taxonID	type	description	language	source
B9771C5A604655E2A6F993D68259115F.taxon	materials_examined	Material examined. Holotype: Mexico - Chiapas • male, tl 51.8 mm; Municipality of Ocosingo, km 140 carretera Palenque-Ocosingo, ca. 5 km NW of the town of Ocosingo, Arroyo La Laja, 24 Jan. 1985, J. L. Villalobos, J. C. Nates, A. Cantu leg.; CNCR 3650. Paratypes: 1 female, tl 42.7 mm, allotype; same data as for holotype; CNCR 3650 • 16 males, tl 44.3 - 57.0 mm, 7 females, 26.2 to 46.0 mm, 2 ovigerous females, 38.1, 42.7 mm; same date as for holotype; CNCR 3650 a.	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
B9771C5A604655E2A6F993D68259115F.taxon	materials_examined	Additional material examined. Mexico - Chiapas • 9 males, tl 25.0 - 45.8 mm, 30 females, 36.8 - 39.2 mm; Municipality of Ocosingo, km 140 carretera Palenque-Ocosingo, ca. 5 km NW of the town of Ocosingo, Arroyo La Laja; 07 Aug 1983; J. L. Villalobos, J. C. Nates, A. Cantu leg.; CNCR 2940 • 16 males, tl 24.3 - 54.5 mm, 23 females, tl 17.1 - 27.4 mm; Municipality of Yajalon, carretera Palenque-Ocosingo, Arroyo Yajalon; 07 Aug. 1983; J. L. Villalobos, J. C. Nates, A. Cantu; CNCR 2941.	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
B9771C5A604655E2A6F993D68259115F.taxon	description	Description. Rostrum. Short, straight, reaching slightly beyond first third of third article of antennular peduncle; upper margin with 6 - 9 regularly spaced teeth, first one behind posterior edge of orbit; lower margin with 1 - 3 teeth. Cephalon. Scaphocerite nearly 2.6 x as long as wide, outer margin straight. Carapace. Smooth; antennal spine small, slightly overreaching lower portion of orbit; hepatic spine absent. Lower orbital angle obtuse, moderately pronounced. Pereiopods. P 1: slender, reaching with distal third of carpus beyond scaphocerite; carpus slightly longer than merus; chelae 0.68 length of carpus. P 2: moderately robust, with small spines, equal in form and size, reaching with distal third of merus beyond scaphocerite; ischium 0.75 length of merus; merus as long as carpus; carpus as long as palm, with slight basal constriction; propodus 2.5 x as long as dactylus, and 1.6 x as long as carpus; palm compressed, nearly 5 x as long as high; fingers 0.62 length of palm, with numerous spinules, not gapping, tips crossing, cutting edge with 3 - 6 teeth on proximal third in both fingers. P 3 - P 5 with all joints covered with rows of small spinules. P 3 reaching with entire dactylus beyond scaphocerite, propodus 2.5 x as long as dactylus, propodus nearly 2 x as long as carpus, propodus slightly longer than merus. P 4 reaching with entire dactylus beyond scaphocerite, propodus 3 x as long as dactylus, nearly 2 x as long as carpus, propodus slightly longer than merus. P 5 reaching with half-length of dactylus end of scaphocerite, propodus 3 x as long as dactylus, propodus nearly 2 x as long as carpus, propodus slightly longer than merus. Pleon. Smooth; somite 5 with posteroventral angle of pleuron acute; abdominal somite merely 2 x as long as somite 5. Inter-uropodal sclerite without keel-shaped pre-anal carinae. Pleopods. PL 2 with appendix masculina less than 2 x length of appendix interna. Uropods. Exopodite with mobile spines as long as spiniform projection of outer margin. Telson. Broad, smooth, slightly longer than abdominal somite 6, bearing two pairs of dorsal spinules close to posterior margin. Posterior margin ending in moderately acute triangular point; two pairs of posterior spinules with several plumose setae, inner pair overreaching distal margin of telson.	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
B9771C5A604655E2A6F993D68259115F.taxon	etymology	Etymology. Villalobos Hiriart et al. (1989) dedicated this species to Dr. Alejandro Villalobos Figueroa, eminent Mexican carcinologist and founder of the CNCR. We maintain this homage by just adding the first part of his name to the specific epithet.	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
B9771C5A604655E2A6F993D68259115F.taxon	description	Size. See in material examined.	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
