58) vs. W. phormictes (Figs 150, 153) – the former species lacks the device, whereas the latter possesses it (see Table 1). Little has been known about the biology of species grouped into Hallodapini as most available specimens were collected using UV light traps. The immature forms of a few Asian members were found to inhabit and / or propagate on inflorescence and leaves (e. g., Wygomiris kaliyahae, as in Figs 17 – 18), leaves (Cleotomiroides tobii, Fig. 12), tree bark or branches (Acrorrhinium spp.; see YASUNAGA et al. 2013 a, DUWAL et al. 2017) of broadleaf trees (YASUNAGA et al. 2013 a, DUWAL et al. 2017). Thus far as known, most Asian species of Alloeomimella and Hallodapus were confirmed to be epigeic and have the distinct stridulatory device (Table 1). The females of these hallodapines are predominantly brachypterous (forewing coleoptery with rudimentary hindwings, cf. Figs 104, 216 – 217); the females of Systellonotus species are exceptionally staphylinoidy (YASUNAGA 2001; WYNIGER 2006). Due to sexual dimorphism of the wings (brachypterous females that cannot take flight), the majority of individuals attracted to UV light are male as a matter of course. The brachypterous males are known in European hallodapines (e. g., Hallodapus montandoni Reuter, 1895 and Omphalonotus quadriguttatus (Kirschbaum, 1856), see WYNIGER 2006). KMENT & BAŇAŘ (2012) also reported that Hallodapus montandoni is an epigeic species inhabiting xerothermic habitats, mostly on limestone, rarely on sandy ground and possibly associated with ants, especially Myrmica spp. YASUNAGA & DUWAL (2016) discussed a spine on the genital segment (cf. Figs 30, 124, 130, 136) that is recognized in some members of Hallodapini; however, the spine is usually possessed only by males (Table 1). Within Asian hallodapines, presence of the spine in each sex (both P- and O-spines) was confirmed in Clapmarius thailandana and Peniculimiris meniscus. Although the P- and O-spines of these taxa were assumed to stabilize the mating position, the actual functions of these spines are yet to be demonstrated (YASUNAGA & DUWAL 2016). On the other hand, the stridulatory device (FWS + MFP) is currently assumed to be synapomorphy for the four Asian genera (Alloeomimella + Hallodapus + Cleotomiris + Wygomiris) plus two Afrotropical Laemocoris + Trichophthalmocapsus. The well-developed device has hitherto been found only in ground-inhabiting hallodapines. The devices in Cleotomiris miyamotoi (Figs 66 – 67) and Wygomiris paveli (Figs 55, 141 – 142) are obviously reduced and seems to lose its original function. Incidentally, the scent efferent system is noticeably small in two epigeic genera, Alloeomimella and Hallodapus, whereas it is enlarged in Cleotomiris and Wygomiris. The current evidence suggests that the ground-inhabiting hallodapines (in Alloeomimella and Hallodapus) may utilize sound for intraspecific communication rather than pheromones (from the scent gland). In addition, the parempodia of all the epigeic Alloeomimella and Hallodapus species we examined are setiform; such species frequently slip and walk clumsily on smooth surface (e. g., acrylic or glass Petri dish) but the simplified pretarsal structure possibly enables agile and comfortable movement on the chaotic, messy ground of deep meadows. Nonetheless, much broader morphological and ecological survey treating all hallodapine taxa is required to elucidate the evolution and phylogeny of Hallodapini.	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC59FFF4FCF1FC00FC48FEE3.taxon	type_taxon	Type species. Alloeomimus muiri Schuh, 1984, present designation.	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC59FFF4FCF1FC00FC48FEE3.taxon	diagnosis	Diagnosis. Distinguished from the most closely related genus, Alloeomimus Reuter, 1910, by the following combination of characters: Body generally slender, constricted at middle of hemelytron, myrmecomorphic (Figs 9 – 10); female sometimes brachypterous (Fig. 59); dorsum dark gray, generally matte; head bulbous, rounded, with eye removed from pronotal collar (Figs 9, 62); pronotum trapezoidal, weakly constricted anteriorly (Fig. 62); scutellum weakly swollen, without any projection; metathoracic scent efferent system reduced, small (Figs 59 – 60); hemelytron with a large, white triangular macula on middle corium across embolium and a wedge-shaped macula at apex of corium across embolium along anterior margin of cuneus (Fig. 9); forewing margin and metafemur equipped with a distinct stridulatory device (Figs 54, 63 – 64); endosoma slender, C-shaped, similar in overall appearance to that of Hallodapus (cf. SCHUH 1984, YASUNAGA et al. 2013 a); bursa copulatrix and ovipositors rather toughened (Fig. 210 – 211), with sclerotized rings small, ovoid (Fig. 210). Description. Male. Macropterous; body small, slender, myrmecomorphic (Figs 9 – 10); dorsum dark gray, matte, with uniformly distributed, short, reclining setae and sparsely distributed, stiff, erect setae (Fig. 62); head bulbous, rounded, less prognathous (Fig. 56); eye removed from pronotal collar but diameter of neck shorter than length of an eye in dorsal view (Figs 9, 62). Antenna generally slender, slightly longer than body; segment I short, about as long as height of an eye; segment II longer than III; segment IV about 3 times as long as I. Labium long, exceeding apex of metacoxa, reaching abdominal sternum IV. Pronotum matte, trapeziform, weakly constricted anteriorly (Fig. 62); metathoracic scent efferent system reduced, narrower than each coxa (Fig. 56); hemelytron with a large, white triangular macula on middle corium across embolium and a wedge-shaped macula at apex of corium across embolium along anterior margin of cuneus (Fig. 9); embolium with distinct FWS (Fig. 60). Legs long; metafemur equipped with distinct MFP; total length of metafemur + metatibia longer than body; parempodia setiform (Fig. 58). Male genitalia (Fig. 61): Pygophore smooth, narrow, with the apically tapered phallotheca; left paramere with stout sensory lobe; endosoma slender, C-shaped (cf. YASUNAGA et al. 2013 a). Macropterous female. Almost similar to male (Fig. 10). Stridulatory device as in male. Female genitalia (Figs 210 − 211): Bursa copulatrix relatively narrow, widely spinulate, with small, ovoid sclerotized rings (Fig. 210); ovipositors (gonapophyses) rather short; gonapophysis II sword-like, not expanded subapically (Fig. 211). Brachypterous female. As in Fig. 59. Apex of forewing reaching anterior margin of abdominal tergum VI. Stridulatory device present (Figs 63 − 64).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC59FFF4FCF1FC00FC48FEE3.taxon	etymology	Etymology. From the hallodapine generic name Alloeomimus combined with a diminutive suffix (- ella); gender feminine.	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC59FFF4FCF1FC00FC48FEE3.taxon	distribution	Distribution. Oriental Region: Indonesia (Java, Lombok), Laos and Thailand (SCHUH 1984, YASUNAGA 2013 a); new record from Lombok Island, or Wallacea, based on two female specimens collected at Dasambaru, Lombok on 5 Mar 2005 by Ishikawa (deposited in TUAK).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC59FFF4FCF1FC00FC48FEE3.taxon	discussion	Discussion. SCHUH (1984) described Alloeomimus muiri, based on Indochinese and Indonesian specimens. Alloeomimus is an Old World genus, currently composed of six species (including A. muiri) but five congeners (incl. type species of the genus) are restricted to the western Palearctic and Ethiopian Regions (Mediterranean, Middle East and Africa, cf. SCHUH 2013). Our present examination suggested that the Schuh’s species (muiri) can be evidently separated from the members of Alloeomimus in having some different characters mentioned above as well as disjunct distribution in the Oriental Region. We therefore propose a new genus Alloeomimella for it, and a new combination is accordingly established: Alloeomimella muiri (Schuh, 1984) comb. nov. (see SCHUH 1984 and YASUNAGA et al. 2013 a for detailed diagnostic characters of this species). Alloeomimella muiri was confirmed to be epigeic; several individuals including brachypterous females were found from the ground at tropical grasslands (cf. Fig. 2). On the other hand, Alloeomimus unifasciatus (Reuter, 1879) was reported to be associated with Ononis natrix L. (Fabaceae) (SCHUH 2013). In addition, a congener from the Middle East, A. kurdus Hoberlandt, 1953, was reported to occasionally inhabit galls of aphids (WAGNER 1974), which implies that Alloeomimus members may be associated with aerial parts of plants and may not prefer an epigeic habitat. Color habitus images of the five Alloeomimus species are available on a website (https: // www. discoverlife. org / mp / 20 q? search = Alloeomimus). Based on the developed stridulatory device (Figs 63 − 64), reduced scent efferent system (Figs 59 − 60), setiform parempodia, similar male and female genitalic structures and habitat preference, Alloeomimella is assumed to be most closely allied to Hallodapus rather than Alloeomimus.	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC57FFF4FC63FEA0FB95F8D2.taxon	diagnosis	Diagnosis. General shape rather ant-like (Fig. 13); small to moderate size (total length 3.0 – 4.5 mm); brown to fuscous basic coloration; more or less terete antennal segments II – IV; weakly swollen, trapezoidal pronotum that is not much constricted anteriorly (Fig. 65); ear-like ostiolar peritreme (Figs 71, 131); presence of white fascia on clavus just posterior to scutellum (Fig. 13); possession of stridulatory device (currently confirmed in two species, see Table 1 and Figs 66 – 67, 132 – 133); weakly fleshy, apically convergent parempodia (Fig. 68); and short endosoma with an apical spine of various length and a developed secondary gonopore. Further diagnostic characters and detailed description were provided by SCHUH (1984).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC57FFF4FC63FEA0FB95F8D2.taxon	distribution	Distribution. Known widely from the Sundaland west of the Wallacea, Indochina, southern China, Philippines and Japanese Okinawa Island (YASUNAGA 2001, 2012).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC57FFF4FC63FEA0FB95F8D2.taxon	discussion	Discussion. Although this genus is superficially similar to Cleotomiroides, the male and female genitalic structures of these genera are significantly different from each other (see YASUNAGA 2012, DUWAL et al. 2017). YASUNAGA (2012) suggested that the similarity between Cleotomiris and Cleotomiroides was only superficial. Present observation also confirmed that some species of Cleotomiroides and Wygomiris are associated with broadleaf trees (on which immature forms were found, cf. Figs 12, 17) and lack the stridulatory mechanism. General shape of the female genitalia of Cleotomiris rather resembles that of Wygomiris. Almost all of the available specimens of these three genera were collected using UV light traps at well preserved subtropical or tropical forests. Therefore, members of these genera are currently considered to be arboreal. Our examination by a SEM could find out the stridulatory devices in two Cleotomiris species, C. miyamotoi Yasunaga (Figs 66 – 67) from Okinawa Island, Japan and an (presumably) undescribed species currently represented by a single female from Yunnan Province, China (NMPC). This unique Chinese species has rather developed FWS + MFP (Figs 132 – 133) which appear to retain the original function (stridulation) although it is superficially most similar to C. miyamotoi or C. schneirlai Schuh, 1984 (Philippines). From southern China (Yunnan, Sichuan) to northern Indochina, more than a few undetermined specimens representing at least three closely related species of Cleotomiris or Cleotomiroides are present. Therefore, definitive determination of these taxa is beyond a scope of the present study.	