B9771C5A604655E2A6F993D68259115F.taxon	distribution	Distribution. Mexico, Chiapas, in the Valle de Ocosingo, Rio La Virgen, Arroyos La Laja, Maravilla, Pasila, and Yajalon (Villalobos Hiriart et al. 1989).	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
05F2E07B7D9E500A87B25F1F4EDC92CB.taxon	materials_examined	Material examined. Chile - Coquimbo • 2 males, cl 28.2, cl 36.4 mm; rio Limari, Jul. 2006; C. Gaymer leg.; CCDB 1870 • 2 males, cl 30.3, cl 86.5 mm; Limari, rio Puente; 19 Oct. 2007; L. G. Pileggi, E. C. Mossolin leg.; CCDB 2146 • 2 males, cl 5.4, cl 5.7 mm, 4 females, cl 5.6 to 10.1 mm, 2 ovigerous females, cl 14.9, cl 16.8 mm, 8 juveniles, cl 3.4 to 4.8 mm; La Serena, Playa el Faro, Avenida de Mar; 18 Oct. 2007; F. L. Mantelatto, L. G. Pileggi, E. C. Mossolin; CCDB 2327.	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
05F2E07B7D9E500A87B25F1F4EDC92CB.taxon	description	Description. Rostrum. Straight, short, nearly reaching first article of antennular peduncle; upper margin with 6 - 8 teeth, regularly spaced, one and / or two behind posterior margin of orbit; lower margin with 0 - 4 teeth. Cephalon. Scaphocerite 2 x as long as wide; outer margin convex proximally. Carapace. Smooth, with strong, acute antennal spine; hepatic spine absent. Lower orbital angle obtuse, moderately pronounced. Pereiopods. P 1 slender, reaching with most of carpus beyond scaphocerite; fingers slightly longer than palm; carpus slightly shorter than chelae; ischium and merus distinctly spinulated; carpus and chelae smooth. P 2 strong, with many spines, strong heterochely; largest cheliped reaching with half-length of merus beyond scaphocerite; ischium larger than half-length of merus; merus longer than carpus; carpus short, slightly shorter than half length of palm, with strong basal constriction; propodus 2.1 x as long as dactylus, 3.3 x as long as carpus; palm slightly inflated, more than 2.3 x as long as high; fingers shorter than palm, with numerous small spinules, cutting edges with 4 - 7 denticles of equal size. P 3 - P 5 smooth, except for sparse setae and spinules along lower margin of propodus; propodus nearly 2 x as long as carpus; propodus slightly shorter than merus; P 3 reaching with half-length of dactylus beyond scaphocerite, propodus 2 x as long as dactylus; P 4 reaching with tip of dactylus end of scaphocerite, propodus 1.5 x as long as dactylus; P 5 reaching with tip of dactylus half-length of scaphocerite, propodus 1.5 x as long as dactylus. Pleon. Smooth. Somite 5 with posteroventral angle of pleuron acute; somite 6 slightly longer than somite 5. Inter-uropodal sclerite with strong, keel-shaped pre-anal carinae. Pleopods. PL 2 with appendix masculina 2 x as long as appendix interna. Uropods. Exopodite with mobile spines slightly longer than spiniform projection of outer margin. Telson. Broad, smooth; 1.5 x as long as abdominal somite 6, bearing 2 pairs of dorsal spinules, first pair located in middle of telson, second pair located 3 / 4 of length of telson. Posterior margin rounded, ending in truncated tip, with several plumose setae and two pairs of posterior spinules, inner pair not reaching end of telson.	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
05F2E07B7D9E500A87B25F1F4EDC92CB.taxon	description	Size. See in material examined.	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
05F2E07B7D9E500A87B25F1F4EDC92CB.taxon	distribution	Distribution. Pacific coastal river basins from Peru and Chile (Holthuis 1952 a, b; Jara et al. 2006; Morales and Meruane 2013).	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
F55D7A704C735A8AA5BFCA002EE520C9.taxon	materials_examined	Material examined. Holotype: Brazil - Distrito Federal • male, cl 18.2 mm; Brasilia, riacho da Granja do Ipe; 13 Sep. 1966; Emilio Varolli (SUDEPE) leg.; M. M. G. Correa det.; MNRJ 903.	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