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC57FFF5FC56F851FD0EF894.taxon	diagnosis	Diagnosis. Externally very similar to Cleotomiris, from which this genus can be distinguished by slender and almost linear antennal segments II – IV (Figs 11 – 12), strongly protruding evaporative area of scent efferent system (Figs 74 – 75), remarkably broadened endosoma that is more or less elaborated apically with some appendages (Fig. 195), and small but thick-rimmed sclerotized ring (see YASUNAGA 2012). Detailed diagnosis and description were provided by SCHUH (1984).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC57FFF5FC56F851FD0EF894.taxon	discussion	Discussion. See discussion under Cleotomiris.	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC56FFF7FF33F8BCFDDBFB43.taxon	description	(Figs 33 – 35, 74 − 76, 192 – 195)	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC56FFF7FF33F8BCFDDBFB43.taxon	materials_examined	Type material. HOLOTYPE: ♁, INDONESIA: JAVA: Malang, Buring, S 07 ° 59 ′ 40.0 ″ – 42.0 ″ E 112 ° 39 ′ 38.0 ″ – 39.1 ″, 513 – 518 m alt., 24 Aug 2005 (afternoon), T. Ishikawa (TUAK) (AMNH _ PBI 00380631).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC56FFF7FF33F8BCFDDBFB43.taxon	diagnosis	Differential diagnosis. Recognized by its relatively large size; fuscous basic coloration contrasting with rusty apical half of the clavus (Fig. 33); conspicuous white fascia and maculae on hemelytron (Figs 33, 35); and form of male genitalia (Figs 192 − 195). This new species can be distinguished from two known congeners by the fuscous general coloration, broad hypophysis of the left paramere (Fig. 193), and elongate apical appendage and spinulate flap-like process of the endosoma (Fig. 195).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC56FFF7FF33F8BCFDDBFB43.taxon	description	Description. Male. Body generally blackish brown, relatively small, myrmecomorphic; dorsal surface weakly shining, widely shagreened or roughened, with sparsely distributed, woolly semierect setae and with more sparsely distributed, longer, upright setae (Fig. 35). Head dull brown, oblique; eyes large; vertex narrowly carinate basally. Antenna dull yellowish brown, almost linear; segment II slightly incrassate toward apex; segments III and IV brown, weakly terete. Labium shiny dark brown, reaching subapical part of mesocoxa. Pronotum with a narrow collar, narrower than all antennal segments; pleura shiny fuscous, minutely rugose; ostiolar peritreme strongly protruding medially, with ivory white posterior margin of evaporative area. Hemelytron with a white, continuous fascia posterior to scutellum across clavus, corium and exocorium (Fig. 33); apical 1 / 4 of corium rusty; membrane smoky brown, except for pale, translucent base. Coxae and legs brown or darker; meso- and metacoxae widely pale brown; pretarsus with rather fleshy, apically convergent parempodia (Fig. 76). Abdomen wholly dark brown, somewhat shagreened (Fig. 34). Male genitalia (Figs 192 – 195): Genital segment with a ventral spine (Fig. 192). Left paramere with rather stout hypophysis (Fig. 193). Phallotheca almost straight, tapered, with a spine near apex (Fig. 194). Endosoma J-shaped, stout, with an elongate apical appendage and a weakly sclerotized, rounded flap-like process beside thick-rimmed secondary gonopore (Fig. 196). Measurements. Male (holotype): Total length of body 3.33; head width including eyes 0.72; vertex width 0.21; lengths of antennal segments I – IV 0.30, 1.07, 0.60, 0.44; total labial length 1.28; basal width of pronotum 1.02; maximum width across hemelytron 1.05; and lengths of metafemur, tibia and tarsus 1.14, 1.70, 0.30. Female. Unknown.	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC56FFF7FF33F8BCFDDBFB43.taxon	etymology	Etymology. Named after a Japanese heteropterist, Dr. Tadashi (nickname Chu) Ishikawa who collected the holotype specimen of this new species; a noun in genitive case.	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC56FFF7FF33F8BCFDDBFB43.taxon	biology_ecology	Biology. Unknown; Dr. Ishikawa collected the holotype female by sweep-netting an unidentified broadleaf tree.	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC56FFF7FF33F8BCFDDBFB43.taxon	distribution	Distribution. Indonesia: Java (this paper).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC54FFF7FEB8FAC0FBDAFA03.taxon	diagnosis	Diagnosis. Body rather small, elongate, slender but female predominantly brachypterous (Figs 19 – 24); dorsum weakly shining, impunctate, with sparsely distributed, pale, simple, upright setae; general coloration basically brown, with pale or white maculae on corium and / or clavus, lacking greenish tinge; more or less prognathous head; eye small, contiguous to flattened pronotum collar; stridulatory device involving dorsobasal surface of metafemur and embolial edge present both in macropterous and brachypterous forms; scent efferent system small; parempodia setiform (Figs 91, 129); pygophore usually smooth, rarely with a pygophoral spine (in H. spinosus sp. nov. and H. susurratus sp. nov.); left paramere developed (Figs 85, 130); endosoma slender, elongate or sometimes very long, with well-marked secondary gonopore (Figs 94, 107, 178 – 180); ovipositors short, with developed apices; posterior wall simple; sclerotized ring small but clearly rimmed (Figs 201 – 209).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC54FFF7FEB8FAC0FBDAFA03.taxon	distribution	Distribution. Known widely from the Old World (Palearctic Region to southern Africa, northeast to Japan, Korea and Russian Far East, and Oriental Region southeast across Wallacea to New Guinea and northern Australia); most speciose in the Ethiopian and Oriental Regions (SCHUH 2013).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC54FFF7FEB8FAC0FBDAFA03.taxon	discussion	Discussion. This is the most speciose genus in the tribe Hallodapini, including about 50 valid species from the Old World. Majority of them are thermophilic, known predominantly from tropical and subtropical climate zones. All Hallodapus species we have examined (Table 1) are equipped with the distinct stridulatory device (e. g., Figs 41 – 53, 77 – 78, 81 – 84). Our recent attempt successfully detected stridulation in two Japanese species, H. centrimaculatus (approx. 2,100 Hz) and H. ravenar (800 Hz), in laboratory observation (Fig. 32). As exhibiting significant interspecific variation, extent and shape of MFP can be regarded as the effective key character for species identification (as in Figs 40 – 53). The stridulatory mechanism is equally possessed by both male and female adults (cf. Figs 40 – 46, 48 – 49) but absent in immature forms (Fig. 50). In addition to stridulation, we observed and videoed remarkable intraspecific conflict occasionally occurring between males of each Hallodapus centrimaculatus and H. ravenar in laboratory condition. Thus far as we recognized, no previous work has ever reported such fighting behavior for the Miridae. We have not yet clarified when and why they fight, and further continuing investigation is required to elucidate the meaning of their conflict. Some of our sample movies are available from the the websites mentioned in Online supplimentary data (p. 98). All Hallodapus congeners are presumed to be epigeic, inhabiting ground densely covered with weeds and / or shrubs (cf. Figs 1 − 2). YASUNAGA (2001) considered that (at least Japanese) species of Hallodapus are predaceous. During our laboratory observation, four Japanese species, H. centrimaculatus, H. kyushuensis, H. linnavuori, and H. ravenar, did not attack or prey on live tiny arthropods but prefer to feed on the cadavers of various insects and spiders (sometimes much larger than these hallodapines) as well as birds’ droppings. This evidence could imply that Hallodapus species are scavengers instead of predators. The eggs and oviposition were for the first time confirmed for H. centrimaculatus and H. ravenar (Figs 7 – 8, 108 – 109); the female adults of both species were found to oviposit near the roots of weeds (Fig. 6).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC54FFEFFCE2FA38FD80FB63.taxon	description	(Figs 25 – 27, 52, 116 – 118, 174 – 176)	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC54FFEFFCE2FA38FD80FB63.taxon	materials_examined	Type material. HOLOTYPE: ♁, TAIWAN: PINGDONG (= PINGTUNG): Sheding, 21 ° 57 ′ 24.7 ″ N, 120 ° 49 ′ 40 ″ E, UV lighting, 22 Sep 2012, J. F. Tsai & Y. C. Lan (NMNS) (AMNH _ PBI 00380632).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC54FFEFFCE2FA38FD80FB63.taxon	diagnosis	Differential diagnosis. Recognized by its tiny size; relatively ovoid body; yellowish brown antenna with reddish extreme base of segment I; narrowly distributed MFP (Fig. 52); triangular pygophore (Fig. 174); roundly inflated sensory lobe of left paramere (Fig. 175); and C-shaped endosoma, with an apically bulbous process near apex (Fig. 176). Combination of these characters enables this new species to be distinguished from other Asian congeners.	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC54FFEFFCE2FA38FD80FB63.taxon	description	Description. Male (holotype). Body generally coffee brown, small, rather ovoid (Figs 25 – 26); dorsal surface weakly shining, with sparsely distributed, pale, simple, semierect setae. Head short, with reddish basal margin; eyes small, less than half as wide as vertex in dorsal view. Antenna almost uniformly yellowish brown; extreme base of segment I reddish brown; segment II slightly longer than III. Labium reddish brown, exceeding apex of metacoxa, reaching abdominal sternum VIII. Pronotum weakly shining, trapezoidal, not strongly constricted anteriorly; pleura shiny dark brown; scent efferent system somber pale brown, small. Hemelytron with two pairs of white maculae as in Fig. 25 (anterior macula across clavus, corium and embolium rectangular and posterior macula across apices of corium and embolium triangular); FWS roundly notched (Figs 27, 117); cuneus brown, narrowed; membrane pale smoky brown. Coxae and legs pale brown; apical half of metafemur and basal 1 / 3 of each tibia obscure; MFP distributed narrowly on median part of metafemur, with each plectrum tiny, circular (Figs 52, 118). Abdomen wholly fuscous, short. Male genitalia (Figs 174 – 176): Pygophore triangular (Fig. 174); left paramere with roundly expanded sensory lobe (Fig. 175); endosoma C-shaped, with an apically bulbous process near apex (Fig. 176). Measurements. Male (holotype, mm): Total length of body 2.65; head width including eyes 0.51; vertex width 0.29; lengths of antennal segments I – IV 0.30, 0.97, 0.90, 0.48; total labial length 1.28; basal width of pronotum 0.78; maximum width across hemelytron 0.93; and lengths of metafemur, tibia and tarsus 1.02, 1.47, 0.41. Female. Unknown.	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC54FFEFFCE2FA38FD80FB63.taxon	etymology	Etymology. Named after a Taiwanese heteropterist, Dr. Jing-Fu Tsai who collected a valuable specimen herein designated as the holotype.	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC54FFEFFCE2FA38FD80FB63.taxon	biology_ecology	Biology. Unknown; the holotype specimen was collected by a UV light trap at rather sparse subtropical forest in Sheding National Park, southernmost part of Taiwan.	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC54FFEFFCE2FA38FD80FB63.taxon	distribution	Distribution. Taiwan (this paper).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC4CFFEFFF31FAD8FAE6F9A3.taxon	description	(Figs 19, 28 – 31, 47, 119 – 124)	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC4CFFEFFF31FAD8FAE6F9A3.taxon	materials_examined	Type material. HOLOTYPE: ♁, THAILAND: NAKHON NAYOK: Sarika (at garden lawn), 14 ° 18 ′ 07 ″ N 101 ° 18 ′ 09 ″ E, at FL light, 20 Dec 2010, T. Yasunaga (AMNH _ PBI 00380633) (DOAT). Additional specimen examined. THAILAND: NAKHON RATCHASIMA: Wang Nam Khiew, Sakaerat Environmental Research Station, 14 ° 30 ′ 27 ″ N, 101 ° 55 ′ 39 ″ E, 410 m alt., UV light trap, 20 Aug 2008, T. Yasunaga, 1 ♁ (TYCN).