F55D7A704C735A8AA5BFCA002EE520C9.taxon	materials_examined	Additional material examined. Brazil - Distrito Federal • 1 ovigerous female, cl 15.6 mm, [allotype]: Brasilia, riacho da Granja do Ipe; 13 Sep. 1966; Emilio Varolli (SUDEPE) leg.; M. M. G. Correa det.; MNRJ 6464 • 1 male, cl 17.93 mm, 2 females, cl 15.3, cl 15.3 mm; Brasilia, riacho da Granja do Ipe; 23 Feb. 1972; M. M. G. Correa leg.; MNRJ 2668 • 1 male, cl 14 mm, 2 females, cl 12.6, cl 13.8 mm; Brasilia, corrego Taquara, Reserva Ecologica do IBGE (Instituto Brasileiro de Geografia e Estatistica); 05 Aug. 2014, F. L. Mantelatto, L. G. Pileggi, F. L. Carvalho leg.; CCDB 5894 • 2 males, cl 22.9, cl 23.9 mm; Brasilia, corrego Onca, upper Parana River basin; 18 Aug. 1988; E. C. Lopes leg.; CCDB 5895 • 2 males, cl 21.7, cl 24.4 mm; Brasilia, corrego Taquara, Onca, upper Parana River basin; 18 Aug. 1988; E. C. Lopes leg.; CCDB 5896 • 2 females, cl 13.8, cl 14 mm, 5 juveniles, cl 4.4 to 6.2 mm; Brasilia, riacho da Granja do Ipe, rio Roncador, Reserva Ecologica do IBGE (Instituto Brasileiro de Geografia e Estatistica); 05 Aug. 2014, F. L. Mantelatto, L. G. Pileggi, F. L. Carvalho leg.; CCDB 5897 • 2 males, cl 9.49, cl 10.23 mm, 2 females, cl 11.34, cl 11.70 mm, 4 juveniles, cl 4.55 to 6.30 mm; Brasilia, riacho da Granja do Ipe, 22 Feb. 2008, F. L. Mantelatto, E. C. Mossolin leg.; CCDB 2195.	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
F55D7A704C735A8AA5BFCA002EE520C9.taxon	description	Description of the holotype. Rostrum. Moderately high, nearly straight, distal end slightly directed upwards, reaching end of antennular peduncle, and little before the distal margin of scaphocerite; upper margin convex over orbit, with seven teeth, first and sometimes the second, slightly behind posterior edge of orbit; lower margin with one tooth. Cephalon. Scaphocerite 2.5 x as long as wide; outer margin slightly concave. Epistome forming two lobes; lobes with laterally arranged carinae. Carapace. Anterolateral region slightly roughened; hepatic spine absent. Lower orbital angle obtuse, strongly pronounced. Pereiopods. P 1 reaching with almost half length of carpus beyond scaphocerite; fingers as long as palm; carpus 1.5 x as long as chelae, 1.5 x as long as merus; articles with scattered setae, fingers with tufts of setae. P 2 similar in shape, different in size; largest one reaching with distal portion of merus beyond scaphocerite; smallest one reaching with distal end of carpus beyond scaphocerite, with fingers as long as palm; all articles with sparse setae and spines. Larger cheliped with ischium nearly as long as merus, with spinulation as in palm; merus as long as carpus, swollen, with spinulation as in palm; carpus slightly shorter than palm, swollen, with strong basal constriction; spinulation as in palm; propodus 2.5 x as long as dactylus, 2 x as long as carpus; palm with upper surface slightly compressed, somewhat swollen, covered with spinules, nearly 3 x as long as high; fingers 2 / 3 as long as palm, with numerous spinules; cutting edge of dactylus with large tooth in proximal third, slightly lower tooth in between large tooth and proximal part; cutting edge of fixed finger with tooth opposing two teeth of dactylus, with row of three denticles between proximal part and this tooth. P 3 - P 5 smooth, except for sparse setae and spinules along lower margin of propodus; propodus nearly 2 x as long as carpus; propodus slightly shorter than merus; P 3 reaching with half-length of dactylus beyond scaphocerite, propodus 2 x as long as dactylus; P 4 reaching with tip of dactylus end of scaphocerite, propodus 2.5 x as long as dactylus; P 5 reaching with tip of dactylus half-length of scaphocerite, propodus 2.5 x as long as dactylus. Pleon. Smooth. Somite 5 with posteroventral angle of pleuron rectangular, not spinose; somite 6 1.5 x as long as somite 5. Inter-uropodal sclerite with strong, keel-shaped pre-anal carinae. Pleopods. PL 2 with appendix masculina 2 x as long as appendix interna. Uropods. Exopodite with mobile spines slightly shorter than spiniform projection of outer margin. Telson. Broad, smooth, 1.5 x as long as abdominal somite 6, bearing two pairs of dorsal spinules, first pair slightly behind middle portion of telson, second pair located closer to first pair than to posterior margin. Posterior margin distinct, ending in acute point, with several plumose setae and two pairs of posterior spinules, inner pair reaching end of telson.	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