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC4CFFEFFF31FAD8FAE6F9A3.taxon	diagnosis	Differential diagnosis. Superficially most similar to H. brunneus (Poppius) (Fig. 20), from which the present new species (Fig. 19) can be distinguished by the larger anterior pair of white maculae contiguous to each other (Fig. 28), totally fuscous femora, narrowly distributed MFP with each plectrum somewhat ovoid (Figs 47, 123), presence of pygophoral spine (Figs 30, 124), and C-shaped endosoma (Fig. 30) with an apically bulbous process near the apex (Fig. 31). In H. brunneus, the maculae of the anterior pair are smaller and separated each other, bases of meso- and metafemora are creamy white, FWS is rather sparsely distributed (Fig. 110), MFP is more widely distributed and each plectrum squared (Figs 40, 111), pygophore lacks any spine, and endosomal subapical process is short and thickened (Fig. 178).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC4CFFEFFF31FAD8FAE6F9A3.taxon	description	Description. Male (holotype). Macropterous; body generally dark brown, almost medium-sized, elongate, parallel-sided (Figs 19, 28); dorsal surface weakly shining, with uniformly distributed, woolly, reclining setae and sparsely distributed, dark, simple, semierect setae. Head tinged with red, pointed in front; eyes small, less than half as wide as vertex in dorsal view. Antenna pale brown; segment I dark brown, with yellow fascia inward; segment II slightly darkened, inwardly with a weak stripe that is nearly invisible in dry-preserved specimen, subequal in length to III; segment IV about twice as long as I. Labium reddish brown, reaching but not exceeding apex of metacoxa; segment III and base of IV pale brown. Pronotum weakly shining, weakly constricted laterally at calli; pleura shiny dark reddish brown; scent efferent system with knob-like ostiolar peritreme (Fig. 121). Hemelytron with two pairs of white maculae as in Fig. 28 (anterior pair of maculae larger, contiguous to each other and posterior macula squared); FWS as in Figs 29, 122; membrane pale smoky brown. Coxae and femora dark brown, except for meso- and metacoxae yellowish brown; MFP rather narrowly distributed, with each plectrum rectangular (Figs 47, 123). Abdomen wholly reddish dark brown, relatively long. Male genitalia (Figs 30 – 31, 124): Pygophore with a distinct pygophoral spine (Fig. 30); endosoma C-shaped (Figs 30, 124), with an apically bulbous process near the apex (Fig. 31). Measurements. Male (holotype, mm): Total length of body 2.94; head width including eyes 0.55; vertex width 0.32; lengths of antennal segments I – IV 0.35, 1.05, 1.01, 0.69; total labial length 1.25; basal width of pronotum 0.81; maximum width across hemelytron 0.92; and lengths of metafemur, tibia and tarsus 1.05, 1.52, 0.39. Female. Unknown.	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC4CFFEFFF31FAD8FAE6F9A3.taxon	etymology	Etymology. From Latin, spinosus [= spiny, thorny], referring to a characteristic ventral spine on the pygophore of this new species; an adjective.	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC4CFFEFFF31FAD8FAE6F9A3.taxon	biology_ecology	Biology. Unknown; habitat environment as in Fig. 2.	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC4CFFEFFF31FAD8FAE6F9A3.taxon	discussion	Comment. YASUNAGA et al. (2013 b) reported H. brunneus from Thailand, but the present examination revealed the Thai specimen represents an undescribed species herein documented as H. spinosus sp. nov. which is unique by having a noticeable pygophoral spine (see Table 1).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC4CFFE8FCC8F9E0FD90F903.taxon	description	(Figs 36 – 39, 125 – 130, 181 – 185)	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC4CFFE8FCC8F9E0FD90F903.taxon	materials_examined	Type material. HOLOTYPE: ♁, THAILAND: CHIANG MAI: Chom Thong, 18 ˚ 25 ′ N, 98 ˚ 40 ′ E, 24 – 27 Apr 1991, J. Horák (NMPC) (AMNH _ PBI 00380644). PARATYPES: THAILAND: CHIANG MAI: Same locality as for holotype, 23 – 27 Apr 1991, S. Bílý (NMPC).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC4CFFE8FCC8F9E0FD90F903.taxon	diagnosis	Differential diagnosis. Recognized by its small size (<2.5 mm) comparable with H. centrimaculatus; widely darkened dorsum; notched inner margin of anterior white macula on corium (Fig. 36); rather granulate, rounded MFP (Fig. 127); enlarged, stout left paramere (Fig. 183); presence of distinct spine and dorsal knob on pygophore (Figs 130, 181); slender phallotheca (Fig. 184); and rather sigmoid endosoma (Fig. 185). This new species is most similar to or conspecific with a New Guinean species identified by SCHUH (1984) as H. ‘ near’ pseudosimilis (Schuh, 1974) originally described from South Africa (SCHUH 1974, Fig. 28); however, H. pseudosimilis is obviously larger in size (total length 3.2 mm and maximum width across hemelytra 1.04 mm), and the color pattern (e. g. broadly pale corium and pale basal 1 / 3 of metafemur) significantly differs from that exhibited by H. susurratus (cf. SCHUH 1974).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC4CFFE8FCC8F9E0FD90F903.taxon	description	Description. Male (holotype). Macropterous; body generally coffee brown, small-sized (<2.5 mm), nearly parallelsided (Fig. 36); dorsal surface matte, with sparsely distributed, woolly, reclining setae and dark, simple, erect setae. Head weakly shining. Antenna creamy white, partly tinged with red; segment I with pale brown basal half and reddish extreme apex. Labium reddish brown, long, exceeding apex of metacoxa, reaching apical margin of abdominal sternum VII; segment III and base of IV pale brown. Pronotum weakly shining, almost trapezoidal (Fig. 125); pleura shiny dark reddish brown; scent efferent system yellowish brown, with ostiolar peritreme not produced (Fig. 128). Hemelytron with two pairs of white maculae as in Fig. 36 (anterior pair of maculae larger, separated from each other and notched inward); FWS as in Fig. 126; membrane pale smoky brown, with slightly darkened veins. All coxae and trochanters yellowish brown, except for basal part of each coxa more or less obscure; all femora reddish or chocolate brown, with pale extreme apices; MFP rather granulate, with each plectrum rounded (Fig. 127); tibiae creamy white; parempodia setiform, long (Fig. 129). Abdomen wholly reddish dark brown, shining. Male genitalia (Figs 180 – 185): Pygophore with a distinct pygophoral spine and a dorsal knob-like protuberance (Figs 130, 181); left paramere enlarged, stout (Fig. 183); right paramere tiny (Fig. 182); phallotheca slender (Fig. 184); endosoma slender, sigmoid, tapered apically, without noticeable process or branch (Fig. 185). Measurements. Male (holotype, mm): Total length of body 2.32 – 2.45; head width including eyes 0.45 – 0.47; vertex width 0.18 – 0.19; lengths of antennal segments I – IV 0.24 – 0.25, 0.75 – 0.76, 0.54 – 0.57,?; total labial length 1.05 – 1.17; basal width of pronotum 0.69 – 0.74; maximum width across hemelytron 0.72 – 0.75; and lengths of metafemur, tibia and tarsus 0.90, 1.36 – 1.37,?. Female. Unknown.	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC4CFFE8FCC8F9E0FD90F903.taxon	etymology	Etymology. From Latin, susurratus [= muttering, whispering], referring to the small size and rather granulate metafemoral plectrum of this new species; adjective.	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC4CFFE8FCC8F9E0FD90F903.taxon	biology_ecology	Biology. Unknown; two available specimens were collected at montane zone of northern Thailand.	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC4CFFE8FCC8F9E0FD90F903.taxon	distribution	Distribution. Thailand (this paper).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC4BFFE8FEC2F900FAB5F9A3.taxon	diagnosis	Diagnosis. Always macropterous in both sexes; body elongate oval, nearly parallel-sided, small to moderate in size (total length 3 – 5 mm), not very ant-like (rather conventional habitus in the Miridae); basic coloration brown to fuscous; dorsum weakly shining or matte, usually with pale, simple, upright setae and silvery or golden, reclining setae; head short, rather vertical; antenna uniformly thick; pronotum more or less shining; metathoracic scent efferent system large, usually produced at middle; stridulatory device (FWS + MFP) missing (currently possessed only by W. paveli and W. phormictes described below); pygophoral spine (PS) present; endosoma J- or L-shaped, usually with simple apical spine and small secondary gonopore; and bursa copulatrix with enlarged, ovoid sclerotized ring. Further diagnostic characters were provided by SCHUH (1984) and YASUNAGA (2012).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC4BFFE8FEC2F900FAB5F9A3.taxon	distribution	Distribution. Oriental Region: SE China (including Hong Kong), Laos, Nepal, Philippines, Taiwan, Thailand, Vietnam (DUWAL et al. 2017). Several undescribed species of Wygomiris appear to occur in India, the Philippines and New Guinea (SCHUH 1984).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC4BFFE8FEC2F900FAB5F9A3.taxon	discussion	Discussion. Wygomiris is now represented by nine species (including two new species below) from the Oriental Region. Members of this genus appear to be associated with aerial parts of plants (possibly broadleaf trees as mentioned in above tribal discussion). Currently, the stridulatory device is confirmed only in W. paveli sp. nov. and W. phormictes sp. nov.; however, FWS and MFP of them are both reduced (Figs 141 – 142, 150, 152 – 153), similar in overall appearance to those possessed by Cleotomiris miyamotoi (Figs 66 – 67). Judging from the condition of the stridulatory device, we presume that (1) modern members of Alloeomimella, Cleotomiris, Hallodapus and Wygomiris had been derived from the epigeic common ancestor migrating from aerial parts of plants, (2) the derived element of Cleotomiris + Wygomiris subsequently returned to the original habitat (aerial parts of plants), and (3) the stridulatory device may be obsoleted or reduced in modern Cleotomiris and Wygomiris, as seen in the species of Acrorrhinium, Cleotomiroides and Systellonotus malaisei Lindberg, 1934 which inhabit the aerial parts of plants and lack the stridulatory device. During examining specimens or photographic images, teratological antennal segments (possibly fused segments III + IV) were confirmed in the fifth instar nymph of Wygomiris kaliyahae collected in Nakhon Ratchasima, Thailand (Fig. 18, left antenna) and adult male of W. paveli sp. nov. from Pingtung, Taiwan (AMNH _ PBI 00380635) (Figs 14 – 15, 166 – 167, right). Although teratological specimens occur less frequently in Cimicomorpha than in the Pentatomomorpha (WHEELER 2001), a similar antennal aberration in Apolygus nigrovirens (Kerzhner, 1988) (Mirinae: Mirini) was recently documented by YASUNAGA (2018).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC4BFFEAFC37F989FB05F950.taxon	description	(Figs 14 – 15, 140 – 143, 164 – 167)	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC4BFFEAFC37F989FB05F950.taxon	materials_examined	Type material examined. HOLOTYPE: ♀, THAILAND: NAKHON NAYOK: Sarika (lodge garden), 14 ° 18 ′ 39 ″ N 101 ° 18 ′ 00 ″ E, at light, 13 Mar 2009, T. Yasunaga (AMNH _ PBI 00379518) (DOAT). Additional material examined. VIETNAM: SON LA PROVINCE: Moc Chau, 940 m alt., 20 ° 49 ′ N 104 ° 42 ′ E, light trap, 19 Jun 1997, M. Tomokuni, 2 ♁♁ (NSMT); Dao Cao Pha, 420 m E of Ban Song, 21 ° 09 ′ 39 ″ N 103 ° 35 ′ 46 ″ E, 23 Jun 1997, M. Tomokuni, 1 ♀ (NSMT).