F55D7A704C735A8AA5BFCA002EE520C9.taxon	etymology	Etymology. The specific epithet Cryphiops brasiliensis was used by Gomes Correa (1973) to refer to the type locality of the species, Brasilia, the capital of Brazil. To keep that intention, we rename the species using the word Macrobrachium candango, a demonym referring to those who are native to Brasilia.	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
F55D7A704C735A8AA5BFCA002EE520C9.taxon	description	Size. See in material examined.	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
F55D7A704C735A8AA5BFCA002EE520C9.taxon	distribution	Distribution. Endemic of inland waters from Central Brazil (Distrito Federal) (Gomes Correa 1973; present paper).	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
CD5806490B795C4CA81D7FDE955D1687.taxon	materials_examined	Material examined. Mexico - Chiapas • 8 males, tl 30.5 - 49.4 mm, 15 ovigerous females, tl 30.8 - 46.3 mm; Municipality of La Trinitaria, Rancho de San Rafael del Arco, Gruta del Arco; 07 Apr. 1986; J. L. Villalobos leg.; CNCR 5759.	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
CD5806490B795C4CA81D7FDE955D1687.taxon	description	Description. Rostrum. Short, directed slightly downwards, tip directed slightly upwards, reaching or slightly overreaching joint between second and third article of antennular peduncle, and at level of distal third of scaphocerite; upper margin convex over orbit, with 5 - 8 teeth regularly spaced, first over or slightly behind posterior edge of orbit; lower margin with none or one tooth. Cephalon. Scaphocerite 2.5 x as long as wide; outer margin straight. Carapace. Smooth, with minute punctuations; antennal spine small, slightly overreaching lower portion of orbit; hepatic spine absent. Lower orbital angle obtuse, moderately pronounced. Pereiopods. P 1 slender, reaching with nearly entire chelae beyond scaphocerite; fingers slightly longer than palm; chelae 2 / 3 length of carpus. P 2 moderately robust, with several spines, equal in form and size, reaching with proximal third of carpus beyond scaphocerite; ischium evidently shorter than merus; merus as long as carpus; carpus as long as palm, with basal constriction; propodus 2 x as long as dactylus, 2 x as long as carpus; palm inflated, nearly 3 x as long as high; fingers as long as palm, with numerous small spinules; cutting edge with two denticles of same size in both teeth. P 3 - P 5 with all joints covered with row of small spinules on the lower margin; P 3 reaching with entire dactylus beyond scaphocerite, propodus 2 x as long as dactylus, propodus slightly longer than merus; P 4 reaching with tip of dactylus end of scaphocerite, propodus 3 x as long as dactylus, propodus slightly longer than merus; P 5 reaching with tip of dactylus half-length of scaphocerite, propodus 3 x as long as dactylus, propodus slightly longer than merus. Pleon. Smooth. Somite 5 with posteroventral angle of pleuron acute; somite 6 nearly 2 x as long as somite 5. Inter-uropodal sclerite without, keel-shaped pre-anal carinae. Pleopods. PL 2 with appendix masculina 2 x as long as appendix interna. Uropods. Exopodite with mobile spines as long as spiniform projection of outer margin. Telson. Broad, smooth, slightly longer than abdominal somite 6, bearing two pairs of dorsal spinules closer to posterior margin of telson. Posterior margin ending in moderately acute triangular point, with several plumose setae and two pairs of posterior spinules, inner pair overreaching end of telson.	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
CD5806490B795C4CA81D7FDE955D1687.taxon	description	Size. See in material examined.	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
CD5806490B795C4CA81D7FDE955D1687.taxon	distribution	Distribution. Only known from the type locality (Holthuis 1973; present paper).	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
F151D75D37E856578F7D52F400C2FFD9.taxon	materials_examined	Material examined. Mexico - Chiapas • 16 males, tl 31.1 - 43.3 mm, 16 ovigerous females, tl 21.6 - 35.5 mm; Municipality of La Trinitaria, Ruinas de Chincultik, Cenote La Cueva; 07 Apr. 1986; J. L. Villalobos-Hiriart, J. C. Nates-Rodriguez, A. Cantu-Diaz Barriga leg; CNCR 7898.	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