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC4BFFEAFC37F989FB05F950.taxon	diagnosis	Differential diagnosis. Most closely related to P. mingorum Schuh, 1984; distinguished by its uniformly dark brown body; smaller eyes; wider vertex; weakly shining, matte pronotum; densely distributed, silvery, reclining setae on hemelytron (Figs 161 − 163); tapered apical part of left paramere hypophysis (Fig. 188) and longer apical branches on endosoma (Fig. 190), in addition to developed female genital chamber (Fig. 191). Although YASUNAGA (2012) supposed W. nanae as the closest relative of W. indochinensis Schuh, 1984, W. mingorum is now considered to be sister to W. nanae, judging from the similar shape of endosoma bearing three apical branches. In external appearance, W. nanae can be readily distinguished from W. indochinensis and W. mingorum by the uniformly dark brown coloration and dense dorsal vestiture (Figs 161, 163); the latter two species have contrasting two-tone pattern (SCHUH 1984, YASUNAGA 2012) as seen in W. phormictes sp. nov. (Fig. 170).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC4BFFEAFC37F989FB05F950.taxon	description	Redescription. Male. General coloration and shape similar to those of female. Macropterous; body almost uniformly chestnut or dark brown, elongate oval, comparatively large in size; dorsal surface rather matte, with simple, semierect setae, long, erect setae and wooly, reclining setae. Head tinged with red, weakly shining. Antenna reddish brown. Labium reddish brown, reaching apex of mesocoxa. Pronotum somewhat roughened, with uniformly distributed, simple, semierect setae; collar pruinose, narrower than base of antennal segment II; scent efferent system widely ivory white, with a median knob. Hemelytra somewhat lighter than pronotum and scutellum, with uniformly distributed, silvery, reclining setae and sparsely distributed, simple, long, erect setae; membrane smoky brown. All coxae and legs brown, partly tinged with red; meso- and metacoxae lighter. Abdomen shiny chestnut brown, partly reddish except for posterior margin of each sternum (Fig. 161). Male genitalia (Figs 186 – 190): Pygophore with distinct PS (Fig. 186). Left paramere with apically tapered hypophysis (Fig. 188); endosoma three-branched, with a wide membranous area along secondary gonopore (Fig. 190). Measurements (♁♁ / ♀♀, mm) [holotype from Thailand]. Total length of body 3.43 – 3.78 / 3.43 [3.60]; head width including eyes 0.72 – 0.75 / 0.71 [0.71]; vertex width 0.30 / 0.33 [0.33]; lengths of antennal segments I – IV 0.25 – 0.26, 0.93, 0.60, 0.48 / 0.24 [0.19], 0.87 [0.90], 0.57 [0.57], 0.51 [?]; labial length 1.20 – 1.22 / 1.08 [1.20]; basal width of pronotum 1.09 – 1.16 / 1.19 [1.08]; maximum width across hemelytron 1.32 – 1.35 / 1.49 [1.39]; and lengths of metafemur, tibia and tarsus 1.09 – 1.14, 1.65, 0.30 / 1.10 [1.13], 1.59 [1.50], 0.29 [0.30].	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC4BFFEAFC37F989FB05F950.taxon	biology_ecology	Biology. Unknown; all available specimens were collected by UV lightings around arable lands or secondary forests.	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC4BFFEAFC37F989FB05F950.taxon	distribution	Distribution. Thailand (Nakhon Nayok) (YASUNAGA 2012), Vietnam (Son La) (this paper).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC49FFE7FCE8F8D3FD50FD63.taxon	description	(Figs 14 – 15, 36 – 39, 140 – 143, 196 – 200)	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC49FFE7FCE8F8D3FD50FD63.taxon	materials_examined	Type material. HOLOTYPE: ♁, TAIWAN: PINGDONG (= PINGTUNG): Siangjiao Bay, 21 ° 55 ′ 42.7 ″ N, 120 ° 49 ′ 40 ″ E, UV lighting, 16 Feb 2011, C. T. Chiu & Y. C. Lan (AMNH _ PBI 00380634) (NMNS). PARATYPES: TAIWAN: PINGDONG: Same locality as for holotype, 14 Mar 2012, Y. C. Lan & J. Z. Wang, 1 ♁ (NMNS); Nanrenshan, UV lighting, 26 Apr 2011, Y. C. Lan & Y. H. Peng, 1 ♁ (NMNS); Mutan, Cacevakan, 22 ° 06 ′ 34.9 ″ N, 120 ° 48 ′ 06.1 ″ E, UV lighting, 16 Mar 2017, T. Yasunaga, 1 ♁ (00380635) (TYCN). HUALIEN: Yuri, Chohsishan, Malaise trap, 24 Apr 2009 – 12 Dec 2012, W. T. Yang & K. W. Huang, 5 ♁♁, 2 ♀♀ (NMNS); Hsiulin, Wenlan, Malaise trap, 24 Jun – 25 Aug 2009, K. W. Huang & W. T. Yang, 1 ♁ (00380636) (NMNS); same data except for date 13 Sep – 24 Nov 2010, 1 ♁ (NMNS). CHIAYI: Chungpu, Yunshui Station, Mercury lighting, 7 – 8 Nov 2006, Lin, Chan, Liang & Lan, 1 ♁ (00380637) (NMNS).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC49FFE7FCE8F8D3FD50FD63.taxon	diagnosis	Differential diagnosis. Recognized readily by its moderate size; generally castaneous basic coloration (Figs 14 – 15); rather small eyes; rather slender antenna; weakly shining pronotum; yellowish brown fascia on hemelytron; possession of stridulatory device (FWS + MFP, Figs 55, 141 – 142); and contrastingly darkened abdomen (Figs 164 – 165). Based on the similar genitalic structure (endosomal shape in particular), this new species is apparently sister to W. indochinensis which however has larger eye (Fig. 144) and rather terete antennal segments, and lacks the stridulatory device (Fig. 145) (see YASUNAGA 2012 and DUWAL et al. 2017, for detailed diagnosis and color habitus images of W. indochinensis).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC49FFE7FCE8F8D3FD50FD63.taxon	description	Description. Male. Macropterous; body generally castaneous, elongate oval, parallel-sided, moderate in size; dorsal surface weakly shining, with sparsely distributed, simple upright setae and woolly (not scalelike) reclining setae. Head somewhat produced anteriorly, with pale, short, erect setae. Antenna pale reddish brown, rather slender, not significantly terete; apex of segment II slightly inflated, about as long as basal width of pronotum; two distal segments reddish brown. Labium shiny pale reddish brown, slightly exceeding apex of mesocoxa, slightly longer than antennal segment II or metafemur; segment IV dark reddish brown. Pronotum weakly shining, not polished; pleura including scent efferent system yellowish brown. Hemelytron chestnut brown, with a yellowish transverse fascia posterior to scutellum; FWS weak, minutely sutured (Figs 55, 141); membrane smoky brown, with pale, translucent basal half. Coxae and legs pale brown; each femur and tibia partly obscure; MFP minutely scattered (Figs 55, 142). Abdomen widely shiny dark brown. Male genitalia (Figs 196 – 198): Pygophore with a small pygophoral spine (Figs 143, 196). Left paramere with tumid, bulbous sensory lobe (Fig. 197). Endosoma L-shaped, with a long, tapered apical process (Fig. 198). Female. Similar in overall appearance to male; body somewhat rounded laterally. Female genitalia (Figs 199 – 200): Bursa copulatrix rather narrow, not strongly sclerotized; sclerotized ring thickened posteriorly (Fig. 199); ovipositors (gonapophyses I and II) rather elongate. Measurements (♁♁ / ♀♀, mm). Total length of body 3.18 – 3.26 / 3.18 – 3.43; head width including eyes 0.67 – 0.70 / 0.64 – 0.70; vertex width 0.28 – 0.30 / 0.33 – 0.38; lengths of antennal segments I – IV 0.25 – 0.27, 1.03 – 1.05, 0.57 – 0.60, 0.45 – 0.47 / 0.24 – 0.30, 0.91 – 1.00, 0.54,? [in aberrant ♁ specimen (Figs 14 – 15, 142 – 143): 0.29, 1.19, 0.60, 0.51 (right) and 0.29, 1.35, 0.86 (left)]; labial length 1.21 – 1.23 / 1.36 – 1.37; basal width of pronotum 1.03 – 1.05 / 1.00 – 1.08; maximum width across hemelytron 1.06 – 1.16 / 1.21 – 1.32; and lengths of metafemur, tibia and tarsus 1.03 – 1.05, 1.66 – 1.68, 0.30 – 0.32 / 1.22, 1.53, 0.32.	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC49FFE7FCE8F8D3FD50FD63.taxon	etymology	Etymology. Named in honor of the eminent Czech heteropterist, Dr. Pavel Štys (former head of Department of Zoology of the Charles University, Praha), who regrettably passed away in 2018.	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC49FFE7FCE8F8D3FD50FD63.taxon	biology_ecology	Biology. Adults were occasionally attracted to UV light at night. Collection records suggest that this mirid has two or three generations per year.	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC49FFE7FCE8F8D3FD50FD63.taxon	distribution	Distribution. Taiwan: Hualien, Pingtung (this paper).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC44FFE7FF29FD20FA1BFD03.taxon	description	(Figs 146 – 154, 170 – 173)	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC44FFE7FF29FD20FA1BFD03.taxon	materials_examined	Type material. HOLOTYPE: ♀, THAILAND: NAKHON RATCHASIMA: Wang Nam Khieo, Sakaerat Environmental Research Station, 14 ° 30 ′ 27 ″ N, 101 ° 55 ′ 39 ″ E, 410 m alt., UV light trap, 22 Mar 2014, T. Yasunaga (DOAT) (AMNH _ PBI 00380645).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC44FFE7FF29FD20FA1BFD03.taxon	diagnosis	Differential diagnosis. Most similar in general appearance and similarly small size (body length <3.0 mm) to W. kaliyahae Yasunaga, 2012, from which this new species is readily distinguished by the following characters: pale brown antennal segments I and II (Fig. 170); short antennal segment II that is shorter than head width across eyes; relatively shiny pronotum; golden brown anterior part of hemelytron (Fig. 171); constricted middle margin of embolium (Figs 149, 171); possession of noticeable stridulatory device (FWS + MFP, Figs 149 – 150, 152 – 153); pale coxae (Fig. 172); short metafemur almost equal in length to mesofemur (Fig. 173); mesotibia longer than mesofemur; and metatarsomere II as long as III (Fig. 151).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC44FFE7FF29FD20FA1BFD03.taxon	description	Description. Female (holotype). Macropterous; body small (2.7 mm); dorsal surface relatively shining, with sparsely distributed, simple upright setae and woolly reclining setae. Head dark brown, weakly shining. Antenna pale brown, partly tinged with red, generally short; segments III and IV dark brown. Labium shiny reddish brown, slightly exceeding apex of mesocoxa. Pronotum and propleuron shining, dark brown; epimeron and episternum shiny reddish brown; scent efferent system pale somber brown; scutellum dark brown, somewhat shagreened, flat. Hemelytron dark brown, with anterior part contrastingly golden brown as in Figs 170 – 171, constricted at middle; embolium with rather developed FWS (Figs 149 – 150); membrane smoky brown, with pale, translucent basal 1 / 3. Coxae and legs pale brown; protibia, base of mesotibia, apical half of metafemur and whole metatibia reddish brown (Fig. 173); mesofemur as long as metafemur; mesotibia slightly longer than metatibia; metatibia somewhat inflated; MFP distributed on median metafemur (Fig. 152), with each plectrum hemispherical (Fig. 153). Abdomen widely shiny dark brown; abdominal sterna II – IV paler. Female genitalia: Not examined. Measurements (mm). Female (holotype): Total length of body 2.70; head width including eyes 0.60; vertex width 0.29; lengths of antennal segments I – IV 0.15, 0.53, 0.35, 0.35; labial length 0.98; basal width of pronotum 0.92; maximum width across hemelytron 1.08; lengths of mesofemur and tibia 0.78 and 1.05; and lengths of metafemur, tibia and tarsus 0.78, 1.10, 0.27. Male. Unknown.	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC44FFE7FF29FD20FA1BFD03.taxon	etymology	Etymology. From Greek, phormictes (or phormiktes = harper), referring to presence of the stridulatory device of this new species; noun in apposition.	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC44FFE7FF29FD20FA1BFD03.taxon	biology_ecology	Biology. A female adult (Fig. 170) was attracted to UV light trap at tropical dry-evergreen forest zone (cf. https: // www. tistr. or. th / sakaerat / sakaeratE / Environment / environment. htm).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
03CDBD54EC44FFE7FF29FD20FA1BFD03.taxon	distribution	Distribution. Thailand: Nakhon Ratchasima (this paper).	en	Yasunaga, Tomohide, Tamada, Yui, Hinami, Haruka, Miyazaki, Ayana, Duwal, Ram Keshari, Nagashima, Tetsuya (2019): Taxonomic review for the Asian taxa of plant bug tribe Hallodapini, with emphasis on stridulatory mechanism (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (1): 71-99, DOI: 10.2478/aemnp-2019-0007, URL: http://dx.doi.org/10.2478/aemnp-2019-0007