F151D75D37E856578F7D52F400C2FFD9.taxon	description	Description. Rostrum. Short, directed downwards, reaching joint between second and third articles of antennular peduncle; upper margin with 5 - 8 teeth regularly spaced, first one at level or slightly behind posterior edge of orbit; lower margin with 1 - 3 teeth. Cephalon. Scaphocerite 2.6 x as long as wide, outer margin straight. Carapace: Smooth, with minute punctuations; antennal spine small, slightly overreaching lower portion of orbit; hepatic spine absent. Lower orbital angle subacute, moderately pronounced. Pereiopods. P 1 slender, reaching with entire chelae or small part of carpus beyond scaphocerite; fingers slightly longer than palm; chelae 2 / 3 length of carpus. P 2 moderately robust, with spines, equal in form and size, reaching with proximal third of carpus beyond scaphocerite; ischium evidently shorter than merus; merus as long as carpus; carpus as long as palm, with basal constriction; propodus 2.2 x as long as dactylus, 2 x as long as carpus; palm inflated, nearly 3 x as long as high; fingers slightly shorter (0.8) than palm, with numerous small spinules, not gaping, tips crossing, cutting edges with two similar denticles closer to proximal portion. P 3 - P 5 with all joints covered with row of small spinules on lower margin; P 3 reaching with entire dactylus beyond scaphocerite, propodus 2 x as long as dactylus, propodus nearly 2 x as long as carpus, propodus slightly longer than merus; P 4 reaching with tip of dactylus end of scaphocerite, propodus 3 x as long as dactylus, propodus nearly 2 x as long as carpus, propodus slightly longer than merus; P 5 reaching with tip of dactylus half-length of scaphocerite, propodus 3 x as long as dactylus, propodus nearly 2 x as long as carpus, propodus slightly longer than merus. Pleon. Smooth, somite 5 with posteroventral angle of pleuron acute; somite 6 nearly 2 x as long as somite 5. Inter-uropodal sclerite without keel-shaped pre-anal carinae. Pleopods. PL 2 with appendix masculina nearly 2 x as long as appendix interna. Uropods. Exopodite with mobile spines as long as spiniform projection of outer margin. Telson. Broad, smooth, slightly longer than abdominal somite 6, bearing two pairs of dorsal spinules close to posterior margin of telson. Posterior margin ending in moderately acute triangular point, with several plumose setae and two pairs of posterior spinules, inner pair overreaching end of telson.	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
F151D75D37E856578F7D52F400C2FFD9.taxon	description	Size. See in material examined.	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
F151D75D37E856578F7D52F400C2FFD9.taxon	distribution	Distribution. Only known from the type locality (Holthuis 1977; present paper).	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
AE8D4FB6FE9F54C88519F6F4B6A3D63C.taxon	diagnosis	Diagnosis (modified from Holthuis 1950, 1952 a). Body compressed, generally robust, sometimes slender. Rostrum well developed, compressed, toothed, size varying from shorter to longer than distal margin of scaphocerite. Carapace with anterolateral portion smooth or bearing numerous small spinules. Carapace armed with antennal spine; hepatic spines present in most species, branchiostegal groove present, distinct. Mandible with 3 - segmented palp. All maxillipeds with well-developed exopods. Pleurobranchs on third maxilliped and all pereiopods. P 1 slender. P 2 more robust than other pereiopods, usually longer than entire body in adult males, left and right legs often equal in size and shape or markedly different in several species. P 3 - P 5 with dactylus simple. P 5 with propodus bearing numerous transverse rows of setae in distal part of posterior margin. PL 1 with endopod much smaller than exopod, endopod of male without appendix interna. Pleon with pleurae smooth in most species or with small spinules. Uropods overreaching telson; exopod with distolateral spine, endopod unarmed. Telson elongate, subtriangular, narrowing posteriorly, with two pairs of dorsal spines, anterior pair placed in middle, posterior pair usually placed midway between anterior pair and posterior margin; posterior margin ending in sharp median point, flanked by two pairs of spines, outer pair usually shorter than inner one, inner pair overreaching apex of telson in most species. See detailed description in Holthuis (1950).	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
29ABADC237C658B283FC64B499510C4D.taxon	materials_examined	Material examined. Holotype: Mexico - Chiapas • male, cl 25 mm; Las Margaritas, Cueva Chamburro; 01 Mar. 2001; V. Sbordoni leg.; CNCR 25106. Paratypes: 1 ovigerous female, cl 22.5 mm, allotype; same data as for holotype; CNCR 25107 • 1 female, cl 12.3 mm; same data as for holotype; CNCR 25108.	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
29ABADC237C658B283FC64B499510C4D.taxon	description	Description. Rostrum. Short, straight, tip not reaching distal border of scaphocerite, almost reaching third article of antennular peduncle; upper margin bearing eight teeth, lower margin smooth. Cephalon. Eyes reduced, globular cornea with facets, pigmented area reduced to a black point. Scaphocerite 2.4 x as long as wide. Carapace. Smooth, maximum length 25 mm, with only antennal spine; branchiostegal groove shallow; hepatic spine absent. Pereiopods. P 1: slender, smooth, with few tufts of setae on both fingers; palm surpassing distal margin of scaphocerite; palm slightly compressed, as long as dactylus; carpus 1.75 x as long as palm, 1.12 x as long as merus. P 2: subequal in size, subequal in size, reaching with half of carpus beyond scaphocerite, without spines; ischium 0.9 x merus; carpus 0.8 x as long as merus, 0.85 x as long as palm; propodus 1.5 x as long as dactylus, 2.5 x as long as carpus; palm semi-cylindrical, 3.3 x as long as high, with dispersed tufts of setae, 0.8 x as long as dactylus; fingers elongated, not gaping, cutting edges covered with tufts of setae, dactylus without teeth. P 3: propodus, dactylus with several short setae, row of seven spines on ventral margin, propodus 3 x as long as dactylus, 2.05 x as long as carpus. P 4: sparsely pilose, propodus 3.4 x as long as dactylus, 1.8 x as long as carpus, propodus with row of nine movable spines on ventral margin, propodus-dactylus articulation with pair of setae. P 5: longest, propodus, carpus pilose, with longitudinal row of 12 movable spines, distal four close together, propodus-dactylus articulation with one spine; propodus 4 x as long as dactylus, 2.1 x as long as carpus. Pleon. Smooth; somites 1 - 3 with pleura broadly rounded; somites 4 and 5 with posteroventral margin of pleura rounded; pleura of all somites bearing setae on ventral border; somite 6 nearly 1.5 x as long as somite 5. Inter-uropodal sclerite without keel-shaped pre-anal carinae. Telson. Nearly 1.5 x longer than abdominal somite 6, shorter than uropodal rami, bearing two pairs of dorsal spines, first pair on distal fifth, second pair on middle section, with a single spine in the middle on left side; posterior margin broadly triangular bearing two pairs of lateral spines, inner pair 5 x longer than outer one, with plumose setae between inner spines, center ending in acute tip.	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
29ABADC237C658B283FC64B499510C4D.taxon	etymology	Etymology. Baldari et al. (2010) named this species in honor of Prof. Valerio Sbordoni, a studious of the cave fauna of Chiapas, Mexico, and collector of the specimens. We maintained this homage by forming the specific epithet with parts of his first and last name.	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
29ABADC237C658B283FC64B499510C4D.taxon	description	Size. See in material examined.	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
29ABADC237C658B283FC64B499510C4D.taxon	distribution	Distribution. Only known from the type locality (Baldari et al. 2010).	en	Mantelatto, Fernando L., Pileggi, Leonardo G., Pantaleao, Joao A. F., Magalhaes, Celio, Villalobos, Jose Luis, Alvarez, Fernando (2021): Multigene phylogeny and taxonomic revision of American shrimps of the genus Cryphiops Dana, 1852 (Decapoda, Palaemonidae) implies a proposal for reversal of precedence with Macrobrachium Spence Bate, 1868. ZooKeys 1047: 155-198, DOI: http://dx.doi.org/10.3897/zookeys.1047.66933, URL: http://dx.doi.org/10.3897/zookeys.1047.66933
